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Page last updated
Wed 06 Nov 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Ambleminae | Lampsilini

genus Ortmanniana Frierson, 1927: cladomics
     2 valid species

Cladomics Summary

number of species = 2

analyzed in 53 cladograms in 32 publications
maximum number of terminals in an analysis = 2, mean = 1.09

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 5
maximum number of included terminals = 2, mean = 2.0
proportion of cladograms that support basic monophyly = 0


Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ tree


Zanatta & Murphy, 2006 [Publ 2672], fig. 1

iT tT bi riS oT roT roS iD
2 23 0 2 54 46 22 0.0

*** Top Monophyly Cladogram *** *** Top Sister-Group Cladogram ***

                                   +-- Cyrtonaias             
  +--------------------------------+                          
  |                                +-- Glebula                
  |                                                           
--+  +-------------------------------- Paetulunio             
  |  |                                                        
  |  |                             +-- Obliquaria             
  +--+  +--------------------------+                          
     |  |                          +-- Toxolasma              
     |  |                                                     
     |  |  +-------------------------- Sagittunio             
     +--+  |                                                  
        |  |              +----------- Potamilus              
        |  |              |                                   
        |  |  +-----------+        +-- Medionidus_conradicus  
        |  |  |           |  +-----+                          
        +--+  |           |  |     +-- Ptychobranchus         
           |  |           +--+                                
           |  |              |  +----- Medionidus_acutissimus 
           |  |              |  |                             
           |  |              +--+----- Lemiox                 
           |  |                 |                             
           |  |                 |  +-- Dromus                 
           +--+                 +--+                          
              |                    +-- Cyprogenia             
              |                                               
              |                 +----- Lampsilis_teres        
              |  +--------------+                             
              |  |              |  +-- Truncilla              
              |  |              +--+                          
              |  |                 +-- Ellipsaria             
              |  |                                            
              +--+  +----------------- Leaunio                
                 |  |                                         
                 |  |     +----------- Hamiota                
                 |  |     |                                   
                 |  |  +--+        +-- Lampsilis_fasciola     
                 +--+  |  |  +-----+                          
                    |  |  |  |     +-- Ortmanniana_pectorosa **  
                    |  |  +--+                                
                    |  |     |  +----- Lampsilis_ornata       
                    |  |     +--+                             
                    |  |        |  +-- Lampsilis_ovata        
                    +--+        +--+                          
                       |           +-- Lampsilis_cardium      
                       |                                      
                       |        +----- Venustaconcha          
                       |  +-----+                             
                       |  |     |  +-- Obovaria               
                       |  |     +--+                          
                       +--+        +-- Epioblasma             
                          |                                   
                          |  +-------- Lampsilis_siliquoidea  
                          +--+                                
                             |  +----- Ortmanniana_ligamentina **
                             +--+                             
                                |  +-- Cambarunio             
                                +--+                          
                                   +-- Ligumia                

Campbell et al., 2005 [Publ 2660], fig. 2

iT tT bi riS oT roT roS iD
2 38 0 2 100 41 22 34.0
           +----- Obliquaria                    
  +--------+                                    
  |        |  +-- Cyrtonaias                    
  |        +--+                                 
  |           +-- Glebula                       
  |                                             
  |  +----------- Sagittunio                    
  |  |                                          
  |  +----------- Leaunio                       
  |  |                                          
  |  +----------- Potamilus                     
  |  |                                          
  |  +----------- Hamiota                       
  |  |                                          
  |  |        +-- Ellipsaria                    
--+  +--------+                                 
  |  |        +-- Truncilla                     
  |  |                                          
  |  |     +----- Lampsilis_teres               
  |  +-----+                                    
  |  |     |  +-- Ortmanniana_ligamentina **       
  |  |     +--+                                 
  |  |        +-- Lampsilis_siliquoidea         
  |  |                                          
  |  |     +----- Villosa                       
  |  +-----+                                    
  |  |     |  +-- Ligumia                       
  |  |     +--+                                 
  |  |        +-- Cambarunio                    
  |  |                                          
  +--+  +-------- Ortmanniana_pectorosa **         
     +--+                                       
     |  |  +----- Lampsilis_ornata              
     |  +--+                                    
     |     |  +-- Lampsilis_cardium             
     |     +--+                                 
     |        +-- Lampsilis_ovata               
     |                                          
     |  +-------- Obovaria_olivaria             
     |  |                                       
     |  +-------- Venustaconcha_ellipsiformis__2
     +--+                                       
     |  +-------- Epioblasma                    
     |  |                                       
     |  |  +----- Obovaria_unicolor             
     |  +--+                                    
     |     |  +-- Obovaria_arkansasensis        
     |     +--+                                 
     |        +-- Obovaria_subrotunda           
     |                                          
     |        +-- Ptychobranchus                
     |  +-----+                                 
     |  |     +-- Venustaconcha_ellipsiformis__1
     |  |                                       
     +--+  +----- Medionidus_conradicus         
        |  |                                    
        |  |  +-- Lemiox                        
        +--+--+                                 
           |  +-- Medionidus_acutissimus        
           |                                    
           |  +-- Dromus                        
           +--+                                 
              +-- Cyprogenia                    

