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Page last updated
Wed 06 Nov 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Unionoida | Unionoidea | Unionidae | Gonideinae | Pseudodontini | Pseudodontina

genus Bineurus Simpson, 1900: cladomics
     5 valid species

Cladomics Summary

number of species = 5

analyzed in 35 cladograms in 22 publications
maximum number of terminals in an analysis = 5, mean = 2.63

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 29
maximum number of included terminals = 5, mean = 2.97
proportion of cladograms that support basic monophyly = 0.9

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ tree

Bolotov et al., 2023 [Publ 4312], fig. S2

iT tT bi riS oT roT roS iD
5 5 3 5 36 31 7 0.0

*** Top Monophyly Cladogram *** *** Top Sister-Group Cladogram ***

  +-------------------------------- Songkhlanaia                   
  |                                                                
  |              +----------------- Nyeinchanconcha                
  |              |                                                 
  |  +-----------+  +-------------- Sundadontina_tumida            
  |  |           |  |                                              
  |  |           |  |        +----- Sundadontina_tanintharyiensis  
  |  |           +--+  +-----+                                     
--+  |              |  |     |  +-- Sundadontina_plugpomenica      
  |  |              |  |     +--+                                  
  |  |              +--+        +-- Sundadontina_cumingii          
  |  |                 |                                           
  |  |                 |  +-------- Sundadontina_sulcata           
  |  |                 +--+                                        
  |  |                    |  +----- Sundadontina_harmandi          
  |  |                    +--+                                     
  +--+                       |  +-- Sundadontina_taskaevi          
     |                       +--+                                  
     |                          +-- Sundadontina_brandti           
     |                                                             
     |           +----------------- Namkongnaia                    
     |  +--------+                                                 
     |  |        |  +-------------- Thaiconcha                     
     |  |        +--+                                              
     |  |           |  +----------- Bineurus_loeiensis **             
     |  |           +--+                                           
     |  |              |  +-------- Bineurus_panhai **                
     |  |              +--+                                        
     +--+                 |  +----- Bineurus_anodontinum **           
        |                 +--+                                     
        |                    |  +-- Bineurus_exilis **                
        |                    +--+                                  
        |                       +-- Bineurus_mouhotii **              
        |                                                          
        |  +----------------------- Pseudodon                      
        |  |                                                       
        +--+  +-------------------- Pilsbryoconcha_carinifera      
           |  |                                                    
           +--+  +----------------- Pilsbryoconcha_linguaeformis__2
              |  |                                                 
              +--+  +-------------- Pilsbryoconcha_linguaeformis__1
                 |  |                                              
                 +--+           +-- Pilsbryoconcha_schomburgki     
                    |  +--------+                                  
                    |  |        +-- Pilsbryoconcha_exilis          
                    +--+                                           
                       |  +-------- Pilsbryoconcha_hoikaab         
                       +--+                                        
                          |  +----- Pilsbryoconcha_mekongiana      
                          +--+                                     
                             |  +-- Pilsbryoconcha_acuta           
                             +--+                                  
                                +-- Pilsbryoconcha_kittitati

Jeratthitikul & Sutcharit, 2023 [Publ 4289], fig. 1

iT tT bi riS oT roT roS iD
5 5 3 5 8 6 6 6.0 
     +----------- Bineurus_mouhotii **   
  +--+                                
  |  |  +-------- Bineurus_loeiensis **  
  |  +--+                             
  |     |  +----- Bineurus_panhai **     
  |     +--+                          
  |        |  +-- Bineurus_anodontinum **
--+        +--+                       
  |           +-- Bineurus_exilis **     
  |                                   
  |        +----- Namkongnaia         
  |     +--+                          
  |     |  |  +-- Sundadontina        
  |     |  +--+                       
  +-----+     +-- Nyeinchanconcha     
        |                             
        |  +----- Pseudodon           
        +--+                          
           |  +-- Pilsbryoconcha      
           +--+                       
              +-- Thaiconcha

