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Page last updated
Fri 07 Aug 2020

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

The nominal species (available and otherwise) in this synonymy are listed by usage rather than strictly by publication date. The primary sorting order is by the most recent year (since 1969) that the species name was treated as valid, and then nominal species are sorted secondarily by publication year.

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Rectidentinae | Contradentini | Contradens

species Contradens contradens (Lea, 1838): synonymy
     454 specimens available.

Unio contradens Lea, 1838
Lea, 1836, Synopsis Fam. Naiades: addenda. [nomen nudum]
Lea, 1838, Trans. Amer. Phil. Soc. 6 [O. 2]: 75, pl. 18, fig. 58.
Type(s): HOLOTYPE USNM_85185, [Java].

Unio contradensLea (1838), Hanley (1843), Reeve (1865)
Margarita (Unio) contradensLea (1836)
Margaron (Unio) contradensLea (1852), Lea (1870)
Unio (Naidea) contradensAdams & Adams (1857)
Nodularia (s.s.) contradensSimpson (1900), Simpson (1914)
Contradens contradensHaas (1913), Thiele (1934), van Benthem Jutting (1953), Haas (1969), Graf & Cummings (2006), Graf & Cummings (2007), Whelan et al. (2011), Affandi et al. (2013), Pfeiffer & Graf (2015), Zieritz et al. (2016), Lopes-Lima et al. (2017), Bolotov et al. (2017), Bolotov et al. (2017), Konopleva et al. (2017), Razak et al. (2018), Zieritz et al. (2018), Do et al. (2018), Bolotov et al. (2018), Pfeiffer et al. (2018), Huang et al. (2019), Konopleva et al. (2019), Jeratthitikul et al. (2019), Konopleva et al. (2019), Pfeiffer et al. (2019), Bolotov et al. (2020), Zieritz et al. (2020), Lopes-Lima et al. (2020)
Contradens (s.s.) contradensModell (1942), Haas (1969)
Uniandra contradens contradensBrandt (1974)
Uniandra contradensGraf (2002), Hoeh et al. (2009), Bogan (2010), IUCN (2012), Bogan & Do (2014)
Uniandra sp.Sri-aroon et al. (2007)
Contortus contortusAraujo et al. (2017)
Lens contradensPfeiffer et al. (2020)

Unio lugens Drouët & Chaper, 1892
Drouët & Chaper, 1892, Mém. Soc. Zool. France 5: 147, pl. 5, figs. 1-3.
Drouët, 1893, J. de Conch. 41: 44.
Type(s): SYNTYPE MNHN_MP_0139, Borneo.

Unio lugensDrouët & Chaper (1892)
Nodularia (s.s.) lugensSimpson (1900), Simpson (1914)
Contradens dimotus lugensHaas (1913)
Contradens ascia lugensHaas (1924)
Contradens asciaHaas (1969)
Lens lugensPfeiffer et al. (2020)

Unio dimotus Lea, 1870
Lea, 1870, Synopsis Fam. Naiades: 30.
(new name for Unio sumatrensis Lea, 1859)

Unio dimotusLea (1870)
Margaron (Unio) dimotusLea (1870)
Nodularia (s.s.) dimotaSimpson (1900), Simpson (1914)
Contradens dimotus dimotusHaas (1913)
Contradens ascia dimotusHaas (1924), van Benthem Jutting (1953), Marwoto (1987)
Contradens asciaHaas (1969)
Uniandra contradens dimotusBrandt (1974)
Lens contradensPfeiffer et al. (2020)

Unio ascia Hanley, 1856
‘Benson, MS’ Hanley, 1856, Cat. Rec. Bivalve Shells, Suppl.: 385, pl. 23, fig. 20.
Type(s): LECTOTYPE MCZ_17046, Penang, India.

