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Page last updated
Thu 04 Apr 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Parreysiinae | Indochinellini

genus Indonaia Prashad, 1918: cladomics
     12 valid species

Cladomics Summary

number of species = 12

analyzed in 28 cladograms in 17 publications
maximum number of terminals in an analysis = 8, mean = 3.86

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 19
maximum number of included terminals = 8, mean = 5.21
proportion of cladograms that support basic monophyly = 0.95

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ tree

Bolotov et al., 2022 [Publ 4252], fig. 2

iT tT bi riS oT roT roS iD
8 8 6 8 53 19 6 0.0

*** Top Monophyly Cladogram *** *** Top Sister-Group Cladogram ***

                 +----------------- Indonaia_cylindrica **     
                 |                                          
  +--------------+  +-------------- Indonaia_gratiosa **       
  |              |  |                                       
  |              +--+  +----------- Indonaia_andersoniana **   
  |                 |  |                                    
  |                 +--+        +-- Indonaia_caerulea **       
  |                    |  +-----+                           
  |                    |  |     +-- Indonaia_subclathrata **   
  |                    +--+                                 
  |                       |  +----- Indonaia_rugosa **         
--+                       +--+                              
  |                          |  +-- Indonaia_bonneaudii **     
  |                          +--+                           
  |                             +-- Indonaia_shurtleffiana **  
  |                                                         
  |  +----------------------------- Indochinella_pugio      
  |  |                                                      
  |  |                 +----------- Radiatula_crispisulcata 
  |  |                 |                                    
  |  |                 |     +----- Radiatula_chaudhurii    
  +--+  +--------------+  +--+                              
     |  |              |  |  |  +-- Radiatula_myitkyinae    
     |  |              |  |  +--+                           
     |  |              +--+     +-- Radiatula_kachinensis   
     |  |                 |                                 
     |  |                 |  +----- Radiatula_chindwinensis 
     +--+                 +--+                              
        |                    |  +-- Radiatula_myitthanensis 
        |                    +--+                           
        |                       +-- Radiatula_mouhoti       
        |                                                   
        |  +----------------------- Scabiellus_songkramensis
        |  |                                                
        +--+  +-------------------- Harmandia_somboriensis  
           |  |                                             
           +--+  +----------------- Unionetta_fabagina      
              |  |                                          
              +--+              +-- Scabies_crispata        
                 |  +-----------+                           
                 |  |           +-- Scabies_mandarinus      
                 +--+                                       
                    |  +----------- Scabies_nucleus         
                    |  |                                    
                    +--+        +-- Scabies_humilis         
                       |  +-----+                           
                       |  |     +-- Scabies_pilata          
                       +--+                                 
                          |  +----- Scabies_phaselus        
                          +--+                              
                             |  +-- Scabies_anceps          
                             +--+                           
                                +-- Scabies_scobinatus

Bolotov et al., 2022 [Publ 4239], fig. 2

iT tT bi riS oT roT roS iD
8 8 6 8 50 18 6 6.0 
                                      +-- Indonaia_cylindrica **     
                             +--------+                           
                             |        +-- Indonaia_gratiosa **       
                             |                                    
  +--------------------------+     +----- Indonaia_andersoniana **   
  |                          |  +--+                              
  |                          |  |  |  +-- Indonaia_caerulea **       
  |                          |  |  +--+                           
  |                          +--+     +-- Indonaia_subclathrata **   
  |                             |                                 
  |                             |  +----- Indonaia_shurtleffiana **  
  |                             +--+                              
--+                                |  +-- Indonaia_bonneaudii **     
  |                                +--+                           
  |                                   +-- Indonaia_rugosa **         
  |                                                               
  |  +----------------------------------- Indochinella_pugio      
  |  |                                                            
  |  |                       +----------- Radiatula_crispisulcata 
  |  |                       |                                    
  +--+  +--------------------+        +-- Radiatula_chaudhurii    
     |  |                    |  +-----+                           
     |  |                    |  |     +-- Radiatula_myitkyinae    
     |  |                    +--+                                 
     |  |                       |  +----- Radiatula_mouhoti       
     +--+                       +--+                              
        |                          |  +-- Radiatula_chindwinensis 
        |                          +--+                           
        |                             +-- Radiatula_myitthanensis 
        |                                                         
        |  +----------------------------- Scabiellus_songkramensis
        +--+                                                      
           |  +-------------------------- Harmandia_somboriensis  
           +--+                                                   
              |  +----------------------- Unionetta_fabagina      
              +--+                                                
                 |  +-------------------- Scabies_crispata        
                 +--+                                             
                    |  +----------------- Scabies_mandarinus      
                    +--+                                          
                       |  +-------------- Scabies_nucleus         
                       +--+                                       
                          |  +----------- Scabies_pilata          
                          +--+                                    
                             |  +-------- Scabies_humilis         
                             +--+                                 
                                |  +----- Scabies_anceps          
                                +--+                              
                                   |  +-- Scabies_phaselus        
                                   +--+                           
                                      +-- Scabies_scobinatus