Bolotov et al., 2023 [Publ 4312], fig. 2

iT tT bi riS oT roT roS iD
4 4 2 4 35 30 7 3.0 
  +-------------------------- Pilsbryoconcha               
  |                                                        
  |  +----------------------- Songkhlanaia                 
--+  |                                                     
  |  |                    +-- Namkongnaia                  
  |  |  +-----------------+                                
  +--+  |                 +-- Pseudodon                    
     |  |                                                  
     |  |     +-------------- Nyeinchanconcha              
     |  |  +--+                                            
     |  |  |  |  +----------- Thaiconcha                   
     +--+  |  +--+                                         
        |  |     |  +-------- Bineurus_loeiensis **           
        |  |     +--+                                      
        |  |        |  +----- Bineurus_mouhotii **            
        |  |        +--+                                   
        |  |           |  +-- Bineurus_anodontinum **         
        +--+           +--+                                
           |              +-- Bineurus_exilis **              
           |                                               
           |        +-------- Sundadontina_sulcata         
           |     +--+                                      
           |     |  |  +----- Sundadontina_harmandi        
           |     |  +--+                                   
           |     |     |  +-- Sundadontina_brandti         
           +-----+     +--+                                
                 |        +-- Sundadontina_taskaevi        
                 |                                         
                 |  +-------- Sundadontina_tumida          
                 +--+                                      
                    |  +----- Sundadontina_tanintharyiensis
                    +--+                                   
                       |  +-- Sundadontina_cumingii        
                       +--+                                
                          +-- Sundadontina_plugpomenica

Bolotov et al., 2023 [Publ 4312], fig. S1

iT tT bi riS oT roT roS iD
4 4 2 4 35 30 7 3.0 
  +-------------------------- Pilsbryoconcha               
  |                                                        
  |           +-------------- Nyeinchanconcha              
  |  +--------+                                            
  |  |        |  +----------- Thaiconcha                   
--+  |        +--+                                         
  |  |           |  +-------- Bineurus_loeiensis **           
  |  |           +--+                                      
  |  |              |  +----- Bineurus_mouhotii **            
  |  |              +--+                                   
  +--+                 |  +-- Bineurus_anodontinum **         
     |                 +--+                                
     |                    +-- Bineurus_exilis **              
     |                                                     
     |  +-------------------- Songkhlanaia                 
     |  |                                                  
     |  |                 +-- Namkongnaia                  
     +--+  +--------------+                                
        |  |              +-- Pseudodon                    
        |  |                                               
        +--+  +-------------- Sundadontina_tumida          
           |  |                                            
           |  |        +----- Sundadontina_tanintharyiensis
           +--+  +-----+                                   
              |  |     |  +-- Sundadontina_cumingii        
              |  |     +--+                                
              +--+        +-- Sundadontina_plugpomenica    
                 |                                         
                 |  +-------- Sundadontina_sulcata         
                 +--+                                      
                    |  +----- Sundadontina_harmandi        
                    +--+                                   
                       |  +-- Sundadontina_brandti         
                       +--+                                
                          +-- Sundadontina_taskaevi

Konopleva et al., 2021 [Publ 4213], fig. 1

iT tT bi riS oT roT roS iD
4 4 2 4 23 15 6 3.0 
                 +----- Pilsbryoconcha_exilis__2
  +--------------+                              
  |              |  +-- Pilsbryoconcha_exilis__1
  |              +--+                           
  |                 +-- Namkongnaia             
--+                                             
  |                 +-- Pseudodon               
  |  +--------------+                           
  |  |              +-- Sundadontina            
  +--+                                          
     |  +-------------- Nyeinchanconcha         
     +--+                                       
        |  +----------- Thaiconcha              
        +--+                                    
           |  +-------- Bineurus_loeiensis **      
           +--+                                 
              |  +----- Bineurus_mouhotii **       
              +--+                              
                 |  +-- Bineurus_anodontinum **    
                 +--+                           
                    +-- Bineurus_exilis **