Smith et al., 2019 [Publ 3951], fig. 2

iT tT bi riS oT roT roS iD
2 6 0 2 39 34 19 23.0
  +----------------------- Obliquaria             
  |                                               
  |                    +-- Medionidus             
  |                 +--+                          
  |                 |  +-- Lemiox                 
--+  +--------------+                             
  |  |              |  +-- Cyprogenia             
  |  |              +--+                          
  |  |                 +-- Ptychobranchus         
  |  |                                            
  +--+           +-------- Potamilus              
     |  +--------+                                
     |  |        |  +----- Atlanticoncha          
     |  |        +--+                             
     |  |           |  +-- Ellipsaria             
     |  |           +--+                          
     +--+              +-- Truncilla              
        |                                         
        |              +-- Hamiota                
        |  +-----------+                          
        |  |           +-- Villosa                
        |  |                                      
        +--+  +----------- Ligumia                
           |  |                                   
           |  |     +----- Venustaconcha          
           +--+  +--+                             
              |  |  |  +-- Obovaria               
              |  |  +--+                          
              +--+     +-- Epioblasma             
                 |                                
                 |  +----- Ortmanniana_pectorosa **  
                 +--+                             
                    |  +-- Ortmanniana_ligamentina **
                    +--+                          
                       +-- Lampsilis              

Zanatta & Murphy, 2006 [Publ 2672], fig. 2

iT tT bi riS oT roT roS iD
2 25 0 2 55 46 22 12.0
                                      +-- Paetulunio             
  +-----------------------------------+                          
  |                                   +-- Toxolasma              
  |                                                              
  |                                +----- Obliquaria             
  |  +-----------------------------+                             
--+  |                             |  +-- Glebula                
  |  |                             +--+                          
  |  |                                +-- Cyrtonaias             
  |  |                                                           
  |  |                             +----- Ptychobranchus         
  |  |                          +--+                             
  +--+                          |  |  +-- Dromus                 
     |                          |  +--+                          
     |  +-----------------------+     +-- Cyprogenia             
     |  |                       |                                
     |  |                       |  +----- Medionidus_conradicus  
     |  |                       +--+                             
     |  |                          |  +-- Medionidus_acutissimus 
     |  |                          +--+                          
     +--+                             +-- Lemiox                 
        |                                                        
        |  +----------------------------- Sagittunio             
        |  |                                                     
        |  +----------------------------- Potamilus              
        |  |                                                     
        |  |                          +-- Truncilla              
        +--+  +-----------------------+                          
           |  |                       +-- Ellipsaria             
           |  |                                                  
           |  |                       +-- Ortmanniana_pectorosa **  
           |  |                 +-----+                          
           |  |                 |     +-- Lampsilis_fasciola     
           +--+  +--------------+                                
              |  |              |  +----- Lampsilis_ornata       
              |  |              +--+                             
              |  |                 |  +-- Lampsilis_ovata        
              |  |                 +--+                          
              |  |                    +-- Lampsilis_cardium      
              +--+                                               
                 |                    +-- Ligumia                
                 |                 +--+                          
                 |                 |  +-- Cambarunio             
                 |  +--------------+                             
                 |  |              |  +-- Lampsilis_teres        
                 |  |              +--+                          
                 +--+                 +-- Hamiota                
                    |                                            
                    |  +----------------- Leaunio                
                    |  |                                         
                    +--+  +-------------- Ortmanniana_ligamentina **
                       |  |                                      
                       +--+  +----------- Lampsilis_siliquoidea  
                          |  |                                   
                          +--+  +-------- Obovaria_olivaria      
                             |  |                                
                             |  +-------- Venustaconcha          
                             +--+                                
                                +-------- Epioblasma             
                                |                                
                                |  +----- Obovaria_unicolor      
                                +--+                             
                                   |  +-- Obovaria_subrotunda    
                                   +--+                          
                                      +-- Obovaria_arkansasensis 

