Unio crassus Retzius, 1788
Schröter, 1779, Gesch. Flußconch.: 182, pl. 2, fig. 2 [Mya testa crassa]. [non-binomial]
Philipsson in Retzius, 1788, Diss. Nat.-Hist. Test. Gen.: 17.

Unio crassus — Retzius (1788), Spengler (1793), Nilsson (1822), Menke (1828), ..., Lopes-Lima et al. (2024), Wu et al. (2024), Zieritz et al. (2024), Dai et al. (2024)
Margarita (Unio) litoralis — Lea (1836), Lea (1838)
Mysca crassa — Schlüter (1838)
Margaron (Unio) crassus — Lea (1852), Lea (1870)
Unio (Potamida) crassus — Mörch (1853)
Unio (Lymnium) crassus — Moquin-Tandon (1855), Drouët (1857)
Unio (Bariosta) crassus — Adams & Adams (1857)
Unio crassus s.s. — Westerlund (1867)
Unio batavus var. crassus — Clessin (1877)
Unio (Crassiana) crassus — Servain (1882), Servain (1888), Locard (1889), Vinarski & Kantor (2016)
Unio (Groupe de l’U. crassus) crassus — Locard (1882)
Unio (sec. Hyridella) crassus — Paetel (1890)
Unio (Groupe de l’U. feliciani) crassus — Locard (1893)
Margaritana crassa — Simpson (1900), Simpson (1914)
Unio (s.s.) crassus — Zhadin (1938)
Unio crassus crassus — Haas (1940), Haas (1969), Fauna Europaea (2012), Prié & Puillandre (2014), Klishko et al. (2017), Lopes-Lima et al. (2017), Bogatov (2022)
Crassiana crassa crassa — Akromovski (1976), Kantor & Sysoev (2005), Kantor et al. (2010)
Batavusiana (Crassunio) crassa — Zatravkin (1983), Stadnichenko (1984), Zatravkin (1987)
Unio (Crassunio) crassus — Falkner (1994)
Batavusiana (Crassiana) crassus — Korniushin (1998)
Unio (Crassiana) crassus crassus — Falkner et al. (2001), Falkner et al. (2002)
Crassiana (s.s.) crassa — Starobogatov et al. (2004)
Unio c. crassus — Príe et al. (2021)

Unio musivus Spengler, 1793
Spengler, 1793, Skr. Naturhist. Selsk. 3: 67, no. 32.

Unio musivus — Spengler (1793), Simpson (1900)
Unio crassus musivus — Ortmann (1912), Bogatov (2022)
Unio (sec. Lymnium) batavus — Simpson (1914)
Unio crassus crassus — Haas (1940), Haas (1969), Fauna Europaea (2012), Klishko et al. (2017)
Batavusiana (s.s.) musiva — Zatravkin (1983), Zatravkin (1987)
Batavusiana (s.s.) musivus [??] — Korniushin (1998)
Crassiana (Bataviana) musiva — Starobogatov et al. (2004)
Crassiana musiva musiva — Kantor & Sysoev (2005), Kantor et al. (2010)
Unio crassus — Graf (2007), Anisimova et al. (2018), Bogatov et al. (2018), Lopes-Lima et al. (2024)
Unio (Crassiana) musivus — Vinarski & Kantor (2016)
Unio crassus cf. courtillieri — Klishko et al. (2017)

Unio courtillieri Hattemann, 1859
Hattemann, 1859, Ann. Soc. Linn. Maine-et-Loire 3: 232.

Unio courtillieri — Hattemann (1859)
Unio batavus var. courtilieri — de Joannis (1859)
Unio (Fusculusiana) fusculus — Locard (1889), Germain (1904)
Unio crassus var. piscinalis — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969), Lopes-Lima et al. (2017)
Unio (Crassiana) crassus courtillieri — Falkner et al. (2001), Falkner et al. (2002)
Unio crassus courtillieri — Prié & Puillandre (2014), Lopes-Lima et al. (2017)
Unio crassus cf. courtillieri — Klishko et al. (2017)
Unio crassus courtillierii — Araujo et al. (2018)
Unio crassus — Anisimova et al. (2018), Bogatov et al. (2018), Lopes-Lima et al. (2024)
Unio c. courtillieri — Príe et al. (2021)
Unio cf. courtillieri — Lopes-Lima et al. (2021)

Mya batava Maton & Rackett, 1807
Schröter, 1779, Gesch. Flußconch.: pl. 3, figs. 2, 4-5.
Martini & Chemnitz, 1782, Conchyl. Cab. 6: 19, pl. 1, fig. 6.
Maton & Rackett, 1807, Trans. Linn. Soc. London 8: 37.

Mya batava — Maton & Rackett (1807), Dillwyn (1817), Wood (1825)
Unio batava — Lamarck (1819), Pfeiffer (1821), Bory de St. Vincent (1827), Kickx (1830), ..., Krynicki (1837), Morelet (1853), Sowerby (1866), Malzine (1867)
Unio batavus — Nilsson (1822), Wood (1828), Menke (1828), Menke (1830), ..., Westerlund (1890), Kobelt & Haas (1910), Haas (1924), Germain (1931)
Unio pictorum — Lea (1829), Leach (1852)
Margarita (Unio) batava — Lea (1836), Lea (1838)
Mysca batava — Schlüter (1838)
Mysca (s.s.) batava — Swainson (1840)
Margaron (Unio) batavus — Lea (1852), Lea (1870)
Unio (Batavi) batavus — Dupuy (1852)
Unio (Lymnium) batavus — Moquin-Tandon (1855), Drouët (1857)
Unio (Hyridella) batavus — Adams & Adams (1857)
Unio crassus var. batavus — Westerlund (1867)
Unio (Groupe de l’U. Batavus) batavus — Locard (1882)
Unio (Bataviana) batavus — Servain (1888)
Unio (Batavusiana) batavus — Locard (1889), Germain (1904)
Unio (sec. Hyridella) batavus — Paetel (1890)
Unio (Groupe de l’U. batavus) batavus — Locard (1893)
Unio (sec. Lymnium) batavus — Simpson (1900), Simpson (1914)
Unio (Groupe des Batavusiana) batavus — Caziot (1907)
Unio crassus musivus — Ortmann (1912)
Unio crassus batavus — Haas (1940), Haas (1969), Fauna Europaea (2012), Lopes-Lima et al. (2017)
Batavusiana (s.s.) musiva — Zatravkin (1983), Zatravkin (1987)
Unio (Crassiana) crassus — Vinarski & Kantor (2016)
Unio crassus — Araujo et al. (2018)

Unio cyprinorum Locard, 1882
Bourguignat in Locard, 1882, Prodr. Malac. Franç. 1: 289, 358.
‘Berthier’ Locard, 1889, Contr. Faune Malacol. Franc. 13: 142 [36].
‘Berthier’ Locard, 1893, Coqu. Eau Douce Saumâtre France: 168.
Type(s): IDIOTYPE MNHN_22185, l’Ornain (Meuse).

Unio cyprinorum — Locard (1882)
Unio (Bataviana) cyprinorum — Servain (1882), Servain (1888)
Unio (Groupe de l’U. Batavus) cyprinorum — Locard (1882)
Unio (Batavusiana) cyprinorum — Locard (1889)
Unio batavus var. cyprinorum — Westerlund (1890)
Unio (Groupe de l’U. batavus) cyprinorum — Locard (1893)
Unio batavus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Crassiana (Bataviana) cyprinorum — Starobogatov et al. (2004)
Crassiana cyprinorum — Kantor & Sysoev (2005), Kantor et al. (2010)
Unio crassus — Graf (2007), Lopes-Lima et al. (2024)
Unio (Crassiana) cyprinorum — Vinarski & Kantor (2016)
Unio crassus musivus — Bogatov (2022)

Unio ater var. irgizlaicus Lindholm, 1904
Lindholm, 1904, Ann. Mus. St. Pétersbourg 8(3-4): 343.
(intro'd as intra-sp. of Unio ater Nilsson, 1822)

Unio ater var. irgizlaicus — Lindholm (1904)
Unio crassus crassus — Haas (1969), Klishko et al. (2017)
Batavusiana (s.s.) irgislaica — Zatravkin (1987)
Crassiana (Bataviana) irgizlaica — Starobogatov et al. (2004)
Crassiana irgizlaica — Kantor & Sysoev (2005), Kantor et al. (2010)
Unio crassus — Graf (2007), Lopes-Lima et al. (2024)
Unio (Crassiana) irgizlaicus — Vinarski & Kantor (2016)
Unio crassus musivus — Bogatov (2022)

Unio kungurensis irenjensis Kobelt, 1912
Kobelt, 1912, Icon. Land- und Süssw.-Moll. 18: 26, pl. 491, fig. 2626.
(intro'd as intra-sp. of Unio kungurensis Kobelt, 1912)

Unio kungurensis irenjensis — Kobelt (1912)
Unio (s.s.) crassus var. chlebnikovi — Zhadin (1938)
Unio crassus gontieri — Haas (1940), Haas (1969), Graf (2007)
Batavusiana (s.s.) irenjensis — Zatravkin (1983), Stadnichenko (1984), Zatravkin (1987)
Crassiana irenjensis — Kantor & Sysoev (2005), Kantor et al. (2010)
Unio (Crassiana) irenjensis — Vinarski & Kantor (2016)
Unio crassus — Lopes-Lima et al. (2024)

Unio littoralis var. minor Rossmässler, 1842
Rossmässler, 1842, Icon. Land- und Süssw.-Moll. 2 (5): 14, pl. 55, figs. 744. [in part]
(misidentified reference to Unio littoralis var. minor Rossmässler, 1842)

Unio littoralis var. minor — Rossmässler (1842)
Unio (sec. Lymnium) batavus — Simpson (1900), Simpson (1914)
Unio (Tumidusiana) tumidus var. minor — Stadnichenko (1984)

Mya ovalis Lightfoot, 1786
Lister, 1685, Conch.: pl. 146, fig. 1. [pre-linnaean]
‘S.’ Lightfoot, 1786, Catalogue of the Portland Museum: 134, no. 2983.

Mya ovalis — Lightfoot (1786)
Unio ovalis — Retzius (1788)
Mya batava — Dillwyn (1817)
Margarita (Unio) batava — Lea (1836), Lea (1838)
Margaron (Unio) batavus — Lea (1852), Lea (1870)
Unio (Mysca) limosus — Mörch (1863)

Mya pictorum Gmelin, 1791
Lister, 1678, Hist. Anim. Anglic.: pl. 2, fig. 30; [app.] pl. 1, fig. 4. [pre-linnaean]
Lister, 1685, Conch.: pl. 146, fig. 1; pl. 147, figs. 2-3. [pre-linnaean]
Schröter, 1779, Gesch. Flußconch.: pl. 2, fig. 2; pl. 3, figs. 2-5; pl. 4, fig. 6.
Martini & Chemnitz, 1782, Conchyl. Cab. 6: 19, pl. 1, fig. 6.
Gmelin, 1791, Syst. Nat. 1(6): 3218, sp. 3.
(misidentified reference to Mya pictorum Linnaeus, 1758)

Mya pictorum — Gmelin (1791), Schreibers (1793)
Unio pictorum — Bosc (1801), Deshayes (1832), Küster (1853), Malzine (1867)
Margarita (Unio) batava — Lea (1836), Lea (1838)
Margaron (Unio) batavus — Lea (1852), Lea (1870)
Unio (Lymnium) batavus — Moquin-Tandon (1855), Drouët (1857)
Unio (sec. Lymnium) batavus — Simpson (1900), Simpson (1914)

Unio margaritifera Draparnaud, 1806
‘Linn.’ Draparnaud, 1806, Hist. Moll. France: pl. 11, fig. 5.
(misidentified reference to Mya margaritifera Linnaeus, 1758)

Unio margaritifera — Draparnaud (1806)
Unio ater — Nilsson (1822)
Unio sinuatus — Gassies (1849)
Unio margaritifer — Küster (1855)
Unio (Groupe de l’U. Philippei) philippei — Locard (1882)
Unio (Ateriana) brevierei — Locard (1889)
Unio consentaneus — Germain (1931)

Unio pictorum Draparnaud, 1806
‘Linn.’ Draparnaud, 1806, Hist. Moll. France: 131, pl. 11, figs. 3 [var. B]. [in part]
(misidentified reference to Mya pictorum Linnaeus, 1758)

Unio pictorum — Draparnaud (1806), Deshayes (1832)
Unio batava — Pfeiffer (1821), Held (1836)
Unio batavus — Rossmässler (1835), Clessin (1877), Germain (1931)
Margarita (Unio) batava — Lea (1836), Lea (1838)
Unio requienii — Gassies (1849)
Margaron (Unio) batavus — Lea (1852), Lea (1870)
Unio (Batavi) batavus — Dupuy (1852)
Unio (Lymnium) batavus — Moquin-Tandon (1855), Drouët (1857)
Unio (Batavusiana) seneauxi — Locard (1889), Germain (1904)
Unio (sec. Lymnium) batavus — Simpson (1900), Simpson (1914)

Unio lemovicincae Férussac, 1807
‘Linn.’ Draparnaud, 1806, Hist. Moll. France: pl. 11, fig. 5.
Férussac, 1807, Ess. méth.: 114 [as ‘lemovicincæ’].

Unio lemovicincae — Férussac (1807), Menke (1828), Menke (1830), Küster (1861), Westerlund (1890)
Unio margaritifera — Pfeiffer (1821), Kickx (1830)
Unio (Margaritana) margaritifer — Drouët (1857)
Margaron (Unio) pictorum — Lea (1870)

Unio litoralis Pfeiffer, 1821
Pfeiffer, 1821, Natur. Deutsch. Land Sussw. Moll. 1: 117, pl. 5, fig. 12.
Rossmässler, 1853, Z. Malak. 10: 92.
Mörch, 1863, Vidensk. Medd. Naturhist. Foren. Kjöben. (17-22): .
(misidentified reference to Unio littoralis Cuvier, 1798; error for Unio littoralis Cuvier, 1798)

Unio litoralis — Pfeiffer (1821)
Unio crassus — Nilsson (1822), Menke (1828), Menke (1830), Rossmässler (1835), Küster (1854)
Margarita (Unio) litoralis — Lea (1836), Lea (1838)
Margaron (Unio) litoralis — Lea (1852), Lea (1870)
Unio (Lymnium) crassus — Moquin-Tandon (1855), Drouët (1857)
Unio batavus var. crassus — Clessin (1877)
Unio batavus var. pseudo-litoralis — Clessin (1877)
Unio crassus var. maximus — Kobelt (1884)
Unio pseudolitoralis — Westerlund (1890)
Margaritana crassa — Simpson (1900), Simpson (1914)
Batavusiana (Crassunio) crassa — Stadnichenko (1984)
Crassiana crassa crassa — Kantor & Sysoev (2005), Kantor et al. (2010)

Unio ater Nilsson, 1822
‘Linn.’ Draparnaud, 1806, Hist. Moll. France: pl. 11, fig. 5 [as ‘Unio margaritifera’].
Nilsson, 1822, Hist. Moll. Svec.: 107.

