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Page last updated
Wed 04 Dec 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

This page documents data for Wu et al. (2024), such as genera and species attributed to that publication (including known misspellings, misidentifications, and other unavailable names) as well as taxonomic opinions, if available. Taxonomic opinions refer to uses of genera and species as valid and invalid in that work.

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Publication information

Wu, R., L. Zhang, L. Liu, J. Jia & X. Liu. 2024. Unraveling the phylogenetic relationships and taxonomic status of a puzzling freshwater mussel genus Inversidens (Bivalvia, Unionidae) through multilocus phylogeny and mitochondrial phylogenomics. Journal of Zoological Systematics and Evolutionary Research 2024, 1499508: 1-18.

Click here for online access.

Cladogram Figures

(click the figure number to see the cladogram)

fig. 5a
fig. 5b
fig. 5c
fig. 6-7
fig. 8a_S1
fig. 8b_S2

Attributed Species Names

Unio crassus Soroka, 2005
‘Philippson’ of Authors.
Soroka, 2005, Zool. Sci. 22: 1138, Table 1.
Soroka, 2010, Folia Malac. 18(4): Table 2. [in part]
Reis et al., 2013, Graellsia 69(1): 22, Table 2.
Prié & Puillandre, 2014, Hydrobiologia 735(1): online resource 1 [as ‘Unio crassus crassus’].
Douda et al., 2014, J. Appl. Ecol. 51: 1087. [in part]
Froufe et al., 2016, Org Divers Evol 16(3): 606, fig. 3. [in part]
Klishko et al., 2017, Zootaxa 4286(1): 98, Table 3.
Soroka & Burzynski, 2017, mtDNA B 2(2): 863.
Burzynski et al., 2017, MPE 107: 606.
Lopes-Lima et al., 2017, Biol. Rev. 92(1): Table S1 [as ‘Unio crassus crassus’]. [in part]
Araujo et al., 2018, J. Linn. Soc., Zool. 182(2): supplementary table. [in part]
Klishko et al., 2018, Systemat. Biodiv. 16(3): 291.
Burzynski & Soroka, 2018, PeerJ 6, e5573: 3.
Wang et al., 2018, Conserv. Genet. Resour. 10(4): 611.
Wang et al., 2018, Conserv. Genet. Resour. 11: 140.
Cha et al., 2018, mtDNA B 3(2): 946.
Aunins et al., 2018, mtDNA B 3(2): 1125.
Wu et al., 2018, PLoS One 13(8), e0200956: 6. [in part]
Klishko et al., 2019, Ruthenica 29(1): 57, Table 1.
Wu et al., 2019, ZooKeys 812: 29.
Huang et al., 2019, MPE 130: Table S5. [in part]
Fukata & Iigo, 2019, mtDNA B 4(2): 3489.
Guerra et al., 2019, BMC Evol. Biol. 19 (229): Table S1.
Froufe et al., 2019, Heredity 124: 185.
Soroka, 2020, J. Zool. Syst. Evol. Res. 58(2): Table 2.
Fukata & Iigo, 2020, mtDNA B 5(2): 1216.
Sano et al., 2020, Plankton Benthos Res. 15(2): 99.
Wu et al., 2020, Invert. Syst. 34: Table S2.
Kilikowska et al., 2020, Life 10(7): 1. [in part]
Lopes-Lima et al., 2021, MPE 163: fig. 6 [by voucher number]. [in part]
Sayenko et al., 2021, Systemat. Biodiv. 19: 58, Table 1.
Wu et al., 2021, J. Lake Sci. 33(6): attached Table 2.
Zieritz et al., 2021, Zool. J. Linn. Soc. 193(3): 1092.
Kim et al., 2021, mtDNA B 6(1): 80.
Pfarrer et al., 2022, Hydrobiologia 849: ESM1.
Wu et al., 2022, Zool. Scripta 51: Table S2.
Dai et al., 2023, Invert. Syst. 37(2): Table 2.
Chen et al., 2023, ZooKeys 1173: Table 1.
Wu et al., 2023, Org Divers Evol (2023): Table 2.
Dai et al., 2024, Zool. Scripta 53: Table 1.
Wu et al., 2024, J. Zool. Syst. Evol. Res. 2024, 1499508: Table 2.
Wu et al., 2024, Zool. Scripta early online: Table S3.
Dai et al., 2024, Invert. Syst. 38(6): Table 3. [in part]
(misidentified reference to Unio crassus Retzius, 1788)

Lamellidens marginalis Bolotov et al., 2017
‘Lamarck’ of Authors.
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Lamellidens marginalis sp. 4’].
Konopleva et al., 2017, Systemat. Biodiv. 15(3): 207. [in part]
Pavan-Kumar et al., 2022, Mol. Biol. Rep. 49: Table S1.
Wu et al., 2023, Org Divers Evol (2023): Table 2.
Wu et al., 2024, J. Zool. Syst. Evol. Res. 2024, 1499508: Table 2.
(misidentified reference to Unio marginalis Lamarck, 1819)