Konopleva et al., 2023 [Publ 4262], fig. 8

iT tT bi riS oT roT roS iD
4 4 2 4 78 22 7 3.0 
                             +-- Pilsbryoconcha_schomburgki  
                       +-----+                               
                       |     +-- Pilsbryoconcha_exilis__1    
  +--------------------+                                     
  |                    |  +----- Pilsbryoconcha_linguaeformis
  |                    +--+                                  
  |                       |  +-- Pilsbryoconcha_exilis__2    
--+                       +--+                               
  |                          +-- Pilsbryoconcha_carinifera   
  |                                                          
  |  +-------------------------- Songkhlanaia                
  +--+                                                       
     |  +----------------------- Namkongnaia                 
     +--+                                                    
        |  +-------------------- Pseudodon                   
        +--+                                                 
           |  +----------------- Sundadontina                
           +--+                                              
              |  +-------------- Nyeinchanconcha             
              +--+                                           
                 |  +----------- Thaiconcha                  
                 +--+                                        
                    |  +-------- Bineurus_loeiensis **          
                    +--+                                     
                       |  +----- Bineurus_mouhotii **           
                       +--+                                  
                          |  +-- Bineurus_anodontinum **        
                          +--+                               
                             +-- Bineurus_exilis **

Bolotov et al., 2018 [Publ 3826], fig. 2

iT tT bi riS oT roT roS iD
3 8 0 3 80 5 2 8.0 
  +-- Bineurus_mouhotii **         
  |                             
  +-- Pseudodon_vagulus         
  |                             
  +-- Pseudodon_mekongi         
  |                             
  +-- Pilsbryoconcha_exilis     
--+                             
  +-- Pilsbryoconcha_carinifera 
  |                             
  +-- Bineurus_loeiensis **        
  |                             
  +-- Bineurus_exilis **           
  |                             
  +-- Pseudodon_vondembuschianus

Huang et al., 2019 [Publ 3843], fig. S4

iT tT bi riS oT roT roS iD
3 3 1 3 195 9 5 2.0 
  +-------- Nyeinchanconcha       
  |                               
  +-------- Thaiconcha            
  |                               
  +-------- Sundadontina_cumingii 
  |                               
  +-------- Sundadontina_tumida__1
--+                               
  +-------- Sundadontina_tumida__2
  |                               
  +-------- Pilsbryoconcha        
  |                               
  +-------- Pseudodon             
  |                               
  |  +----- Bineurus_loeiensis **    
  +--+                            
     |  +-- Bineurus_mouhotii **     
     +--+                         
        +-- Bineurus_exilis **

Huang et al., 2019 [Publ 3843], fig. 2

iT tT bi riS oT roT roS iD
3 3 1 3 195 9 5 2.0 
  +----------------- Pilsbryoconcha    
  |                                    
--+              +-- Pseudodon         
  |  +-----------+                     
  |  |           +-- Sundadontina      
  +--+                                 
     |  +----------- Nyeinchanconcha   
     +--+                              
        |  +-------- Thaiconcha        
        +--+                           
           |  +----- Bineurus_loeiensis **
           +--+                        
              |  +-- Bineurus_mouhotii ** 
              +--+                     
                 +-- Bineurus_exilis **

Huang et al., 2019 [Publ 3843], fig. S3

iT tT bi riS oT roT roS iD
3 3 1 3 193 9 5 2.0 
  +----------------- Pilsbryoconcha       
  |                                       
  |              +-- Sundadontina_tumida  
--+  +-----------+                        
  |  |           +-- Pseudodon            
  |  |                                    
  +--+           +-- Nyeinchanconcha      
     |  +--------+                        
     |  |        +-- Sundadontina_cumingii
     +--+                                 
        |  +-------- Thaiconcha           
        +--+                              
           |  +----- Bineurus_loeiensis **   
           +--+                           
              |  +-- Bineurus_mouhotii **    
              +--+                        
                 +-- Bineurus_exilis **

Bolotov et al., 2017 [Publ 3761], fig. S2

iT tT bi riS oT roT roS iD
3 3 1 3 136 10 5 4.0 
  +-------- Thaiconcha        
--+                           
  |  +----- Bineurus_exilis **   
  +--+                        
     |  +-- Bineurus_mouhotii ** 
     +--+                     
        +-- Bineurus_loeiensis **