Unio ater — Nilsson (1822), Rossmässler (1835), Küster (1854), Drouët (1881), Paetel (1890), Westerlund (1890)
Unio lemovicincae — Menke (1828), Menke (1830)
Margarita (Unio) batava — Lea (1836), Lea (1838)
Unio atra — Krynicki (1837)
Unio ater var. ? — Rossmässler (1838)
Margaron (Margaritana) margaritifera — Lea (1852)
Unio (Lymnium) ater — Moquin-Tandon (1855), Drouët (1857)
Unio (Mysca) tumidus limicola — Mörch (1863)
Unio crassus var. ater — Westerlund (1867)
Margaron (Unio) batavus — Lea (1870)
Unio batavus var. ater — Clessin (1877)
Unio (Groupe de l’U. ater) ater — Locard (1882), Locard (1893)
Unio (Ateriana) ater — Locard (1889)
Unio (sec. Lymnium) batavus — Simpson (1900), Simpson (1914)
Unio (s.s.) ater — Zhadin (1938)
Unio crassus crassus — Haas (1940), Haas (1969)
Batavusiana (s.s.) musiva — Zatravkin (1987)
Unio (Crassiana) crassus — Vinarski & Kantor (2016)
Unio crassus — Lopes-Lima et al. (2024)

Unio batavus Nilsson, 1822
‘Lamarck’ Nilsson, 1822, Hist. Moll. Svec.: 112.
(reference to Mya batava Maton & Rackett, 1807)

Unio batavus — Nilsson (1822), Clessin (1877), Zittel (1881)
Unio steveniana — Krynicki (1837)
Unio batava — Malzine (1867)

Unio crassus Nilsson, 1822
‘Retzius’ Nilsson, 1822, Hist. Moll. Svec.: 108.
(reference to Unio crassus Retzius, 1788)

Unio crassus — Nilsson (1822), Küster (1854)
Margarita (Unio) litoralis — Lea (1838)
Margaron (Unio) litoralis — Lea (1852)
Unio (Lymnium) crassus — Drouët (1857)
Margaron (Unio) crassus — Lea (1870)
Unio batavus var. crassus — Clessin (1877)
Unio (sec. Lymnium) batavus — Simpson (1900), Simpson (1914)

Unio rubens Menke, 1828
Menke, 1828, Syn. Method. Moll.: 64, 90.
Menke, 1830, Syn. Method. Moll.: 107, 149.

Unio rubens — Menke (1828), Menke (1830), Rossmässler (1837)
Margaron (Unio) batavus — Lea (1852), Lea (1870)
Unio crassus — Küster (1854), Lopes-Lima et al. (2024)
Unio (Lymnium) crassus var. — Drouët (1857)
Unio crassus var. rubens — Westerlund (1890)
Quadrula (sec. Fusconaia) coccinea — Simpson (1900), Simpson (1914)
Quadrula (Obliquata) coccinea — Frierson (1927)
Unio crassus crassus — Haas (1940), Haas (1969)
Pleurobema sintoxia — Parmalee & Bogan (1998), Williams et al. (2008)
Unio (Crassiana) crassus — Vinarski & Kantor (2016)

Unio rugatus Menke, 1828
Menke, 1828, Syn. Method. Moll.: 64, 90.
Menke, 1830, Syn. Method. Moll.: 107, 149.

Unio rugatus — Menke (1828), Menke (1830), Rossmässler (1837)
Unio batavus — Rossmässler (1835), Drouët (1889)
Margaron (Unio) batavus — Lea (1852), Lea (1870)
Unio batavus var. crassus — Clessin (1877)
Unio crassus var. rugatus — Westerlund (1890)
Unio (sec. Lymnium) batavus — Simpson (1900), Simpson (1914)
Unio consentaneus var. — Kobelt & Haas (1910)
Unio crassus crassus — Haas (1940), Haas (1969)
Unio (Crassiana) crassus — Vinarski & Kantor (2016)
Unio crassus — Lopes-Lima et al. (2024)

Unio lemovicensis Kickx, 1830
‘Daud.’ Kickx, 1830, Synopsin Molluscorum Brabantiae: 82.
(error for Unio lemovicincae Férussac, 1807)

Unio lemovicensis — Kickx (1830)

Unio rubens Menke, 1830
Menke, 1830, Syn. Method. Moll.: 149.
(reference to Unio rubens Menke, 1828)

Unio rubens — Menke (1830)
Unio (sec. Lymnium) littoralis — Simpson (1900), Simpson (1914)

Unio badius Kokeil, 1831
Kokeil, 1831, Mich. Comp.: pl. 16, fig. 36.

Unio badius — Kokeil (1831), J.F. Schmidt (1840)
Margaron (Unio) batavus — Lea (1852), Lea (1870)
Unio (s.s.) badius — Paetel (1890)
Unio batavus — Westerlund (1890)
Unio (sec. Lymnium) batavus — Simpson (1900), Simpson (1914)

Unio carynthiacus Lea, 1836
‘Zeigler’ of Authors.
Lea, 1836, Synopsis Fam. Naiades: 27.
Schlüter, 1838, Kurzgef. Syst. Verz. Conch. Samml.: 33.
Lea, 1838, Synopsis Fam. Naiades: 20.
J.F. Schmidt, 1840, Bull. Soc. imp. Nat. Moscou 13: 435.
C.B. Adams, 1847, Cat. Gen. Sp. Adams: 14.
Lea, 1852, Synopsis Fam. Naiades: 30.
Lea, 1870, Synopsis Fam. Naiades: 47.
Schröder, 1885, Bull. Soc. Malac. France 2: 229.
Servain, 1888, Bull. Soc. Malac. France 5: 317.
Locard, 1889, Contr. Faune Malacol. Franc. 13: 141 [35].
Germain, 1904, Bull. Soc. Sci. Nat. Ouest France 4: 264.
Germain, 1931, Faune de France 22: 728.
(error for Unio carinthiacus Rossmässler, 1836)

Unio carynthiacus — Lea (1836)

Unio fuscus Lea, 1836
‘Zieg.’ Lea, 1836, Synopsis Fam. Naiades: 27.
(intro'd as syn of Mya batava Maton & Rackett, 1807)

Unio fuscus — Lea (1836)
Margarita (Unio) batava — Lea (1836), Lea (1838)
Margaron (Unio) batavus — Lea (1852), Lea (1870)

Unio gibba Lea, 1836
‘Mühl. Pfeif.’ Lea, 1836, Synopsis Fam. Naiades: 27.
(intro'd as syn of Mya batava Maton & Rackett, 1807)

Unio gibba — Lea (1836)
Margarita (Unio) batava — Lea (1836), Lea (1838)
Margaron (Unio) batavus — Lea (1852), Lea (1870)

Unio acutus Rossmässler, 1837
Rossmässler, 1837, Icon. Land- und Süssw.-Moll. 1 (5-6): 56, pl 29, fig. 413 [as ‘Unio an n. sp.?’]. [nomen nudum]

Unio acutus — Rossmässler (1837)
Unio batavus var. crassus — Clessin (1877)
Unio (sec. Hyridella) crassus — Paetel (1890)
Unio crassus var. acutus — Westerlund (1890)

Mysca batava var. fuscula subvar. pallipes Schlüter, 1838
‘Ziegl.’ Schlüter, 1838, Kurzgef. Syst. Verz. Conch. Samml.: 33. [infra-subsp.]
(intro'd as intra-sp. of Unio fusculus Rossmässler, 1836)

Mysca batava var. fuscula subvar. pallipes — Schlüter (1838)

Mysca reniformis subvar. kokeilli Schlüter, 1838
Schlüter, 1838, Kurzgef. Syst. Verz. Conch. Samml.: 32. [infra-subsp.]
(intro'd as intra-sp. of Unio reniformis Rossmässler, 1836)

Mysca reniformis subvar. kokeilli — Schlüter (1838)

Unio gargottae Rossmässler, 1838
Rossmässler, 1838, Icon. Land- und Süssw.-Moll. 2 (1-2): 26, pl. 35, fig. 493.
(misidentified reference to Unio gargottae Philippi, 1836)

Unio gargottae — Rossmässler (1838)
Unio gargottæ — Bourguignat (1883)
Unio (sec. Lymnium) batavus — Simpson (1900), Simpson (1914)

Unio corrugata Mauduyt, 1839
Mauduyt, 1839, Tableau Indic. Moll. Vienne: 8, pl. 1, fig. 1.

Unio corrugata — Mauduyt (1839)
Unio (Batavi) moulinsianus — Dupuy (1852)
Unio (Lymnium) batavus — Moquin-Tandon (1855)
Unio corrugatus — Westerlund (1890)
Unio (sec. Lymnium) batavus — Simpson (1900), Simpson (1914)

Unio gangrenosus J.F. Schmidt, 1840
‘Ziegler’ J.F. Schmidt, 1840, Bull. Soc. imp. Nat. Moscou 13: 436.
J.F. Schmidt, 1847, Conch. Krain: 26.

Unio gangrenosus — J.F. Schmidt (1840), C.B. Adams (1847)
Unio gangraenosus — Küster (1855)
Unio (Mancusiana) gangrenosus — Locard (1889)
Unio gangrænosus — Westerlund (1890)
Unio (sec. Lymnium) batavus — Simpson (1900), Simpson (1914)
Unio crassus cytherea — Haas (1940), Haas (1969)

Unio crassissimus Hanley, 1843
Schlüter, 1838, Kurzgef. Syst. Verz. Conch. Samml.: 33 [as ‘Mysca tumida var. crassissima’]. [nomen nudum]
‘Fer.’ Hanley, 1843, Illustr. Descr. Cat. Rec. Bivalve Shells: 209.

Unio crassissimus — Hanley (1843)
Mysca tumida var. crassissima — Schlüter (1838)
Margaron (Unio) crassus — Lea (1870)
Margaritana crassa — Simpson (1900), Simpson (1914)

Unio manca Millet, 1843
‘Lamarck’ of Authors.
Millet, 1843, Mag. Zool.: 4, pl. 64, fig. 2.
(misidentified reference to Unio manca Lamarck, 1819)

Unio manca — Millet (1843)
Unio (Mancusiana) mancus — Locard (1889), Germain (1904)
Unio pruinosus — Drouët (1889)
Unio (sec. Lymnium) batavus — Simpson (1900), Simpson (1914)
Unio mancus — Germain (1931)

Unio requieni Rossmässler, 1844
‘Mich.’ Rossmässler, 1844, Icon. Land- und Süssw.-Moll. 2 (6): 29, pl. 57, fig. 757-761.
(misidentified reference to Unio requienii Michaud, 1831)

Unio requieni — Rossmässler (1844)
Unio requienii — Küster (1855)
Margaron (Unio) pictorum — Lea (1870)
Unio subtilis — Drouët (1889)
Unio meretricis — Westerlund (1890)
Unio (sec. Lymnium) batavus — Simpson (1900)
Unio (sec. Lymnium) batavus var. requieni — Simpson (1914)

Unio batavus var. pusillus Rossmässler, 1844
Rossmässler, 1844, Icon. Land- und Süssw.-Moll. 2 (6): .
(intro'd as intra-sp. of Mya batava Maton & Rackett, 1807; homonym of Unio pusillus Lea, 1840)

Unio batavus var. pusillus — Rossmässler (1844)
Unio (Lymnium) batavus var. L nanus — Moquin-Tandon (1855)
Unio batavus var. crassus — Clessin (1877)
Unio amnicus — Westerlund (1890)

Unio glaucinus Stabile, 1845
Stabile, 1845, Faun. Lug.: f. 75.
(misidentified reference to Unio glaucinus Porro, 1838)

Unio glaucinus — Stabile (1845)
Unio (Limnium) requienii var. vulgaris — Stabile (1859)
Unio potamius — Bourguignat (1883)
Unio batavus var. potamius — Westerlund (1890)
Unio (sec. Lymnium) batavus — Simpson (1900)

Unio piscinalis J.F. Schmidt, 1847
J.F. Schmidt, 1847, Conch. Krain: 27.
(misidentified reference to Unio piscinalis Rossmässler, 1836)

Unio piscinalis — J.F. Schmidt (1847)
Unio reniformis — Küster (1854)

Unio retzii C.B. Adams, 1847
‘Küst.’ C.B. Adams, 1847, Cat. Gen. Sp. Adams: 14. [nomen nudum]

Unio retzii — C.B. Adams (1847)
Margaron (Unio) batavus — Lea (1852), Lea (1870)

Unio ripario Philippi, 1847
‘Pfr’ Philippi, 1847, Z. Malak. 4: 95.
(error for Unio riparia Pfeiffer, 1821)

Unio ripario — Philippi (1847)

Unio drouetii Dupuy, 1849
Dupuy, 1849, Cat. Extramar. Galliae Test.: no. 326.
Dupuy, 1852, Hist. Natur. Moll. France (6): 639, pl. 24, fig. 12.

