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Page last updated
Thu 05 Sep 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Gonideinae | Contradentini

genus Trapezoideus Simpson, 1900: cladomics
     5 valid species

Cladomics Summary

number of species = 5

analyzed in 25 cladograms in 16 publications
maximum number of terminals in an analysis = 5, mean = 1.52

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 7
maximum number of included terminals = 5, mean = 2.86
proportion of cladograms that support basic monophyly = 1.0

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ tree

Konopleva et al., 2023 [Publ 4262], fig. 8

iT tT bi riS oT roT roS iD
5 5 3 5 77 14 3 0.0

*** Top Monophyly Cladogram *** 

              +----- Physunio_modelli          
        +-----+                                
        |     |  +-- Physunio_superbus         
        |     +--+                             
        |        +-- Physunio_massini          
  +-----+                                      
  |     |        +-- Trapezoideus_thachiadensis **
  |     |  +-----+                             
  |     |  |     +-- Trapezoideus_subviridis **   
  |     +--+                                   
  |        |  +----- Trapezoideus_mitanensis **   
  |        +--+                                
  |           |  +-- Trapezoideus_foliaceus **    
--+           +--+                             
  |              +-- Trapezoideus_lenya **        
  |                                            
  |              +-- Yaukthwa_nesemanni        
  |  +-----------+                             
  |  |           +-- Yaukthwa_elongatula       
  |  |                                         
  |  |        +----- Lens_misellus             
  +--+  +-----+                                
     |  |     |  +-- Lens_contradens           
     |  |     +--+                             
     |  |        +-- Lens_micropterus          
     +--+                                      
        |     +----- Lens_pallegoixi           
        |  +--+                                
        |  |  |  +-- Lens_novoselovi           
        |  |  +--+                             
        +--+     +-- Lens_comptus              
           |                                   
           |  +----- Lens_maenamensis          
           +--+                                
              |  +-- Lens_eximius              
              +--+                             
                 +-- Lens_rolfbrandti

Pfeiffer et al., 2021 [Publ 4126], fig. 9

iT tT bi riS oT roT roS iD
2 2 0 2 27 13 4 --

*** Top Sister-Group Cladogram ***

                    +-- Trapezoideus_subviridis **
              +-----+                          
              |     +-- Trapezoideus_foliaceus ** 
  +-----------+                                
  |           |  +----- Physunio_superbus      
  |           +--+                             
  |              |  +-- Physunio_modelli       
  |              +--+                          
--+                 +-- Physunio_pellucidus    
  |                                            
  |  +----------------- Yaukthwa_paiensis      
  |  |                                         
  |  |  +-------------- Solenaia_emarginata    
  +--+  |                                      
     |  |     +-------- Lens_misellus          
     |  |  +--+                                
     +--+  |  |  +----- Lens_inornatus         
        |  |  +--+                             
        |  |     |  +-- Lens_contradens        
        |  |     +--+                          
        +--+        +-- Lens_micropterus       
           |                                   
           |        +-- Lens_maenamensis       
           |     +--+                          
           |     |  +-- Lens_eximius           
           +-----+                             
                 |  +-- Lens_pallegoixi        
                 +--+                          
                    +-- Lens_comptus

Pfeiffer et al., 2021 [Publ 4126], fig. 18

iT tT bi riS oT roT roS iD
3 3 1 3 5 5 2 4.0 
  +-------------- Yaukthwa_paiensis      
  |                                      
  |        +----- Trapezoideus_foliaceus ** 
--+  +-----+                             
  |  |     |  +-- Trapezoideus_lenya **     
  |  |     +--+                          
  |  |        +-- Trapezoideus_subviridis **
  +--+                                   
     |        +-- Physunio_massini       
     |  +-----+                          
     |  |     +-- Physunio_superbus      
     +--+                                
        |  +----- Physunio_modelli__3    
        |  |                             
        +--+  +-- Physunio_modelli__1    
           |  |                          
           +--+-- Physunio_modelli__2    
              |                          
              +-- Physunio_pellucidus

Bolotov et al., 2022 [Publ 4250], fig. S5

iT tT bi riS oT roT roS iD
3 3 1 3 12 12 2 4.0 
                 +-- Physunio_modelli       
           +-----+                          
           |     +-- Physunio_superbus      
  +--------+                                
  |        |  +----- Trapezoideus_foliaceus ** 
  |        +--+                             
  |           |  +-- Trapezoideus_lenya **     
  |           +--+                          
  |              +-- Trapezoideus_mitanensis **
--+                                         
  |              +-- Yaukthwa_elongatula    
  |  +-----------+                          
  |  |           +-- Yaukthwa_baniensis     
  |  |                                      
  |  |     +-------- Yaukthwa_panhai        
  +--+  +--+                                
     |  |  |  +----- Yaukthwa_rectangularis 
     |  |  +--+                             
     |  |     |  +-- Yaukthwa_nesemanni     
     +--+     +--+                          
        |        +-- Yaukthwa_zayleymanensis
        |                                   
        |     +----- Yaukthwa_dalliana      
        |     |                             
        +-----+  +-- Yaukthwa_paiensis      
              |  |                          
              +--+-- Yaukthwa_inlenensis    
                 |                          
                 +-- Yaukthwa_avaensis

