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Page last updated
Wed 27 Sep 2023

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Parreysiinae | Lamellidentini

genus Lamellidens Simpson, 1900: cladomics
     13 valid species

Cladomics Summary

number of species = 13

analyzed in 36 cladograms in 22 publications
maximum number of terminals in an analysis = 10, mean = 3.97

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 20
maximum number of included terminals = 10, mean = 6.35
proportion of cladograms that support basic monophyly = 0.9

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ tree

Bolotov et al., 2022 [Publ 4252], fig. S1

iT tT bi riS oT roT roS iD
10 10 8 10 51 6 1 0.0

*** Top Monophyly Cladogram *** *** Top Sister-Group Cladogram ***

     +-------------- Trapezidens_mogaungensis   
  +--+                                          
  |  |  +----------- Trapezidens_scutum         
  |  +--+                                       
  |     |  +-------- Trapezidens_exolescens     
  |     +--+                                    
  |        |  +----- Trapezidens_yeti           
  |        +--+                                 
  |           |  +-- Trapezidens_dolichorhynchus
--+           +--+                              
  |              +-- Trapezidens_angustior      
  |                                             
  |        +-------- Lamellidens_savadiensis **    
  |  +-----+                                    
  |  |     |  +----- Lamellidens_chindwinensis **  
  |  |     +--+                                 
  |  |        |  +-- Lamellidens_corrianus **      
  |  |        +--+                              
  +--+           +-- Lamellidens_nongyangensis **  
     |                                          
     |           +-- Lamellidens_marginalis **     
     |  +--------+                              
     |  |        +-- Lamellidens_ferrugineus **    
     +--+                                       
        |  +-------- Lamellidens_generosus **      
        +--+                                    
           |  +----- Lamellidens_mainwaringi **    
           +--+                                 
              |  +-- Lamellidens_jenkinsianus **   
              +--+                              
                 +-- Lamellidens_lamellatus **

Bolotov et al., 2022 [Publ 4252], fig. 2

iT tT bi riS oT roT roS iD
10 10 7 10 51 6 1 7.0 
        +----------- Trapezidens_mogaungensis   
        |                                       
  +-----+  +-------- Trapezidens_exolescens     
  |     |  |                                    
  |     +--+  +----- Trapezidens_scutum         
  |        |  |                                 
  |        +--+  +-- Trapezidens_angustior      
  |           |  |                              
--+           +--+-- Trapezidens_dolichorhynchus
  |              |                              
  |              +-- Trapezidens_yeti           
  |                                             
  |  +-------------- Lamellidens_marginalis **     
  |  |                                          
  |  |  +----------- Lamellidens_ferrugineus **    
  +--+  |                                       
     |  |  +-------- Lamellidens_generosus **      
     |  +--+                                    
     |  |  |  +----- Lamellidens_mainwaringi **    
     +--+  +--+                                 
        |     |  +-- Lamellidens_lamellatus **     
        |     +--+                              
        |        +-- Lamellidens_jenkinsianus **   
        |                                       
        |  +-------- Lamellidens_nongyangensis **  
        +--+                                    
           |  +----- Lamellidens_savadiensis **    
           +--+                                 
              |  +-- Lamellidens_corrianus **      
              +--+                              
                 +-- Lamellidens_chindwinensis **

