| Page last updated
Thu 15 May 2025 |
The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!
This page documents data for Dai et al. (2023), such as genera and species attributed to that publication (including known misspellings, misidentifications, and other unavailable names) as well as taxonomic opinions, if available. Taxonomic opinions refer to uses of genera and species as valid and invalid in that work.
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Publication information
Dai, Y., X.-C. Huang, C.-H.-Z. Wu, Z.-G. Chen, L. Guo et al. 2023. Multilocus and mitogenomic phylogenetic analyses reveal a new genus and species of freshwater mussel (Bivalvia: Unionidae) from Guangxi, China. Invertebrate Systematics 37(2): 152-166.
Click here for online access.
Cladogram Figures
(click the figure number to see the cladogram) fig. 3-BI
fig. 3-ML
fig. 5
Attributed Genus Names
Attributed Species Names
Unio crassus Soroka, 2005
‘Philippson’ of Authors.
Soroka, 2005, Zool. Sci. 22: 1138, Table 1.
Soroka, 2010, Folia Malac. 18(4): Table 2. [in part]
Reis et al., 2013, Graellsia 69(1): 22, Table 2.
Prié & Puillandre, 2014, Hydrobiologia 735(1): online resource 1 [as ‘Unio crassus crassus’].
Douda et al., 2014, J. Appl. Ecol. 51: 1087. [in part]
Froufe et al., 2016, Org Divers Evol 16(3): 606, fig. 3. [in part]
Klishko et al., 2017, Zootaxa 4286(1): 98, Table 3.
Soroka & Burzynski, 2017, mtDNA B 2(2): 863.
Burzynski et al., 2017, MPE 107: 606.
Lopes-Lima et al., 2017, Biol. Rev. 92(1): Table S1 [as ‘Unio crassus crassus’]. [in part]
Araujo et al., 2018, J. Linn. Soc., Zool. 182(2): supplementary table. [in part]
Klishko et al., 2018, Systemat. Biodiv. 16(3): 291.
Burzynski & Soroka, 2018, PeerJ 6, e5573: 3.
Wang et al., 2018, Conserv. Genet. Resour. 10(4): 611.
Wang et al., 2018, Conserv. Genet. Resour. 11: 140.
Cha et al., 2018, mtDNA B 3(2): 946.
Aunins et al., 2018, mtDNA B 3(2): 1125.
Wu et al., 2018, PLoS One 13(8), e0200956: 6. [in part]
Klishko et al., 2019, Ruthenica 29(1): 57, Table 1.
Wu et al., 2019, ZooKeys 812: 29.
Huang et al., 2019, MPE 130: Table S5. [in part]
Fukata & Iigo, 2019, mtDNA B 4(2): 3489.
Guerra et al., 2019, BMC Evol. Biol. 19 (229): Table S1.
Froufe et al., 2019, Heredity 124: 185.
Soroka, 2020, J. Zool. Syst. Evol. Res. 58(2): Table 2.
Fukata & Iigo, 2020, mtDNA B 5(2): 1216.
Sano et al., 2020, Plankton Benthos Res. 15(2): 99.
Wu et al., 2020, Invert. Syst. 34: Table S2.
Kilikowska et al., 2020, Life 10(7): 1. [in part]
Lopes-Lima et al., 2021, MPE 163: fig. 6 [by voucher number]. [in part]
Sayenko et al., 2021, Systemat. Biodiv. 19: 58, Table 1.
Wu et al., 2021, J. Lake Sci. 33(6): attached Table 2.
Zieritz et al., 2021, Zool. J. Linn. Soc. 193(3): 1092.
Kim et al., 2021, mtDNA B 6(1): 80.
Pfarrer et al., 2022, Hydrobiologia 849: ESM1.
Wu et al., 2022, Zool. Scripta 51: Table S2.
Dai et al., 2023, Invert. Syst. 37(2): Table 2.
Chen et al., 2023, ZooKeys 1173: Table 1.
Wu et al., 2023, Org Divers Evol (2023): Table 2.
Dai et al., 2024, Zool. Scripta 53: Table 1.
Wu et al., 2024, J. Zool. Syst. Evol. Res. 2024, 1499508: Table 2.
