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Page last updated
Sun 06 Nov 2022

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Ambleminae | Lampsilini

genus Ptychobranchus Simpson, 1900: cladomics
     8 valid species

Cladomics Summary

number of species = 8

analyzed in 31 cladograms in 18 publications
maximum number of terminal species in an analysis = 6, mean = 1.16

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 1
maximum number of included species = 6, mean = 6.0
proportion of cladograms that support basic monophyly = 1.0


Roe, 2013, fig. 4

iT tT bi riS oT roT roS iD
6 6 4 6 6 6 5 0.0

*** Top Monophyly Cladogram ***

  +-------------------- Medionidus_conradicus         
  |                                                   
--+                 +-- Cyprogenia_aberti             
  |  +--------------+                                 
  |  |              +-- Dromus_dromas                 
  +--+                                                
     |  +-------------- Ptychobranchus_subtentus **      
     +--+                                             
        |  +----------- Ptychobranchus_jonesi **         
        +--+                                          
           |  +-------- Ptychobranchus_greenii **        
           +--+                                       
              |  +----- Ptychobranchus_occidentalis[2] **
              +--+                                    
                 |  +-- Ptychobranchus_occidentalis[1] **
                 +--+                                 
                    +-- Ptychobranchus_fasciolaris **    

Campbell et al., 2005, fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 116 48 22 --

*** Top Sister-Group Cladogram ***

     +-- Sagittunio_nasutus         
  +--+                              
  |  +-- Medionidus_conradicus      
--+                                 
  |  +-- Ptychobranchus_fasciolaris ** 
  +--+                              
     +-- Venustaconcha_ellipsiformis

Graf & Cummings, 2006, fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 46 7 7 --
  +-------------- Obliquaria_reflexa        
  |                                         
  +-------------- Truncilla_truncata        
--+                                         
  +-------------- Ptychobranchus_fasciolaris **
  |                                         
  |  +----------- Ligumia_recta             
  +--+                                      
     |  +-------- Cambarunio_iris           
     +--+                                   
        |  +----- Epioblasma_triquetra      
        +--+                                
           |  +-- Lampsilis_cardium         
           +--+                             
              +-- Ortmanniana_ligamentina   

Graf & Ó Foighil, 2000, fig. 2A

iT tT bi riS oT roT roS iD
1 1 0 1 29 11 8 --
  +----------- Ligumia_recta             
  |                                      
  +----------- Truncilla_truncata        
  |                                      
  |        +-- Ptychobranchus_fasciolaris **
  +--------+                             
--+        +-- Sagittunio_nasutus        
  |                                      
  |     +----- Lampsilis_siliquoidea     
  |  +--+                                
  |  |  |  +-- Epioblasma_triquetra      
  |  |  +--+                             
  |  |     +-- Epioblasma_brevidens      
  +--+                                   
     |  +----- Ortmanniana_ligamentina   
     |  |                                
     |  +----- Cambarunio_iris           
     +--+                                
        +----- Leaunio_vanuxemensis      
        |                                
        |  +-- Lampsilis_cardium         
        +--+                             
           +-- Lampsilis_fasciola        

Graf & Ó Foighil, 2000, fig. 2B

iT tT bi riS oT roT roS iD
1 1 0 1 29 11 8 --
              +-- Ptychobranchus_fasciolaris **
  +-----------+                             
  |           +-- Sagittunio_nasutus        
  |                                         
--+  +----------- Lampsilis_siliquoidea     
  |  |                                      
  |  |     +----- Ortmanniana_ligamentina   
  +--+  +--+                                
     |  |  |  +-- Lampsilis_cardium         
     |  |  +--+                             
     |  |     +-- Lampsilis_fasciola        
     +--+                                   
        |     +-- Epioblasma_brevidens      
        |  +--+                             
        |  |  +-- Epioblasma_triquetra      
        +--+                                
           |  +-- Cambarunio_iris           
           +--+                             
              +-- Leaunio_vanuxemensis      

Lydeard et al., 1996, fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 26 8 6 --
  +-------------- Potamilus_fragilis      
  |                                       
--+  +----------- Ptychobranchus_subtentus **
  |  |                                    
  +--+  +-------- Medionidus_conradicus   
     |  |                                 
     +--+  +----- Potamilus_purpuratus    
        |  |                              
        |  |  +-- Ellipsaria_lineolata    
        +--+--+                           
           |  +-- Villosa_delumbis        
           |                              
           |  +-- Lampsilis_teres         
           +--+                           
              +-- Lampsilis_siliquoidea   

