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Page last updated
Thu 07 Dec 2023

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Unionoida | Unionoidea | Unionidae | Gonideinae | Pseudodontini | Pseudodontina

genus Bineurus Simpson, 1900: cladomics
     5 valid species

Cladomics Summary

number of species = 5

analyzed in 29 cladograms in 19 publications
maximum number of terminals in an analysis = 5, mean = 2.76

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 23
maximum number of included terminals = 5, mean = 3.22
proportion of cladograms that support basic monophyly = 0.91

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ tree

Bolotov et al., 2023 [Publ 4312], fig. S2

iT tT bi riS oT roT roS iD
5 5 3 5 37 32 7 0.0

*** Top Monophyly Cladogram *** *** Top Sister-Group Cladogram ***

                 +-- Namkongnaia_inkhavilayi
  +--------------+                          
  |              +-- Namkongnaia_lemeslei   
  |                                         
--+              +-- Thaiconcha_callifera   
  |              |                          
  |  +-----------+-- Thaiconcha_thaiensis   
  |  |           |                          
  +--+           +-- Thaiconcha_munelliptica
     |                                      
     |  +----------- Bineurus_loeiensis **     
     +--+                                   
        |  +-------- Bineurus_panhai **        
        +--+                                
           |  +----- Bineurus_anodontinum **   
           +--+                             
              |  +-- Bineurus_exilis **        
              +--+                          
                 +-- Bineurus_mouhotii **

Jeratthitikul & Sutcharit, 2023 [Publ 4289], fig. 1

iT tT bi riS oT roT roS iD
5 5 3 5 8 6 6 6.0 
     +----------- Bineurus_mouhotii **         
  +--+                                      
  |  |  +-------- Bineurus_loeiensis **        
  |  +--+                                   
  |     |  +----- Bineurus_panhai **           
  |     +--+                                
  |        |  +-- Bineurus_anodontinum **      
--+        +--+                             
  |           +-- Bineurus_exilis **           
  |                                         
  |        +----- Namkongnaia_inkhavilayi   
  |     +--+                                
  |     |  |  +-- Sundadontina_cumingii     
  |     |  +--+                             
  +-----+     +-- Nyeinchanconcha_nyeinchani
        |                                   
        |  +----- Pseudodon_vondembuschianus
        +--+                                
           |  +-- Pilsbryoconcha_exilis     
           +--+                             
              +-- Thaiconcha_callifera

Konopleva et al., 2021 [Publ 4213], fig. 1

iT tT bi riS oT roT roS iD
4 4 2 4 24 16 6 3.0 
  +-------------- Nyeinchanconcha_nyeinchani
  |                                         
  |           +-- Thaiconcha_munelliptica   
--+           |                             
  |  +--------+-- Thaiconcha_thaiensis      
  |  |        |                             
  +--+        +-- Thaiconcha_callifera      
     |                                      
     |  +-------- Bineurus_loeiensis **        
     +--+                                   
        |  +----- Bineurus_mouhotii **         
        +--+                                
           |  +-- Bineurus_anodontinum **      
           +--+                             
              +-- Bineurus_exilis **

Konopleva et al., 2023 [Publ 4262], fig. 8

iT tT bi riS oT roT roS iD
4 4 2 4 80 23 7 3.0 
  +-------------- Nyeinchanconcha_nyeinchani
  |                                         
  |        +----- Thaiconcha_thaiensis      
--+  +-----+                                
  |  |     |  +-- Thaiconcha_munelliptica   
  |  |     +--+                             
  +--+        +-- Thaiconcha_callifera      
     |                                      
     |  +-------- Bineurus_loeiensis **        
     +--+                                   
        |  +----- Bineurus_mouhotii **         
        +--+                                
           |  +-- Bineurus_anodontinum **      
           +--+                             
              +-- Bineurus_exilis **

