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Page last updated
Thu 21 Mar 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Unionoida | Unionoidea | Unionidae | Unioninae | Anodontini | Cristariina

genus Beringiana Starobogatov in Zatravkin, 1983: cladomics
     4 valid species

Cladomics Summary

number of species = 4

analyzed in 31 cladograms in 21 publications
maximum number of terminals in an analysis = 20, mean = 2.68

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 14
maximum number of included terminals = 20, mean = 4.71
proportion of cladograms that support basic monophyly = 0.86


Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ tree


Mabuchi & Nishida, 2020 [Publ 4120], fig. 1

iT tT bi riS oT roT roS iD
20 28 3 20 17 10 2 0.0

*** Top Monophyly Cladogram ***

  +-------------- Cristaria_plicata       
  |                                       
  |  +----------- Sinanodonta_tumens      
  |  |                                    
  |  |        +-- Sinanodonta_lauta[5]    
  |  |        |                           
--+  |        +-- Sinanodonta_lauta[6]    
  |  |        |                           
  |  |        +-- Beringiana_japonica[15] ** 
  |  |        |                           
  |  |        +-- Beringiana_japonica[16] ** 
  |  |  +-----+                           
  |  |  |     +-- Beringiana_japonica[17] ** 
  +--+  |     |                           
     |  |     +-- Beringiana_japonica[18] ** 
     |  |     |                           
     |  |     +-- Beringiana_japonica[19] ** 
     |  |     |                           
     |  |     +-- Beringiana_japonica[20] ** 
     |  |                                 
     |  |  +----- Beringiana_japonica[1] **  
     |  |  |                              
     |  |  +----- Beringiana_japonica[2] **  
     |  |  |                              
     +--+  +----- Beringiana_japonica[3] **  
        |  |                              
        |  +----- Beringiana_japonica[4] **  
        |  |                              
        |  +----- Beringiana_japonica[5] **  
        |  |                              
        |  +----- Sinanodonta_lauta[1]    
        |  |                              
        |  +----- Beringiana_japonica[6] **  
        |  |                              
        |  +----- Sinanodonta_lauta[2]    
        |  |                              
        |  +----- Sinanodonta_calipygos[1]
        |  |                              
        +--+----- Beringiana_japonica[7] **  
           |                              
           +----- Sinanodonta_lauta[4]    
           |                              
           +----- Beringiana_japonica[8] **  
           |                              
           +----- Beringiana_japonica[9] **  
           |                              
           +----- Beringiana_japonica[10] ** 
           |                              
           +----- Beringiana_japonica[11] ** 
           |                              
           +----- Beringiana_japonica[12] ** 
           |                              
           +----- Beringiana_japonica[13] ** 
           |                              
           +----- Beringiana_japonica[14] ** 
           |                              
           |  +-- Sinanodonta_lauta[3]    
           +--+                           
              +-- Sinanodonta_calipygos[2]

Bogan et al., 2023 [Publ 4295], fig. 3

iT tT bi riS oT roT roS iD
4 4 2 4 27 23 6 28.0

*** Top Sister-Group Cladogram ***

                       +-- Pletholophus_honglinhensis
  +--------------------+                             
  |                    +-- Pletholophus_tenuis       
  |                                                  
  |              +-------- Beringiana_japonica **       
--+  +-----------+                                   
  |  |           |  +----- Beringiana_beringiana **     
  |  |           +--+                                
  |  |              |  +-- Beringiana_fukuharai **      
  |  |              +--+                             
  +--+                 +-- Beringiana_gosannensis **    
     |                                               
     |                 +-- Sinanodonta_angula        
     |  +--------------+                             
     |  |              +-- Sinanodonta_lucida        
     |  |                                            
     +--+  +-------------- Sinanodonta_schrenkii     
        |  |                                         
        |  |           +-- Sinanodonta_calipygos     
        +--+-----------+                             
           |           +-- Sinanodonta_tumens        
           |                                         
           |  +----------- Sinanodonta_woodiana[2]   
           +--+                                      
              |  +-------- Sinanodonta_jourdyi       
              +--+                                   
                 |  +----- Sinanodonta_elliptica     
                 +--+                                
                    |  +-- Sinanodonta_lauta         
                    +--+                             
                       +-- Sinanodonta_woodiana[1]   

