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Page last updated
Wed 06 Nov 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

The nominal species (available and otherwise) in this synonymy are listed by usage rather than strictly by publication date. The primary sorting order is by the most recent year (since 1969) that the species name was treated as valid, and then nominal species are sorted secondarily by publication year.

The taxonomic history of the valid species (since 1900) is listed at the bottom of the page.

Search the database pages:

Unionoida | Unionoidea | Unionidae | Unioninae | Unionini | Nodularia

species Nodularia douglasiae (Griffith & Pidgeon, 1833): nominal species
     273 specimens available.

Unio douglasiae Griffith & Pidgeon, 1833
Griffith & Pidgeon, 1833, [in] Cuvier, Animal Kingdom 12(38): pl. 21, fig. 2 [as ‘douglasiæ’].
‘Gray’ Griffith & Pidgeon, 1834, [in] Cuvier, Animal Kingdom 12(39-40): 601 [as ‘douglasiæ’].

Unio douglasiaeGriffith & Pidgeon (1833), Sowerby (1866), Heude (1885), Neumayr (1899), ..., Wang et al. (2018), Aunins et al. (2018), Wang et al. (2018), Araujo et al. (2018)
Margarita (Unio) merchisonianusLea (1836)
Margarita (Unio) murchisonianusLea (1838)
Unio murchisonianusHanley (1843)
Margaron (Unio) murchisonianusLea (1852), Lea (1870)
Nodularia douglasiæConrad (1853)
Unio osbeckiSowerby (1868)
Nodularia (s.s.) douglasiaeSimpson (1900), Preston (1912), Simpson (1914), Haas (1924), Korniushin (1998)
Unio (Nodularia) douglasiaeMartens (1905), Zhadin (1938)
Nodularia (s.s.) douglasiae douglasiaeHaas (1911)
Nodularia douglasiaeHaas (1920), Prozorova et al. (2005), Graf & Cummings (2007), Graf (2007), ..., Wu et al. (2024), Dai et al. (2024), Wu et al. (2024), Wu et al. (2024)
Unio douglasiae douglasiaeHaas (1969)
Nodularidia douglasiaeHe & Zhuang (2013)
Nodularia cf. douglasiaeHuang et al. (2019)
Nodularia douglasiae Clade CChoi et al. (2020)
Nodularia douglasiae Clade BChoi et al. (2020)
Nodularia douglasiae Clade AChoi et al. (2020)
Nodularia douglasiae douglasiaeBogatov (2022)
Nodularia sp. 2Dai et al. (2024)

Unio pictorum var. amurensis Mousson, 1887
Mousson, 1887, J. de Conch. 35: 26, pl. 1, fig. 8.
(intro'd as intra-sp. of Mya pictorum Linnaeus, 1758)

Unio pictorum var. amurensisMousson (1887)
Unio (s.s.) pictorumPaetel (1890)
Nodularia (s.s.) douglasiae var. shanghaiensisSimpson (1900), Simpson (1914)
Nodularia (s.s.) douglasiae douglasiaeHaas (1911)
Unio douglasiae amurensisHaas (1969)
Nodularia amurensisMoskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Nodularia (s.s.) amurensisZatravkin & Bogatov (1987), Starobogatov et al. (2004), Bogatov (2012)
Nodularia douglasiaeGraf (2007), Klishko et al. (2018), Lopes-Lima et al. (2020), Bolotov et al. (2020), Sayenko et al. (2021)
Nodularidia douglasiaeHe & Zhuang (2013)
Nodularia douglasiae amurensisBogatov (2022)

Unio middendorffi Westerlund, 1890
Middendorff, 1851, Reise Nord. Ost. Sibiriens 2(1): 276-277, pl. 28, figs. 1-3 [as Unio pictorum var. longirostris].
Westerlund, 1890, Fauna Paläarct. Binnenconch. 2(7): 113.
Type(s): NEOTYPE BIV229_5, Gladkaya River, Primorye Region, Russia.

Unio middendorffiWesterlund (1890)
Unio pictorum middendorffiHaas (1940)
Unio douglasiae amurensisHaas (1969)
Nodularia middendorffiMoskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Nodularia (s.s.) middendorffiZatravkin & Bogatov (1987), Starobogatov et al. (2004), Bogatov (2012)
Nodularia douglasiaeGraf (2007), Klishko et al. (2018), Lopes-Lima et al. (2020), Bolotov et al. (2020)
Nodularidia douglasiaeHe & Zhuang (2013)
Nodularia douglasiae middendorffiBogatov (2022)

Unio schrencki Westerlund, 1897
Westerlund, 1897, Ann. Mus. St. Pétersbourg 1897: 131.
Type(s): PARATYPE SMF_3663, Amur - Fluss.

Unio schrenckiWesterlund (1897)
Nodularia (s.s.) douglasiaeSimpson (1900), Simpson (1914)
Unio (Nodularia) douglasiae var. schrenkiZhadin (1938)
Unio douglasiae var. schrenckiZhadin (1965)
Unio douglasiae amurensisHaas (1969)
Nodularia schrenckiMoskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010), Bogatov & Prozorova (2017)
Nodularia (s.s.) schrenckiZatravkin & Bogatov (1987), Starobogatov et al. (2004), Bogatov (2012)
Nodularia douglasiaeGraf (2007), Klishko et al. (2018), Lopes-Lima et al. (2020), Bolotov et al. (2020), Sayenko et al. (2021)
Nodularidia douglasiaeHe & Zhuang (2013)
Nodularia douglasiae schrenckiBogatov (2022)

Middendorffinaia (Middendorffinaia) ochotica Bogatov, 2000
Bogatov, 2000, Zool. Zhur. 79: 861, text fig. a-r.