Unio drouetii — Dupuy (1849)
Unio (Batavi) drouetii — Dupuy (1852)
Unio (Lymnium) batavus var. I drouëtii — Moquin-Tandon (1855)
Unio (Groupe de l’U. Batavus) droueti — Locard (1882)
Unio (Batavusiana) droueti — Locard (1889)
Unio (sec. Hyridella) batavus vr. drouëti — Paetel (1890)
Unio batavus var. drouëti — Westerlund (1890)
Unio (Groupe de l’U. batavus) droueti — Locard (1893)
Unio (sec. Lymnium) batavus — Simpson (1900), Simpson (1914)
Unio batavus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969), Fauna Europaea (2012)
Unio crassus — Lopes-Lima et al. (2024)

Unio moquinianus Sandberger & Koch, 1851
Sandberger & Koch, 1851, Jb. Nassau. Ver. Naturk. 7: 245.
(misidentified reference to Unio moquinianus Dupuy, 1843)

Unio moquinianus — Sandberger & Koch (1851)
Unio kochi — Kobelt (1886)

Unio argens Lea, 1852
‘Küst.’ Lea, 1852, Synopsis Fam. Naiades: 30.
(intro'd as syn of Mya batava Maton & Rackett, 1807)

Unio argens — Lea (1852)
Margaron (Unio) batavus — Lea (1852), Lea (1870)

Unio drouatii Lea, 1852
‘Dup.’ Lea, 1852, Synopsis Fam. Naiades: 39.
(error for Unio drouetii Dupuy, 1849)

Unio drouatii — Lea (1852)

Unio kurrii Lea, 1852
‘Küst.’ Lea, 1852, Synopsis Fam. Naiades: 30.
(intro'd as syn of Mya batava Maton & Rackett, 1807)

Unio kurrii — Lea (1852)
Margaron (Unio) batavus — Lea (1852), Lea (1870)

Unio moquinianus Drouët, 1852
Drouët, 1852, Etudes Nayades de la France: pl. 6, fig. 3.
(misidentified reference to Unio moquinianus Dupuy, 1843)

Unio moquinianus — Drouët (1852)
Unio (Ateriana) lambottei — Locard (1889)
Unio consentaneus — Germain (1931)

Unio nanus Dupuy, 1852
‘Lamarck’ Dupuy, 1852, Hist. Natur. Moll. France (6): pl. 25, fig. 16.
(misidentified reference to Unio nana Lamarck, 1819)

Unio nanus — Dupuy (1852), Kobelt (1879)
Unio (sec. Lymnium) batavus — Simpson (1900)
Unio (Rhomboïdiana) nana — Germain (1904)
Unio nana — Kobelt & Haas (1910)
Unio (sec. Lymnium) batavus var. requieni — Simpson (1914)
Unio mancus var. nanus — Germain (1931)

Unio ovalis Dupuy, 1852
Dupuy, 1852, Hist. Natur. Moll. France (6): pl. 25, fig. 13.
(misidentified reference to Mya ovalis Montagu, 1803)

Unio ovalis — Dupuy (1852)
Unio (Lymnium) batavus var. ovalis — Drouët (1857)
Unio (Batavusiana) besnardianus — Locard (1889)
Unio besnardianus — Westerlund (1890)
Unio (sec. Lymnium) batavus — Simpson (1900), Simpson (1914)
Unio (Batavusiana) besnardi — Germain (1904)
Unio batavus — Germain (1931)

Unio pescinalis Lea, 1852
‘Zieg.’ Lea, 1852, Synopsis Fam. Naiades: 30.
(error for Unio piscinalis Rossmässler, 1836)

Unio pescinalis — Lea (1852)

Unio ventricosus Scholtz, 1853
Scholtz, 1853, unknown publication: .
(homonym of Unio ventricosus Barnes, 1823)

Unio ventricosus — Scholtz (1853)
Unio batavus var. ventricosus — Westerlund (1890)

Unio crassus Küster, 1854
‘Retz.’ Küster, 1854, Conchyl. Cab. 9(2): 113, pl. 31, fig. 5. [in part]
(reference to Unio crassus Retzius, 1788)

Unio crassus — Küster (1854), Drouët (1881)
Unio batavus var. crassus — Clessin (1877)
Lampsilis (sec. Eurynia) radiatus — Simpson (1900)
Lampsilis (Eurynia) radiata — Simpson (1914)

Unio ater Moquin-Tandon, 1855
‘Nilss.’ Moquin-Tandon, 1855, Hist. Natur. Moll. France 2: 570.
(reference to Unio ater Nilsson, 1822)

Unio ater — Moquin-Tandon (1855)
Unio (Melasiana) melas — Locard (1889)

Unio capigliolo Küster, 1855
‘Payraudeau’ of Authors.
Küster, 1855, Conchyl. Cab. 9(2): 125, pl. 34, figs. 5-6. [in part]
Sowerby, 1868, Conch. Icon. 16: pl. 75, sp. 392.
(misidentified reference to Unio capigliolo Payraudeau, 1826)

Unio capigliolo — Küster (1855)
Unio (sec. Lymnium) batavus — Simpson (1900), Simpson (1914)

Unio gangraenosus Küster, 1855
‘Ziegler’ of Authors.
Küster, 1855, Conchyl. Cab. 9(2): 124.
Westerlund, 1890, Fauna Paläarct. Binnenconch. 2(7): 81.
Haas, 1940, Zool. Ser. Field Mus. Nat. Hist. 24: 130.
Haas, 1969, Das Tierreich 88: 46.
(error for Unio gangrenosus J.F. Schmidt, 1840)

Unio gangraenosus — Küster (1855)
Unio (Crassiana) crassus — Vinarski & Kantor (2016)

Unio rivularis Moquin-Tandon, 1855
Rossmässler, 1844, Icon. Land- und Süssw.-Moll. 2 (6): fig. 414.
Moquin-Tandon, 1855, Hist. Natur. Moll. France 2: 571.
(homonym of Unio rivularis Conrad, 1853)

Unio rivularis — Moquin-Tandon (1855)
Unio (Lymnium) batavus var. B rivularis — Moquin-Tandon (1855)
Unio (sec. Hyridella) batavus vr. rivularis — Paetel (1890)
Unio batavus — Westerlund (1890)
Unio (Crassiana) crassus — Vinarski & Kantor (2016)

Unio batavus var. sarratinus Moquin-Tandon, 1855
‘St. Germain, in litt.’ Moquin-Tandon, 1855, Hist. Natur. Moll. France 2: 571.
(intro'd as intra-sp. of Mya batava Maton & Rackett, 1807)

Unio batavus var. sarratinus — Moquin-Tandon (1855), de Joannis (1859), Westerlund (1890)
Unio (Lymnium) batavus var. G sarratinus — Moquin-Tandon (1855)
Unio (sec. Hyridella) batavus vr. sarratianus — Paetel (1890)

Unio ater Drouët, 1857
‘Nilss.’ Drouët, 1857, Étud. Naïades. France, Unio: 72, pl. 4, fig. 1.
(reference to Unio ater Nilsson, 1822)

Unio ater — Drouët (1857)
Unio (Ateriana) ignari — Locard (1889)
Unio ignari — Westerlund (1890)
Unio consentaneus — Germain (1931)

Unio nanus Drouët, 1857
‘Lam.’ Drouët, 1857, Étud. Naïades. France, Unio: pl. 5, fig. 2.
(misidentified reference to Unio nana Lamarck, 1819)

Unio nanus — Drouët (1857)
Unio (Nanusiana) nanus — Locard (1889)
Unio (Rhomboïdiana) nana — Germain (1904)
Unio nana — Kobelt & Haas (1910)
Unio mancus var. nanus — Germain (1931)

Unio platyrhynchoideus Drouët, 1857
‘Dup.’ Drouët, 1857, Étud. Naïades. France, Unio: 100, pl. 9, fig. 1.
(misidentified reference to Unio platyrinchoideus Dupuy, 1849)

Unio platyrhynchoideus — Drouët (1857)
Unio (Ateriana) ignariformis — Locard (1889)
Unio ignariformis — Westerlund (1890)

Unio batavus de Joannis, 1859
de Joannis, 1859, Ann. Soc. Linn. Maine-et-Loire 3: pl. 9, fig. 2. [in part]
(reference to Mya batava Maton & Rackett, 1807)

Unio batavus — de Joannis (1859)
Unio (Batavusiana) batavus radiata — Germain (1904)

Unio batavus var. acutus de Joannis, 1859
de Joannis, 1859, Ann. Soc. Linn. Maine-et-Loire 3: 287, pl. 10, fig. 1.
(homonym of Unio acutus Rossmässler, 1837)

Unio batavus var. acutus — de Joannis (1859)

Unio batavus var. baraceus de Joannis, 1859
de Joannis, 1859, Ann. Soc. Linn. Maine-et-Loire 3: 285, pl. 9, fig. 7.
(intro'd as intra-sp. of Mya batava Maton & Rackett, 1807)

Unio batavus var. baraceus — de Joannis (1859), Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio batavus var. courtilieri de Joannis, 1859
de Joannis, 1859, Ann. Soc. Linn. Maine-et-Loire 3: pl. 9, fig. 3.
(reference to Unio courtillieri Hattemann, 1859; error for Unio courtillieri Hattemann, 1859)

Unio batavus var. courtilieri — de Joannis (1859)
Unio (Batavusiana) batavus courtilieri — Germain (1904)
Unio batavus — Germain (1931)

Unio batavus var. longus de Joannis, 1859
de Joannis, 1859, Ann. Soc. Linn. Maine-et-Loire 3: 288, pl. 10, fig. 3.
(intro'd as intra-sp. of Mya batava Maton & Rackett, 1807; homonym of Unio longus Philippi, 1869)

Unio batavus var. longus — de Joannis (1859)
Unio batavus — Germain (1931)

Unio batavus var. minimus de Joannis, 1859
de Joannis, 1859, Ann. Soc. Linn. Maine-et-Loire 3: 289, pl. 9, fig. 6.
(intro'd as intra-sp. of Mya batava Maton & Rackett, 1807; homonym of Unio requienii var. minima Drouët, 1857)

Unio batavus var. minimus — de Joannis (1859)
Unio mancus var. amnicus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio batavus var. nanus de Joannis, 1859
de Joannis, 1859, Ann. Soc. Linn. Maine-et-Loire 3: pl. 9, fig. 8. [in part]
(misidentified reference to Unio nana Lamarck, 1819)

Unio batavus var. nanus — de Joannis (1859)
Unio (Melasiana) melas — Germain (1904)
Unio mancus var. amnicus — Germain (1931)

Unio batavus var. nanus de Joannis, 1859
de Joannis, 1859, Ann. Soc. Linn. Maine-et-Loire 3: pl. 9, fig. 10. [in part]
(misidentified reference to Unio nana Lamarck, 1819)

Unio batavus var. nanus — de Joannis (1859)
Unio (Rhomboïdiana) nana — Germain (1904)

Unio batavus var. ovalis de Joannis, 1859
de Joannis, 1859, Ann. Soc. Linn. Maine-et-Loire 3: pl. 10, fig. 2.
(misidentified reference to Mya ovalis Montagu, 1803)

Unio batavus var. ovalis — de Joannis (1859)
Unio (Batavusiana) ligericus — Locard (1889), Germain (1904)
Unio batavus — Germain (1931)

Unio batavus var. senotius de Joannis, 1859
de Joannis, 1859, Ann. Soc. Linn. Maine-et-Loire 3: 288, pl. 10, fig. 4.

Unio batavus var. senotius — de Joannis (1859)

Unio orientalis Küster, 1862
‘Parreis in sched.’ Küster, 1862, Conchyl. Cab. 9(2): 267.
(intro'd as syn of Unio kotschyi Küster, 1862)

Unio orientalis — Küster (1862)
Unio kotschyi — Küster (1862)

Unio sanderi Bourguignat, 1862
Bourguignat, 1862, Malacol. Lac Quatre-Cantons: 55.
Bourguignat, 1864, Malac. Aix-les-Baines: 76.

Unio sanderi — Bourguignat (1862)
Unio sandriopsis — Servain (1885), Westerlund (1890)
Unio (Sandriana) sandriopsis — Locard (1889)
Unio (Groupe de l’U. macrorhynchus) sandriopsis — Locard (1893)
Unio mancus var. macrorhynchus — Germain (1931)

Unio ater Bourguignat, 1864
Bourguignat, 1864, Malac. Aix-les-Baines: 73.
(reference to Unio ater Nilsson, 1822)

Unio ater — Bourguignat (1864)
Unio (Groupe de l’U. sandrii) macrorhynchus — Locard (1882)
Unio (Sandriana) macrorhynchus — Locard (1889)
Unio macrorhynchus — Westerlund (1890)
Unio mancus var. macrorhynchus — Germain (1931)

Unio merdiger Reeve, 1865
Jay, 1839, Cat. Rec. Shells: 25, sp. 1065.
‘Walsh.’ Lea, 1852, Synopsis Fam. Naiades: . [nomen nudum]
‘Watts’ Reeve, 1865, Conch. Icon. 16: pl. 28, sp. 145.

Unio merdiger — Reeve (1865)
Margaron (Unio) batavus — Lea (1852), Lea (1870)
Unio (sec. Lymnium) batavus — Simpson (1900), Simpson (1914)

Unio lambottei Malzine, 1867
Malzine, 1867, Essai faun. malac. Belg.: 32, pl. 2, figs, 3-4.

Unio lambottei — Malzine (1867), Colbeau (1868), Westerlund (1890)
Unio (Groupe de l’U. crassus) lambottei — Locard (1882)
Unio (Ateriana) lambottei — Locard (1889), Germain (1904)
Unio berilloni — Westerlund (1890)
Unio (Groupe de l’U. ater) lamboltei — Locard (1893)
Unio consentaneus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio ryckholtii Malzine, 1867
Malzine, 1867, Essai faun. malac. Belg.: 32, pl. 2, figs. 1-2.

