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Page last updated
Wed 18 Jan 2023

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Parreysiinae | Lamellidentini

genus Lamellidens Simpson, 1900: cladomics
     13 valid species

Cladomics Summary

number of species = 13

analyzed in 35 cladograms in 22 publications
maximum number of terminal species in an analysis = 10, mean = 3.97

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 19
maximum number of included species = 10, mean = 6.47
proportion of cladograms that support basic monophyly = 0.89


Bolotov et al., 2022, fig. 2

iT tT bi riS oT roT roS iD
10 10 7 10 51 6 1 0.0

*** Top Monophyly Cladogram *** *** Top Sister-Group Cladogram ***

        +----------- Trapezidens_mogaungensis   
        |                                       
  +-----+  +-------- Trapezidens_exolescens     
  |     |  |                                    
  |     +--+  +----- Trapezidens_scutum         
  |        |  |                                 
  |        +--+  +-- Trapezidens_angustior      
  |           |  |                              
--+           +--+-- Trapezidens_dolichorhynchus
  |              |                              
  |              +-- Trapezidens_yeti           
  |                                             
  |  +-------------- Lamellidens_marginalis **     
  |  |                                          
  |  |  +----------- Lamellidens_ferrugineus **    
  +--+  |                                       
     |  |  +-------- Lamellidens_generosus **      
     |  +--+                                    
     |  |  |  +----- Lamellidens_mainwaringi **    
     +--+  +--+                                 
        |     |  +-- Lamellidens_lamellatus **     
        |     +--+                              
        |        +-- Lamellidens_jenkinsianus **   
        |                                       
        |  +-------- Lamellidens_nongyangensis **  
        +--+                                    
           |  +----- Lamellidens_savadiensis **    
           +--+                                 
              |  +-- Lamellidens_corrianus **      
              +--+                              
                 +-- Lamellidens_chindwinensis **  

Bolotov et al., 2022, fig. S1

iT tT bi riS oT roT roS iD
10 10 7 10 51 6 1 6.0
     +-------------- Trapezidens_mogaungensis   
  +--+                                          
  |  |  +----------- Trapezidens_scutum         
  |  +--+                                       
  |     |  +-------- Trapezidens_exolescens     
  |     +--+                                    
  |        |  +----- Trapezidens_yeti           
  |        +--+                                 
  |           |  +-- Trapezidens_dolichorhynchus
--+           +--+                              
  |              +-- Trapezidens_angustior      
  |                                             
  |        +-------- Lamellidens_savadiensis **    
  |  +-----+                                    
  |  |     |  +----- Lamellidens_chindwinensis **  
  |  |     +--+                                 
  |  |        |  +-- Lamellidens_corrianus **      
  |  |        +--+                              
  +--+           +-- Lamellidens_nongyangensis **  
     |                                          
     |           +-- Lamellidens_marginalis **     
     |  +--------+                              
     |  |        +-- Lamellidens_ferrugineus **    
     +--+                                       
        |  +-------- Lamellidens_generosus **      
        +--+                                    
           |  +----- Lamellidens_mainwaringi **    
           +--+                                 
              |  +-- Lamellidens_jenkinsianus **   
              +--+                              
                 +-- Lamellidens_lamellatus **     