Taxonomic Opinions

Aculamprotula coreana

Aculamprotula polysticta

Aculamprotula scripta

Aculamprotula tientsinensis

Aculamprotula tortuosa

Acuticosta chinensis

Alasmidonta varicosa

Amblema plicata

Anemina arcaeformis

Anemina euscaphys

Anodonta anatina

Anodonta cygnea

Anodonta exulcerata

Anodonta nuttalliana

Anodontites trapesialis

Beringiana fukuharai

Bineurus anodontinum (Rochebrune, 1882)

Bineurus loeiensis Konopleva et al., 2021

Chamberlainia somsakpanhai Kongim et al., 2023

Cristaria plicata

Cuneopsis celtiformis

Cuneopsis demangei

Cuneopsis heudei

Cuneopsis rufescens

Echyridella menziesii

Elliptio complanata

Ensidens sagittarius (Lea, 1856)

Gonidea angulata (Lea, 1838)

Hyriopsis bialata Simpson, 1900

Hyriopsis desowitzi Brandt, 1974

Indopseudodon bogani (Bolotov et al., 2017)

Indopseudodon kayinensis (Bolotov et al., 2020)
+ Pseudodon crebristriatus Pfeiffer et al., 2019

Inversidens brandtii (Kobelt, 1879)

Inversidens rentianensis Wu & Wu, 2021

Khairuloconcha lunbawangorum Zieritz et al., 2021

Khairuloconcha sahanae Zieritz et al., 2021

Koreosolenaia sitgyensis Lee et al., 2020

Lamellidens marginalis

Lamprotula caveata

Lamprotula cornuumlunae

Lamprotula leaii (Gray, 1833)

Lampsilis cardium

Lampsilis cariosa

Lampsilis ornata

Lampsilis powellii

Lampsilis siliquoidea

Lanceolaria gladiola

Lanceolaria grayii

Lanceolaria lanceolata

Lasmigona complanata

Leaunio lienosus

Lens contradens (Lea, 1838)

Lens eximius (Lea, 1856)

Lepidodesma aligera

Lepidodesma languilati

Margaritifera dahurica (Middendorff, 1850)

Margaritifera margaritifera (Linnaeus, 1758)

Microcondylaea bonellii (FĂ©russac, 1827)

Mutela dubia

Neotrigonia margaritacea

Nodularia breviconcha

Nodularia douglasiae

Parvasolenaia rivularis (Heude, 1877)

Physunio superbus (Lea, 1843)

Pilsbryoconcha exilis (Lea, 1838)

Platynaias compressa

Pleurobema oviforme

Popenaias popeii

Postolata guangxiensis Dai et al., 2023

Potamilus alatus

Potamilus leptodon

Potamilus streckersoni

Potomida littoralis (Cuvier, 1798)

Prolasmidonta heterodon

Pronodularia japanensis

Pseudanodonta complanata

Pseudocuneopsis capitata

Pseudocuneopsis sichuanensis

Pseudodon mekongi (Bolotov et al., 2020)

Pseudodon vondembuschianus (Lea, 1840)

Ptychorhynchus pfisteri (Heude, 1874)

Pyganodon grandis

Quadrula quadrula

Rectidens sumatrensis (Dunker, 1852)

Schistodesmus lampreyanus

Schistodesmus sp.

Schistodesmus spinosus

Sinanodonta lucida

Sinanodonta tumens

Sinanodonta woodiana

Sinohyriopsis cumingii

Sinohyriopsis schlegelii

Sinosolenaia carinata (Heude, 1877)

Sinosolenaia iridinea

Sinosolenaia oleivora

Tchangsinaia piscicula

Thaiconcha callifera (Martens, 1860)

Toxolasma parvum

Trapezoideus foliaceus (Gould, 1843)

