MUSSELp
 
MUSSELpdb
 
 
Taxonomy
 
 
Geography
 
 
 

Page last updated
Wed 05 Jun 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Parreysiinae

tribe COELATURINI Modell, 1942: cladomics
     10 genera

Cladomics Summary

number of species = 41
number of subtaxa (rank: genus) = 10

analyzed in 47 cladograms in 30 publications
maximum number of terminals in an analysis = 27, mean = 4.09
maximum number of subtaxa (genus) in an analysis = 7, mean = 2.11

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 25
maximum number of included terminals = 27, mean = 6.8
maximum number of subtaxa (genus) = 7, mean = 3.08
proportion of cladograms that support basic monophyly = 1.0

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 25
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (tribe).
maximum number of included terminals = 27, mean = 6.8
maximum number of subtaxa (genus) = 7, mean = 3.08
proportion of cladograms that support rigorous monophyly = 1.0


Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ tree


Ortiz-Sepulveda et al., 2020 [Publ 4101], fig. 1

iT tT bi riS oT roT roS iD
27 27 16 7 9 9 2 0.0

*** Top Monophyly Cladogram ***

  +-------------------------------------- Leoparreysia_tavoyensis      
  |                                                                    
  |                          +----------- Indonaia_andersoniana        
  |                          |                                         
  |  +-----------------------+  +-------- Indochinella_pugio           
  |  |                       |  |                                      
--+  |                       |  |  +----- Radiatula_mouhoti            
  |  |                       +--+  |                                   
  |  |                          |  +----- Radiatula_myitkyinae         
  |  |                          |  |                                   
  |  |                          +--+----- Radiatula_chaudhurii         
  |  |                             |                                   
  |  |                             |  +-- Scabies_humilis              
  +--+                             |  |                                
     |                             +--+-- Scabies_phaselus             
     |                                |                                
     |                                +-- Scabies_mandarinus           
     |                                                                 
     |  +-------------------------------- Grandidieria_burtoni **         
     |  |                                                              
     |  |                             +-- Pseudospatha_tanganyicensis **  
     |  |  +--------------------------+                                
     +--+  |                          +-- Moncetia_anceyi **              
        |  |                                                           
        |  |                          +-- Nitia_acuminata **              
        |  |                          |                                
        +--+  +-----------------------+-- Nitia_monceti **                
           |  |                       |                                
           |  |                       +-- Nitia_teretiuscula **           
           |  |                                                        
           |  |                       +-- Coelatura_aegyptiaca **         
           |  |                       |                                
           +--+                       +-- Coelatura_alluaudi **           
              |                       |                                
              |  +--------------------+-- Coelatura_bakeri **             
              |  |                    |                                
              |  |                    +-- Coelatura_hauttecoeuri **       
              |  |                    |                                
              |  |                    +-- Coelatura_stuhlmanni **         
              +--+                                                     
                 |                    +-- Coelatura_leopoldvillensis **   
                 |  +-----------------+                                
                 |  |                 +-- Coelatura_mesafricana **        
                 |  |                                                  
                 |  |                 +-- Coelatura_horei **              
                 +--+                 |                                
                    |  +--------------+-- Coelatura_ratidota **           
                    |  |              |                                
                    |  |              +-- Coelatura **                    
                    +--+                                               
                       |  +-------------- Coelatura_gabonensis **         
                       |  |                                            
                       |  +-------------- Coelatura_stanleyvillensis **   
                       +--+                                            
                          |  +----------- Coelatura_kunenensis **         
                          |  |                                         
                          |  |     +----- Prisodontopsis_aviculaeformis **
                          +--+  +--+                                   
                             |  |  |  +-- Coelatura_briarti **            
                             |  |  +--+                                
                             |  |     +-- Prisodontopsis_mweruensis **    
                             +--+                                      
                                |     +-- Coelatura_choziensis **         
                                |     |                                
                                |     +-- Coelatura_hypsiprymna **        
                                |     |                                
                                +-----+-- Coelatura_luapulaensis **       
                                      |                                
                                      +-- Coelatura_mossambicensis **     
                                      |                                
                                      +-- Nyassunio_nyassaensis **        

Bolotov et al., 2024 [Publ 4369], fig. 2

iT tT bi riS oT roT roS iD
4 4 2 3 58 58 4 23.0

*** Top Sister-Group Cladogram ***

                                      +-- Leoparreysia_burmana    
                                +-----+                           
                                |     +-- Leoparreysia_whitteni   
                       +--------+                                 
                       |        |  +----- Leoparreysia_olivacea   
                       |        +--+                              
                       |           |  +-- Leoparreysia_canefrii   
  +--------------------+           +--+                           
  |                    |              +-- Leoparreysia_tavoyensis 
  |                    |                                          
  |                    |  +-------------- Assamnaia_involuta      
  |                    |  |                                       
  |                    +--+           +-- Parreysia_keralaensis   
  |                       |  +--------+                           
  |                       |  |        +-- Parreysia_marcens       
  |                       +--+                                    
  |                          |  +-------- Parreysia_rakhinensis   
  |                          +--+                                 
  |                             |  +----- Parreysia_corrugata     
  |                             +--+                              
  |                                |  +-- Parreysia_plagiosoma    
--+                                +--+                           
  |                                   +-- Parreysia_nagpoorensis  
  |                                                               
  |                             +-------- Nitia_teretiuscula **      
  |  +--------------------------+                                 
  |  |                          |  +----- Coelatura_aegyptiaca **    
  |  |                          +--+                              
  |  |                             |  +-- Coelatura_kunenensis **    
  |  |                             +--+                           
  |  |                                +-- Nyassunio_nyassaensis **   
  |  |                                                            
  |  |           +----------------------- Pseudoscabies_scobina   
  |  |           |                                                
  |  |  +--------+  +-------------------- Indonaia_cylindrica     
  +--+  |        |  |                                             
     |  |        +--+  +----------------- Indonaia_gratiosa       
     |  |           |  |                                          
     |  |           +--+  +-------------- Indonaia_corbis         
     |  |              |  |                                       
     |  |              +--+  +----------- Indonaia_andersoniana   
     |  |                 |  |                                    
     |  |                 +--+        +-- Indonaia_caerulea       
     |  |                    |  +-----+                           
     |  |                    |  |     +-- Indonaia_subclathrata   
     +--+                    +--+                                 
        |                       |  +----- Indonaia_rugosa         
        |                       +--+                              
        |                          |  +-- Indonaia_bonneaudii     
        |                          +--+                           
        |                             +-- Indonaia_shurtleffiana  
        |                                                         
        |  +----------------------------- Indochinella_pugio      
        |  |                                                      
        |  |                 +----------- Radiatula_crispisulcata 
        |  |                 |                                    
        |  |                 |     +----- Radiatula_chaudhurii    
        +--+  +--------------+  +--+                              
           |  |              |  |  |  +-- Radiatula_myitkyinae    
           |  |              |  |  +--+                           
           |  |              +--+     +-- Radiatula_kachinensis   
           |  |                 |                                 
           |  |                 |  +----- Radiatula_chindwinensis 
           +--+                 +--+                              
              |                    |  +-- Radiatula_myitthanensis 
              |                    +--+                           
              |                       +-- Radiatula_mouhoti       
              |                                                   
              |  +----------------------- Scabiellus_songkramensis
              |  |                                                
              +--+  +-------------------- Harmandia_somboriensis  
                 |  |                                             
                 +--+  +----------------- Unionetta_fabagina      
                    |  |                                          
                    +--+              +-- Scabies_crispata        
                       |  +-----------+                           
                       |  |           +-- Scabies_mandarinus      
                       +--+                                       
                          |  +----------- Scabies_nucleus         
                          |  |                                    
                          +--+        +-- Scabies_humilis         
                             |  +-----+                           
                             |  |     +-- Scabies_pilata          
                             +--+                                 
                                |  +----- Scabies_phaselus        
                                +--+                              
                                   |  +-- Scabies_anceps          
                                   +--+                           
                                      +-- Scabies_scobinatus      

