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Page last updated
Mon 24 Mar 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Ambleminae | Lampsilini

genus Glebula Conrad, 1853: cladomics
     1 valid species

Cladomics Summary

number of species = 1

analyzed in 22 cladograms in 16 publications
maximum number of terminals in an analysis = 1, mean = 1.0


Zanatta & Murphy, 2006 [Publ 2672], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 55 47 22 --

*** Top Monophyly Cladogram *** *** Top Sister-Group Cladogram ***

                                   +-- Cyrtonaias             
  +--------------------------------+                          
  |                                +-- Glebula_rotundata **      
  |                                                           
--+  +-------------------------------- Paetulunio             
  |  |                                                        
  |  |                             +-- Obliquaria             
  +--+  +--------------------------+                          
     |  |                          +-- Toxolasma              
     |  |                                                     
     |  |  +-------------------------- Sagittunio             
     +--+  |                                                  
        |  |              +----------- Potamilus              
        |  |              |                                   
        |  |  +-----------+        +-- Medionidus_conradicus  
        |  |  |           |  +-----+                          
        +--+  |           |  |     +-- Ptychobranchus         
           |  |           +--+                                
           |  |              |  +----- Medionidus_acutissimus 
           |  |              |  |                             
           |  |              +--+----- Lemiox                 
           |  |                 |                             
           |  |                 |  +-- Dromus                 
           +--+                 +--+                          
              |                    +-- Cyprogenia             
              |                                               
              |                 +----- Lampsilis_teres        
              |  +--------------+                             
              |  |              |  +-- Truncilla              
              |  |              +--+                          
              |  |                 +-- Ellipsaria             
              |  |                                            
              +--+  +----------------- Leaunio                
                 |  |                                         
                 |  |     +----------- Hamiota                
                 |  |     |                                   
                 |  |  +--+        +-- Lampsilis_fasciola     
                 +--+  |  |  +-----+                          
                    |  |  |  |     +-- Ortmanniana_pectorosa  
                    |  |  +--+                                
                    |  |     |  +----- Lampsilis_ornata       
                    |  |     +--+                             
                    |  |        |  +-- Lampsilis_ovata        
                    +--+        +--+                          
                       |           +-- Lampsilis_cardium      
                       |                                      
                       |        +----- Venustaconcha          
                       |  +-----+                             
                       |  |     |  +-- Obovaria               
                       |  |     +--+                          
                       +--+        +-- Epioblasma             
                          |                                   
                          |  +-------- Lampsilis_siliquoidea  
                          +--+                                
                             |  +----- Ortmanniana_ligamentina
                             +--+                             
                                |  +-- Cambarunio             
                                +--+                          
                                   +-- Ligumia                

Inoue et al., 2018 [Publ 3858], fig. 1_3a-c

iT tT bi riS oT roT roS iD
1 1 0 1 103 10 7 --
-- Glebula_rotundata

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 33 5 5 --
  +-- Glebula_rotundata **
--+                    
  +-- Cyrtonaias       

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

iT tT bi riS oT roT roS iD
1 1 0 1 30 5 5 --
           +-- Cyrtonaias       
  +--------+                    
  |        +-- Glebula_rotundata **
--+                             
  |  +-------- Toxolasma        
  +--+                          
     |  +----- Ortmanniana      
     +--+                       
        |  +-- Ligumia          
        +--+                    
           +-- Potamilus        

Chapman et al., 2008 [Publ 2667], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 20 12 10 --
  +-------------------------- Toxolasma          
  |                                              
--+  +----------------------- Obliquaria         
  |  |                                           
  |  |                    +-- Cyrtonaias         
  +--+  +-----------------+                      
     |  |                 +-- Glebula_rotundata **  
     |  |                                        
     +--+                 +-- Ptychobranchus     
        |  +--------------+                      
        |  |              +-- Lemiox             
        |  |                                     
        +--+              +-- Venustaconcha      
           |  +-----------+                      
           |  |           +-- Obovaria           
           +--+                                  
              |  +----------- Hamiota            
              +--+                               
                 |  +-------- Lampsilis_ovata    
                 +--+                            
                    |  +----- Ortmanniana        
                    +--+                         
                       |  +-- Lampsilis_straminea
                       +--+                      
                          +-- Lampsilis_hydiana  