Unio crassus

Unio delphinus

Unio elongatulus

Unio mancus

Unio pictorum

Unio tumidus

Uniomerus tetralasmus

Utterbackia imbecillis

Utterbackia peninsularis

Venustaconcha ellipsiformis

Yaukthwa elongatula Bolotov et al., 2019

Yaukthwa inlenensis Konopleva et al., 2019

Yaukthwa paiensis Konopleva et al., 2019

Cladogram Figure 5a
                             +-- Margaritifera_dahurica      
  +--------------------------+                               
  |                          +-- Margaritifera_margaritifera 
  |                                                          
  |  +-------------------------- Chamberlainia_somsakpanhai  
  |  |                                                       
  |  |                       +-- Khairuloconcha_lunbawangorum
  |  |        +--------------+                               
--+  |        |              +-- Khairuloconcha_sahanae      
  |  |        |                                              
  |  |        |        +-------- Ensidens_sagittarius        
  |  |  +-----+  +-----+                                     
  |  |  |     |  |     |  +----- Rectidens_sumatrensis       
  |  |  |     |  |     +--+                                  
  |  |  |     |  |        |  +-- Hyriopsis_bialata           
  |  |  |     |  |        +--+                               
  +--+  |     +--+           +-- Hyriopsis_desowitzi         
     |  |        |                                           
     |  |        |           +-- Physunio_superbus           
     |  |        |  +--------+                               
     |  |        |  |        +-- Trapezoideus_foliaceus      
     |  |        |  |                                        
     |  |        +--+        +-- Lens_contradens             
     |  |           |  +-----+                               
     |  |           |  |     +-- Lens_eximius                
     |  |           +--+                                     
     |  |              |  +----- Yaukthwa_elongatula         
     |  |              +--+                                  
     +--+                 |  +-- Yaukthwa_inlenensis         
        |                 +--+                               
        |                    +-- Yaukthwa_paiensis           
        |                                                    
        |     +----------------- Schepmania_nieuwenhuisi     
        |     |                                              
        |  +--+              +-- Indopseudodon_bogani        
        |  |  |  +-----------+                               
        |  |  |  |           +-- Indopseudodon_kayinensis    
        |  |  +--+                                           
        |  |     |  +----------- Pilsbryoconcha_exilis       
        |  |     |  |                                        
        |  |     +--+        +-- Pseudodon_mekongi           
        |  |        |  +-----+                               
        |  |        |  |     +-- Pseudodon_vondembuschianus  
        |  |        +--+                                     
        |  |           |  +----- Thaiconcha_callifera        
        +--+           +--+                                  
           |              |  +-- Bineurus_anodontinum        
           |              +--+                               
           |                 +-- Bineurus_loeiensis          
           |                                                 
           |                 +-- Inversidens_brandtii        
           |           +-----+                               
           |           |     +-- Inversidens_rentianensis    
           |  +--------+                                     
           |  |        |  +----- Potomida_littoralis         
           |  |        +--+                                  
           |  |           |  +-- Lamprotula_caveata          
           +--+           +--+                               
              |              +-- Lamprotula_leaii            
              |                                              
              |  +-------------- Sinosolenaia_carinata       
              |  |                                           
              +--+           +-- Gonidea_angulata            
                 |  +--------+                               
                 |  |        +-- Microcondylaea_bonellii     
                 +--+                                        
                    |  +-------- Parvasolenaia_rivularis     
                    +--+                                     
                       |  +----- Koreosolenaia_sitgyensis    
                       +--+                                  
                          |  +-- Postolata_guangxiensis      
                          +--+                               
                             +-- Ptychorhynchus_pfisteri     
Cladogram Figure 5b
                                      +-- Margaritifera_dahurica      
  +-----------------------------------+                               
  |                                   +-- Margaritifera_margaritifera 
  |                                                                   
  |  +----------------------------------- Schepmania_nieuwenhuisi     
  |  |                                                                
  |  |                                +-- Indopseudodon_bogani        
--+  |                    +-----------+                               
  |  |                    |           +-- Indopseudodon_kayinensis    
  |  |  +-----------------+                                           
  |  |  |                 |  +----------- Pilsbryoconcha_exilis       
  |  |  |                 |  |                                        
  |  |  |                 +--+        +-- Bineurus_anodontinum        
  |  |  |                    |  +-----+                               
  +--+  |                    |  |     +-- Bineurus_loeiensis          
     |  |                    +--+                                     
     |  |                       |  +----- Thaiconcha_callifera        
     |  |                       +--+                                  
     |  |                          |  +-- Pseudodon_mekongi           
     |  |                          +--+                               
     |  |                             +-- Pseudodon_vondembuschianus  
     |  |                                                             
     |  |     +-------------------------- Hyriopsis_bialata           
     |  |     |                                                       
     +--+     |                    +----- Rectidens_sumatrensis       
        |  +--+  +-----------------+                                  
        |  |  |  |                 |  +-- Khairuloconcha_lunbawangorum
        |  |  |  |                 +--+                               
        |  |  +--+                    +-- Khairuloconcha_sahanae      
        |  |     |                                                    
        |  |     |  +-------------------- Ensidens_sagittarius        
        |  |     |  |                                                 
        |  |     +--+  +----------------- Hyriopsis_desowitzi         
        |  |        |  |                                              
        |  |        +--+  +-------------- Yaukthwa_elongatula         
        |  |           |  |                                           
        |  |           +--+  +----------- Trapezoideus_foliaceus      
        |  |              |  |                                        
        |  |              +--+        +-- Physunio_superbus           
        +--+                 |  +-----+                               
           |                 |  |     +-- Yaukthwa_inlenensis         
           |                 +--+                                     
           |                    |  +----- Yaukthwa_paiensis           
           |                    +--+                                  
           |                       |  +-- Lens_contradens             
           |                       +--+                               
           |                          +-- Lens_eximius                
           |                                                          
           |                          +-- Inversidens_brandtii        