Bolotov et al., 2023 [Publ 4312], fig. 2

iT tT bi riS oT roT roS iD
4 4 2 4 36 31 7 3.0 
  +-------------- Nyeinchanconcha_nyeinchani
  |                                         
  |        +----- Thaiconcha_munelliptica   
--+  +-----+                                
  |  |     |  +-- Thaiconcha_callifera      
  |  |     +--+                             
  +--+        +-- Thaiconcha_thaiensis      
     |                                      
     |  +-------- Bineurus_loeiensis **        
     +--+                                   
        |  +----- Bineurus_mouhotii **         
        +--+                                
           |  +-- Bineurus_anodontinum **      
           +--+                             
              +-- Bineurus_exilis **

Bolotov et al., 2023 [Publ 4312], fig. S1

iT tT bi riS oT roT roS iD
4 4 2 4 35 30 7 3.0 
  +-------------- Nyeinchanconcha_nyeinchani
  |                                         
  |        +----- Thaiconcha_thaiensis      
--+  +-----+                                
  |  |     |  +-- Thaiconcha_callifera      
  |  |     +--+                             
  +--+        +-- Thaiconcha_munelliptica   
     |                                      
     |  +-------- Bineurus_loeiensis **        
     +--+                                   
        |  +----- Bineurus_mouhotii **         
        +--+                                
           |  +-- Bineurus_anodontinum **      
           +--+                             
              +-- Bineurus_exilis **

Bolotov et al., 2017 [Publ 3761], fig. S1

iT tT bi riS oT roT roS iD
3 3 1 3 138 11 5 4.0 
           +-- Nyeinchanconcha_nyeinchani
  +--------+                             
  |        +-- Sundadontina_cumingii     
--+                                      
  |  +-------- Thaiconcha_callifera      
  +--+                                   
     |  +----- Bineurus_exilis **           
     +--+                                
        |  +-- Bineurus_mouhotii **         
        +--+                             
           +-- Bineurus_loeiensis **

Bolotov et al., 2017 [Publ 3761], fig. S2

iT tT bi riS oT roT roS iD
3 3 1 3 138 12 5 4.0 
  +-------- Thaiconcha_callifera
--+                             
  |  +----- Bineurus_exilis **     
  +--+                          
     |  +-- Bineurus_mouhotii **   
     +--+                       
        +-- Bineurus_loeiensis **

Bolotov et al., 2017 [Publ 3764], fig. S1

iT tT bi riS oT roT roS iD
3 3 1 3 128 11 5 4.0 
              +-- Nyeinchanconcha_nyeinchani
  +-----------+                             
  |           +-- Sundadontina_cumingii     
  |                                         
  |        +----- Bineurus_exilis **           
  +--------+                                
  |        |  +-- Bineurus_mouhotii **         
  |        +--+                             
  |           +-- Bineurus_loeiensis **        
  |                                         
--+     +-------- Sundadontina_tumida       
  +-----+                                   
  |     |  +----- Pseudodon_vagulus         
  |     +--+                                
  |        |  +-- Pseudodon_mekongi         
  |        +--+                             
  |           +-- Pseudodon_vondembuschianus
  |                                         
  |  +----------- Thaiconcha_callifera      
  +--+                                      
     |  +-------- Pilsbryoconcha_exilis[3]  
     +--+                                   
        |  +----- Pilsbryoconcha_exilis[2]  
        +--+                                
           |  +-- Pilsbryoconcha_exilis[1]  
           +--+                             
              +-- Pilsbryoconcha_carinifera

Bolotov et al., 2018 [Publ 3826], fig. 2

iT tT bi riS oT roT roS iD
3 8 0 3 82 5 2 8.0 
  +-- Bineurus_mouhotii **         
  |                             
  +-- Pseudodon_vagulus         
  |                             
  +-- Pseudodon_mekongi         
  |                             
  +-- Pilsbryoconcha_exilis     
--+                             
  +-- Pilsbryoconcha_carinifera 
  |                             
  +-- Bineurus_loeiensis **        
  |                             
  +-- Bineurus_exilis **           
  |                             
  +-- Pseudodon_vondembuschianus

