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Sun 07 Sep 2025 |
The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!
The nominal species (available and otherwise) in this synonymy are listed by usage rather than strictly by publication date. The primary sorting order is by the most recent year (since 1969) that the species name was treated as valid, and then nominal species are sorted secondarily by publication year. The taxonomic history of the valid species (since 1900) is listed at the bottom of the page. The MUSSEL Project Database has moved to a new server, with a new URL! The new address for this page is: https://musselpdb.org/db.php?ty=validsp_syn&id=5703
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Unionoidea | Unionidae | Ambleminae | Quadrulini | Quadrulina | Pustulosa species Pustulosa kieneriana (Lea, 1852): nominal species
3 specimens available.
Unio kienerianus Lea, 1852
Lea, 1852, Trans. Amer. Phil. Soc. 10 [O. 5]: 281 [37], pl. 23, fig. 40 [as ‘keineriana’].
(correct name for Unio keinerianus Lea, 1852)
Unio kienerianus — Lea (1852), B.H. Wright (1888), Singley (1893)
Margaron (Unio) keinerianus — Lea (1852)
Unio keinerianus — Conrad (1853)
Unio (Rotundaria) kieneriana — Adams & Adams (1857)
Margaron (Unio) kienerianus — Lea (1870)
Quadrula (sec. Theliderma) pustulosa var. keineriana — Simpson (1900)
Quadrula (sec. Theliderma) pustulosa var. kieneriana — Simpson (1914)
Quadrula (Bullata) pustulosa-keineriana — Frierson (1927)
Quadrula (Pustulosa) pustulosa keineriana — Haas (1969)
Quadrula pustulosa — R.I. Johnson (1980), Howells et al. (1996)
Quadrula keineriana — Serb et al. (2003)
Quadrula kieneriana — Campbell et al. (2005), Williams et al. (2008), Bogan (2010), Szumowski et al. (2012)
Amphinaias keineriana — ITIS (2013)
Cyclonaias kieneriana — Williams et al. (2017), Lopes-Lima et al. (2019), Graf & Cummings (2021), FMCS (2021), Bieler (2021), Hopper et al. (2023), Hauck et al. (2023)
Pustulosa kieneriana — Neemuchwala et al. (2023), FMCS (2025)
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Unio asperatus Lea, 1861
Lea, 1861, Proc. Acad. Nat. Sci. 13: 41.
Lea, 1862, Jour. Acad. Nat. Sci. 5 [O. 8]: 68 [72], pl. 7, fig. 218.
Type(s): HOLOTYPE USNM_84272, Claiborne, Ala.
Unio asperatus — Lea (1861), Sowerby (1868), B.H. Wright (1888), Paetel (1890)
Margaron (Unio) asperatus — Lea (1870)
Unio pustulosus — Call (1895)
Quadrula (sec. Theliderma) pustulosa var. pernodosa — Simpson (1900), Simpson (1914)
Quadrula pustulosa asperata — Utterback (1915)
Quadrula asperata — Ortmann (1923), Turgeon et al. (1988), Williams et al. (1993), Turgeon et al. (1998), ..., Bogan (2010), ITIS (2012), Haag (2012), Szumowski et al. (2012)
Quadrula (Bullata) pustulosa-asperata — Frierson (1927)
Quadrula (Pustulosa) pustulosa pernodosa — Haas (1969)
Quadrula pustulosa — Burch (1975), R.I. Johnson (1980)
Amphinaias asperata — IUCN (1996), Graf & Cummings (2007), Cummings & Graf (2010)
Cyclonaias asperata — Williams et al. (2017), Johnson et al. (2018), Hauck et al. (2023)
Cyclonaias kieneriana — Lopes-Lima et al. (2019), Graf & Cummings (2021), FMCS (2021)
Pustulosa kieneriana — FMCS (2025)
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Quadrula archeri Frierson, 1905
Frierson, 1905, Nautilus 19(2): 13, pl. 1, figs. 1-2.
Type(s): HOLOTYPE UMMZ_113538, Tallapoosa River, Tallassee, Elmore, Alabama.