Wu et al., 2024, Zool. Scripta early online: Table S3.
Dai et al., 2024, Invert. Syst. 38(6): Table 3. [in part]
(misidentified reference to Unio crassus Retzius, 1788)
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Potomida littoralis Whelan et al., 2011
‘Cuvier’ of Authors.
Whelan et al., 2011, MPE 61: 506. [in part]
Bolotov et al., 2020, Sci. Repts. 10: Table S1.
Konopleva et al., 2021, Sci. Repts. 11: Table S1.
Jeratthitikul et al., 2021, Eur. J. Taxonomy 775: 125.
Bolotov et al., 2022, Hindawi 6600359: Table S1.
Jeratthitikul et al., 2022, Sci. Repts. 12: Table S1.
Konopleva et al., 2023, Diversity 15(1), 10: Table S2.
Konopleva et al., 2023, Zoosyst. Evol. 99(1): Table S1.
Dai et al., 2023, Invert. Syst. 37(2): Table 1. [in part]
Kongim et al., 2023, Tropical Nat. Hist. 7: Table 1.
Jeratthitikul et al., 2024, Zoosyst. Evol. 100(4): Table S1.
Dai et al., 2024, Zoosyst. Evol. 100(4): Table 2.
Jeratthitikul et al., 2025, ZooKeys 1224: Table S1.
(misidentified reference to Unio littoralis Cuvier, 1798)
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Pseudodon cambodjensis Pfeiffer & Graf, 2013
‘Petit’ of Authors.
Pfeiffer & Graf, 2013, J. Moll. Studies 79: 251.
Pfeiffer & Graf, 2015, Zool. J. Linn. Soc. 175(2): 310.
Zieritz et al., 2016, Sci. Total Environ. 571: 1071.
Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Monodontina cambodjensis’].
Lopes-Lima et al., 2017, MPE 106: 177.
Konopleva et al., 2017, Systemat. Biodiv. 15(3): 207.
Bolotov et al., 2018, Sci. Repts. 8: Table S1 [as ‘Monodontina cambodjensis’].
Razak et al., 2018, Conserv. Genet. Resour. 11: 459.
Huang et al., 2019, MPE 130: Table S4 [as ‘Monodontina cambodjensis’].
Konopleva et al., 2019, Sci. Repts. 9: Table S1 [as ‘Monodontina cambodjensis’].
Bolotov et al., 2020, Sci. Repts. 10: Table S1 [as ‘Monodontina cambodiensis’ (sic)].
Konopleva et al., 2021, Sci. Repts. 11: Table S1 [as ‘Monodontina cambodiensis’ (sic)].
Jeratthitikul et al., 2021, Eur. J. Taxonomy 775: 123 [as ‘Monodontina cambodiensis’ (sic)].
Pfeiffer et al., 2021, Invert. Syst. 35: 453 [as ‘Monodontina cambodjensis’].
Wu et al., 2021, ZooKeys 1054: 87 [as ‘Monodontina cambodjensis’].
Wu et al., 2021, J. Lake Sci. 33(6): attached Table 1.
Konopleva et al., 2023, Diversity 15(1), 10: Table S2 [as ‘Monodontina cambodjensis’].
Dai et al., 2023, Invert. Syst. 37(2): Table 1 [as ‘Monodontina cambodjensis’].
Dai et al., 2024, Zoosyst. Evol. 100(4): Table 2.
(misidentified reference to Monocondylea cambodjensis Petit de la Saussaye, 1865)
Type(s): SEQUENCED SPECIMEN UMMZ_304350, Pursat River, just downstream (north) of Highway 5 bridge, Pursat City, Cambodia.
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Chamberlainia hainesiana Kongim et al., 2015
‘Lea’ of Authors.
Kongim et al., 2015, ZooKeys 514: 98, fig. 2A.
Lopes-Lima et al., 2017, MPE 106: 177.
Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Froufe et al., 2019, Heredity 124: 185.
Huang et al., 2019, MPE 130: Table S4.
Chen et al., 2020, mtDNA B 5(3): 2800.
Teiga-Teixeira et al., 2020, mtDNA B 5(3): 2978.
Zieritz et al., 2020, Biodivers. Conserv. 29: 2244.