Bolotov et al., 2017, fig. S1

iT tT bi riS oT roT roS iD
9 38 27 9 116 3 1 41.0
                       +----- Trapezidens_exolescens     
              +--------+                                 
              |        |  +-- Trapezidens_dolichorhynchus
              |        +--+                              
  +-----------+           +-- Trapezidens_angustior      
  |           |                                          
  |           |  +----------- Lamellidens_lamellatus **     
  |           |  |                                       
  |           +--+  +-------- Lamellidens_jenkinsianus **   
  |              |  |                                    
  |              |  |  +----- Lamellidens_generosus **      
  |              +--+  |                                 
  |                 |  +----- Lamellidens_mainwaringi **    
  |                 |  |                                 
  |                 +--+  +-- Lamellidens_ferrugineus **    
  |                    +--+                              
  |                    |  +-- Lamellidens_marginalis **     
--+                    |                                 
  |                    |  +-- Lamellidens_savadiensis **    
  |                    +--+                              
  |                       +-- Lamellidens_corrianus[1] **   
  |                                                      
  |                 +-------- Parreysia_keralaensis      
  |  +--------------+                                    
  |  |              |  +----- Parreysia_plagiosoma       
  |  |              +--+                                 
  |  |                 |  +-- Parreysia_corrugata        
  |  |                 +--+                              
  |  |                    +-- Parreysia_nagpoorensis     
  |  |                                                   
  |  |              +-------- Leoparreysia_olivacea      
  +--+  +-----------+                                    
     |  |           |  +----- Leoparreysia_tavoyensis[2] 
     |  |           +--+                                 
     |  |              |  +-- Leoparreysia_canefrii      
     |  |              +--+                              
     |  |                 +-- Leoparreysia_tavoyensis[1] 
     |  |                                                
     |  |                 +-- Nitia_teretiuscula         
     |  |                 |                              
     +--+                 +-- Coelatura_gabonensis       
        |                 |                              
        |                 +-- Prisodontopsis_mweruensis  
        |  +--------------+                              
        |  |              +-- Coelatura_kunenensis       
        |  |              |                              
        |  |              +-- Coelatura_aegyptiaca[1]    
        |  |              |                              
        |  |              +-- Coelatura_aegyptiaca[2]    
        |  |                                             
        +--+        +-------- Indonaia_shurtleffiana     
           |        |                                    
           |  +-----+  +----- Indonaia_rugosa            
           |  |     |  |                                 
           |  |     |  |  +-- Indonaia_andersoniana      
           |  |     +--+--+                              
           |  |        |  +-- Indonaia_caerulea          
           |  |        |                                 
           +--+        |  +-- Indonaia_bonneaudii        
              |        +--+                              
              |           +-- Lamellidens_corrianus[2] **   
              |                                          
              |  +----------- Indochinella_pugio         
              |  |                                       
              |  |        +-- Scabies_humilis            
              +--+        |                              
                 |  +-----+-- Scabies_phaselus           
                 |  |     |                              
                 +--+     +-- Scabies_mandarinus         
                    |                                    
                    |  +----- Radiatula_mouhoti          
                    +--+                                 
                       |  +-- Radiatula_myitkyinae       
                       +--+                              
                          +-- Radiatula_chaudhurii       

Huang et al., 2019, fig. 2

iT tT bi riS oT roT roS iD
9 9 7 9 190 3 1 10.0
                    +----- Trapezidens_exolescens     
  +-----------------+                                 
  |                 |  +-- Trapezidens_dolichorhynchus
  |                 +--+                              
--+                    +-- Trapezidens_angustior      
  |                                                   
  |  +-------------------- Lamellidens_corrianus **      
  |  |                                                
  +--+  +----------------- Lamellidens_savadiensis **    
     |  |                                             
     +--+              +-- Lamellidens_lamellatus **     
        |  +-----------+                              
        |  |           +-- Lamellidens_jenkinsianus **   
        +--+                                          
           |  +----------- Lamellidens_generosus **      
           +--+                                       
              |  +-------- Lamellidens_mainwaringi[2] ** 
              +--+                                    
                 |  +----- Lamellidens_mainwaringi[1] ** 
                 +--+                                 
                    |  +-- Lamellidens_ferrugineus **    
                    +--+                              
                       +-- Lamellidens_marginalis **     

Huang et al., 2019, fig. S3

iT tT bi riS oT roT roS iD
9 9 3 9 188 3 1 9.0
        +----- Trapezidens_exolescens     
  +-----+                                 
  |     |  +-- Trapezidens_angustior      
  |     +--+                              
  |        +-- Trapezidens_dolichorhynchus
  |                                       
--+        +-- Lamellidens_corrianus **      
  |  +-----+                              
  |  |     +-- Lamellidens_savadiensis **    
  |  |                                    
  |  |  +----- Lamellidens_lamellatus **     
  +--+  |                                 
     |  +----- Lamellidens_jenkinsianus **   
     |  |                                 
     |  |  +-- Lamellidens_marginalis **     
     +--+--+                              
        |  +-- Lamellidens_ferrugineus **    
        |                                 
        |  +-- Lamellidens_mainwaringi[1] ** 
        |  |                              
        +--+-- Lamellidens_mainwaringi[2] ** 
           |                              
           +-- Lamellidens_generosus **      