Bolotov et al., 2022 [Publ 4239], fig. S1

iT tT bi riS oT roT roS iD
8 8 6 8 50 18 6 0.0 
                 +----------------- Indonaia_cylindrica **     
                 |                                          
  +--------------+  +-------------- Indonaia_gratiosa **       
  |              |  |                                       
  |              +--+  +----------- Indonaia_andersoniana **   
  |                 |  |                                    
  |                 +--+        +-- Indonaia_caerulea **       
  |                    |  +-----+                           
  |                    |  |     +-- Indonaia_subclathrata **   
  |                    +--+                                 
  |                       |  +----- Indonaia_rugosa **         
--+                       +--+                              
  |                          |  +-- Indonaia_bonneaudii **     
  |                          +--+                           
  |                             +-- Indonaia_shurtleffiana **  
  |                                                         
  |  +----------------------------- Indochinella_pugio      
  |  |                                                      
  |  |                    +-------- Radiatula_crispisulcata 
  |  |                    |                                 
  |  |  +-----------------+     +-- Radiatula_chaudhurii    
  +--+  |                 |  +--+                           
     |  |                 |  |  +-- Radiatula_myitkyinae    
     |  |                 +--+                              
     |  |                    |  +-- Radiatula_myitthanensis 
     |  |                    |  |                           
     +--+                    +--+-- Radiatula_mouhoti       
        |                       |                           
        |                       +-- Radiatula_chindwinensis 
        |                                                   
        |  +----------------------- Scabiellus_songkramensis
        |  |                                                
        +--+  +-------------------- Harmandia_somboriensis  
           |  |                                             
           +--+  +----------------- Unionetta_fabagina      
              |  |                                          
              +--+              +-- Scabies_crispata        
                 |  +-----------+                           
                 |  |           +-- Scabies_mandarinus      
                 +--+                                       
                    |  +----------- Scabies_nucleus         
                    |  |                                    
                    +--+        +-- Scabies_humilis         
                       |  +-----+                           
                       |  |     +-- Scabies_pilata          
                       +--+                                 
                          |  +----- Scabies_phaselus        
                          +--+                              
                             |  +-- Scabies_anceps          
                             +--+                           
                                +-- Scabies_scobinatus

Bolotov et al., 2022 [Publ 4239], fig. S2

iT tT bi riS oT roT roS iD
8 8 6 8 50 18 6 6.0 
                                   +-- Indonaia_cylindrica **     
                          +--------+                           
                          |        +-- Indonaia_gratiosa **       
                          |                                    
  +-----------------------+     +----- Indonaia_shurtleffiana **  
  |                       |  +--+                              
  |                       |  |  |  +-- Indonaia_bonneaudii **     
  |                       |  |  +--+                           
  |                       +--+     +-- Indonaia_rugosa **         
  |                          |                                 
  |                          |  +----- Indonaia_andersoniana **   
  |                          +--+                              
--+                             |  +-- Indonaia_caerulea **       
  |                             +--+                           
  |                                +-- Indonaia_subclathrata **   
  |                                                            
  |  +-------------------------------- Indochinella_pugio      
  |  |                                                         
  |  |                    +----------- Radiatula_crispisulcata 
  |  |                    |                                    
  |  |  +-----------------+        +-- Radiatula_chaudhurii    
  +--+  |                 |  +-----+                           
     |  |                 |  |     +-- Radiatula_myitkyinae    
     |  |                 +--+                                 
     |  |                    |  +----- Radiatula_mouhoti       
     |  |                    +--+                              
     +--+                       |  +-- Radiatula_myitthanensis 
        |                       +--+                           
        |                          +-- Radiatula_chindwinensis 
        |                                                      
        |  +-------------------------- Scabiellus_songkramensis
        |  |                                                   
        +--+  +----------------------- Harmandia_somboriensis  
           |  |                                                
           +--+  +-------------------- Unionetta_fabagina      
              |  |                                             
              +--+  +----------------- Scabies_crispata        
                 |  |                                          
                 +--+              +-- Scabies_humilis         
                    |  +-----------+                           
                    |  |           +-- Scabies_mandarinus      
                    +--+                                       
                       |  +----------- Scabies_nucleus         
                       +--+                                    
                          |  +-------- Scabies_pilata          
                          +--+                                 
                             |  +----- Scabies_anceps          
                             +--+                              
                                |  +-- Scabies_phaselus        
                                +--+                           
                                   +-- Scabies_scobinatus

