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Page last updated
Wed 04 Dec 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Parreysiinae | Indochinellini

genus Radiatula Simpson, 1900: cladomics
     7 valid species

Cladomics Summary

number of species = 7

analyzed in 34 cladograms in 22 publications
maximum number of terminals in an analysis = 7, mean = 3.47

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 26
maximum number of included terminals = 7, mean = 4.23
proportion of cladograms that support basic monophyly = 0.85


Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ tree


Bolotov et al., 2024 [Publ 4369], fig. 2

iT tT bi riS oT roT roS iD
7 7 5 7 55 22 7 0.0

*** Top Monophyly Cladogram *** *** Top Sister-Group Cladogram ***

                    +-- Pseudoscabies          
  +-----------------+                          
  |                 +-- Indonaia               
  |                                            
--+  +----------------- Indochinella           
  |  |                                         
  |  |        +-------- Scabiellus             
  |  |  +-----+                                
  +--+  |     |  +----- Harmandia              
     |  |     +--+                             
     |  |        |  +-- Unionetta              
     |  |        +--+                          
     +--+           +-- Scabies                
        |                                      
        |  +----------- Radiatula_crispisulcata **
        |  |                                   
        |  |     +----- Radiatula_chaudhurii **   
        +--+  +--+                             
           |  |  |  +-- Radiatula_myitkyinae **   
           |  |  +--+                          
           +--+     +-- Radiatula_kachinensis **  
              |                                
              |  +----- Radiatula_chindwinensis **
              +--+                             
                 |  +-- Radiatula_myitthanensis **
                 +--+                          
                    +-- Radiatula_mouhoti **      

Bolotov et al., 2022 [Publ 4252], fig. 2

iT tT bi riS oT roT roS iD
7 7 5 7 54 20 6 0.0
  +-------------------- Indonaia               
  |                                            
  |  +----------------- Indochinella           
--+  |                                         
  |  |        +-------- Scabiellus             
  |  |  +-----+                                
  +--+  |     |  +----- Harmandia              
     |  |     +--+                             
     |  |        |  +-- Unionetta              
     |  |        +--+                          
     +--+           +-- Scabies                
        |                                      
        |  +----------- Radiatula_crispisulcata **
        |  |                                   
        |  |     +----- Radiatula_chaudhurii **   
        +--+  +--+                             
           |  |  |  +-- Radiatula_myitkyinae **   
           |  |  +--+                          
           +--+     +-- Radiatula_kachinensis **  
              |                                
              |  +----- Radiatula_chindwinensis **
              +--+                             
                 |  +-- Radiatula_myitthanensis **
                 +--+                          
                    +-- Radiatula_mouhoti **      

Bolotov et al., 2024 [Publ 4369], fig. S1

iT tT bi riS oT roT roS iD
7 7 5 7 56 22 7 6.0
                       +-- Pseudoscabies          
  +--------------------+                          
  |                    +-- Indonaia               
  |                                               
--+  +-------------------- Indochinella           
  |  |                                            
  |  |           +-------- Scabiellus             
  |  |  +--------+                                
  +--+  |        |  +----- Harmandia              
     |  |        +--+                             
     |  |           |  +-- Unionetta              
     |  |           +--+                          
     +--+              +-- Scabies                
        |                                         
        |              +-- Radiatula_myitkyinae **   
        |  +-----------+                          
        |  |           +-- Radiatula_kachinensis **  
        +--+                                      
           |  +----------- Radiatula_chaudhurii **   
           +--+                                   
              |  +-------- Radiatula_crispisulcata **
              +--+                                
                 |  +----- Radiatula_mouhoti **      
                 +--+                             
                    |  +-- Radiatula_myitthanensis **
                    +--+                          
                       +-- Radiatula_chindwinensis **