Unio asciaHanley (1856), Preston (1912)
Contradens ascia asciaHaas (1924)
Ensidens (Uniandra) asciaPrashad (1932)
Contradens asciaHaas (1952), Haas (1969)
Uniandra contradens asciaBrandt (1974)
Contradens contradensJeratthitikul et al. (2019)
Lens contradensPfeiffer et al. (2020)

Unio rusticus Lea, 1856
Lea, 1856, Proc. Acad. Nat. Sci. 8: 93.
Lea, 1857, Jour. Acad. Nat. Sci. 3 [O. 6]: 294 [14], pl. 25, fig. 7.
Type(s): LECTOTYPE USNM_84082, Siam.

Unio rusticusLea (1856), Martens (1860), Sowerby (1868), Morelet (1875), Morlet (1889)
Margaron (Unio) rusticusLea (1870)
Nodularia (s.s.) rusticaSimpson (1900)
Contradens rusticusHaas (1914), Ortmann (1917), Haas (1952), Haas (1969)
Nodularia (s.s.) rusticusSimpson (1914)
Contradens (Sprickia) rusticusModell (1942)
Uniandra contradens rusticaBrandt (1974)
Contradens contradensJeratthitikul et al. (2019)
Lens contradensPfeiffer et al. (2020)

Unio tumidulus Lea, 1856
Lea, 1856, Proc. Acad. Nat. Sci. 8: 93.
Lea, 1857, Jour. Acad. Nat. Sci. 3 [O. 6]: 295 [15], pl. 25, fig. 9.
Type(s): LECTOTYPE USNM_84712, Siam.

Unio tumidulusLea (1856), Martens (1860), Sowerby (1868), Morelet (1875)
Unio rusticusSowerby (1868)
Margaron (Unio) tumidulusLea (1870)
Nodularia (s.s.) tumidulaSimpson (1900), Simpson (1914)
Ensidens (s.s.) tumidulusHaas (1913)
Ensidens (s.s.) ingallsianusHaas (1969)
Ensidens ingallsianusF.R. Woodward (1969)
Uniandra contradens tumidulaBrandt (1974)
Contradens contradensDo et al. (2018), Jeratthitikul et al. (2019)
Lens contradensPfeiffer et al. (2020)

Unio fischerianus Morlet, 1883
Morlet, 1883, J. de Conch. 31: 109, pl. 4, fig. 6.
Type(s): SYNTYPE MNHN_MP_0019, Cambodge.

Unio fischerianusMorlet (1883), Morlet (1889)
Nodularia (s.s.) fischerianiaSimpson (1900), Simpson (1914)
Contradens dimotus fischerianusHaas (1913)
Contradens ascia fischerianusHaas (1924), Haas (1952)
Contradens asciaHaas (1969)
Uniandra contradens fischerianaBrandt (1974)
Contradens contradensJeratthitikul et al. (2019)
Lens micropterusPfeiffer et al. (2020)

Unio asperulus Lea, 1866
Lea, 1866, Proc. Acad. Nat. Sci. 18: 133.
Lea, 1868, Jour. Acad. Nat. Sci. 6 [O. 12]: 280, pl. 38, fig. 94.
Type(s): HOLOTYPE USNM_84078, Siam.

Unio asperulusLea (1866)
Margaron (Unio) asperulusLea (1870)
Nodularia (s.s.) asperulaSimpson (1900), Simpson (1914)
Ensidens (sec. Uniandra) asperulusHaas (1913)
Ensidens (Uniandra) asperulusPrashad (1932), Haas (1952), Haas (1969)
Uniandra contradens tumidulaBrandt (1974)
Lens contradensPfeiffer et al. (2020)

Unio dautzenbergi Morlet, 1889
Morlet, 1889, J. de Conch. 37: 163, 190, pl. 8, fig. 5.