Bolotov et al., 2020, Sci. Repts. 10: Table S1.
Lopes-Lima et al., 2020, MPE 146: Table 1.
Zieritz et al., 2021, Zool. J. Linn. Soc. 193(3): Table 1.
Zieritz et al., 2021, Aquatic Conserv.: Mar. Freshw. Ecosyst. 31: 3176.
Jeratthitikul et al., 2021, Eur. J. Taxonomy 775: 125.
Wu et al., 2021, ZooKeys 1054: 87.
Wu et al., 2021, J. Lake Sci. 33(6): attached Table 1.
Wu et al., 2022, Zool. Scripta 51: Table S2.
Pavan-Kumar et al., 2022, Mol. Biol. Rep. 49: Table S1.
Jeratthitikul et al., 2022, Sci. Repts. 12: Table S1.
Dai et al., 2023, Invert. Syst. 37(2): Tables 1-2.
Konopleva et al., 2023, Zoosyst. Evol. 99(1): Table S1.
Wu et al., 2023, Org Divers Evol (2023): Table 2.
Dai et al., 2024, Zool. Scripta 53: Table 1.
Dai et al., 2024, Invert. Syst. 38(6): Table 3.
(misidentified reference to Unio hainesianus Lea, 1856)
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Pilsbryoconcha exilis Pfeiffer & Graf, 2015
‘Lea’ of Authors.
Pfeiffer & Graf, 2015, Zool. J. Linn. Soc. 175(2): 310.
Ng et al., 2016, PLoS One 11(8), e0161130: [5]. [in part]
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘P. aff. exilis sp. 1’].
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘P. aff. exilis sp. 1’].
Konopleva et al., 2017, Systemat. Biodiv. 15(3): 207.
Bolotov et al., 2018, Sci. Repts. 8: Table S1 [as ‘P. aff. exilis sp. 1’].
Huang et al., 2019, MPE 130: Table S4 [as ‘P. aff. exilis sp. 1’].
Konopleva et al., 2019, Sci. Repts. 9: Table S1 [as ‘P. aff. exilis sp. 1’].
Lopes-Lima et al., 2020, MPE 146: Table 1 [as ‘P. aff. exilis sp. 1’].
Zieritz et al., 2020, Biodivers. Conserv. 29: 2244.
Wu et al., 2021, ZooKeys 1054: 87.
Wu et al., 2021, J. Lake Sci. 33(6): attached Table 1.
Wu et al., 2021, J. Lake Sci. 33(6): attached Table 1.
Konopleva et al., 2023, Diversity 15(1), 10: Table S2 [as ‘P. aff. exilis’].
Dai et al., 2023, Invert. Syst. 37(2): Tables 1-2. [in part]
(misidentified reference to Anodonta exilis Lea, 1838)
Type(s): SEQUENCED SPECIMEN UMMZ_304647, Beung Dang Teuk Leach Community Fishery Pursat Province, Cambodia.
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Pseudodon avae Bolotov et al., 2017
‘Theobald’ of Authors.
Bolotov et al., 2017, Sci. Repts. 7: Table S1 [as ‘Pseudodon cf. avae’].
Bolotov et al., 2017, Sci. Repts. 7: Table S1.
Bolotov et al., 2018, Sci. Repts. 8: Table S1.
Konopleva et al., 2019, Ecologica Montenegrina 24: Appendix 1 [as ‘Pseudodon cf. avae’].
Huang et al., 2019, MPE 130: Table S4.
Konopleva et al., 2019, Sci. Repts. 9: Table S1.
Zieritz et al., 2020, Biodivers. Conserv. 29: 2244.
Lopes-Lima et al., 2020, MPE 146: Table 1.
Dai et al., 2023, Invert. Syst. 37(2): 155, Table 1.
(misidentified reference to Monocondylaea avae Theobald, 1873)
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Postolata guangxiensis Dai et al., 2023
Dai, Huang, Guo & Wu in Dai et al., 2023, Invert. Syst. 37(2): 161. figs. 1c, 2.