Campbell et al., 2005 [Publ 2660], fig. 1

iT tT bi riS oT roT roS iD
2 14 0 2 100 41 22 20.0
                       +-------- Toxolasma                     
  +--------------------+                                       
  |                    |  +----- Obliquaria                    
  |                    +--+                                    
  |                       |  +-- Cyrtonaias                    
  |                       +--+                                 
  |                          +-- Glebula                       
  |                                                            
  |              +-------------- Potamilus                     
  |              |                                             
  |              |     +-------- Medionidus_acutissimus        
--+              |  +--+                                       
  |  +-----------+  |  |  +----- Lemiox                        
  |  |           |  |  +--+                                    
  |  |           |  |     |  +-- Dromus                        
  |  |           |  |     +--+                                 
  |  |           +--+        +-- Cyprogenia                    
  |  |              |                                          
  |  |              |        +-- Sagittunio                    
  |  |              |     +--+                                 
  |  |              |     |  +-- Medionidus_conradicus         
  +--+              +-----+                                    
     |                    |  +-- Ptychobranchus                
     |                    +--+                                 
     |                       +-- Venustaconcha_ellipsiformis__2
     |                                                         
     |                       +-- Ellipsaria                    
     |  +--------------------+                                 
     |  |                    +-- Truncilla                     
     |  |                                                      
     |  |  +-------------------- Villosa                       
     +--+  |                                                   
        |  |                 +-- Obovaria_olivaria             
        |  |        +--------+                                 
        |  |        |        +-- Venustaconcha_ellipsiformis__1
        |  |  +-----+                                          
        +--+  |     |  +-------- Epioblasma                    
           |  |     +--+                                       
           |  |        |  +----- Obovaria_unicolor             
           |  |        +--+                                    
           |  |           |  +-- Obovaria_arkansasensis        
           |  |           +--+                                 
           |  |              +-- Obovaria_subrotunda           
           +--+                                                
              |        +-------- Hamiota                       
              |        |                                       
              |  +-----+     +-- Lampsilis_cardium             
              |  |     |  +--+                                 
              |  |     |  |  +-- Lampsilis_ovata               
              |  |     +--+                                    
              |  |        |  +-- Ortmanniana_pectorosa **         
              +--+        +--+                                 
                 |           +-- Lampsilis_ornata              
                 |                                             
                 |           +-- Ligumia                       
                 |  +--------+                                 
                 |  |        +-- Cambarunio                    
                 +--+                                          
                    |  +-------- Lampsilis_teres               
                    +--+                                       
                       |  +----- Ortmanniana_ligamentina **       
                       +--+                                    
                          |  +-- Lampsilis_siliquoidea         
                          +--+                                 
                             +-- Leaunio                       

Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11

iT tT bi riS oT roT roS iD
1 1 0 1 24 7 7 --
  +----------- Truncilla              
  |                                   
--+        +-- Lampsilis              
  |  +-----+                          
  |  |     +-- Epioblasma             
  +--+                                
     |  +----- Ligumia                
     +--+                             
        |  +-- Obliquaria             
        +--+                          
           +-- Ortmanniana_ligamentina **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iT tT bi riS oT roT roS iD
1 1 0 1 75 2 2 --
  +----- Ortmanniana_ligamentina **
--+                             
  |  +-- Cambarunio             
  +--+                          
     +-- Lampsilis              

Walker et al., 2006 [Publ 2666], fig. 6-M

iT tT bi riS oT roT roS iD
1 1 0 1 13 4 4 --
  +----------- Potamilus              
--+                                   
  |  +-------- Cyrtonaias             
  +--+                                
     |  +----- Ligumia                
     +--+                             
        |  +-- Lampsilis              
        +--+                          
           +-- Ortmanniana_ligamentina **

Inoue et al., 2020 [Publ 4128], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 24 10 9 --
  +-- Ortmanniana_ligamentina **
--+                          
  +-- Lampsilis              

Inoue et al., 2020 [Publ 3941], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 41 26 6 --
  +----------------------------------- Toxolasma              
  |                                                           
--+  +-------------------------------- Potamilus              
  |  |                                                        
  |  |                             +-- Ellipsaria             
  +--+  +--------------------------+                          
     |  |                          +-- Truncilla              
     |  |                                                     
     |  |  +-------------------------- Leaunio                
     +--+  |                                                  
        |  |     +-------------------- Lampsilis_bracteata    
        |  |     |                                            
        |  |  +--+  +----------------- Lampsilis_fasciola     
        |  |  |  |  |                                         
        +--+  |  +--+  +-------------- Lampsilis_ornata       
           |  |     |  |                                      
           |  |     +--+           +-- Lampsilis_ovata        
           |  |        |  +--------+                          
           |  |        |  |        +-- Lampsilis_cariosa      
           |  |        +--+                                   
           |  |           |  +-------- Lampsilis_cardium__3   
           +--+           +--+                                
              |              |  +----- Lampsilis_cardium__2   
              |              +--+                             
              |                 |  +-- Lampsilis_satura       
              |                 +--+                          
              |                    +-- Lampsilis_cardium__1   
              |                                               
              |                    +-- Lampsilis_higginsii    
              |     +--------------+                          
              |     |              +-- Ortmanniana_ligamentina **
              +-----+                                         
                    |  +-------------- Lampsilis_teres        
                    |  |                                      
                    +--+           +-- Lampsilis_radiata      
                       |  +--------+                          
                       |  |        +-- Lampsilis_siliquoidea  
                       +--+                                   
                          |  +-------- Lampsilis_bergmanni    
                          +--+                                
                             |  +----- Lampsilis_hydiana      
                             +--+                             
                                |  +-- Lampsilis_virescens    
                                +--+                          
                                   +-- Lampsilis_straminea    