Unio dautzenbergiMorlet (1889)
Nodularia (s.s.) dautzenbergiSimpson (1900), Simpson (1914)
Unio (Nodularia) dautzenbergiDautzenberg & Fischer (1905)
Contradens dautzenbergiHaas (1914), Haas (1969)
Uniandra contradens rusticaBrandt (1974)
Contradens contradensDo et al. (2018)
Lens contradensPfeiffer et al. (2020)

Unio hageni Strubell, 1897
Strubell, 1897, Nachr. Bl. Dtsch. Malak. Ges. 29: 10.
Type(s): TYPE SMF_3603, S. Sumatra.

Unio hageniStrubell (1897)
Contradens hageniHaas (1913), Ortmann (1917), Haas (1969)
Nodularia (s.s.) dimotaSimpson (1914)
Uniandra contradens dimotusBrandt (1974)
Lens contradensPfeiffer et al. (2020)

Unio verbecki Martens, 1897
‘Bött.’ Martens, 1897, Zool. Erg. Nied. Ost. Ind. 4: 89, pl. 5, figs. 1-2, 4-5.

Unio verbeckiMartens (1897)
Nodularia (s.s.) verbeckiSimpson (1900), Simpson (1914)
Contradens verbeekiHaas (1914), Haas (1969)
Contradens verbeckiOrtmann (1917)
Contradens (Sprickia) verbeekiModell (1942), Haas (1969)
Uniandra contradens dimotusBrandt (1974)
Lens contradensPfeiffer et al. (2020)

Unio pajakomboensis Bullen, 1906
Bullen, 1906, Proc. Malac. Soc. London 7: 15, pl. 2, figs. 9-11.
Type(s): HOLOTYPE BMNH_1906-1-16-52, Pajakombo, Sumatra.

Unio pajakomboensisBullen (1906), Simpson (1914)
Schizocleithrum pajacombo├źnseHaas (1914)
Contradens hageniHaas (1969)
Uniandra contradens dimotusBrandt (1974)
Lens contradensPfeiffer et al. (2020)

Unio javanus Lea, 1840
Lea, 1840, Proc. Amer. Phil. Soc. 1: 285.
Lea, 1842, Trans. Amer. Phil. Soc. 8 [O. 3]: 220, pl. 18, fig. 37.
Type(s): HOLOTYPE USNM_85187, Java.

Unio javanusLea (1840), Küster (1856), Küster (1861), Sowerby (1868)
Margaron (Unio) javanusLea (1852)
Unio (Hyridella) javanusAdams & Adams (1857)
Margaron (Unio) contradensLea (1870)
Nodularia (s.s.) contradensSimpson (1900), Simpson (1914)
Contradens contradensHaas (1913), van Benthem Jutting (1953), Haas (1969)
Uniandra contradens contradensBrandt (1974)
Lens contradensPfeiffer et al. (2020)

Unio exilis Dunker, 1846
Dunker, 1846, Z. Malak. 3: 109.
Mousson, 1849, Land-Süßw.-Moll. Java: 92, pl. 16, fig. 3.

Unio exilisDunker (1846), Mousson (1849)
Margaron (Unio) exilisLea (1852)
Unio (Hyridella) exilisAdams & Adams (1857)
Margaron (Unio) contradensLea (1870)
Nodularia (s.s.) contradensSimpson (1900), Simpson (1914)
Contradens contradensHaas (1913), van Benthem Jutting (1953), Haas (1969)
Uniandra contradens contradensBrandt (1974)
Lens contradensPfeiffer et al. (2020)

Unio mutatus Mousson, 1849
Mousson, 1849, Land-Süßw.-Moll. Java: 92, pl. 16, figs. 1-2.