Sano et al., 2020 [Publ 4138], fig. 1

iT tT bi riS oT roT roS iD
5 34 5 5 43 11 5 24.0
  +-------- Lanceolaria_grayii[1]      
  |                                    
  +-------- Lanceolaria_grayii[2]      
  |                                    
  +-------- Cristaria_plicata[1]       
  |                                    
  +-------- Cristaria_plicata[2]       
  |                                    
  +-------- Sinanodonta_lauta          
  |                                    
  +-------- Beringiana_japonica **        
  |                                    
  +-------- Beringiana_fukuharai[1] **    
  |                                    
  +-------- Beringiana_beringiana **      
  |                                    
  +-------- Beringiana_fukuharai[2] **    
  |                                    
  +-------- Beringiana_fukuharai[3] **    
  |                                    
  +-------- Buldowskia_iwakawai[1]     
  |                                    
  +-------- Buldowskia_iwakawai[2]     
  |                                    
  +-------- Sinanodonta_tumens[1]      
  |                                    
  +-------- Sinanodonta_tumens[2]      
  |                                    
--+-------- Pletholophus_tenuis        
  |                                    
  +-------- Anemina_arcaeformis        
  |                                    
  +-------- Lanceolaria_lanceolata     
  |                                    
  +-------- Cristaria_plicata[3]       
  |                                    
  +-------- Lanceolaria_gladiola       
  |                                    
  +-------- Sinanodonta_angula         
  |                                    
  +-------- Lepidodesma_languilati     
  |                                    
  |     +-- Aculamprotula_tortuosa     
  +-----+                              
  |     +-- Acuticosta_chinensis       
  |                                    
  |  +----- Cuneopsis_celtiformis      
  |  |                                 
  +--+  +-- Nodularia_douglasiae[1]    
  |  |  |                              
  |  +--+-- Nodularia_douglasiae[2]    
  |     |                              
  |     +-- Nodularia_nipponensis      
  |                                    
  |  +----- Schistodesmus_lampreyanus  
  |  |                                 
  |  |  +-- Inversiunio_reinianus[1]   
  +--+  |                              
     |  +-- Inversiunio_reinianus[2]   
     |  |                              
     |  +-- Inversiunio_jokohamensis[1]
     +--+                              
        +-- Inversiunio_jokohamensis[2]
        |                              
        +-- Inversiunio_jokohamensis[3]
        |                              
        +-- Inversiunio_jokohamensis[4]

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iT tT bi riS oT roT roS iD
4 4 2 4 118 20 6 28.0
  +-------------------- Pletholophus_tenuis    
  |                                            
  |           +-------- Beringiana_japonica **    
  |  +--------+                                
--+  |        |  +----- Beringiana_beringiana **  
  |  |        +--+                             
  |  |           |  +-- Beringiana_fukuharai **   
  |  |           +--+                          
  +--+              +-- Beringiana_gosannensis ** 
     |                                         
     |              +-- Sinanodonta_angula     
     |  +-----------+                          
     |  |           +-- Sinanodonta_lucida     
     |  |                                      
     +--+  +----------- Sinanodonta_schrenkii  
        |  |                                   
        |  |        +-- Sinanodonta_calipygos  
        |  +--------+                          
        +--+        +-- Sinanodonta_tumens     
           |                                   
           |  +-------- Sinanodonta_woodiana[2]
           |  |                                
           +--+  +----- Sinanodonta_elliptica  
              |  |                             
              +--+----- Sinanodonta_jourdyi    
                 |                             
                 |  +-- Sinanodonta_lauta      
                 +--+                          
                    +-- Sinanodonta_woodiana[1]

Lopes-Lima et al., 2020 [Publ 4080], fig. 7

iT tT bi riS oT roT roS iD
4 4 2 4 4 2 1 28.0
           +-- Sinanodonta_woodiana  
  +--------+                         
  |        +-- Sinanodonta_tumens    
--+                                  
  |  +-------- Beringiana_gosannensis **
  +--+                               
     |  +----- Beringiana_fukuharai **  
     +--+                            
        |  +-- Beringiana_beringiana ** 
        +--+                         
           +-- Beringiana_japonica **   

