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Page last updated
Fri 02 Sep 2022

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Parreysiinae | Indochinellini

genus Radiatula Simpson, 1900: cladomics
     7 valid species

Cladomics Summary

number of species = 7

analyzed in 26 cladograms in 17 publications
maximum number of terminal species in an analysis = 7, mean = 3.38

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 20
maximum number of included species = 7, mean = 4.1
proportion of cladograms that support basic monophyly = 0.85


Bolotov et al., 2022, fig. 2

iT tT bi riS oT roT roS iD
7 7 5 7 54 20 6 0.0

*** Top Monophyly Cladogram *** *** Top Sister-Group Cladogram ***

  +----------------------------- Indochinella_pugio      
  |                                                      
  |                 +----------- Radiatula_crispisulcata ** 
  |                 |                                    
  |                 |     +----- Radiatula_chaudhurii **    
--+  +--------------+  +--+                              
  |  |              |  |  |  +-- Radiatula_myitkyinae **    
  |  |              |  |  +--+                           
  |  |              +--+     +-- Radiatula_kachinensis **   
  |  |                 |                                 
  |  |                 |  +----- Radiatula_chindwinensis ** 
  +--+                 +--+                              
     |                    |  +-- Radiatula_myitthanensis ** 
     |                    +--+                           
     |                       +-- Radiatula_mouhoti **       
     |                                                   
     |  +----------------------- Scabiellus_songkramensis
     |  |                                                
     +--+  +-------------------- Harmandia_somboriensis  
        |  |                                             
        +--+  +----------------- Unionetta_fabagina      
           |  |                                          
           +--+              +-- Scabies_crispata        
              |  +-----------+                           
              |  |           +-- Scabies_mandarinus      
              +--+                                       
                 |  +----------- Scabies_nucleus         
                 |  |                                    
                 +--+        +-- Scabies_humilis         
                    |  +-----+                           
                    |  |     +-- Scabies_pilata          
                    +--+                                 
                       |  +----- Scabies_phaselus        
                       +--+                              
                          |  +-- Scabies_anceps          
                          +--+                           
                             +-- Scabies_scobinatus      

Bolotov et al., 2022, fig. S1

iT tT bi riS oT roT roS iD
7 7 4 7 54 20 6 3.0
  +-------------------------------- Indochinella_pugio      
  |                                                         
  |                          +----- Radiatula_chaudhurii **    
  |                    +-----+                              
  |                    |     |  +-- Radiatula_kachinensis **   
  |                    |     +--+                           
--+  +-----------------+        +-- Radiatula_myitkyinae **    
  |  |                 |                                    
  |  |                 |  +-------- Radiatula_crispisulcata ** 
  |  |                 +--+                                 
  |  |                    |  +----- Radiatula_mouhoti **       
  |  |                    +--+                              
  +--+                       |  +-- Radiatula_myitthanensis ** 
     |                       +--+                           
     |                          +-- Radiatula_chindwinensis ** 
     |                                                      
     |  +-------------------------- Scabiellus_songkramensis
     |  |                                                   
     +--+  +----------------------- Harmandia_somboriensis  
        |  |                                                
        +--+  +-------------------- Unionetta_fabagina      
           |  |                                             
           +--+  +----------------- Scabies_crispata        
              |  |                                          
              +--+              +-- Scabies_humilis         
                 |  +-----------+                           
                 |  |           +-- Scabies_mandarinus      
                 +--+                                       
                    |  +----------- Scabies_nucleus         
                    +--+                                    
                       |  +-------- Scabies_pilata          
                       +--+                                 
                          |  +----- Scabies_anceps          
                          +--+                              
                             |  +-- Scabies_phaselus        
                             +--+                           
                                +-- Scabies_scobinatus      