Unio ryckholtii — Malzine (1867), Colbeau (1868)
Unio (Groupe de l’U. Batavus) ryckholti — Locard (1882)
Unio (Ellipsopsisiana) ryckholtii — Locard (1889)
Unio batavus var. ryckholtii — Westerlund (1890)
Unio (Groupe de l’U. batavus) ryckholti — Locard (1893)
Unio (Batavusiana) ryckholtii — Germain (1904)
Unio batavus — Germain (1931)

Unio batavus var. squamosus Brot, 1867
Brot, 1867, Assoc. Zool. Léman Ann. (1866): pl. 9, fig. 1.
(intro'd as intra-sp. of Mya batava Maton & Rackett, 1807; misidentified reference to Unio batavus var. squamosus Charpentier, 1837)

Unio batavus var. squamosus — Brot (1867)
Unio sandriopsis — Servain (1885), Westerlund (1890)
Unio (Ateriana) ater — Locard (1889)
Unio consentaneus — Germain (1931)

Unio ryckholti var. cuneata Malzine, 1867
Malzine, 1867, Essai faun. malac. Belg.: 32.
(homonym of Unio cuneatus Jacquemin, 1835; intro'd as intra-sp. of Unio ryckholtii Malzine, 1867)

Unio ryckholti var. cuneata — Malzine (1867)
Unio ryckholtii var. cuneatus — Colbeau (1868)
Unio batavus — Germain (1931)

Unio (tumidus) elegans Westerlund, 1867
Westerlund, 1867, Malak. Blatt. 14: 202, 206.
(homonym of Unio elegans Lea, 1831)

Unio (tumidus) elegans — Westerlund (1867)
Unio ater var. elegans — Westerlund (1890)
Unio (Crassiana) crassus — Vinarski & Kantor (2016)

Unio batavus var. belgicus Colbeau, 1868
Colbeau, 1868, Ann. Soc. Malac. Belgique 3: 106, pl. 4, fig. 3.
(intro'd as intra-sp. of Mya batava Maton & Rackett, 1807)

Unio batavus var. belgicus — Colbeau (1868)
Unio (Groupe de l’U. crassus) incurvatus — Locard (1882)
Unio (sec. Hyridella) batavus vr. belgicus — Paetel (1890)
Unio batavus var. incurvatus — Westerlund (1890)

Unio batavus var. fontainei Colbeau, 1868
‘Lecomte’ Colbeau, 1868, Ann. Soc. Malac. Belgique 3: 106. [nomen nudum]
(intro'd as intra-sp. of Mya batava Maton & Rackett, 1807)

Unio batavus var. fontainei — Colbeau (1868)
Unio (sec. Hyridella) batavus vr. fontainei — Paetel (1890)

Unio labacencis Lea, 1870
‘Pfeif. Mühl.’ Lea, 1870, Synopsis Fam. Naiades: 47.
(error for Unio labacensis Rossmässler, 1835)

Unio labacencis — Lea (1870)

Unio maceerthyanus Lea, 1870
‘Bourg.’ Lea, 1870, Synopsis Fam. Naiades: 48.
(error for Unio maccarthyanus Bourguignat, 1866; misidentified reference to Unio maccarthyanus Bourguignat, 1866)

Unio maceerthyanus — Lea (1870)

Unio crassus var. maximus Kobelt, 1872
Kobelt, 1872, Malak. Blatt. 19: 142, pl. 5.
Kobelt, 1884, Icon. Land- und Süssw.-Moll. 1: 67, pl. 29, figs. 218-219.
(intro'd as intra-sp. of Unio crassus Retzius, 1788; homonym of Unio pictorum maxima Mörch, 1863)

Unio crassus var. maximus — Kobelt (1872), Kobelt (1884)
Unio batavus var. pseudo-litoralis — Clessin (1877)
Unio crassus crassus — Haas (1940), Haas (1969)
Unio (Crassiana) crassus — Vinarski & Kantor (2016)
Unio crassus — Lopes-Lima et al. (2024)

Unio batavus Miller, 1873
Miller, 1873, Schalth. Bodensees: pl. 2, fig. 8.
(reference to Mya batava Maton & Rackett, 1807)

Unio batavus — Miller (1873)
Unio (Socardiana) subrobustus — Locard (1889)
Unio crassus — Germain (1931)

Unio pseudolitoralis Clessin, 1877
Clessin, 1875, Verh. Ver. Naturgesch. Hamburg 2: 258 [as ‘pseudo-litoralis’]. [nomen nudum]
Clessin, 1877, Deutsche Excurs. Moll. Fauna: 469, text fig. 305 [as ‘pseudo-litoralis’].

Unio pseudolitoralis — Clessin (1877), Westerlund (1890)
Unio batavus var. pseudo-litoralis — Clessin (1877)
Unio crassus var. maximus — Kobelt (1884)
Unio (s.s.) crassus var. pseudolitoralis — Zhadin (1938)
Unio crassus crassus — Haas (1940), Haas (1969)
Unio crassus var. pseudolitoralis — Zhadin (1965)
Batavusiana (Crassunio) crassa — Stadnichenko (1984), Zatravkin (1987)
Crassiana crassa crassa — Kantor & Sysoev (2005), Kantor et al. (2010)
Unio (Crassiana) crassus — Vinarski & Kantor (2016)
Unio crassus — Lopes-Lima et al. (2024)

Unio subtilis Drouët, 1879
Drouët, 1879, J. de Conch. 27: 142.

Unio subtilis — Drouët (1879), Kobelt (1886), Drouët (1889), Westerlund (1890)
Unio (Groupe de l’U. Brevierei) subtilis — Locard (1882)
Unio (Mancusiana) subtilis — Locard (1889)
Unio (Groupe de l’U. mancus) subtilis — Locard (1893)
Unio (Groupe des Mancusiana) subtilis — Caziot (1907)
Unio mancus var. amnicus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio vescoi var. conemenosi Westerlund & Blanc, 1879
Blanc in Westerlund & Blanc, 1879, Aperçu Faune Malac. Grèce: 145.
(intro'd as intra-sp. of Unio vescoi Bourguignat, 1856)

Unio vescoi var. conemenosi — Westerlund & Blanc (1879), Westerlund (1890)

Unio moquinianus Brevière, 1880
Brevière, 1880, Cat. moll. Nièvre: 26.
(misidentified reference to Unio moquinianus Dupuy, 1843)

Unio moquinianus — Brevière (1880)
Unio (Groupe de l’U. Brevierei) brevierei — Locard (1882)
Unio (Ateriana) brevierei — Locard (1889)
Unio consentaneus — Germain (1931)

Unio requieni Brevière, 1880
‘Mich.’ Brevière, 1880, Cat. moll. Nièvre: 26.
(misidentified reference to Unio requienii Michaud, 1831)

Unio requieni — Brevière (1880)
Unio (Groupe de l’U. Brevierei) brevierei — Locard (1882)
Unio (Ateriana) brevierei — Locard (1889)
Unio consentaneus — Germain (1931)

Unio diagonalis Drouët, 1881
‘Held’ Drouët, 1881, J. de Conch. 29: 27.
(intro'd as syn of Unio batavus var. squamosus Charpentier, 1837)

Unio diagonalis — Drouët (1881)
Unio squamosus — Drouët (1881), Drouët (1889)

Unio andegavensis Locard, 1882
‘Servain’ Locard, 1882, Prodr. Malac. Franç. 1: 289, 359.
‘Servain’ Locard, 1889, Contr. Faune Malacol. Franc. 13: 142 [36].
‘Servain’ Locard, 1893, Coqu. Eau Douce Saumâtre France: 167.
Type(s): POSSIBLE SYNTYPE MNHN_22100, La Maine à Angers.

Unio andegavensis — Locard (1882)
Unio (Groupe de l’U. Batavus) andegavensis — Locard (1882)
Unio (Bataviana) andegavensis — Servain (1888)
Unio (Batavusiana) andegavensis — Locard (1889), Germain (1904)
Unio batavus var. andegavensis — Westerlund (1890)
Unio (Groupe de l’U. batavus) andegavensis — Locard (1893)
Unio batavus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio arenarum Locard, 1882
Bourguignat in Locard, 1882, Prodr. Malac. Franç. 1: 289, 358.
Bourguignat in Locard, 1889, Contr. Faune Malacol. Franc. 13: 144 [38].
Bourguignat in Locard, 1893, Coqu. Eau Douce Saumâtre France: 171.
Type(s): IDIOTYPE MNHN_22110, Basse Indre (Loire-Inf.).

Unio arenarum — Locard (1882)
Unio (Groupe de l’U. Batavus) arenarum — Locard (1882)
Unio (Bataviana) arenarum — Servain (1888)
Unio (Batavusiana) arenarum — Locard (1889), Germain (1904)
Unio batavus var. arenarum — Westerlund (1890)
Unio (Groupe de l’U. batavus) arenarum — Locard (1893)
Unio batavus var. baraceus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio berthelini Locard, 1882
Bourguignat in Locard, 1882, Prodr. Malac. Franç. 1: 290, 359.
Bourguignat in Locard, 1889, Contr. Faune Malacol. Franc. 13: 137 [31].
Bourguignat in Locard, 1893, Coqu. Eau Douce Saumâtre France: 163.
Type(s): IDIOTYPE MNHN_22132, La Marne, à Charenton (Seine).

Unio berthelini — Locard (1882), Westerlund (1890)
Unio (Groupe de l’U. fusculus) berthelini — Locard (1882)
Unio (Amniciana) berthelini — Servain (1888)
Unio (Amnicusiana) berthelini — Locard (1889)
Unio (Groupe de l’U. amnicus) berthelini — Locard (1893)
Unio mancus var. amnicus form riparius — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio brevierei Locard, 1882
‘Bourguignat’ Locard, 1882, Prodr. Malac. Franç. 1: 286 [as ‘brevieri’], 356.
‘Bourguignat’ Locard, 1889, Contr. Faune Malacol. Franc. 13: 148 [42].
‘Bourguignat’ Locard, 1893, Coqu. Eau Douce Saumâtre France: 178.
(correct name for Unio brevieri Locard, 1882)
Type(s): LECTOTYPE MNHN_22145, La Canne, à Goulnot.

Unio brevierei — Locard (1882), Westerlund (1890)
Unio (Groupe de l’U. Brevierei) brevierei — Locard (1882)
Unio (Ateriana) brevierei — Locard (1889)
Unio (Groupe de l’U. ater) brevierei — Locard (1893)
Unio (Ateriana) brevieri — Germain (1904)
Unio (Groupe des Ateriana) brevierei — Caziot (1907)
Unio consentaneus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio brevieri Locard, 1882
‘Bourguignat in Locard’ of Authors.
Locard, 1882, Prodr. Malac. Franç. 1: 286.
Germain, 1904, Bull. Soc. Sci. Nat. Ouest France 4: 265.
Germain, 1931, Faune de France 22: 732.
Haas, 1940, Zool. Ser. Field Mus. Nat. Hist. 24: 130.
Haas, 1969, Das Tierreich 88: 49.
(incorrect original spelling of Unio brevierei Locard, 1882)

Unio brevieri — Locard (1882)
Unio crassus — Lopes-Lima et al. (2024)

Unio carantoni Locard, 1882
Bourguignat in Locard, 1882, Prodr. Malac. Franç. 1: 295, 364.
‘Coutagne’ Locard, 1889, Contr. Faune Malacol. Franc. 13: 129 [23].
‘Coutagne’ Locard, 1893, Coqu. Eau Douce Saumâtre France: 157.
Type(s): IDIOTYPE MNHN_22162, [illegible] (Charente).

Unio carantoni — Locard (1882), Westerlund (1890), Rochebrune (1903)
Unio (Groupe de l’U. falsus) carantoni — Locard (1882)
Unio (Mancusiana) carantoni — Locard (1889)
Unio (Groupe de l’U. mancus) carantoni — Locard (1893)
Unio mancus var. amnicus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio crassatellus Locard, 1882
Bourguignat in Locard, 1882, Prodr. Malac. Franç. 1: 286, 356.
Bourguignat in Locard, 1889, Contr. Faune Malacol. Franc. 13: 136 [30].
Bourguignat in Locard, 1893, Coqu. Eau Douce Saumâtre France: 162.
Type(s): LECTOTYPE MNHN_22182, La Canne, à Jailly (Nièvre).

Unio crassatellus — Locard (1882), Rochebrune (1903)
Unio (Groupe de l’U. crassus) crassatellus — Locard (1882)
Unio (Crassiana) crassatellus — Locard (1889)
Unio (Groupe de l’U. feliciani) crassatellus — Locard (1893)
Unio crassus — Germain (1931), Lopes-Lima et al. (2024)
Unio crassus batavus — Haas (1940), Haas (1969)

Unio dubisanus Locard, 1882
‘Coutagne’ Locard, 1882, Prodr. Malac. Franç. 1: 291.
(incorrect original spelling of Unio dubisianus Locard, 1882)

Unio dubisanus — Locard (1882)

Unio dubisopsis Locard, 1882
Locard, 1882, Prodr. Malac. Franç. 1: 291.
(incorrect original spelling of Unio dubisiopsis Locard, 1882)

Unio dubisopsis — Locard (1882)

Unio feliciani Locard, 1882
Bourguignat in Locard, 1882, Prodr. Malac. Franç. 1: 285, 355.
Bourguignat in Locard, 1889, Contr. Faune Malacol. Franc. 13: 134 [28].
Bourguignat in Locard, 1893, Coqu. Eau Douce Saumâtre France: 161, fig. 175.
Bourguignat in Locard, 1899, Conchyliol. Portug.: 217.
Type(s): POSSIBLE SYNTYPE MNHN_22215, La Moselle.

Unio feliciani — Locard (1882)
Unio (Crassiana) feliciani — Servain (1882)
Unio (Groupe de l’U. crassus) feliciani — Locard (1882)
Unio (Socardiana) feliciani — Locard (1889), Germain (1904)
Unio (Groupe de l’U. feliciani) feliciani — Locard (1893)
Unio (groupe de l’U. feliciani) feliciani — Locard (1899)
Unio batavus var. baraceus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio lagnisicus Locard, 1882
Bourguignat in Locard, 1882, Prodr. Malac. Franç. 1: 291, 359.
‘Bourguignat’ Locard, 1889, Contr. Faune Malacol. Franc. 13: 127 [21] [as ‘lagnysicus’].
‘Bourguignat’ Locard, 1893, Coqu. Eau Douce Saumâtre France: 154 [as ‘lagnysicus’].
Type(s): SYNTYPE MNHN_22303, Condal (Saône-et-Loire).

Unio lagnisicus — Locard (1882)
Unio (Groupe de l’U. mancus) lagnisicus — Locard (1882)
Unio (Nanusiana) lagnysicus — Locard (1889)
Unio lagnysicus — Westerlund (1890), Rochebrune (1903), Kobelt & Haas (1910)
Unio (Groupe de l’U. nanus) lagnysicus — Locard (1893)
Unio (groupe de l’U. nanus) lagnysicus — Dumas (1895)
Unio mancus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio mancus mancus — Prie (2012)
Unio crassus — Lopes-Lima et al. (2024)

Unio ligericus Locard, 1882
Bourguignat in Locard, 1882, Prodr. Malac. Franç. 1: 289, 358.
‘Bourguignat’ Locard, 1889, Contr. Faune Malacol. Franc. 13: 142 [36].
‘Bourguignat’ Locard, 1893, Coqu. Eau Douce Saumâtre France: 169.
Type(s): IDIOTYPE MNHN_22310, La Loire à Ingrandes.

Unio ligericus — Locard (1882)
Unio (Groupe de l’U. Batavus) ligericus — Locard (1882)
Unio (Bataviana) ligericus — Servain (1888)
Unio (Batavusiana) ligericus — Locard (1889), Germain (1904)
Unio batavus var. ligericus — Westerlund (1890)
Unio (Groupe de l’U. batavus) ligericus — Locard (1893)
Unio batavus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio locardianus Locard, 1882
Bourguignat in Locard, 1882, Prodr. Malac. Franç. 1: 287, 356.
‘Bourguignat’ Locard, 1889, Contr. Faune Malacol. Franc. 13: 138 [32].
‘Bourguignat’ Locard, 1893, Coqu. Eau Douce Saumâtre France: 164, fig. 178.
Type(s): IDIOTYPE MNHN_22316, l’Albane (Côte-d’Or).