Smith et al., 2019 [Publ 3951], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 40 35 19 --
  +-- Glebula_rotundata **
--+                    
  +-- Cyrtonaias       

Hoeh et al., 2001 [Publ 2652], fig. 14f2

iT tT bi riS oT roT roS iD
1 1 0 1 30 5 5 --
  +----------- Ortmanniana      
--+                             
  |  +-------- Glebula_rotundata **
  +--+                          
     |  +----- Potamilus        
     +--+                       
        |  +-- Ligumia          
        +--+                    
           +-- Toxolasma        

Zanatta & Murphy, 2006 [Publ 2672], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 56 47 22 --
                                      +-- Paetulunio             
  +-----------------------------------+                          
  |                                   +-- Toxolasma              
  |                                                              
  |                                +----- Obliquaria             
  |  +-----------------------------+                             
--+  |                             |  +-- Glebula_rotundata **      
  |  |                             +--+                          
  |  |                                +-- Cyrtonaias             
  |  |                                                           
  |  |                             +----- Ptychobranchus         
  |  |                          +--+                             
  +--+                          |  |  +-- Dromus                 
     |                          |  +--+                          
     |  +-----------------------+     +-- Cyprogenia             
     |  |                       |                                
     |  |                       |  +----- Medionidus_conradicus  
     |  |                       +--+                             
     |  |                          |  +-- Medionidus_acutissimus 
     |  |                          +--+                          
     +--+                             +-- Lemiox                 
        |                                                        
        |  +----------------------------- Sagittunio             
        |  |                                                     
        |  +----------------------------- Potamilus              
        |  |                                                     
        |  |                          +-- Truncilla              
        +--+  +-----------------------+                          
           |  |                       +-- Ellipsaria             
           |  |                                                  
           |  |                       +-- Ortmanniana_pectorosa  
           |  |                 +-----+                          
           |  |                 |     +-- Lampsilis_fasciola     
           +--+  +--------------+                                
              |  |              |  +----- Lampsilis_ornata       
              |  |              +--+                             
              |  |                 |  +-- Lampsilis_ovata        
              |  |                 +--+                          
              |  |                    +-- Lampsilis_cardium      
              +--+                                               
                 |                    +-- Ligumia                
                 |                 +--+                          
                 |                 |  +-- Cambarunio             
                 |  +--------------+                             
                 |  |              |  +-- Lampsilis_teres        
                 |  |              +--+                          
                 +--+                 +-- Hamiota                
                    |                                            
                    |  +----------------- Leaunio                
                    |  |                                         
                    +--+  +-------------- Ortmanniana_ligamentina
                       |  |                                      
                       +--+  +----------- Lampsilis_siliquoidea  
                          |  |                                   
                          +--+  +-------- Obovaria_olivaria      
                             |  |                                
                             |  +-------- Venustaconcha          
                             +--+                                
                                +-------- Epioblasma             
                                |                                
                                |  +----- Obovaria_unicolor      
                                +--+                             
                                   |  +-- Obovaria_subrotunda    
                                   +--+                          
                                      +-- Obovaria_arkansasensis 

Curole & Kocher, 2005 [Publ 2682], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 10 4 3 --
-- Glebula_rotundata

Pfeiffer et al., 2016 [Publ 3660], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 37 5 5 --
        +-- Cyrtonaias       
  +-----+                    
  |     +-- Glebula_rotundata **
  |                          
--+     +-- Lampsilis        
  |  +--+                    
  |  |  +-- Leaunio          
  +--+                       
     |  +-- Truncilla        
     +--+                    
        +-- Potamilus        

Pfeiffer et al., 2019 [Publ 3866], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 34 11 10 --
  +-- Cyrtonaias       
--+                    
  +-- Glebula_rotundata **

Burlakova et al., 2019 [Publ 3899], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 32 25 17 --
  +-- Cyrtonaias       
--+                    
  +-- Glebula_rotundata **