Sano et al., 2022 [Publ 4311], fig. 2

iT tT bi riS oT roT roS iD
4 4 2 4 10 10 4 28.0
              +-------- Pletholophus_tenuis   
  +-----------+                               
  |           |  +----- Cristaria_plicata     
  |           +--+                            
  |              |  +-- Buldowskia_iwakawai   
  |              +--+                         
  |                 +-- Buldowskia_kamiyai    
--+                                           
  |           +-------- Beringiana_fukuharai **  
  |  +--------+                               
  |  |        |  +----- Beringiana_japonica **   
  |  |        +--+                            
  |  |           |  +-- Beringiana_gosannensis **
  +--+           +--+                         
     |              +-- Beringiana_beringiana ** 
     |                                        
     |  +-------------- Sinanodonta_tumens    
     +--+                                     
        |  +----------- Sinanodonta[2]        
        +--+                                  
           |  +-------- Sinanodonta_calipygos 
           +--+                               
              |  +----- Sinanodonta_lauta     
              +--+                            
                 |  +-- Sinanodonta_woodiana  
                 +--+                         
                    +-- Sinanodonta[1]        

Sano et al., 2022 [Publ 4311], fig. 3

iT tT bi riS oT roT roS iD
4 4 2 4 14 14 4 28.0
                 +-------- Pletholophus_tenuis    
  +--------------+                                
  |              |  +----- Cristaria_plicata      
  |              +--+                             
  |                 |  +-- Buldowskia_kamiyai     
  |                 +--+                          
  |                    +-- Buldowskia_iwakawai    
--+                                               
  |              +-------- Beringiana_fukuharai **   
  |  +-----------+                                
  |  |           |  +----- Beringiana_gosannensis ** 
  |  |           +--+                             
  |  |              |  +-- Beringiana_japonica **    
  |  |              +--+                          
  +--+                 +-- Beringiana_beringiana **  
     |                                            
     |                 +-- Sinanodonta_angula     
     |  +--------------+                          
     |  |              +-- Sinanodonta_lucida     
     |  |                                         
     +--+           +----- Sinanodonta_schrenkii  
        |  +--------+                             
        |  |        |  +-- Sinanodonta_tumens     
        |  |        +--+                          
        +--+           +-- Sinanodonta_calipygos  
           |                                      
           |  +----------- Sinanodonta_woodiana[2]
           +--+                                   
              |  +-------- Sinanodonta_jourdyi    
              +--+                                
                 |  +----- Sinanodonta_elliptica  
                 +--+                             
                    |  +-- Sinanodonta_woodiana[1]
                    +--+                          
                       +-- Sinanodonta_lauta      

Bulakhova et al., 2023 [Publ 4325], fig. 5

iT tT bi riS oT roT roS iD
4 4 2 4 0 0 0 28.0
     +-- Beringiana_beringiana ** 
  +--+                         
  |  +-- Beringiana_japonica **   
--+                            
  |  +-- Beringiana_gosannensis **
  +--+                         
     +-- Beringiana_fukuharai **  

Sano et al., 2017 [Publ 3799], fig. 1

iT tT bi riS oT roT roS iD
3 3 1 3 24 5 4 28.0
  +----------- Pletholophus_tenuis  
  |                                 
--+        +-- Sinanodonta_tumens   
  |  +-----+                        
  |  |     +-- Sinanodonta_lauta    
  +--+                              
     |  +----- Beringiana_fukuharai ** 
     +--+                           
        |  +-- Beringiana_japonica **  
        +--+                        
           +-- Beringiana_beringiana **

Sano et al., 2017 [Publ 3799], fig. 2

iT tT bi riS oT roT roS iD
3 3 1 3 33 6 4 28.0
              +-- Cristaria_plicata    
  +-----------+                        
  |           +-- Buldowskia_iwakawai  
  |                                    
--+        +----- Beringiana_fukuharai ** 
  |  +-----+                           
  |  |     |  +-- Beringiana_japonica **  
  |  |     +--+                        
  +--+        +-- Beringiana_beringiana **
     |                                 
     |  +-------- Pletholophus_tenuis  
     +--+                              
        |  +----- Sinanodonta_woodiana 
        +--+                           
           |  +-- Sinanodonta_lauta    
           +--+                        
              +-- Sinanodonta_tumens   

Sano et al., 2020 [Publ 4138], fig. 2

iT tT bi riS oT roT roS iD
3 3 1 3 70 9 5 28.0
  +-------------- Pletholophus_tenuis  
  |                                    
  |        +----- Beringiana_fukuharai ** 
--+  +-----+                           
  |  |     |  +-- Beringiana_japonica **  
  |  |     +--+                        
  +--+        +-- Beringiana_beringiana **
     |                                 
     |  +-------- Sinanodonta_lucida   
     +--+                              
        |  +----- Sinanodonta_tumens   
        +--+                           
           |  +-- Sinanodonta_lauta    
           +--+                        
              +-- Sinanodonta_woodiana 