Middendorffinaia (Middendorffinaia) ochoticaBogatov (2000)
Middendorffinaia (s.s.) ochoticaStarobogatov et al. (2004), Bogatov (2012), Vinarski & Kantor (2016), Bogatov & Prozorova (2017)
Middendorffinaia ochoticaKantor & Sysoev (2005), Kantor et al. (2010)
Unio crassus mongolicusGraf (2007)
Middendorffinaia mongolicaKlishko et al. (2019)
Nodularia douglasiaeLopes-Lima et al. (2020), Bolotov et al. (2020)
Middendorffinaia mongolica ochoticaBogatov (2022)

Unio jourdyi Morlet, 1886
Morlet, 1886, J. de Conch. 34: 76, pl. 13, figs. 5-5a.
Type(s): SYNTYPES MNHN_MP_0150, Bac Hat etang de la riviere Claire.

Unio jourdyiMorlet (1886), Mabille (1887), Fischer (1891), Dautzenberg & Fischer (1905), Dautzenberg & Fischer (1905)
Nodularia (s.s.) jourdyiSimpson (1900), Simpson (1914)
Oxynaia jourdyiHaas (1913), Thiele (1934), Haas (1969), Haas (1969), Dang et al. (1980), Graf & Cummings (2007), Bogan (2010), IUCN (2011), Do et al. (2018), Zieritz et al. (2018)
Nodularia jourdyiBolotov et al. (2018), Pfeiffer et al. (2019), Graf & Cummings (2021), Sayenko et al. (2021)
Nodularia douglasiaeWu et al. (2024)

Nodularia denserugata Haas, 1910
Haas, 1910, Ann. Mag. Nat. Hist. 6: 496.
Haas, 1910, Conch. Cab. 9 (pt. 2, h. 42): pl. 8, figs. 3-4.
Haas, 1911, Conch. Cab. 9 (pt. 2, h. 44): 83, pl. 8, figs. 3-4.

Nodularia denserugataHaas (1910), Graf & Cummings (2007), Bogan (2010)
Nodularia (s.s.) denserugataHaas (1911), Simpson (1914)
Unio denserugatusHaas (1969)
Middendorffinaia (Sujfununio) denserugataMoskvicheva & Starobogatov (1973)
Ptychorhynchus denserugataHe & Zhuang (2013), Zieritz et al. (2018)
Ptychorhynchus denserugatumGraf & Cummings (2021)
Nodularia douglasiaeWu et al. (2024)

Unio biwae Kobelt, 1879
Kobelt, 1879, Abh. Senckenb. Naturf. Ges. 11: 425, pl. 23, figs. 2-4.
Type(s): LECTOTYPE SMF_3650, Biwa-See, Japan.

Unio biwaeKobelt (1879), Paetel (1890), Ihering (1893), Haas (1969), Habe (1977), Habe (1991), Slugina (2006)
Nodularia (s.s.) biwaeSimpson (1900), Haas (1911), Simpson (1914)
Unio douglasiae biwaeKondo (1997), Kondo (2008)
Nodularia biwaeBogan (2010)
Nodularidia douglasiaeHe & Zhuang (2013)
Nodularia douglasiae biwaeSano et al. (2017), Mabuchi & Nishida (2020)
Nodularia douglasiaeLopes-Lima et al. (2020), Mabuchi et al. (2021)

Acuticosta jianghanensis He & Zhuang, 2013
He & Zhuang, 2013, Freshwater Bivalves China: 23, fig. 47-48.

Acuticosta jianghanensisHe & Zhuang (2013), Zieritz et al. (2018), Liu et al. (2020)
Nodularia douglasiaeBogatov & Prozorova (2017)

Nodularia douglasiae crassidens Haas, 1910
Haas, 1910, Ann. Mag. Nat. Hist. 6: 499.
Haas, 1910, Conch. Cab. 9 (pt. 2, h. 42): pl. 7, figs. 4-5.
Haas, 1911, Conch. Cab. 9 (pt. 2, h. 44): 75.
(intro'd as intra-sp. of Unio douglasiae Griffith & Pidgeon, 1833)
Type(s): TYPE SMF_3659, Hunan, M. China.

Nodularia douglasiae crassidensHaas (1910), Dang et al. (1980), Do et al. (2018)
Nodularia (s.s.) douglasiae crassidensHaas (1911)
Nodularia (s.s.) douglasiae var. crassidensSimpson (1914)
Unio douglasiae douglasiaeHaas (1969)
Nodularia crassidensProzorova et al. (2005)
Nodularia dorriIUCN (2012)
Nodularidia douglasiaeHe & Zhuang (2013)

Nodularia sujfunica Moskvicheva, 1973
Moskvicheva, 1973, Zool. Zhur. 52(10): 1463, figs. 1 с-т.

Nodularia sujfunicaMoskvicheva (1973), Zatravkin (1983)
Nodularia (s.s.) sujfunicaZatravkin & Bogatov (1987)
Nodularia (s.s.) vladivostokensisStarobogatov et al. (2004)
Nodularia vladivostokensisKantor & Sysoev (2005)
Nodularidia douglasiaeHe & Zhuang (2013)
Nodularia suifunicaKlishko et al. (2018)
Nodularia douglasiaeLopes-Lima et al. (2020), Bolotov et al. (2020)
Nodularia douglasiae schrenckiBogatov (2022)

Nodularia vladivostokensis Moskvicheva, 1973
Moskvicheva, 1973, Zool. Zhur. 52(10): 1463, figs. 1 п-р.

Nodularia vladivostokensisMoskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010), Klishko et al. (2018)
Nodularia (s.s.) vladivostokensisZatravkin & Bogatov (1987), Starobogatov et al. (2004)
Nodularia douglasiaeGraf (2007), Lopes-Lima et al. (2020), Bolotov et al. (2020), Sayenko et al. (2021)
Nodularidia douglasiaeHe & Zhuang (2013)
Nodularia douglasiae douglasiaeBogatov (2022)

Nodularia lebedevi Zatravkin & Starobogatov, 1984
Zatravkin & Starobogatov, 1984, Zool. Zhur. 63(12): 1787, figs. 5-8.