Ortiz-Sepulveda et al., 2020 [Publ 4101], fig. 2

iT tT bi riS oT roT roS iD
27 27 16 7 9 9 2 5.0
  +-------------------------------- Leoparreysia_tavoyensis      
  |                                                              
  |                          +----- Radiatula_mouhoti            
  |                          |                                   
  |                          +----- Radiatula_myitkyinae         
  |                          |                                   
  |                          +----- Radiatula_chaudhurii         
  |                          |                                   
--+  +-----------------------+----- Indochinella_pugio           
  |  |                       |                                   
  |  |                       +----- Indonaia_andersoniana        
  |  |                       |                                   
  |  |                       |  +-- Scabies_humilis              
  |  |                       |  |                                
  |  |                       +--+-- Scabies_phaselus             
  |  |                          |                                
  +--+                          +-- Scabies_mandarinus           
     |                                                           
     |                       +----- Grandidieria_burtoni **         
     |  +--------------------+                                   
     |  |                    |  +-- Pseudospatha_tanganyicensis **  
     |  |                    +--+                                
     |  |                       +-- Moncetia_anceyi **              
     |  |                                                        
     |  |                       +-- Nitia_acuminata **              
     +--+                       |                                
        |  +--------------------+-- Nitia_monceti **                
        |  |                    |                                
        |  |                    +-- Nitia_teretiuscula **           
        |  |                                                     
        |  |                    +-- Coelatura_aegyptiaca **         
        |  |                    |                                
        +--+                    +-- Coelatura_alluaudi **           
           |                    |                                
           |  +-----------------+-- Coelatura_bakeri **             
           |  |                 |                                
           |  |                 +-- Coelatura_hauttecoeuri **       
           |  |                 |                                
           |  |                 +-- Coelatura_stuhlmanni **         
           +--+                                                  
              |                 +-- Coelatura_leopoldvillensis **   
              |  +--------------+                                
              |  |              +-- Coelatura_mesafricana **        
              |  |                                               
              |  |              +-- Coelatura_horei **              
              +--+              |                                
                 |  +-----------+-- Coelatura_ratidota **           
                 |  |           |                                
                 |  |           +-- Coelatura **                    
                 |  |                                            
                 +--+           +-- Coelatura_gabonensis **         
                    |  +--------+                                
                    |  |        +-- Coelatura_stanleyvillensis **   
                    |  |                                         
                    +--+  +-------- Coelatura_kunenensis **         
                       |  |                                      
                       |  |     +-- Coelatura_briarti **            
                       |  |     |                                
                       +--+  +--+-- Prisodontopsis_mweruensis **    
                          |  |  |                                
                          |  |  +-- Prisodontopsis_aviculaeformis **
                          |  |                                   
                          +--+  +-- Coelatura_choziensis **         
                             |  |                                
                             |  +-- Coelatura_hypsiprymna **        
                             |  |                                
                             +--+-- Coelatura_luapulaensis **       
                                |                                
                                +-- Coelatura_mossambicensis **     
                                |                                
                                +-- Nyassunio_nyassaensis **        

Graf et al., 2014 [Publ 3591], fig. 3

iT tT bi riS oT roT roS iD
24 24 11 3 3 1 1 24.0
  +----------------------------- Leoparreysia_tavoyensis         
  |                                                              
--+  +-------------------------- Nitia_teretiuscula **              
  |  |                                                           
  +--+  +----------------------- Coelatura_aegyptiaca[2] **         
     |  |                                                        
     +--+  +-------------------- Coelatura_aegyptiaca[1] **         
        |  |                                                     
        |  |  +----------------- Coelatura_gabonensis **            
        +--+  |                                                  
           |  |  +-------------- Coelatura_kunenensis **            
           |  |  |                                               
           |  |  |           +-- Coelatura_choziensis[1] **         
           +--+  |           |                                   
              |  |           +-- Coelatura_choziensis[2] **         
              |  |           |                                   
              |  |           +-- Coelatura_luapulaensis[1] **       
              |  |           |                                   
              +--+  +--------+-- Coelatura_luapulaensis[2] **       
                 |  |        |                                   
                 |  |        +-- Coelatura_luapulaensis[3] **       
                 |  |        |                                   
                 |  |        +-- Coelatura_luapulaensis[4] **       
                 |  |        |                                   
                 |  |        +-- Coelatura_luapulaensis[5] **       
                 |  |                                            
                 +--+     +----- Prisodontopsis_mweruensis[2] **    
                    |     |                                      
                    |     +----- Prisodontopsis_mweruensis[3] **    
                    |  +--+                                      
                    |  |  |  +-- Prisodontopsis_mweruensis[4] **    
                    |  |  |  |                                   
                    |  |  +--+-- Prisodontopsis_mweruensis[5] **    
                    |  |     |                                   
                    +--+     +-- Prisodontopsis_aviculaeformis[7] **
                       |                                         
                       |  +----- Prisodontopsis_aviculaeformis[1] **
                       |  |                                      
                       |  |  +-- Prisodontopsis_aviculaeformis[2] **
                       +--+  |                                   
                          |  +-- Prisodontopsis_aviculaeformis[3] **
                          |  |                                   
                          |  +-- Prisodontopsis_aviculaeformis[4] **
                          +--+                                   
                             +-- Prisodontopsis_aviculaeformis[5] **
                             |                                   
                             +-- Prisodontopsis_aviculaeformis[6] **
                             |                                   
                             +-- Prisodontopsis_mweruensis[1] **    

Bolotov et al., 2017 [Publ 3764], fig. S1

iT tT bi riS oT roT roS iD
9 9 0 3 123 33 4 26.0
              +-------- Leoparreysia_olivacea        
  +-----------+                                      
  |           |  +----- Leoparreysia_tavoyensis[2]   
  |           +--+                                   
  |              |  +-- Leoparreysia_canefrii        
  |              +--+                                
  |                 +-- Leoparreysia_tavoyensis[1]   
  |                                                  
  |                 +-- Nitia_teretiuscula **           
  |                 |                                
  |                 +-- Coelatura_gabonensis **         
  |                 |                                
  |                 +-- Prisodontopsis_mweruensis **    
--+                 |                                
  |                 +-- Coelatura_kunenensis **         
  |                 |                                
  |  +--------------+-- Coelatura_aegyptiaca[1] **      
  |  |              |                                
  |  |              +-- Coelatura_aegyptiaca[2] **      
  |  |              |                                
  |  |              +-- Prisodontopsis_aviculaeformis **
  |  |              |                                
  |  |              +-- Coelatura_luapulaensis **       
  |  |              |                                
  |  |              +-- Coelatura_choziensis **         
  |  |                                               
  +--+     +----------- Indochinella_pugio           
     |     |                                         
     |     |        +-- Scabies_humilis              
     |  +--+        |                                
     |  |  |  +-----+-- Scabies_phaselus             
     |  |  |  |     |                                
     |  |  +--+     +-- Scabies_mandarinus           
     |  |     |                                      
     |  |     |  +----- Radiatula_mouhoti            
     |  |     +--+                                   
     +--+        |  +-- Radiatula_myitkyinae         
        |        +--+                                
        |           +-- Radiatula_chaudhurii         
        |                                            
        |     +-------- Indonaia_shurtleffiana       
        |     |                                      
        |     |     +-- Indonaia_andersoniana        
        +-----+  +--+                                
              |  |  +-- Indonaia_caerulea            
              |  |                                   
              |  |  +-- Indonaia_bonneaudii          
              +--+--+                                
                 |  +-- Lamellidens_corrianus[2]     
                 |                                   
                 |  +-- Pseudoscabies_scobina        
                 +--+                                
                    +-- Indonaia_rugosa              