Bolotov et al., 2022, fig. 2

iT tT bi riS oT roT roS iD
6 6 4 6 52 20 6 3.0
  +----------------------------------- Indochinella_pugio      
  |                                                            
  |                       +----------- Radiatula_crispisulcata ** 
  |                       |                                    
--+  +--------------------+        +-- Radiatula_chaudhurii **    
  |  |                    |  +-----+                           
  |  |                    |  |     +-- Radiatula_myitkyinae **    
  |  |                    +--+                                 
  |  |                       |  +----- Radiatula_mouhoti **       
  +--+                       +--+                              
     |                          |  +-- Radiatula_chindwinensis ** 
     |                          +--+                           
     |                             +-- Radiatula_myitthanensis ** 
     |                                                         
     |  +----------------------------- Scabiellus_songkramensis
     +--+                                                      
        |  +-------------------------- Harmandia_somboriensis  
        +--+                                                   
           |  +----------------------- Unionetta_fabagina      
           +--+                                                
              |  +-------------------- Scabies_crispata        
              +--+                                             
                 |  +----------------- Scabies_mandarinus      
                 +--+                                          
                    |  +-------------- Scabies_nucleus         
                    +--+                                       
                       |  +----------- Scabies_pilata          
                       +--+                                    
                          |  +-------- Scabies_humilis         
                          +--+                                 
                             |  +----- Scabies_anceps          
                             +--+                              
                                |  +-- Scabies_phaselus        
                                +--+                           
                                   +-- Scabies_scobinatus      

Bolotov et al., 2022, fig. S1

iT tT bi riS oT roT roS iD
6 6 3 6 52 20 6 2.0
  +----------------------------- Indochinella_pugio      
  |                                                      
  |                    +-------- Radiatula_crispisulcata ** 
  |                    |                                 
  |  +-----------------+     +-- Radiatula_chaudhurii **    
--+  |                 |  +--+                           
  |  |                 |  |  +-- Radiatula_myitkyinae **    
  |  |                 +--+                              
  |  |                    |  +-- Radiatula_myitthanensis ** 
  |  |                    |  |                           
  +--+                    +--+-- Radiatula_mouhoti **       
     |                       |                           
     |                       +-- Radiatula_chindwinensis ** 
     |                                                   
     |  +----------------------- Scabiellus_songkramensis
     |  |                                                
     +--+  +-------------------- Harmandia_somboriensis  
        |  |                                             
        +--+  +----------------- Unionetta_fabagina      
           |  |                                          
           +--+              +-- Scabies_crispata        
              |  +-----------+                           
              |  |           +-- Scabies_mandarinus      
              +--+                                       
                 |  +----------- Scabies_nucleus         
                 |  |                                    
                 +--+        +-- Scabies_humilis         
                    |  +-----+                           
                    |  |     +-- Scabies_pilata          
                    +--+                                 
                       |  +----- Scabies_phaselus        
                       +--+                              
                          |  +-- Scabies_anceps          
                          +--+                           
                             +-- Scabies_scobinatus      

Bolotov et al., 2022, fig. S2

iT tT bi riS oT roT roS iD
6 6 4 6 52 20 6 3.0
  +-------------------------------- Indochinella_pugio      
  |                                                         
  |                    +----------- Radiatula_crispisulcata ** 
  |                    |                                    
  |  +-----------------+        +-- Radiatula_chaudhurii **    
--+  |                 |  +-----+                           
  |  |                 |  |     +-- Radiatula_myitkyinae **    
  |  |                 +--+                                 
  |  |                    |  +----- Radiatula_mouhoti **       
  |  |                    +--+                              
  +--+                       |  +-- Radiatula_myitthanensis ** 
     |                       +--+                           
     |                          +-- Radiatula_chindwinensis ** 
     |                                                      
     |  +-------------------------- Scabiellus_songkramensis
     |  |                                                   
     +--+  +----------------------- Harmandia_somboriensis  
        |  |                                                
        +--+  +-------------------- Unionetta_fabagina      
           |  |                                             
           +--+  +----------------- Scabies_crispata        
              |  |                                          
              +--+              +-- Scabies_humilis         
                 |  +-----------+                           
                 |  |           +-- Scabies_mandarinus      
                 +--+                                       
                    |  +----------- Scabies_nucleus         
                    +--+                                    
                       |  +-------- Scabies_pilata          
                       +--+                                 
                          |  +----- Scabies_anceps          
                          +--+                              
                             |  +-- Scabies_phaselus        
                             +--+                           
                                +-- Scabies_scobinatus      