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Taxonomic Opinions
Aculamprotula tientsinensis (Crosse & Debeaux, 1863)
Acuticosta chinensis (Lea, 1868)
Amblema plicata (Say, 1817)
Anemina arcaeformis (Heude, 1877)
Anodonta cygnea (Linnaeus, 1758)
Bineurus exilis
Bineurus loeiensis
Bineurus mouhotii
Chamberlainia hainesiana (Lea, 1856)
Cristaria plicata (Leach, 1814)
Cuneopsis celtiformis (Heude, 1874)
Elliptio complanata (Lightfoot, 1786)
Ensidens ingallsianus
Gibbosula laosensis
Gibbosula rochechouartii (Heude, 1875)
Gonidea angulata
Hyriopsis bialata Simpson, 1900
Inversidens brandtii
Inversidens rentianensis
Koreosolenaia sitgyensis
Lamprotula caveata (Heude, 1877)
Lamprotula leaii (Griffith & Pidgeon, 1833)
Lampsilis cardium Rafinesque, 1820
Lampsilis ornata (Conrad, 1835)
Lampsilis powellii (Lea, 1852)
Lampsilis siliquoidea (Barnes, 1823)
Lanceolaria grayii (Gray, 1833) [sic]
Lasmigona compressa (Lea, 1829)
Leaunio lienosus (Conrad, 1834)
Leguminaia wheatleyi
Lens comptus
Lens contradens (Lea, 1838)
Lepidodesma languilati (Heude, 1874)
Margaritifera dahurica (Middendorff, 1850)
Margaritifera margaritifera
Microcondylaea bonellii (Férussac, 1827)
Monodontina cambodjensis
Monodontina lenyanensis
Monodontina mekongi
Monodontina vondembuschiana (Lea, 1840)
Nodularia douglasiae (Griffith & Pidge) [sic]
Nyeinchanconcha nyeinchani
Obovalis omiensis
Parvasolenaia rivularis (Heude, 1877)
Physunio superbus (Lea, 1843)
Pilsbryoconcha compressa
Pilsbryoconcha exilis (Lea, 1838)
Pleurobema oviforme (Conrad, 1834)
Popenaias popeii (Lea, 1857)
Postolata guangxiensis Dai, Huang, Guo & Wu gen. nov., sp. nov.
Potamilus alatus (Say, 1817)
Potamilus leptodon (Rafinesque, 1820)
Potomida littoralis (Cuvier, 1798)
Pronodularia japanensis (Lea, 1859)
Pseudocuneopsis capitata (Heude, 1874)
Pseudodon avae
Pseudodon bogani
Pseudodon manueli
Ptychorhynchus pfisteri (Heude, 1874)
Pyganodon grandis (Say, 1829)
Quadrula quadrula (Rafinesque, 1820)
Rectidens sumatrensis (Dunker, 1852)
Schistodesmus lampreyanus (Baird & Adams, 1867)
Sinanodonta woodiana (Lea, 1834)
Sinohyriopsis cumingii (Lea, 1852)
Sinohyriopsis schlegelii (Martens, 1861)
Sinosolenaia carinata (Heude, 1877)
Sinosolenaia iridinea
Sinosolenaia oleivora (Heude, 1877)
Sinosolenaia recognita
Sundadontina brandti
Sundadontina tanintharyiensis
Sundadontina taskaevi
Tchangsinaia piscicula (Heude, 1874)
Thaiconcha callifera
Thaiconcha munelliptica
Toxolasma parvum (Barnes, 1823)
Unio crassus Retzius, 1788
Uniomerus tetralasmus (Say, 1831)
Utterbackia imbecillis (Say, 1829)
Venustaconcha ellipsiformis (Conrad, 1836)
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Cladogram Figure 3-BI
+-- Gibbosula_laosensis
+-----------------------+
| +-- Margaritifera_margaritifera
|
| +----- Indopseudodon_indawgyiensis
| +--------+
| | | +-- Indopseudodon_bogani
| | +--+
| | +-- Indopseudodon_salwenianus
| |
| | +-------- Nyeinchanconcha_nyeinchani
| | +--+
| | | | +----- Bineurus_loeiensis