Bolotov et al., 2022 [Publ 4252], fig. S1

iT tT bi riS oT roT roS iD
7 7 5 7 54 20 6 4.0
  +-------------------- Indonaia               
  |                                            
  |  +----------------- Indochinella           
--+  |                                         
  |  |        +-------- Scabiellus             
  |  |  +-----+                                
  +--+  |     |  +----- Harmandia              
     |  |     +--+                             
     |  |        |  +-- Unionetta              
     |  |        +--+                          
     +--+           +-- Scabies                
        |                                      
        |        +----- Radiatula_chaudhurii **   
        |  +-----+                             
        |  |     |  +-- Radiatula_kachinensis **  
        |  |     +--+                          
        +--+        +-- Radiatula_myitkyinae **   
           |                                   
           |  +-------- Radiatula_crispisulcata **
           +--+                                
              |  +----- Radiatula_mouhoti **      
              +--+                             
                 |  +-- Radiatula_myitthanensis **
                 +--+                          
                    +-- Radiatula_chindwinensis **

Bolotov et al., 2022 [Publ 4250], fig. 8

iT tT bi riS oT roT roS iD
6 6 4 6 43 3 2 3.0
  +----------------- Indonaia               
  |                                         
--+  +-------------- Indochinella           
  |  |                                      
  +--+  +----------- Radiatula_crispisulcata **
     |  |                                   
     +--+        +-- Radiatula_chaudhurii **   
        |  +-----+                          
        |  |     +-- Radiatula_myitkyinae **   
        +--+                                
           |  +----- Radiatula_mouhoti **      
           +--+                             
              |  +-- Radiatula_chindwinensis **
              +--+                          
                 +-- Radiatula_myitthanensis **

Bolotov et al., 2022 [Publ 4239], fig. 2

iT tT bi riS oT roT roS iD
6 6 4 6 52 20 6 3.0
  +-------------------- Indonaia               
  |                                            
--+  +----------------- Indochinella           
  |  |                                         
  |  |        +-------- Scabiellus             
  +--+  +-----+                                
     |  |     |  +----- Harmandia              
     |  |     +--+                             
     |  |        |  +-- Unionetta              
     +--+        +--+                          
        |           +-- Scabies                
        |                                      
        |  +----------- Radiatula_crispisulcata **
        |  |                                   
        +--+        +-- Radiatula_chaudhurii **   
           |  +-----+                          
           |  |     +-- Radiatula_myitkyinae **   
           +--+                                
              |  +----- Radiatula_mouhoti **      
              +--+                             
                 |  +-- Radiatula_chindwinensis **
                 +--+                          
                    +-- Radiatula_myitthanensis **

Bolotov et al., 2022 [Publ 4250], fig. S2

iT tT bi riS oT roT roS iD
6 6 4 6 9 6 2 1.0
  +----------------- Indonaia               
  |                                         
--+  +-------------- Indochinella           
  |  |                                      
  +--+  +----------- Radiatula_crispisulcata **
     |  |                                   
     +--+        +-- Radiatula_chaudhurii **   
        |  +-----+                          
        |  |     +-- Radiatula_myitkyinae **   
        +--+                                
           |  +----- Radiatula_chindwinensis **
           +--+                             
              |  +-- Radiatula_mouhoti **      
              +--+                          
                 +-- Radiatula_myitthanensis **

Bolotov et al., 2022 [Publ 4239], fig. S2

iT tT bi riS oT roT roS iD
6 6 4 6 52 20 6 3.0
  +-------------------- Indonaia               
  |                                            
--+  +----------------- Indochinella           
  |  |                                         
  |  |        +-------- Scabiellus             
  +--+  +-----+                                
     |  |     |  +----- Harmandia              
     |  |     +--+                             
     |  |        |  +-- Unionetta              
     +--+        +--+                          
        |           +-- Scabies                
        |                                      
        |  +----------- Radiatula_crispisulcata **
        |  |                                   
        +--+        +-- Radiatula_chaudhurii **   
           |  +-----+                          
           |  |     +-- Radiatula_myitkyinae **   
           +--+                                
              |  +----- Radiatula_mouhoti **      
              +--+                             
                 |  +-- Radiatula_myitthanensis **
                 +--+                          
                    +-- Radiatula_chindwinensis **