Porto-Hannes et al., 2021 [Publ 4227], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 14 8 3 --
           +-- Obovaria               
  +--------+                          
  |        +-- Ortmanniana_ligamentina **
  |                                   
  |        +-- Obliquaria             
  +--------+                          
--+        +-- Lampsilis_ornata       
  |                                   
  |     +----- Lampsilis_radiata__2   
  |  +--+                             
  |  |  |  +-- Lampsilis_siliquoidea  
  |  |  +--+                          
  +--+     +-- Lampsilis_radiata__1   
     |                                
     |  +----- Lampsilis_virescens    
     +--+                             
        |  +-- Lampsilis_hydiana      
        +--+                          
           +-- Lampsilis_straminea    

Bolotov et al., 2018 [Publ 3826], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 82 2 2 --
-- Ortmanniana_ligamentina

Hoeh et al., 2002 [Publ 2653], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 30 5 5 --
  +----- Potamilus              
--+                             
  |  +-- Ligumia                
  +--+                          
     +-- Ortmanniana_ligamentina **

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

iT tT bi riS oT roT roS iD
1 1 0 1 29 11 8 --
  +----------- Ligumia                
  |                                   
  +----------- Truncilla              
  |                                   
  |        +-- Ptychobranchus         
  +--------+                          
--+        +-- Sagittunio             
  |                                   
  |        +-- Lampsilis_siliquoidea  
  |  +-----+                          
  |  |     +-- Epioblasma             
  |  |                                
  +--+  +----- Ortmanniana_ligamentina **
     |  |                             
     |  +----- Cambarunio             
     +--+                             
        +----- Leaunio                
        |                             
        |  +-- Lampsilis_cardium      
        +--+                          
           +-- Lampsilis_fasciola     

Boyer et al., 2011 [Publ 2718], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 37 15 9 --
  +----------------- Potamilus              
  |                                         
  |              +-- Ellipsaria             
--+  +-----------+                          
  |  |           +-- Truncilla              
  |  |                                      
  +--+  +----------- Lampsilis_cardium      
     |  |                                   
     |  |     +----- Ligumia                
     |  +-----+                             
     +--+     |  +-- Obovaria               
        |     +--+                          
        |        +-- Epioblasma             
        |                                   
        |  +-------- Lampsilis_teres        
        +--+                                
           |  +----- Lampsilis_siliquoidea  
           +--+                             
              |  +-- Ortmanniana_ligamentina **
              +--+                          
                 +-- Lampsilis_higginsii    

Krebs et al., 2013 [Publ 3515], fig. 2A

iT tT bi riS oT roT roS iD
1 1 0 1 4 4 1 --
  +-------- Lampsilis_radiata      
  |                                
--+     +-- Lampsilis_ovata        
  |  +--+                          
  |  |  +-- Ortmanniana_ligamentina **
  +--+                             
     |  +-- Lampsilis_hydiana      
     +--+                          
        +-- Lampsilis_straminea    

Hoeh et al., 2001 [Publ 2652], fig. 14f2

iT tT bi riS oT roT roS iD
1 1 0 1 30 5 5 --
  +----------- Ortmanniana_ligamentina **
--+                                   
  |  +-------- Glebula                
  +--+                                
     |  +----- Potamilus              
     +--+                             
        |  +-- Ligumia                
        +--+                          
           +-- Toxolasma              

Hoeh et al., 2001 [Publ 2652], fig. 14f4

iT tT bi riS oT roT roS iD
1 1 0 1 30 5 5 --
           +-- Cyrtonaias             
  +--------+                          
  |        +-- Glebula                
--+                                   
  |  +-------- Toxolasma              
  +--+                                
     |  +----- Potamilus              
     +--+                             
        |  +-- Ligumia                
        +--+                          
           +-- Ortmanniana_ligamentina **

Huang et al., 2019 [Publ 3843], fig. S4

iT tT bi riS oT roT roS iD
1 1 0 1 197 13 11 --
  +----- Ellipsaria             
  |                             
  +----- Potamilus_leptodon     
  |                             
  +----- Medionidus             
  |                             
  +----- Potamilus_alatus       
  |                             
  +----- Ptychobranchus         
--+                             
  +----- Truncilla              
  |                             
  +----- Venustaconcha          
  |                             
  +----- Cambarunio             
  |                             
  +----- Leaunio                
  |                             
  |  +-- Ortmanniana_ligamentina **
  +--+                          
     +-- Lampsilis              