Unio locardianus — Locard (1882), Westerlund (1890)
Unio (Groupe de l’U. locardianus) locardianus — Locard (1882), Locard (1893)
Unio (Locardiana) locardianus — Locard (1889)
Unio crassus — Germain (1931), Lopes-Lima et al. (2024)
Unio crassus batavus — Haas (1940), Haas (1969)

Unio marcellinus Locard, 1882
Bourguignat in Locard, 1882, Prodr. Malac. Franç. 1: 285, 355.
‘Berthier’ Locard, 1889, Contr. Faune Malacol. Franc. 13: 149 [43].
‘Berthier’ Locard, 1893, Coqu. Eau Douce Saumâtre France: 180.

Unio marcellinus — Locard (1882)
Unio (Groupe de l’U. Philippei) marcellinus — Locard (1882)
Unio (Ateriana) marcellinus — Locard (1889), Germain (1904)
Unio (Groupe de l’U. ater) marcellinus — Locard (1893)
Unio consentaneus var. moquini — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio mancus moquinianus — Prie (2012)
Unio crassus — Lopes-Lima et al. (2024)

Unio matronicus Locard, 1882
Bourguignat in Locard, 1882, Prodr. Malac. Franç. 1: 289, 358.
‘Bourguignat’ Locard, 1889, Contr. Faune Malacol. Franc. 13: 143 [37].
‘Bourguignat’ Locard, 1893, Coqu. Eau Douce Saumâtre France: 170.
Type(s): LECTOTYPE MNHN_22355, La Marne à Lagny (Seine-et-Marne).

Unio matronicus — Locard (1882)
Unio (Bataviana) matronicus — Servain (1882), Servain (1888)
Unio (Groupe de l’U. Batavus) matronicus — Locard (1882)
Unio (Batavusiana) matronicus — Locard (1889), Germain (1904)
Unio batavus var. matronicus — Westerlund (1890)
Unio (Groupe de l’U. batavus) matronicus — Locard (1893)
Unio batavus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio minutus Locard, 1882
‘Ray’ Locard, 1882, Prodr. Malac. Franç. 1: 290. [nomen nudum]

Unio minutus — Locard (1882)
Unio (Groupe de l’U. fusculus) minutus — Locard (1882)
Unio (Amniciana) minutulus — Servain (1888)
Unio (Amnicusiana) minutulus — Locard (1889)
Unio mancus var. amnicus — Germain (1931)
Unio crassus batavus — Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio oxyrhynchus Locard, 1882
Bourguignat in Locard, 1882, Prodr. Malac. Franç. 1: 285, 355.
‘Brevière’ Locard, 1889, Contr. Faune Malacol. Franc. 13: 149 [43].
‘Brevière’ Locard, 1893, Coqu. Eau Douce Saumâtre France: 180.
(homonym of Unio oxyrhynchus Martens, 1861)
Type(s): LECTOTYPE MNHN_22438, La Canne, dans la Nièvre.

Unio oxyrhynchus — Locard (1882)
Unio (Groupe de l’U. crassus) oxyrhynchus — Locard (1882)
Unio (Ateriana) oxyrhynchus — Locard (1889)
Unio (Groupe de l’U. ater) oxyrhynchus — Locard (1893)
Unio consentaneus var. oxyrhynchus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio pilloti Locard, 1882
Bourguignat in Locard, 1882, Prodr. Malac. Franç. 1: 291, 360.
‘Bourguignat’ Locard, 1889, Contr. Faune Malacol. Franc. 13: 128 [22].
‘Bourguignat’ Locard, 1893, Coqu. Eau Douce Saumâtre France: 157.
Type(s): IDIOTYPE MNHN_22460, Passavant (Haute-Saône).

Unio pilloti — Locard (1882), Kobelt & Haas (1910)
Unio (Groupe de l’U. mancus) pilloti — Locard (1882), Locard (1893)
Unio (Mancusiana) pilloti — Locard (1889)
Unio mancus var. pilloti — Westerlund (1890)
Unio mancus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio mancus mancus — Prie (2012)
Unio crassus — Lopes-Lima et al. (2024)

Unio rayi Locard, 1882
Bourguignat in Locard, 1882, Prodr. Malac. Franç. 1: 291, 360.
‘Bourguignat’ Locard, 1889, Contr. Faune Malacol. Franc. 13: 127 [21].
‘Bourguignat’ Locard, 1893, Coqu. Eau Douce Saumâtre France: 155.
Type(s): LECTOTYPE MNHN_22483, Gigny (Jura).

Unio rayi — Locard (1882), Servain (1885), Westerlund (1890), Kobelt & Haas (1910)
Unio (Groupe de l’U. mancus) rayi — Locard (1882)
Unio (Nanusiana) rayi — Locard (1889)
Unio (Groupe de l’U. nanus) rayi — Locard (1893)
Unio (Groupe des Ninusiana) rayi — Caziot (1907)
Unio mancus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio mancus mancus — Prie (2012)
Unio crassus — Lopes-Lima et al. (2024)

Unio sandrii Locard, 1882
Locard, 1882, Prodr. Malac. Franç. 1: 292.
(misidentified reference to Unio sandrii Rossmässler, 1844)

Unio sandrii — Locard (1882)
Unio (Sandriana) sandriopsis — Locard (1889)

Unio septentrionalis Locard, 1882
Bourguignat in Locard, 1882, Prodr. Malac. Franç. 1: 284, 354.
‘Bourguignat’ Locard, 1889, Contr. Faune Malacol. Franc. 13: 149 [43].
‘Bourguignat’ Locard, 1893, Coqu. Eau Douce Saumâtre France: 181, fig. 194.
Type(s): IDIOTYPE MNHN_22528, le Mandrezey, à Saulcy-sur-Meurthe (Vosges).

Unio septentrionalis — Locard (1882)
Unio (Groupe de l’U. ater) septentrionalis — Locard (1882), Locard (1893)
Unio (Ateriana) septentrionalis — Locard (1889)
Unio consentaneus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)

Unio sequanicus Locard, 1882
Bourguignat in Locard, 1882, Prodr. Malac. Franç. 1: 289, 358.
‘Coutagne’ Locard, 1889, Contr. Faune Malacol. Franc. 13: 142 [36].
‘Coutagne’ Locard, 1893, Coqu. Eau Douce Saumâtre France: 169, fig. 183.
Type(s): LECTOTYPE MNHN_22529, La Marne à Lagny (Seine-et-Marne).

Unio sequanicus — Locard (1882)
Unio (Groupe de l’U. Batavus) sequanicus — Locard (1882)
Unio (Bataviana) sequanicus — Servain (1888)
Unio (Batavusiana) sequanicus — Locard (1889)
Unio batavus var. sequanicus — Westerlund (1890)
Unio (Groupe de l’U. batavus) sequanicus — Locard (1893)
Unio batavus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio batavellus Locard, 1885
‘Letourneux’ Locard, 1885, Bull. Soc. Sci. Nat. Rouen 21: 11.
‘Letourneux’ Locard, 1889, Contr. Faune Malacol. Franc. 13: 144 [38].
‘Letourneux’ Locard, 1893, Coqu. Eau Douce Saumâtre France: 170, fig. 184.
Type(s): LECTOTYPE MNHN_22127, La Bouille, près de Rouen (Seine-Inf.).

Unio batavellus — Locard (1885), Westerlund (1890), Rochebrune (1903), Kobelt & Haas (1910)
Unio (Bataviana) batavellus — Servain (1888)
Unio (Batavusiana) batavellus — Locard (1889), Germain (1904)
Unio (Groupe de l’U. batavus) batavellus — Locard (1893)
Unio batavus var. baraceus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio hueti Kobelt, 1886
‘Bourg.’ Kobelt, 1886, Icon. Land- und Süssw.-Moll. 2: 22, pl. 41, fig. 255.
(misidentified reference to Unio hueti Bourguignat, 1855)

Unio hueti — Kobelt (1886)
Unio (sec. Lymnium) pseudonymus — Simpson (1900), Simpson (1914)

Unio neocomiensis Kobelt, 1886
‘Drouët’ Kobelt, 1886, Icon. Land- und Süssw.-Moll. 2: 34.
(error for Unio neocomensis Drouët, 1881)

Unio neocomiensis — Kobelt (1886)

Unio sieversi Kobelt, 1886
Kobelt, 1886, Icon. Land- und Süssw.-Moll. 2: pl. 53, fig. 286.
(misidentified reference to Unio sieversi Drouët, 1881)

Unio sieversi — Kobelt (1886)
Unio sieversi var. kobelti — Westerlund (1890)

Unio baroni Servain, 1887
Servain, 1887, Bull. Soc. Malac. France 4: 254.

Unio baroni — Servain (1887), Westerlund (1890)
Unio (Penchinatiana) baroni — Locard (1889)
Unio (Groupe de l’U. fusculus) baroni — Locard (1893)

Unio carcasinus Sourbieu, 1887
Sourbieu, 1887, Bull. Soc. Malac. France 4: 235.

Unio carcasinus — Sourbieu (1887), Westerlund (1890), Baichère (1890)
Unio (Fusculusiana) carcasinus — Locard (1889)
Unio crassus var. piscinalis — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)

Unio baudoni Folin, 1888
Folin, 1888, Le Naturaliste 10: 273, 4 text figs.
(homonym of Unio baudoni Folin & Bérillon, 1874)

Unio baudoni — Folin (1888), Westerlund (1890), Kobelt (1902)
Unio crassus batavus — Haas (1940), Haas (1969)

Unio crassulus Drouët, 1888
Drouët, 1888, J. de Conch. 36: 106.
Drouët, 1889, Mém. Acad. Sci. Dijon 1: 66, pl. 2, fig. 5.
(homonym of Unio requienii crassulus Drouët, 1867)
Type(s): SYNTYPE MNHN_MP_0027, Saone.

Unio crassulus — Drouët (1888), Drouët (1889)
Unio (Ateriana) brevierei — Locard (1889)
Unio brevierei — Westerlund (1890)
Unio consentaneus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio dilophius Servain, 1888
‘Bourguignat’ Servain, 1888, Bull. Soc. Malac. France 5: 316, footnote. [nomen nudum]
Locard, 1889, Contr. Faune Malacol. Franc. 13: 136 [30], footnote. [nomen nudum]

Unio dilophius — Servain (1888)
Unio (Amniciana) dilophius — Servain (1888)
Unio (Amnicusiana) dilophius — Locard (1889)

Unio hamburgiensis Servain, 1888
Servain, 1888, Bull. Soc. Malac. France 5: 315.

Unio hamburgiensis — Servain (1888), Westerlund (1890)
Unio (Amniciana) hamburgiensis — Servain (1888)
Unio (Crassiana) hamburgiensis — Locard (1889)
Unio (Groupe de l’U. feliciani) hamburgiensis — Locard (1893)
Unio crassus — Germain (1931), Lopes-Lima et al. (2024)
Unio crassus crassus — Haas (1940), Haas (1969)
Unio (Crassiana) crassus — Vinarski & Kantor (2016)

Unio minutulus Servain, 1888
‘Ray’ Locard, 1882, Prodr. Malac. Franç. 1: 290 [as ‘Unio minutus’]. [nomen nudum]
‘Ray’ Servain, 1888, Bull. Soc. Malac. France 5: 316.

Unio minutulus — Servain (1888), Westerlund (1890)
Unio (Amniciana) minutulus — Servain (1888)
Unio (Amnicusiana) minutulus — Locard (1889)
Unio (Groupe de l’U. amnicus) minutulus — Locard (1893)
Unio mancus var. amnicus — Germain (1931)
Unio crassus batavus — Haas (1940)

Unio vegesackensis Servain, 1888
Bourguignat in Servain, 1888, Bull. Soc. Malac. France 5: 317.

Unio vegesackensis — Servain (1888), Westerlund (1890)
Unio (Bataviana) vegesackensis — Servain (1888)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio visurgisinus Servain, 1888
Servain, 1888, Bull. Soc. Malac. France 5: 316.

Unio visurgisinus — Servain (1888), Westerlund (1890)
Unio (Bataviana) visurgisinus — Servain (1888)
Unio (Batavusiana) visurgicus — Locard (1889), Germain (1904)
Unio (Groupe de l’U. batavus) visurgicus — Locard (1893)
Unio batavus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio adonus Locard, 1889
‘Servain’ Locard, 1889, Contr. Faune Malacol. Franc. 13: 146 [40], 211 [105].
‘Servain’ Locard, 1893, Coqu. Eau Douce Saumâtre France: 175, fig. 188.
Type(s): LECTOTYPE MNHN_22085, La Loire a Basse Indre (Loire-Inf.).

Unio adonus — Locard (1889), Westerlund (1890)
Unio (Pisaniana) adonus — Locard (1889)
Unio (Groupe de l’U. adonus) adonus — Locard (1893)
Unio (Lemotheuxiana) adonus — Germain (1904)
Unio batavus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio ampulodon Locard, 1889
‘Coutagne Mss. teste Bourguignat’ Locard, 1889, Contr. Faune Malacol. Franc. 13: 128 [22]. [nomen nudum]
(intro'd as syn of Unio dubisianopsis Locard, 1889)

Unio ampulodon — Locard (1889)
Unio (Melasiana) dubisianopsis — Locard (1889)
Unio mancus var. amnicus — Germain (1931)

Unio andeliacus Locard, 1889
Bourguignat in Locard, 1889, Contr. Faune Malacol. Franc. 13: 138 [32], 196 [90].
Bourguignat in Locard, 1893, Coqu. Eau Douce Saumâtre France: 165.
Type(s): LECTOTYPE MNHN_22101, le Gouet, à Saint-Brieue (Côtes-du-Nord).

Unio andeliacus — Locard (1889), Westerlund (1890)
Unio (Nubilusiana) andeliacus — Locard (1889)
Unio (Groupe de l’U. locardianus) andeliacus — Locard (1893)
Unio mancus var. elongatulus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio aturicus Locard, 1889
Locard, 1889, Contr. Faune Malacol. Franc. 13: 129 [23], 188 [82].
Locard, 1893, Coqu. Eau Douce Saumâtre France: 157.
Type(s): HOLOTYPE (MONOTYPY) MNHN_22118, Bahus (Landes).

Unio aturicus — Locard (1889), Westerlund (1890)
Unio (Mancusiana) aturicus — Locard (1889)
Unio (Groupe de l’U. mancus) aturicus — Locard (1893)
Unio mancus var. amnicus form riparius — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)

Unio balbignyanus Locard, 1889
Locard, 1889, Contr. Faune Malacol. Franc. 13: 147 [41], 217 [111].
Locard, 1893, Coqu. Eau Douce Saumâtre France: 178.
Type(s): LECTOTYPE MNHN_22122, La Loire, à Balbigny (Loire).