Bolotov et al., 2017 [Publ 3764], fig. S1

iT tT bi riS oT roT roS iD
3 3 1 3 122 9 5 4.0 
        +-- Nyeinchanconcha      
  +-----+                        
  |     +-- Sundadontina_cumingii
  |                              
  |     +-- Sundadontina_tumida  
  +-----+                        
  |     +-- Pseudodon            
--+                              
  |     +-- Thaiconcha           
  +-----+                        
  |     +-- Pilsbryoconcha       
  |                              
  |  +----- Bineurus_exilis **      
  +--+                           
     |  +-- Bineurus_mouhotii **    
     +--+                        
        +-- Bineurus_loeiensis **

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

iT tT bi riS oT roT roS iD
3 3 0 3 33 15 6 3.0 
              +-- Pilsbryoconcha_exilis       
  +-----------+                               
  |           +-- Pilsbryoconcha_linguaeformis
  |                                           
--+        +----- Sundadontina                
  |  +-----+                                  
  |  |     |  +-- Namkongnaia                 
  |  |     +--+                               
  +--+        +-- Pseudodon                   
     |                                        
     |  +-------- Nyeinchanconcha             
     |  |                                     
     +--+  +----- Thaiconcha                  
        |  |                                  
        +--+  +-- Bineurus_panhai **             
           |  |                               
           +--+-- Bineurus_exilis **             
              |                               
              +-- Bineurus_mouhotii **

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

iT tT bi riS oT roT roS iD
3 3 1 3 33 15 6 4.0 
                 +-- Pilsbryoconcha_exilis       
  +--------------+                               
  |              +-- Pilsbryoconcha_linguaeformis
  |                                              
  |              +-- Namkongnaia                 
  |        +-----+                               
--+        |     +-- Pseudodon                   
  |  +-----+                                     
  |  |     |  +----- Sundadontina_tumida         
  |  |     +--+                                  
  |  |        |  +-- Sundadontina_brandti        
  |  |        +--+                               
  +--+           +-- Sundadontina_taskaevi       
     |                                           
     |           +-- Nyeinchanconcha             
     |  +--------+                               
     |  |        +-- Sundadontina_cumingii       
     +--+                                        
        |  +-------- Thaiconcha                  
        +--+                                     
           |  +----- Bineurus_mouhotii **           
           +--+                                  
              |  +-- Bineurus_panhai **             
              +--+                               
                 +-- Bineurus_exilis **

Konopleva et al., 2019 [Publ 3900], fig. 1

iT tT bi riS oT roT roS iD
3 8 0 3 49 5 2 8.0 
  +-- Bineurus_mouhotii **         
  |                             
  +-- Pseudodon_vagulus         
  |                             
  +-- Pseudodon_mekongi         
  |                             
  +-- Pilsbryoconcha_exilis     
--+                             
  +-- Pilsbryoconcha_carinifera 
  |                             
  +-- Bineurus_loeiensis **        
  |                             
  +-- Bineurus_exilis **           
  |                             
  +-- Pseudodon_vondembuschianus

Dai et al., 2023 [Publ 4285], fig. 3-BI

iT tT bi riS oT roT roS iD
3 3 1 3 42 11 5 2.0 
           +-- Thaiconcha        
  +--------+                     
  |        +-- Sundadontina      
  |                              
  |        +-- Pilsbryoconcha    
--+--------+                     
  |        +-- Pseudodon         
  |                              
  |  +-------- Nyeinchanconcha   
  +--+                           
     |  +----- Bineurus_loeiensis **
     +--+                        
        |  +-- Bineurus_mouhotii ** 
        +--+                     
           +-- Bineurus_exilis **

Bolotov et al., 2020 [Publ 4100], fig. 1

iT tT bi riS oT roT roS iD
3 3 1 3 115 14 5 2.0 
           +----- Bineurus_loeiensis **      
  +--------+                              
  |        |  +-- Bineurus_exilis **         
  |        +--+                           
--+           +-- Bineurus_mouhotii **       
  |                                       
  |  +----------- Nyeinchanconcha         
  |  |                                    
  +--+        +-- Pseudodon               
     |  +-----+                           
     |  |     +-- Sundadontina            
     +--+                                 
        |  +----- Thaiconcha              
        +--+                              
           |  +-- Pilsbryoconcha_exilis__2
           +--+                           
              +-- Pilsbryoconcha_exilis__1