Nodularia lebedeviZatravkin & Starobogatov (1984), Kantor & Sysoev (2005), Kantor et al. (2010)
Nodularia sp.n.Zatravkin (1983)
Nodularia (Amurunio) lebedeviZatravkin & Bogatov (1987), Korniushin (1998), Starobogatov et al. (2004), Klishko et al. (2018)
Nodularia douglasiaeGraf (2007), Lopes-Lima et al. (2020), Bolotov et al. (2020)
Nodularia (s.s.) middendorffiBogatov (2012)
Nodularidia douglasiaeHe & Zhuang (2013)
Nodularia middendorffiVinarski & Kantor (2016)
Nodularia douglasiae middendorffiBogatov (2022)

Nodularia sakhalinensis Bogatov, 2001
Bogatov, 2001, Byull. Dal'nev. Malak.Obshch. 5: 75.

Nodularia sakhalinensisBogatov (2001), Kantor & Sysoev (2005), Kantor et al. (2010), Klishko et al. (2018)
Nodularia (s.s.) sakhalinensisStarobogatov et al. (2004)
Nodularia douglasiaeGraf (2007), Lopes-Lima et al. (2020), Bolotov et al. (2020)
Nodularidia douglasiaeHe & Zhuang (2013)
Nodularia douglasiae schrenckiBogatov (2022)

Scabies longata Liu et al., 1991
Liu et al., 1991, Acta Zootax. Sinica 16(3): 279 [281], fig’d.

Scabies longataLiu et al. (1991), Hu (2005), Graf & Cummings (2007), Bogan (2010), WoRMS (2017)
Scabies crispataHe & Zhuang (2013), Graf & Cummings (2021)
Nodularia douglasiaeDai et al. (2024)

Nodularia (Magadaninaia) extremalis Martynov & Chernyshev, 1992
Martynov & Chernyshev, 1992, Zool. Zhur. 71: 19, figs. 1-3.

Nodularia (Magadaninaia) extremalisMartynov & Chernyshev (1992), Starobogatov et al. (2004)
Nodularia extremalisKantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Nodularia douglasiaeGraf (2007), Klishko et al. (2018), Lopes-Lima et al. (2020), Bolotov et al. (2020)
Nodularidia douglasiaeHe & Zhuang (2013)
Nodularia douglasiae schrenckiBogatov (2022)

Nodularia (s.s.) moskvichevae Bogatov & Starobogatov, 1992
Bogatov & Starobogatov, 1992, Zool. Zhur. 71: 134, fig’d.

Nodularia (s.s.) moskvichevaeBogatov & Starobogatov (1992), Starobogatov et al. (2004)
Nodularia moskvichevaeKantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Nodularia douglasiaeGraf (2007), Klishko et al. (2018), Lopes-Lima et al. (2020), Bolotov et al. (2020)
Nodularia douglasiae middendorffiBogatov (2022)

Unio abbreviatus Westerlund, 1897
Westerlund, 1897, Ann. Mus. St. Pétersbourg 1897: 132.
(homonym of Unio requienii var. abbreviatus de Joannis, 1859)
Type(s): PARATYPE SMF_3662, Ussuri fl. bei Noor, Amur gebiet.

Unio abbreviatusWesterlund (1897)
Nodularia (s.s.) douglasiaeSimpson (1900), Preston (1912), Simpson (1914)
Nodularia (s.s.) douglasiae douglasiaeHaas (1911)
Unio (Nodularia) douglasiae var. schrenkiZhadin (1938)
Unio douglasiae var. abbreviatusZhadin (1965)
Unio douglasiae amurensisHaas (1969)
Nodularia abbreviataMoskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010)
Nodularia (s.s.) abbreviataZatravkin & Bogatov (1987), Starobogatov et al. (2004)
Nodularia douglasiaeGraf (2007), Lopes-Lima et al. (2020), Bolotov et al. (2020)
Nodularia (s.s.) schrenckiBogatov (2012)
Nodularidia douglasiaeHe & Zhuang (2013)
Nodularia douglasiae schrenckiBogatov (2022)

Nodularia douglasiae flavoviridis Haas, 1910
Haas, 1910, Conch. Cab. 9 (pt. 2, h. 41): 71, pl. 6, fig. 2.
(intro'd as intra-sp. of Unio douglasiae Griffith & Pidgeon, 1833)
Type(s): TYPE SMF_3654, W. Ufer d. Sees Buirnor, O. Mongolei, Amur-Geb.

Nodularia douglasiae flavoviridisHaas (1910)
Unio douglasiae amurensisHaas (1969)
Nodularia flavoviridisMoskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010)
Nodularia (s.s.) flavoviridisZatravkin & Bogatov (1987), Starobogatov et al. (2004)
Nodularia douglasiaeGraf (2007), Klishko et al. (2018), Lopes-Lima et al. (2020), Bolotov et al. (2020)
Nodularia (s.s.) schrenckiBogatov (2012)
Nodularidia douglasiaeHe & Zhuang (2013)

Middendorffinaia (s.s.) arsenievi Moskvicheva & Starobogatov, 1973
Moskvicheva & Starobogatov, 1973, Byull. Mosk. Obshch.. Ispyt. Prir. 78(2): 29, figs. 3м-п.
(correct name for Middendorffinaia arsenjevi Moskvicheva & Starobogatov, 1973)

Middendorffinaia (s.s.) arsenieviMoskvicheva & Starobogatov (1973), Zatravkin & Bogatov (1987), Starobogatov et al. (2004)
Middendorffinaia (s.s.) arsenjeviZatravkin (1983)
Middendorffinaia arsenieviKantor & Sysoev (2005), Kantor et al. (2010)
Unio crassus mongolicusGraf (2007)
Middendorffinaia (s.s.) ussuriensisBogatov (2012), Vinarski & Kantor (2016)
Middendorffinaia mongolicaKlishko et al. (2019)
Nodularia douglasiaeLopes-Lima et al. (2020), Bolotov et al. (2020)
Middendorffinaia mongolica ussuriensisBogatov (2022)

Unio sculptus Deshayes, 1873
Deshayes, 1873, Nouvelles archives 9: 9, pl. 1, figs. 3-3a.
Type(s): SYNTYPES MNHN-IM-2000-1779, Petchily, Chine Centrale.

