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Page last updated
Thu 02 Feb 2023

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

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Bivalvia | Unionoida | Unionoidea | Unionidae | Parreysiinae | Lamellidentini

genus Trapezidens Bolotov, Vikhrev & Konopleva in Bolotov et al., 2017: cladomics
     6 valid species

Cladomics Summary

number of species = 6

analyzed in 27 cladograms in 17 publications
maximum number of terminal species in an analysis = 6, mean = 2.81

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 18
maximum number of included species = 6, mean = 3.72
proportion of cladograms that support basic monophyly = 1.0


Bolotov et al., 2022, fig. S1

iT tT bi riS oT roT roS iD
6 6 4 6 55 10 1 0.0

*** Top Monophyly Cladogram *** *** Top Sister-Group Cladogram ***

     +-------------- Trapezidens_mogaungensis **   
  +--+                                          
  |  |  +----------- Trapezidens_scutum **         
  |  +--+                                       
  |     |  +-------- Trapezidens_exolescens **     
  |     +--+                                    
  |        |  +----- Trapezidens_yeti **           
  |        +--+                                 
  |           |  +-- Trapezidens_dolichorhynchus **
--+           +--+                              
  |              +-- Trapezidens_angustior **      
  |                                             
  |        +-------- Lamellidens_savadiensis    
  |  +-----+                                    
  |  |     |  +----- Lamellidens_chindwinensis  
  |  |     +--+                                 
  |  |        |  +-- Lamellidens_corrianus      
  |  |        +--+                              
  +--+           +-- Lamellidens_nongyangensis  
     |                                          
     |           +-- Lamellidens_marginalis     
     |  +--------+                              
     |  |        +-- Lamellidens_ferrugineus    
     +--+                                       
        |  +-------- Lamellidens_generosus      
        +--+                                    
           |  +----- Lamellidens_mainwaringi    
           +--+                                 
              |  +-- Lamellidens_jenkinsianus   
              +--+                              
                 +-- Lamellidens_lamellatus     

Bolotov et al., 2022, fig. 2

iT tT bi riS oT roT roS iD
6 6 3 6 55 10 1 3.0
        +----------- Trapezidens_mogaungensis **   
        |                                       
  +-----+  +-------- Trapezidens_exolescens **     
  |     |  |                                    
  |     +--+  +----- Trapezidens_scutum **         
  |        |  |                                 
  |        +--+  +-- Trapezidens_angustior **      
  |           |  |                              
--+           +--+-- Trapezidens_dolichorhynchus **
  |              |                              
  |              +-- Trapezidens_yeti **           
  |                                             
  |  +-------------- Lamellidens_marginalis     
  |  |                                          
  |  |  +----------- Lamellidens_ferrugineus    
  +--+  |                                       
     |  |  +-------- Lamellidens_generosus      
     |  +--+                                    
     |  |  |  +----- Lamellidens_mainwaringi    
     +--+  +--+                                 
        |     |  +-- Lamellidens_lamellatus     
        |     +--+                              
        |        +-- Lamellidens_jenkinsianus   
        |                                       
        |  +-------- Lamellidens_nongyangensis  
        +--+                                    
           |  +----- Lamellidens_savadiensis    
           +--+                                 
              |  +-- Lamellidens_corrianus      
              +--+                              
                 +-- Lamellidens_chindwinensis  

Konopleva et al., 2020, fig. 2

iT tT bi riS oT roT roS iD
5 5 3 5 7 4 1 1.0
        +-------- Lamellidens_savadiensis    
  +-----+                                    
  |     |  +----- Lamellidens_generosus      
  |     +--+                                 
  |        |  +-- Lamellidens_mainwaringi    
--+        +--+                              
  |           +-- Lamellidens_ferrugineus    
  |                                          
  |  +----------- Trapezidens_scutum **         
  +--+                                       
     |  +-------- Trapezidens_exolescens **     
     +--+                                    
        |  +----- Trapezidens_yeti **           
        +--+                                 
           |  +-- Trapezidens_angustior **      
           +--+                              
              +-- Trapezidens_dolichorhynchus **