Bolotov et al., 2017 [Publ 3761], fig. S1

iT tT bi riS oT roT roS iD
6 6 4 3 135 23 4 25.0
                    +-- Leoparreysia_tavoyensis  
  +-----------------+                            
  |                 +-- Leoparreysia_olivacea    
  |                                              
  |     +-------------- Nitia_teretiuscula **       
  |  +--+                                        
  |  |  |  +----------- Coelatura_aegyptiaca[2] **  
--+  |  +--+                                     
  |  |     |  +-------- Coelatura_aegyptiaca[1] **  
  |  |     +--+                                  
  |  |        |  +----- Prisodontopsis_mweruensis **
  |  |        +--+                               
  |  |           |  +-- Coelatura_gabonensis **     
  +--+           +--+                            
     |              +-- Coelatura_kunenensis **     
     |                                           
     |        +-------- Indonaia_shurtleffiana   
     |  +-----+                                  
     |  |     |  +----- Indonaia_andersoniana    
     |  |     +--+                               
     |  |        |  +-- Indonaia_rugosa          
     |  |        +--+                            
     +--+           +-- Indonaia_bonneaudii      
        |                                        
        |           +-- Indochinella_pugio       
        |  +--------+                            
        |  |        +-- Scabies_crispata         
        |  |                                     
        +--+        +-- Radiatula_myitkyinae     
           |  +-----+                            
           |  |     +-- Radiatula_chaudhurii     
           +--+                                  
              |  +----- Scabies_mandarinus       
              +--+                               
                 |  +-- Scabies_humilis          
                 +--+                            
                    +-- Scabies_phaselus         

Bolotov et al., 2017 [Publ 3761], fig. S2

iT tT bi riS oT roT roS iD
6 6 4 3 135 23 4 23.0
                    +-- Leoparreysia_olivacea    
  +-----------------+                            
  |                 +-- Leoparreysia_tavoyensis  
  |                                              
  |     +-------------- Nitia_teretiuscula **       
  |  +--+                                        
  |  |  |  +----------- Coelatura_aegyptiaca[1] **  
--+  |  +--+                                     
  |  |     |  +-------- Coelatura_aegyptiaca[2] **  
  |  |     +--+                                  
  |  |        |  +----- Coelatura_gabonensis **     
  |  |        +--+                               
  |  |           |  +-- Coelatura_kunenensis **     
  +--+           +--+                            
     |              +-- Prisodontopsis_mweruensis **
     |                                           
     |        +-------- Indonaia_shurtleffiana   
     |  +-----+                                  
     |  |     |  +----- Indonaia_andersoniana    
     |  |     +--+                               
     |  |        |  +-- Indonaia_rugosa          
     |  |        +--+                            
     +--+           +-- Indonaia_bonneaudii      
        |                                        
        |           +-- Indochinella_pugio       
        |  +--------+                            
        |  |        +-- Scabies_crispata         
        |  |                                     
        +--+        +-- Radiatula_myitkyinae     
           |  +-----+                            
           |  |     +-- Radiatula_chaudhurii     
           +--+                                  
              |  +----- Scabies_mandarinus       
              +--+                               
                 |  +-- Scabies_humilis          
                 +--+                            
                    +-- Scabies_phaselus         

Huang et al., 2019 [Publ 3843], fig. 2

iT tT bi riS oT roT roS iD
6 6 2 3 199 31 4 25.0
                    +-- Leoparreysia_tavoyensis  
                    |                            
  +-----------------+-- Leoparreysia_canefrii    
  |                 |                            
  |                 +-- Leoparreysia_olivacea    
  |                                              
  |           +-------- Coelatura_aegyptiaca[2] **  
  |           |                                  
  |  +--------+  +----- Coelatura_aegyptiaca[1] **  
--+  |        |  |                               
  |  |        |  +----- Nitia_teretiuscula **       
  |  |        +--+                               
  |  |           |  +-- Coelatura_gabonensis **     
  |  |           |  |                            
  |  |           +--+-- Coelatura_kunenensis **     
  |  |              |                            
  |  |              +-- Prisodontopsis_mweruensis **
  +--+                                           
     |        +-------- Indonaia_shurtleffiana   
     |        |                                  
     |  +-----+  +----- Indonaia_bonneaudii      
     |  |     |  |                               
     |  |     |  |  +-- Indonaia_rugosa          
     |  |     +--+--+                            
     |  |        |  +-- Pseudoscabies_scobina    
     |  |        |                               
     +--+        |  +-- Indonaia_andersoniana    
        |        +--+                            
        |           +-- Indonaia_caerulea        
        |                                        
        |  +----------- Indochinella_pugio       
        |  |                                     
        |  |     +----- Scabies_humilis          
        +--+  +--+                               
           |  |  |  +-- Scabies_phaselus         
           |  |  +--+                            
           +--+     +-- Scabies_mandarinus       
              |                                  
              |  +----- Radiatula_myitkyinae     
              +--+                               
                 |  +-- Radiatula_chaudhurii     
                 +--+                            
                    +-- Radiatula_mouhoti        

Huang et al., 2019 [Publ 3843], fig. S3

iT tT bi riS oT roT roS iD
6 6 4 3 196 31 4 27.0
                 +----- Leoparreysia_canefrii    
  +--------------+                               
  |              |  +-- Leoparreysia_tavoyensis  
  |              +--+                            
  |                 +-- Leoparreysia_olivacea    
  |                                              
  |        +----------- Coelatura_aegyptiaca[2] **  
  |        |                                     
  |  +-----+        +-- Coelatura_aegyptiaca[1] **  
--+  |     |  +-----+                            
  |  |     |  |     +-- Nitia_teretiuscula **       
  |  |     +--+                                  
  |  |        |  +----- Coelatura_kunenensis **     
  |  |        +--+                               
  |  |           |  +-- Coelatura_gabonensis **     
  |  |           +--+                            
  |  |              +-- Prisodontopsis_mweruensis **
  +--+                                           
     |        +-------- Indonaia_shurtleffiana   
     |        |                                  
     |  +-----+  +----- Indonaia_bonneaudii      
     |  |     |  |                               
     |  |     |  |  +-- Indonaia_rugosa          
     |  |     +--+--+                            
     |  |        |  +-- Pseudoscabies_scobina    
     |  |        |                               
     +--+        |  +-- Indonaia_andersoniana    
        |        +--+                            
        |           +-- Indonaia_caerulea        
        |                                        
        |  +----------- Indochinella_pugio       
        |  |                                     
        |  |     +----- Scabies_phaselus         
        +--+  +--+                               
           |  |  |  +-- Scabies_humilis          
           |  |  +--+                            
           +--+     +-- Scabies_mandarinus       
              |                                  
              |  +----- Radiatula_mouhoti        
              +--+                               
                 |  +-- Radiatula_myitkyinae     
                 +--+                            
                    +-- Radiatula_chaudhurii     