Huang et al., 2019 [Publ 3843], fig. 2

iT tT bi riS oT roT roS iD
3 3 1 3 201 12 5 2.0 
  +----------- Nyeinchanconcha_nyeinchani
--+                                      
  |  +-------- Thaiconcha_callifera      
  +--+                                   
     |  +----- Bineurus_loeiensis **        
     +--+                                
        |  +-- Bineurus_mouhotii **         
        +--+                             
           +-- Bineurus_exilis **

Huang et al., 2019 [Publ 3843], fig. S3

iT tT bi riS oT roT roS iD
3 3 1 3 198 12 5 2.0 
           +-- Nyeinchanconcha_nyeinchani
  +--------+                             
  |        +-- Sundadontina_cumingii     
--+                                      
  |  +-------- Thaiconcha_callifera      
  +--+                                   
     |  +----- Bineurus_loeiensis **        
     +--+                                
        |  +-- Bineurus_mouhotii **         
        +--+                             
           +-- Bineurus_exilis **

Huang et al., 2019 [Publ 3843], fig. S4

iT tT bi riS oT roT roS iD
3 3 1 3 199 11 5 2.0 
  +-------- Nyeinchanconcha_nyeinchani
  |                                   
  +-------- Thaiconcha_callifera      
  |                                   
  +-------- Sundadontina_cumingii     
  |                                   
  +-------- Sundadontina_tumida[1]    
  |                                   
  +-------- Sundadontina_tumida[2]    
  |                                   
  |  +----- Pilsbryoconcha_exilis[2]  
--+--+                                
  |  |  +-- Pilsbryoconcha_carinifera 
  |  +--+                             
  |     +-- Pilsbryoconcha_exilis[1]  
  |                                   
  |  +----- Pseudodon_vagulus         
  +--+                                
  |  |  +-- Pseudodon_mekongi         
  |  +--+                             
  |     +-- Pseudodon_vondembuschianus
  |                                   
  |  +----- Bineurus_loeiensis **        
  +--+                                
     |  +-- Bineurus_mouhotii **         
     +--+                             
        +-- Bineurus_exilis **

Konopleva et al., 2019 [Publ 3900], fig. 1

iT tT bi riS oT roT roS iD
3 8 0 3 49 5 2 8.0 
  +-- Bineurus_mouhotii **         
  |                             
  +-- Pseudodon_vagulus         
  |                             
  +-- Pseudodon_mekongi         
  |                             
  +-- Pilsbryoconcha_exilis     
--+                             
  +-- Pilsbryoconcha_carinifera 
  |                             
  +-- Bineurus_loeiensis **        
  |                             
  +-- Bineurus_exilis **           
  |                             
  +-- Pseudodon_vondembuschianus

Bolotov et al., 2020 [Publ 4100], fig. 1

iT tT bi riS oT roT roS iD
3 3 1 3 117 15 5 2.0 
                 +----- Bineurus_loeiensis **           
  +--------------+                                   
  |              |  +-- Bineurus_exilis **              
  |              +--+                                
  |                 +-- Bineurus_mouhotii **            
  |                                                  
--+  +----------------- Nyeinchanconcha_nyeinchani   
  |  |                                               
  |  |              +-- Thaiconcha_callifera         
  |  |        +-----+                                
  |  |        |     +-- Thaiconcha_munelliptica      
  +--+  +-----+                                      
     |  |     |  +----- Pilsbryoconcha_exilis[2]     
     |  |     +--+                                   
     |  |        |  +-- Pilsbryoconcha_exilis[1]     
     |  |        +--+                                
     |  |           +-- Pilsbryoconcha               
     +--+                                            
        |     +-------- Pseudodon_lenyanensis        
        |  +--+                                      
        |  |  |  +----- Pseudodon_vagulus            
        |  |  +--+                                   
        |  |     |  +-- Pseudodon_vondembuschianus   
        |  |     +--+                                
        +--+        +-- Pseudodon_mekongi            
           |                                         
           |        +-- Sundadontina_taskaevi        
           |  +-----+                                
           |  |     +-- Sundadontina_brandti         
           +--+                                      
              |  +----- Sundadontina_tumida          
              +--+                                   
                 |  +-- Sundadontina_cumingii        
                 +--+                                
                    +-- Sundadontina_tanintharyiensis