Porto-Hannes et al., 2019 [Publ 3969], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 17 16 5 --
  +----------------- Lampsilis_teres        
  |                                         
  +----------------- Obovaria               
  |                                         
  |              +-- Leaunio                
  +--------------+                          
  |              +-- Cambarunio             
  |                                         
  |        +-------- Ortmanniana_ligamentina **
  |        |                                
--+--------+     +-- Lampsilis_hydiana      
  |        |  +--+                          
  |        |  |  +-- Lampsilis_virescens    
  |        +--+                             
  |           |  +-- Lampsilis_siliquoidea  
  |           +--+                          
  |              +-- Lampsilis_radiata      
  |                                         
  |  +-------------- Lampsilis_bracteata    
  |  |                                      
  +--+  +----------- Lampsilis_ornata       
     |  |                                   
     +--+  +-------- Lampsilis_ovata        
        |  |                                
        +--+  +----- Lampsilis_cardium__3   
           |  |                             
           +--+  +-- Lampsilis_satura       
              |  |                          
              +--+-- Lampsilis_cardium__1   
                 |                          
                 +-- Lampsilis_cardium__2   

Walker et al., 2006 [Publ 2666], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 14 4 4 --
  +----------- Cyrtonaias             
--+                                   
  |  +-------- Potamilus              
  +--+                                
     |  +----- Ligumia                
     +--+                             
        |  +-- Ortmanniana_ligamentina **
        +--+                          
           +-- Lampsilis              

Porto-Hannes et al., 2019 [Publ 3969], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 17 16 5 --
  +----------------------- Obliquaria             
  |                                               
  |                    +-- Cambarunio             
  |  +-----------------+                          
  |  |                 +-- Leaunio                
--+  |                                            
  |  |        +----------- Lampsilis_teres        
  |  |        |                                   
  |  +--------+  +-------- Ortmanniana_ligamentina **
  |  |        |  |                                
  |  |        +--+     +-- Lampsilis_hydiana      
  +--+           |  +--+                          
     |           |  |  +-- Lampsilis_virescens    
     |           +--+                             
     |              |  +-- Lampsilis_siliquoidea  
     |              +--+                          
     |                 +-- Lampsilis_radiata      
     |                                            
     |  +----------------- Obovaria               
     |  |                                         
     +--+  +-------------- Lampsilis_bracteata    
        |  |                                      
        +--+  +----------- Lampsilis_ornata       
           |  |                                   
           +--+  +-------- Lampsilis_ovata        
              |  |                                
              +--+  +----- Lampsilis_cardium__3   
                 |  |                             
                 +--+  +-- Lampsilis_satura       
                    |  |                          
                    +--+-- Lampsilis_cardium__1   
                       |                          
                       +-- Lampsilis_cardium__2   

Graf & Cummings, 2006 [Publ 1968], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 46 7 7 --
  +-------------- Obliquaria             
  |                                      
  +-------------- Truncilla              
--+                                      
  +-------------- Ptychobranchus         
  |                                      
  |  +----------- Ligumia                
  +--+                                   
     |  +-------- Cambarunio             
     +--+                                
        |  +----- Epioblasma             
        +--+                             
           |  +-- Lampsilis              
           +--+                          
              +-- Ortmanniana_ligamentina **

Walker et al., 2006 [Publ 2666], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 14 4 4 --
  +----------- Potamilus              
--+                                   
  |  +-------- Cyrtonaias             
  +--+                                
     |  +----- Ligumia                
     +--+                             
        |  +-- Ortmanniana_ligamentina **
        +--+                          
           +-- Lampsilis              

Grobler et al., 2011 [Publ 3359], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 16 15 9 --
        +-------- Lemiox                 
  +-----+                                
  |     |  +----- Medionidus             
  |     +--+                             
  |        |  +-- Dromus                 
  |        +--+                          
--+           +-- Cyprogenia             
  |                                      
  |  +----------- Potamilus              
  |  |                                   
  |  |     +----- Lampsilis_siliquoidea  
  |  |  +--+                             
  +--+  |  |  +-- Lampsilis_higginsii    
     |  |  +--+                          
     |  |     +-- Ortmanniana_ligamentina **
     |  |                                
     |  |  +----- Lampsilis_fasciola     
     +--+--+                             
        |  |  +-- Lampsilis_cardium      
        |  +--+                          
        |     +-- Lampsilis_ovata        
        |                                
        |     +-- Cambarunio             
        |     |                          
        +-----+-- Ligumia                
              |                          
              +-- Epioblasma             