Bolotov et al., 2022, fig. 2

iT tT bi riS oT roT roS iD
5 5 2 5 53 9 1 4.0
                    +-- Trapezidens_exolescens **     
              +-----+                              
              |     +-- Trapezidens_scutum **         
  +-----------+                                    
  |           |  +----- Trapezidens_angustior **      
  |           +--+                                 
  |              |  +-- Trapezidens_dolichorhynchus **
--+              +--+                              
  |                 +-- Trapezidens_yeti **           
  |                                                
  |  +----------------- Lamellidens_corrianus      
  |  |                                             
  +--+  +-------------- Lamellidens_nongyangensis  
     |  |                                          
     +--+  +----------- Lamellidens_savadiensis    
        |  |                                       
        |  |        +-- Lamellidens_ferrugineus    
        +--+  +-----+                              
           |  |     +-- Lamellidens_marginalis     
           |  |                                    
           +--+     +-- Lamellidens_lamellatus     
              |  +--+                              
              |  |  +-- Lamellidens_jenkinsianus   
              +--+                                 
                 |  +-- Lamellidens_generosus      
                 +--+                              
                    +-- Lamellidens_mainwaringi    

Bolotov et al., 2022, fig. S1

iT tT bi riS oT roT roS iD
5 5 2 5 53 9 1 4.0
           +-------- Trapezidens_exolescens **     
           |                                    
  +--------+  +----- Trapezidens_scutum **         
  |        |  |                                 
  |        +--+  +-- Trapezidens_angustior **      
  |           |  |                              
  |           +--+-- Trapezidens_dolichorhynchus **
  |              |                              
--+              +-- Trapezidens_yeti **           
  |                                             
  |           +----- Lamellidens_savadiensis    
  |  +--------+                                 
  |  |        |  +-- Lamellidens_corrianus      
  |  |        +--+                              
  |  |           +-- Lamellidens_nongyangensis  
  +--+                                          
     |           +-- Lamellidens_ferrugineus    
     |  +--------+                              
     |  |        +-- Lamellidens_marginalis     
     +--+                                       
        |  +-------- Lamellidens_lamellatus     
        +--+                                    
           |  +----- Lamellidens_jenkinsianus   
           +--+                                 
              |  +-- Lamellidens_generosus      
              +--+                              
                 +-- Lamellidens_mainwaringi    

Bolotov et al., 2022, fig. S2

iT tT bi riS oT roT roS iD
5 5 3 5 53 9 1 1.0
        +----------- Trapezidens_scutum **         
  +-----+                                       
  |     |  +-------- Trapezidens_exolescens **     
  |     +--+                                    
  |        |  +----- Trapezidens_yeti **           
  |        +--+                                 
  |           |  +-- Trapezidens_dolichorhynchus **
  |           +--+                              
--+              +-- Trapezidens_angustior **      
  |                                             
  |           +----- Lamellidens_savadiensis    
  |  +--------+                                 
  |  |        |  +-- Lamellidens_corrianus      
  |  |        +--+                              
  |  |           +-- Lamellidens_nongyangensis  
  +--+                                          
     |           +-- Lamellidens_marginalis     
     |  +--------+                              
     |  |        +-- Lamellidens_ferrugineus    
     +--+                                       
        |  +-------- Lamellidens_lamellatus     
        +--+                                    
           |  +----- Lamellidens_jenkinsianus   
           +--+                                 
              |  +-- Lamellidens_mainwaringi    
              +--+                              
                 +-- Lamellidens_generosus      

Bolotov et al., 2022, fig. 8

iT tT bi riS oT roT roS iD
5 5 3 5 45 4 1 3.0
        +-------- Lamellidens_savadiensis    
  +-----+                                    
  |     |  +----- Lamellidens_ferrugineus    
  |     +--+                                 
  |        |  +-- Lamellidens_generosus      
--+        +--+                              
  |           +-- Lamellidens_mainwaringi    
  |                                          
  |  +----------- Trapezidens_exolescens **     
  +--+                                       
     |  +-------- Trapezidens_scutum **         
     +--+                                    
        |  +----- Trapezidens_yeti **           
        +--+                                 
           |  +-- Trapezidens_angustior **      
           +--+                              
              +-- Trapezidens_dolichorhynchus **

Bolotov et al., 2020, fig. 1

iT tT bi riS oT roT roS iD
4 4 2 4 117 4 1 4.0
     +-------- Trapezidens_exolescens **     
  +--+                                    
  |  |  +----- Trapezidens_scutum **         
  |  +--+                                 
  |     |  +-- Trapezidens_dolichorhynchus **
--+     +--+                              
  |        +-- Trapezidens_angustior **      
  |                                       
  |  +-------- Lamellidens_savadiensis    
  +--+                                    
     |  +----- Lamellidens_generosus      
     +--+                                 
        |  +-- Lamellidens_ferrugineus    
        +--+                              
           +-- Lamellidens_mainwaringi    