Unio balbignyanus — Locard (1889), Westerlund (1890)
Unio (Ateriana) balbignyanus — Locard (1889)
Unio (Groupe de l’U. ater) balbignyanus — Locard (1893)
Unio consentaneus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio besnardianus Locard, 1889
Servain, 1888, Bull. Soc. Malac. France 5: 317. [nomen nudum]
Bourguignat in Locard, 1889, Contr. Faune Malacol. Franc. 13: 141 [35], 206 [100].
‘Servain’ Locard, 1893, Coqu. Eau Douce Saumâtre France: 172, fig. 186 [as ‘besnardinus’].
(renamed Unio besnardi Germain, 1904)
Type(s): LECTOTYPE MNHN_22133, La Loire, à Saint-Gemmes (Maine-et-Loire).

Unio besnardianus — Locard (1889), Westerlund (1890)
Unio (Bataviana) besnardianus — Servain (1888)
Unio (Batavusiana) besnardianus — Locard (1889)
Unio (Groupe de l’U. batavus) besnardinus — Locard (1893)
Unio (Batavusiana) besnardi — Germain (1904)
Unio batavus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio bouchardi Locard, 1889
‘Bourguignat’ Locard, 1889, Contr. Faune Malacol. Franc. 13: 149 [43].
‘Bourguignat’ Locard, 1893, Coqu. Eau Douce Saumâtre France: 180.
(new name for Unio arcuata Bouchard-Chantereaux, 1838)
Type(s): LECTOTYPE FIGURED SPECIMEN, ditches of Saint-Omer, fed by the Aa.

Unio bouchardi — Locard (1889)
Unio (Ateriana) bouchardi — Locard (1889)
Unio arcuatus — Westerlund (1890)
Unio (Groupe de l’U. ater) bouchardi — Locard (1893)
Unio consentaneus var. moquini — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio mancus moquinianus — Prie (2012)
Unio crassus — Lopes-Lima et al. (2024)

Unio caroliensis Locard, 1889
Bourguignat in Locard, 1889, Contr. Faune Malacol. Franc. 13: 159 [53], 237 [131].
‘F. Pacome’ Locard, 1893, Coqu. Eau Douce Saumâtre France: 172.
Type(s): LECTOTYPE MNHN_22167, Charolles (Saône-et-Loire).

Unio caroliensis — Locard (1889), Westerlund (1890)
Unio (Mariana) caroliensis — Locard (1889)
Unio (Groupe de l’U. batavus) caroliensis — Locard (1893)
Unio batavus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio catalaunicus Locard, 1889
Bourguignat in Locard, 1889, Contr. Faune Malacol. Franc. 13: 133 [27], 192 [86].
‘Coutagne’ Locard, 1893, Coqu. Eau Douce Saumâtre France: 160.
Type(s): LECTOTYPE MNHN_22171, La Marne, à Charenton (Seine).

Unio catalaunicus — Locard (1889), Westerlund (1890)
Unio (Fusculusiana) catalaunicus — Locard (1889)
Unio (Groupe de l’U. fusculus) catalaunicus — Locard (1893)
Unio crassus var. piscinalis — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio caumonti Locard, 1889
Bourguignat in Locard, 1889, Contr. Faune Malacol. Franc. 13: 145 [39], 207 [101].
Bourguignat in Locard, 1893, Coqu. Eau Douce Saumâtre France: 171.

Unio caumonti — Locard (1889), Westerlund (1890)
Unio (Batavusiana) caumonti — Locard (1889)
Unio (Groupe de l’U. batavus) caumonti — Locard (1893)
Unio batavus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio diptychus Locard, 1889
Bourguignat in Locard, 1889, Contr. Faune Malacol. Franc. 13: 143 [37], 201 [95].
‘Surault’ Locard, 1893, Coqu. Eau Douce Saumâtre France: 169.
Type(s): POSSIBLE SYNTYPE MNHN_22192, La Loire à Ingrandes.

Unio diptychus — Locard (1889), Westerlund (1890)
Unio (Batavusiana) diptychus — Locard (1889), Germain (1904)
Unio (Groupe de l’U. batavus) diptychus — Locard (1893)
Unio batavus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio dubisianopsis Locard, 1889
Locard, 1889, Contr. Faune Malacol. Franc. 13: 128 [22].
Locard, 1893, Coqu. Eau Douce Saumâtre France: 155.
(error for Unio dubisiopsis Locard, 1882)

Unio dubisianopsis — Locard (1889)

Unio financei Locard, 1889
Locard, 1889, Contr. Faune Malacol. Franc. 13: 143 [37], 205 [99].
Locard, 1893, Coqu. Eau Douce Saumâtre France: 171.
Type(s): IDIOTYPE MNHN_22217, La Marne à Châlons (Marne).

Unio financei — Locard (1889), Westerlund (1890)
Unio (Batavusiana) financei — Locard (1889)
Unio (Groupe de l’U. batavus) financei — Locard (1893)
Unio batavus — Germain (1931)

Unio hattemani Locard, 1889
Bourguignat in Locard, 1889, Contr. Faune Malacol. Franc. 13: 146 [40], 212 [106].
‘Bourguignat’ Locard, 1893, Coqu. Eau Douce Saumâtre France: 176.
Type(s): SYNTYPE MNHN_22254, La Loire, à Basse-Indre.

Unio hattemani — Locard (1889), Westerlund (1890)
Unio (Pisaniana) hattemani — Locard (1889)
Unio (Groupe de l’U. adonus) hattemani — Locard (1893)
Unio (Lemotheuxiana) hattemanni — Germain (1904)
Unio batavus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio ignariformis Locard, 1889
‘Dup.’ Drouët, 1857, Étud. Naïades. France, Unio: 100, pl. 9, fig. 1 [as ‘U. platyrhynchoideus’].
Bourguignat in Locard, 1889, Contr. Faune Malacol. Franc. 13: 147 [41], 214 [108].
‘Bourguignat’ Locard, 1893, Coqu. Eau Douce Saumâtre France: 177.

Unio ignariformis — Locard (1889), Westerlund (1890)
Unio (Ateriana) ignariformis — Locard (1889)
Unio (Groupe de l’U. ater) ignariformis — Locard (1893)
Unio consentaneus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)

Unio ingrandiensis Locard, 1889
Bourguignat in Locard, 1889, Contr. Faune Malacol. Franc. 13: 143 [37], 202 [96].
‘Sarault’ Locard, 1893, Coqu. Eau Douce Saumâtre France: 173.
Type(s): POSSIBLE SYNTYPE MNHN_22282, Ingrandes (Maine-et-Loire).

Unio ingrandiensis — Locard (1889), Westerlund (1890)
Unio (Batavusiana) ingrandiensis — Locard (1889), Germain (1904)
Unio (Groupe de l’U. batavus) ingrandiensis — Locard (1893)
Unio batavus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio lemotheuxi Locard, 1889
Bourguignat in Locard, 1889, Contr. Faune Malacol. Franc. 13: 145 [39], 208 [102].
‘Servain’ Locard, 1893, Coqu. Eau Douce Saumâtre France: 175, fig. 189.
Type(s): LECTOTYPE MNHN_22306, le Reyran (Var).

Unio lemotheuxi — Locard (1889), Westerlund (1890)
Unio (Lemotheuxiana) lemotheuxi — Locard (1889), Germain (1904), Caziot (1910)
Unio (Groupe de l’U. adonus) lemotheuxi — Locard (1893)
Unio mancus var. elongatulus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio mariae Locard, 1889
Bourguignat in Locard, 1889, Contr. Faune Malacol. Franc. 13: 159 [53], 236 [130] [as ‘mariæ’].
‘F. Pacome’ Locard, 1893, Coqu. Eau Douce Saumâtre France: 172 [as ‘mariæ’].
Type(s): IDIOTYPE MNHN_22342, La Loire à Nantes (Loire-Infereur).

Unio mariae — Locard (1889)
Unio (Mariana) mariæ — Locard (1889)
Unio mariæ — Westerlund (1890)
Unio (Groupe de l’U. batavus) mariæ — Locard (1893)
Unio batavus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio materniacus Locard, 1889
Locard, 1889, Contr. Faune Malacol. Franc. 13: 143 [37], 203 [97].
Locard, 1893, Coqu. Eau Douce Saumâtre France: 168, fig. 182.
Type(s): LECTOTYPE MNHN_22347, la Somme à Abbeville.

Unio materniacus — Locard (1889), Westerlund (1890), Rochebrune (1903)
Unio (Batavusiana) materniacus — Locard (1889)
Unio (Groupe de l’U. batavus) materniacus — Locard (1893)
Unio batavus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio melantatus Locard, 1889
Locard, 1889, Contr. Faune Malacol. Franc. 13: 147 [41], 216 [110].
Locard, 1893, Coqu. Eau Douce Saumâtre France: 177.
Type(s): LECTOTYPE MNHN_22360, La Loire à Balbigny (Loire).

Unio melantatus — Locard (1889), Westerlund (1890)
Unio (Ateriana) melantatus — Locard (1889)
Unio (Groupe de l’U. ater) melantatus — Locard (1893)
Unio consentaneus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio necomensis Locard, 1889
‘Drouët’ of Authors.
Locard, 1889, Contr. Faune Malacol. Franc. 13: 132 [26].
Locard, 1893, Coqu. Eau Douce Saumâtre France: 159.
Germain, 1931, Faune de France 22: 725.
(error for Unio neocomensis Drouët, 1881)

Unio necomensis — Locard (1889)
Unio mancus var. macrorhynchus — Germain (1931)

Unio nicolloni Locard, 1889
Locard, 1889, Contr. Faune Malacol. Franc. 13: 141 [35], 200 [94].
Locard, 1893, Coqu. Eau Douce Saumâtre France: 173.
Type(s): HOLOTYPE (MONOTYPY) MNHN_22404, Nantes.

Unio nicolloni — Locard (1889), Westerlund (1890)
Unio (Sperchinusiana) nicolloni — Locard (1889)
Unio (Groupe de l’U. batavus) nicolloni — Locard (1893)
Unio (Batavusiana) nicolloni — Germain (1904)
Unio batavus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio occidentalis Locard, 1889
‘Bourguignat’ Locard, 1889, Contr. Faune Malacol. Franc. 13: 149 [43], 220 [114].
‘Bourguignat’ Locard, 1893, Coqu. Eau Douce Saumâtre France: 180.
‘Bourguignat’ Locard, 1899, Conchyliol. Portug.: 220.
(homonym of Unio occidentalis Conrad, 1836)
Type(s): LECTOTYPE MNHN_22424, Balbigny (Loire).

Unio occidentalis — Locard (1889), Westerlund (1890)
Unio (Ateriana) occidentalis — Locard (1889)
Unio (Groupe de l’U. ater) occidentalis — Locard (1893)
Unio (groupe de l’U. ater) occidentalis — Locard (1899)
Unio consentaneus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio seneauxi Locard, 1889
‘Linn.’ Draparnaud, 1806, Hist. Moll. France: 131, pl. 11, figs. 3 [as ‘Unio pictorum var. B’].
Bourguignat in Locard, 1889, Contr. Faune Malacol. Franc. 13: 145 [39], 208 [102].
‘Bourguignat’ Locard, 1893, Coqu. Eau Douce Saumâtre France: 174, fig. 187 [as ‘senauxi’].
Type(s): LECTOTYPE MNHN_22523, La Loire à Orléans.

Unio seneauxi — Locard (1889)
Unio (Batavusiana) seneauxi — Locard (1889), Germain (1904)
Unio caumonti — Westerlund (1890)
Unio (Groupe de l’U. batavus) senauxi — Locard (1893)
Unio batavus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio sperchinus Locard, 1889
‘Thièse’ Locard, 1889, Contr. Faune Malacol. Franc. 13: 141 [35], footnote. [nomen nudum]

Unio sperchinus — Locard (1889)
Unio (Sperchinusiana) sperchinus — Locard (1889)

Unio subamnicus Locard, 1889
Locard, 1889, Contr. Faune Malacol. Franc. 13: 137 [31], 194 [88].
Locard, 1893, Coqu. Eau Douce Saumâtre France: 163.
Type(s): SYNTYPE MNHN_22559, La Saône, à Auxonne (Côte-d’Or).

Unio subamnicus — Locard (1889), Westerlund (1890), Kobelt & Haas (1910)
Unio (Amnicusiana) subamnicus — Locard (1889)
Unio (Groupe de l’U. amnicus) subamnicus — Locard (1893)
Unio mancus var. amnicus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio surraulti Locard, 1889
Bourguignat in Locard, 1889, Contr. Faune Malacol. Franc. 13: 144 [38], 204 [98].
‘Servain’ Locard, 1893, Coqu. Eau Douce Saumâtre France: 174 [as ‘suraulti’].
Type(s): SYNTYPE MNHN_22569, La Loire à Ingrandes.

Unio surraulti — Locard (1889), Westerlund (1890)
Unio (Batavusiana) surraulti — Locard (1889), Germain (1904)
Unio (Groupe de l’U. batavus) suraulti — Locard (1893)
Unio batavus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio visurgicus Locard, 1889
‘Servain’ of Authors.
Locard, 1889, Contr. Faune Malacol. Franc. 13: 141 [35].
Locard, 1893, Coqu. Eau Douce Saumâtre France: 173.
Germain, 1904, Bull. Soc. Sci. Nat. Ouest France 4: 262.
(error for Unio visurgisinus Servain, 1888)

Unio visurgicus — Locard (1889)
Unio visurgisinus — Westerlund (1890)
Unio batavus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio pilloti var. minor Locard, 1889
Locard, 1889, Contr. Faune Malacol. Franc. 13: 129 [23]. [nomen nudum]
(intro'd as intra-sp. of Unio pilloti Locard, 1882)

Unio pilloti var. minor — Locard (1889)
Unio (Mancusiana) pilloti — Locard (1889)

Unio suborbicularis var. minor Locard, 1889
Locard, 1889, Contr. Faune Malacol. Franc. 13: 127 [21]. [nomen nudum]
Westerlund, 1890, Fauna Paläarct. Binnenconch. 2(7): 84. [nomen nudum]
(intro'd as intra-sp. of Unio suborbicularis Drouët, 1888)

Unio suborbicularis var. minor — Locard (1889)
Unio (Nanusiana) lagnysicus — Locard (1889)
Unio lagnysicus — Westerlund (1890)

Unio subrobustus var. curta Locard, 1889
Locard, 1889, Contr. Faune Malacol. Franc. 13: 135 [29].
(homonym of Unio curtus Lea, 1859; intro'd as intra-sp. of Unio subrobustus Servain, 1885)

Unio subrobustus var. curta — Locard (1889)
Unio (Socardiana) subrobustus var. curta — Locard (1889)

Unio courtilieri Locard, 1889
‘Hattemann’ of Authors.
Locard, 1889, Contr. Faune Malacol. Franc. 13: 132 [26].
‘Hattemann’ Germain, 1904, Bull. Soc. Sci. Nat. Ouest France 4: 256 [18].
‘Hattmann’ Germain, 1931, Faune de France 22: 724, 727.
Lopes-Lima et al., 2024, MPE 195: suppl. append. 2, p. 3.
(error for Unio courtillieri Hattemann, 1859)

Unio courtilieri — Locard (1889)

Unio sieversi var. kobelti Westerlund, 1890
Kobelt, 1886, Icon. Land- und Süssw.-Moll. 2: pl. 53, fig. 286 [as ‘U. sieversi’].
Westerlund, 1890, Fauna Paläarct. Binnenconch. 2(7): 151.
(intro'd as intra-sp. of Unio sieversi Drouët, 1881; homonym of Unio kobelti Neumayr, 1899)

Unio sieversi var. kobelti — Westerlund (1890)

Unio ampullaceus Locard, 1893
Locard, 1893, Coqu. Eau Douce Saumâtre France: 174.
(homonym of Unio ampullaceus Lea, 1866)
Type(s): LECTOTYPE MNHN_22098, Le Mans (Sarthe).