Huang et al., 2019, fig. S4

iT tT bi riS oT roT roS iD
9 9 3 9 190 3 1 7.0
           +----- Trapezidens_exolescens     
  +--------+                                 
  |        |  +-- Trapezidens_angustior      
  |        +--+                              
--+           +-- Trapezidens_dolichorhynchus
  |                                          
  |  +----------- Lamellidens_corrianus **      
  |  |                                       
  +--+  +-------- Lamellidens_savadiensis **    
     |  |                                    
     |  |  +----- Lamellidens_marginalis **     
     +--+  |                                 
        |  +----- Lamellidens_ferrugineus **    
        |  |                                 
        +--+  +-- Lamellidens_mainwaringi[1] ** 
           |  |                              
           |  +-- Lamellidens_mainwaringi[2] ** 
           |  |                              
           +--+-- Lamellidens_lamellatus **     
              |                              
              +-- Lamellidens_generosus **      
              |                              
              +-- Lamellidens_jenkinsianus **   

Bolotov et al., 2022, fig. 2

iT tT bi riS oT roT roS iD
9 9 7 9 49 5 1 8.0
                    +-- Trapezidens_exolescens     
              +-----+                              
              |     +-- Trapezidens_scutum         
  +-----------+                                    
  |           |  +----- Trapezidens_angustior      
  |           +--+                                 
  |              |  +-- Trapezidens_dolichorhynchus
--+              +--+                              
  |                 +-- Trapezidens_yeti           
  |                                                
  |  +----------------- Lamellidens_corrianus **      
  |  |                                             
  +--+  +-------------- Lamellidens_nongyangensis **  
     |  |                                          
     +--+  +----------- Lamellidens_savadiensis **    
        |  |                                       
        |  |        +-- Lamellidens_ferrugineus **    
        +--+  +-----+                              
           |  |     +-- Lamellidens_marginalis **     
           |  |                                    
           +--+     +-- Lamellidens_lamellatus **     
              |  +--+                              
              |  |  +-- Lamellidens_jenkinsianus **   
              +--+                                 
                 |  +-- Lamellidens_generosus **      
                 +--+                              
                    +-- Lamellidens_mainwaringi **    

Bolotov et al., 2022, fig. S1

iT tT bi riS oT roT roS iD
9 9 6 9 49 5 1 9.0
           +-------- Trapezidens_exolescens     
           |                                    
  +--------+  +----- Trapezidens_scutum         
  |        |  |                                 
  |        +--+  +-- Trapezidens_angustior      
  |           |  |                              
  |           +--+-- Trapezidens_dolichorhynchus
  |              |                              
--+              +-- Trapezidens_yeti           
  |                                             
  |           +----- Lamellidens_savadiensis **    
  |  +--------+                                 
  |  |        |  +-- Lamellidens_corrianus **      
  |  |        +--+                              
  |  |           +-- Lamellidens_nongyangensis **  
  +--+                                          
     |           +-- Lamellidens_ferrugineus **    
     |  +--------+                              
     |  |        +-- Lamellidens_marginalis **     
     +--+                                       
        |  +-------- Lamellidens_lamellatus **     
        +--+                                    
           |  +----- Lamellidens_jenkinsianus **   
           +--+                                 
              |  +-- Lamellidens_generosus **      
              +--+                              
                 +-- Lamellidens_mainwaringi **    

Bolotov et al., 2022, fig. S2

iT tT bi riS oT roT roS iD
9 9 6 9 49 5 1 9.0
        +----------- Trapezidens_scutum         
  +-----+                                       
  |     |  +-------- Trapezidens_exolescens     
  |     +--+                                    
  |        |  +----- Trapezidens_yeti           
  |        +--+                                 
  |           |  +-- Trapezidens_dolichorhynchus
  |           +--+                              
--+              +-- Trapezidens_angustior      
  |                                             
  |           +----- Lamellidens_savadiensis **    
  |  +--------+                                 
  |  |        |  +-- Lamellidens_corrianus **      
  |  |        +--+                              
  |  |           +-- Lamellidens_nongyangensis **  
  +--+                                          
     |           +-- Lamellidens_marginalis **     
     |  +--------+                              
     |  |        +-- Lamellidens_ferrugineus **    
     +--+                                       
        |  +-------- Lamellidens_lamellatus **     
        +--+                                    
           |  +----- Lamellidens_jenkinsianus **   
           +--+                                 
              |  +-- Lamellidens_mainwaringi **    
              +--+                              
                 +-- Lamellidens_generosus **      

Bolotov et al., 2017, fig. S1

iT tT bi riS oT roT roS iD
6 6 4 6 132 2 1 7.0
              +-- Trapezidens_dolichorhynchus
  +-----------+                              
  |           +-- Trapezidens_exolescens     
--+                                          
  |  +----------- Lamellidens_corrianus **      
  |  |                                       
  +--+  +-------- Lamellidens_savadiensis **    
     |  |                                    
     +--+     +-- Lamellidens_jenkinsianus **   
        |  +--+                              
        |  |  +-- Lamellidens_mainwaringi **    
        +--+                                 
           |  +-- Lamellidens_ferrugineus **    
           +--+                              
              +-- Lamellidens_marginalis **     