Campbell et al., 2005 [Publ 2660], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 101 42 22 --
                       +-------- Toxolasma                     
  +--------------------+                                       
  |                    |  +----- Obliquaria                    
  |                    +--+                                    
  |                       |  +-- Cyrtonaias                    
  |                       +--+                                 
  |                          +-- Glebula_rotundata **             
  |                                                            
  |              +-------------- Potamilus                     
  |              |                                             
  |              |     +-------- Medionidus_acutissimus        
--+              |  +--+                                       
  |  +-----------+  |  |  +----- Lemiox                        
  |  |           |  |  +--+                                    
  |  |           |  |     |  +-- Dromus                        
  |  |           |  |     +--+                                 
  |  |           +--+        +-- Cyprogenia                    
  |  |              |                                          
  |  |              |        +-- Sagittunio                    
  |  |              |     +--+                                 
  |  |              |     |  +-- Medionidus_conradicus         
  +--+              +-----+                                    
     |                    |  +-- Ptychobranchus                
     |                    +--+                                 
     |                       +-- Venustaconcha_ellipsiformis__2
     |                                                         
     |                       +-- Ellipsaria                    
     |  +--------------------+                                 
     |  |                    +-- Truncilla                     
     |  |                                                      
     |  |  +-------------------- Villosa                       
     +--+  |                                                   
        |  |                 +-- Obovaria_olivaria             
        |  |        +--------+                                 
        |  |        |        +-- Venustaconcha_ellipsiformis__1
        |  |  +-----+                                          
        +--+  |     |  +-------- Epioblasma                    
           |  |     +--+                                       
           |  |        |  +----- Obovaria_unicolor             
           |  |        +--+                                    
           |  |           |  +-- Obovaria_arkansasensis        
           |  |           +--+                                 
           |  |              +-- Obovaria_subrotunda           
           +--+                                                
              |        +-------- Hamiota                       
              |        |                                       
              |  +-----+     +-- Lampsilis_cardium             
              |  |     |  +--+                                 
              |  |     |  |  +-- Lampsilis_ovata               
              |  |     +--+                                    
              |  |        |  +-- Ortmanniana_pectorosa         
              +--+        +--+                                 
                 |           +-- Lampsilis_ornata              
                 |                                             
                 |           +-- Ligumia                       
                 |  +--------+                                 
                 |  |        +-- Cambarunio                    
                 +--+                                          
                    |  +-------- Lampsilis_teres               
                    +--+                                       
                       |  +----- Ortmanniana_ligamentina       
                       +--+                                    
                          |  +-- Lampsilis_siliquoidea         
                          +--+                                 
                             +-- Leaunio                       

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

iT tT bi riS oT roT roS iD
1 1 0 1 30 5 5 --
  +----------- Ortmanniana      
--+                             
  |  +-------- Glebula_rotundata **
  +--+                          
     |  +----- Potamilus        
     +--+                       
        |  +-- Ligumia          
        +--+                    
           +-- Toxolasma        

Campbell & Lydeard, 2012 [Publ 3309], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 72 14 10 --
  +-- Glebula_rotundata **
--+                    
  +-- Paetulunio       

Curole & Kocher, 2005 [Publ 2682], fig. 1-M

iT tT bi riS oT roT roS iD
1 1 0 1 11 4 3 --
     +-- Potamilus        
  +--+                    
  |  +-- Glebula_rotundata **
--+                       
  |  +-- Leaunio          
  +--+                    
     +-- Lampsilis        

Chapman et al., 2008 [Publ 2744], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 20 12 10 --
  +-------------------------- Toxolasma          
  |                                              
--+  +----------------------- Obliquaria         
  |  |                                           
  |  |                    +-- Cyrtonaias         
  +--+  +-----------------+                      
     |  |                 +-- Glebula_rotundata **  
     |  |                                        
     +--+                 +-- Ptychobranchus     
        |  +--------------+                      
        |  |              +-- Lemiox             
        |  |                                     
        +--+              +-- Venustaconcha      
           |  +-----------+                      
           |  |           +-- Obovaria           
           +--+                                  
              |  +----------- Hamiota            
              +--+                               
                 |  +-------- Lampsilis_ovata    
                 +--+                            
                    |  +----- Ortmanniana        
                    +--+                         
                       |  +-- Lampsilis_straminea
                       +--+                      
                          +-- Lampsilis_hydiana  