Dai et al., 2023 [Publ 4285], fig. 3-ML

iT tT bi riS oT roT roS iD
3 3 1 3 42 11 5 2.0 
     +-------- Nyeinchanconcha   
  +--+                           
  |  |  +----- Bineurus_loeiensis **
  |  +--+                        
  |     |  +-- Bineurus_mouhotii ** 
  |     +--+                     
--+        +-- Bineurus_exilis **   
  |                              
  |        +-- Thaiconcha        
  |     +--+                     
  |     |  +-- Sundadontina      
  +-----+                        
        |  +-- Pilsbryoconcha    
        +--+                     
           +-- Pseudodon

Bolotov et al., 2020 [Publ 4100], fig. S1

iT tT bi riS oT roT roS iD
3 3 1 3 109 14 5 2.0 
           +----- Bineurus_loeiensis **      
  +--------+                              
  |        |  +-- Bineurus_exilis **         
  |        +--+                           
--+           +-- Bineurus_mouhotii **       
  |                                       
  |  +----------- Nyeinchanconcha         
  |  |                                    
  +--+        +-- Pseudodon               
     |  +-----+                           
     |  |     +-- Sundadontina            
     +--+                                 
        |  +----- Thaiconcha              
        +--+                              
           |  +-- Pilsbryoconcha_exilis__2
           +--+                           
              +-- Pilsbryoconcha_exilis__1

Bolotov et al., 2017 [Publ 3761], fig. S1

iT tT bi riS oT roT roS iD
3 3 1 3 136 9 5 4.0 
  +----------------- Pilsbryoconcha       
  |                                       
  |              +-- Sundadontina_tumida  
--+  +-----------+                        
  |  |           +-- Pseudodon            
  |  |                                    
  +--+           +-- Nyeinchanconcha      
     |  +--------+                        
     |  |        +-- Sundadontina_cumingii
     +--+                                 
        |  +-------- Thaiconcha           
        +--+                              
           |  +----- Bineurus_exilis **      
           +--+                           
              |  +-- Bineurus_mouhotii **    
              +--+                        
                 +-- Bineurus_loeiensis **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iT tT bi riS oT roT roS iD
2 2 0 2 74 13 6 -- 
        +----- Pilsbryoconcha   
  +-----+                       
  |     |  +-- Namkongnaia      
  |     +--+                    
  |        +-- Pseudodon        
--+                             
  |        +-- Nyeinchanconcha  
  |  +-----+                    
  |  |     +-- Sundadontina     
  +--+                          
     |  +----- Thaiconcha       
     +--+                       
        |  +-- Bineurus_panhai **  
        +--+                    
           +-- Bineurus_mouhotii **

Zieritz et al., 2024 [Publ 4381], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 44 6 4 -- 
           +-- Pilsbryoconcha      
  +--------+                       
  |        +-- Pseudodon           
--+                                
  |  +-------- Sundadontina        
  +--+                             
     |  +----- Thaiconcha          
     +--+                          
        |  +-- Bineurus_loeiensis **  
        +--+                       
           +-- Bineurus_anodontinum **

Zieritz et al., 2021 [Publ 4218], fig. 5

iT tT bi riS oT roT roS iD
2 2 0 2 30 5 4 -- 
  +-------------- Pilsbryoconcha      
--+                                   
  |  +----------- Pseudodon           
  +--+                                
     |  +-------- Sundadontina        
     +--+                             
        |  +----- Thaiconcha          
        +--+                          
           |  +-- Bineurus_loeiensis **  
           +--+                       
              +-- Bineurus_anodontinum **

Wu et al., 2024 [Publ 4350], fig. 5b

iT tT bi riS oT roT roS iD
2 2 0 2 35 4 3 -- 
  +-------- Pilsbryoconcha      
  |                             
--+     +-- Bineurus_anodontinum **
  |  +--+                       
  |  |  +-- Bineurus_loeiensis **  
  +--+                          
     |  +-- Thaiconcha          
     +--+                       
        +-- Pseudodon