| +--------+ | +--+
| | | | | +-- Bineurus_mouhotii
| | | | +--+
--+ | | | +-- Bineurus_exilis
| | | |
| | | | +-- Thaiconcha_callifera
| | | | +-----+
| | | | | +-- Thaiconcha_munelliptica
| | +--+--+
| | | | +----- Sundadontina_tanintharyiensis
| | | +--+
| | | | +-- Sundadontina_brandti
| | | +--+
| | | +-- Sundadontina_taskaevi
| | |
| | | +-- Pilsbryoconcha_carinifera
| | | +-----+
| | | | +-- Pilsbryoconcha_exilis
| | +--+
+--+ | +----- Pseudodon_lenyanensis
| +--+
| | +-- Pseudodon_mekongi
| +--+
| +-- Pseudodon_vagulus
|
| +-- Hyriopsis_bialata
| +-----+
| | +-- Ensidens_ingallsianus
| +-----------+
| | | +----- Physunio_superbus
| | +--+
| | | +-- Lens_contradens
| | +--+
| | +-- Lens_comptus
| |
| | +-- Inversidens_brandtii
| | +--------------+
| | | +-- Inversidens_rentianensis
+--+ |
| | +----- Chamberlainia_somsakpanhai
| | +-----+
| | | | +-- Sinohyriopsis_cumingii
| | | +--+
| +-----+ +-- Sinohyriopsis_schlegelii
| | |
| | | +-------- Potomida_semirugata
| | +--+
| | | +----- Pronodularia_japanensis
+--+ +--+
| | +-- Lamprotula_leaii
| +--+
| +-- Lamprotula_caveata
|
| +-------- Sinosolenaia_carinata
| +-----+
| | | +----- Sinosolenaia_recognita
| | +--+
| | | +-- Sinosolenaia_iridinea
| | +--+
| | +-- Sinosolenaia_oleivora
+--+
| +----- Gonidea_angulata
| +-----+
| | | +-- Leguminaia_wheatleyi
| | +--+
| | +-- Microcondylaea_bonellii
+--+
| +-- Koreosolenaia_sitgyensis
| +-----+
| | +-- Parvasolenaia_rivularis
+--+
| +----- Postolata_guangxiensis
+--+
| +-- Ptychorhynchus_pfisteri
+--+
+-- Obovalis_omiensis
Cladogram Figure 3-ML
+-- Gibbosula_laosensis
+-----------------------+
| +-- Margaritifera_margaritifera
|
| +----- Indopseudodon_indawgyiensis
| +-----------+
| | | +-- Indopseudodon_bogani
| | +--+
| | +-- Indopseudodon_salwenianus
| |
| | +-------- Nyeinchanconcha_nyeinchani
| +-----+ +-----+
| | | | | +----- Bineurus_loeiensis
| | | | +--+
| | | | | +-- Bineurus_mouhotii
| | | | +--+
--+ | | | +-- Bineurus_exilis
| | | |
| | +--+ +-- Thaiconcha_callifera
| | | +-----+
| | | | +-- Thaiconcha_munelliptica
| | | +--+
| | | | | +----- Sundadontina_tanintharyiensis
| | | | +--+
| | | | | +-- Sundadontina_brandti
| | | | +--+
| | +--+ +-- Sundadontina_taskaevi
| | |
| | | +-- Pilsbryoconcha_carinifera
| | | +-----+
| | | | +-- Pilsbryoconcha_exilis
+--+ +--+
| | +----- Pseudodon_lenyanensis
| +--+
| | +-- Pseudodon_mekongi
| +--+
| +-- Pseudodon_vagulus
|
| +-- Hyriopsis_bialata
| +-----+
| | +-- Ensidens_ingallsianus
| +-----------+
| | | +----- Physunio_superbus
| | +--+
| | | +-- Lens_contradens
| | +--+
| | +-- Lens_comptus
| |
| | +-- Inversidens_brandtii
| | +-----------+
| | | +-- Inversidens_rentianensis
+--+ |
| +--+ +----- Chamberlainia_somsakpanhai
| | | +-----+
| | | | | +-- Sinohyriopsis_cumingii
| | | | +--+
| | +--+ +-- Sinohyriopsis_schlegelii
| | |
| | | +-------- Potomida_semirugata
| | +--+
| | | +----- Pronodularia_japanensis
| | +--+
+--+ | +-- Lamprotula_leaii