Huang et al., 2019 [Publ 3843], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 197 13 11 --
  +-------------- Potamilus_leptodon     
  |                                      
  |  +----------- Potamilus_alatus       
--+  |                                   
  |  |  +-------- Venustaconcha          
  |  |  |                                
  +--+  +-------- Leaunio                
     |  |                                
     |  |     +-- Ellipsaria             
     |  +-----+                          
     +--+     +-- Truncilla              
        |                                
        |     +-- Medionidus             
        +-----+                          
        |     +-- Ptychobranchus         
        |                                
        |  +----- Cambarunio             
        +--+                             
           |  +-- Ortmanniana_ligamentina **
           +--+                          
              +-- Lampsilis              

Huang et al., 2019 [Publ 3843], fig. S3

iT tT bi riS oT roT roS iD
1 1 0 1 195 13 11 --
  +-------------------- Potamilus_leptodon     
  |                                            
--+  +----------------- Potamilus_alatus       
  |  |                                         
  |  |              +-- Medionidus             
  +--+  +-----------+                          
     |  |           +-- Ptychobranchus         
     |  |                                      
     +--+           +-- Ellipsaria             
        |  +--------+                          
        |  |        +-- Truncilla              
        +--+                                   
           |  +-------- Leaunio                
           |  |                                
           +--+     +-- Venustaconcha          
              |  +--+                          
              |  |  +-- Cambarunio             
              +--+                             
                 |  +-- Ortmanniana_ligamentina **
                 +--+                          
                    +-- Lampsilis              

Bolotov et al., 2017 [Publ 3761], fig. S2

iT tT bi riS oT roT roS iD
1 1 0 1 138 2 2 --
  +----- Cambarunio             
--+                             
  |  +-- Ortmanniana_ligamentina **
  +--+                          
     +-- Lampsilis              

Bolotov et al., 2017 [Publ 3764], fig. S1

iT tT bi riS oT roT roS iD
1 1 0 1 124 2 2 --
-- Ortmanniana_ligamentina

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iT tT bi riS oT roT roS iD
1 1 0 1 79 2 2 --
  +----- Cambarunio             
--+                             
  |  +-- Lampsilis              
  +--+                          
     +-- Ortmanniana_ligamentina **

Szumowski et al., 2012 [Publ 3569], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 15 14 9 --
  +----------------------- Ellipsaria             
  |                                               
--+  +-------------------- Truncilla              
  |  |                                            
  +--+  +----------------- Obliquaria             
     |  |                                         
     |  |  +-------------- Potamilus              
     +--+  |                                      
        |  |        +----- Lampsilis_ornata       
        |  |  +-----+                             
        |  |  |     |  +-- Lampsilis_cardium      
        +--+  |     +--+                          
           |  |        +-- Lampsilis_ovata        
           |  |                                   
           |  |  +-------- Lampsilis_teres        
           |  +--+                                
           +--+  |  +----- Lampsilis_siliquoidea  
              |  +--+                             
              |     |  +-- Ortmanniana_ligamentina **
              |     +--+                          
              |        +-- Lampsilis_higginsii    
              |                                   
              |     +----- Hamiota                
              |     |                             
              +-----+  +-- Epioblasma             
                    |  |                          
                    +--+-- Cambarunio             
                       |                          
                       +-- Ligumia                

Inoue et al., 2020 [Publ 3941], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 40 25 6 --
  +----------------------------------- Toxolasma              
  |                                                           
--+  +-------------------------------- Ellipsaria             
  |  |                                                        
  +--+  +----------------------------- Truncilla              
     |  |                                                     
     |  |  +-------------------------- Potamilus              
     +--+  |                                                  
        |  |  +----------------------- Leaunio                
        |  |  |                                               
        +--+  |     +----------------- Lampsilis_bracteata    
           |  |     |                                         
           |  |  +--+              +-- Lampsilis_ornata       
           |  |  |  |  +-----------+                          
           +--+  |  |  |           +-- Lampsilis_fasciola     
              |  |  +--+                                      
              |  |     |  +----------- Lampsilis_ovata        
              |  |     +--+                                   
              |  |        |  +-------- Lampsilis_cardium__3   
              |  |        +--+                                
              +--+           |  +----- Lampsilis_cardium__2   
                 |           +--+                             
                 |              |  +-- Lampsilis_satura       
                 |              +--+                          
                 |                 +-- Lampsilis_cardium__1   
                 |                                            
                 |     +-------------- Lampsilis_teres        
                 |     |                                      
                 |     |           +-- Lampsilis_higginsii    
                 +-----+  +--------+                          
                       |  |        +-- Ortmanniana_ligamentina **
                       |  |                                   
                       +--+        +-- Lampsilis_radiata      
                          |  +-----+                          
                          |  |     +-- Lampsilis_siliquoidea  
                          |  |                                
                          +--+     +-- Lampsilis_hydiana      
                             |  +--+                          
                             |  |  +-- Lampsilis_bergmanni    
                             +--+                             
                                |  +-- Lampsilis_virescens    
                                +--+                          
                                   +-- Lampsilis_straminea    