Unio sculptusDeshayes (1873), Heude (1875), Gredler (1881), Neumayr (1899), Yu et al. (2001)
Unio sculptus, var.Heude (1877)
Nodularia (s.s.) douglasiaeSimpson (1900), Preston (1912), Simpson (1914)
Nodularia (s.s.) douglasiae douglasiaeHaas (1911)
Unio douglasiae douglasiaeHaas (1969)
Unio douglasiaeLiu et al. (1979)
Nodularia douglasiaeProzorova et al. (2005), Schneider et al. (2013), Lopes-Lima et al. (2020)
Nodularidia douglasiaeHe & Zhuang (2013)

Unio pliculosus Martens, 1894
Martens, 1894, S.-Ber. Ges. Naturf. Freunde zu Berlin 1894: 216.
Martens, 1905, Zool. Jb., Jena, Suppl. 8: 60, pl. 3, figs. 3-3b.
Type(s): TYPE ZMB_55620, Singei, Korea.

Unio pliculosusMartens (1894)
Unio (Nodularia) pliculosusMartens (1905)
Nodularia (s.s.) pliculosusHaas (1911)
Nodularia (s.s.) pliculosaSimpson (1914)
Unio douglasiae pliculosusHaas (1969)
Unio douglasiaeHabe (1977)
Nodularidia douglasiaeHe & Zhuang (2013)
Nodularia douglasiaeLopes-Lima et al. (2020)

Unio verrucifer Martens, 1894
Martens, 1894, S.-Ber. Ges. Naturf. Freunde zu Berlin 1894: 216.
Martens, 1905, Zool. Jb., Jena, Suppl. 8: 61, pl. 3, figs. 2-2b.
Type(s): TYPE ZMB_55622, Hangang.

Unio verruciferMartens (1894)
Unio (Nodularia) verruciferMartens (1905)
Nodularia (s.s.) verruciferaHaas (1911), Simpson (1914)
Unio douglasiae verruciferHaas (1969)
Nodularidia douglasiaeHe & Zhuang (2013)
Nodularia douglasiaeLopes-Lima et al. (2020)

Nodularia douglasiae taiwanica Pilsbry & Hirase, 1905
Pilsbry & Hirase, 1905, Proc. Acad. Nat. Sci. 57: 750.
(intro'd as intra-sp. of Unio douglasiae Griffith & Pidgeon, 1833)
Type(s): HOLOTYPE ANSP_58170, Formosa.

Nodularia douglasiae taiwanicaPilsbry & Hirase (1905)
Nodularia (s.s.) douglasiae var. taiwanicaSimpson (1914)
Nodularia (s.s.) douglasiae taiwanicaHaas (1924)
Unio douglasiae taiwanicusKuroda (1941)
Unio douglasiae taiwanicaHaas (1969)
Unio douglasiaeHabe (1977), Liu et al. (1979), Wu (1982)
Nodularidia douglasiaeHe & Zhuang (2013)

Unio murchisonianus Lea, 1834
Lea, 1834, Trans. Amer. Phil. Soc. 5 [O. 1]: 33, pl. 3, fig. 6.
Type(s): HOLOTYPE (MONOTYPY) USNM_84156, Calcutta, India [penciled out].

Unio murchisonianusLea (1834), Hanley (1843), Küster (1861), Ihering (1893)
Margarita (Unio) merchisonianusLea (1836)
Margarita (Unio) murchisonianusLea (1838)
Margaron (Unio) murchisonianusLea (1852), Lea (1870)
Unio (Diplodon) murchisonianusAdams & Adams (1857)
Unio douglasiaeSowerby (1866)
Nodularia (s.s.) douglasiaeSimpson (1900), Preston (1912), Simpson (1914)
Unio (Nodularia) douglasiaeMartens (1905)
Nodularia (s.s.) douglasiae douglasiaeHaas (1911)
Unio douglasiae douglasiaeHaas (1969)
Nodularidia douglasiaeHe & Zhuang (2013)
Nodularia douglasiaeLopes-Lima et al. (2020)

Margarita (Unio) merchisonianus Lea, 1836
Lea, 1836, Synopsis Fam. Naiades: 14 [corrected in the errata].
(error for Unio murchisonianus Lea, 1834)

Margarita (Unio) merchisonianusLea (1836)

Unio osbeckii Philippi, 1844
Philippi, 1844, Z. Malak. 1: 164.
Philippi, 1847, Abbild. Beschr. Conch. 3: 45, pl. 3, fig. 1.

Unio osbeckiiPhilippi (1844), Philippi (1847), Küster (1861)
Margaron (Unio) osbeckiiLea (1852), Lea (1870)
Unio (Diplodon) osbeckiiAdams & Adams (1857)
Unio osbeckiSowerby (1868)
Unio douglasiae var. osbeckiiHeude (1885)
Nodularia (s.s.) douglasiaeSimpson (1900), Preston (1912), Simpson (1914)
Unio (Nodularia) douglasiaeMartens (1905)
Nodularia (s.s.) douglasiae douglasiaeHaas (1911)
Unio douglasiae douglasiaeHaas (1969)
Unio douglasiaeLiu et al. (1979)
Nodularia douglasiaeProzorova et al. (2005), Lopes-Lima et al. (2020)
Nodularidia douglasiaeHe & Zhuang (2013)

Unio pictorum var. longirostris Middendorff, 1851
Middendorff, 1851, Reise Nord. Ost. Sibiriens 2(1): 276-277, pl. 28, figs. 1-3.
Schrenck, 1867, Reise Forsch. Amurlande 2: 696.
(misidentified reference to Unio longirostris Rossmässler, 1836)

Unio pictorum var. longirostrisMiddendorff (1851)
Unio middendorffiWesterlund (1890)
Nodularia middendorffiMoskvicheva (1973), Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Nodularia (s.s.) middendorffiZatravkin & Bogatov (1987), Bogatov (2012)
Nodularidia douglasiaeHe & Zhuang (2013)
Nodularia douglasiaeLopes-Lima et al. (2020)

Unio shanghaiensis Lea, 1859
Lea, 1859, Proc. Acad. Nat. Sci. 11: 153.
Lea, 1860, Jour. Acad. Nat. Sci. 4 [O. 7]: 242, pl. 36, fig. 121.
Type(s): HOLOTYPE BMNH_1965171, Shanghai, China.