Unio mutatusMousson (1849)
Margaron (Unio) mutatusLea (1852)
Unio (Hyridella) mutatusAdams & Adams (1857)
Unio javanusSowerby (1868)
Margaron (Unio) contradensLea (1870)
Nodularia (s.s.) contradensSimpson (1900), Simpson (1914)
Contradens contradensHaas (1913), van Benthem Jutting (1953), Haas (1969)
Uniandra contradens contradensBrandt (1974)
Lens contradensPfeiffer et al. (2020)

Unio mutatus form curvatus Mousson, 1849
Mousson, 1849, Land-Süßw.-Moll. Java: 92 pl. 16, fig. 2. [infra-subsp.]
(homonym of Unio bonelli var. curvata Rossmässler, 1842; intro'd as intra-sp. of Unio mutatus Mousson, 1849)

Unio mutatus form curvatusMousson (1849)
Unio mutatusMousson (1849)
Contradens contradensvan Benthem Jutting (1953)

Unio mutatus form inflatus Mousson, 1849
Mousson, 1849, Land-Süßw.-Moll. Java: 92. [infra-subsp.]
(homonym of Unio inflatus Studer, 1820; intro'd as intra-sp. of Unio mutatus Mousson, 1849)

Unio mutatus form inflatusMousson (1849)
Unio mutatusMousson (1849)
Contradens contradensvan Benthem Jutting (1953)

Unio mutatus form obscurus Mousson, 1849
Mousson, 1849, Land-Süßw.-Moll. Java: 92, pl. 16, fig. 1. [infra-subsp.]
(homonym of Unio obscurus Lea, 1838; intro'd as intra-sp. of Unio mutatus Mousson, 1849)

Unio mutatus form obscurusMousson (1849)
Unio mutatusMousson (1849)
Contradens contradensvan Benthem Jutting (1953)

Unio sumatrensis Lea, 1859
Lea, 1859, Proc. Acad. Nat. Sci. 11: 153.
Lea, 1860, Jour. Acad. Nat. Sci. 4 [O. 7]: 239 [57], pl. 34, fig. 118.
(homonym of Unio sumatrensis Dunker, 1852; renamed Unio dimotus Lea, 1870)
Type(s): HOLOTYPE USNM_84059, Sumatra.

Unio sumatrensisLea (1859), Reeve (1865), Crosse & Fischer (1876), Bullen (1906)
Margaron (Unio) dimotusLea (1870)
Nodularia (s.s.) dimotaSimpson (1900), Simpson (1914)
Contradens dimotus dimotusHaas (1913)
Contradens asciaHaas (1969)
Uniandra contradens dimotusBrandt (1974)
Lens contradensPfeiffer et al. (2020)

Unio mederianus Küster, 1861
‘Mousson’ Küster, 1861, Conchyl. Cab. 9(2): 242, pl. 80, fig. 7.

Unio mederianusKüster (1861)
Nodularia (s.s.) contradensSimpson (1900), Simpson (1914)
Contradens contradensHaas (1913), van Benthem Jutting (1953), Haas (1969)
Uniandra contradens contradensBrandt (1974)
Lens contradensPfeiffer et al. (2020)

Unio inornatus Reeve, 1865
‘Lea’ Reeve, 1865, Conch. Icon. 16: pl. 29, sp. 147; errata.
(misidentified reference to Unio inornatus Lea, 1856)

Unio inornatusReeve (1865), Morelet (1875), Morlet (1889)
Margaron (Unio) versusLea (1870)
Nodularia (s.s.) asperulaSimpson (1900), Simpson (1914)
Ensidens (Uniandra) asperulusHaas (1969)
Uniandra contradens tumidulaBrandt (1974)
Lens micropterusPfeiffer et al. (2020)

Unio paivanus Morelet, 1865
Morelet, 1865, J. de Conch. 13: 227.
Type(s): HOLOTYPE BMNH_93-2-4-1738, Siam.

Unio paivanusMorelet (1865)
Unio rusticusMorelet (1875), Morlet (1889)
Nodularia (s.s.) rusticaSimpson (1900)
Contradens rusticusHaas (1914), Haas (1969)
Nodularia (s.s.) rusticusSimpson (1914)
Uniandra contradens rusticaBrandt (1974)
Lens contradensPfeiffer et al. (2020)

Unio cambojensis Sowerby, 1866
Sowerby, 1866, Conch. Icon. 16: pl. 42, sp. 231.
Type(s): LECTOTYPE BMNH_1965148, Camboja.