Bolotov et al., 2017 [Publ 3764], fig. S1

iT tT bi riS oT roT roS iD
9 41 28 9 119 3 1 43.0 
                       +----- Trapezidens_exolescens       
              +--------+                                   
              |        |  +-- Trapezidens_dolichorhynchus  
              |        +--+                                
  +-----------+           +-- Trapezidens_angustior        
  |           |                                            
  |           |  +----------- Lamellidens_lamellatus **       
  |           |  |                                         
  |           +--+  +-------- Lamellidens_jenkinsianus **     
  |              |  |                                      
  |              |  |  +----- Lamellidens_generosus **        
  |              +--+  |                                   
  |                 |  +----- Lamellidens_mainwaringi **      
  |                 |  |                                   
  |                 +--+  +-- Lamellidens_ferrugineus **      
  |                    +--+                                
  |                    |  +-- Lamellidens_marginalis **       
  |                    |                                   
--+                    |  +-- Lamellidens_savadiensis **      
  |                    +--+                                
  |                       +-- Lamellidens_corrianus[1] **     
  |                                                        
  |                 +-------- Parreysia_keralaensis        
  |  +--------------+                                      
  |  |              |  +----- Parreysia_plagiosoma         
  |  |              +--+                                   
  |  |                 |  +-- Parreysia_corrugata          
  |  |                 +--+                                
  |  |                    +-- Parreysia_nagpoorensis       
  |  |                                                     
  |  |              +-------- Leoparreysia_olivacea        
  |  |  +-----------+                                      
  |  |  |           |  +----- Leoparreysia_tavoyensis[2]   
  +--+  |           +--+                                   
     |  |              |  +-- Leoparreysia_canefrii        
     |  |              +--+                                
     |  |                 +-- Leoparreysia_tavoyensis[1]   
     |  |                                                  
     |  |                 +-- Nitia_teretiuscula           
     |  |                 |                                
     |  |                 +-- Coelatura_gabonensis         
     |  |                 |                                
     |  |                 +-- Prisodontopsis_mweruensis    
     +--+                 |                                
        |                 +-- Coelatura_kunenensis         
        |                 |                                
        |  +--------------+-- Coelatura_aegyptiaca[1]      
        |  |              |                                
        |  |              +-- Coelatura_aegyptiaca[2]      
        |  |              |                                
        |  |              +-- Prisodontopsis_aviculaeformis
        |  |              |                                
        |  |              +-- Coelatura_luapulaensis       
        |  |              |                                
        |  |              +-- Coelatura_choziensis         
        +--+                                               
           |        +-------- Indonaia_shurtleffiana       
           |        |                                      
           |  +-----+  +----- Indonaia_rugosa              
           |  |     |  |                                   
           |  |     |  |  +-- Indonaia_andersoniana        
           |  |     +--+--+                                
           |  |        |  +-- Indonaia_caerulea            
           |  |        |                                   
           +--+        |  +-- Indonaia_bonneaudii          
              |        +--+                                
              |           +-- Lamellidens_corrianus[2] **     
              |                                            
              |  +----------- Indochinella_pugio           
              |  |                                         
              |  |        +-- Scabies_humilis              
              +--+        |                                
                 |  +-----+-- Scabies_phaselus             
                 |  |     |                                
                 +--+     +-- Scabies_mandarinus           
                    |                                      
                    |  +----- Radiatula_mouhoti            
                    +--+                                   
                       |  +-- Radiatula_myitkyinae         
                       +--+                                
                          +-- Radiatula_chaudhurii

Huang et al., 2019 [Publ 3843], fig. 2

iT tT bi riS oT roT roS iD
9 9 7 9 192 3 1 9.0 
                    +----- Trapezidens_exolescens     
  +-----------------+                                 
  |                 |  +-- Trapezidens_dolichorhynchus
  |                 +--+                              
--+                    +-- Trapezidens_angustior      
  |                                                   
  |  +-------------------- Lamellidens_corrianus **      
  |  |                                                
  +--+  +----------------- Lamellidens_savadiensis **    
     |  |                                             
     +--+              +-- Lamellidens_lamellatus **     
        |  +-----------+                              
        |  |           +-- Lamellidens_jenkinsianus **   
        +--+                                          
           |  +----------- Lamellidens_generosus **      
           +--+                                       
              |  +-------- Lamellidens_mainwaringi[2] ** 
              +--+                                    
                 |  +----- Lamellidens_mainwaringi[1] ** 
                 +--+                                 
                    |  +-- Lamellidens_ferrugineus **    
                    +--+                              
                       +-- Lamellidens_marginalis **