Bolotov et al., 2017, fig. S2

iT tT bi riS oT roT roS iD
6 6 3 6 132 2 1 6.0
           +-- Trapezidens_dolichorhynchus
  +--------+                              
  |        +-- Trapezidens_exolescens     
  |                                       
--+  +-------- Lamellidens_savadiensis **    
  |  |                                    
  |  +-------- Lamellidens_corrianus **      
  +--+                                    
     |     +-- Lamellidens_marginalis **     
     |  +--+                              
     |  |  +-- Lamellidens_ferrugineus **    
     +--+                                 
        |  +-- Lamellidens_jenkinsianus **   
        +--+                              
           +-- Lamellidens_mainwaringi **    

Bolotov et al., 2018, fig. 2

iT tT bi riS oT roT roS iD
4 4 2 4 81 2 1 7.0
           +-- Trapezidens_exolescens     
  +--------+                              
  |        +-- Trapezidens_dolichorhynchus
--+                                       
  |  +-------- Lamellidens_savadiensis **    
  +--+                                    
     |  +----- Lamellidens_ferrugineus **    
     +--+                                 
        |  +-- Lamellidens_generosus **      
        +--+                              
           +-- Lamellidens_mainwaringi **    

Konopleva et al., 2020, fig. 2

iT tT bi riS oT roT roS iD
4 4 2 4 8 5 1 9.0
        +-------- Lamellidens_savadiensis **    
  +-----+                                    
  |     |  +----- Lamellidens_generosus **      
  |     +--+                                 
  |        |  +-- Lamellidens_mainwaringi **    
--+        +--+                              
  |           +-- Lamellidens_ferrugineus **    
  |                                          
  |  +----------- Trapezidens_scutum         
  +--+                                       
     |  +-------- Trapezidens_exolescens     
     +--+                                    
        |  +----- Trapezidens_yeti           
        +--+                                 
           |  +-- Trapezidens_angustior      
           +--+                              
              +-- Trapezidens_dolichorhynchus

Bolotov et al., 2020, fig. 1

iT tT bi riS oT roT roS iD
4 4 2 4 117 4 1 9.0
     +-------- Trapezidens_exolescens     
  +--+                                    
  |  |  +----- Trapezidens_scutum         
  |  +--+                                 
  |     |  +-- Trapezidens_dolichorhynchus
--+     +--+                              
  |        +-- Trapezidens_angustior      
  |                                       
  |  +-------- Lamellidens_savadiensis **    
  +--+                                    
     |  +----- Lamellidens_generosus **      
     +--+                                 
        |  +-- Lamellidens_ferrugineus **    
        +--+                              
           +-- Lamellidens_mainwaringi **    

Bolotov et al., 2020, fig. S1

iT tT bi riS oT roT roS iD
4 4 2 4 111 4 1 9.0
     +-------- Trapezidens_exolescens     
  +--+                                    
  |  |  +----- Trapezidens_scutum         
  |  +--+                                 
  |     |  +-- Trapezidens_dolichorhynchus
--+     +--+                              
  |        +-- Trapezidens_angustior      
  |                                       
  |  +-------- Lamellidens_savadiensis **    
  +--+                                    
     |  +----- Lamellidens_generosus **      
     +--+                                 
        |  +-- Lamellidens_ferrugineus **    
        +--+                              
           +-- Lamellidens_mainwaringi **    

Bolotov et al., 2022, fig. 8

iT tT bi riS oT roT roS iD
4 4 2 4 46 5 1 7.0
        +-------- Lamellidens_savadiensis **    
  +-----+                                    
  |     |  +----- Lamellidens_ferrugineus **    
  |     +--+                                 
  |        |  +-- Lamellidens_generosus **      
--+        +--+                              
  |           +-- Lamellidens_mainwaringi **    
  |                                          
  |  +----------- Trapezidens_exolescens     
  +--+                                       
     |  +-------- Trapezidens_scutum         
     +--+                                    
        |  +----- Trapezidens_yeti           
        +--+                                 
           |  +-- Trapezidens_angustior      
           +--+                              
              +-- Trapezidens_dolichorhynchus