Bolotov et al., 2022 [Publ 4239], fig. S1

iT tT bi riS oT roT roS iD
6 6 3 6 52 20 6 2.0
  +----------------- Indonaia               
  |                                         
--+  +-------------- Indochinella           
  |  |                                      
  |  |     +-------- Scabiellus             
  +--+  +--+                                
     |  |  |  +----- Harmandia              
     |  |  +--+                             
     |  |     |  +-- Unionetta              
     +--+     +--+                          
        |        +-- Scabies                
        |                                   
        |  +-------- Radiatula_crispisulcata **
        |  |                                
        +--+     +-- Radiatula_chaudhurii **   
           |  +--+                          
           |  |  +-- Radiatula_myitkyinae **   
           +--+                             
              |  +-- Radiatula_myitthanensis **
              |  |                          
              +--+-- Radiatula_mouhoti **      
                 |                          
                 +-- Radiatula_chindwinensis **

Bolotov et al., 2019 [Publ 3974], fig. 1

iT tT bi riS oT roT roS iD
5 5 3 5 13 1 1 2.0
  +----------- Indochinella           
  |                                   
--+        +-- Radiatula_myitkyinae **   
  |  +-----+                          
  |  |     +-- Radiatula_chaudhurii **   
  +--+                                
     |  +----- Radiatula_chindwinensis **
     +--+                             
        |  +-- Radiatula_mouhoti **      
        +--+                          
           +-- Radiatula_myitthanensis **

Bolotov et al., 2020 [Publ 4100], fig. S1

iT tT bi riS oT roT roS iD
4 15 12 4 108 16 6 14.0
  +-------------------- Indonaia               
  |                                            
--+  +----------------- Indochinella           
  |  |                                         
  |  |           +----- Radiatula_mouhoti **      
  +--+  +--------+                             
     |  |        |  +-- Radiatula_myitkyinae **   
     |  |        +--+                          
     +--+           +-- Radiatula_chaudhurii **   
        |                                      
        |  +----------- Radiatula_crispisulcata **
        +--+                                   
           |  +-------- Scabiellus             
           +--+                                
              |  +----- Harmandia              
              +--+                             
                 |  +-- Unionetta              
                 +--+                          
                    +-- Scabies                

Bolotov et al., 2020 [Publ 4100], fig. 1

iT tT bi riS oT roT roS iD
4 4 2 4 114 17 6 3.0
  +----------------- Indonaia               
  |                                         
--+  +-------------- Indochinella           
  |  |                                      
  |  |     +-------- Radiatula_crispisulcata **
  +--+  +--+                                
     |  |  |  +----- Radiatula_mouhoti **      
     |  |  +--+                             
     |  |     |  +-- Radiatula_chaudhurii **   
     +--+     +--+                          
        |        +-- Radiatula_myitkyinae **   
        |                                   
        |  +-------- Scabiellus             
        +--+                                
           |  +----- Scabies                
           +--+                             
              |  +-- Unionetta              
              +--+                          
                 +-- Harmandia              

Huang et al., 2019 [Publ 3843], fig. S4

iT tT bi riS oT roT roS iD
3 3 1 3 195 9 3 4.0
  +-------------- Indonaia            
--+                                   
  |  +----------- Indochinella        
  +--+                                
     |  +-------- Scabies             
     +--+                             
        |  +----- Radiatula_mouhoti **   
        +--+                          
           |  +-- Radiatula_myitkyinae **
           +--+                       
              +-- Radiatula_chaudhurii **

Ortiz-Sepulveda et al., 2020 [Publ 4101], fig. 1

iT tT bi riS oT roT roS iD
3 6 1 3 33 5 3 8.0
  +-------- Indonaia            
  |                             
--+  +----- Indochinella        
  |  |                          
  +--+  +-- Radiatula_mouhoti **   
     |  |                       
     |  +-- Radiatula_myitkyinae **
     +--+                       
        +-- Radiatula_chaudhurii **
        |                       
        +-- Scabies             