Unio cambojensisSowerby (1866)
Nodularia (s.s.) rusticaSimpson (1900)
Contradens rusticusHaas (1914)
Nodularia (s.s.) rusticusSimpson (1914)
Contradens cambojensisOrtmann (1917)
Uniandra contradens rusticaBrandt (1974)
Lens micropterusPfeiffer et al. (2020)

Margaron (Unio) versus Lea, 1870
‘Lea’ Reeve, 1865, Conch. Icon. 16: pl. 29, sp. 147 [as ‘Unio inornatus’].
Lea, 1870, Synopsis Fam. Naiades: 46.

Margaron (Unio) versusLea (1870)
Nodularia (s.s.) asperulaSimpson (1900), Simpson (1914)
Uniandra contradens tumidulaBrandt (1974)
Lens micropterusPfeiffer et al. (2020)

Unio paivaeanus Morelet, 1875
Morelet, 1875, Séries Conch. 4: 353 [as ‘paivæanus’].
Morlet, 1889, J. de Conch. 37: 164 [as ‘paivæanus’].
(error for Unio paivanus Morelet, 1865)

Unio paivaeanusMorelet (1875)

Unio semidecoratus Morlet, 1889
Morlet, 1889, J. de Conch. 37: 164, 192, pl. 8, fig. 4.

Unio semidecoratusMorlet (1889)
Nodularia (s.s.) semidecorataSimpson (1900)
Ensidens (sec. Uniandra) inaequalisHaas (1913)
Nodularia (s.s.) inaequalisSimpson (1914)
Ensidens (Uniandra) inaequalisHaas (1969)
Uniandra contradens tumidulaBrandt (1974)
Contradens contradensDo et al. (2018)
Lens contradensPfeiffer et al. (2020)

Unio siamensis Morlet, 1889
Morlet, 1889, J. de Conch. 37: 164, 194, pl. 7, fig. 2.
Morlet, 1904, Mission Pavie 3: 381, pl. 2, fig. 8.
(homonym of Unio siamensis Lea, 1866; renamed Unio soboles Fischer, 1891)

Unio siamensisMorlet (1889)
Nodularia (s.s.) sobolusSimpson (1900)
Contradens rusticusHaas (1914), Haas (1969)
Nodularia (s.s.) sobolesSimpson (1914)
Uniandra contradens rusticaBrandt (1974)
Lens micropterusPfeiffer et al. (2020)

Unio soboles Fischer, 1891
Fischer, 1891, Bull. Soc. Hist. Nat. Autun 4: 227.
(new name for Unio siamensis Morlet, 1889)

Unio sobolesFischer (1891)
Nodularia (s.s.) sobolusSimpson (1900)
Contradens rusticusHaas (1914), Haas (1969)
Nodularia (s.s.) sobolesSimpson (1914)
Uniandra contradens rusticaBrandt (1974)
Lens micropterusPfeiffer et al. (2020)

Unio saccellus Drouët & Chaper, 1892
Drouët & Chaper, 1892, Mém. Soc. Zool. France 5: 148, pl. 5, figs. 4-6.
Drouët, 1893, J. de Conch. 41: 43.
Type(s): SYNTYPES MNHN_MP_0093, Borneo.

Unio saccellusDrouët & Chaper (1892)
Nodularia (s.s.) sacellusSimpson (1900), Simpson (1914)
Contradens dimotus saccellusHaas (1913)
Contradens ascia sacellusHaas (1924)
Contradens asciaHaas (1969)
Lens lugensPfeiffer et al. (2020)

Microcondylaea bicristata Strubell, 1897
Strubell, 1897, Nachr. Bl. Dtsch. Malak. Ges. 29: 9.

