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Thu 15 May 2025 |
The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!
This page documents data for Pfeiffer et al. (2021), such as genera and species attributed to that publication (including known misspellings, misidentifications, and other unavailable names) as well as taxonomic opinions, if available. Taxonomic opinions refer to uses of genera and species as valid and invalid in that work.
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Publication information
Pfeiffer, J.M., D.L. Graf, K.S. Cummings & L.M. Page. 2021. Taxonomic revision of a radiation of Southeast Asian freshwater mussels (Unionidae: Gonideinae: Contradentini+Rectidentini). Invertebrate Systematics 35: 394-470.
Click here for online access.
Cladogram Figures
(click the figure number to see the cladogram) fig. 12
fig. 18
fig. 25
fig. 27
fig. 30
fig. 9
Attributed Species Names
Pseudodon cambodjensis Pfeiffer & Graf, 2013
‘Petit’ of Authors.
Pfeiffer & Graf, 2013, J. Moll. Studies 79: 251.
Pfeiffer & Graf, 2015, Zool. J. Linn. Soc. 175(2): 310.
Zieritz et al., 2016, Sci. Total Environ. 571: 1071.
Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Monodontina cambodjensis’].
Lopes-Lima et al., 2017, MPE 106: 177.
Konopleva et al., 2017, Systemat. Biodiv. 15(3): 207.
Bolotov et al., 2018, Sci. Repts. 8: Table S1 [as ‘Monodontina cambodjensis’].
Razak et al., 2018, Conserv. Genet. Resour. 11: 459.
Huang et al., 2019, MPE 130: Table S4 [as ‘Monodontina cambodjensis’].
Konopleva et al., 2019, Sci. Repts. 9: Table S1 [as ‘Monodontina cambodjensis’].
Bolotov et al., 2020, Sci. Repts. 10: Table S1 [as ‘Monodontina cambodiensis’ (sic)].
Konopleva et al., 2021, Sci. Repts. 11: Table S1 [as ‘Monodontina cambodiensis’ (sic)].
Jeratthitikul et al., 2021, Eur. J. Taxonomy 775: 123 [as ‘Monodontina cambodiensis’ (sic)].
Pfeiffer et al., 2021, Invert. Syst. 35: 453 [as ‘Monodontina cambodjensis’].
Wu et al., 2021, ZooKeys 1054: 87 [as ‘Monodontina cambodjensis’].
Wu et al., 2021, J. Lake Sci. 33(6): attached Table 1.
Konopleva et al., 2023, Diversity 15(1), 10: Table S2 [as ‘Monodontina cambodjensis’].
Dai et al., 2023, Invert. Syst. 37(2): Table 1 [as ‘Monodontina cambodjensis’].
Dai et al., 2024, Zoosyst. Evol. 100(4): Table 2.
(misidentified reference to Monocondylea cambodjensis Petit de la Saussaye, 1865)
Type(s): SEQUENCED SPECIMEN UMMZ_304350, Pursat River, just downstream (north) of Highway 5 bridge, Pursat City, Cambodia.
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Yaukthwa dalliana Konopleva et al., 2019
‘Frierson’ of Authors.
Bolotov et al., 2019, Sci. Repts. 9: Table S1.
Konopleva et al., 2019, Sci. Repts. 9: 7.
Konopleva et al., 2019, Ecologica Montenegrina 24: Appendix 1.
Zieritz et al., 2020, Biodivers. Conserv. 29: 2243.
Bolotov et al., 2020, Sci. Repts. 10: Table S1.
Pfeiffer et al., 2021, Invert. Syst. 35: Table S2.
(misidentified reference to Parreysia dalliana Frierson, 1913)
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Ctenodesma borneensis Zieritz et al., 2020
‘Issel’ of Authors.
Zieritz et al., 2020, Biodivers. Conserv. 29: 2242, Table 2.
Pfeiffer et al., 2021, Invert. Syst. 35: Table S2. [in part]
(misidentified reference to Unio borneensis Issel, 1874)
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Lens maenamensis Pfeiffer et al., 2021
Pfeiffer et al., 2021, Invert. Syst. 35: 416, figs. 1, 9, 11-13.