           |        +-----------------+                               
           |        |                 +-- Inversidens_rentianensis    
           |        |                                                 
           |        |                 +-- Chamberlainia_somsakpanhai  
           +--------+              +--+                               
                    |              |  +-- Potomida_littoralis         
                    |  +-----------+                                  
                    |  |           |  +-- Lamprotula_caveata          
                    |  |           +--+                               
                    +--+              +-- Lamprotula_leaii            
                       |                                              
                       |  +-------------- Sinosolenaia_carinata       
                       |  |                                           
                       +--+           +-- Gonidea_angulata            
                          |  +--------+                               
                          |  |        +-- Microcondylaea_bonellii     
                          +--+                                        
                             |  +-------- Koreosolenaia_sitgyensis    
                             +--+                                     
                                |  +----- Parvasolenaia_rivularis     
                                +--+                                  
                                   |  +-- Postolata_guangxiensis      
                                   +--+                               
                                      +-- Ptychorhynchus_pfisteri     
Cladogram Figure 5c
                                +-- Margaritifera_dahurica      
  +-----------------------------+                               
  |                             +-- Margaritifera_margaritifera 
  |                                                             
  |  +----------------------------- Schepmania_nieuwenhuisi__1  
  |  |                                                          
  |  +----------------------------- Schepmania_nieuwenhuisi__2  
  |  |                                                          
--+  |                          +-- Indopseudodon_bogani        
  |  |              +-----------+                               
  |  |              |           +-- Indopseudodon_kayinensis    
  |  |  +-----------+                                           
  |  |  |           |  +----------- Pilsbryoconcha_exilis       
  |  |  |           |  |                                        
  |  |  |           +--+  +-------- Bineurus_anodontinum        
  +--+  |              |  |                                     
     |  |              +--+-------- Bineurus_loeiensis          
     |  |                 |                                     
     |  |                 |  +----- Thaiconcha_callifera        
     |  |                 +--+                                  
     |  |                    |  +-- Pseudodon_mekongi           
     |  |                    +--+                               
     |  |                       +-- Pseudodon_vondembuschianus  
     |  |                                                       
     |  |                 +-------- Ensidens_sagittarius        
     |  |                 |                                     
     |  |                 +-------- Hyriopsis_desowitzi         
     +--+                 |                                     
        |                 +-------- Rectidens_sumatrensis       
        |                 |                                     
        |                 +-------- Hyriopsis_bialata           
        |                 |                                     
        |  +--------------+     +-- Khairuloconcha_lunbawangorum
        |  |              +-----+                               
        |  |              |     +-- Khairuloconcha_sahanae      
        |  |              |                                     
        |  |              |  +----- Physunio_superbus           
        |  |              |  |                                  
        |  |              |  +----- Yaukthwa_paiensis           
        |  |              |  |                                  
        |  |              |  +----- Yaukthwa_inlenensis         
        |  |              +--+                                  
        |  |                 +----- Trapezoideus_foliaceus      
        +--+                 |                                  
           |                 +----- Yaukthwa_elongatula         
           |                 |                                  
           |                 |  +-- Lens_contradens             
           |                 +--+                               
           |                    +-- Lens_eximius                
           |                                                    
           |                    +-- Inversidens_rentianensis    
           |  +-----------------+                               
           |  |                 +-- Inversidens_brandtii        
           |  |                                                 
           |  |                 +-- Chamberlainia_somsakpanhai  
           +--+              +--+                               
              |              |  +-- Potomida_littoralis         
              |  +-----------+                                  
              |  |           |  +-- Lamprotula_caveata          
              |  |           +--+                               
              +--+              +-- Lamprotula_leaii            
                 |                                              
                 |  +-------------- Sinosolenaia_carinata       
                 |  |                                           
                 +--+  +----------- Gonidea_angulata            
                    |  |                                        
                    +--+----------- Microcondylaea_bonellii     
                       |                                        
                       |  +-------- Koreosolenaia_sitgyensis    
                       +--+                                     
                          |  +----- Parvasolenaia_rivularis     
                          +--+                                  
                             |  +-- Postolata_guangxiensis      
                             +--+                               
                                +-- Ptychorhynchus_pfisteri     
Cladogram Figure 6-7
                                +-- Margaritifera_dahurica      
  +-----------------------------+                               
  |                             +-- Margaritifera_margaritifera 
  |                                                             
  |  +----------------------------- Chamberlainia_somsakpanhai  
  |  |                                                          
  |  |                          +-- Khairuloconcha_lunbawangorum
  |  |           +--------------+                               
--+  |           |              +-- Khairuloconcha_sahanae      
  |  |           |                                              
  |  |           |        +-------- Ensidens_sagittarius        
  |  |  +--------+  +-----+                                     
  |  |  |        |  |     |  +----- Rectidens_sumatrensis       
  |  |  |        |  |     +--+                                  
  |  |  |        |  |        |  +-- Hyriopsis_bialata           
  +--+  |        |  |        +--+                               
     |  |        +--+           +-- Hyriopsis_desowitzi         
     |  |           |                                           
     |  |           |           +-- Physunio_superbus           
     |  |           |  +--------+                               
     |  |           |  |        +-- Trapezoideus_foliaceus      
     |  |           |  |                                        
     |  |           +--+        +-- Lens_contradens             
     |  |              |  +-----+                               
     |  |              |  |     +-- Lens_eximius                
     |  |              +--+                                     
     |  |                 |  +----- Yaukthwa_elongatula         
