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Page last updated
Sat 17 Feb 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Hyrioidea | Hyriidae | Hyriinae

tribe HYRIDELLINI McMichael, 1956 (1934): cladomics
     6 genera

Cladomics Summary

number of species = 17
number of subtaxa (rank: genus) = 6

analyzed in 44 cladograms in 26 publications
maximum number of terminals in an analysis = 4, mean = 1.57
maximum number of subtaxa (genus) in an analysis = 2, mean = 1.09

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 15
maximum number of included terminals = 4, mean = 2.67
maximum number of subtaxa (genus) = 2, mean = 1.27
proportion of cladograms that support basic monophyly = 0.6

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 4
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (tribe).
maximum number of included terminals = 4, mean = 3.5
maximum number of subtaxa (genus) = 2, mean = 2.0
proportion of cladograms that support rigorous monophyly = 0.5


Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ tree


Santos-Neto et al., 2016 [Publ 3739], fig. 2a,b

iT tT bi riS oT roT roS iD
4 23 20 2 20 13 4 0.0

*** Top Monophyly Cladogram *** *** Top Sister-Group Cladogram ***

                 +-------- Westralunio_carteri        
                 |                                    
  +--------------+     +-- Hyridella_drapeta **          
  |              |  +--+                              
  |              |  |  +-- Hyridella_depressa **         
  |              +--+                                 
  |                 |  +-- Diplodon_gratus            
  |                 +--+                              
  |                    +-- Diplodon_suavidicus[2]     
  |                                                   
  |                    +-- Hyridella_australis **        
  |     +--------------+                              
  |     |              +-- Cucumerunio_novaehollandiae **
  +-----+                                             
  |     |  +-------------- Echyridella_menziesii      
--+     +--+                                          
  |        |  +----------- Velesunio_ambiguus         
  |        +--+                                       
  |           |  +-------- Microdontia_anodontaeformis
  |           +--+                                    
  |              |  +----- Lortiella                  
  |              +--+                                 
  |                 |  +-- Alathyria_jacksoni         
  |                 +--+                              
  |                    +-- Velesunio_angasi           
  |                                                   
  |  +-------------------- Diplodon_suavidicus[1]     
  |  |                                                
  +--+                 +-- Triplodon_corrugatus       
     |  +--------------+                              
     |  |              +-- Prisodon_syrmatophorus     
     +--+                                             
        |  +-------------- Castalia_stevensi          
        |  |                                          
        +--+           +-- Callonaia_duprei           
           |  +--------+                              
           |  |        +-- Castalia_ambigua[1]        
           +--+                                       
              |  +-------- Castalia_ambigua[4]        
              +--+                                    
                 |  +----- Castalia_ambigua[2]        
                 +--+                                 
                    |  +-- Castaliella_quadrata       
                    +--+                              
                       +-- Castalia_ambigua[3]        

Graf et al., 2015 [Publ 3619], fig. 1

iT tT bi riS oT roT roS iD
4 4 2 2 16 3 3 19.0
     +----- Diplodon_suavidicus        
  +--+                                 
  |  |  +-- Triplodon_corrugatus       
  |  +--+                              
  |     +-- Castalia_ambigua           
--+                                    
  |     +-- Hyridella_depressa **         
  |  +--+                              
  |  |  +-- Hyridella_drapeta **          
  +--+                                 
     |  +-- Hyridella_australis **        
     +--+                              
        +-- Cucumerunio_novaehollandiae **

Santos-Neto et al., 2016 [Publ 3739], fig. 4

iT tT bi riS oT roT roS iD
4 13 11 2 20 13 4 16.0
     +----------------------- Diplodon_suavidicus[2]     
     |                                                   
  +--+                    +-- Triplodon_corrugatus       
  |  |  +-----------------+                              
  |  |  |                 +-- Prisodon_syrmatophorus     
  |  +--+                                                
  |     |  +----------------- Castalia_ambigua[4]        
  |     +--+                                             
  |        |  +-------------- Callonaia_duprei           
  |        +--+                                          
  |           |  +----------- Castalia_ambigua[3]        
  |           +--+                                       
  |              |  +-------- Castalia_stevensi          
  |              +--+                                    
--+                 |  +----- Castalia_ambigua[2]        
  |                 +--+                                 
  |                    |  +-- Castaliella_quadrata       
  |                    +--+                              
  |                       +-- Castalia_ambigua[1]        
  |                                                      
  |                 +-------- Westralunio_carteri        
  |                 |                                    
  |     +-----------+     +-- Hyridella_drapeta **          
  |     |           |  +--+                              
  |     |           |  |  +-- Hyridella_depressa **         
  |     |           +--+                                 
  |     |              |  +-- Diplodon_suavidicus[1]     
  +-----+              +--+                              
        |                 +-- Diplodon_gratus            
        |                                                
        |  +----------------- Echyridella_menziesii      
        |  |                                             
        +--+              +-- Cucumerunio_novaehollandiae **
           |  +-----------+                              
           |  |           +-- Hyridella_australis **        
           +--+                                          
              |  +----------- Velesunio_ambiguus         
              +--+                                       
                 |  +-------- Alathyria_jacksoni         
                 +--+                                    
                    |  +----- Velesunio_angasi           
                    +--+                                 
                       |  +-- Lortiella                  
                       +--+                              
                          +-- Microdontia_anodontaeformis

