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Page last updated
Wed 03 Aug 2022

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Parreysiinae | Indochinellini

genus Indonaia Prashad, 1918: cladomics
     12 valid species

Cladomics Summary

number of species = 12

analyzed in 17 cladograms in 11 publications
maximum number of terminal species in an analysis = 7, mean = 3.65

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 12
maximum number of included species = 7, mean = 4.75
proportion of cladograms that support basic monophyly = 0.67


Huang et al., 2019, fig. 2

iT tT bi riS oT roT roS iD
7 37 29 7 194 7 3 0.0

*** Top Monophyly Cladogram ***

                       +----- Trapezidens_exolescens     
     +-----------------+                                 
     |                 |  +-- Trapezidens_dolichorhynchus
     |                 +--+                              
     |                    +-- Trapezidens_angustior      
  +--+                                                   
  |  |                    +-- Indonaia_shurtleffiana[4] **  
  |  |  +-----------------+                              
  |  |  |                 +-- Lamellidens_corrianus      
  |  +--+                                                
  |     |  +----------------- Lamellidens_savadiensis    
  |     |  |                                             
  |     +--+              +-- Lamellidens_marginalis[3]  
  |        |  +-----------+                              
  |        |  |           +-- Lamellidens_jenkinsianus   
  |        +--+                                          
  |           |  +----------- Lamellidens_generosus[2]   
  |           +--+                                       
--+              |  +-------- Lamellidens_marginalis[2]  
  |              +--+                                    
  |                 |  +----- Lamellidens_marginalis[1]  
  |                 +--+                                 
  |                    |  +-- Lamellidens_generosus[1]   
  |                    +--+                              
  |                       +-- Lamellidens_marginalis[4]  
  |                                                      
  |  +----------------------- Parreysia_corrugata        
  |  |                                                   
  |  |                 +----- Leoparreysia_canefrii      
  |  |                 |                                 
  |  |  +--------------+----- Leoparreysia_olivacea      
  |  |  |              |                                 
  +--+  |              |  +-- Leoparreysia_bhamoensis    
     |  |              +--+                              
     |  |                 +-- Leoparreysia_tavoyensis    
     |  |                                                
     |  |           +-------- Coelatura_aegyptiaca[2]    
     |  |           |                                    
     +--+  +--------+  +----- Coelatura_aegyptiaca[1]    
        |  |        |  |                                 
        |  |        |  +----- Nitia_teretiuscula         
        |  |        +--+                                 
        |  |           |  +-- Coelatura_gabonensis       
        |  |           |  |                              
        |  |           +--+-- Coelatura_kunenensis       
        |  |              |                              
        |  |              +-- Prisodontopsis_mweruensis  
        +--+                                             
           |        +-------- Indonaia_shurtleffiana[1] **  
           |        |                                    
           |  +-----+  +----- Indonaia_shurtleffiana[3] **  
           |  |     |  |                                 
           |  |     |  |  +-- Indonaia_shurtleffiana[2] **  
           |  |     +--+--+                              
           |  |        |  +-- Indonaia_rugosa **            
           |  |        |                                 
           |  |        |  +-- Indonaia_andersoniana **      
           +--+        +--+                              
              |           +-- Indonaia_caerulea **          
              |                                          
              |           +-- Indochinella_pugio[1]      
              |           |                              
              |  +--------+-- Indochinella_pugio[2]      
              |  |        |                              
              +--+        +-- Indochinella               
                 |                                       
                 |  +-------- Scabies_humilis            
                 +--+                                    
                    |  +----- Radiatula_myitkyinae       
                    +--+                                 
                       |  +-- Radiatula_chaudhurii       
                       +--+                              
                          +-- Radiatula_mouhoti          