Quadrula archeri — Frierson (1905), Burch (1975)
Quadrula (sec. Theliderma) archeri — Simpson (1914)
Quadrula (Bullata) archeri — Frierson (1927)
Quadrula (Pustulosa) archeri — Haas (1969)
Amphinaias archeri — Graf & Cummings (2007), Cummings & Graf (2010)
Quadrula asperata — Williams et al. (2008)
Cyclonaias archeri — Williams et al. (2017), FMCS (2021), Hopper et al. (2023)
Cyclonaias pustulosa — Lopes-Lima et al. (2019), Graf & Cummings (2021)
Pustulosa kieneriana — FMCS (2025)
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Unio keinerianus Lea, 1852
 
Lea, 1852, Proc. Amer. Phil. Soc. 5: 251. [nomen nudum]
Lea, 1852, Trans. Amer. Phil. Soc. 10 [O. 5]: 281 [37], pl. 23, fig. 40.
(incorrect original spelling of Unio kienerianus Lea, 1852)
Type(s): HOLOTYPE USNM_84273, Coosawattee R., Ga.
Unio keinerianus — Lea (1852)
Quadrula (sec. Theliderma) pustulosa var. kieneriana — Simpson (1914)
Cyclonaias kieneriana — Bieler (2021)
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Unio asperatus Sowerby, 1868
‘Lea’ Sowerby, 1868, Conch. Icon. 16: pl. 85, sp. 450.
(reference to Unio asperatus Lea, 1861)
Unio asperatus — Sowerby (1868), Paetel (1890)
Quadrula (sec. Theliderma) pustulosa var. pernodosa — Simpson (1900)
Quadrula (Striata) striata — Frierson (1927)
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Taxonomic History (since 1900)
Quadrula (sec. Theliderma) pustulosa var. keineriana — Simpson (1900)
Quadrula (sec. Theliderma) pustulosa var. pernodosa — Simpson (1900), Simpson (1914)
Quadrula archeri — Frierson (1905), Burch (1975)
Quadrula (sec. Theliderma) pustulosa var. kieneriana — Simpson (1914)
Quadrula (sec. Theliderma) archeri — Simpson (1914)
Quadrula pustulosa asperata — Utterback (1915)
Quadrula asperata — Ortmann (1923), Turgeon et al. (1988), Williams et al. (1993), Turgeon et al. (1998), ..., Bogan (2010), ITIS (2012), Haag (2012), Szumowski et al. (2012)
Quadrula (Bullata) pustulosa-keineriana — Frierson (1927)
Quadrula (Bullata) pustulosa-asperata — Frierson (1927)
Quadrula (Striata) striata — Frierson (1927)
Quadrula (Bullata) archeri — Frierson (1927)
Quadrula (Pustulosa) pustulosa keineriana — Haas (1969)
Quadrula (Pustulosa) pustulosa pernodosa — Haas (1969)
Quadrula (Pustulosa) archeri — Haas (1969)
Quadrula pustulosa — Burch (1975), R.I. Johnson (1980), Howells et al. (1996)
Amphinaias asperata — IUCN (1996), Graf & Cummings (2007), Cummings & Graf (2010)
Quadrula keineriana — Serb et al. (2003)
Quadrula kieneriana — Campbell et al. (2005), Williams et al. (2008), Bogan (2010), Szumowski et al. (2012)
Amphinaias archeri — Graf & Cummings (2007), Cummings & Graf (2010)
Amphinaias keineriana — ITIS (2013)
Cyclonaias kieneriana — Williams et al. (2017), Lopes-Lima et al. (2019), Graf & Cummings (2021), FMCS (2021), Bieler (2021), Hopper et al. (2023), Hauck et al. (2023)
Cyclonaias asperata — Williams et al. (2017), Johnson et al. (2018), Hauck et al. (2023)
Cyclonaias archeri — Williams et al. (2017), FMCS (2021), Hopper et al. (2023)
Cyclonaias pustulosa — Lopes-Lima et al. (2019), Graf & Cummings (2021)
Pustulosa kieneriana — Neemuchwala et al. (2023), FMCS (2025)
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Page updated: 08:28:45 Sun 07 Sep 2025 |
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