Bolotov et al., 2022, fig. 8

iT tT bi riS oT roT roS iD
6 6 4 6 44 3 2 3.0
                 +-- Indonaia_andersoniana  
  +--------------+                          
  |              +-- Indonaia_subclathrata  
--+                                         
  |  +-------------- Indochinella_pugio     
  |  |                                      
  +--+  +----------- Radiatula_crispisulcata **
     |  |                                   
     +--+        +-- Radiatula_chaudhurii **   
        |  +-----+                          
        |  |     +-- Radiatula_myitkyinae **   
        +--+                                
           |  +----- Radiatula_mouhoti **      
           +--+                             
              |  +-- Radiatula_chindwinensis **
              +--+                          
                 +-- Radiatula_myitthanensis **

Bolotov et al., 2019, fig. 1

iT tT bi riS oT roT roS iD
5 5 2 5 13 1 1 2.0
  +----------- Indochinella_pugio     
  |                                   
--+        +-- Radiatula_myitkyinae **   
  |  +-----+                          
  |  |     +-- Radiatula_chaudhurii **   
  +--+                                
     |  +----- Radiatula_chindwinensis **
     +--+                             
        |  +-- Radiatula_mouhoti **      
        +--+                          
           +-- Radiatula_myitthanensis **

Bolotov et al., 2020, fig. 1

iT tT bi riS oT roT roS iD
4 4 2 4 118 17 6 3.0
  +-------------------------- Indochinella_pugio      
  |                                                   
  |                 +-------- Radiatula_crispisulcata ** 
  |  +--------------+                                 
--+  |              |  +----- Radiatula_mouhoti **       
  |  |              +--+                              
  |  |                 |  +-- Radiatula_chaudhurii **    
  |  |                 +--+                           
  +--+                    +-- Radiatula_myitkyinae **    
     |                                                
     |  +-------------------- Scabiellus_songkramensis
     |  |                                             
     |  |                 +-- Unionetta_fabagina      
     +--+  +--------------+                           
        |  |              +-- Harmandia_somboriensis  
        |  |                                          
        +--+  +-------------- Scabies_nucleus         
           |  |                                       
           |  |           +-- Scabies_crispata        
           +--+  +--------+                           
              |  |        +-- Scabies_mandarinus      
              |  |                                    
              +--+        +-- Scabies_humilis         
                 |  +-----+                           
                 |  |     +-- Scabies_pilata          
                 +--+                                 
                    |  +----- Scabies_anceps          
                    +--+                              
                       |  +-- Scabies_phaselus        
                       +--+                           
                          +-- Scabies_scobinatus      

Bolotov et al., 2020, fig. S1

iT tT bi riS oT roT roS iD
4 15 11 4 112 16 6 14.0
                             +-------- Indonaia_shurtleffiana  
  +--------------------------+                                 
  |                          |  +----- Indonaia_andersoniana   
  |                          +--+                              
  |                             |  +-- Indonaia_caerulea       
  |                             +--+                           
--+                                +-- Indonaia_subclathrata   
  |                                                            
  |  +-------------------------------- Indochinella_pugio      
  |  |                                                         
  |  |                          +----- Radiatula_mouhoti **       
  +--+  +-----------------------+                              
     |  |                       |  +-- Radiatula_myitkyinae **    
     |  |                       +--+                           
     +--+                          +-- Radiatula_chaudhurii **    
        |                                                      
        |  +-------------------------- Radiatula_crispisulcata ** 
        |  |                                                   
        +--+  +----------------------- Scabiellus_songkramensis
           |  |                                                
           +--+  +-------------------- Harmandia_somboriensis  
              |  |                                             
              +--+  +----------------- Unionetta_fabagina      
                 |  |                                          
                 +--+  +-------------- Scabies_mandarinus      
                    |  |                                       
                    +--+  +----------- Scabies_crispata        
                       |  |                                    
                       +--+  +-------- Scabies_nucleus         
                          |  |                                 
                          +--+     +-- Scabies_humilis         
                             |  +--+                           
                             |  |  +-- Scabies_pilata          
                             +--+                              
                                |  +-- Scabies_phaselus        
                                |  |                           
                                +--+-- Scabies_scobinatus      
                                   |                           
                                   +-- Scabies_anceps          