Bolotov et al., 2022 [Publ 4250], fig. 8

iT tT bi riS oT roT roS iD
3 3 1 3 46 9 1 2.0 
              +----- Trapezoideus_mitanensis **
  +-----------+                             
  |           |  +-- Trapezoideus_foliaceus ** 
  |           +--+                          
  |              +-- Trapezoideus_lenya **     
--+                                         
  |              +-- Yaukthwa_elongatula    
  |  +-----------+                          
  |  |           +-- Yaukthwa_baniensis     
  |  |                                      
  +--+        +----- Yaukthwa_inlenensis    
     |  +-----+                             
     |  |     |  +-- Yaukthwa_paiensis      
     |  |     +--+                          
     +--+        +-- Yaukthwa_avaensis      
        |                                   
        |  +-------- Yaukthwa_panhai        
        +--+                                
           |  +----- Yaukthwa_rectangularis 
           +--+                             
              |  +-- Yaukthwa_nesemanni     
              +--+                          
                 +-- Yaukthwa_zayleymanensis

Bolotov et al., 2020 [Publ 4100], fig. S1

iT tT bi riS oT roT roS iD
2 2 0 2 110 21 3 -- 
                    +----- Physunio               
  +-----------------+                             
  |                 |  +-- Physunio_modelli       
  |                 +--+                          
  |                    +-- Physunio_superbus      
  |                                               
  |                    +-- Trapezoideus_lenya **     
  |  +-----------------+                          
--+  |                 +-- Trapezoideus_foliaceus ** 
  |  |                                            
  |  |              +----- Lens_misellus          
  |  |        +-----+                             
  |  |        |     |  +-- Lens_contradens        
  |  |        |     +--+                          
  |  |        |        +-- Lens_micropterus       
  +--+  +-----+                                   
     |  |     |     +----- Lens_maenamensis       
     |  |     |  +--+                             
     |  |     |  |  |  +-- Lens_eximius           
     |  |     |  |  +--+                          
     |  |     +--+     +-- Lens_rolfbrandti       
     |  |        |                                
     |  |        |  +----- Lens_pallegoixi        
     +--+        +--+                             
        |           |  +-- Lens_novoselovi        
        |           +--+                          
        |              +-- Lens_comptus           
        |                                         
        |              +-- Yaukthwa_elongatula    
        |  +-----------+                          
        |  |           +-- Yaukthwa_baniensis     
        |  |                                      
        +--+        +----- Yaukthwa_inlenensis    
           |  +-----+                             
           |  |     |  +-- Yaukthwa_avaensis      
           |  |     +--+                          
           +--+        +-- Yaukthwa_paiensis      
              |                                   
              |  +-------- Yaukthwa_panhai        
              +--+                                
                 |  +----- Yaukthwa_rectangularis 
                 +--+                             
                    |  +-- Yaukthwa_zayleymanensis
                    +--+                          
                       +-- Yaukthwa_nesemanni

Bolotov et al., 2020 [Publ 4100], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 116 21 3 -- 
                    +-- Trapezoideus_lenya **     
              +-----+                          
              |     +-- Trapezoideus_foliaceus ** 
  +-----------+                                
  |           |  +----- Physunio_superbus      
  |           +--+                             
  |              |  +-- Physunio_modelli       
  |              +--+                          
  |                 +-- Physunio               
  |                                            
  |                 +-- Yaukthwa_elongatula    
  |     +-----------+                          
  |     |           +-- Yaukthwa_baniensis     
--+     |                                      
  |  +--+        +----- Yaukthwa_inlenensis    
  |  |  |  +-----+                             
  |  |  |  |     |  +-- Yaukthwa_avaensis      
  |  |  |  |     +--+                          
  |  |  +--+        +-- Yaukthwa_paiensis      
  |  |     |                                   
  |  |     |  +-------- Yaukthwa_panhai        
  |  |     +--+                                
  |  |        |  +----- Yaukthwa_rectangularis 
  +--+        +--+                             
     |           |  +-- Yaukthwa_zayleymanensis
     |           +--+                          
     |              +-- Yaukthwa_nesemanni     
     |                                         
     |           +----- Lens_misellus          
     |     +-----+                             
     |     |     |  +-- Lens_contradens        
     |     |     +--+                          
     |     |        +-- Lens_micropterus       
     +-----+                                   
           |     +----- Lens_pallegoixi        
           |  +--+                             
           |  |  |  +-- Lens_comptus           
           |  |  +--+                          
           +--+     +-- Lens_novoselovi        
              |                                
              |  +----- Lens_maenamensis       
              +--+                             
                 |  +-- Lens_eximius           
                 +--+                          
                    +-- Lens_rolfbrandti