Graf & Ó Foighil, 2000 [Publ 2649], fig. 1, 2

iT tT bi riS oT roT roS iD
3 4 2 1 9 3 3 23.0
           +-- Diplodon_chilensis    
  +--------+                         
  |        +-- Castalia              
--+                                  
  |  +-------- Hyridella_australis[2] **
  +--+                               
     |  +----- Hyridella_australis[1] **
     +--+                            
        |  +-- Echyridella_menziesii 
        +--+                         
           +-- Hyridella_depressa **    

Huang et al., 2019 [Publ 3843], fig. 2

iT tT bi riS oT roT roS iD
3 3 1 1 201 4 4 20.0
  +----------- Echyridella_menziesii
  |                                 
  |     +----- Diplodon_suavidicus  
--+  +--+                           
  |  |  |  +-- Castalia_ambigua     
  |  |  +--+                        
  +--+     +-- Triplodon_corrugatus 
     |                              
     |  +----- Hyridella_australis **  
     +--+                           
        |  +-- Hyridella_depressa **   
        +--+                        
           +-- Hyridella_drapeta **    

Huang et al., 2019 [Publ 3843], fig. S3

iT tT bi riS oT roT roS iD
3 3 1 1 198 4 4 20.0
  +----------- Echyridella_menziesii
  |                                 
  |     +----- Diplodon_suavidicus  
--+  +--+                           
  |  |  |  +-- Castalia_ambigua     
  |  |  +--+                        
  +--+     +-- Triplodon_corrugatus 
     |                              
     |  +----- Hyridella_australis **  
     +--+                           
        |  +-- Hyridella_depressa **   
        +--+                        
           +-- Hyridella_drapeta **    

Huang et al., 2019 [Publ 3843], fig. S4

iT tT bi riS oT roT roS iD
3 3 1 1 199 4 4 20.0
  +----- Hyridella_australis **
--+                         
  |  +-- Hyridella_depressa ** 
  +--+                      
     +-- Hyridella_drapeta **  

Graf & Cummings, 2006 [Publ 1968], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 1 45 4 3 --
  +----------- Echyridella_menziesii
  |                                 
--+        +-- Hyridella_depressa **   
  |  +-----+                        
  |  |     +-- Hyridella_australis **  
  +--+                              
     |  +----- Castalia             
     +--+                           
        |  +-- Diplodon_chilensis   
        +--+                        
           +-- Diplodon_gratus      

Walker et al., 2006 [Publ 2666], fig. 6

iT tT bi riS oT roT roS iD
2 10 0 1 36 3 3 20.5
  +----------------- Castalia_stevensi    
  |                                       
  +----------------- Diplodon_gratus      
--+                                       
  |  +-------------- Hyridella_depressa **   
  |  |                                    
  |  +-------------- Hyridella_australis **  
  +--+                                    
     |  +----------- Echyridella_menziesii
     |  |                                 
     |  |        +-- Velesunio_ambiguus   
     +--+  +-----+                        
        |  |     +-- Velesunio[3]         
        |  |                              
        +--+  +----- Velesunio[2]         
           |  |                           
           |  |  +-- Alathyria_jacksoni   
           +--+--+                        
              |  +-- Velesunio[1]         
              |                           
              |  +-- Velesunio_angasi     
              +--+                        
                 +-- Lortiella_rugata     

Baker et al., 2004 [Publ 2685], fig. 2-3

iT tT bi riS oT roT roS iD
2 3 0 1 10 3 3 22.0
  +----- Hyridella_depressa ** 
--+                         
  |  +-- Hyridella_australis **
  +--+                      
     +-- Diplodon_gratus    

Baker et al., 2003 [Publ 2690], fig. 5

iT tT bi riS oT roT roS iD
2 7 0 1 6 3 3 23.0
           +-- Hyridella_australis **  
  +--------+                        
  |        +-- Diplodon_gratus      
--+                                 
  |  +-------- Hyridella_depressa **   
  |  |                              
  +--+     +-- Velesunio_angasi     
     |  +--+                        
     |  |  +-- Velesunio_ambiguus   
     +--+                           
        |  +-- Echyridella_menziesii
        +--+                        
           +-- Castalia_stevensi    

Hoeh et al., 2009 [Publ 2716], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 1 45 4 3 --
  +----------- Echyridella_menziesii
  |                                 
--+        +-- Hyridella_depressa **   
  |  +-----+                        
  |  |     +-- Hyridella_australis **  
  +--+                              
     |  +----- Castalia             
     +--+                           
        |  +-- Diplodon_chilensis   
        +--+                        
           +-- Diplodon_gratus      