Unio ampullaceus — Locard (1893)
Unio (Groupe de l’U. batavus) ampullaceus — Locard (1893)
Unio (Batavusiana) ampullaceus — Germain (1904)
Unio batavus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio besnardinus Locard, 1893
‘Servain’ Locard, 1893, Coqu. Eau Douce Saumâtre France: 172.
(error for Unio besnardianus Locard, 1889)

Unio besnardinus — Locard (1893)
Unio (Batavusiana) besnardi — Germain (1904)

Unio chardoni Locard, 1893
‘Bourguignat’ Locard, 1893, Coqu. Eau Douce Saumâtre France: 173.

Unio chardoni — Locard (1893)
Unio (Groupe de l’U. batavus) chardoni — Locard (1893)
Unio batavus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio gliscerus Locard, 1893
Locard, 1893, Coqu. Eau Douce Saumâtre France: 168.
Type(s): LECTOTYPE MNHN_22241, La Loire à Nantes (Loire-Infereur).

Unio gliscerus — Locard (1893)
Unio (Groupe de l’U. batavus) gliscerus — Locard (1893)
Unio (Batavusiana) gliscerus — Germain (1904)
Unio batavus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio juranus Locard, 1893
‘Locard’ of Authors.
Locard, 1893, Coqu. Eau Douce Saumâtre France: 157.
Germain, 1931, Faune de France 22: 725.
(error for Unio jurianus Locard, 1889)

Unio juranus — Locard (1893)

Unio lamboltei Locard, 1893
‘Malzine’ of Authors.
Locard, 1893, Coqu. Eau Douce Saumâtre France: 177.
Germain, 1931, Faune de France 22: 732.
Haas, 1940, Zool. Ser. Field Mus. Nat. Hist. 24: 130.
(error for Unio lambottei Malzine, 1867)

Unio lamboltei — Locard (1893)

Unio lancelevei Locard, 1893
Locard, 1893, Bull. Soc. Sci. Nat. Elbeuf 12: 1.
Locard, 1893, Coqu. Eau Douce Saumâtre France: 170.
Type(s): SYNTYPE MNHN_22305, le Seine au nord d. Rouen.

Unio lancelevei — Locard (1893)
Unio (Groupe de l’U. batavus) lancelevei — Locard (1893)
Unio batavus var. baraceus — Germain (1931)
Unio crassus batavus — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio senauxi Locard, 1893
‘Bourguignat’ Locard, 1893, Coqu. Eau Douce Saumâtre France: 174.
(error for Unio seneauxi Locard, 1889)

Unio senauxi — Locard (1893)

Unio suraulti Locard, 1893
‘Servain in Locard’ Locard, 1893, Coqu. Eau Douce Saumâtre France: 174.
(error for Unio surraulti Locard, 1889)

Unio suraulti — Locard (1893)

Unio lepidulus Drouët, 1898
Drouët, 1898, Mém. Acad. Sci. Dijon ??: 60, pl. 1, fig. 7.

Unio lepidulus — Drouët (1898)
Unio mancus var. elongatulus — Germain (1931)

Unio batavus var. major Drouët, 1898
Drouët, 1898, Mém. Acad. Sci. Dijon ??: 61.
(intro'd as intra-sp. of Mya batava Maton & Rackett, 1807; homonym of Unio (Anodontes) sirionos var. major d’Orbigny, 1835)

Unio batavus var. major — Drouët (1898)
Unio batavus var. baraceus — Germain (1931)

Unio ellipsopsis Locard, 1899
Bourguignat in Locard, 1889, Contr. Faune Malacol. Franc. 13: 140 [34]. [nomen nudum]
Bourguignat in Locard, 1899, Conchyliol. Portug.: 218.

Unio ellipsopsis — Locard (1899)
Unio (Ellipsopsisiana) ellipsopsis — Locard (1889)
Unio (groupe de l’U. ellipsopsis) ellipsopsis — Locard (1899)

Unio musicus Simpson, 1900
‘Spengler’ Simpson, 1900, Proc. U.S. Nat. Mus. 22: 744.
(error for Unio musivus Spengler, 1793)

Unio musicus — Simpson (1900)
Unio (sec. Lymnium) batavus — Simpson (1914)

Unio besnardi Germain, 1904
Germain, 1904, Bull. Soc. Sci. Nat. Ouest France 4: 261 [23].
(new name for Unio besnardianus Locard, 1889)

Unio besnardi — Germain (1904)
Unio (Batavusiana) besnardi — Germain (1904)

Unio hattemanni Germain, 1904
‘Bourguignat’ Germain, 1904, Bull. Soc. Sci. Nat. Ouest France 4: 265 [27].
(error for Unio hattemani Locard, 1889)

Unio hattemanni — Germain (1904)

Unio batavellus var. perconvexus Germain, 1904
Germain, 1904, Bull. Soc. Sci. Nat. Ouest France 4: 260 [22].
(intro'd as intra-sp. of Unio batavellus Locard, 1885)

Unio batavellus var. perconvexus — Germain (1904)
Unio (Batavusiana) batavellus perconvexus — Germain (1904)

Unio batavus var. radiata Germain, 1904
Germain, 1904, Bull. Soc. Sci. Nat. Ouest France 4: 257 [19].
(intro'd as intra-sp. of Mya batava Maton & Rackett, 1807; homonym of Unio (Elliptio) viridis var. radiata Rafinesque, 1820)

Unio batavus var. radiata — Germain (1904)
Unio (Batavusiana) batavus radiata — Germain (1904)

Unio ligericus var. subsinuatus Germain, 1904
Germain, 1904, Bull. Soc. Sci. Nat. Ouest France 4: 259 [21].
(intro'd as intra-sp. of Unio ligericus Locard, 1882)

Unio ligericus var. subsinuatus — Germain (1904)
Unio (Batavusiana) ligericus subsinuatus — Germain (1904)

Unio pseudolittoralis Riemenschneider, 1907
‘Clessin’ of Authors.
Riemenschneider, 1907, Nachr. Bl. Dtsch. Malak. Ges. 39: 150.
Vinarski & Kantor, 2016, Analyt. Cat. Fr. Brack. Moll. Russia: 25.
(error for Unio pseudolitoralis Clessin, 1877)

Unio pseudolittoralis — Riemenschneider (1907)

Unio pseudolittoralis [sic] var. curonicus Riemenschneider, 1907
Riemenschneider, 1907, Nachr. Bl. Dtsch. Malak. Ges. 39: 150.
(intro'd as intra-sp. of Unio pseudolitoralis Clessin, 1877)

Unio pseudolittoralis var. curonicus — Riemenschneider (1907)
Unio crassus crassus — Haas (1940), Haas (1969)
Unio (Crassiana) crassus — Vinarski & Kantor (2016)
Unio crassus — Lopes-Lima et al. (2024)

Unio consentaneus marisensis Kobelt & Haas, 1910
Kobelt & Haas, 1910, Icon. Land- und Süssw.-Moll. 17: pl. 479 [caption].
(incorrect original spelling of Unio consentaneus marisaensis Kobelt & Haas, 1910)

Unio consentaneus marisensis — Kobelt & Haas (1910)

Unio crassus var. okae Kobelt & Haas, 1910
Kobelt & Haas, 1910, Icon. Land- und Süssw.-Moll. 17: 58, index, pl. 480, fig. 2567.
(intro'd as intra-sp. of Unio crassus Retzius, 1788)

Unio crassus var. okae — Kobelt & Haas (1910)
Unio (s.s.) crassus var. okae — Zhadin (1938)
Unio crassus gontieri — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio crassus heimburgi Haas, 1911
Haas, 1911, Verh. Naturhist. Ver. Preuß. Rheinl. 68: 527, pl. 6, fig. 20.
(intro'd as intra-sp. of Unio crassus Retzius, 1788)

Unio crassus heimburgi — Haas (1911)
Unio crassus crassus — Haas (1940), Haas (1969)
Unio (Crassiana) crassus — Vinarski & Kantor (2016)
Unio crassus — Lopes-Lima et al. (2024)

Unio kungurensis Kobelt, 1912
Kobelt, 1912, Icon. Land- und Süssw.-Moll. 18: 24, pl. 490, figs. 2622-2623.

Unio kungurensis — Kobelt (1912)
Unio (s.s.) crassus kungurensis — Zhadin (1938)
Unio crassus gontieri — Haas (1940), Haas (1969)
Unio (Crassiana) crassus — Vinarski & Kantor (2016)
Unio crassus — Lopes-Lima et al. (2024)

Unio roseni Kobelt, 1912
Kobelt, 1912, Icon. Land- und Süssw.-Moll. 18: 27, pl. 492, fig. 2627.

Unio roseni — Kobelt (1912)
Unio (s.s.) crassus var. roseni — Zhadin (1938)
Unio crassus gontieri — Haas (1940), Haas (1969)
Unio crassus — Lopes-Lima et al. (2024)

Unio kungurensis chlebnikowi Kobelt, 1912
Kobelt, 1912, Icon. Land- und Süssw.-Moll. 18: 26, pl. 491, fig. 2624.
(intro'd as intra-sp. of Unio kungurensis Kobelt, 1912; renamed Unio crassus var. chlebnikovi Zhadin, 1938)

Unio kungurensis chlebnikowi — Kobelt (1912)
Unio (s.s.) crassus var. chlebnikovi — Zhadin (1938)
Unio crassus gontieri — Haas (1940), Haas (1969)
Unio (Crassiana) crassus — Vinarski & Kantor (2016)
Unio crassus — Lopes-Lima et al. (2024)

Unio kungurensis sylwensis Kobelt, 1912
Kobelt, 1912, Icon. Land- und Süssw.-Moll. 18: 26, pl. 491, fig. 2625.
(intro'd as intra-sp. of Unio kungurensis Kobelt, 1912)

Unio kungurensis sylwensis — Kobelt (1912)
Unio (s.s.) crassus var. chlebnikovi — Zhadin (1938)
Unio crassus gontieri — Haas (1940), Haas (1969)
Unio (Crassiana) crassus — Vinarski & Kantor (2016)
Unio crassus — Lopes-Lima et al. (2024)

Unio crassus thuringiacus Israel, 1913
Israel, 1913, [in] Thüring. Lehrerverein Naturk.: 28, pl. 6, figs. 4-6.
(intro'd as intra-sp. of Unio crassus Retzius, 1788)

Unio crassus thuringiacus — Israel (1913)
Unio crassus crassus — Haas (1940), Haas (1969)
Unio (Crassiana) crassus — Vinarski & Kantor (2016)
Unio crassus — Lopes-Lima et al. (2024)

Unio crassus ornatus Polinksi, 1917
Polinksi, 1917, C. R. Soc. Sci. Varsovie 10: 904, pl. ??, fig. 2.
(intro'd as intra-sp. of Unio crassus Retzius, 1788; homonym of Unio ovatus var. ornatus Conrad, 1835)

Unio crassus ornatus — Polinksi (1917)
Unio crassus crassus — Haas (1940), Haas (1969)
Unio (Crassiana) crassus — Vinarski & Kantor (2016)
Unio crassus — Lopes-Lima et al. (2024)

Unio crassus polonicus Polinksi, 1917
Polinksi, 1917, C. R. Soc. Sci. Varsovie 10: 902, pl. ??, figs. 1, 3.
(intro'd as intra-sp. of Unio crassus Retzius, 1788)

Unio crassus polonicus — Polinksi (1917)
Unio crassus crassus — Haas (1940), Haas (1969)
Unio (Crassiana) crassus — Vinarski & Kantor (2016)
Unio crassus — Lopes-Lima et al. (2024)

Unio cantianus Kennard & Woodward, 1924
Kennard & Woodward, 1924, Proc. Malac. Soc. London 16: 136, text figs. 8-9.