Bolotov et al., 2022 [Publ 4252], fig. S1

iT tT bi riS oT roT roS iD
8 8 6 8 53 19 6 6.0 
                                   +-- Indonaia_cylindrica **     
                          +--------+                           
                          |        +-- Indonaia_gratiosa **       
                          |                                    
  +-----------------------+     +----- Indonaia_shurtleffiana **  
  |                       |  +--+                              
  |                       |  |  |  +-- Indonaia_bonneaudii **     
  |                       |  |  +--+                           
  |                       +--+     +-- Indonaia_rugosa **         
  |                          |                                 
  |                          |  +----- Indonaia_andersoniana **   
  |                          +--+                              
  |                             |  +-- Indonaia_caerulea **       
--+                             +--+                           
  |                                +-- Indonaia_subclathrata **   
  |                                                            
  |  +-------------------------------- Indochinella_pugio      
  |  |                                                         
  |  |                          +----- Radiatula_chaudhurii    
  |  |                    +-----+                              
  |  |                    |     |  +-- Radiatula_kachinensis   
  |  |                    |     +--+                           
  +--+  +-----------------+        +-- Radiatula_myitkyinae    
     |  |                 |                                    
     |  |                 |  +-------- Radiatula_crispisulcata 
     |  |                 +--+                                 
     |  |                    |  +----- Radiatula_mouhoti       
     |  |                    +--+                              
     +--+                       |  +-- Radiatula_myitthanensis 
        |                       +--+                           
        |                          +-- Radiatula_chindwinensis 
        |                                                      
        |  +-------------------------- Scabiellus_songkramensis
        |  |                                                   
        +--+  +----------------------- Harmandia_somboriensis  
           |  |                                                
           +--+  +-------------------- Unionetta_fabagina      
              |  |                                             
              +--+  +----------------- Scabies_crispata        
                 |  |                                          
                 +--+              +-- Scabies_humilis         
                    |  +-----------+                           
                    |  |           +-- Scabies_mandarinus      
                    +--+                                       
                       |  +----------- Scabies_nucleus         
                       +--+                                    
                          |  +-------- Scabies_pilata          
                          +--+                                 
                             |  +----- Scabies_anceps          
                             +--+                              
                                |  +-- Scabies_phaselus        
                                +--+                           
                                   +-- Scabies_scobinatus

Bolotov et al., 2017 [Publ 3764], fig. S1

iT tT bi riS oT roT roS iD
5 6 3 5 126 7 3 9.0 
        +-------- Indonaia_shurtleffiana **  
        |                                 
  +-----+  +----- Indonaia_rugosa **         
  |     |  |                              
  |     |  |  +-- Indonaia_andersoniana **   
  |     +--+--+                           
  |        |  +-- Indonaia_caerulea **       
  |        |                              
--+        |  +-- Indonaia_bonneaudii **     
  |        +--+                           
  |           +-- Lamellidens_corrianus[2]
  |                                       
  |  +----------- Indochinella_pugio      
  |  |                                    
  |  |        +-- Scabies_humilis         
  +--+        |                           
     |  +-----+-- Scabies_phaselus        
     |  |     |                           
     +--+     +-- Scabies_mandarinus      
        |                                 
        |  +----- Radiatula_mouhoti       
        +--+                              
           |  +-- Radiatula_myitkyinae    
           +--+                           
              +-- Radiatula_chaudhurii