Huang et al., 2019 [Publ 3843], fig. S4

iT tT bi riS oT roT roS iD
6 6 2 3 197 31 4 24.0
                 +----- Leoparreysia_tavoyensis  
  +--------------+                               
  |              |  +-- Leoparreysia_canefrii    
  |              +--+                            
  |                 +-- Leoparreysia_olivacea    
  |                                              
  |           +-------- Coelatura_aegyptiaca[2] **  
  |           |                                  
  |           +-------- Nitia_teretiuscula **       
--+  +--------+                                  
  |  |        |  +----- Coelatura_aegyptiaca[1] **  
  |  |        |  |                               
  |  |        +--+  +-- Coelatura_gabonensis **     
  |  |           |  |                            
  |  |           +--+-- Coelatura_kunenensis **     
  |  |              |                            
  |  |              +-- Prisodontopsis_mweruensis **
  |  |                                           
  +--+        +-------- Indonaia_shurtleffiana   
     |        |                                  
     |        |     +-- Indonaia_andersoniana    
     |  +-----+-----+                            
     |  |     |     +-- Indonaia_caerulea        
     |  |     |                                  
     |  |     |  +----- Indonaia_bonneaudii      
     |  |     +--+                               
     |  |        |  +-- Indonaia_rugosa          
     +--+        +--+                            
        |           +-- Pseudoscabies_scobina    
        |                                        
        |  +----------- Indochinella_pugio       
        |  |                                     
        |  |     +----- Scabies_humilis          
        +--+  +--+                               
           |  |  |  +-- Scabies_phaselus         
           |  |  +--+                            
           +--+     +-- Scabies_mandarinus       
              |                                  
              |  +----- Radiatula_mouhoti        
              +--+                               
                 |  +-- Radiatula_myitkyinae     
                 +--+                            
                    +-- Radiatula_chaudhurii     

Whelan et al., 2011 [Publ 3380], fig. 1 ML

iT tT bi riS oT roT roS iD
5 5 3 3 32 5 3 24.0
  +----------------- Leoparreysia_tavoyensis      
  |                                               
--+              +-- Radiatula_chaudhurii         
  |  +-----------+                                
  |  |           +-- Indochinella_pugio           
  +--+                                            
     |  +----------- Nitia_teretiuscula **           
     +--+                                         
        |  +-------- Coelatura_aegyptiaca[2] **      
        +--+                                      
           |  +----- Coelatura_aegyptiaca[1] **      
           +--+                                   
              |  +-- Coelatura_gabonensis **         
              +--+                                
                 +-- Prisodontopsis_aviculaeformis **

Whelan et al., 2011 [Publ 3380], fig. 1 MP

iT tT bi riS oT roT roS iD
5 5 2 3 32 5 3 24.0
  +-------------- Leoparreysia_tavoyensis      
  |                                            
--+           +-- Radiatula_chaudhurii         
  |  +--------+                                
  |  |        +-- Indochinella_pugio           
  +--+                                         
     |  +-------- Nitia_teretiuscula **           
     |  |                                      
     +--+  +----- Coelatura_aegyptiaca[1] **      
        |  |                                   
        +--+----- Coelatura_aegyptiaca[2] **      
           |                                   
           |  +-- Coelatura_gabonensis **         
           +--+                                
              +-- Prisodontopsis_aviculaeformis **

Bolotov et al., 2022 [Publ 4239], fig. 2

iT tT bi riS oT roT roS iD
4 4 2 3 54 50 4 23.0
                                   +----------- Parreysia_keralaensis   
                                +--+                                    
                                |  |  +-------- Parreysia_plagiosoma    
                                |  +--+                                 
                                |     |  +----- Parreysia_nagpoorensis  
                                |     +--+                              
                                |        |  +-- Parreysia_corrugata     
  +-----------------------------+        +--+                           
  |                             |           +-- Parreysia_rakhinensis   
  |                             |                                       
  |                             |           +-- Leoparreysia_burmana    
  |                             |     +-----+                           
  |                             |     |     +-- Leoparreysia_whitteni   
  |                             +-----+                                 
  |                                   |  +----- Leoparreysia_olivacea   
  |                                   +--+                              
  |                                      |  +-- Leoparreysia_canefrii   
  |                                      +--+                           
--+                                         +-- Leoparreysia_tavoyensis 
  |                                                                     
  |                                   +-------- Nitia_teretiuscula **      
  |  +--------------------------------+                                 
  |  |                                |  +----- Coelatura_aegyptiaca **    
  |  |                                +--+                              
  |  |                                   |  +-- Coelatura_kunenensis **    
  |  |                                   +--+                           
  |  |                                      +-- Nyassunio_nyassaensis **   
  |  |                                                                  
  |  |                                      +-- Indonaia_cylindrica     
  |  |                             +--------+                           
  +--+                             |        +-- Indonaia_gratiosa       
     |                             |                                    
     |  +--------------------------+     +----- Indonaia_andersoniana   
     |  |                          |  +--+                              
     |  |                          |  |  |  +-- Indonaia_caerulea       
     |  |                          |  |  +--+                           
     |  |                          +--+     +-- Indonaia_subclathrata   
     |  |                             |                                 
     |  |                             |  +----- Indonaia_shurtleffiana  
     |  |                             +--+                              
     +--+                                |  +-- Indonaia_bonneaudii     
        |                                +--+                           
        |                                   +-- Indonaia_rugosa         
        |                                                               
        |  +----------------------------------- Indochinella_pugio      
        |  |                                                            
        |  |                       +----------- Radiatula_crispisulcata 
        |  |                       |                                    
        +--+  +--------------------+        +-- Radiatula_chaudhurii    
           |  |                    |  +-----+                           
           |  |                    |  |     +-- Radiatula_myitkyinae    
           |  |                    +--+                                 
           |  |                       |  +----- Radiatula_mouhoti       
           +--+                       +--+                              
              |                          |  +-- Radiatula_chindwinensis 
              |                          +--+                           
              |                             +-- Radiatula_myitthanensis 
              |                                                         
              |  +----------------------------- Scabiellus_songkramensis
              +--+                                                      
                 |  +-------------------------- Harmandia_somboriensis  
                 +--+                                                   
                    |  +----------------------- Unionetta_fabagina      
                    +--+                                                
                       |  +-------------------- Scabies_crispata        
                       +--+                                             
                          |  +----------------- Scabies_mandarinus      
                          +--+                                          
                             |  +-------------- Scabies_nucleus         
                             +--+                                       
                                |  +----------- Scabies_pilata          
                                +--+                                    
                                   |  +-------- Scabies_humilis         
                                   +--+                                 
                                      |  +----- Scabies_anceps          
                                      +--+                              
                                         |  +-- Scabies_phaselus        
                                         +--+                           
                                            +-- Scabies_scobinatus      

Bolotov et al., 2022 [Publ 4239], fig. S1

iT tT bi riS oT roT roS iD
4 4 2 3 54 50 4 23.0
                                      +-- Leoparreysia_burmana    
                                +-----+                           
                                |     +-- Leoparreysia_whitteni   
  +-----------------------------+                                 
  |                             |  +----- Leoparreysia_olivacea   
  |                             +--+                              
  |                                |  +-- Leoparreysia_canefrii   
  |                                +--+                           
  |                                   +-- Leoparreysia_tavoyensis 
  |                                                               
  |                             +-------- Nitia_teretiuscula **      
--+  +--------------------------+                                 
  |  |                          |  +----- Coelatura_aegyptiaca **    
  |  |                          +--+                              
  |  |                             |  +-- Coelatura_kunenensis **    
  |  |                             +--+                           
  |  |                                +-- Nyassunio_nyassaensis **   
  |  |                                                            
  |  |                 +----------------- Indonaia_cylindrica     
  |  |                 |                                          
  +--+  +--------------+  +-------------- Indonaia_gratiosa       
     |  |              |  |                                       
     |  |              +--+  +----------- Indonaia_andersoniana   
     |  |                 |  |                                    
     |  |                 +--+        +-- Indonaia_caerulea       
     |  |                    |  +-----+                           
     |  |                    |  |     +-- Indonaia_subclathrata   
     |  |                    +--+                                 
     |  |                       |  +----- Indonaia_rugosa         
     +--+                       +--+                              
        |                          |  +-- Indonaia_bonneaudii     
        |                          +--+                           
        |                             +-- Indonaia_shurtleffiana  
        |                                                         
        |  +----------------------------- Indochinella_pugio      
        |  |                                                      
        |  |                    +-------- Radiatula_crispisulcata 
        |  |                    |                                 
        |  |  +-----------------+     +-- Radiatula_chaudhurii    
        +--+  |                 |  +--+                           
           |  |                 |  |  +-- Radiatula_myitkyinae    
           |  |                 +--+                              
           |  |                    |  +-- Radiatula_myitthanensis 
           |  |                    |  |                           
           +--+                    +--+-- Radiatula_mouhoti       
              |                       |                           
              |                       +-- Radiatula_chindwinensis 
              |                                                   
              |  +----------------------- Scabiellus_songkramensis
              |  |                                                
              +--+  +-------------------- Harmandia_somboriensis  
                 |  |                                             
                 +--+  +----------------- Unionetta_fabagina      
                    |  |                                          
                    +--+              +-- Scabies_crispata        
                       |  +-----------+                           
                       |  |           +-- Scabies_mandarinus      
                       +--+                                       
                          |  +----------- Scabies_nucleus         
                          |  |                                    
                          +--+        +-- Scabies_humilis         
                             |  +-----+                           
                             |  |     +-- Scabies_pilata          
                             +--+                                 
                                |  +----- Scabies_phaselus        
                                +--+                              
                                   |  +-- Scabies_anceps          
                                   +--+                           
                                      +-- Scabies_scobinatus      