Unio cantianus — Kennard & Woodward (1924)
Unio crassus batavus — Haas (1940), Haas (1969)

Unio carioliensis Germain, 1931
‘Pacôme’ Germain, 1931, Faune de France 22: 728.
(error for Unio caroliensis Locard, 1889)

Unio carioliensis — Germain (1931)

Unio nubilius Germain, 1931
‘Locard’ Germain, 1931, Faune de France 22: 727.
(error for Unio nubilus Locard, 1889)

Unio nubilius — Germain (1931)

Unio valleriacus Germain, 1931
‘Bourg. in Loc.’ Germain, 1931, Faune de France 22: 727.
(error for Unio valliericus Locard, 1889)

Unio valleriacus — Germain (1931)

Unio visursisinus Germain, 1931
‘Servain’ Germain, 1931, Faune de France 22: 728.
(error for Unio visurgisinus Servain, 1888)

Unio visursisinus — Germain (1931)

Unio crassus consentancus Zhadin, 1938
‘Ziegler’ Zhadin, 1938, Faune USSR Mollusques 4(1): 91.
(error for Unio consentaneus Rossmässler, 1836)

Unio crassus consentancus — Zhadin (1938)

Unio crassus var. chlebnikovi Zhadin, 1938
‘Kobelt’ Zhadin, 1938, Faune USSR Mollusques 4(1): 89.
(new name for Unio kungurensis chlebnikowi Kobelt, 1912)

Unio crassus var. chlebnikovi — Zhadin (1938)

Unio crassus var. ishmensis Zhadin, 1938
Zhadin, 1938, Faune USSR Mollusques 4(1): 160.
(intro'd as intra-sp. of Unio crassus Retzius, 1788)

Unio crassus var. ishmensis — Zhadin (1938)
Unio (s.s.) crassus var. ishmensis — Zhadin (1938)
Unio crassus crassus — Haas (1940), Haas (1969)
Unio (Crassiana) crassus — Vinarski & Kantor (2016)
Unio crassus — Lopes-Lima et al. (2024)

Unio crassus var. stepanovi Zhadin, 1938
‘Drouët’ Zhadin, 1938, Faune USSR Mollusques 4(1): 90.
(error for Unio stepanoffi Drouët, 1881)

Unio crassus var. stepanovi — Zhadin (1938)

Unio raddei var. kutaisiana Zhadin, 1938
‘Kobelt’ of Authors.
Zhadin, 1938, Faune USSR Mollusques 4(1): 94.
Zatravkin, 1987, Byull. Mosk. Obshch.. Ispyt. Prir. 92(4): 49.
(error for Unio raddei var. koutaisiana Kobelt, 1886)

Unio raddei var. kutaisiana — Zhadin (1938)

Unio hattmanni Haas, 1940
‘Bourguignat’ Haas, 1940, Zool. Ser. Field Mus. Nat. Hist. 24: 131.
(error for Unio hattemani Locard, 1889)

Unio hattmanni — Haas (1940)

Unio kungurensis sylvensis Haas, 1940
‘Kobelt’ Haas, 1940, Zool. Ser. Field Mus. Nat. Hist. 24: 132.
‘Kobelt’ Haas, 1969, Das Tierreich 88: 53.
Vinarski & Kantor, 2016, Analyt. Cat. Fr. Brack. Moll. Russia: 25.
(error for Unio kungurensis sylwensis Kobelt, 1912)

Unio kungurensis sylvensis — Haas (1940)

Unio raddei var. kutaisanus Haas, 1940
‘Kobelt’ Haas, 1940, Zool. Ser. Field Mus. Nat. Hist. 24: 131.
(error for Unio raddei var. koutaisiana Kobelt, 1886)

Unio raddei var. kutaisanus — Haas (1940)

Unio lancelevi Haas, 1969
‘Locard’ Haas, 1969, Das Tierreich 88: 51.
(error for Unio lancelevei Locard, 1893)

Unio lancelevi — Haas (1969)

Unio raddei var. koutaisanus Haas, 1969
‘Kobelt’ Haas, 1969, Das Tierreich 88: 53.
(error for Unio raddei var. koutaisiana Kobelt, 1886)

Unio raddei var. koutaisanus — Haas (1969)

Unio byzantius Zatravkin, 1983
‘Drouet’ Zatravkin, 1983, [in] I.M. Likharev, Molluscs: 40.
(error for Unio byzantinus Drouët, 1879)

Unio byzantius — Zatravkin (1983)

Unio crassus var. irgislaicus Zatravkin, 1987
‘Lindholm’ Zatravkin, 1987, Byull. Mosk. Obshch.. Ispyt. Prir. 92(4): 57.
(error for Unio ater var. irgizlaicus Lindholm, 1904)

Unio crassus var. irgislaicus — Zatravkin (1987)

Unio mingrelicus var. stevenianiformes Zatravkin, 1987
‘Shadin’ Zatravkin, 1987, Byull. Mosk. Obshch.. Ispyt. Prir. 92(4): 56.
(error for Unio mingrelicus var. stevenianiformis Zhadin, 1938)

Unio mingrelicus var. stevenianiformes — Zatravkin (1987)

Unio stevenianius Bába, 2000
‘Kyrnicky’ Bába, 2000, Boll. Malac. 36(5-8): 135.
(error for Unio stevenianus Drouët, 1881)

Unio stevenianius — Bába (2000)

Eolymnium byzantinium Graf, 2007
Graf, 2007, Proc. Acad. Nat. Sci. 156: 84.
(error for Unio byzantinus Drouët, 1879)

Eolymnium byzantinium — Graf (2007)
Unio (Eolymnium) byzantinus — Vinarski & Kantor (2016)

Taxonomic History (since 1900)

Margaritana crassa — Simpson (1900), Simpson (1914)
Unio (sec. Lymnium) batavus — Simpson (1900), Simpson (1914)
Unio musivus — Simpson (1900)
Quadrula (sec. Fusconaia) coccinea — Simpson (1900), Simpson (1914)
Unio (sec. Lymnium) littoralis — Simpson (1900), Simpson (1914)
Lampsilis (sec. Eurynia) radiatus — Simpson (1900)
Unio (sec. Lymnium) pseudonymus — Simpson (1900), Simpson (1914)
Unio musicus — Simpson (1900)
Unio baudoni — Kobelt (1902)
Unio carantoni — Rochebrune (1903)
Unio crassatellus — Rochebrune (1903)
Unio lagnysicus — Rochebrune (1903), Kobelt & Haas (1910)
Unio batavellus — Rochebrune (1903), Kobelt & Haas (1910)
Unio materniacus — Rochebrune (1903)
Unio (Batavusiana) seneauxi — Germain (1904)
Unio (Batavusiana) batavus — Germain (1904)
Unio (Mancusiana) mancus — Germain (1904)
Unio (Rhomboïdiana) nana — Germain (1904)
Unio (Batavusiana) besnardi — Germain (1904)
Unio (Batavusiana) batavus radiata — Germain (1904)
Unio (Fusculusiana) fusculus — Germain (1904)
Unio (Batavusiana) batavus courtilieri — Germain (1904)
Unio (Melasiana) melas — Germain (1904)
Unio (Batavusiana) ligericus — Germain (1904)
Unio (Ateriana) lambottei — Germain (1904)
Unio (Batavusiana) ryckholtii — Germain (1904)
Unio (Batavusiana) andegavensis — Germain (1904)
Unio (Batavusiana) arenarum — Germain (1904)
Unio (Ateriana) brevieri — Germain (1904)
Unio (Socardiana) feliciani — Germain (1904)
Unio (Ateriana) marcellinus — Germain (1904)
Unio (Batavusiana) matronicus — Germain (1904)
Unio (Batavusiana) batavellus — Germain (1904)
Unio (Batavusiana) visurgicus — Germain (1904)
Unio (Lemotheuxiana) adonus — Germain (1904)
Unio (Batavusiana) diptychus — Germain (1904)
Unio (Lemotheuxiana) hattemanni — Germain (1904)
Unio (Batavusiana) ingrandiensis — Germain (1904)
Unio (Lemotheuxiana) lemotheuxi — Germain (1904), Caziot (1910)
Unio (Batavusiana) nicolloni — Germain (1904)
Unio (Batavusiana) surraulti — Germain (1904)
Unio (Batavusiana) ampullaceus — Germain (1904)
Unio (Batavusiana) gliscerus — Germain (1904)
Unio besnardi — Germain (1904)
Unio hattemanni — Germain (1904)
Unio ater var. irgizlaicus — Lindholm (1904)
Unio batavellus var. perconvexus — Germain (1904)
Unio (Batavusiana) batavellus perconvexus — Germain (1904)
Unio batavus var. radiata — Germain (1904)
Unio ligericus var. subsinuatus — Germain (1904)
Unio (Batavusiana) ligericus subsinuatus — Germain (1904)
Unio (Groupe des Batavusiana) batavus — Caziot (1907)
Unio (Groupe des Mancusiana) subtilis — Caziot (1907)
Unio (Groupe des Ateriana) brevierei — Caziot (1907)
Unio (Groupe des Ninusiana) rayi — Caziot (1907)
Unio pseudolittoralis — Riemenschneider (1907)
Unio pseudolittoralis var. curonicus — Riemenschneider (1907)
Unio crassus — Kobelt & Haas (1910), Ortmann (1912), Ortmann (1914), Germain (1931), ..., Lopes-Lima et al. (2024), Wu et al. (2024), Zieritz et al. (2024), Dai et al. (2024)
Unio batavus — Kobelt & Haas (1910), Haas (1924), Germain (1931)
Unio consentaneus var. — Kobelt & Haas (1910)
Unio nana — Kobelt & Haas (1910)
Unio pilloti — Kobelt & Haas (1910)
Unio rayi — Kobelt & Haas (1910)
Unio subamnicus — Kobelt & Haas (1910)
Unio consentaneus marisensis — Kobelt & Haas (1910)
Unio crassus var. okae — Kobelt & Haas (1910)
Unio crassus heimburgi — Haas (1911)
Unio crassus musivus — Ortmann (1912), Bogatov (2022)
Unio kungurensis — Kobelt (1912)
Unio roseni — Kobelt (1912)
Unio kungurensis chlebnikowi — Kobelt (1912)
Unio kungurensis irenjensis — Kobelt (1912)
Unio kungurensis sylwensis — Kobelt (1912)
Unio crassus thuringiacus — Israel (1913)
Unio (sec. Lymnium) batavus var. requieni — Simpson (1914)
Lampsilis (Eurynia) radiata — Simpson (1914)
Unio crassus ornatus — Polinksi (1917)
Unio crassus polonicus — Polinksi (1917)
Unio cantianus — Kennard & Woodward (1924)
Quadrula (Obliquata) coccinea — Frierson (1927)
Unio consentaneus — Germain (1931)
Unio mancus — Germain (1931)
Unio mancus var. nanus — Germain (1931)
Unio crassus var. piscinalis — Germain (1931)
Unio batavus var. baraceus — Germain (1931)
Unio mancus var. amnicus — Germain (1931)
Unio mancus var. macrorhynchus — Germain (1931)
Unio mancus var. amnicus form riparius — Germain (1931)
Unio consentaneus var. moquini — Germain (1931)
Unio consentaneus var. oxyrhynchus — Germain (1931)
Unio mancus var. elongatulus — Germain (1931)
Unio carioliensis — Germain (1931)
Unio nubilius — Germain (1931)
Unio valleriacus — Germain (1931)
Unio visursisinus — Germain (1931)
Unio (s.s.) crassus — Zhadin (1938)
Unio (s.s.) ater — Zhadin (1938)
Unio (s.s.) crassus var. pseudolitoralis — Zhadin (1938)
Unio (s.s.) crassus var. okae — Zhadin (1938)
Unio (s.s.) crassus kungurensis — Zhadin (1938)
Unio (s.s.) crassus var. roseni — Zhadin (1938)
Unio (s.s.) crassus var. chlebnikovi — Zhadin (1938)
Unio crassus consentancus — Zhadin (1938)
Unio crassus var. chlebnikovi — Zhadin (1938)
Unio crassus var. ishmensis — Zhadin (1938)
Unio (s.s.) crassus var. ishmensis — Zhadin (1938)
Unio crassus var. stepanovi — Zhadin (1938)
Unio raddei var. kutaisiana — Zhadin (1938)
Unio crassus crassus — Haas (1940), Haas (1969), Fauna Europaea (2012), Prié & Puillandre (2014), Klishko et al. (2017), Lopes-Lima et al. (2017), Bogatov (2022)
Unio crassus batavus — Haas (1940), Haas (1969), Fauna Europaea (2012), Lopes-Lima et al. (2017)
Unio crassus cytherea — Haas (1940), Haas (1969)
Unio crassus gontieri — Haas (1940), Haas (1969), Graf (2007)
Unio hattmanni — Haas (1940)
Unio kungurensis sylvensis — Haas (1940)
Unio raddei var. kutaisanus — Haas (1940)
Unio crassus var. pseudolitoralis — Zhadin (1965)
Unio lancelevi — Haas (1969)
Unio raddei var. koutaisanus — Haas (1969)
Crassiana crassa crassa — Akromovski (1976), Kantor & Sysoev (2005), Kantor et al. (2010)
Batavusiana (Crassunio) crassa — Zatravkin (1983), Stadnichenko (1984), Zatravkin (1987)
Batavusiana (s.s.) musiva — Zatravkin (1983), Zatravkin (1987)
Batavusiana (s.s.) irenjensis — Zatravkin (1983), Stadnichenko (1984), Zatravkin (1987)
Unio byzantius — Zatravkin (1983)
Unio (Tumidusiana) tumidus var. minor — Stadnichenko (1984)
Batavusiana (s.s.) irgislaica — Zatravkin (1987)
Unio crassus var. irgislaicus — Zatravkin (1987)
Unio mingrelicus var. stevenianiformes — Zatravkin (1987)
Unio (Crassunio) crassus — Falkner (1994)
Batavusiana (Crassiana) crassus — Korniushin (1998)
Batavusiana (s.s.) musivus [??] — Korniushin (1998)
Pleurobema sintoxia — Parmalee & Bogan (1998), Williams et al. (2008)
Unio stevenianius — Bába (2000)
Unio (Crassiana) crassus crassus — Falkner et al. (2001), Falkner et al. (2002)
Unio (Crassiana) crassus courtillieri — Falkner et al. (2001), Falkner et al. (2002)
Crassiana (s.s.) crassa — Starobogatov et al. (2004)
Crassiana (Bataviana) musiva — Starobogatov et al. (2004)
Crassiana (Bataviana) cyprinorum — Starobogatov et al. (2004)
Crassiana (Bataviana) irgizlaica — Starobogatov et al. (2004)
Crassiana musiva musiva — Kantor & Sysoev (2005), Kantor et al. (2010)
Crassiana cyprinorum — Kantor & Sysoev (2005), Kantor et al. (2010)
Crassiana irgizlaica — Kantor & Sysoev (2005), Kantor et al. (2010)
Crassiana irenjensis — Kantor & Sysoev (2005), Kantor et al. (2010)
Eolymnium byzantinium — Graf (2007)
Unio mancus mancus — Prie (2012)
Unio mancus moquinianus — Prie (2012)
Unio crassus courtillieri — Prié & Puillandre (2014), Lopes-Lima et al. (2017)
Unio (Crassiana) crassus — Vinarski & Kantor (2016)
Unio (Crassiana) musivus — Vinarski & Kantor (2016)
Unio (Crassiana) cyprinorum — Vinarski & Kantor (2016)
Unio (Crassiana) irgizlaicus — Vinarski & Kantor (2016)
Unio (Crassiana) irenjensis — Vinarski & Kantor (2016)
Unio (Eolymnium) byzantinus — Vinarski & Kantor (2016)
Unio crassus cf. courtillieri — Klishko et al. (2017)
Unio crassus courtillierii — Araujo et al. (2018)
Unio c. crassus — Príe et al. (2021)
Unio c. courtillieri — Príe et al. (2021)
Unio cf. courtillieri — Lopes-Lima et al. (2021)