Huang et al., 2019 [Publ 3843], fig. S3

iT tT bi riS oT roT roS iD
9 9 4 9 189 3 1 7.0 
        +----- Trapezidens_exolescens     
  +-----+                                 
  |     |  +-- Trapezidens_angustior      
  |     +--+                              
  |        +-- Trapezidens_dolichorhynchus
  |                                       
--+        +-- Lamellidens_corrianus **      
  |  +-----+                              
  |  |     +-- Lamellidens_savadiensis **    
  |  |                                    
  |  |  +----- Lamellidens_lamellatus **     
  +--+  |                                 
     |  +----- Lamellidens_jenkinsianus **   
     |  |                                 
     |  |  +-- Lamellidens_marginalis **     
     +--+--+                              
        |  +-- Lamellidens_ferrugineus **    
        |                                 
        |  +-- Lamellidens_mainwaringi[1] ** 
        |  |                              
        +--+-- Lamellidens_mainwaringi[2] ** 
           |                              
           +-- Lamellidens_generosus **

Huang et al., 2019 [Publ 3843], fig. S4

iT tT bi riS oT roT roS iD
9 9 3 9 191 3 1 8.0 
           +----- Trapezidens_exolescens     
  +--------+                                 
  |        |  +-- Trapezidens_angustior      
  |        +--+                              
--+           +-- Trapezidens_dolichorhynchus
  |                                          
  |  +----------- Lamellidens_corrianus **      
  |  |                                       
  +--+  +-------- Lamellidens_savadiensis **    
     |  |                                    
     |  |  +----- Lamellidens_marginalis **     
     +--+  |                                 
        |  +----- Lamellidens_ferrugineus **    
        |  |                                 
        +--+  +-- Lamellidens_mainwaringi[1] ** 
           |  |                              
           |  +-- Lamellidens_mainwaringi[2] ** 
           |  |                              
           +--+-- Lamellidens_lamellatus **     
              |                              
              +-- Lamellidens_generosus **      
              |                              
              +-- Lamellidens_jenkinsianus **

Bolotov et al., 2022 [Publ 4239], fig. 2

iT tT bi riS oT roT roS iD
9 9 7 9 49 5 1 7.0 
                    +-- Trapezidens_exolescens     
              +-----+                              
              |     +-- Trapezidens_scutum         
  +-----------+                                    
  |           |  +----- Trapezidens_angustior      
  |           +--+                                 
  |              |  +-- Trapezidens_dolichorhynchus
--+              +--+                              
  |                 +-- Trapezidens_yeti           
  |                                                
  |  +----------------- Lamellidens_corrianus **      
  |  |                                             
  +--+  +-------------- Lamellidens_nongyangensis **  
     |  |                                          
     +--+  +----------- Lamellidens_savadiensis **    
        |  |                                       
        |  |        +-- Lamellidens_ferrugineus **    
        +--+  +-----+                              
           |  |     +-- Lamellidens_marginalis **     
           |  |                                    
           +--+     +-- Lamellidens_lamellatus **     
              |  +--+                              
              |  |  +-- Lamellidens_jenkinsianus **   
              +--+                                 
                 |  +-- Lamellidens_generosus **      
                 +--+                              
                    +-- Lamellidens_mainwaringi **

Bolotov et al., 2022 [Publ 4239], fig. S1

iT tT bi riS oT roT roS iD
9 9 7 9 49 5 1 5.0 
           +-------- Trapezidens_exolescens     
           |                                    
  +--------+  +----- Trapezidens_scutum         
  |        |  |                                 
  |        +--+  +-- Trapezidens_angustior      
  |           |  |                              
  |           +--+-- Trapezidens_dolichorhynchus
  |              |                              
--+              +-- Trapezidens_yeti           
  |                                             
  |           +----- Lamellidens_savadiensis **    
  |  +--------+                                 
  |  |        |  +-- Lamellidens_corrianus **      
  |  |        +--+                              
  |  |           +-- Lamellidens_nongyangensis **  
  +--+                                          
     |           +-- Lamellidens_ferrugineus **    
     |  +--------+                              
     |  |        +-- Lamellidens_marginalis **     
     +--+                                       
        |  +-------- Lamellidens_lamellatus **     
        +--+                                    
           |  +----- Lamellidens_jenkinsianus **   
           +--+                                 
              |  +-- Lamellidens_generosus **      
              +--+                              
                 +-- Lamellidens_mainwaringi **