     +--+                 +--+                                  
        |                    |  +-- Yaukthwa_inlenensis         
        |                    +--+                               
        |                       +-- Yaukthwa_paiensis           
        |                                                       
        |        +----------------- Schepmania_nieuwenhuisi     
        |        |                                              
        |  +-----+              +-- Indopseudodon_bogani        
        |  |     |  +-----------+                               
        |  |     |  |           +-- Indopseudodon_kayinensis    
        |  |     +--+                                           
        |  |        |  +----------- Pilsbryoconcha_exilis       
        |  |        |  |                                        
        |  |        +--+        +-- Pseudodon_mekongi           
        |  |           |  +-----+                               
        |  |           |  |     +-- Pseudodon_vondembuschianus  
        |  |           +--+                                     
        +--+              |  +----- Thaiconcha_callifera        
           |              +--+                                  
           |                 |  +-- Bineurus_anodontinum        
           |                 +--+                               
           |                    +-- Bineurus_loeiensis          
           |                                                    
           |                 +----- Potomida_littoralis         
           |  +--------------+                                  
           |  |              |  +-- Lamprotula_caveata          
           |  |              +--+                               
           |  |                 +-- Lamprotula_leaii            
           +--+                                                 
              |                 +-- Inversidens_brandtii        
              |  +--------------+                               
              |  |              +-- Inversidens_rentianensis    
              +--+                                              
                 |  +-------------- Sinosolenaia_carinata       
                 |  |                                           
                 +--+           +-- Gonidea_angulata            
                    |  +--------+                               
                    |  |        +-- Microcondylaea_bonellii     
                    +--+                                        
                       |  +-------- Koreosolenaia_sitgyensis    
                       +--+                                     
                          |  +----- Parvasolenaia_rivularis     
                          +--+                                  
                             |  +-- Postolata_guangxiensis      
                             +--+                               
                                +-- Ptychorhynchus_pfisteri     
Cladogram Figure 8a_S1
  +-------------------------------------------------- Neotrigonia_margaritacea    
  |                                                                               
  |  +----------------------------------------------- Echyridella_menziesii       
  |  |                                                                            
--+  |                                            +-- Anodontites_trapesialis     
  |  |  +-----------------------------------------+                               
  |  |  |                                         +-- Mutela_dubia                
  |  |  |                                                                         
  |  |  |                                         +-- Margaritifera_dahurica      
  +--+  |  +--------------------------------------+                               
     |  |  |                                      +-- Margaritifera_margaritifera 
     |  |  |                                                                      
     |  |  |                                      +-- Quadrula_quadrula           
     |  |  |           +--------------------------+                               
     |  |  |           |                          +-- Uniomerus_tetralasmus       
     |  |  |           |                                                          
     +--+  |  +--------+                          +-- Elliptio_complanata         
        |  |  |        |  +-----------------------+                               
        |  |  |        |  |                       +-- Pleurobema_oviforme         
        |  |  |        +--+                                                       
        |  |  |           |  +----------------------- Amblema_plicata             
        |  |  |           |  |                                                    
        |  |  |           +--+  +-------------------- Popenaias_popeii            
        |  |  |              |  |                                                 
        |  |  |              +--+  +----------------- Toxolasma_parvum            
        |  |  |                 |  |                                              
        |  |  |                 |  |           +----- Potamilus_leptodon          
        +--+  |                 +--+  +--------+                                  
           |  |                    |  |        |  +-- Potamilus_alatus            
           |  |                    |  |        +--+                               
           |  |                    |  |           +-- Potamilus_streckersoni      
           |  |                    +--+                                           
           |  |                       |  +----------- Leaunio_lienosus            
           |  |                       |  |                                        
           |  |                       |  |     +----- Lampsilis_ornata            
           |  |                       +--+  +--+                                  
           |  |                          |  |  |  +-- Lampsilis_cardium           
           |  |                          |  |  +--+                               
           |  |                          +--+     +-- Lampsilis_cariosa           
           |  |                             |                                     
           |  |                             |  +----- Venustaconcha_ellipsiformis 
           |  |                             +--+                                  
           |  |                                |  +-- Lampsilis_powellii          
           |  |                                +--+                               
           |  |                                   +-- Lampsilis_siliquoidea       
           |  |                                                                   
           +--+                                +----- Chamberlainia_somsakpanhai  
              |           +--------------------+                                  
              |           |                    |  +-- Sinohyriopsis_cumingii      
              |           |                    +--+                               
              |           |                       +-- Sinohyriopsis_schlegelii    
              |           |                                                       
              |           |                       +-- Hyriopsis_bialata           
              |           |                    +--+                               
              |  +--------+                    |  +-- Rectidens_sumatrensis       
              |  |        |  +-----------------+                                  
              |  |        |  |                 |  +-- Lens_contradens             
              |  |        |  |                 +--+                               
              |  |        |  |                    +-- Physunio_superbus           
              |  |        |  |                                                    
              |  |        |  |           +----------- Potomida_littoralis         
              |  |        +--+  +--------+                                        
              |  |           |  |        |  +-------- Pronodularia_japanensis     
              |  |           |  |        +--+                                     
              |  |           |  |           |  +----- Lamprotula_leaii            