Bolotov et al., 2020, fig. 1

iT tT bi riS oT roT roS iD
5 5 3 5 117 16 6 36.0

*** Top Sister-Group Cladogram ***

                    +----------- Indonaia_shurtleffiana **  
  +-----------------+                                    
  |                 |  +-------- Indonaia_rugosa **         
  |                 +--+                                 
  |                    |  +----- Indonaia_andersoniana **   
  |                    +--+                              
  |                       |  +-- Indonaia_subclathrata **   
--+                       +--+                           
  |                          +-- Indonaia_caerulea **       
  |                                                      
  |  +-------------------------- Indochinella_pugio      
  |  |                                                   
  |  |                 +-------- Radiatula_crispisulcata 
  |  |  +--------------+                                 
  +--+  |              |  +----- Radiatula_mouhoti       
     |  |              +--+                              
     |  |                 |  +-- Radiatula_chaudhurii    
     |  |                 +--+                           
     +--+                    +-- Radiatula_myitkyinae    
        |                                                
        |  +-------------------- Scabiellus_songkramensis
        |  |                                             
        |  |                 +-- Unionetta_fabagina      
        +--+  +--------------+                           
           |  |              +-- Harmandia_somboriensis  
           |  |                                          
           +--+  +-------------- Scabies_nucleus         
              |  |                                       
              |  |           +-- Scabies_crispata        
              +--+  +--------+                           
                 |  |        +-- Scabies_mandarinus      
                 |  |                                    
                 +--+        +-- Scabies_humilis         
                    |  +-----+                           
                    |  |     +-- Scabies_pilata          
                    +--+                                 
                       |  +----- Scabies_anceps          
                       +--+                              
                          |  +-- Scabies_phaselus        
                          +--+                           
                             +-- Scabies_scobinatus      

Huang et al., 2019, fig. S3

iT tT bi riS oT roT roS iD
7 35 28 7 190 5 3 13.0
                       +----- Trapezidens_exolescens     
                 +-----+                                 
                 |     |  +-- Trapezidens_angustior      
                 |     +--+                              
                 |        +-- Trapezidens_dolichorhynchus
                 |                                       
  +--------------+     +----- Lamellidens_savadiensis    
  |              |  +--+                                 
  |              |  |  |  +-- Indonaia_shurtleffiana[4] **  
  |              |  |  +--+                              
  |              |  |     +-- Lamellidens_corrianus      
  |              |  |                                    
  |              +--+  +----- Lamellidens_marginalis[3]  
  |                 |  |                                 
  |                 |  +----- Lamellidens_jenkinsianus   
  |                 |  |                                 
  |                 |  |  +-- Lamellidens_marginalis[4]  
  |                 +--+--+                              
--+                    |  +-- Lamellidens_generosus[1]   
  |                    |                                 
  |                    |  +-- Lamellidens_marginalis[1]  
  |                    |  |                              
  |                    +--+-- Lamellidens_marginalis[2]  
  |                       |                              
  |                       +-- Lamellidens_generosus[2]   
  |                                                      
  |  +----------------------- Parreysia_corrugata        
  |  |                                                   
  |  |              +-------- Leoparreysia_canefrii      
  |  |  +-----------+                                    
  |  |  |           |  +----- Leoparreysia_olivacea      
  +--+  |           +--+                                 
     |  |              |  +-- Leoparreysia_bhamoensis    
     |  |              +--+                              
     |  |                 +-- Leoparreysia_tavoyensis    
     |  |                                                
     |  |        +----------- Coelatura_aegyptiaca[2]    
     +--+        |                                       
        |  +-----+        +-- Coelatura_aegyptiaca[1]    
        |  |     |  +-----+                              
        |  |     |  |     +-- Nitia_teretiuscula         
        |  |     +--+                                    
        |  |        |  +----- Coelatura_kunenensis       
        |  |        +--+                                 
        |  |           |  +-- Coelatura_gabonensis       
        |  |           +--+                              
        +--+              +-- Prisodontopsis_mweruensis  
           |                                             
           |     +----------- Indochinella_pugio         
           |  +--+                                       
           |  |  |  +-------- Scabies_humilis            
           |  |  +--+                                    
           |  |     |  +----- Radiatula_mouhoti          
           |  |     +--+                                 
           +--+        |  +-- Radiatula_myitkyinae       
              |        +--+                              
              |           +-- Radiatula_chaudhurii       
              |                                          
              |     +-------- Indonaia_shurtleffiana[1] **  
              |     |                                    
              +-----+  +----- Indonaia_shurtleffiana[3] **  
                    |  |                                 
                    |  |  +-- Indonaia_shurtleffiana[2] **  
                    +--+--+                              
                       |  +-- Indonaia_rugosa **            
                       |                                 
                       |  +-- Indonaia_andersoniana **      
                       +--+                              
                          +-- Indonaia_caerulea **          