Konopleva et al., 2017, fig. 5

iT tT bi riS oT roT roS iD
3 3 1 3 2 2 1 7.0
        +-- Trapezidens_exolescens     
  +-----+                              
  |     +-- Trapezidens_dolichorhynchus
--+                                    
  |  +----- Lamellidens_savadiensis **    
  +--+                                 
     |  +-- Lamellidens_mainwaringi **    
     +--+                              
        +-- Lamellidens_jenkinsianus **   

Konopleva et al., 2017, fig. 6

iT tT bi riS oT roT roS iD
3 3 1 3 28 2 1 7.0
        +-- Trapezidens_exolescens     
  +-----+                              
  |     +-- Trapezidens_dolichorhynchus
--+                                    
  |  +----- Lamellidens_savadiensis **    
  +--+                                 
     |  +-- Lamellidens_jenkinsianus **   
     +--+                              
        +-- Lamellidens_mainwaringi **    

Sonowal et al., 2021, fig. 4

iT tT bi riS oT roT roS iD
2 10 0 2 14 0 0 16.0
  +-------------------------- Lamellidens_mainwaringi **    
--+                                                      
  |  +----------------------- Lamprotula_leaii           
  +--+                                                   
     |  +-------------------- Potomida_littoralis        
     +--+                                                
        |  +----------------- Lampsilis_cardium          
        +--+                                             
           |  +-------------- Elliptio_complanata        
           +--+                                          
              |  +----------- Aculamprotula_tientsinensis
              +--+                                       
                 |  +-------- Unio_pictorum              
                 +--+                                    
                    |  +----- Anodonta_cygnea            
                    +--+                                 
                       |  +-- Unio_ravoisieri            
                       +--+                              
                          +-- Lamellidens_corrianus **      

Whelan et al., 2011, fig. 1 ML

iT tT bi riS oT roT roS iD
1 1 0 1 35 1 1 --
  +-- Lamellidens_savadiensis **    
--+                              
  +-- Trapezidens_dolichorhynchus

Whelan et al., 2011, fig. 1 MP

iT tT bi riS oT roT roS iD
1 1 0 1 35 1 1 --
  +-- Lamellidens_savadiensis **    
--+                              
  +-- Trapezidens_dolichorhynchus

Pfeiffer & Graf, 2013, fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 34 0 0 --
-- Lamellidens_savadiensis

Pfeiffer & Graf, 2015, fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 39 1 1 --
  +-- Trapezidens_dolichorhynchus
--+                              
  +-- Lamellidens_savadiensis **    

Jadhav & Jamkhedkar, 2009, fig. 11

iT tT bi riS oT roT roS iD
1 1 0 1 25 0 0 --
-- Lamellidens_corrianus

Araujo et al., 2017, fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 13 0 0 --
-- Lamellidens_savadiensis

Pfeiffer et al., 2018, fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 28 1 1 --
  +-- Trapezidens_dolichorhynchus
--+                              
  +-- Lamellidens_savadiensis **    

Jeratthitikul et al., 2019, fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 22 1 1 --
  +-- Lamellidens_mainwaringi **
--+                          
  +-- Trapezidens_exolescens 

Pfeiffer et al., 2019, fig. 4-5

iT tT bi riS oT roT roS iD
1 1 0 1 56 1 1 --
  +-- Trapezidens_dolichorhynchus
--+                              
  +-- Lamellidens_savadiensis **    

Pfeiffer et al., 2019, fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 56 1 1 --
  +-- Trapezidens_dolichorhynchus
--+                              
  +-- Lamellidens_savadiensis **    

Choi et al., 2020, fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 8 0 0 --
-- Lamellidens_generosus

Sonowal et al., 2021, fig. 3a

iT tT bi riS oT roT roS iD
1 1 0 1 14 0 0 --
-- Lamellidens_corrianus

Sonowal et al., 2021, fig. 3b

iT tT bi riS oT roT roS iD
1 1 0 1 14 0 0 --
-- Lamellidens_mainwaringi

Pavan-Kumar et al., 2022, fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 35 0 0 --
-- Lamellidens_marginalis

Pavan-Kumar et al., 2022, fig. S5

iT tT bi riS oT roT roS iD
1 1 0 1 38 0 0 --
-- Lamellidens_marginalis

Konopleva et al., 2023, fig. S1, 4

iT tT bi riS oT roT roS iD
1 1 0 1 55 1 1 --
  +-- Trapezidens_exolescens 
--+                          
  +-- Lamellidens_savadiensis **

Page updated: 08:56:47 Wed 18 Jan 2023

 
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