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Physunio pellucidus Pfeiffer et al., 2021
Pfeiffer et al., 2021, Invert. Syst. 35: 424, figs. 1, 9, 17-19.
Type(s): HOLOTYPE (ORIG. DESIG.) UF_564750, Thuai River, Rt. 212 bridge S of Tha Uthen, Nakhon Phanom Province, Thailand.
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Trapezoideus subviridis Pfeiffer et al., 2021
Pfeiffer et al., 2021, Invert. Syst. 35: [62], figs. 1, 9, 20, 22.
Type(s): HOLOTYPE (ORIG. DESIG.) UF_564751, Patong River, downstream of Khao Soi Dao Wildlife Sanctuary, Soi Dao District, Chanthaburi Province, Thailand.
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Ensidens spiculus Pfeiffer et al., 2021
Pfeiffer et al., 2021, Invert. Syst. 35: 448, figs. 1, 9, 29-30, 32.
Type(s): HOLOTYPE (ORIG. DESIG.) UF_564752, Mekong River, along Rt. 211, Sangkhom District, Nong Khai Province, Thailand.
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Ensidens telus Pfeiffer et al., 2021
Pfeiffer et al., 2021, Invert. Syst. 35: 448, figs. 1, 9, 29-31.
Type(s): HOLOTYPE (ORIG. DESIG.) UF_564753, Tonle Sap River, near Phumi Trâpeang Préal, Kampong Chhnang Province, Cambodia.
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Hyriopsis kratiensis Pfeiffer et al., 2021
Pfeiffer et al., 2021, Invert. Syst. 35: 454, figs. 1, 9, 27, 33, 36.
Type(s): HOLOTYPE (ORIG. DESIG.) UF_564754, Mekong River, 1.4 miles upstream of Sambour, Sambour, Kratié Province, Cambodia.
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Hyriopsis phuphaniensis Pfeiffer et al., 2021
Pfeiffer et al., 2021, Invert. Syst. 35: 454, figs. 1, 27, 33, 36.
Type(s): HOLOTYPE (MONOTYPY) UF_564755, Mekong River, Mukdahan, at Muk River mouth, Mukdahan Province, Thailand.
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Hyriopsis khoratensis Pfeiffer et al., 2021
Pfeiffer et al., 2021, Invert. Syst. 35: 456, figs. 1, 9, 27, 33, 36.
Type(s): HOLOTYPE (ORIG. DESIG.) UF_564756, Pao River, Kamalasai, at Rt 214 bridge, Kamalasai District, Kalasin Province, Thailand.
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Hyriopsis sakhonensis Pfeiffer et al., 2021
Pfeiffer et al., 2021, Invert. Syst. 35: 456, figs. 1, 9, 27, 33, 36.
Type(s): HOLOTYPE (ORIG. DESIG.) UF_564757, Songkhram River, near confluence with Un River at bridge N of Ban Si Songkhram, Nakhon Phanom Province, Thailand.
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Taxonomic Opinions
Balwantia soleniformis (Benson, 1836)
Ctenodesma borneensis (Issel, 1874)
+ Unio plicatulus Lea, 1859
+ Unio pensitatus Fischer & Crosse, 1894
Ctenodesma scheibeneri Haas, 1927
Elongaria orientalis (Lea, 1840)
+ Unio productus Mousson, 1849
+ Unio productior Lea, 1852
+ Unio bithynicus Kobelt, 1893
Elongaria trompi (Drouët & Chaper, 1892)
Ensidens dugasti (Morlet, 1892), stat. rev.
+ Ensidens sagittarius Muanta et al., 2019
Ensidens ingallsianus (Lea, 1852)
+ Unio pazii Lea, 1862
Ensidens jaculus (Rochebrune, 1882), stat. rev.
+ Ensidens sagittarius Muanta et al., 2019
Ensidens sagittarius (Lea, 1856)
Ensidens spiculus sp. nov.
+ Ensidens sagittarius Bolotov et al., 2017
Ensidens telus sp. nov.