Lydeard et al., 1996, fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 27 8 6 --
  +-------------------- Medionidus_conradicus   
--+                                             
  |  +----------------- Ptychobranchus_subtentus **
  +--+                                          
     |  +-------------- Potamilus_fragilis      
     +--+                                       
        |  +----------- Potamilus_purpuratus    
        +--+                                    
           |  +-------- Ellipsaria_lineolata    
           +--+                                 
              |  +----- Villosa_delumbis        
              +--+                              
                 |  +-- Lampsilis_teres         
                 +--+                           
                    +-- Lampsilis_siliquoidea   

Huang et al., 2002, fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 33 7 6 --
  +-------------- Potamilus_fragilis      
--+                                       
  |  +----------- Ptychobranchus_subtentus **
  +--+                                    
     |  +-------- Potamilus_purpuratus    
     +--+                                 
        |  +----- Lampsilis_teres         
        +--+                              
           |  +-- Villosa_delumbis        
           +--+                           
              +-- Ellipsaria_lineolata    

Huang et al., 2002, fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 33 7 6 --
  +-- Villosa_delumbis        
  |                           
  +-- Ellipsaria_lineolata    
  |                           
--+-- Lampsilis_teres         
  |                           
  +-- Potamilus_purpuratus    
  |                           
  +-- Ptychobranchus_subtentus **

Campbell et al., 2005, fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 114 48 22 --
           +-- Ptychobranchus_fasciolaris ** 
  +--------+                              
  |        +-- Venustaconcha_ellipsiformis
  |                                       
--+  +-------- Medionidus_conradicus      
  |  |                                    
  |  |     +-- Lemiox_rimosus             
  +--+-----+                              
     |     +-- Medionidus_acutissimus     
     |                                    
     |  +----- Dromus_dromas              
     +--+                                 
        |  +-- Cyprogenia_aberti          
        +--+                              
           +-- Cyprogenia_stegaria        

Chapman et al., 2008, fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 20 12 10 --
                 +-- Ptychobranchus_fasciolaris ** 
  +--------------+                              
  |              +-- Lemiox_rimosus             
  |                                             
--+              +-- Venustaconcha_ellipsiformis
  |  +-----------+                              
  |  |           +-- Obovaria_olivaria          
  +--+                                          
     |  +----------- Hamiota_subangulata        
     +--+                                       
        |  +-------- Lampsilis_ovata            
        +--+                                    
           |  +----- Ortmanniana_ligamentina    
           +--+                                 
              |  +-- Lampsilis_straminea        
              +--+                              
                 +-- Lampsilis_hydiana          

Zanatta & Murphy, 2006, fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 56 48 22 --
        +-- Medionidus_conradicus     
  +-----+                             
  |     +-- Ptychobranchus_fasciolaris **
--+                                   
  |  +----- Medionidus_acutissimus    
  |  |                                
  +--+----- Lemiox_rimosus            
     |                                
     |  +-- Dromus_dromas             
     +--+                             
        +-- Cyprogenia_stegaria       

Zanatta & Murphy, 2006, fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 57 48 22 --
     +----- Ptychobranchus_fasciolaris **
  +--+                                
  |  |  +-- Dromus_dromas             
  |  +--+                             
--+     +-- Cyprogenia_stegaria       
  |                                   
  |  +----- Medionidus_conradicus     
  +--+                                
     |  +-- Medionidus_acutissimus    
     +--+                             
        +-- Lemiox_rimosus            

Serb & Barnhart, 2008, fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 12 12 4 --
  +----------------------- Obliquaria_reflexa        
  |                                                  
--+  +-------------------- Ptychobranchus_fasciolaris **
  |  |                                               
  +--+                 +-- Cyprogenia_aberti[6]      
     |  +--------------+                             
     |  |              +-- Dromus_dromas             
     +--+                                            
        |  +-------------- Cyprogenia_aberti[4]      
        |  |                                         
        +--+  +----------- Cyprogenia_aberti[3]      
           |  |                                      
           +--+  +-------- Cyprogenia_aberti[5]      
              |  |                                   
              +--+  +----- Cyprogenia_stegaria[2]    
                 |  |                                
                 +--+----- Cyprogenia_aberti[2]      
                    |                                
                    |  +-- Cyprogenia_stegaria[1]    
                    +--+                             
                       +-- Cyprogenia_aberti[1]      

Serb & Barnhart, 2008, fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 12 12 4 --
  +----------------------- Obliquaria_reflexa        
  |                                                  
  |                    +-- Lampsilis_ornata          
  +--------------------+                             
--+                    +-- Lampsilis_siliquoidea     
  |                                                  
  |  +-------------------- Ptychobranchus_fasciolaris **
  |  |                                               
  +--+                 +-- Cyprogenia_aberti[6]      
     |  +--------------+                             
     |  |              +-- Dromus_dromas             
     +--+                                            
        |  +-------------- Cyprogenia_aberti[5]      
        |  |                                         
        +--+  +----------- Cyprogenia_aberti[4]      
           |  |                                      
           +--+  +-------- Cyprogenia_aberti[3]      
              |  |                                   
              +--+  +----- Cyprogenia_stegaria[2]    
                 |  |                                
                 +--+----- Cyprogenia_aberti[2]      
                    |                                
                    |  +-- Cyprogenia_stegaria[1]    
                    +--+                             
                       +-- Cyprogenia_aberti[1]      