Hoeh et al., 2009 [Publ 2716], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 1 44 4 3 --
  +----------- Echyridella_menziesii
  |                                 
--+        +-- Hyridella_depressa **   
  |  +-----+                        
  |  |     +-- Hyridella_australis **  
  +--+                              
     |  +----- Castalia             
     +--+                           
        |  +-- Diplodon_chilensis   
        +--+                        
           +-- Diplodon_gratus      

Hoeh et al., 2009 [Publ 2716], fig. 5

iT tT bi riS oT roT roS iD
2 2 0 1 44 4 3 --
  +----------- Echyridella_menziesii
  |                                 
--+        +-- Hyridella_depressa **   
  |  +-----+                        
  |  |     +-- Hyridella_australis **  
  +--+                              
     |  +----- Castalia             
     +--+                           
        |  +-- Diplodon_chilensis   
        +--+                        
           +-- Diplodon_gratus      

Marshall et al., 2014 [Publ 3593], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 12 5 2 --
     +-- Cucumerunio_novaehollandiae **
  +--+                              
  |  +-- Hyridella_australis **        
--+                                 
  |  +-- Diplodon_chilensis         
  +--+                              
     +-- Diplodon_gratus            

Graf, 2000 [Publ 2647], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 17 2 2 --
-- Hyridella

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

iT tT bi riS oT roT roS iD
1 1 0 1 29 0 0 --
-- Hyridella_depressa

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

iT tT bi riS oT roT roS iD
1 1 0 1 29 0 0 --
-- Hyridella_depressa

Graf, 2002 [Publ 2651], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 17 2 2 --
  +----- Hyridella_depressa **
--+                        
  |  +-- Diplodon_chilensis
  +--+                     
     +-- Castalia          

Walker et al., 2006 [Publ 2666], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 14 0 0 --
-- Hyridella

Walker et al., 2006 [Publ 2666], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 14 0 0 --
-- Hyridella

Walker et al., 2006 [Publ 2666], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 14 0 0 --
-- Hyridella

Whelan et al., 2011 [Publ 3380], fig. 1 ML

iT tT bi riS oT roT roS iD
1 1 0 1 36 2 2 --
  +----- Hyridella_australis ** 
--+                          
  |  +-- Castalia_ambigua    
  +--+                       
     +-- Triplodon_corrugatus

Whelan et al., 2011 [Publ 3380], fig. 1 MP

iT tT bi riS oT roT roS iD
1 1 0 1 36 2 2 --
  +----- Hyridella_australis ** 
--+                          
  |  +-- Castalia_ambigua    
  +--+                       
     +-- Triplodon_corrugatus

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 39 2 2 --
  +----- Hyridella_australis ** 
--+                          
  |  +-- Triplodon_corrugatus
  +--+                       
     +-- Castalia_ambigua    

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iT tT bi riS oT roT roS iD
1 1 0 1 80 0 0 --
-- Hyridella_depressa

Combosch et al., 2017 [Publ 3806], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 14 1 1 --
-- Hyridella_australis

Combosch et al., 2017 [Publ 3806], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 14 1 1 --
  +-- Triplodon_corrugatus
--+                       
  +-- Hyridella_australis ** 

Huang et al., 2018 [Publ 3828], fig. 3, 5

iT tT bi riS oT roT roS iD
1 1 0 1 28 1 1 --
-- Hyridella_australis

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 27 1 1 --
-- Hyridella_australis

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 27 1 1 --
-- Hyridella_australis

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iT tT bi riS oT roT roS iD
1 1 0 1 56 0 0 --
-- Hyridella_australis

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 56 0 0 --
-- Hyridella_australis

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iT tT bi riS oT roT roS iD
1 1 0 1 121 0 0 --
-- Hyridella_depressa

Klunzinger et al., 2021 [Publ 4086], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 5 0 0 --
-- Cucumerunio_novaehollandiae

Wu et al., 2020 [Publ 4121], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 33 1 1 --
-- Hyridella_australis

Wu et al., 2020 [Publ 4121], fig. S1

iT tT bi riS oT roT roS iD
1 1 0 1 33 1 1 --
-- Hyridella_australis

Sano et al., 2020 [Publ 4138], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 72 1 1 --
-- Hyridella_australis

Sano et al., 2020 [Publ 4138], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 71 1 1 --
-- Hyridella_australis

Sano et al., 2020 [Publ 4138], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 71 1 1 --
-- Hyridella_australis

Sonowal et al., 2021 [Publ 4221], fig. 3a

iT tT bi riS oT roT roS iD
1 1 0 1 14 1 1 --
-- Hyridella_australis

Sonowal et al., 2021 [Publ 4221], fig. 3b

iT tT bi riS oT roT roS iD
1 1 0 1 14 1 1 --
-- Hyridella_australis

Sonowal et al., 2021 [Publ 4221], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 15 1 1 --
-- Hyridella_australis

Klunzinger et al., 2022 [Publ 4259], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 3 0 0 --
-- Cucumerunio_novaehollandiae

Page updated: 10:58:18 Sat 17 Feb 2024

 
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