Unio shanghaiensisLea (1859), Reeve (1865), Neumayr (1899)
Margaron (Unio) osbeckiiLea (1870)
Nodularia (s.s.) douglasiae var. shanghaiensisSimpson (1900), Preston (1912), Simpson (1914)
Unio (Nodularia) douglasiaeMartens (1905)
Nodularia (s.s.) douglasiae shanghaiensisHaas (1911)
Nodularia douglasiaeHaas (1920), Lopes-Lima et al. (2020)
Unio douglasiae douglasiaeHaas (1969)
Unio douglasiaeLiu et al. (1979)
Nodularidia douglasiaeHe & Zhuang (2013)

Unio murchinsonianus Küster, 1862
‘Lea’ Küster, 1862, Conchyl. Cab. 9(2): 309.
(error for Unio murchisonianus Lea, 1834)

Unio murchinsonianusKüster (1862)

Unio wrightii Lea, 1865
Lea, 1865, Proc. Acad. Nat. Sci. 17: 75.
Lea, 1868, Jour. Acad. Nat. Sci. 6 [O. 12]: 283, pl. 39, fig. 97.
Type(s): HOLOTYPE USNM_84129, China.

Unio wrightiiLea (1865)
Margaron (Unio) wrightiiLea (1870)
Nodularia (s.s.) douglasiaeSimpson (1900), Simpson (1914)
Unio (Nodularia) douglasiaeMartens (1905)
Nodularia (s.s.) douglasiae douglasiaeHaas (1911)
Unio wrigtiide Gregorio (1914)
Unio douglasiae douglasiaeHaas (1969)
Nodularidia douglasiaeHe & Zhuang (2013)
Nodularia douglasiaeLopes-Lima et al. (2020)

Unio pictorum Schrenck, 1867
‘L.’ Schrenck, 1867, Reise Forsch. Amurlande 2: 696-699.
(misidentified reference to Mya pictorum Linnaeus, 1758)

Unio pictorumSchrenck (1867)
Margaron (Unio) pictorumLea (1870)
Nodularia middendorffiMoskvicheva (1973)

Unio pictorum var. longirostris Westerlund, 1877
Westerlund, 1877, Kongl. Svenska Vet. Akad. Handl. 14(12): 74.
(misidentified reference to Unio longirostris Rossmässler, 1836)

Unio pictorum var. longirostrisWesterlund (1877)
Nodularia (s.s.) douglasiaeSimpson (1900), Preston (1912), Simpson (1914)
Nodularia (s.s.) douglasiae douglasiaeHaas (1911)
Unio douglasiae amurensisHaas (1969)
Nodularia amurensisVinarski & Kantor (2016)

Unio nipponensis Kobelt, 1879
Kobelt, 1879, Abh. Senckenb. Naturf. Ges. 11: 422, pl. 12, fig. 3.
(misidentified reference to Unio nipponensis Martens, 1877)

Unio nipponensisKobelt (1879)
Unio murchisonianusIhering (1893)
Nodularia (s.s.) douglasiae var. nipponensisSimpson (1900)
Unio (Nodularia) nipponensisMartens (1905)

Unio douglasiae chloreus Heude, 1885
Heude, 1885, Conch. Fluv. Nanking (9): pl. 65 bis, fig. 128c.
(intro'd as intra-sp. of Unio douglasiae Griffith & Pidgeon, 1833)

Unio douglasiae chloreusHeude (1885)
Unio douglasiae var. chloreusHeude (1885)
Nodularia (s.s.) douglasiaeSimpson (1900)
Unio douglasiaeLiu et al. (1979)
Nodularia douglasiaeProzorova et al. (2005), Lopes-Lima et al. (2020)
Nodularidia douglasiaeHe & Zhuang (2013)

Unio douglasiae dactylinus Heude, 1885
Heude, 1885, Conch. Fluv. Nanking (9): pl. 65, fig. 128e.
(intro'd as intra-sp. of Unio douglasiae Griffith & Pidgeon, 1833)

Unio douglasiae dactylinusHeude (1885)
Unio douglasiae var. dactylinusHeude (1885)
Nodularia (s.s.) douglasiaeSimpson (1900), Preston (1912), Simpson (1914)
Nodularia (s.s.) douglasiae douglasiaeHaas (1911)
Unio douglasiae douglasiaeHaas (1969)
Unio douglasiaeLiu et al. (1979)
Nodularia douglasiaeProzorova et al. (2005), Lopes-Lima et al. (2020)
Nodularidia douglasiaeHe & Zhuang (2013)

Unio douglasiae deltae Heude, 1885
Heude, 1885, Conch. Fluv. Nanking (9): pl. 65 bis, fig. 128d [as ‘deltæ’].
(intro'd as intra-sp. of Unio douglasiae Griffith & Pidgeon, 1833; homonym of Unio favidens var. deltae Benson, 1862)

Unio douglasiae deltaeHeude (1885)
Unio douglasiae var. deltæHeude (1885)
Unio douglasiaeLiu et al. (1979)
Nodularia douglasiaeProzorova et al. (2005), Lopes-Lima et al. (2020)
Nodularidia douglasiaeHe & Zhuang (2013)

Unio douglasiae firmus Heude, 1885
Heude, 1885, Conch. Fluv. Nanking (9): pl. 65 bis, figs. 128b-b’.
(intro'd as intra-sp. of Unio douglasiae Griffith & Pidgeon, 1833; homonym of Unio firmus Lea, 1866)