Bolotov et al., 2022 [Publ 4239], fig. S2

iT tT bi riS oT roT roS iD
9 9 7 9 49 5 1 5.0 
        +----------- Trapezidens_scutum         
  +-----+                                       
  |     |  +-------- Trapezidens_exolescens     
  |     +--+                                    
  |        |  +----- Trapezidens_yeti           
  |        +--+                                 
  |           |  +-- Trapezidens_dolichorhynchus
  |           +--+                              
--+              +-- Trapezidens_angustior      
  |                                             
  |           +----- Lamellidens_savadiensis **    
  |  +--------+                                 
  |  |        |  +-- Lamellidens_corrianus **      
  |  |        +--+                              
  |  |           +-- Lamellidens_nongyangensis **  
  +--+                                          
     |           +-- Lamellidens_marginalis **     
     |  +--------+                              
     |  |        +-- Lamellidens_ferrugineus **    
     +--+                                       
        |  +-------- Lamellidens_lamellatus **     
        +--+                                    
           |  +----- Lamellidens_jenkinsianus **   
           +--+                                 
              |  +-- Lamellidens_mainwaringi **    
              +--+                              
                 +-- Lamellidens_generosus **

Bolotov et al., 2017 [Publ 3761], fig. S1

iT tT bi riS oT roT roS iD
6 6 4 6 132 2 1 6.0 
              +-- Trapezidens_dolichorhynchus
  +-----------+                              
  |           +-- Trapezidens_exolescens     
--+                                          
  |  +----------- Lamellidens_corrianus **      
  |  |                                       
  +--+  +-------- Lamellidens_savadiensis **    
     |  |                                    
     +--+     +-- Lamellidens_jenkinsianus **   
        |  +--+                              
        |  |  +-- Lamellidens_mainwaringi **    
        +--+                                 
           |  +-- Lamellidens_ferrugineus **    
           +--+                              
              +-- Lamellidens_marginalis **

Bolotov et al., 2017 [Publ 3761], fig. S2

iT tT bi riS oT roT roS iD
6 6 3 6 132 2 1 5.0 
           +-- Trapezidens_dolichorhynchus
  +--------+                              
  |        +-- Trapezidens_exolescens     
  |                                       
--+  +-------- Lamellidens_savadiensis **    
  |  |                                    
  |  +-------- Lamellidens_corrianus **      
  +--+                                    
     |     +-- Lamellidens_marginalis **     
     |  +--+                              
     |  |  +-- Lamellidens_ferrugineus **    
     +--+                                 
        |  +-- Lamellidens_jenkinsianus **   
        +--+                              
           +-- Lamellidens_mainwaringi **

Bolotov et al., 2018 [Publ 3826], fig. 2

iT tT bi riS oT roT roS iD
4 4 2 4 81 2 1 6.0 
           +-- Trapezidens_exolescens     
  +--------+                              
  |        +-- Trapezidens_dolichorhynchus
--+                                       
  |  +-------- Lamellidens_savadiensis **    
  +--+                                    
     |  +----- Lamellidens_ferrugineus **    
     +--+                                 
        |  +-- Lamellidens_generosus **      
        +--+                              
           +-- Lamellidens_mainwaringi **