              |  |           |  |           +--+                                  
              |  |           |  |              |  +-- Lamprotula_caveata          
              |  |           +--+              +--+                               
              |  |              |                 +-- Lamprotula_cornuumlunae     
              |  |              |                                                 
              |  |              |                 +-- Pseudodon_vondembuschianus  
              |  |              |  +--------------+                               
              |  |              |  |              +-- Pilsbryoconcha_exilis       
              |  |              +--+                                              
              |  |                 |  +-------------- Inversidens_rentianensis    
              |  |                 |  |                                           
              +--+                 +--+           +-- Parvasolenaia_rivularis     
                 |                    |  +--------+                               
                 |                    |  |        +-- Ptychorhynchus_pfisteri     
                 |                    +--+                                        
                 |                       |  +-------- Microcondylaea_bonellii     
                 |                       +--+                                     
                 |                          |  +----- Sinosolenaia_carinata       
                 |                          +--+                                  
                 |                             |  +-- Sinosolenaia_iridinea       
                 |                             +--+                               
                 |                                +-- Sinosolenaia_oleivora       
                 |                                                                
                 |  +-------------------------------- Lamellidens_jenkinsianus    
                 |  |                                                             
                 |  |                             +-- Lepidodesma_aligera         
                 |  |  +--------------------------+                               
                 |  |  |                          +-- Lepidodesma_languilati      
                 |  |  |                                                          
                 |  |  |                    +-------- Aculamprotula_tortuosa      
                 |  |  |                    |                                     
                 |  |  |     +--------------+     +-- Aculamprotula_coreana       
                 |  |  |     |              |  +--+                               
                 +--+  |     |              |  |  +-- Aculamprotula_tientsinensis 
                    |  |     |              +--+                                  
                    |  |     |                 |  +-- Aculamprotula_polysticta    
                    |  |     |                 +--+                               
                    |  |     |                    +-- Aculamprotula_scripta       
                    |  |     |                                                    
                    |  |  +--+        +-------------- Unio_tumidus                
                    |  |  |  |  +-----+                                           
                    |  |  |  |  |     |  +----------- Unio_nanus                  
                    |  |  |  |  |     +--+                                        
                    +--+  |  |  |        |  +-------- Unio_elongatulus            
                       |  |  |  |        +--+                                     
                       |  |  |  |           |  +----- Unio_pictorum               
                       |  |  |  |           +--+                                  
                       |  |  |  |              |  +-- Unio_delphinus              
                       |  |  +--+              +--+                               
                       |  |     |                 +-- Unio_mancus                 
                       |  |     |                                                 
                       |  |     |                 +-- Nodularia_breviconcha       
                       |  |     |  +--------------+                               
                       |  |     |  |              +-- Nodularia_douglasiae        
                       |  |     |  |                                              
                       |  |     |  |              +-- Pseudocuneopsis_capitata    
                       |  |     |  +--------------+                               
                       |  |     +--+              +-- Pseudocuneopsis_sichuanensis
                       |  |        |                                              
                       |  |        |              +-- Schistodesmus_lampreyanus   
                       |  |        |  +-----------+                               
                       +--+        |  |           +-- Schistodesmus_spinosus      
                          |        +--+                                           
                          |           |  +----------- Tchangsinaia_piscicula      
                          |           +--+                                        
                          |              |  +-------- Cuneopsis_rufescens         
                          |              +--+                                     
                          |                 |  +----- Cuneopsis_celtiformis       
                          |                 +--+                                  
                          |                    |  +-- Cuneopsis_demangei          
                          |                    +--+                               
                          |                       +-- Cuneopsis_heudei            
                          |                                                       
                          |                 +-------- Acuticosta_chinensis        
                          |  +--------------+                                     
                          |  |              |  +----- Lanceolaria_gladiola        
                          |  |              +--+                                  
                          |  |                 |  +-- Lanceolaria_grayii          
                          |  |                 +--+                               
                          |  |                    +-- Lanceolaria_lanceolata      
                          |  |                                                    
                          |  |        +-------------- Cristaria_plicata           
                          |  |        |                                           
                          +--+  +-----+           +-- Anemina_arcaeformis         
                             |  |     |  +--------+                               
                             |  |     |  |        +-- Anemina_euscaphys           
                             |  |     +--+                                        
                             |  |        |  +-------- Beringiana_fukuharai        
                             |  |        +--+                                     
                             |  |           |  +----- Sinanodonta_lucida          
                             |  |           +--+                                  
                             |  |              |  +-- Sinanodonta_tumens          
                             +--+              +--+                               
                                |                 +-- Sinanodonta_woodiana        
                                |                                                 
                                |           +-------- Pseudanodonta_complanata    
                                |  +--------+                                     
                                |  |        |  +----- Anodonta_anatina            
                                |  |        +--+                                  
                                |  |           |  +-- Anodonta_cygnea             
                                |  |           +--+                               