Campbell & Lydeard, 2012 [Publ 3309], fig. 1-2

iT tT bi riS oT roT roS iD
1 1 0 1 72 14 10 --
  +-------------- Potamilus_alatus      
  |                                     
  +-------------- Potamilus_capax       
  |                                     
  +-------------- Glebula_rotundata **     
  |                                     
  |           +-- Toxolasma             
  +-----------+                         
--+           +-- Paetulunio            
  |                                     
  |           +-- Dromus                
  +-----------+                         
  |           +-- Lemiox                
  |                                     
  |           +-- Lampsilis_ovata       
  |  +--------+                         
  |  |        +-- Obliquaria            
  +--+                                  
     |  +-------- Lampsilis_virescens   
     |  |                               
     +--+     +-- Epioblasma            
        |  +--+                         
        |  |  +-- Obovaria_arkansasensis
        +--+                            
           |  +-- Obovaria_olivaria     
           +--+                         
              +-- Venustaconcha         

Inoue et al., 2020 [Publ 4128], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 25 11 10 --
  +-- Glebula_rotundata **
--+                    
  +-- Cyrtonaias       

Hoeh et al., 2001 [Publ 2652], fig. 14f4

iT tT bi riS oT roT roS iD
1 1 0 1 30 5 5 --
           +-- Cyrtonaias       
  +--------+                    
  |        +-- Glebula_rotundata **
--+                             
  |  +-------- Toxolasma        
  +--+                          
     |  +----- Potamilus        
     +--+                       
        |  +-- Ligumia          
        +--+                    
           +-- Ortmanniana      

Campbell et al., 2005 [Publ 2660], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 101 42 22 --
           +----- Obliquaria                    
  +--------+                                    
  |        |  +-- Cyrtonaias                    
  |        +--+                                 
  |           +-- Glebula_rotundata **             
  |                                             
  |  +----------- Sagittunio                    
  |  |                                          
  |  +----------- Leaunio                       
  |  |                                          
  |  +----------- Potamilus                     
  |  |                                          
  |  +----------- Hamiota                       
  |  |                                          
  |  |        +-- Ellipsaria                    
--+  +--------+                                 
  |  |        +-- Truncilla                     
  |  |                                          
  |  |     +----- Lampsilis_teres               
  |  +-----+                                    
  |  |     |  +-- Ortmanniana_ligamentina       
  |  |     +--+                                 
  |  |        +-- Lampsilis_siliquoidea         
  |  |                                          
  |  |     +----- Villosa                       
  |  +-----+                                    
  |  |     |  +-- Ligumia                       
  |  |     +--+                                 
  |  |        +-- Cambarunio                    
  |  |                                          
  +--+  +-------- Ortmanniana_pectorosa         
     +--+                                       
     |  |  +----- Lampsilis_ornata              
     |  +--+                                    
     |     |  +-- Lampsilis_cardium             
     |     +--+                                 
     |        +-- Lampsilis_ovata               
     |                                          
     |  +-------- Obovaria_olivaria             
     |  |                                       
     |  +-------- Venustaconcha_ellipsiformis__2
     +--+                                       
     |  +-------- Epioblasma                    
     |  |                                       
     |  |  +----- Obovaria_unicolor             
     |  +--+                                    
     |     |  +-- Obovaria_arkansasensis        
     |     +--+                                 
     |        +-- Obovaria_subrotunda           
     |                                          
     |        +-- Ptychobranchus                
     |  +-----+                                 
     |  |     +-- Venustaconcha_ellipsiformis__1
     |  |                                       
     +--+  +----- Medionidus_conradicus         
        |  |                                    
        |  |  +-- Lemiox                        
        +--+--+                                 
           |  +-- Medionidus_acutissimus        
           |                                    
           |  +-- Dromus                        
           +--+                                 
              +-- Cyprogenia                    

Hoeh et al., 2002 [Publ 2653], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 30 5 5 --
  +-- Glebula_rotundata **
--+                    
  +-- Cyrtonaias       

Page updated: 17:20:11 Mon 24 Mar 2025

 
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