Bolotov et al., 2017, fig. S1

iT tT bi riS oT roT roS iD
3 3 1 3 125 10 3 4.0
  +----------- Indochinella_pugio  
  |                                
  |        +-- Scabies_humilis[1]  
--+        |                       
  |  +-----+-- Scabies_humilis[2]  
  |  |     |                       
  +--+     +-- Scabies_mandarinus  
     |                             
     |  +----- Radiatula_mouhoti **   
     +--+                          
        |  +-- Radiatula_myitkyinae **
        +--+                       
           +-- Radiatula_chaudhurii **

Bolotov et al., 2018, fig. 2

iT tT bi riS oT roT roS iD
3 3 1 3 81 7 3 6.0
  +----------- Indochinella_pugio  
  |                                
--+        +-- Scabies_mandarinus  
  |  +-----+                       
  |  |     +-- Scabies_humilis     
  +--+                             
     |  +----- Radiatula_myitkyinae **
     +--+                          
        |  +-- Radiatula_mouhoti **   
        +--+                       
           +-- Radiatula_chaudhurii **

Huang et al., 2019, fig. 2

iT tT bi riS oT roT roS iD
3 3 1 3 200 12 3 6.0
           +-- Indochinella_pugio[1]
           |                        
  +--------+-- Indochinella_pugio[2]
  |        |                        
  |        +-- Indochinella         
  |                                 
--+     +----- Scabies_humilis[1]   
  |  +--+                           
  |  |  |  +-- Scabies_humilis[2]   
  |  |  +--+                        
  +--+     +-- Scabies_mandarinus   
     |                              
     |  +----- Radiatula_myitkyinae ** 
     +--+                           
        |  +-- Radiatula_chaudhurii ** 
        +--+                        
           +-- Radiatula_mouhoti **    

Huang et al., 2019, fig. S3

iT tT bi riS oT roT roS iD
3 3 1 3 196 10 3 4.0
  +----------- Indochinella_pugio  
  |                                
  |     +----- Scabies_humilis[2]  
--+  +--+                          
  |  |  |  +-- Scabies_humilis[1]  
  |  |  +--+                       
  +--+     +-- Scabies_mandarinus  
     |                             
     |  +----- Radiatula_mouhoti **   
     +--+                          
        |  +-- Radiatula_myitkyinae **
        +--+                       
           +-- Radiatula_chaudhurii **

Huang et al., 2019, fig. S4

iT tT bi riS oT roT roS iD
3 3 1 3 199 10 3 4.0
  +----------- Indochinella_pugio  
  |                                
  |     +----- Scabies_humilis[1]  
--+  +--+                          
  |  |  |  +-- Scabies_humilis[2]  
  |  |  +--+                       
  +--+     +-- Scabies_mandarinus  
     |                             
     |  +----- Radiatula_mouhoti **   
     +--+                          
        |  +-- Radiatula_myitkyinae **
        +--+                       
           +-- Radiatula_chaudhurii **

Konopleva et al., 2019, fig. 1

iT tT bi riS oT roT roS iD
3 3 1 3 49 9 3 4.0
  +----------- Indochinella_pugio  
  |                                
  |        +-- Scabies_humilis     
--+        |                       
  |  +-----+-- Scabies_mandarinus  
  |  |     |                       
  +--+     +-- Scabies_phaselus    
     |                             
     |  +----- Radiatula_mouhoti **   
     +--+                          
        |  +-- Radiatula_myitkyinae **
        +--+                       
           +-- Radiatula_chaudhurii **