+ Ensidens sagittarius Pfeiffer & Graf, 2015
+ Ensidens ingallsianus Muanta et al., 2019
Hyriopsis bialata Simpson, 1900
+ Unio delphinus Lea, 1840
+ Unio delphinus Gruner, 1841
+ Unio megapterus Morelet, 1864
+ Unio delphinopterus Dautzenberg & Fischer, 1905
+ Hyriopsis gracilis Haas, 1910
Hyriopsis delaportei (Crosse & Fischer, 1876)
Hyriopsis desowitzi Brandt, 1974
Hyriopsis khoratensis sp. nov.
Hyriopsis kratiensis sp. nov.
Hyriopsis myersiana (Lea, 1856)
+ Unio housei Lea, 1856
+ Unio sutrangensis Morlet, 1889
+ Unio stungtrangensis Fischer & Dautzenberg, 1904
+ Unio patrangensis Morlet, 1904
+ Hyriopsis delaportei Muanta et al., 2019
Hyriopsis phuphaniensis sp. nov.
Hyriopsis sakhonensis sp. nov.
Hyriopsis undescribed
Hyriopsis velthuizeni (Schepman, 1896)
Lamprotula cornuumlunae
Lens comptus (Deshayes in Deshayes & Jullien 1876), comb. nov.
+ Harmandia munensis Brandt, 1974
+ Contradens semmelinki fultoni Lopes-Lima et al., 2017
Lens contradens (Lea, 1838), comb. nov.
+ Unio javanus Lea, 1840
+ Unio exilis Dunker, 1846
+ Unio mutatus Mousson, 1849
+ Unio tumidulus Lea, 1856
+ Unio cambodiensis Lea, 1856
+ Unio ascia Hanley, 1856
+ Unio rusticus Lea, 1856
+ Unio sumatrensis Lea, 1859
+ Unio mederianus Küster, 1861
+ Unio paivanus Morelet, 1865
+ Unio asperulus Lea, 1866
+ Margaron (Unio) dimotus Lea, 1870
+ Unio semidecoratus Morlet, 1889
+ Unio dautzenbergi Morlet, 1889
+ Unio verbecki Martens, 1897
+ Unio hageni Strubell, 1897
+ Unio laticeps Martens, 1900
+ Trapezoideus peninsularis Simpson, 1900
+ Unio pajakomboensis Bullen, 1906
+ Unio verbeeki Haas, 1914
+ Unio thaiensis Habe, 1964
+ Uniandra contradens rusticoides Brandt, 1974
+ Trapezoideus peninsularius Graf & Cummings, 2007
+ Contortus contortus Araujo et al., 2017
Lens eximius (Lea, 1856)
+ Unio semiquadrata Sowerby, 1866
Lens inornatus (Lea, 1856), comb. nov.
Lens lugens (Drouët & Chaper, 1892), comb. nov.
+ Unio saccellus Drouët & Chaper, 1892
+ Unio fulvaster Drouët & Chaper, 1892
Lens maenamensis sp. nov.
+ Contradens crossei Jeratthitikul et al., 2019
Lens micropterus (Morelet, 1866), comb. nov.
+ Unio inornatus Reeve, 1865
+ Unio cambojensis Sowerby, 1866
+ Margaron (Unio) versus Lea, 1870
+ Unio crossei Deshayes & Jullien, 1876
+ Unio semialatus Deshayes & Jullien, 1876
+ Diplodon ludovicianum Rochebrune, 1882
+ Unio inaequalis Rochebrune, 1882
+ Unio fischerianus Morlet, 1883
+ Unio siamensis Morlet, 1889
+ Unio soboles Fischer, 1891
+ Contradens contradens Pfeiffer & Graf, 2015
Lens misellus (Morelet, 1865)
+ Unio siamensis Lea, 1866
Lens novoselovi (Konopleva et al., 2019)
Lens pallegoixi (Sowerby, 1867)
Lens rolfbrandti (Jeratthitikul & Panha, 2019)
Lens semmelinki (von Martens, 1891)
+ Unio semmelincki Haas, 1914
Leoparreysia subcircularis (Brandt, 1974)
Monodontina cambodjensis (Petit, 1865)
Parvasolenaia neotriangularis
Parvasolenaia rivularis
Parvasolenaia triangularis
Physunio massini (Morelet, 1864)
Physunio modelli Brandt, 1974
Physunio pellucidus sp. nov.