Unio douglasiae firmusHeude (1885)
Unio douglasiae var. firmusHeude (1885)
Nodularia (s.s.) douglasiaeSimpson (1900)
Unio douglasiaeLiu et al. (1979)
Nodularia douglasiaeProzorova et al. (2005), Lopes-Lima et al. (2020)
Nodularidia douglasiaeHe & Zhuang (2013)

Unio douglasiae hainanensis Heude, 1885
Heude, 1885, Conch. Fluv. Nanking (9): pl. 65, fig. 128f.
(intro'd as intra-sp. of Unio douglasiae Griffith & Pidgeon, 1833)

Unio douglasiae hainanensisHeude (1885)
Unio douglasiae var. hainanensisHeude (1885)
Unio douglasiaeLiu et al. (1979)
Nodularia douglasiaeProzorova et al. (2005), Lopes-Lima et al. (2020)
Nodularidia douglasiaeHe & Zhuang (2013)

Unio douglasiae var. osbeckii Heude, 1885
Heude, 1885, Conch. Fluv. Nanking (9): pl. 65, fig. 128i.
(reference to Unio osbeckii Philippi, 1844)

Unio douglasiae var. osbeckiiHeude (1885)
Unio (Nodularia) douglasiae var. sinuolatusMartens (1905)

Unio boettgeri Neumayr, 1899
Neumayr, 1899, Wiss. Erg. Reise Szechény Ost-Asien 2: 645, pl. 2, fig. 3 [as ‘böttgeri’].
(homonym of Unio firmus var. boettgeri Ihering, 1893)

Unio boettgeriNeumayr (1899)
Unio böttgeriNeumayr (1899)
Nodularia (s.s.) boettgeriSimpson (1914)
Nodularia douglasiaeHaas (1920)
Nodularia (s.s.) douglasiaeHaas (1924)
Nodularidia douglasiaeHe & Zhuang (2013)

Unio kobelti Neumayr, 1899
Neumayr, 1899, Wiss. Erg. Reise Szechény Ost-Asien 2: 645, pl. 3, fig. 1.
(homonym of Unio kobelti Kobelt & Rolle, 1895)

Unio kobeltiNeumayr (1899), Simpson (1914)
Nodularia douglasiaeHaas (1920)
Nodularia (s.s.) douglasiaeHaas (1924)
Unio douglasiaeLiu et al. (1979)
Nodularidia douglasiaeHe & Zhuang (2013)

Unio schanghaiensis Neumayr, 1899
‘Lea’ Neumayr, 1899, Wiss. Erg. Reise Szechény Ost-Asien 2: 645.
(error for Unio shanghaiensis Lea, 1859)

Unio schanghaiensisNeumayr (1899)

Unio jourdyi var. corrugata Dautzenberg & Fischer, 1905
Dautzenberg & Fischer, 1905, J. de Conch. 53: 205.
(homonym of Unio corrugata Mauduyt, 1839; intro'd as intra-sp. of Unio jourdyi Morlet, 1886)
Type(s): SYNTYPE MNHN_MP_0149, Bas-Luc Nam.

Unio jourdyi var. corrugataDautzenberg & Fischer (1905)
Nodularia (s.s.) jourdyi var. corrugataSimpson (1914)

Unio jourdyi var. ponderosa Dautzenberg & Fischer, 1905
Dautzenberg & Fischer, 1905, J. de Conch. 53: 453.
(homonym of Mya ponderosa Dillwyn, 1817; intro'd as intra-sp. of Unio jourdyi Morlet, 1886)
Type(s): SYNTYPE MNHN_MP_0148, Tonkin, Grand Lac, Hanoi.

Unio jourdyi var. ponderosaDautzenberg & Fischer (1905)
Nodularia (s.s.) jourdyi var. ponderosaSimpson (1914)

Nodularia verrucosa Haas, 1910
Haas, 1910, Nachr. Bl. Dtsch. Malak. Ges. 42: 99.
Type(s): TYPE SMF_3660, Hunan, M. China.

Nodularia verrucosaHaas (1910), Simpson (1914)
Nodularidia douglasiaeHe & Zhuang (2013)

Unio wrigtii de Gregorio, 1914
‘Lea’ de Gregorio, 1914, Il Nat. Siciliano 2: 40.
(error for Unio wrightii Lea, 1865)

Unio wrigtiide Gregorio (1914)

Unio (Nodularia) douglasiae var. schrenki Zhadin, 1938
‘Westerlund’ Zhadin, 1938, Faune USSR Mollusques 4(1): 99.
(error for Unio schrencki Westerlund, 1897)

Unio (Nodularia) douglasiae var. schrenkiZhadin (1938)

Middendorffinaia arsenjevi Moskvicheva & Starobogatov, 1973
Moskvicheva & Starobogatov, 1973, Byull. Mosk. Obshch.. Ispyt. Prir. 78(2): 22, fig. 1.
‘Moskv. et Star.’ Zatravkin, 1983, [in] I.M. Likharev, Molluscs: 41.
(incorrect original spelling of Middendorffinaia (s.s.) arsenievi Moskvicheva & Starobogatov, 1973)

Middendorffinaia arsenjeviMoskvicheva & Starobogatov (1973)

Unio taiwanensis Habe, 1977
‘Pilsbry’ Habe, 1977, Mollusc. Japan: 114.
(error for Nodularia douglasiae taiwanica Pilsbry & Hirase, 1905)

Unio taiwanensisHabe (1977)

Unio firnus Liu et al., 1979
Liu et al., 1979, Econ. Fauna China: 74.
(error for Unio douglasiae firmus Heude, 1885)

Unio firnusLiu et al. (1979)

Unio douglasiae chlorens Liu et al., 1979
Liu et al., 1979, Econ. Fauna China: 74.
(error for Unio douglasiae chloreus Heude, 1885)

Unio douglasiae chlorensLiu et al. (1979)

Unio douglasiae cleltae Liu et al., 1979
Liu et al., 1979, Econ. Fauna China: 74.
(error for Unio douglasiae deltae Heude, 1885)

Unio douglasiae cleltaeLiu et al. (1979)