Porto-Hannes et al., 2021 [Publ 4227], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 10 6 2 --
  +----- Obovaria               
--+                             
  |  +-- Ortmanniana_ligamentina **
  +--+                          
     +-- Lampsilis              

Hoeh et al., 2009 [Publ 2716], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 45 7 7 --
  +----------------- Ptychobranchus         
--+                                         
  |  +-------------- Truncilla              
  +--+                                      
     |  +----------- Ortmanniana_ligamentina **
     +--+                                   
        |  +-------- Lampsilis              
        +--+                                
           |  +----- Cambarunio             
           +--+                             
              |  +-- Ligumia                
              +--+                          
                 +-- Epioblasma             

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 33 5 5 --
  +----- Potamilus              
--+                             
  |  +-- Ortmanniana_ligamentina **
  +--+                          
     +-- Ligumia                

Hoeh et al., 2009 [Publ 2716], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 46 7 7 --
  +----------------- Ptychobranchus         
--+                                         
  |  +-------------- Truncilla              
  +--+                                      
     |  +----------- Ortmanniana_ligamentina **
     +--+                                   
        |  +-------- Lampsilis              
        +--+                                
           |  +----- Cambarunio             
           +--+                             
              |  +-- Ligumia                
              +--+                          
                 +-- Epioblasma             

Walker et al., 2006 [Publ 2666], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 35 4 4 --
  +-------- Potamilus              
--+                                
  |  +----- Lampsilis              
  +--+                             
     |  +-- Ligumia                
     +--+                          
        +-- Ortmanniana_ligamentina **

Hoeh et al., 2002 [Publ 2673], fig. 1-F

iT tT bi riS oT roT roS iD
1 1 0 1 8 4 4 --
  +----------- Cyrtonaias             
--+                                   
  |  +-------- Ortmanniana_ligamentina **
  +--+                                
     |  +----- Lampsilis              
     +--+                             
        |  +-- Potamilus              
        +--+                          
           +-- Ligumia                

Hoeh et al., 2002 [Publ 2673], fig. 1-M

iT tT bi riS oT roT roS iD
1 1 0 1 8 4 4 --
  +----------- Potamilus              
--+                                   
  |  +-------- Cyrtonaias             
  +--+                                
     |  +----- Ligumia                
     +--+                             
        |  +-- Lampsilis              
        +--+                          
           +-- Ortmanniana_ligamentina **

Walker et al., 2006 [Publ 2666], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 14 4 4 --
  +----------- Cyrtonaias             
--+                                   
  |  +-------- Potamilus              
  +--+                                
     |  +----- Ligumia                
     +--+                             
        |  +-- Ortmanniana_ligamentina **
        +--+                          
           +-- Lampsilis              

Stodola et al., 2021 [Publ 4266], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 9 9 1 --
              +-- Ortmanniana_ligamentina **
              |                          
  +-----------+-- Lampsilis_abrupta      
  |           |                          
  |           +-- Lampsilis_higginsii    
--+                                      
  |  +----------- Lampsilis_sietmani     
  |  |                                   
  |  |        +-- Lampsilis_siliquoidea  
  +--+  +-----+                          
     |  |     +-- Lampsilis_radiata      
     |  |                                
     +--+  +----- Lampsilis_hydiana__2   
        |  |                             
        |  +----- Lampsilis_straminea__2 
        |  |                             
        +--+  +-- Lampsilis_hydiana__1   
           +--+                          
           |  +-- Lampsilis_bergmanni    
           |                             
           |  +-- Lampsilis_straminea__1 
           +--+                          
              +-- Lampsilis_virescens    

Chapman et al., 2008 [Publ 2667], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 20 12 10 --
  +-------------------------- Toxolasma              
  |                                                  
--+  +----------------------- Obliquaria             
  |  |                                               
  |  |                    +-- Cyrtonaias             
  +--+  +-----------------+                          
     |  |                 +-- Glebula                
     |  |                                            
     +--+                 +-- Ptychobranchus         
        |  +--------------+                          
        |  |              +-- Lemiox                 
        |  |                                         
        +--+              +-- Venustaconcha          
           |  +-----------+                          
           |  |           +-- Obovaria               
           +--+                                      
              |  +----------- Hamiota                
              +--+                                   
                 |  +-------- Lampsilis_ovata        
                 +--+                                
                    |  +----- Ortmanniana_ligamentina **
                    +--+                             
                       |  +-- Lampsilis_straminea    
                       +--+                          
                          +-- Lampsilis_hydiana      