Physunio superbus (Lea, 1843)
+ Unio gravidus Lea, 1856
+ Unio velaris Hanley, 1856
+ Unio abnormis Morelet, 1862
+ Unio superbus Sowerby, 1867
Pressidens exanthematicus (Küster, 1861)
+ Pseudodon insularis Drouët, 1894
+ Pressidens moellendorffi Haas, 1910
Prisodon obliquus Schumacher, 1817
+ Unio macropterus Dunker, 1846
Prohyriopsis stolata (von Martens, 1900)
Radiatula gratiosus (Philippi, 1843) comb. nov.
Rectidens sumatrensis (Dunker, 1852)
+ Unio perakensis Morgan, 1885
+ Unio lingulatus Drouët & Chaper, 1892
+ Unio prolongatus Drouët, 1894
+ Microcondylaea gracilis Strubell, 1897
+ Unio palembangensis Strubell, 1897
+ Unio pahangensis E.A. Smith, 1899
+ Unio palembangensis var. pahumbaensis Martens, 1900
+ Unio pressirostris Martens, 1900
+ Unio perakensis Preston, 1912
+ Rectidens prestoni Simpson, 1914
Solenaia carinata
Solenaia emarginata (Lea, 1860)
+ Solenaia khwaenoiensis Deein et al., 2004
Solenaia oleivora
+ Mycetopus iridineus Heude, 1874
Trapezoideus foliaceus (Gould, 1843)
Trapezoideus lenya Bolotov, Konopleva, Vikhrev, Gofarov, Lopes-Lima, Bogan, Lunn, Chan, Win, Aksenova, Tomilova, Tanmuangpak, Tumpeesuwan & Kondakov, 2020
Trapezoideus subviridis sp. nov.
Yaukthwa avaensis Bolotov, Konopleva, Vikhrev, Gofarov, Lopes-Lima, Bogan, Lunn, Chan, Win, Aksenova, Tomilova, Tanmuangpak, Tumpeesuwan & Kondakov, 2020
Yaukthwa baniensis (Bolotov, Konopleva, Vikhrev, Gofarov, Lopes-Lima, Bogan, Lunn, Chan, Win, Aksenova, Tomilova, Tanmuangpak, Tumpeesuwan & Kondakov, 2020), comb. nov.
+ Balwantia myanmarensis Bolotov et al., 2020
Yaukthwa dalliana (Frierson, 1913)
Yaukthwa elongatula Bolotov, Konopleva, Vikhrev, Lopes-Lima, Bogan, Lunn, Chan, Win, Aksenova, Gofarov, Tomilova & Kondakov, 2019, comb. res.
Yaukthwa inlenensis Konopleva, Pfeiffer, Vikhrev, Kondakov, Gofarov, Aksenova, Lunn, Chan & Bolotov, 2019
Yaukthwa nesemanni (Konopleva, Vikhrev & Bolotov in Bolotov et al., 2017)
Yaukthwa paiensis Konopleva, Pfeiffer, Vikhrev, Kondakov, Gofarov, Aksenova, Lunn, Chan & Bolotov, 2019
Yaukthwa panhai (Konopleva, Bolotov & Kondakov in Bolotov et al., 2017)
Yaukthwa peguensis (Anthony, 1865)
Yaukthwa rectangularis (Tapparone-Canefri, 1889), comb. nov.