Bolotov et al., 2022 [Publ 4239], fig. S2

iT tT bi riS oT roT roS iD
4 4 2 3 54 50 4 23.0
                                +----------- Leoparreysia_burmana    
  +-----------------------------+                                    
  |                             |  +-------- Leoparreysia_whitteni   
  |                             +--+                                 
  |                                |  +----- Leoparreysia_olivacea   
  |                                +--+                              
  |                                   |  +-- Leoparreysia_canefrii   
  |                                   +--+                           
  |                                      +-- Leoparreysia_tavoyensis 
  |                                                                  
--+                                +-------- Nitia_teretiuscula **      
  |  +-----------------------------+                                 
  |  |                             |  +----- Coelatura_aegyptiaca **    
  |  |                             +--+                              
  |  |                                |  +-- Coelatura_kunenensis **    
  |  |                                +--+                           
  |  |                                   +-- Nyassunio_nyassaensis **   
  |  |                                                               
  |  |                                   +-- Indonaia_cylindrica     
  |  |                          +--------+                           
  +--+                          |        +-- Indonaia_gratiosa       
     |                          |                                    
     |  +-----------------------+     +----- Indonaia_shurtleffiana  
     |  |                       |  +--+                              
     |  |                       |  |  |  +-- Indonaia_bonneaudii     
     |  |                       |  |  +--+                           
     |  |                       +--+     +-- Indonaia_rugosa         
     |  |                          |                                 
     |  |                          |  +----- Indonaia_andersoniana   
     |  |                          +--+                              
     +--+                             |  +-- Indonaia_caerulea       
        |                             +--+                           
        |                                +-- Indonaia_subclathrata   
        |                                                            
        |  +-------------------------------- Indochinella_pugio      
        |  |                                                         
        |  |                    +----------- Radiatula_crispisulcata 
        |  |                    |                                    
        |  |  +-----------------+        +-- Radiatula_chaudhurii    
        +--+  |                 |  +-----+                           
           |  |                 |  |     +-- Radiatula_myitkyinae    
           |  |                 +--+                                 
           |  |                    |  +----- Radiatula_mouhoti       
           |  |                    +--+                              
           +--+                       |  +-- Radiatula_myitthanensis 
              |                       +--+                           
              |                          +-- Radiatula_chindwinensis 
              |                                                      
              |  +-------------------------- Scabiellus_songkramensis
              |  |                                                   
              +--+  +----------------------- Harmandia_somboriensis  
                 |  |                                                
                 +--+  +-------------------- Unionetta_fabagina      
                    |  |                                             
                    +--+  +----------------- Scabies_crispata        
                       |  |                                          
                       +--+              +-- Scabies_humilis         
                          |  +-----------+                           
                          |  |           +-- Scabies_mandarinus      
                          +--+                                       
                             |  +----------- Scabies_nucleus         
                             +--+                                    
                                |  +-------- Scabies_pilata          
                                +--+                                 
                                   |  +----- Scabies_anceps          
                                   +--+                              
                                      |  +-- Scabies_phaselus        
                                      +--+                           
                                         +-- Scabies_scobinatus      

Bolotov et al., 2022 [Publ 4252], fig. 2

iT tT bi riS oT roT roS iD
4 4 2 3 57 53 4 23.0
                                      +-- Leoparreysia_burmana    
                                +-----+                           
                                |     +-- Leoparreysia_whitteni   
  +-----------------------------+                                 
  |                             |  +----- Leoparreysia_olivacea   
  |                             +--+                              
  |                                |  +-- Leoparreysia_canefrii   
  |                                +--+                           
  |                                   +-- Leoparreysia_tavoyensis 
  |                                                               
  |                             +-------- Nitia_teretiuscula **      
--+  +--------------------------+                                 
  |  |                          |  +----- Coelatura_aegyptiaca **    
  |  |                          +--+                              
  |  |                             |  +-- Nyassunio_nyassaensis **   
  |  |                             +--+                           
  |  |                                +-- Coelatura_kunenensis **    
  |  |                                                            
  |  |                 +----------------- Indonaia_cylindrica     
  |  |                 |                                          
  +--+  +--------------+  +-------------- Indonaia_gratiosa       
     |  |              |  |                                       
     |  |              +--+  +----------- Indonaia_andersoniana   
     |  |                 |  |                                    
     |  |                 +--+        +-- Indonaia_caerulea       
     |  |                    |  +-----+                           
     |  |                    |  |     +-- Indonaia_subclathrata   
     |  |                    +--+                                 
     |  |                       |  +----- Indonaia_rugosa         
     +--+                       +--+                              
        |                          |  +-- Indonaia_bonneaudii     
        |                          +--+                           
        |                             +-- Indonaia_shurtleffiana  
        |                                                         
        |  +----------------------------- Indochinella_pugio      
        |  |                                                      
        |  |                 +----------- Radiatula_crispisulcata 
        |  |                 |                                    
        |  |                 |     +----- Radiatula_chaudhurii    
        +--+  +--------------+  +--+                              
           |  |              |  |  |  +-- Radiatula_myitkyinae    
           |  |              |  |  +--+                           
           |  |              +--+     +-- Radiatula_kachinensis   
           |  |                 |                                 
           |  |                 |  +----- Radiatula_chindwinensis 
           +--+                 +--+                              
              |                    |  +-- Radiatula_myitthanensis 
              |                    +--+                           
              |                       +-- Radiatula_mouhoti       
              |                                                   
              |  +----------------------- Scabiellus_songkramensis
              |  |                                                
              +--+  +-------------------- Harmandia_somboriensis  
                 |  |                                             
                 +--+  +----------------- Unionetta_fabagina      
                    |  |                                          
                    +--+              +-- Scabies_crispata        
                       |  +-----------+                           
                       |  |           +-- Scabies_mandarinus      
                       +--+                                       
                          |  +----------- Scabies_nucleus         
                          |  |                                    
                          +--+        +-- Scabies_humilis         
                             |  +-----+                           
                             |  |     +-- Scabies_pilata          
                             +--+                                 
                                |  +----- Scabies_phaselus        
                                +--+                              
                                   |  +-- Scabies_anceps          
                                   +--+                           
                                      +-- Scabies_scobinatus      