Huang et al., 2019, fig. S4

iT tT bi riS oT roT roS iD
7 35 25 7 193 5 3 11.0
  +----------------------- Parreysia_corrugata        
  |                                                   
  |                 +----- Trapezidens_exolescens     
  |        +--------+                                 
  |        |        |  +-- Trapezidens_angustior      
  |        |        +--+                              
  |        |           +-- Trapezidens_dolichorhynchus
  +--------+                                          
  |        |           +-- Indonaia_shurtleffiana[4] **  
  |        |  +--------+                              
  |        |  |        +-- Lamellidens_corrianus      
  |        |  |                                       
  |        +--+  +-------- Lamellidens_savadiensis    
  |           |  |                                    
  |           |  |  +----- Lamellidens_marginalis[4]  
  |           +--+  |                                 
  |              |  +----- Lamellidens_generosus[2]   
  |              |  |                                 
  |              +--+  +-- Lamellidens_marginalis[1]  
--+                 |  |                              
  |                 |  +-- Lamellidens_marginalis[2]  
  |                 |  |                              
  |                 +--+-- Lamellidens_marginalis[3]  
  |                    |                              
  |                    +-- Lamellidens_generosus[1]   
  |                    |                              
  |                    +-- Lamellidens_jenkinsianus   
  |                                                   
  |                    +-- Leoparreysia_bhamoensis    
  |                 +--+                              
  |                 |  +-- Leoparreysia_tavoyensis    
  |  +--------------+                                 
  |  |              |  +-- Leoparreysia_canefrii      
  |  |              +--+                              
  |  |                 +-- Leoparreysia_olivacea      
  |  |                                                
  |  |           +-------- Coelatura_aegyptiaca[2]    
  |  |           |                                    
  |  |           +-------- Nitia_teretiuscula         
  +--+  +--------+                                    
     |  |        |  +----- Coelatura_aegyptiaca[1]    
     |  |        |  |                                 
     |  |        +--+  +-- Coelatura_gabonensis       
     |  |           |  |                              
     |  |           +--+-- Coelatura_kunenensis       
     |  |              |                              
     |  |              +-- Prisodontopsis_mweruensis  
     +--+                                             
        |     +----------- Indochinella_pugio         
        |  +--+                                       
        |  |  |  +-------- Scabies_humilis            
        |  |  +--+                                    
        |  |     |  +----- Radiatula_mouhoti          
        |  |     +--+                                 
        |  |        |  +-- Radiatula_myitkyinae       
        +--+        +--+                              
           |           +-- Radiatula_chaudhurii       
           |                                          
           |     +-------- Indonaia_shurtleffiana[1] **  
           |     |                                    
           |     |     +-- Indonaia_andersoniana **      
           +-----+-----+                              
                 |     +-- Indonaia_caerulea **          
                 |                                    
                 |  +----- Indonaia_shurtleffiana[3] **  
                 +--+                                 
                    |  +-- Indonaia_shurtleffiana[2] **  
                    +--+                              
                       +-- Indonaia_rugosa **            

Bolotov et al., 2017, fig. S1

iT tT bi riS oT roT roS iD
6 6 3 6 116 5 3 32.0
     +----------- Indochinella_pugio       
  +--+                                     
  |  |  +-------- Scabies_humilis          
  |  +--+                                  
  |     |  +----- Radiatula_mouhoti        
  |     +--+                               
--+        |  +-- Radiatula_myitkyinae     
  |        +--+                            
  |           +-- Radiatula_chaudhurii     
  |                                        
  |     +-------- Indonaia_shurtleffiana[1] **
  |     |                                  
  +-----+  +----- Indonaia_shurtleffiana[3] **
        |  |                               
        |  |  +-- Indonaia_andersoniana **    
        +--+--+                            
           |  +-- Indonaia_caerulea **        
           |                               
           |  +-- Indonaia_rugosa **          
           +--+                            
              +-- Indonaia_shurtleffiana[2] **

Konopleva et al., 2019, fig. 1

iT tT bi riS oT roT roS iD
5 5 3 5 47 7 3 36.0
     +----------- Indonaia_shurtleffiana **
  +--+                                  
  |  |  +-------- Indonaia_rugosa **       
  |  +--+                               
  |     |  +----- Indonaia_andersoniana ** 
  |     +--+                            
  |        |  +-- Indonaia_caerulea **     
--+        +--+                         
  |           +-- Indonaia_subclathrata ** 
  |                                     
  |  +----------- Indochinella_pugio    
  |  |                                  
  |  |        +-- Scabies_humilis[1]    
  +--+        |                         
     |  +-----+-- Scabies_humilis[2]    
     |  |     |                         
     +--+     +-- Scabies_phaselus      
        |                               
        |  +----- Radiatula_mouhoti     
        +--+                            
           |  +-- Radiatula_myitkyinae  
           +--+                         
              +-- Radiatula_chaudhurii  