Sano et al., 2020 [Publ 4138], fig. 3

iT tT bi riS oT roT roS iD
3 3 1 3 69 9 5 28.0
     +-------- Pletholophus_tenuis  
  +--+                              
  |  |  +----- Beringiana_fukuharai ** 
  |  +--+                           
  |     |  +-- Beringiana_japonica **  
--+     +--+                        
  |        +-- Beringiana_beringiana **
  |                                 
  |  +-------- Sinanodonta_lucida   
  +--+                              
     |  +----- Sinanodonta_tumens   
     +--+                           
        |  +-- Sinanodonta_woodiana 
        +--+                        
           +-- Sinanodonta_lauta    

Sano et al., 2020 [Publ 4138], fig. 4

iT tT bi riS oT roT roS iD
3 3 1 3 69 9 5 28.0
     +-------- Sinanodonta_lucida   
  +--+                              
  |  |  +----- Sinanodonta_tumens   
  |  +--+                           
  |     |  +-- Sinanodonta_woodiana 
--+     +--+                        
  |        +-- Sinanodonta_lauta    
  |                                 
  |  +-------- Pletholophus_tenuis  
  +--+                              
     |  +----- Beringiana_fukuharai ** 
     +--+                           
        |  +-- Beringiana_japonica **  
        +--+                        
           +-- Beringiana_beringiana **

Lopes-Lima et al., 2020 [Publ 4080], fig. 10

iT tT bi riS oT roT roS iD
2 2 0 2 12 10 1 --
                 +-- Beringiana_japonica **    
  +--------------+                          
  |              +-- Beringiana_beringiana **  
  |                                         
--+              +-- Sinanodonta_angula     
  |  +-----------+                          
  |  |           +-- Sinanodonta_lucida     
  |  |                                      
  +--+  +----------- Sinanodonta_schrenkii  
     |  |                                   
     |  |        +-- Sinanodonta_calipygos  
     |  +--------+                          
     +--+        +-- Sinanodonta_tumens     
        |                                   
        |  +-------- Sinanodonta_woodiana[2]
        |  |                                
        +--+-------- Sinanodonta_jourdyi    
           |                                
           |  +----- Sinanodonta_elliptica  
           +--+                             
              |  +-- Sinanodonta_lauta      
              +--+                          
                 +-- Sinanodonta_woodiana[1]

Chong et al., 2008 [Publ 2668], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 20 1 1 --
  +-- Beringiana_beringiana **
--+                        
  +-- Sinanodonta_woodiana 

Gustafson & Iwamoto, 2005 [Publ 2741], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 4 0 0 --
-- Beringiana_beringiana

Soroka, 2010 [Publ 3404], fig. 9a

iT tT bi riS oT roT roS iD
1 1 0 1 4 4 2 --
  +----------- Buldowskia_flavotincta
--+                                  
  |  +-------- Beringiana_beringiana ** 
  +--+                               
     |  +----- Sinanodonta_schrenkii 
     +--+                            
        |  +-- Sinanodonta_lauta     
        +--+                         
           +-- Sinanodonta_woodiana  

Klishko et al., 2014 [Publ 3666], fig. 8

iT tT bi riS oT roT roS iD
1 1 0 1 3 3 2 --
     +-- Cristaria_plicata    
  +--+                        
  |  +-- Cristaria_radiata    
--+                           
  |  +-- Beringiana_beringiana **
  +--+                        
     +-- Sinanodonta_woodiana 

Klishko et al., 2016 [Publ 3733], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 3 3 2 --
     +-- Cristaria_plicata    
  +--+                        
  |  +-- Cristaria_radiata    
--+                           
  |  +-- Sinanodonta_woodiana 
  +--+                        
     +-- Beringiana_beringiana **

Sayenko et al., 2017 [Publ 3760], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 3 3 1 --
  +-------- Beringiana_beringiana **
--+                              
  |  +----- Sinanodonta_schrenkii
  +--+                           
     |  +-- Sinanodonta_woodiana 
     +--+                        
        +-- Sinanodonta_lauta    

Froufe et al., 2017 [Publ 3802], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 11 9 3 --
                 +-- Cristaria_plicata      
  +--------------+                          
  |              +-- Cristaria_radiata      
--+                                         
  |  +-------------- Beringiana_beringiana **  
  |  |                                      
  +--+  +----------- Sinanodonta_lucida     
     |  |                                   
     +--+  +-------- Sinanodonta_schrenkii  
        |  |                                
        +--+  +----- Sinanodonta_woodiana[1]
           |  |                             
           +--+  +-- Sinanodonta_lauta      
              |  |                          
              +--+-- Sinanodonta_elliptica  
                 |                          
                 +-- Sinanodonta_woodiana[2]