Yaukthwa zayleymanensis (Preston, 1912)
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Cladogram Figure 12
+-------------- Solenaia_emarginata
|
| +-- Lens_contradens
| +--------+
--+ | +-- Lens_micropterus
| |
| | +-- Lens_inornatus
| +--------+
+--+ +-- Lens_misellus
|
| +----- Lens_maenamensis
| +--+
| | | +-- Lens_rolfbrandti
| | +--+
+--+ +-- Lens_eximius
|
| +----- Lens_pallegoixi
+--+
| +-- Lens_comptus
+--+
+-- Lens_novoselovi
Cladogram Figure 18
+-------------- Yaukthwa_paiensis
|
| +----- Trapezoideus_foliaceus
--+ +-----+
| | | +-- Trapezoideus_lenya
| | +--+
| | +-- Trapezoideus_subviridis
+--+
| +-- Physunio_massini
| +-----+
| | +-- Physunio_superbus
+--+
| +----- Physunio_modelli__3
| |
+--+ +-- Physunio_modelli__1
| |
+--+-- Physunio_modelli__2
|
+-- Physunio_pellucidus
Cladogram Figure 25
+-------------- Solenaia_emarginata
|
| +-- Yaukthwa_elongatula
--+ +--------+
| | +-- Yaukthwa_baniensis
| |
+--+ +----- Yaukthwa_panhai
| +--+
| | | +-- Yaukthwa_rectangularis
| | +--+
+--+ +-- Yaukthwa_nesemanni
|
| +----- Yaukthwa_avaensis
+--+
| +-- Yaukthwa_inlenensis
+--+
+-- Yaukthwa_paiensis
Cladogram Figure 27
+----------------- Khairuloconcha_lunbawangorum
|
--+ +-------------- Rectidens_sumatrensis
| |
| | +-- Hyriopsis_delaportei
+--+ +--------+
| | +-- Hyriopsis_desowitzi
| |
+--+ +-------- Hyriopsis_khoratensis
| |
| | +-- Hyriopsis_kratiensis
| +-----+
+--+ +-- Hyriopsis_phuphaniensis
|
| +----- Hyriopsis_sakhonensis
| |
+--+ +-- Hyriopsis_myersiana
| |
+--+-- Hyriopsis_panhai
|
+-- Hyriopsis_bialata
Cladogram Figure 30
+-------------- Rectidens_sumatrensis
|
--+ +-- Ensidens_telus
| +--------+
| | +-- Ensidens_ingallsianus
+--+
| +-------- Ensidens_dugasti
+--+
| +----- Ensidens_sagittarius
+--+
| +-- Ensidens_spiculus
+--+
+-- Ensidens_jaculus
Cladogram Figure 9
+-------------------------- Lamprotula_cornuumlunae
|
| +-- Ensidens_ingallsianus
| +--------+
| | +-- Ensidens_telus
| +--------+
| | | +-------- Ensidens_spiculus
| | +--+
| | | +----- Ensidens_dugasti
| | +--+
| +--+ | +-- Ensidens_jaculus
--+ | | +--+
| | | +-- Ensidens_sagittarius
| | |
| | | +----------------- Rectidens_sumatrensis
| | +--+
| | | +-------------- Hyriopsis_delaportei
| | +--+
| | | +----------- Hyriopsis_myersiana
| | +--+
| | | +-------- Hyriopsis_bialata
| | +--+
| | | +----- Hyriopsis_khoratensis
+--+ +--+
| | +-- Hyriopsis_kratiensis
| +--+
| +-- Hyriopsis_sakhonensis
|
| +-- Trapezoideus_subviridis
| +-----+
| | +-- Trapezoideus_foliaceus
| +-----------+
| | | +----- Physunio_superbus
| | +--+
| | | +-- Physunio_modelli
| | +--+
+--+ +-- Physunio_pellucidus
|
| +----------------- Yaukthwa_paiensis
| |
| | +-------------- Solenaia_emarginata
+--+ |
| | +-------- Lens_misellus
| | +--+
+--+ | | +----- Lens_inornatus
| | +--+
| | | +-- Lens_contradens
| | +--+
+--+ +-- Lens_micropterus
|
| +-- Lens_maenamensis
| +--+
| | +-- Lens_eximius
+-----+
| +-- Lens_pallegoixi
+--+
+-- Lens_comptus
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Page updated: 08:05:45 Thu 15 May 2025 |
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