Bolotov et al., 2018, fig. 2

iT tT bi riS oT roT roS iD
4 4 2 4 78 5 3 35.0
        +-------- Indonaia_shurtleffiana **
  +-----+                               
  |     |  +----- Indonaia_rugosa **       
  |     +--+                            
  |        |  +-- Indonaia_caerulea **     
--+        +--+                         
  |           +-- Indonaia_andersoniana ** 
  |                                     
  |  +----------- Indochinella_pugio    
  +--+                                  
     |  +-------- Scabies_humilis       
     +--+                               
        |  +----- Radiatula_myitkyinae  
        +--+                            
           |  +-- Radiatula_mouhoti     
           +--+                         
              +-- Radiatula_chaudhurii  

Bolotov et al., 2020, fig. S1

iT tT bi riS oT roT roS iD
4 4 2 4 112 16 6 35.0
                             +-------- Indonaia_shurtleffiana **  
  +--------------------------+                                 
  |                          |  +----- Indonaia_andersoniana **   
  |                          +--+                              
  |                             |  +-- Indonaia_caerulea **       
  |                             +--+                           
--+                                +-- Indonaia_subclathrata **   
  |                                                            
  |  +-------------------------------- Indochinella_pugio      
  |  |                                                         
  |  |                          +----- Radiatula_mouhoti       
  +--+  +-----------------------+                              
     |  |                       |  +-- Radiatula_myitkyinae    
     |  |                       +--+                           
     +--+                          +-- Radiatula_chaudhurii    
        |                                                      
        |  +-------------------------- Radiatula_crispisulcata 
        |  |                                                   
        +--+  +----------------------- Scabiellus_songkramensis
           |  |                                                
           +--+  +-------------------- Harmandia_somboriensis  
              |  |                                             
              +--+  +----------------- Unionetta_fabagina      
                 |  |                                          
                 +--+  +-------------- Scabies_mandarinus      
                    |  |                                       
                    +--+  +----------- Scabies_crispata        
                       |  |                                    
                       +--+  +-------- Scabies_nucleus         
                          |  |                                 
                          +--+     +-- Scabies_humilis         
                             |  +--+                           
                             |  |  +-- Scabies_pilata          
                             +--+                              
                                |  +-- Scabies_phaselus        
                                |  |                           
                                +--+-- Scabies_scobinatus      
                                   |                           
                                   +-- Scabies_anceps          

Bolotov et al., 2017, fig. S1

iT tT bi riS oT roT roS iD
3 3 1 3 133 6 3 35.0
        +----- Indonaia_shurtleffiana[1] **
  +-----+                               
  |     |  +-- Indonaia_caerulea **        
  |     +--+                            
  |        +-- Indonaia_shurtleffiana[2] **
--+                                     
  |        +-- Indochinella_pugio       
  |  +-----+                            
  |  |     +-- Scabies_crispata[1]      
  |  |                                  
  +--+     +-- Scabies_humilis          
     |  +--+                            
     |  |  +-- Scabies_crispata[2]      
     +--+                               
        |  +-- Radiatula_myitkyinae     
        +--+                            
           +-- Radiatula_chaudhurii     

Bolotov et al., 2017, fig. S2

iT tT bi riS oT roT roS iD
3 3 1 3 133 6 3 35.0
        +----- Indonaia_shurtleffiana[1] **
  +-----+                               
  |     |  +-- Indonaia_caerulea **        
  |     +--+                            
  |        +-- Indonaia_shurtleffiana[2] **
--+                                     
  |        +-- Indochinella_pugio       
  |  +-----+                            
  |  |     +-- Scabies_crispata[2]      
  |  |                                  
  +--+     +-- Scabies_humilis          
     |  +--+                            
     |  |  +-- Scabies_crispata[1]      
     +--+                               
        |  +-- Radiatula_myitkyinae     
        +--+                            
           +-- Radiatula_chaudhurii     