Jeratthitikul et al., 2019, fig. 1

iT tT bi riS oT roT roS iD
3 3 1 3 20 15 6 4.0
  +----------------------------------- Indochinella_pugio      
  |                                                            
  |                             +----- Radiatula_mouhoti **       
--+  +--------------------------+                              
  |  |                          |  +-- Radiatula_myitkyinae **    
  |  |                          +--+                           
  +--+                             +-- Radiatula_chaudhurii **    
     |                                                         
     |  +----------------------------- Scabiellus_songkramensis
     +--+                                                      
        |  +-------------------------- Harmandia_somboriensis  
        +--+                                                   
           |  +----------------------- Unionetta_fabagina      
           +--+                                                
              |  +-------------------- Scabies_crispata        
              +--+                                             
                 |  +----------------- Scabies_mandarinus      
                 +--+                                          
                    |  +-------------- Scabies_pilata          
                    +--+                                       
                       |  +----------- Scabies_humilis[2]      
                       +--+                                    
                          |  +-------- Scabies_humilis[1]      
                          +--+                                 
                             |  +----- Scabies_scobinatus      
                             +--+                              
                                |  +-- Scabies_phaselus        
                                +--+                           
                                   +-- Scabies_anceps          

Ortiz-Sepulveda et al., 2020, fig. 1

iT tT bi riS oT roT roS iD
3 4 0 3 31 3 3 6.0
  +-------- Indonaia_andersoniana
  |                              
--+  +----- Indochinella_pugio   
  |  |                           
  +--+  +-- Radiatula_mouhoti **    
     |  |                        
     |  +-- Scabies_humilis      
     +--+                        
        +-- Radiatula_myitkyinae ** 
        |                        
        +-- Radiatula_chaudhurii ** 

Ortiz-Sepulveda et al., 2020, fig. 2

iT tT bi riS oT roT roS iD
3 6 0 3 31 3 3 8.0
  +-- Radiatula_mouhoti **    
  |                        
  +-- Scabies_humilis      
  |                        
  +-- Radiatula_myitkyinae ** 
--+                        
  +-- Radiatula_chaudhurii ** 
  |                        
  +-- Indochinella_pugio   
  |                        
  +-- Indonaia_andersoniana

Bolotov et al., 2017, fig. S1

iT tT bi riS oT roT roS iD
2 2 0 2 136 7 3 --
        +-- Indochinella_pugio  
  +-----+                       
  |     +-- Scabies_crispata    
  |                             
--+     +-- Scabies_humilis     
  |  +--+                       
  |  |  +-- Scabies_mandarinus  
  +--+                          
     |  +-- Radiatula_myitkyinae **
     +--+                       
        +-- Radiatula_chaudhurii **

Bolotov et al., 2017, fig. S2

iT tT bi riS oT roT roS iD
2 2 0 2 136 7 3 --
        +-- Indochinella_pugio  
  +-----+                       
  |     +-- Scabies_crispata    
  |                             
--+     +-- Scabies_humilis     
  |  +--+                       
  |  |  +-- Scabies_mandarinus  
  +--+                          
     |  +-- Radiatula_myitkyinae **
     +--+                       
        +-- Radiatula_chaudhurii **

Whelan et al., 2011, fig. 1 ML

iT tT bi riS oT roT roS iD
1 1 0 1 35 1 1 --
  +-- Radiatula_chaudhurii **
--+                       
  +-- Indochinella_pugio  

Whelan et al., 2011, fig. 1 MP

iT tT bi riS oT roT roS iD
1 1 0 1 35 1 1 --
  +-- Radiatula_chaudhurii **
--+                       
  +-- Indochinella_pugio  

Pfeiffer & Graf, 2015, fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 39 2 2 --
  +----- Indochinella_pugio  
--+                          
  |  +-- Radiatula_chaudhurii **
  +--+                       
     +-- Scabies_mandarinus  

Konopleva et al., 2017, fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 30 2 2 --
  +----- Indochinella_pugio  
--+                          
  |  +-- Radiatula_chaudhurii **
  +--+                       
     +-- Scabies_mandarinus  

Lopes-Lima et al., 2020, fig. 2-4

iT tT bi riS oT roT roS iD
1 1 0 1 122 0 0 --
No taxon cladogram to show.

Choi et al., 2020, fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 8 0 0 --
No taxon cladogram to show.

Page updated: 13:40:15 Fri 02 Sep 2022

 
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