Huang et al., 2019 [Publ 3843], fig. S3

iT tT bi riS oT roT roS iD
3 3 1 3 193 9 3 4.0
  +-------------- Indonaia            
--+                                   
  |  +----------- Indochinella        
  +--+                                
     |  +-------- Scabies             
     +--+                             
        |  +----- Radiatula_mouhoti **   
        +--+                          
           |  +-- Radiatula_myitkyinae **
           +--+                       
              +-- Radiatula_chaudhurii **

Wu et al., 2021 [Publ 4366], fig. 2_I

iT tT bi riS oT roT roS iD
3 3 1 3 88 2 2 6.0
  +----------- Indonaia            
--+                                
  |  +-------- Scabies             
  +--+                             
     |  +----- Radiatula_myitkyinae **
     +--+                          
        |  +-- Radiatula_mouhoti **   
        +--+                       
           +-- Radiatula_chaudhurii **

Jeratthitikul et al., 2019 [Publ 3922], fig. 1

iT tT bi riS oT roT roS iD
3 3 1 3 20 15 6 4.0
  +----------------- Indonaia            
  |                                      
--+  +-------------- Indochinella        
  |  |                                   
  |  |        +----- Radiatula_mouhoti **   
  +--+  +-----+                          
     |  |     |  +-- Radiatula_myitkyinae **
     |  |     +--+                       
     +--+        +-- Radiatula_chaudhurii **
        |                                
        |  +-------- Scabiellus          
        +--+                             
           |  +----- Harmandia           
           +--+                          
              |  +-- Unionetta           
              +--+                       
                 +-- Scabies             

Konopleva et al., 2019 [Publ 3900], fig. 1

iT tT bi riS oT roT roS iD
3 3 1 3 49 9 3 4.0
  +-------------- Indonaia            
--+                                   
  |  +----------- Indochinella        
  +--+                                
     |  +-------- Scabies             
     +--+                             
        |  +----- Radiatula_mouhoti **   
        +--+                          
           |  +-- Radiatula_myitkyinae **
           +--+                       
              +-- Radiatula_chaudhurii **

Bolotov et al., 2017 [Publ 3764], fig. S1

iT tT bi riS oT roT roS iD
3 3 1 3 122 9 3 4.0
  +----------- Indochinella        
--+                                
  |  +-------- Scabies             
  +--+                             
     |  +----- Radiatula_mouhoti **   
     +--+                          
        |  +-- Radiatula_myitkyinae **
        +--+                       
           +-- Radiatula_chaudhurii **

Bolotov et al., 2018 [Publ 3826], fig. 2

iT tT bi riS oT roT roS iD
3 3 1 3 80 8 3 6.0
  +-------------- Indonaia            
--+                                   
  |  +----------- Indochinella        
  +--+                                
     |  +-------- Scabies             
     +--+                             
        |  +----- Radiatula_myitkyinae **
        +--+                          
           |  +-- Radiatula_mouhoti **   
           +--+                       
              +-- Radiatula_chaudhurii **

Ortiz-Sepulveda et al., 2020 [Publ 4101], fig. 2

iT tT bi riS oT roT roS iD
3 8 1 3 33 5 3 10.0
  +-- Radiatula_mouhoti **   
  |                       
  +-- Radiatula_myitkyinae **
  |                       
  +-- Radiatula_chaudhurii **
--+                       
  +-- Indochinella        
  |                       
  +-- Indonaia            
  |                       
  +-- Scabies             

Wu et al., 2021 [Publ 4366], fig. 6

iT tT bi riS oT roT roS iD
3 4 2 3 88 2 2 7.0
  +----------- Indonaia            
--+                                
  |  +-------- Radiatula_myitkyinae **
  +--+                             
     |  +----- Scabies             
     +--+                          
        |  +-- Radiatula_mouhoti **   
        +--+                       
           +-- Radiatula_chaudhurii **