Serb, 2006, fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 12 12 4 --
  +-------------- Obliquaria_reflexa        
  |                                         
  |        +----- Ptychobranchus_fasciolaris **
  +--------+                                
  |        |  +-- Lampsilis_ornata          
--+        +--+                             
  |           +-- Lampsilis_siliquoidea     
  |                                         
  |           +-- Cyprogenia_aberti[6]      
  |  +--------+                             
  |  |        +-- Dromus_dromas             
  +--+                                      
     |  +-------- Cyprogenia_aberti[5]      
     |  |                                   
     +--+  +----- Cyprogenia_aberti[4]      
        |  |                                
        |  |  +-- Cyprogenia_stegaria[1]    
        +--+  |                             
           |  +-- Cyprogenia_stegaria[2]    
           |  |                             
           +--+-- Cyprogenia_aberti[1]      
              |                             
              +-- Cyprogenia_aberti[2]      
              |                             
              +-- Cyprogenia_aberti[3]      

Serb, 2006, fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 11 11 4 --
  +-------------------- Obliquaria_reflexa        
  |                                               
  |                 +-- Lampsilis_ornata          
  +-----------------+                             
--+                 +-- Lampsilis_siliquoidea     
  |                                               
  |  +----------------- Ptychobranchus_fasciolaris **
  |  |                                            
  +--+              +-- Cyprogenia_aberti[5]      
     |  +-----------+                             
     |  |           +-- Dromus_dromas             
     +--+                                         
        |  +----------- Cyprogenia_aberti[4]      
        |  |                                      
        +--+  +-------- Cyprogenia_aberti[3]      
           |  |                                   
           +--+  +----- Cyprogenia_stegaria[2]    
              |  |                                
              +--+----- Cyprogenia_aberti[2]      
                 |                                
                 |  +-- Cyprogenia_stegaria[1]    
                 +--+                             
                    +-- Cyprogenia_aberti[1]      

Serb, 2006, fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 11 11 4 --
  +-------------------- Obliquaria_reflexa        
  |                                               
  |                 +-- Lampsilis_ornata          
  +-----------------+                             
--+                 +-- Lampsilis_siliquoidea     
  |                                               
  |  +----------------- Ptychobranchus_fasciolaris **
  |  |                                            
  +--+              +-- Cyprogenia_aberti[5]      
     |  +-----------+                             
     |  |           +-- Dromus_dromas             
     +--+                                         
        |  +----------- Cyprogenia_aberti[4]      
        |  |                                      
        +--+  +-------- Cyprogenia_aberti[3]      
           |  |                                   
           +--+  +----- Cyprogenia_stegaria[2]    
              |  |                                
              +--+----- Cyprogenia_aberti[2]      
                 |                                
                 |  +-- Cyprogenia_stegaria[1]    
                 +--+                             
                    +-- Cyprogenia_aberti[1]      

Krebs et al., 2003, fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 33 10 7 --
           +----- Ptychobranchus_subtentus **
  +--------+                              
  |        |  +-- Potamilus_fragilis      
  |        +--+                           
  |           +-- Medionidus_conradicus   
--+                                       
  |           +-- Villosa_delumbis        
  |  +--------+                           
  |  |        +-- Ellipsaria_lineolata    
  +--+                                    
     |  +-------- Lampsilis_teres         
     +--+                                 
        |  +----- Lampsilis_siliquoidea   
        +--+                              
           |  +-- Potamilus_purpuratus    
           +--+                           
              +-- Potamilus_alatus        

Hoeh et al., 2009, fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 46 7 7 --
  +----------------- Ptychobranchus_fasciolaris **
--+                                            
  |  +-------------- Truncilla_truncata        
  +--+                                         
     |  +----------- Ortmanniana_ligamentina   
     +--+                                      
        |  +-------- Lampsilis_cardium         
        +--+                                   
           |  +----- Cambarunio_iris           
           +--+                                
              |  +-- Ligumia_recta             
              +--+                             
                 +-- Epioblasma_triquetra      