Bolotov et al., 2020, fig. S1

iT tT bi riS oT roT roS iD
4 4 2 4 111 4 1 4.0
     +-------- Trapezidens_exolescens **     
  +--+                                    
  |  |  +----- Trapezidens_scutum **         
  |  +--+                                 
  |     |  +-- Trapezidens_dolichorhynchus **
--+     +--+                              
  |        +-- Trapezidens_angustior **      
  |                                       
  |  +-------- Lamellidens_savadiensis    
  +--+                                    
     |  +----- Lamellidens_generosus      
     +--+                                 
        |  +-- Lamellidens_ferrugineus    
        +--+                              
           +-- Lamellidens_mainwaringi    

Bolotov et al., 2017, fig. S1

iT tT bi riS oT roT roS iD
3 3 1 3 122 9 1 3.0
           +----- Trapezidens_exolescens **     
  +--------+                                 
  |        |  +-- Trapezidens_dolichorhynchus **
  |        +--+                              
--+           +-- Trapezidens_angustior **      
  |                                          
  |  +----------- Lamellidens_lamellatus     
  |  |                                       
  +--+  +-------- Lamellidens_jenkinsianus   
     |  |                                    
     |  |  +----- Lamellidens_generosus      
     +--+  |                                 
        |  +----- Lamellidens_mainwaringi    
        |  |                                 
        +--+  +-- Lamellidens_ferrugineus    
           +--+                              
           |  +-- Lamellidens_marginalis     
           |                                 
           |  +-- Lamellidens_savadiensis    
           +--+                              
              +-- Lamellidens_corrianus[1]   

Huang et al., 2019, fig. 2

iT tT bi riS oT roT roS iD
3 3 1 3 196 9 1 3.0
                    +----- Trapezidens_exolescens **     
  +-----------------+                                 
  |                 |  +-- Trapezidens_dolichorhynchus **
  |                 +--+                              
--+                    +-- Trapezidens_angustior **      
  |                                                   
  |  +-------------------- Lamellidens_corrianus      
  |  |                                                
  +--+  +----------------- Lamellidens_savadiensis    
     |  |                                             
     +--+              +-- Lamellidens_lamellatus     
        |  +-----------+                              
        |  |           +-- Lamellidens_jenkinsianus   
        +--+                                          
           |  +----------- Lamellidens_generosus      
           +--+                                       
              |  +-------- Lamellidens_mainwaringi[2] 
              +--+                                    
                 |  +----- Lamellidens_mainwaringi[1] 
                 +--+                                 
                    |  +-- Lamellidens_ferrugineus    
                    +--+                              
                       +-- Lamellidens_marginalis     

Huang et al., 2019, fig. S3

iT tT bi riS oT roT roS iD
3 3 1 3 194 9 1 3.0
        +----- Trapezidens_exolescens **     
  +-----+                                 
  |     |  +-- Trapezidens_angustior **      
  |     +--+                              
  |        +-- Trapezidens_dolichorhynchus **
  |                                       
--+        +-- Lamellidens_corrianus      
  |  +-----+                              
  |  |     +-- Lamellidens_savadiensis    
  |  |                                    
  |  |  +----- Lamellidens_lamellatus     
  +--+  |                                 
     |  +----- Lamellidens_jenkinsianus   
     |  |                                 
     |  |  +-- Lamellidens_marginalis     
     +--+--+                              
        |  +-- Lamellidens_ferrugineus    
        |                                 
        |  +-- Lamellidens_mainwaringi[1] 
        |  |                              
        +--+-- Lamellidens_mainwaringi[2] 
           |                              
           +-- Lamellidens_generosus      

Huang et al., 2019, fig. S4

iT tT bi riS oT roT roS iD
3 3 1 3 196 9 1 3.0
           +----- Trapezidens_exolescens **     
  +--------+                                 
  |        |  +-- Trapezidens_angustior **      
  |        +--+                              
--+           +-- Trapezidens_dolichorhynchus **
  |                                          
  |  +----------- Lamellidens_corrianus      
  |  |                                       
  +--+  +-------- Lamellidens_savadiensis    
     |  |                                    
     |  |  +----- Lamellidens_marginalis     
     +--+  |                                 
        |  +----- Lamellidens_ferrugineus    
        |  |                                 
        +--+  +-- Lamellidens_mainwaringi[1] 
           |  |                              
           |  +-- Lamellidens_mainwaringi[2] 
           |  |                              
           +--+-- Lamellidens_lamellatus     
              |                              
              +-- Lamellidens_generosus      
              |                              
              +-- Lamellidens_jenkinsianus   