Liu et al., 2024 [Publ 4380], fig. 5

iT tT bi riS oT roT roS iD
2 2 0 2 35 4 3 -- 
  +----------- Pilsbryoconcha      
--+                                
  |  +-------- Pseudodon           
  +--+                             
     |  +----- Thaiconcha          
     +--+                          
        |  +-- Bineurus_anodontinum **
        +--+                       
           +-- Bineurus_loeiensis **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iT tT bi riS oT roT roS iD
2 2 0 2 74 13 6 -- 
        +----- Pilsbryoconcha   
  +-----+                       
  |     |  +-- Namkongnaia      
  |     +--+                    
  |        +-- Pseudodon        
--+                             
  |        +-- Nyeinchanconcha  
  |  +-----+                    
  |  |     +-- Sundadontina     
  +--+                          
     |  +----- Thaiconcha       
     +--+                       
        |  +-- Bineurus_panhai **  
        +--+                    
           +-- Bineurus_mouhotii **

Wu et al., 2024 [Publ 4350], fig. 6-7

iT tT bi riS oT roT roS iD
2 2 0 2 35 4 3 -- 
  +----------- Pilsbryoconcha      
--+                                
  |  +-------- Pseudodon           
  +--+                             
     |  +----- Thaiconcha          
     +--+                          
        |  +-- Bineurus_anodontinum **
        +--+                       
           +-- Bineurus_loeiensis **

Wu et al., 2024 [Publ 4350], fig. 5a

iT tT bi riS oT roT roS iD
2 2 0 2 35 4 3 -- 
  +----------- Pilsbryoconcha      
--+                                
  |  +-------- Pseudodon           
  +--+                             
     |  +----- Thaiconcha          
     +--+                          
        |  +-- Bineurus_anodontinum **
        +--+                       
           +-- Bineurus_loeiensis **

Wu et al., 2024 [Publ 4350], fig. 5c

iT tT bi riS oT roT roS iD
2 5 0 2 35 4 3 6.0 
  +-------- Pilsbryoconcha      
  |                             
--+  +----- Bineurus_anodontinum **
  |  |                          
  +--+----- Bineurus_loeiensis **  
     |                          
     |  +-- Thaiconcha          
     +--+                       
        +-- Pseudodon

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 56 3 3 -- 
  +-------- Pilsbryoconcha      
--+                             
  |  +----- Bineurus_anodontinum **
  +--+                          
     |  +-- Pseudodon           
     +--+                       
        +-- Sundadontina

Konopleva et al., 2017 [Publ 3807], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 30 5 4 -- 
  +-------- Sundadontina   
  |                        
--+     +-- Nyeinchanconcha
  |  +--+                  
  |  |  +-- Bineurus_exilis **
  +--+                     
     |  +-- Pilsbryoconcha 
     +--+                  
        +-- Pseudodon

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iT tT bi riS oT roT roS iD
1 1 0 1 79 4 3 -- 
  +-------- Pilsbryoconcha 
--+                        
  |  +----- Pseudodon      
  +--+                     
     |  +-- Bineurus_exilis **
     +--+                  
        +-- Nyeinchanconcha

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iT tT bi riS oT roT roS iD
1 1 0 1 119 5 4 -- 
  +-------- Pilsbryoconcha   
  |                          
--+  +----- Pseudodon        
  |  |                       
  +--+  +-- Nyeinchanconcha  
     |  |                    
     +--+-- Bineurus_mouhotii **
        |                    
        +-- Sundadontina

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iT tT bi riS oT roT roS iD
1 1 0 1 56 3 3 -- 
  +-------- Bineurus_anodontinum **
--+                             
  |  +----- Pilsbryoconcha      
  +--+                          
     |  +-- Pseudodon           
     +--+                       
        +-- Sundadontina

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 39 5 4 -- 
  +----------- Pilsbryoconcha 
--+                           
  |  +-------- Pseudodon      
  +--+                        
     |  +----- Sundadontina   
     +--+                     
        |  +-- Nyeinchanconcha
        +--+                  
           +-- Bineurus_exilis **

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