Hoeh et al., 2009, fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 45 7 7 --
  +----------------- Ptychobranchus_fasciolaris **
--+                                            
  |  +-------------- Truncilla_truncata        
  +--+                                         
     |  +----------- Ortmanniana_ligamentina   
     +--+                                      
        |  +-------- Lampsilis_cardium         
        +--+                                   
           |  +----- Cambarunio_iris           
           +--+                                
              |  +-- Ligumia_recta             
              +--+                             
                 +-- Epioblasma_triquetra      

Hoeh et al., 2009, fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 45 6 6 --
  +----- Truncilla_truncata        
  |                                
  +----- Ptychobranchus_fasciolaris **
--+                                
  |  +-- Ortmanniana_ligamentina   
  |  |                             
  |  +-- Cambarunio_iris           
  +--+                             
     +-- Ligumia_recta             
     |                             
     +-- Epioblasma_triquetra      

Chapman et al., 2008, fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 20 12 10 --
                 +-- Ptychobranchus_fasciolaris ** 
  +--------------+                              
  |              +-- Lemiox_rimosus             
  |                                             
--+              +-- Venustaconcha_ellipsiformis
  |  +-----------+                              
  |  |           +-- Obovaria_olivaria          
  +--+                                          
     |  +----------- Hamiota_subangulata        
     +--+                                       
        |  +-------- Lampsilis_ovata            
        +--+                                    
           |  +----- Ortmanniana_ligamentina    
           +--+                                 
              |  +-- Lampsilis_straminea        
              +--+                              
                 +-- Lampsilis_hydiana          

Jadhav & Jamkhedkar, 2009, fig. 11

iT tT bi riS oT roT roS iD
1 1 0 1 25 7 7 --
  +-- Cambarunio_iris           
--+                             
  +-- Ptychobranchus_fasciolaris **

Huang et al., 2019, fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 199 13 11 --
  +----------- Venustaconcha_ellipsiformis
  |                                       
  +----------- Leaunio_lienosus           
  |                                       
  |        +-- Ellipsaria_lineolata       
  +--------+                              
--+        +-- Truncilla_truncata         
  |                                       
  |        +-- Medionidus_conradicus      
  +--------+                              
  |        +-- Ptychobranchus_subtentus **   
  |                                       
  |  +-------- Cambarunio_iris            
  +--+                                    
     |  +----- Ortmanniana_ligamentina    
     +--+                                 
        |  +-- Lampsilis_cardium          
        +--+                              
           +-- Lampsilis_ornata           

Huang et al., 2019, fig. S3

iT tT bi riS oT roT roS iD
1 1 0 1 197 13 11 --
                 +-- Medionidus_conradicus      
  +--------------+                              
  |              +-- Ptychobranchus_subtentus **   
  |                                             
--+              +-- Ellipsaria_lineolata       
  |  +-----------+                              
  |  |           +-- Truncilla_truncata         
  +--+                                          
     |  +----------- Leaunio_lienosus           
     |  |                                       
     +--+        +-- Venustaconcha_ellipsiformis
        |  +-----+                              
        |  |     +-- Cambarunio_iris            
        +--+                                    
           |  +----- Ortmanniana_ligamentina    
           +--+                                 
              |  +-- Lampsilis_cardium          
              +--+                              
                 +-- Lampsilis_ornata           

Huang et al., 2019, fig. S4

iT tT bi riS oT roT roS iD
1 1 0 1 200 13 11 --
  +-------- Ellipsaria_lineolata       
  |                                    
  +-------- Potamilus_leptodon         
  |                                    
  +-------- Medionidus_conradicus      
  |                                    
  +-------- Potamilus_alatus           
  |                                    
  +-------- Ptychobranchus_subtentus **   
--+                                    
  +-------- Truncilla_truncata         
  |                                    
  +-------- Venustaconcha_ellipsiformis
  |                                    
  +-------- Cambarunio_iris            
  |                                    
  +-------- Leaunio_lienosus           
  |                                    
  |  +----- Ortmanniana_ligamentina    
  +--+                                 
     |  +-- Lampsilis_cardium          
     +--+                              
        +-- Lampsilis_ornata           

Pfeiffer et al., 2019, fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 38 11 10 --
  +-- Truncilla_truncata        
--+                             
  +-- Ptychobranchus_fasciolaris **

Smith et al., 2019, fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 41 36 19 --
     +-- Medionidus_conradicus     
  +--+                             
  |  +-- Lemiox_rimosus            
--+                                
  |  +-- Cyprogenia_stegaria       
  +--+                             
     +-- Ptychobranchus_fasciolaris **

Inoue et al., 2020, fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 28 11 10 --
  +-- Ptychobranchus_fasciolaris **
--+                             
  +-- Truncilla_truncata        

Inoue et al., 2020, fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 28 10 9 --
No taxon cladogram to show.

Page updated: 10:57:08 Sun 06 Nov 2022

 
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