Bolotov et al., 2022 [Publ 4252], fig. S1

iT tT bi riS oT roT roS iD
4 4 0 3 57 53 4 25.0
                                         +-- Leoparreysia_burmana    
                                         |                           
                                         +-- Leoparreysia_canefrii   
                                         |                           
  +--------------------------------------+-- Leoparreysia_olivacea   
  |                                      |                           
  |                                      +-- Leoparreysia_tavoyensis 
  |                                      |                           
  |                                      +-- Leoparreysia_whitteni   
  |                                                                  
  |                                      +-- Coelatura_aegyptiaca **    
  |                                      |                           
  |                                      +-- Coelatura_kunenensis **    
--+  +-----------------------------------+                           
  |  |                                   +-- Nitia_teretiuscula **      
  |  |                                   |                           
  |  |                                   +-- Nyassunio_nyassaensis **   
  |  |                                                               
  |  |                                   +-- Indonaia_cylindrica     
  |  |                          +--------+                           
  |  |                          |        +-- Indonaia_gratiosa       
  |  |                          |                                    
  +--+  +-----------------------+     +----- Indonaia_shurtleffiana  
     |  |                       |  +--+                              
     |  |                       |  |  |  +-- Indonaia_bonneaudii     
     |  |                       |  |  +--+                           
     |  |                       +--+     +-- Indonaia_rugosa         
     |  |                          |                                 
     |  |                          |  +----- Indonaia_andersoniana   
     |  |                          +--+                              
     |  |                             |  +-- Indonaia_caerulea       
     +--+                             +--+                           
        |                                +-- Indonaia_subclathrata   
        |                                                            
        |  +-------------------------------- Indochinella_pugio      
        |  |                                                         
        |  |                          +----- Radiatula_chaudhurii    
        |  |                    +-----+                              
        |  |                    |     |  +-- Radiatula_kachinensis   
        |  |                    |     +--+                           
        +--+  +-----------------+        +-- Radiatula_myitkyinae    
           |  |                 |                                    
           |  |                 |  +-------- Radiatula_crispisulcata 
           |  |                 +--+                                 
           |  |                    |  +----- Radiatula_mouhoti       
           |  |                    +--+                              
           +--+                       |  +-- Radiatula_myitthanensis 
              |                       +--+                           
              |                          +-- Radiatula_chindwinensis 
              |                                                      
              |  +-------------------------- Scabiellus_songkramensis
              |  |                                                   
              +--+  +----------------------- Harmandia_somboriensis  
                 |  |                                                
                 +--+  +-------------------- Unionetta_fabagina      
                    |  |                                             
                    +--+  +----------------- Scabies_crispata        
                       |  |                                          
                       +--+              +-- Scabies_humilis         
                          |  +-----------+                           
                          |  |           +-- Scabies_mandarinus      
                          +--+                                       
                             |  +----------- Scabies_nucleus         
                             +--+                                    
                                |  +-------- Scabies_pilata          
                                +--+                                 
                                   |  +----- Scabies_anceps          
                                   +--+                              
                                      |  +-- Scabies_phaselus        
                                      +--+                           
                                         +-- Scabies_scobinatus      

Bolotov et al., 2024 [Publ 4369], fig. S1

iT tT bi riS oT roT roS iD
4 4 2 3 59 58 4 23.0
  +-------------------------------------------- Assamnaia_involuta      
  |                                                                     
  |                                   +-------- Nitia_teretiuscula **      
  |  +--------------------------------+                                 
  |  |                                |  +----- Coelatura_aegyptiaca **    
  |  |                                +--+                              
--+  |                                   |  +-- Nyassunio_nyassaensis **   
  |  |                                   +--+                           
  |  |                                      +-- Coelatura_kunenensis **    
  |  |                                                                  
  |  |                    +-------------------- Pseudoscabies_scobina   
  |  |                    |                                             
  |  |  +-----------------+                 +-- Indonaia_cylindrica     
  +--+  |                 |  +--------------+                           
     |  |                 |  |              +-- Indonaia_gratiosa       
     |  |                 +--+                                          
     |  |                    |  +-------------- Indonaia_corbis         
     |  |                    |  |                                       
     |  |                    +--+  +----------- Indonaia_andersoniana   
     |  |                       |  |                                    
     |  |                       +--+        +-- Indonaia_caerulea       
     |  |                          |  +-----+                           
     |  |                          |  |     +-- Indonaia_subclathrata   
     +--+                          +--+                                 
        |                             |  +----- Indonaia_shurtleffiana  
        |                             +--+                              
        |                                |  +-- Indonaia_bonneaudii     
        |                                +--+                           
        |                                   +-- Indonaia_rugosa         
        |                                                               
        |  +----------------------------------- Indochinella_pugio      
        |  |                                                            
        |  |                                +-- Radiatula_myitkyinae    
        |  |                    +-----------+                           
        |  |                    |           +-- Radiatula_kachinensis   
        +--+  +-----------------+                                       
           |  |                 |  +----------- Radiatula_chaudhurii    
           |  |                 +--+                                    
           |  |                    |  +-------- Radiatula_crispisulcata 
           |  |                    +--+                                 
           |  |                       |  +----- Radiatula_mouhoti       
           +--+                       +--+                              
              |                          |  +-- Radiatula_myitthanensis 
              |                          +--+                           
              |                             +-- Radiatula_chindwinensis 
              |                                                         
              |  +----------------------------- Scabiellus_songkramensis
              +--+                                                      
                 |  +-------------------------- Harmandia_somboriensis  
                 +--+                                                   
                    |  +----------------------- Unionetta_fabagina      
                    +--+                                                
                       |  +-------------------- Scabies_crispata        
                       +--+                                             
                          |  +----------------- Scabies_mandarinus      
                          +--+                                          
                             |  +-------------- Scabies_nucleus         
                             +--+                                       
                                |  +----------- Scabies_pilata          
                                +--+                                    
                                   |  +-------- Scabies_humilis         
                                   +--+                                 
                                      |  +----- Scabies_anceps          
                                      +--+                              
                                         |  +-- Scabies_phaselus        
                                         +--+                           
                                            +-- Scabies_scobinatus      

Pfeiffer et al., 2018 [Publ 3827], fig. 2

iT tT bi riS oT roT roS iD
3 3 1 3 26 25 4 24.0
                                      +-- Leoparreysia_tavoyensis      
  +-----------------------------------+                                
  |                                   +-- Leoparreysia_olivacea        
  |                                                                    
  |                                +----- Nitia_teretiuscula **           
--+  +-----------------------------+                                   
  |  |                             |  +-- Coelatura_aegyptiaca **         
  |  |                             +--+                                
  |  |                                +-- Prisodontopsis_aviculaeformis **
  +--+                                                                 
     |                             +----- Indonaia_bonneaudii          
     |  +--------------------------+                                   
     |  |                          |  +-- Indonaia_andersoniana        
     |  |                          +--+                                
     +--+                             +-- Indonaia_caerulea            
        |                                                              
        |  +----------------------------- Indochinella_pugio           
        |  |                                                           
        +--+                          +-- Radiatula_chaudhurii         
           |  +-----------------------+                                
           |  |                       +-- Radiatula_myitkyinae         
           +--+                                                        
              |  +----------------------- Scabiellus_songkramensis     
              |  |                                                     
              +--+  +-------------------- Unionetta_fabagina[2]        
                 |  |                                                  
                 +--+  +----------------- Harmandia_somboriensis       
                    |  |                                               
                    +--+  +-------------- Unionetta_fabagina[1]        
                       |  |                                            
                       +--+           +-- Scabies_crispata             
                          |  +--------+                                
                          |  |        +-- Scabies_mandarinus           
                          +--+                                         
                             |  +-------- Scabies_nucleus              
                             |  |                                      
                             +--+     +-- Scabies_pilata               
                                |  +--+                                
                                |  |  +-- Scabies_humilis              
                                +--+                                   
                                   |  +-- Scabies_phaselus             
                                   |  |                                
                                   +--+-- Scabies_anceps               
                                      |                                
                                      +-- Scabies_scobinatus           