Bolotov et al., 2017, fig. S1

iT tT bi riS oT roT roS iD
2 2 0 2 136 6 1 --
              +-- Trapezidens_dolichorhynchus **
  +-----------+                              
  |           +-- Trapezidens_exolescens **     
--+                                          
  |  +----------- Lamellidens_corrianus      
  |  |                                       
  +--+  +-------- Lamellidens_savadiensis    
     |  |                                    
     +--+     +-- Lamellidens_jenkinsianus   
        |  +--+                              
        |  |  +-- Lamellidens_mainwaringi    
        +--+                                 
           |  +-- Lamellidens_ferrugineus    
           +--+                              
              +-- Lamellidens_marginalis     

Bolotov et al., 2017, fig. S2

iT tT bi riS oT roT roS iD
2 2 0 2 136 6 1 --
           +-- Trapezidens_dolichorhynchus **
  +--------+                              
  |        +-- Trapezidens_exolescens **     
  |                                       
--+  +-------- Lamellidens_savadiensis    
  |  |                                    
  |  +-------- Lamellidens_corrianus      
  +--+                                    
     |     +-- Lamellidens_marginalis     
     |  +--+                              
     |  |  +-- Lamellidens_ferrugineus    
     +--+                                 
        |  +-- Lamellidens_jenkinsianus   
        +--+                              
           +-- Lamellidens_mainwaringi    

Konopleva et al., 2017, fig. 5

iT tT bi riS oT roT roS iD
2 2 0 2 3 3 1 --
        +-- Trapezidens_exolescens **     
  +-----+                              
  |     +-- Trapezidens_dolichorhynchus **
--+                                    
  |  +----- Lamellidens_savadiensis    
  +--+                                 
     |  +-- Lamellidens_mainwaringi    
     +--+                              
        +-- Lamellidens_jenkinsianus   

Konopleva et al., 2017, fig. 6

iT tT bi riS oT roT roS iD
2 2 0 2 29 3 1 --
        +-- Trapezidens_exolescens **     
  +-----+                              
  |     +-- Trapezidens_dolichorhynchus **
--+                                    
  |  +----- Lamellidens_savadiensis    
  +--+                                 
     |  +-- Lamellidens_jenkinsianus   
     +--+                              
        +-- Lamellidens_mainwaringi    

Bolotov et al., 2018, fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 83 4 1 --
           +-- Trapezidens_exolescens **     
  +--------+                              
  |        +-- Trapezidens_dolichorhynchus **
--+                                       
  |  +-------- Lamellidens_savadiensis    
  +--+                                    
     |  +----- Lamellidens_ferrugineus    
     +--+                                 
        |  +-- Lamellidens_generosus      
        +--+                              
           +-- Lamellidens_mainwaringi    

Whelan et al., 2011, fig. 1 ML

iT tT bi riS oT roT roS iD
1 1 0 1 35 1 1 --
  +-- Lamellidens_savadiensis    
--+                              
  +-- Trapezidens_dolichorhynchus **

Whelan et al., 2011, fig. 1 MP

iT tT bi riS oT roT roS iD
1 1 0 1 35 1 1 --
  +-- Lamellidens_savadiensis    
--+                              
  +-- Trapezidens_dolichorhynchus **

Pfeiffer & Graf, 2015, fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 39 1 1 --
  +-- Trapezidens_dolichorhynchus **
--+                              
  +-- Lamellidens_savadiensis    

Lopes-Lima et al., 2017, fig. 1-2

iT tT bi riS oT roT roS iD
1 1 0 1 80 0 0 --
-- Trapezidens_dolichorhynchus

Pfeiffer et al., 2018, fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 28 1 1 --
  +-- Trapezidens_dolichorhynchus **
--+                              
  +-- Lamellidens_savadiensis    

Jeratthitikul et al., 2019, fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 22 1 1 --
  +-- Lamellidens_mainwaringi
--+                          
  +-- Trapezidens_exolescens ** 

Pfeiffer et al., 2019, fig. 4-5

iT tT bi riS oT roT roS iD
1 1 0 1 56 1 1 --
  +-- Trapezidens_dolichorhynchus **
--+                              
  +-- Lamellidens_savadiensis    

Pfeiffer et al., 2019, fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 56 1 1 --
  +-- Trapezidens_dolichorhynchus **
--+                              
  +-- Lamellidens_savadiensis    

Konopleva et al., 2023, fig. S1, 4

iT tT bi riS oT roT roS iD
1 1 0 1 55 1 1 --
  +-- Trapezidens_exolescens ** 
--+                          
  +-- Lamellidens_savadiensis

Page updated: 11:41:26 Thu 02 Feb 2023

 
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