Nodularia doctylina Liu et al., 1980
Liu et al., 1980, Acta Zool. Sinica 26(4): 366.
(error for Unio douglasiae dactylinus Heude, 1885)

Nodularia doctylinaLiu et al. (1980)

Nodularia sachalinensis Bogatov, 2001
Bogatov, 2001, Byull. Dal'nev. Malak.Obshch. 5: 59.
(error for Nodularia sakhalinensis Bogatov, 2001)

Nodularia sachalinensisBogatov (2001)

Unio douglassiae Kantor & Sysoev, 2005
‘Griffith et Pidgeon’ Kantor & Sysoev, 2005, Cat. Moll. Russia: 337.
(error for Unio douglasiae Griffith & Pidgeon, 1833)

Unio douglassiaeKantor & Sysoev (2005)

Nodularia douglasiae nipponensis Sano et al., 2017
‘Martens’ of Authors.
Sano et al., 2017, J. Wat. Res. Prot. 9: Table 2. [in part]
(misidentified reference to Unio nipponensis Martens, 1877)

Nodularia douglasiae nipponensisSano et al. (2017)
Nodularia douglasiaeChoi et al. (2020), Sano et al. (2020)

Nodularia jourdyi Bolotov et al., 2018
Bolotov et al., 2018, Sci. Repts. 8: 4.
Pfeiffer et al., 2019, MPE 137: 117.
(reference to Unio jourdyi Morlet, 1886)

Nodularia jourdyiBolotov et al. (2018)
Nodularia douglasiaeLopes-Lima et al. (2020), Choi et al. (2020), Sayenko et al. (2021), Wu et al. (2024)

Nodularia suifunica Klishko et al., 2018
‘Moskvicheva’ of Authors.
Klishko et al., 2018, Systemat. Biodiv. 16(3): 288.
Bogatov, 2022, Lg. Bivalve Moll. Russia’s Fresh Waters: 59.
(error for Nodularia sujfunica Moskvicheva, 1973)

Nodularia suifunicaKlishko et al. (2018)

Unio osbreckii Lopes-Lima et al., 2020
‘Philippi’ Lopes-Lima et al., 2020, MPE 146: Appendix 1.
(error for Unio osbeckii Philippi, 1844)

Unio osbreckiiLopes-Lima et al. (2020)

Taxonomic History (since 1900)