Huang et al., 2019 [Publ 3843], fig. 2

iT tT bi riS oT roT roS iD
3 3 1 3 195 9 3 6.0
  +-------------- Indonaia            
--+                                   
  |  +----------- Indochinella        
  +--+                                
     |  +-------- Scabies             
     +--+                             
        |  +----- Radiatula_myitkyinae **
        +--+                          
           |  +-- Radiatula_chaudhurii **
           +--+                       
              +-- Radiatula_mouhoti **   

Bolotov et al., 2017 [Publ 3761], fig. S1

iT tT bi riS oT roT roS iD
2 2 0 2 137 9 3 --
  +-------------- Indonaia            
  |                                   
  |           +-- Indochinella        
--+  +--------+                       
  |  |        +-- Scabies_crispata    
  |  |                                
  +--+        +-- Radiatula_myitkyinae **
     |  +-----+                       
     |  |     +-- Radiatula_chaudhurii **
     +--+                             
        |  +----- Scabies_mandarinus  
        +--+                          
           |  +-- Scabies_humilis     
           +--+                       
              +-- Scabies_phaselus    

Pfeiffer et al., 2018 [Publ 3827], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 26 15 6 --
  +-------------------- Indonaia             
  |                                          
--+  +----------------- Indochinella         
  |  |                                       
  +--+              +-- Radiatula_chaudhurii ** 
     |  +-----------+                        
     |  |           +-- Radiatula_myitkyinae ** 
     +--+                                    
        |  +----------- Scabiellus           
        +--+                                 
           |  +-------- Unionetta_fabagina__2
           +--+                              
              |  +----- Harmandia            
              +--+                           
                 |  +-- Unionetta_fabagina__1
                 +--+                        
                    +-- Scabies              

Bolotov et al., 2017 [Publ 3761], fig. S2

iT tT bi riS oT roT roS iD
2 2 0 2 137 9 3 --
  +-------------- Indonaia            
  |                                   
  |           +-- Indochinella        
--+  +--------+                       
  |  |        +-- Scabies_crispata    
  |  |                                
  +--+        +-- Radiatula_myitkyinae **
     |  +-----+                       
     |  |     +-- Radiatula_chaudhurii **
     +--+                             
        |  +----- Scabies_mandarinus  
        +--+                          
           |  +-- Scabies_humilis     
           +--+                       
              +-- Scabies_phaselus    

Choi et al., 2020 [Publ 4169], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 9 0 0 --
-- Radiatula_mouhoti

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iT tT bi riS oT roT roS iD
1 1 0 1 55 2 2 --
  +----- Indonaia         
--+                       
  |  +-- Indochinella     
  +--+                    
     +-- Radiatula_mouhoti **

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iT tT bi riS oT roT roS iD
1 1 0 1 35 1 1 --
  +-- Radiatula_chaudhurii **
--+                       
  +-- Indochinella        

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 39 2 2 --
  +----- Indochinella        
--+                          
  |  +-- Radiatula_chaudhurii **
  +--+                       
     +-- Scabies             

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iT tT bi riS oT roT roS iD
1 1 0 1 119 0 0 --
-- Radiatula_mouhoti

Konopleva et al., 2023 [Publ 4262], fig. 8

iT tT bi riS oT roT roS iD
1 1 0 1 81 13 6 --
  +----------------- Indonaia         
--+                                   
  |  +-------------- Indochinella     
  +--+                                
     |  +----------- Radiatula_mouhoti **
     +--+                             
        |  +-------- Scabiellus       
        +--+                          
           |  +----- Scabies          
           +--+                       
              |  +-- Unionetta        
              +--+                    
                 +-- Harmandia        

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iT tT bi riS oT roT roS iD
1 1 0 1 35 1 1 --
  +-- Radiatula_chaudhurii **
--+                       
  +-- Indochinella        

Konopleva et al., 2017 [Publ 3807], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 30 2 2 --
  +----- Indochinella        
--+                          
  |  +-- Radiatula_chaudhurii **
  +--+                       
     +-- Scabies             

Page updated: 14:57:14 Wed 04 Dec 2024

 
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