Bolotov et al., 2020 [Publ 4100], fig. S1

iT tT bi riS oT roT roS iD
3 3 1 3 111 15 5 2.0 
                 +----- Bineurus_loeiensis **           
  +--------------+                                   
  |              |  +-- Bineurus_exilis **              
  |              +--+                                
  |                 +-- Bineurus_mouhotii **            
  |                                                  
--+  +----------------- Nyeinchanconcha_nyeinchani   
  |  |                                               
  |  |              +-- Thaiconcha_callifera         
  |  |        +-----+                                
  |  |        |     +-- Thaiconcha_munelliptica      
  +--+  +-----+                                      
     |  |     |  +----- Pilsbryoconcha_exilis[2]     
     |  |     +--+                                   
     |  |        |  +-- Pilsbryoconcha_exilis[1]     
     |  |        +--+                                
     |  |           +-- Pilsbryoconcha               
     +--+                                            
        |     +-------- Pseudodon_lenyanensis        
        |  +--+                                      
        |  |  |  +----- Pseudodon_vagulus            
        |  |  +--+                                   
        |  |     |  +-- Pseudodon_vondembuschianus   
        |  |     +--+                                
        +--+        +-- Pseudodon_mekongi            
           |                                         
           |        +-- Sundadontina_tanintharyiensis
           |  +-----+                                
           |  |     +-- Sundadontina_cumingii        
           +--+                                      
              |  +----- Sundadontina_tumida          
              +--+                                   
                 |  +-- Sundadontina_taskaevi        
                 +--+                                
                    +-- Sundadontina_brandti

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

iT tT bi riS oT roT roS iD
3 3 0 3 33 15 6 3.0 
  +-------- Nyeinchanconcha_nyeinchani
  |                                   
--+  +----- Thaiconcha_callifera      
  |  |                                
  +--+  +-- Bineurus_panhai **           
     |  |                             
     +--+-- Bineurus_exilis **           
        |                             
        +-- Bineurus_mouhotii **

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

iT tT bi riS oT roT roS iD
3 3 1 3 33 15 6 4.0 
           +-- Nyeinchanconcha_nyeinchani
  +--------+                             
  |        +-- Sundadontina_cumingii     
--+                                      
  |  +-------- Thaiconcha_callifera      
  +--+                                   
     |  +----- Bineurus_mouhotii **         
     +--+                                
        |  +-- Bineurus_panhai **           
        +--+                             
           +-- Bineurus_exilis **

Dai et al., 2023 [Publ 4285], fig. 3-BI

iT tT bi riS oT roT roS iD
3 3 1 3 42 11 5 2.0 
     +-------- Nyeinchanconcha_nyeinchani   
  +--+                                      
  |  |  +----- Bineurus_loeiensis **           
  |  +--+                                   
  |     |  +-- Bineurus_mouhotii **            
  |     +--+                                
  |        +-- Bineurus_exilis **              
  |                                         
  |        +-- Thaiconcha_callifera         
  |  +-----+                                
  |  |     +-- Thaiconcha_munelliptica      
--+--+                                      
  |  |  +----- Sundadontina_tanintharyiensis
  |  +--+                                   
  |     |  +-- Sundadontina_brandti         
  |     +--+                                
  |        +-- Sundadontina_taskaevi        
  |                                         
  |        +-- Pilsbryoconcha_carinifera    
  |  +-----+                                
  |  |     +-- Pilsbryoconcha_exilis        
  +--+                                      
     |  +----- Pseudodon_lenyanensis        
     +--+                                   
        |  +-- Pseudodon_mekongi            
        +--+                                
           +-- Pseudodon_vagulus