Bolotov et al., 2018 [Publ 3826], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 83 22 4 --
                 +----- Leoparreysia_olivacea  
  +--------------+                             
  |              |  +-- Leoparreysia_canefrii  
  |              +--+                          
  |                 +-- Leoparreysia_tavoyensis
  |                                            
--+                 +-- Coelatura_aegyptiaca **   
  |  +--------------+                          
  |  |              +-- Nitia_teretiuscula **     
  |  |                                         
  |  |        +-------- Indonaia_shurtleffiana 
  +--+  +-----+                                
     |  |     |  +----- Pseudoscabies_scobina  
     |  |     +--+                             
     |  |        |  +-- Indonaia_caerulea      
     |  |        +--+                          
     +--+           +-- Indonaia_andersoniana  
        |                                      
        |  +----------- Indochinella_pugio     
        |  |                                   
        |  |     +----- Radiatula_myitkyinae   
        +--+  +--+                             
           |  |  |  +-- Radiatula_mouhoti      
           |  |  +--+                          
           +--+     +-- Radiatula_chaudhurii   
              |                                
              |  +----- Scabies_mandarinus     
              +--+                             
                 |  +-- Scabies_humilis        
                 +--+                          
                    +-- Scabies_phaselus       

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iT tT bi riS oT roT roS iD
2 2 0 2 55 8 3 --
              +-- Leoparreysia_tavoyensis      
  +-----------+                                
  |           +-- Leoparreysia_olivacea        
  |                                            
--+           +-- Coelatura_choziensis **         
  |  +--------+                                
  |  |        +-- Prisodontopsis_aviculaeformis **
  +--+                                         
     |  +-------- Indonaia_caerulea            
     +--+                                      
        |  +----- Indochinella_pugio           
        +--+                                   
           |  +-- Scabies_phaselus             
           +--+                                
              +-- Harmandia_somboriensis       

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iT tT bi riS oT roT roS iD
2 2 0 2 55 8 3 --
              +-- Leoparreysia_tavoyensis      
  +-----------+                                
  |           +-- Leoparreysia_olivacea        
  |                                            
--+           +-- Coelatura_choziensis **         
  |  +--------+                                
  |  |        +-- Prisodontopsis_aviculaeformis **
  +--+                                         
     |  +-------- Indonaia_caerulea            
     +--+                                      
        |  +----- Indochinella_pugio           
        +--+                                   
           |  +-- Scabies_phaselus             
           +--+                                
              +-- Harmandia_somboriensis       

Bolotov et al., 2019 [Publ 3974], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 16 11 2 --
                 +-- Leoparreysia_whitteni  
           +-----+                          
           |     +-- Leoparreysia_burmana   
  +--------+                                
  |        |  +----- Leoparreysia_olivacea  
  |        +--+                             
  |           |  +-- Leoparreysia_tavoyensis
  |           +--+                          
--+              +-- Leoparreysia_canefrii  
  |                                         
  |              +-- Nitia_teretiuscula **     
  |  +-----------+                          
  |  |           +-- Coelatura_aegyptiaca **   
  +--+                                      
     |  +----------- Indochinella_pugio     
     |  |                                   
     +--+        +-- Radiatula_myitkyinae   
        |  +-----+                          
        |  |     +-- Radiatula_chaudhurii   
        +--+                                
           |  +----- Radiatula_chindwinensis
           +--+                             
              |  +-- Radiatula_mouhoti      
              +--+                          
                 +-- Radiatula_myitthanensis

Bolotov et al., 2020 [Publ 4100], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 118 39 4 --
                          +----------- Parreysia_keralaensis    
                       +--+                                     
                       |  |  +-------- Parreysia_plagiosoma     
                       |  +--+                                  
                       |     |  +----- Parreysia_nagpoorensis[2]
                       |     +--+                               
                       |        |  +-- Parreysia_nagpoorensis[1]
  +--------------------+        +--+                            
  |                    |           +-- Parreysia_rakhinensis    
  |                    |                                        
  |                    |           +-- Leoparreysia_burmana     
  |                    |     +-----+                            
  |                    |     |     +-- Leoparreysia_whitteni    
  |                    +-----+                                  
  |                          |  +----- Leoparreysia_tavoyensis  
  |                          +--+                               
--+                             |  +-- Leoparreysia_olivacea    
  |                             +--+                            
  |                                +-- Leoparreysia_canefrii    
  |                                                             
  |                                +-- Coelatura_aegyptiaca **     
  |  +-----------------------------+                            
  |  |                             +-- Nitia_teretiuscula **       
  |  |                                                          
  |  |                    +----------- Indonaia_shurtleffiana   
  |  |  +-----------------+                                     
  +--+  |                 |  +-------- Pseudoscabies_scobina    
     |  |                 +--+                                  
     |  |                    |  +----- Indonaia_andersoniana    
     |  |                    +--+                               
     |  |                       |  +-- Indonaia_subclathrata    
     +--+                       +--+                            
        |                          +-- Indonaia_caerulea        
        |                                                       
        |  +-------------------------- Indochinella_pugio       
        |  |                                                    
        |  |                 +-------- Radiatula_crispisulcata  
        |  |  +--------------+                                  
        +--+  |              |  +----- Radiatula_mouhoti        
           |  |              +--+                               
           |  |                 |  +-- Radiatula_chaudhurii     
           |  |                 +--+                            
           +--+                    +-- Radiatula_myitkyinae     
              |                                                 
              |  +-------------------- Scabiellus_songkramensis 
              |  |                                              
              |  |                 +-- Unionetta_fabagina       
              +--+  +--------------+                            
                 |  |              +-- Harmandia_somboriensis   
                 |  |                                           
                 +--+  +-------------- Scabies_nucleus          
                    |  |                                        
                    |  |           +-- Scabies_crispata         
                    +--+  +--------+                            
                       |  |        +-- Scabies_mandarinus       
                       |  |                                     
                       +--+        +-- Scabies_humilis          
                          |  +-----+                            
                          |  |     +-- Scabies_pilata           
                          +--+                                  
                             |  +----- Scabies_anceps           
                             +--+                               
                                |  +-- Scabies_phaselus         
                                +--+                            
                                   +-- Scabies_scobinatus       

Bolotov et al., 2020 [Publ 4100], fig. S1

iT tT bi riS oT roT roS iD
2 2 0 2 112 38 4 --
                                +----------- Parreysia_keralaensis    
                             +--+                                     
                             |  |  +-------- Parreysia_nagpoorensis[2]
                             |  +--+                                  
                             |     |  +----- Parreysia_nagpoorensis[1]
                             |     +--+                               
                             |        |  +-- Parreysia_rakhinensis    
  +--------------------------+        +--+                            
  |                          |           +-- Parreysia_plagiosoma     
  |                          |                                        
  |                          |           +-- Leoparreysia_burmana     
  |                          |     +-----+                            
  |                          |     |     +-- Leoparreysia_whitteni    
  |                          +-----+                                  
  |                                |  +----- Leoparreysia_olivacea    
  |                                +--+                               
--+                                   |  +-- Leoparreysia_tavoyensis  
  |                                   +--+                            
  |                                      +-- Leoparreysia_canefrii    
  |                                                                   
  |                                      +-- Coelatura_aegyptiaca **     
  |  +-----------------------------------+                            
  |  |                                   +-- Nitia_teretiuscula **       
  |  |                                                                
  |  |                             +-------- Indonaia_shurtleffiana   
  +--+  +--------------------------+                                  
     |  |                          |  +----- Indonaia_andersoniana    
     |  |                          +--+                               
     |  |                             |  +-- Indonaia_caerulea        
     |  |                             +--+                            
     +--+                                +-- Indonaia_subclathrata    
        |                                                             
        |  +-------------------------------- Indochinella_pugio       
        |  |                                                          
        |  |                          +----- Radiatula_mouhoti        
        +--+  +-----------------------+                               
           |  |                       |  +-- Radiatula_myitkyinae     
           |  |                       +--+                            
           +--+                          +-- Radiatula_chaudhurii     
              |                                                       
              |  +-------------------------- Radiatula_crispisulcata  
              |  |                                                    
              +--+  +----------------------- Scabiellus_songkramensis 
                 |  |                                                 
                 +--+  +-------------------- Harmandia_somboriensis   
                    |  |                                              
                    +--+  +----------------- Unionetta_fabagina       
                       |  |                                           
                       +--+  +-------------- Scabies_mandarinus       
                          |  |                                        
                          +--+  +----------- Scabies_crispata         
                             |  |                                     
                             +--+  +-------- Scabies_nucleus          
                                |  |                                  
                                +--+     +-- Scabies_humilis          
                                   |  +--+                            
                                   |  |  +-- Scabies_pilata           
                                   +--+                               
                                      |  +-- Scabies_phaselus         
                                      |  |                            
                                      +--+-- Scabies_scobinatus       
                                         |                            
                                         +-- Scabies_anceps           