Pfeiffer et al., 2018, fig. 2

iT tT bi riS oT roT roS iD
3 16 12 3 23 13 5 42.0
                                +-- Indonaia_caerulea **       
  +-----------------------------+                           
  |                             +-- Indonaia_shurtleffiana **  
--+                                                         
  |  +----------------------------- Indochinella_pugio      
  |  |                                                      
  +--+  +-------------------------- Indonaia_bonneaudii **     
     |  |                                                   
     +--+  +----------------------- Scabiellus_songkramensis
        |  |                                                
        +--+  +-------------------- Unionetta_fabagina[2]   
           |  |                                             
           +--+  +----------------- Harmandia_somboriensis  
              |  |                                          
              +--+  +-------------- Unionetta_fabagina[1]   
                 |  |                                       
                 +--+           +-- Scabies_crispata        
                    |  +--------+                           
                    |  |        +-- Scabies_mandarinus      
                    +--+                                    
                       |  +-------- Scabies_nucleus         
                       |  |                                 
                       +--+     +-- Scabies_pilata          
                          |  +--+                           
                          |  |  +-- Scabies_humilis         
                          +--+                              
                             |  +-- Scabies_phaselus        
                             |  |                           
                             +--+-- Scabies_anceps          
                                |                           
                                +-- Scabies_scobinatus      

Jeratthitikul et al., 2019, fig. 1

iT tT bi riS oT roT roS iD
3 3 1 3 20 15 6 34.0
                                   +----- Indonaia_rugosa **         
  +--------------------------------+                              
  |                                |  +-- Indonaia_caerulea **       
  |                                +--+                           
  |                                   +-- Indonaia_andersoniana **   
--+                                                               
  |  +----------------------------------- Indochinella_pugio      
  |  |                                                            
  |  |                             +----- Radiatula_mouhoti       
  +--+  +--------------------------+                              
     |  |                          |  +-- Radiatula_myitkyinae    
     |  |                          +--+                           
     +--+                             +-- Radiatula_chaudhurii    
        |                                                         
        |  +----------------------------- Scabiellus_songkramensis
        +--+                                                      
           |  +-------------------------- Harmandia_somboriensis  
           +--+                                                   
              |  +----------------------- Unionetta_fabagina      
              +--+                                                
                 |  +-------------------- Scabies_crispata        
                 +--+                                             
                    |  +----------------- Scabies_mandarinus      
                    +--+                                          
                       |  +-------------- Scabies_pilata          
                       +--+                                       
                          |  +----------- Scabies_humilis[2]      
                          +--+                                    
                             |  +-------- Scabies_humilis[1]      
                             +--+                                 
                                |  +----- Scabies_scobinatus      
                                +--+                              
                                   |  +-- Scabies_phaselus        
                                   +--+                           
                                      +-- Scabies_anceps          

Pfeiffer et al., 2019, fig. 4-5

iT tT bi riS oT roT roS iD
1 1 0 1 56 3 3 --
  +-------- Indonaia_caerulea **     
--+                               
  |  +----- Indochinella_pugio    
  +--+                            
     |  +-- Scabies_phaselus      
     +--+                         
        +-- Harmandia_somboriensis

Pfeiffer et al., 2019, fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 56 3 3 --
  +-------- Indonaia_caerulea **     
--+                               
  |  +----- Indochinella_pugio    
  +--+                            
     |  +-- Scabies_phaselus      
     +--+                         
        +-- Harmandia_somboriensis

Ortiz-Sepulveda et al., 2020, fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 33 5 3 --
  +-------- Indonaia_andersoniana **
  |                              
--+  +----- Indochinella_pugio   
  |  |                           
  +--+  +-- Radiatula_mouhoti    
     |  |                        
     |  +-- Scabies_humilis      
     +--+                        
        +-- Radiatula_myitkyinae 
        |                        
        +-- Radiatula_chaudhurii 

Ortiz-Sepulveda et al., 2020, fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 33 5 3 --
  +-- Radiatula_mouhoti    
  |                        
  +-- Scabies_humilis      
  |                        
  +-- Radiatula_myitkyinae 
--+                        
  +-- Radiatula_chaudhurii 
  |                        
  +-- Indochinella_pugio   
  |                        
  +-- Indonaia_andersoniana **

Wu et al., 2021, fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 23 0 0 --
No taxon cladogram to show.

Page updated: 09:39:46 Wed 03 Aug 2022

 
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