                                +--+              +-- Anodonta_exulcerata         
                                   |                                              
                                   |  +-------------- Anodonta_nuttalliana        
                                   |  |                                           
                                   |  |        +----- Pyganodon_grandis           
                                   +--+  +-----+                                  
                                      |  |     |  +-- Utterbackia_imbecillis      
                                      |  |     +--+                               
                                      +--+        +-- Utterbackia_peninsularis    
                                         |                                        
                                         |  +-------- Lasmigona_complanata        
                                         +--+                                     
                                            |  +----- Prolasmidonta_heterodon     
                                            +--+                                  
                                               |  +-- Alasmidonta_varicosa        
                                               +--+                               
                                                  +-- Platynaias_compressa        
Cladogram Figure 8b_S2
  +-------------------------------------------------- Neotrigonia_margaritacea    
  |                                                                               
  |  +----------------------------------------------- Echyridella_menziesii       
  |  |                                                                            
--+  |                                            +-- Anodontites_trapesialis     
  |  |  +-----------------------------------------+                               
  |  |  |                                         +-- Mutela_dubia                
  |  |  |                                                                         
  |  |  |                                         +-- Margaritifera_dahurica      
  +--+  |  +--------------------------------------+                               
     |  |  |                                      +-- Margaritifera_margaritifera 
     |  |  |                                                                      
     |  |  |                                      +-- Quadrula_quadrula           
     |  |  |           +--------------------------+                               
     |  |  |           |                          +-- Uniomerus_tetralasmus       
     |  |  |           |                                                          
     +--+  |  +--------+                          +-- Elliptio_complanata         
        |  |  |        |  +-----------------------+                               
        |  |  |        |  |                       +-- Pleurobema_oviforme         
        |  |  |        +--+                                                       
        |  |  |           |  +----------------------- Amblema_plicata             
        |  |  |           |  |                                                    
        |  |  |           +--+  +-------------------- Popenaias_popeii            
        |  |  |              |  |                                                 
        |  |  |              +--+  +----------------- Toxolasma_parvum            
        |  |  |                 |  |                                              
        |  |  |                 |  |           +----- Potamilus_leptodon          
        +--+  |                 +--+  +--------+                                  
           |  |                    |  |        |  +-- Potamilus_alatus            
           |  |                    |  |        +--+                               
           |  |                    +--+           +-- Potamilus_streckersoni      
           |  |                       |                                           
           |  |                       |  +----------- Leaunio_lienosus            
           |  |                       |  |                                        
           |  |                       +--+  +-------- Venustaconcha_ellipsiformis 
           |  |                          |  |                                     
           |  |                          |  |     +-- Lampsilis_powellii          
           |  |                          +--+-----+                               
           |  |                             |     +-- Lampsilis_siliquoidea       
           |  |                             |                                     
           |  |                             |  +----- Lampsilis_ornata            
           |  |                             +--+                                  
           |  |                                |  +-- Lampsilis_cardium           
           |  |                                +--+                               
           |  |                                   +-- Lampsilis_cariosa           
           |  |                                                                   
           +--+                                +----- Chamberlainia_somsakpanhai  
              |           +--------------------+                                  
              |           |                    |  +-- Sinohyriopsis_cumingii      
              |           |                    +--+                               
              |           |                       +-- Sinohyriopsis_schlegelii    
              |           |                                                       
              |           |                       +-- Hyriopsis_bialata           
              |           |                    +--+                               
              |  +--------+                    |  +-- Rectidens_sumatrensis       
              |  |        |  +-----------------+                                  
              |  |        |  |                 |  +-- Lens_contradens             
              |  |        |  |                 +--+                               
              |  |        |  |                    +-- Physunio_superbus           
              |  |        |  |                                                    
              |  |        |  |           +----------- Potomida_littoralis         
              |  |        +--+  +--------+                                        
              |  |           |  |        |  +-------- Pronodularia_japanensis     
              |  |           |  |        +--+                                     
              |  |           |  |           |  +----- Lamprotula_leaii            
              |  |           |  |           +--+                                  
              |  |           |  |              |  +-- Lamprotula_caveata          
              |  |           +--+              +--+                               
              |  |              |                 +-- Lamprotula_cornuumlunae     
              |  |              |                                                 
              |  |              |                 +-- Pseudodon_vondembuschianus  
              |  |              |  +--------------+                               
              |  |              |  |              +-- Pilsbryoconcha_exilis       
              |  |              +--+                                              
              |  |                 |  +-------------- Inversidens_rentianensis    
              |  |                 |  |                                           
              +--+                 +--+           +-- Parvasolenaia_rivularis     
                 |                    |  +--------+                               
                 |                    |  |        +-- Ptychorhynchus_pfisteri     
                 |                    +--+                                        
                 |                       |  +-------- Microcondylaea_bonellii     
                 |                       +--+                                     
                 |                          |  +----- Sinosolenaia_carinata       