Boyer et al., 2011 [Publ 2718], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 34 15 9 --
  +----------- Toxolasma              
  |                                   
  |  +-------- Obliquaria             
--+  |                                
  |  |  +----- Obovaria               
  |  |  |                             
  |  |  +----- Lampsilis_cardium      
  +--+  |                             
     |  +----- Potamilus              
     |  |                             
     |  |  +-- Epioblasma             
     |  +--+                          
     |  |  +-- Lampsilis_siliquoidea  
     +--+                             
        |  +-- Ortmanniana_ligamentina **
        +--+                          
        |  +-- Lampsilis_higginsii    
        |                             
        |  +-- Ligumia                
        +--+                          
        |  +-- Lampsilis_teres        
        |                             
        |  +-- Ellipsaria             
        +--+                          
           +-- Truncilla              

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

iT tT bi riS oT roT roS iD
1 1 0 1 30 5 5 --
           +-- Cyrtonaias             
  +--------+                          
  |        +-- Glebula                
--+                                   
  |  +-------- Toxolasma              
  +--+                                
     |  +----- Ortmanniana_ligamentina **
     +--+                             
        |  +-- Ligumia                
        +--+                          
           +-- Potamilus              

Chapman et al., 2008 [Publ 2744], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 20 12 10 --
  +-------------------------- Toxolasma              
  |                                                  
--+  +----------------------- Obliquaria             
  |  |                                               
  |  |                    +-- Cyrtonaias             
  +--+  +-----------------+                          
     |  |                 +-- Glebula                
     |  |                                            
     +--+                 +-- Ptychobranchus         
        |  +--------------+                          
        |  |              +-- Lemiox                 
        |  |                                         
        +--+              +-- Venustaconcha          
           |  +-----------+                          
           |  |           +-- Obovaria               
           +--+                                      
              |  +----------- Hamiota                
              +--+                                   
                 |  +-------- Lampsilis_ovata        
                 +--+                                
                    |  +----- Ortmanniana_ligamentina **
                    +--+                             
                       |  +-- Lampsilis_straminea    
                       +--+                          
                          +-- Lampsilis_hydiana      

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

iT tT bi riS oT roT roS iD
1 1 0 1 29 11 8 --
  +-------------------- Truncilla              
  |                                            
--+  +----------------- Ligumia                
  |  |                                         
  |  |              +-- Ptychobranchus         
  +--+  +-----------+                          
     |  |           +-- Sagittunio             
     |  |                                      
     +--+  +----------- Lampsilis_siliquoidea  
        |  |                                   
        |  |     +----- Ortmanniana_ligamentina **
        +--+  +--+                             
           |  |  |  +-- Lampsilis_cardium      
           |  |  +--+                          
           +--+     +-- Lampsilis_fasciola     
              |                                
              |  +----- Epioblasma             
              +--+                             
                 |  +-- Cambarunio             
                 +--+                          
                    +-- Leaunio                

Hoeh et al., 2009 [Publ 2716], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 45 6 6 --
  +----- Truncilla              
  |                             
  +----- Ptychobranchus         
--+                             
  |  +-- Ortmanniana_ligamentina **
  |  |                          
  |  +-- Cambarunio             
  +--+                          
     +-- Ligumia                
     |                          
     +-- Epioblasma             

Bolotov et al., 2017 [Publ 3761], fig. S1

iT tT bi riS oT roT roS iD
1 1 0 1 138 2 2 --
  +----- Ortmanniana_ligamentina **
--+                             
  |  +-- Lampsilis              
  +--+                          
     +-- Cambarunio             

Bolotov et al., 2020 [Publ 4084], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 35 2 2 --
-- Ortmanniana_ligamentina

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

iT tT bi riS oT roT roS iD
1 1 0 1 30 5 5 --
  +----------- Ortmanniana_ligamentina **
--+                                   
  |  +-------- Glebula                
  +--+                                
     |  +----- Potamilus              
     +--+                             
        |  +-- Ligumia                
        +--+                          
           +-- Toxolasma              

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iT tT bi riS oT roT roS iD
1 1 0 1 75 2 2 --
  +----- Ortmanniana_ligamentina **
--+                             
  |  +-- Lampsilis              
  +--+                          
     +-- Cambarunio             

Pfeiffer et al., 2019 [Publ 3866], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 34 11 10 --
        +-- Lampsilis              
  +-----+                          
  |     +-- Ortmanniana_ligamentina **
--+                                
  |  +----- Cambarunio             
  +--+                             
     |  +-- Epioblasma             
     +--+                          
        +-- Ligumia                

Inoue et al., 2020 [Publ 4128], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 25 11 10 --
-- Ortmanniana_ligamentina

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