Konopleva et al., 2019 [Publ 4041], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 33 18 3 -- 
        +-------- Trapezoideus_foliaceus **
  +-----+                               
  |     |  +----- Physunio_superbus     
  |     +--+                            
  |        |  +-- Physunio_modelli      
  |        +--+                         
  |           +-- Physunio              
--+                                     
  |        +----- Lens_misellus         
  |  +-----+                            
  |  |     |  +-- Lens_contradens       
  |  |     +--+                         
  |  |        +-- Lens_micropterus      
  +--+                                  
     |     +----- Lens_pallegoixi       
     |  +--+                            
     |  |  |  +-- Lens_novoselovi       
     |  |  +--+                         
     +--+     +-- Lens_comptus          
        |                               
        |  +----- Lens_maenamensis      
        +--+                            
           |  +-- Lens_eximius          
           +--+                         
              +-- Lens_rolfbrandti

Jeratthitikul et al., 2022 [Publ 4260], fig. S2, 2

iT tT bi riS oT roT roS iD
1 1 0 1 75 2 2 -- 
  +----- Lens_contradens       
--+                            
  |  +-- Trapezoideus_foliaceus **
  +--+                         
     +-- Physunio_superbus

Zieritz et al., 2024 [Publ 4381], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 45 7 3 -- 
           +-- Physunio_superbus     
  +--------+                         
  |        +-- Trapezoideus_foliaceus **
  |                                  
--+     +----- Yaukthwa_elongatula   
  |  +--+                            
  |  |  |  +-- Yaukthwa_inlenensis   
  |  |  +--+                         
  +--+     +-- Yaukthwa_paiensis     
     |                               
     |  +----- Lens_eximius          
     +--+                            
        |  +-- Lens_contradens       
        +--+                         
           +-- Lens_lugens

Wu et al., 2024 [Publ 4350], fig. 6-7

iT tT bi riS oT roT roS iD
1 1 0 1 36 6 3 -- 
           +-- Physunio_superbus     
  +--------+                         
  |        +-- Trapezoideus_foliaceus **
  |                                  
--+        +-- Lens_contradens       
  |  +-----+                         
  |  |     +-- Lens_eximius          
  +--+                               
     |  +----- Yaukthwa_elongatula   
     +--+                            
        |  +-- Yaukthwa_inlenensis   
        +--+                         
           +-- Yaukthwa_paiensis

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 35 2 2 -- 
  +----- Lens_contradens       
--+                            
  |  +-- Trapezoideus_foliaceus **
  +--+                         
     +-- Physunio_superbus

Konopleva et al., 2019 [Publ 3900], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 51 15 3 -- 
                 +----- Physunio_superbus     
  +--------------+                            
  |              |  +-- Physunio_modelli      
  |              +--+                         
--+                 +-- Physunio              
  |                                           
  |  +----------------- Trapezoideus_foliaceus **
  |  |                                        
  +--+              +-- Lens_contradens       
     |  +-----------+                         
     |  |           +-- Lens_micropterus      
     +--+                                     
        |  +----------- Lens_misellus         
        +--+                                  
           |  +-------- Lens_eximius          
           +--+                               
              |  +----- Lens_pallegoixi       
              +--+                            
                 |  +-- Lens_comptus          
                 +--+                         
                    +-- Lens_novoselovi

Pfeiffer et al., 2018 [Publ 3827], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 6 3 2 -- 
  +----- Physunio       
  |                     
--+----- Trapezoideus **   
  |                     
  |  +-- Lens_comptus   
  +--+                  
     +-- Lens_contradens

Zieritz et al., 2021 [Publ 4218], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 31 6 3 -- 
  +-------------- Physunio_superbus     
  |                                     
--+  +----------- Trapezoideus_foliaceus **
  |  |                                  
  +--+        +-- Lens_contradens       
     |  +-----+                         
     |  |     +-- Lens_eximius          
     +--+                               
        |  +----- Yaukthwa_elongatula   
        +--+                            
           |  +-- Yaukthwa_inlenensis   
           +--+                         
              +-- Yaukthwa_paiensis