| +--+
| +-- Lamprotula_caveata
|
| +-------- Sinosolenaia_carinata
| +-----+
| | | +----- Sinosolenaia_recognita
| | +--+
| | | +-- Sinosolenaia_iridinea
| | +--+
| | +-- Sinosolenaia_oleivora
+--+
| +----- Gonidea_angulata
| +-----+
| | | +-- Leguminaia_wheatleyi
| | +--+
| | +-- Microcondylaea_bonellii
+--+
| +-- Koreosolenaia_sitgyensis
| +-----+
| | +-- Parvasolenaia_rivularis
+--+
| +----- Postolata_guangxiensis
+--+
| +-- Ptychorhynchus_pfisteri
+--+
+-- Obovalis_omiensis
Cladogram Figure 5
+-- Gibbosula_rochechouartii
+--------------------------------+
| +-- Margaritifera_dahurica
|
| +-- Quadrula_quadrula
| +--------------------------+
| | +-- Uniomerus_tetralasmus
| |
| +--+ +-- Elliptio_complanata
| | | +-----------------------+
| | | | +-- Pleurobema_oviforme
| | +--+
| | | +----------------------- Amblema_plicata
| | | |
| | +--+ +-------------------- Popenaias_popeii
| | | |
--+ | +--+ +----------------- Toxolasma_parvum
| | | |
| | +--+ +-- Potamilus_leptodon
| | | +-----------+
| | | | +-- Potamilus_alatus
| | +--+
| | | +----------- Leaunio_lienosus
| | | |
| | +--+ +-- Lampsilis_cardium
| | | +-----+
| | | | +-- Lampsilis_ornata
| | +--+
| | | +----- Venustaconcha_ellipsiformis
| | +--+
| | | +-- Lampsilis_powellii
| | +--+
+--+ +-- Lampsilis_siliquoidea
|
| +-------------------- Lepidodesma_languilati
| |
| | +-------------- Aculamprotula_tientsinensis
| | |
| | +--+ +----------- Unio_nanus
| | | | |
| +--+ | +--+ +-- Pseudocuneopsis_capitata
| | | | | +-----+
| | | | | | +-- Nodularia_douglasiae
| | | | +--+
| | | | | +----- Schistodesmus_lampreyanus
| | | | +--+
| | +--+ | +-- Cuneopsis_celtiformis
| | | +--+
| | | +-- Tchangsinaia_piscicula
| | |
| | | +-- Acuticosta_chinensis
| | | +-----------+
| | | | +-- Lanceolaria_grayii
| | | |
| | +--+ +----- Cristaria_plicata
| | | +-----+
| | | | | +-- Anemina_arcaeformis
+--------+ | | +--+
| +--+ +-- Sinanodonta_woodiana
| |
| | +-------- Anodonta_cygnea
| +--+
| | +----- Platynaias_compressa
| +--+
| | +-- Pyganodon_grandis
| +--+
| +-- Utterbackia_imbecillis
|
| +----- Chamberlainia_somsakpanhai
| +--------------+
| | | +-- Sinohyriopsis_cumingii
| | +--+
| | +-- Sinohyriopsis_schlegelii
| |
| | +-- Hyriopsis_bialata
+--+ +--+
| | +-- Rectidens_sumatrensis
| +-----------+
| | | +-- Lens_contradens
| | +--+
| | +-- Physunio_superbus
| |
+--+ +-------- Potomida_littoralis
| +-----+
| | | +----- Pronodularia_japanensis
| | +--+
| | | +-- Lamprotula_caveata
| | +--+
+--+ +-- Lamprotula_leaii
|
| +-- Pseudodon_vondembuschianus
| +--------+
| | +-- Pilsbryoconcha_exilis
| |
+--+ +----- Parvasolenaia_rivularis
| +--+
| | | +-- Postolata_guangxiensis
| | +--+
+--+ +-- Ptychorhynchus_pfisteri
|
| +----- Microcondylaea_bonellii
+--+
| +-- Sinosolenaia_oleivora
+--+
+-- Sinosolenaia_carinata
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Page updated: 08:05:16 Thu 15 May 2025 |
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