Inoue et al., 2018 [Publ 3858], fig. 1, 3a-c

iT tT bi riS oT roT roS iD
1 1 0 1 204 0 0 --
-- Beringiana_beringiana

Bolotov et al., 2020 [Publ 4084], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 35 8 5 --
  +----------- Cristaria_plicata    
--+                                 
  |  +-------- Beringiana_beringiana **
  +--+                              
     |  +----- Sinanodonta_schrenkii
     +--+                           
        |  +-- Sinanodonta_lauta    
        +--+                        
           +-- Sinanodonta_woodiana 

Kano et al., 2019 [Publ 4194], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 9 2 2 --
  +----- Sinanodonta_lauta  
--+                         
  |  +-- Pletholophus_tenuis
  +--+                      
     +-- Beringiana_japonica **

Konopleva et al., 2023 [Publ 4269], fig. S1, 4

iT tT bi riS oT roT roS iD
1 1 0 1 55 12 5 --
     +-------- Beringiana_beringiana **
  +--+                              
  |  |  +----- Sinanodonta_schrenkii
  |  +--+                           
  |     |  +-- Sinanodonta_woodiana 
--+     +--+                        
  |        +-- Sinanodonta_lauta    
  |                                 
  |  +-------- Cristaria_radiata    
  |  |                              
  +--+     +-- Cristaria_truncata   
     |  +--+                        
     |  |  +-- Cristaria_plicata    
     +--+                           
        |  +-- Cristaria_bellua     
        +--+                        
           +-- Cristaria_clessini   

Chen et al., 2023 [Publ 4302], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 39 5 5 --
  +----- Pletholophus_tenuis  
--+                           
  |  +-- Sinanodonta_jourdyi  
  +--+                        
     +-- Beringiana_beringiana **

Chen et al., 2023 [Publ 4302], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 39 5 5 --
  +----- Pletholophus_tenuis  
--+                           
  |  +-- Sinanodonta_jourdyi  
  +--+                        
     +-- Beringiana_beringiana **

Wu et al., 2023 [Publ 4339], fig. 4, S5-aa

iT tT bi riS oT roT roS iD
1 1 0 1 95 6 3 --
        +----- Cristaria_plicata   
  +-----+                          
  |     |  +-- Anemina_arcaeformis 
  |     +--+                       
--+        +-- Anemina_euscaphys   
  |                                
  |  +-------- Beringiana_fukuharai **
  +--+                             
     |  +----- Sinanodonta_lucida  
     +--+                          
        |  +-- Sinanodonta_tumens  
        +--+                       
           +-- Sinanodonta_woodiana

Wu et al., 2023 [Publ 4339], fig. 4, S5-nt

iT tT bi riS oT roT roS iD
1 1 0 1 96 6 3 --
           +-- Anemina_arcaeformis 
  +--------+                       
  |        +-- Anemina_euscaphys   
--+                                
  |  +-------- Beringiana_fukuharai **
  +--+                             
     |  +----- Sinanodonta_lucida  
     +--+                          
        |  +-- Sinanodonta_tumens  
        +--+                       
           +-- Sinanodonta_woodiana

Wu et al., 2024 [Publ 4350], fig. 8a, S1

iT tT bi riS oT roT roS iD
1 1 0 1 91 6 3 --
           +-- Anemina_arcaeformis 
  +--------+                       
  |        +-- Anemina_euscaphys   
--+                                
  |  +-------- Beringiana_fukuharai **
  +--+                             
     |  +----- Sinanodonta_lucida  
     +--+                          
        |  +-- Sinanodonta_tumens  
        +--+                       
           +-- Sinanodonta_woodiana

Wu et al., 2024 [Publ 4350], fig. 8b, S2

iT tT bi riS oT roT roS iD
1 1 0 1 91 6 3 --
           +-- Anemina_arcaeformis 
  +--------+                       
  |        +-- Anemina_euscaphys   
--+                                
  |  +-------- Beringiana_fukuharai **
  +--+                             
     |  +----- Sinanodonta_lucida  
     +--+                          
        |  +-- Sinanodonta_tumens  
        +--+                       
           +-- Sinanodonta_woodiana

Page updated: 11:02:59 Thu 21 Mar 2024

 
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