Konopleva et al., 2020 [Publ 4072], fig. 2

iT tT bi riS oT roT roS iD
4 4 2 4 8 5 1 8.0 
        +-------- Lamellidens_savadiensis **    
  +-----+                                    
  |     |  +----- Lamellidens_generosus **      
  |     +--+                                 
  |        |  +-- Lamellidens_mainwaringi **    
--+        +--+                              
  |           +-- Lamellidens_ferrugineus **    
  |                                          
  |  +----------- Trapezidens_scutum         
  +--+                                       
     |  +-------- Trapezidens_exolescens     
     +--+                                    
        |  +----- Trapezidens_yeti           
        +--+                                 
           |  +-- Trapezidens_angustior      
           +--+                              
              +-- Trapezidens_dolichorhynchus

Bolotov et al., 2020 [Publ 4100], fig. 1

iT tT bi riS oT roT roS iD
4 4 2 4 117 4 1 8.0 
     +-------- Trapezidens_exolescens     
  +--+                                    
  |  |  +----- Trapezidens_scutum         
  |  +--+                                 
  |     |  +-- Trapezidens_dolichorhynchus
--+     +--+                              
  |        +-- Trapezidens_angustior      
  |                                       
  |  +-------- Lamellidens_savadiensis **    
  +--+                                    
     |  +----- Lamellidens_generosus **      
     +--+                                 
        |  +-- Lamellidens_ferrugineus **    
        +--+                              
           +-- Lamellidens_mainwaringi **

Bolotov et al., 2020 [Publ 4100], fig. S1

iT tT bi riS oT roT roS iD
4 4 2 4 111 4 1 8.0 
     +-------- Trapezidens_exolescens     
  +--+                                    
  |  |  +----- Trapezidens_scutum         
  |  +--+                                 
  |     |  +-- Trapezidens_dolichorhynchus
--+     +--+                              
  |        +-- Trapezidens_angustior      
  |                                       
  |  +-------- Lamellidens_savadiensis **    
  +--+                                    
     |  +----- Lamellidens_generosus **      
     +--+                                 
        |  +-- Lamellidens_ferrugineus **    
        +--+                              
           +-- Lamellidens_mainwaringi **

Bolotov et al., 2022 [Publ 4250], fig. 8

iT tT bi riS oT roT roS iD
4 4 2 4 46 5 1 6.0 
        +-------- Lamellidens_savadiensis **    
  +-----+                                    
  |     |  +----- Lamellidens_ferrugineus **    
  |     +--+                                 
  |        |  +-- Lamellidens_generosus **      
--+        +--+                              
  |           +-- Lamellidens_mainwaringi **    
  |                                          
  |  +----------- Trapezidens_exolescens     
  +--+                                       
     |  +-------- Trapezidens_scutum         
     +--+                                    
        |  +----- Trapezidens_yeti           
        +--+                                 
           |  +-- Trapezidens_angustior      
           +--+                              
              +-- Trapezidens_dolichorhynchus

Bolotov et al., 2022 [Publ 4250], fig. S4

iT tT bi riS oT roT roS iD
4 4 2 4 5 5 1 6.0 
     +-------- Lamellidens_savadiensis **    
  +--+                                    
  |  |  +----- Lamellidens_ferrugineus **    
  |  +--+                                 
  |     |  +-- Lamellidens_mainwaringi **    
--+     +--+                              
  |        +-- Lamellidens_generosus **      
  |                                       
  |  +-------- Trapezidens_scutum         
  |  |                                    
  +--+  +----- Trapezidens_exolescens     
     |  |                                 
     +--+  +-- Trapezidens_dolichorhynchus
        |  |                              
        +--+-- Trapezidens_angustior      
           |                              
           +-- Trapezidens_yeti

Konopleva et al., 2017 [Publ 3807], fig. 5

iT tT bi riS oT roT roS iD
3 3 1 3 2 2 1 7.0 
        +-- Trapezidens_exolescens     
  +-----+                              
  |     +-- Trapezidens_dolichorhynchus
--+                                    
  |  +----- Lamellidens_savadiensis **    
  +--+                                 
     |  +-- Lamellidens_mainwaringi **    
     +--+                              
        +-- Lamellidens_jenkinsianus **