                 |                          +--+                                  
                 |                             |  +-- Sinosolenaia_iridinea       
                 |                             +--+                               
                 |                                +-- Sinosolenaia_oleivora       
                 |                                                                
                 |  +-------------------------------- Lamellidens_jenkinsianus    
                 |  |                                                             
                 |  |                             +-- Lepidodesma_aligera         
                 |  |  +--------------------------+                               
                 |  |  |                          +-- Lepidodesma_languilati      
                 |  |  |                                                          
                 |  |  |                    +-------- Aculamprotula_tortuosa      
                 |  |  |                    |                                     
                 |  |  |     +--------------+     +-- Aculamprotula_coreana       
                 |  |  |     |              |  +--+                               
                 +--+  |     |              |  |  +-- Aculamprotula_tientsinensis 
                    |  |     |              +--+                                  
                    |  |     |                 |  +-- Aculamprotula_polysticta    
                    |  |     |                 +--+                               
                    |  |     |                    +-- Aculamprotula_scripta       
                    |  |     |                                                    
                    |  |  +--+        +-------------- Unio_tumidus                
                    |  |  |  |  +-----+                                           
                    |  |  |  |  |     |  +----------- Unio_nanus                  
                    |  |  |  |  |     +--+                                        
                    +--+  |  |  |        |  +-------- Unio_elongatulus            
                       |  |  |  |        +--+                                     
                       |  |  |  |           |  +----- Unio_pictorum               
                       |  |  |  |           +--+                                  
                       |  |  |  |              |  +-- Unio_delphinus              
                       |  |  +--+              +--+                               
                       |  |     |                 +-- Unio_mancus                 
                       |  |     |                                                 
                       |  |     |                 +-- Nodularia_breviconcha       
                       |  |     |  +--------------+                               
                       |  |     |  |              +-- Nodularia_douglasiae        
                       |  |     |  |                                              
                       |  |     |  |              +-- Pseudocuneopsis_capitata    
                       |  |     |  +--------------+                               
                       |  |     +--+              +-- Pseudocuneopsis_sichuanensis
                       |  |        |                                              
                       |  |        |              +-- Schistodesmus_lampreyanus   
                       +--+        |  +-----------+                               
                          |        |  |           +-- Schistodesmus_spinosus      
                          |        +--+                                           
                          |           |  +----------- Tchangsinaia_piscicula      
                          |           +--+                                        
                          |              |  +-------- Cuneopsis_rufescens         
                          |              +--+                                     
                          |                 |  +----- Cuneopsis_celtiformis       
                          |                 +--+                                  
                          |                    |  +-- Cuneopsis_demangei          
                          |                    +--+                               
                          |                       +-- Cuneopsis_heudei            
                          |                                                       
                          |                 +-------- Acuticosta_chinensis        
                          |  +--------------+                                     
                          |  |              |  +----- Lanceolaria_gladiola        
                          |  |              +--+                                  
                          |  |                 |  +-- Lanceolaria_grayii          
                          |  |                 +--+                               
                          |  |                    +-- Lanceolaria_lanceolata      
                          |  |                                                    
                          |  |        +-------------- Cristaria_plicata           
                          +--+        |                                           
                             |  +-----+           +-- Anemina_arcaeformis         
                             |  |     |  +--------+                               
                             |  |     |  |        +-- Anemina_euscaphys           
                             |  |     +--+                                        
                             |  |        |  +-------- Beringiana_fukuharai        
                             |  |        +--+                                     
                             |  |           |  +----- Sinanodonta_lucida          
                             |  |           +--+                                  
                             +--+              |  +-- Sinanodonta_tumens          
                                |              +--+                               
                                |                 +-- Sinanodonta_woodiana        
                                |                                                 
                                |  +----------------- Anodonta_nuttalliana        
                                |  |                                              
                                |  |        +-------- Pseudanodonta_complanata    
                                |  |  +-----+                                     
                                +--+  |     |  +----- Anodonta_anatina            
                                   |  |     +--+                                  
                                   |  |        |  +-- Anodonta_cygnea             
                                   |  |        +--+                               
                                   +--+           +-- Anodonta_exulcerata         
                                      |                                           
                                      |        +----- Pyganodon_grandis           
                                      |  +-----+                                  
                                      |  |     |  +-- Utterbackia_imbecillis      
                                      |  |     +--+                               
                                      +--+        +-- Utterbackia_peninsularis    
                                         |                                        
                                         |  +-------- Lasmigona_complanata        
                                         +--+                                     
                                            |  +----- Prolasmidonta_heterodon     
                                            +--+                                  
                                               |  +-- Alasmidonta_varicosa        
                                               +--+                               
                                                  +-- Platynaias_compressa        

Page updated: 15:12:30 Wed 04 Dec 2024

 
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