Nodularia (s.s.) douglasiaeSimpson (1900), Preston (1912), Simpson (1914), Haas (1924), Korniushin (1998)
Nodularia (s.s.) douglasiae var. shanghaiensisSimpson (1900), Preston (1912), Simpson (1914)
Nodularia (s.s.) biwaeSimpson (1900), Haas (1911), Simpson (1914)
Nodularia (s.s.) douglasiae var. nipponensisSimpson (1900)
Nodularia (s.s.) jourdyiSimpson (1900), Simpson (1914)
Unio (Nodularia) douglasiaeMartens (1905), Zhadin (1938)
Unio (Nodularia) nipponensisMartens (1905)
Unio (Nodularia) douglasiae var. sinuolatusMartens (1905)
Unio jourdyiDautzenberg & Fischer (1905), Dautzenberg & Fischer (1905)
Unio (Nodularia) pliculosusMartens (1905)
Unio (Nodularia) verruciferMartens (1905)
Nodularia douglasiae taiwanicaPilsbry & Hirase (1905)
Unio jourdyi var. corrugataDautzenberg & Fischer (1905)
Unio jourdyi var. ponderosaDautzenberg & Fischer (1905)
Nodularia denserugataHaas (1910), Graf & Cummings (2007), Bogan (2010)
Nodularia verrucosaHaas (1910), Simpson (1914)
Nodularia douglasiae crassidensHaas (1910), Dang et al. (1980), Do et al. (2018)
Nodularia douglasiae flavoviridisHaas (1910)
Nodularia (s.s.) douglasiae douglasiaeHaas (1911)
Nodularia (s.s.) douglasiae shanghaiensisHaas (1911)
Nodularia (s.s.) pliculosusHaas (1911)
Nodularia (s.s.) verruciferaHaas (1911), Simpson (1914)
Nodularia (s.s.) denserugataHaas (1911), Simpson (1914)
Nodularia (s.s.) douglasiae crassidensHaas (1911)
Oxynaia jourdyiHaas (1913), Thiele (1934), Haas (1969), Haas (1969), Dang et al. (1980), Graf & Cummings (2007), Bogan (2010), IUCN (2011), Do et al. (2018), Zieritz et al. (2018)
Unio wrigtiide Gregorio (1914)
Nodularia (s.s.) pliculosaSimpson (1914)
Nodularia (s.s.) boettgeriSimpson (1914)
Unio kobeltiSimpson (1914)
Nodularia (s.s.) douglasiae var. taiwanicaSimpson (1914)
Nodularia (s.s.) jourdyi var. corrugataSimpson (1914)
Nodularia (s.s.) jourdyi var. ponderosaSimpson (1914)
Nodularia (s.s.) douglasiae var. crassidensSimpson (1914)
Nodularia douglasiaeHaas (1920), Prozorova et al. (2005), Graf & Cummings (2007), Graf (2007), ..., Wu et al. (2024), Dai et al. (2024), Wu et al. (2024), Wu et al. (2024)
Nodularia (s.s.) douglasiae taiwanicaHaas (1924)
Unio (Nodularia) douglasiae var. schrenkiZhadin (1938)
Unio pictorum middendorffiHaas (1940)
Unio douglasiae taiwanicusKuroda (1941)
Unio douglasiaeZhadin (1965), Kil (1976), Habe (1977), Liu et al. (1979), ..., Wang et al. (2018), Aunins et al. (2018), Wang et al. (2018), Araujo et al. (2018)
Unio douglasiae var. abbreviatusZhadin (1965)
Unio douglasiae var. schrenckiZhadin (1965)
Unio douglasiae douglasiaeHaas (1969)
Unio douglasiae amurensisHaas (1969)
Unio biwaeHaas (1969), Habe (1977), Habe (1991), Slugina (2006)
Unio douglasiae pliculosusHaas (1969)
Unio douglasiae verruciferHaas (1969)
Unio douglasiae taiwanicaHaas (1969)
Unio denserugatusHaas (1969)
Nodularia middendorffiMoskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Nodularia amurensisMoskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Nodularia abbreviataMoskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010)
Nodularia schrenckiMoskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010), Bogatov & Prozorova (2017)
Middendorffinaia (Sujfununio) denserugataMoskvicheva & Starobogatov (1973)
Nodularia flavoviridisMoskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010)
Middendorffinaia arsenjeviMoskvicheva & Starobogatov (1973)
Middendorffinaia (s.s.) arsenieviMoskvicheva & Starobogatov (1973), Zatravkin & Bogatov (1987), Starobogatov et al. (2004)
Nodularia sujfunicaMoskvicheva (1973), Zatravkin (1983)
Nodularia vladivostokensisMoskvicheva (1973), Zatravkin (1983), Kantor & Sysoev (2005), Kantor et al. (2010), Klishko et al. (2018)
Unio taiwanensisHabe (1977)
Unio firnusLiu et al. (1979)
Unio douglasiae chlorensLiu et al. (1979)
Unio douglasiae cleltaeLiu et al. (1979)
Nodularia doctylinaLiu et al. (1980)
Middendorffinaia (s.s.) arsenjeviZatravkin (1983)
Nodularia sp.n.Zatravkin (1983)
Nodularia lebedeviZatravkin & Starobogatov (1984), Kantor & Sysoev (2005), Kantor et al. (2010)
Nodularia (s.s.) middendorffiZatravkin & Bogatov (1987), Starobogatov et al. (2004), Bogatov (2012)
Nodularia (s.s.) amurensisZatravkin & Bogatov (1987), Starobogatov et al. (2004), Bogatov (2012)
Nodularia (s.s.) abbreviataZatravkin & Bogatov (1987), Starobogatov et al. (2004)
Nodularia (s.s.) schrenckiZatravkin & Bogatov (1987), Starobogatov et al. (2004), Bogatov (2012)
Nodularia (s.s.) flavoviridisZatravkin & Bogatov (1987), Starobogatov et al. (2004)
Nodularia (s.s.) sujfunicaZatravkin & Bogatov (1987)
Nodularia (s.s.) vladivostokensisZatravkin & Bogatov (1987), Starobogatov et al. (2004)
Nodularia (Amurunio) lebedeviZatravkin & Bogatov (1987), Korniushin (1998), Starobogatov et al. (2004), Klishko et al. (2018)
Scabies longataLiu et al. (1991), Hu (2005), Graf & Cummings (2007), Bogan (2010), WoRMS (2017)
Nodularia (Magadaninaia) extremalisMartynov & Chernyshev (1992), Starobogatov et al. (2004)
Nodularia (s.s.) moskvichevaeBogatov & Starobogatov (1992), Starobogatov et al. (2004)
Unio douglasiae biwaeKondo (1997), Kondo (2008)
Middendorffinaia (Middendorffinaia) ochoticaBogatov (2000)
Unio sculptusYu et al. (2001)
Nodularia sachalinensisBogatov (2001)
Nodularia sakhalinensisBogatov (2001), Kantor & Sysoev (2005), Kantor et al. (2010), Klishko et al. (2018)
Middendorffinaia (s.s.) ochoticaStarobogatov et al. (2004), Bogatov (2012), Vinarski & Kantor (2016), Bogatov & Prozorova (2017)
Nodularia (s.s.) sakhalinensisStarobogatov et al. (2004)
Nodularia crassidensProzorova et al. (2005)
Middendorffinaia arsenieviKantor & Sysoev (2005), Kantor et al. (2010)
Nodularia extremalisKantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Nodularia moskvichevaeKantor & Sysoev (2005), Kantor et al. (2010), Vinarski & Kantor (2016)
Middendorffinaia ochoticaKantor & Sysoev (2005), Kantor et al. (2010)
Unio douglassiaeKantor & Sysoev (2005)
Unio crassus mongolicusGraf (2007)
Nodularia biwaeBogan (2010)
Nodularia dorriIUCN (2012)
Middendorffinaia (s.s.) ussuriensisBogatov (2012), Vinarski & Kantor (2016)
Nodularidia douglasiaeHe & Zhuang (2013)
Ptychorhynchus denserugataHe & Zhuang (2013), Zieritz et al. (2018)
Scabies crispataHe & Zhuang (2013), Graf & Cummings (2021)
Acuticosta jianghanensisHe & Zhuang (2013), Zieritz et al. (2018), Liu et al. (2020)
Nodularia douglasiae biwaeSano et al. (2017), Mabuchi & Nishida (2020)
Nodularia douglasiae nipponensisSano et al. (2017)
Nodularia jourdyiBolotov et al. (2018), Pfeiffer et al. (2019), Graf & Cummings (2021), Sayenko et al. (2021)
Nodularia suifunicaKlishko et al. (2018)
Nodularia cf. douglasiaeHuang et al. (2019)
Middendorffinaia mongolicaKlishko et al. (2019)
Nodularia douglasiae Clade CChoi et al. (2020)
Nodularia douglasiae Clade BChoi et al. (2020)
Nodularia douglasiae Clade AChoi et al. (2020)
Unio osbreckiiLopes-Lima et al. (2020)
Ptychorhynchus denserugatumGraf & Cummings (2021)
Nodularia douglasiae douglasiaeBogatov (2022)
Nodularia douglasiae amurensisBogatov (2022)
Nodularia douglasiae middendorffiBogatov (2022)
Nodularia douglasiae schrenckiBogatov (2022)
Middendorffinaia mongolica ussuriensisBogatov (2022)
Middendorffinaia mongolica ochoticaBogatov (2022)
Nodularia sp. 2Dai et al. (2024)

Page updated: 08:48:05 Wed 06 Nov 2024

 
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