Konopleva et al., 2017 [Publ 3807], fig. 6

iT tT bi riS oT roT roS iD
3 3 1 3 28 2 1 7.0 
        +-- Trapezidens_exolescens     
  +-----+                              
  |     +-- Trapezidens_dolichorhynchus
--+                                    
  |  +----- Lamellidens_savadiensis **    
  +--+                                 
     |  +-- Lamellidens_jenkinsianus **   
     +--+                              
        +-- Lamellidens_mainwaringi **

Sonowal et al., 2021 [Publ 4221], fig. 4

iT tT bi riS oT roT roS iD
2 10 0 2 14 0 0 16.0 
  +-------------------------- Lamellidens_mainwaringi **    
--+                                                      
  |  +----------------------- Lamprotula_leaii           
  +--+                                                   
     |  +-------------------- Potomida_littoralis        
     +--+                                                
        |  +----------------- Lampsilis_cardium          
        +--+                                             
           |  +-------------- Elliptio_complanata        
           +--+                                          
              |  +----------- Aculamprotula_tientsinensis
              +--+                                       
                 |  +-------- Unio_pictorum              
                 +--+                                    
                    |  +----- Anodonta_cygnea            
                    +--+                                 
                       |  +-- Unio_ravoisieri            
                       +--+                              
                          +-- Lamellidens_corrianus **

Whelan et al., 2011 [Publ 3380], fig. 1 ML

iT tT bi riS oT roT roS iD
1 1 0 1 36 1 1 -- 
  +-- Lamellidens_savadiensis **    
--+                              
  +-- Trapezidens_dolichorhynchus

Whelan et al., 2011 [Publ 3380], fig. 1 MP

iT tT bi riS oT roT roS iD
1 1 0 1 36 1 1 -- 
  +-- Lamellidens_savadiensis **    
--+                              
  +-- Trapezidens_dolichorhynchus

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 34 0 0 -- 
-- Lamellidens_savadiensis

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 39 1 1 -- 
  +-- Trapezidens_dolichorhynchus
--+                              
  +-- Lamellidens_savadiensis **

Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11

iT tT bi riS oT roT roS iD
1 1 0 1 25 0 0 -- 
-- Lamellidens_corrianus

Araujo et al., 2017 [Publ 3801], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 13 0 0 -- 
-- Lamellidens_savadiensis

Pfeiffer et al., 2018 [Publ 3827], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 28 1 1 -- 
  +-- Trapezidens_dolichorhynchus
--+                              
  +-- Lamellidens_savadiensis **

Jeratthitikul et al., 2019 [Publ 3922], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 22 1 1 -- 
  +-- Lamellidens_mainwaringi **
--+                          
  +-- Trapezidens_exolescens

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iT tT bi riS oT roT roS iD
1 1 0 1 56 1 1 -- 
  +-- Trapezidens_dolichorhynchus
--+                              
  +-- Lamellidens_savadiensis **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 56 1 1 -- 
  +-- Trapezidens_dolichorhynchus
--+                              
  +-- Lamellidens_savadiensis **

Choi et al., 2020 [Publ 4169], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 8 0 0 -- 
-- Lamellidens_generosus

Sonowal et al., 2021 [Publ 4221], fig. 3a

iT tT bi riS oT roT roS iD
1 1 0 1 14 0 0 -- 
-- Lamellidens_corrianus

Sonowal et al., 2021 [Publ 4221], fig. 3b

iT tT bi riS oT roT roS iD
1 1 0 1 14 0 0 -- 
-- Lamellidens_mainwaringi

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 35 0 0 -- 
-- Lamellidens_marginalis

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iT tT bi riS oT roT roS iD
1 1 0 1 38 0 0 -- 
-- Lamellidens_marginalis

Konopleva et al., 2023 [Publ 4269], fig. S1, 4

iT tT bi riS oT roT roS iD
1 1 0 1 55 1 1 -- 
  +-- Trapezidens_exolescens 
--+                          
  +-- Lamellidens_savadiensis **

Page updated: 10:55:08 Wed 27 Sep 2023
 
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