Microcondylaea bicristataStrubell (1897)
Contradens hageniHaas (1913)
Pseudodon (sec. Bineurus) bicristatusSimpson (1914)
Pseudodon (Bineurus) bicristatusHaas (1920)
Pseudodon (Bineurus) hageniHaas (1969)
Uniandra contradens dimotusBrandt (1974)

Nodularia sobolus Simpson, 1900
‘Fischer’ Simpson, 1900, Proc. U.S. Nat. Mus. 22: 819.
(error for Unio soboles Fischer, 1891)

Nodularia sobolusSimpson (1900)
Contradens rusticusHaas (1914), Haas (1969)
Nodularia (s.s.) sobolesSimpson (1914)

Nodularia (s.s.) fischeriania Simpson, 1900
‘Morlet’ Simpson, 1900, Proc. U.S. Nat. Mus. 22: 818.
‘Morlet’ Simpson, 1914, Descr. Cat. Naiades: 1009.
(error for Unio fischerianus Morlet, 1883)

Nodularia (s.s.) fischerianiaSimpson (1900)

Unio hagnei Simpson, 1900
‘Strubell’ Simpson, 1900, Proc. U.S. Nat. Mus. 22: 862.
(error for Unio hageni Strubell, 1897)

Unio hagneiSimpson (1900)

Unio sacellus Simpson, 1900
‘Drouet & Chaper’ of Authors.
Simpson, 1900, Proc. U.S. Nat. Mus. 22: 818.
Simpson, 1914, Descr. Cat. Naiades: 1011.
Haas, 1924, Abh. Senckenb. Naturf. Ges. 38: 198.
(error for Unio saccellus Drouët & Chaper, 1892)

Unio sacellusSimpson (1900)
Contradens dimotus saccellusHaas (1913)

Schizocleithrum pajacomboense Haas, 1914
Haas, 1914, Conch. Cab. 9 (pt. 2, h. 49): 203 [as ‘pajacomboënse’].
(error for Unio pajakomboensis Bullen, 1906)

Schizocleithrum pajacomboenseHaas (1914)

Unio verbeeki Haas, 1914
‘von Martens’ of Authors.
‘(Boettger) v. Martens’ Haas, 1914, Conch. Cab. 9 (pt. 2, h. 49): 198.
Modell, 1942, Arch. Moll. 74(5/6): 189.
‘Martens’ Haas, 1969, Das Tierreich 88: 101.
‘Martens’ Haas, 1969, Treatise on Invert. Paleo: N420.
‘(Martens)’ Brandt, 1974, Arch. Moll. 105: 289.
Millard, 2001, Class. Moll. (3): 1064.
(error for Unio verbecki Martens, 1897)

Unio verbeekiHaas (1914)
Lens contradensPfeiffer et al. (2020)

Unio thaiensis Habe, 1964
Habe, 1964, Nature & Life in Southeast Asia 3: 62, pl. ?, figs. 3-4.

Unio thaiensisHabe (1964)
Uniandra contradens rusticaBrandt (1974)
Lens contradensPfeiffer et al. (2020)

Uniandra contradens rusticoides Brandt, 1974
Brandt, 1974, Arch. Moll. 105: 290, pl. 24, fig. 50.
(intro'd as intra-sp. of Unio contradens Lea, 1838)
Type(s): HOLOTYPE SMF_220845, Thailand: Klong Min near Chandi; Chawang Distr.; Prov. Nakon Sritammarat.

Uniandra contradens rusticoidesBrandt (1974)
Contradens contradensJeratthitikul et al. (2019)
Lens contradensPfeiffer et al. (2020)

Contortus contortus Araujo et al., 2017
Araujo et al., 2017, Zool. J. Linn. Soc. 179(2): 280.
(error for Unio contradens Lea, 1838)

Contortus contortusAraujo et al. (2017)
Lens contradensPfeiffer et al. (2020)

Page updated: 15:15:20 Fri 07 Aug 2020

 
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