Dai et al., 2023 [Publ 4285], fig. 3-ML

iT tT bi riS oT roT roS iD
3 3 1 3 42 11 5 2.0 
        +-------- Nyeinchanconcha_nyeinchani   
  +-----+                                      
  |     |  +----- Bineurus_loeiensis **           
  |     +--+                                   
  |        |  +-- Bineurus_mouhotii **            
  |        +--+                                
  |           +-- Bineurus_exilis **              
  |                                            
--+           +-- Thaiconcha_callifera         
  |     +-----+                                
  |     |     +-- Thaiconcha_munelliptica      
  |  +--+                                      
  |  |  |  +----- Sundadontina_tanintharyiensis
  |  |  +--+                                   
  |  |     |  +-- Sundadontina_brandti         
  |  |     +--+                                
  +--+        +-- Sundadontina_taskaevi        
     |                                         
     |        +-- Pilsbryoconcha_carinifera    
     |  +-----+                                
     |  |     +-- Pilsbryoconcha_exilis        
     +--+                                      
        |  +----- Pseudodon_lenyanensis        
        +--+                                   
           |  +-- Pseudodon_mekongi            
           +--+                                
              +-- Pseudodon_vagulus

Zieritz et al., 2021 [Publ 4218], fig. 5

iT tT bi riS oT roT roS iD
2 2 0 2 30 5 4 -- 
  +-------- Sundadontina        
--+                             
  |  +----- Thaiconcha_callifera
  +--+                          
     |  +-- Bineurus_loeiensis **  
     +--+                       
        +-- Bineurus_anodontinum **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1, 1

iT tT bi riS oT roT roS iD
2 2 0 2 74 13 6 -- 
        +-- Nyeinchanconcha_nyeinchani
  +-----+                             
  |     +-- Sundadontina_cumingii     
--+                                   
  |  +----- Thaiconcha_callifera      
  +--+                                
     |  +-- Bineurus_panhai **           
     +--+                             
        +-- Bineurus_mouhotii **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2, 2

iT tT bi riS oT roT roS iD
2 2 0 2 74 13 6 -- 
        +-- Nyeinchanconcha_nyeinchani
  +-----+                             
  |     +-- Sundadontina_cumingii     
--+                                   
  |  +----- Thaiconcha_callifera      
  +--+                                
     |  +-- Bineurus_panhai **           
     +--+                             
        +-- Bineurus_mouhotii **

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 39 5 4 -- 
  +----- Sundadontina_tumida       
--+                                
  |  +-- Nyeinchanconcha_nyeinchani
  +--+                             
     +-- Bineurus_exilis **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iT tT bi riS oT roT roS iD
1 1 0 1 80 4 3 -- 
  +----- Pseudodon_vagulus         
--+                                
  |  +-- Bineurus_exilis **           
  +--+                             
     +-- Nyeinchanconcha_nyeinchani

Konopleva et al., 2017 [Publ 3807], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 30 5 4 -- 
        +-- Nyeinchanconcha_nyeinchani
  +-----+                             
  |     +-- Bineurus_exilis **           
--+                                   
  |  +----- Pilsbryoconcha_carinifera 
  +--+                                
     |  +-- Pseudodon_vagulus         
     +--+                             
        +-- Pseudodon_vondembuschianus

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iT tT bi riS oT roT roS iD
1 1 0 1 56 3 2 -- 
  +-------- Bineurus_anodontinum **      
--+                                   
  |  +----- Pilsbryoconcha_exilis     
  +--+                                
     |  +-- Pseudodon_vondembuschianus
     +--+                             
        +-- Pseudodon_inoscularis

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 56 3 2 -- 
  +-------- Pilsbryoconcha_exilis     
--+                                   
  |  +----- Bineurus_anodontinum **      
  +--+                                
     |  +-- Pseudodon_vondembuschianus
     +--+                             
        +-- Pseudodon_inoscularis

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iT tT bi riS oT roT roS iD
1 1 0 1 121 5 4 -- 
  +----- Pseudodon_mekongi         
  |                                
--+  +-- Nyeinchanconcha_nyeinchani
  |  |                             
  +--+-- Bineurus_mouhotii **         
     |                             
     +-- Sundadontina_tumida

Page updated: 09:08:10 Thu 07 Dec 2023
 
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