Wu et al., 2024 [Publ 4350], fig. 5a

iT tT bi riS oT roT roS iD
1 1 0 1 36 6 3 -- 
           +-- Physunio_superbus     
  +--------+                         
  |        +-- Trapezoideus_foliaceus **
  |                                  
--+        +-- Lens_contradens       
  |  +-----+                         
  |  |     +-- Lens_eximius          
  +--+                               
     |  +----- Yaukthwa_elongatula   
     +--+                            
        |  +-- Yaukthwa_inlenensis   
        +--+                         
           +-- Yaukthwa_paiensis

Zieritz et al., 2020 [Publ 4102], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 30 13 3 -- 
                 +-- Yaukthwa_paiensis     
           +-----+                         
           |     +-- Yaukthwa_inlenensis   
  +--------+                               
  |        |  +----- Yaukthwa_panhai       
  |        +--+                            
  |           |  +-- Yaukthwa_rectangularis
--+           +--+                         
  |              +-- Yaukthwa_nesemanni    
  |                                        
  |  +-------------- Trapezoideus_foliaceus **
  |  |                                     
  +--+           +-- Lens_contradens       
     |  +--------+                         
     |  |        +-- Lens_misellus         
     +--+                                  
        |  +-------- Lens_eximius          
        +--+                               
           |  +----- Lens_pallegoixi       
           +--+                            
              |  +-- Lens_comptus          
              +--+                         
                 +-- Lens_novoselovi

Jeratthitkul et al., 2021 [Publ 4212], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 20 9 3 -- 
        +-------- Trapezoideus_foliaceus **
  +-----+                               
  |     |  +----- Physunio_superbus     
  |     +--+                            
  |        |  +-- Physunio_pellucidus   
  |        +--+                         
--+           +-- Physunio_modelli      
  |                                     
  |           +-- Yaukthwa_nesemanni    
  |  +--------+                         
  |  |        +-- Yaukthwa_panhai       
  +--+                                  
     |  +-------- Lens_contradens       
     +--+                               
        |  +----- Lens_novoselovi       
        +--+                            
           |  +-- Lens_rolfbrandti      
           +--+                         
              +-- Lens_maenamensis

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 35 2 2 -- 
  +----- Lens_contradens       
--+                            
  |  +-- Trapezoideus_foliaceus **
  +--+                         
     +-- Physunio_superbus

Wu et al., 2024 [Publ 4350], fig. 5c

iT tT bi riS oT roT roS iD
1 1 0 1 36 6 3 -- 
  +----- Physunio_superbus     
  |                            
  +----- Yaukthwa_paiensis     
  |                            
  +----- Yaukthwa_inlenensis   
--+                            
  +----- Trapezoideus_foliaceus **
  |                            
  +----- Yaukthwa_elongatula   
  |                            
  |  +-- Lens_contradens       
  +--+                         
     +-- Lens_eximius

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iT tT bi riS oT roT roS iD
1 1 0 1 56 4 4 -- 
        +-- Trapezoideus_foliaceus **
  +-----+                         
  |     +-- Physunio_superbus     
--+                               
  |  +----- Yaukthwa_paiensis     
  +--+                            
     |  +-- Lens_contradens       
     +--+                         
        +-- Solenaia_emarginata

Liu et al., 2024 [Publ 4380], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 36 6 3 -- 
           +-- Physunio_superbus     
  +--------+                         
  |        +-- Trapezoideus_foliaceus **
  |                                  
--+        +-- Lens_contradens       
  |  +-----+                         
  |  |     +-- Lens_eximius          
  +--+                               
     |  +----- Yaukthwa_elongatula   
     +--+                            
        |  +-- Yaukthwa_inlenensis   
        +--+                         
           +-- Yaukthwa_paiensis

Wu et al., 2024 [Publ 4350], fig. 5b

iT tT bi riS oT roT roS iD
1 1 0 1 36 6 3 -- 
  +-------------- Yaukthwa_elongatula   
  |                                     
--+  +----------- Trapezoideus_foliaceus **
  |  |                                  
  +--+        +-- Physunio_superbus     
     |  +-----+                         
     |  |     +-- Yaukthwa_inlenensis   
     +--+                               
        |  +----- Yaukthwa_paiensis     
        +--+                            
           |  +-- Lens_contradens       
           +--+                         
              +-- Lens_eximius

Jeratthitikul et al., 2022 [Publ 4260], fig. S1, 1

iT tT bi riS oT roT roS iD
1 1 0 1 75 2 2 -- 
  +----- Lens_contradens       
--+                            
  |  +-- Trapezoideus_foliaceus **
  +--+                         
     +-- Physunio_superbus

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 56 4 4 -- 
        +-- Trapezoideus_foliaceus **
  +-----+                         
  |     +-- Physunio_superbus     
--+                               
  |  +----- Yaukthwa_paiensis     
  +--+                            
     |  +-- Lens_contradens       
     +--+                         
        +-- Solenaia_emarginata

Page updated: 13:22:35 Thu 05 Sep 2024
 
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