Graf & Cummings, 2006 [Publ 1968], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 46 0 0 --
-- Coelatura_aegyptiaca

Graf, 2000 [Publ 2647], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 17 1 1 --
-- Grandidieria

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 33 0 0 --
-- Coelatura_aegyptiaca

Hoeh et al., 2001 [Publ 2652], fig. 14.2

iT tT bi riS oT roT roS iD
1 1 0 1 30 0 0 --
-- Coelatura_aegyptiaca

Hoeh et al., 2001 [Publ 2652], fig. 14.4

iT tT bi riS oT roT roS iD
1 1 0 1 30 0 0 --
-- Coelatura_aegyptiaca

Hoeh et al., 2002 [Publ 2653], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 30 0 0 --
-- Coelatura_aegyptiaca

Roe & Hoeh, 2003 [Publ 2655], fig. 4.2

iT tT bi riS oT roT roS iD
1 1 0 1 30 0 0 --
-- Coelatura_aegyptiaca

Roe & Hoeh, 2003 [Publ 2655], fig. 4.3

iT tT bi riS oT roT roS iD
1 1 0 1 30 0 0 --
-- Coelatura_aegyptiaca

Hoeh et al., 2009 [Publ 2716], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 46 0 0 --
-- Coelatura_aegyptiaca

Hoeh et al., 2009 [Publ 2716], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 45 0 0 --
-- Coelatura_aegyptiaca

Hoeh et al., 2009 [Publ 2716], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 45 0 0 --
-- Coelatura_aegyptiaca

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 34 1 1 --
  +-- Lamellidens_savadiensis
--+                          
  +-- Coelatura_aegyptiaca **   

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 39 7 3 --
           +-- Leoparreysia_olivacea  
  +--------+                          
  |        +-- Leoparreysia_tavoyensis
--+                                   
  |  +-------- Coelatura_aegyptiaca **   
  +--+                                
     |  +----- Indochinella_pugio     
     +--+                             
        |  +-- Radiatula_chaudhurii   
        +--+                          
           +-- Scabies_mandarinus     

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iT tT bi riS oT roT roS iD
1 1 0 1 80 3 3 --
  +----- Leoparreysia_tavoyensis
--+                             
  |  +-- Coelatura_aegyptiaca **   
  +--+                          
     +-- Indochinella_pugio     

Araujo et al., 2017 [Publ 3801], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 13 1 1 --
  +-- Lamellidens_savadiensis
--+                          
  +-- Coelatura_aegyptiaca **   

Konopleva et al., 2017 [Publ 3807], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 30 10 3 --
           +-- Leoparreysia_tavoyensis
  +--------+                          
  |        +-- Leoparreysia_olivacea  
--+                                   
  |  +-------- Coelatura_aegyptiaca **   
  +--+                                
     |  +----- Indochinella_pugio     
     +--+                             
        |  +-- Radiatula_chaudhurii   
        +--+                          
           +-- Scabies_mandarinus     

Jeratthitikul et al., 2019 [Publ 3922], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 22 22 4 --
  +-------------------------------------------- Parreysia_keralaensis   
  |                                                                     
--+  +----------------------------------------- Coelatura_aegyptiaca **    
  |  |                                                                  
  |  |                                   +----- Pseudoscabies_scobina   
  +--+  +--------------------------------+                              
     |  |                                |  +-- Indonaia_caerulea       
     |  |                                +--+                           
     |  |                                   +-- Indonaia_andersoniana   
     +--+                                                               
        |  +----------------------------------- Indochinella_pugio      
        |  |                                                            
        |  |                             +----- Radiatula_mouhoti       
        +--+  +--------------------------+                              
           |  |                          |  +-- Radiatula_myitkyinae    
           |  |                          +--+                           
           +--+                             +-- Radiatula_chaudhurii    
              |                                                         
              |  +----------------------------- Scabiellus_songkramensis
              +--+                                                      
                 |  +-------------------------- Harmandia_somboriensis  
                 +--+                                                   
                    |  +----------------------- Unionetta_fabagina      
                    +--+                                                
                       |  +-------------------- Scabies_crispata        
                       +--+                                             
                          |  +----------------- Scabies_mandarinus      
                          +--+                                          
                             |  +-------------- Scabies_pilata          
                             +--+                                       
                                |  +----------- Scabies_nucleus         
                                +--+                                    
                                   |  +-------- Scabies_humilis         
                                   +--+                                 
                                      |  +----- Scabies_scobinatus      
                                      +--+                              
                                         |  +-- Scabies_phaselus        
                                         +--+                           
                                            +-- Scabies_anceps          

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iT tT bi riS oT roT roS iD
1 1 0 1 121 2 2 --
  +-- Coelatura_aegyptiaca **   
  |                          
--+-- Leoparreysia_tavoyensis
  |                          
  +-- Radiatula_mouhoti      

Chen et al., 2023 [Publ 4302], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 39 2 2 --
  +-- Coelatura_aegyptiaca **  
  |                         
--+-- Indonaia_andersoniana 
  |                         
  +-- Parreysia_nagpoorensis

Chen et al., 2023 [Publ 4302], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 39 2 2 --
  +----- Parreysia_nagpoorensis
--+                            
  |  +-- Coelatura_aegyptiaca **  
  +--+                         
     +-- Indonaia_andersoniana 

Wu et al., 2021 [Publ 4366], fig. 2, I

iT tT bi riS oT roT roS iD
1 1 0 1 90 12 3 --
              +----- Leoparreysia_olivacea  
  +-----------+                             
  |           |  +-- Leoparreysia_canefrii  
  |           +--+                          
--+              +-- Leoparreysia_tavoyensis
  |                                         
  |  +-------------- Coelatura_aegyptiaca **   
  +--+                                      
     |  +----------- Indonaia_andersoniana  
     +--+                                   
        |  +-------- Scabies_crispata       
        +--+                                
           |  +----- Radiatula_myitkyinae   
           +--+                             
              |  +-- Radiatula_mouhoti      
              +--+                          
                 +-- Radiatula_chaudhurii   

Wu et al., 2021 [Publ 4366], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 90 12 3 --
              +----- Leoparreysia_olivacea  
  +-----------+                             
  |           |  +-- Leoparreysia_canefrii  
  |           +--+                          
--+              +-- Leoparreysia_tavoyensis
  |                                         
  |  +-------------- Coelatura_aegyptiaca **   
  +--+                                      
     |  +----------- Indonaia_andersoniana  
     +--+                                   
        |  +-------- Radiatula_myitkyinae   
        +--+                                
           |  +----- Scabies_crispata       
           +--+                             
              |  +-- Radiatula_mouhoti      
              +--+                          
                 +-- Radiatula_chaudhurii   

Page updated: 10:02:44 Wed 05 Jun 2024

 
NSF icon MUSSEL icon
"Making the world a better place, one mollusk at a time."