Huang et al., 2019 [Publ 3843], fig. 2

iT tT bi riS oT roT roS iD
5 5 2 5 199 7 3 8.0 
        +-------- Indonaia_shurtleffiana **
        |                               
  +-----+  +----- Indonaia_bonneaudii **   
  |     |  |                            
  |     +--+----- Indonaia_rugosa **       
  |        |                            
  |        |  +-- Indonaia_andersoniana ** 
--+        +--+                         
  |           +-- Indonaia_caerulea **     
  |                                     
  |  +----------- Indochinella_pugio    
  |  |                                  
  |  |     +----- Scabies_humilis       
  +--+  +--+                            
     |  |  |  +-- Scabies_phaselus      
     |  |  +--+                         
     +--+     +-- Scabies_mandarinus    
        |                               
        |  +----- Radiatula_myitkyinae  
        +--+                            
           |  +-- Radiatula_chaudhurii  
           +--+                         
              +-- Radiatula_mouhoti

Huang et al., 2019 [Publ 3843], fig. S3

iT tT bi riS oT roT roS iD
5 5 2 5 196 7 3 8.0 
        +-------- Indonaia_shurtleffiana **
        |                               
  +-----+  +----- Indonaia_bonneaudii **   
  |     |  |                            
  |     +--+----- Indonaia_rugosa **       
  |        |                            
  |        |  +-- Indonaia_andersoniana ** 
--+        +--+                         
  |           +-- Indonaia_caerulea **     
  |                                     
  |  +----------- Indochinella_pugio    
  |  |                                  
  |  |     +----- Scabies_phaselus      
  +--+  +--+                            
     |  |  |  +-- Scabies_humilis       
     |  |  +--+                         
     +--+     +-- Scabies_mandarinus    
        |                               
        |  +----- Radiatula_mouhoti     
        +--+                            
           |  +-- Radiatula_myitkyinae  
           +--+                         
              +-- Radiatula_chaudhurii

Huang et al., 2019 [Publ 3843], fig. S4

iT tT bi riS oT roT roS iD
5 5 2 5 197 7 3 8.0 
           +----- Indonaia_shurtleffiana **
           |                            
           |  +-- Indonaia_bonneaudii **   
  +--------+--+                         
  |        |  +-- Indonaia_rugosa **       
  |        |                            
  |        |  +-- Indonaia_andersoniana ** 
  |        +--+                         
--+           +-- Indonaia_caerulea **     
  |                                     
  |  +----------- Indochinella_pugio    
  |  |                                  
  |  |     +----- Scabies_humilis       
  +--+  +--+                            
     |  |  |  +-- Scabies_phaselus      
     |  |  +--+                         
     +--+     +-- Scabies_mandarinus    
        |                               
        |  +----- Radiatula_mouhoti     
        +--+                            
           |  +-- Radiatula_myitkyinae  
           +--+                         
              +-- Radiatula_chaudhurii

Konopleva et al., 2019 [Publ 3900], fig. 1

iT tT bi riS oT roT roS iD
5 5 3 5 47 7 3 7.0 
     +----------- Indonaia_shurtleffiana **
  +--+                                  
  |  |  +-------- Indonaia_rugosa **       
  |  +--+                               
  |     |  +----- Indonaia_andersoniana ** 
  |     +--+                            
  |        |  +-- Indonaia_caerulea **     
--+        +--+                         
  |           +-- Indonaia_subclathrata ** 
  |                                     
  |  +----------- Indochinella_pugio    
  |  |                                  
  |  |        +-- Scabies_humilis       
  +--+        |                         
     |  +-----+-- Scabies_mandarinus    
     |  |     |                         
     +--+     +-- Scabies_phaselus      
        |                               
        |  +----- Radiatula_mouhoti     
        +--+                            
           |  +-- Radiatula_myitkyinae  
           +--+                         
              +-- Radiatula_chaudhurii

Bolotov et al., 2020 [Publ 4100], fig. 1

iT tT bi riS oT roT roS iD
5 5 3 5 115 16 6 7.0 
                    +----------- Indonaia_shurtleffiana **  
  +-----------------+                                    
  |                 |  +-------- Indonaia_rugosa **         
  |                 +--+                                 
  |                    |  +----- Indonaia_andersoniana **   
  |                    +--+                              
  |                       |  +-- Indonaia_subclathrata **   
--+                       +--+                           
  |                          +-- Indonaia_caerulea **       
  |                                                      
  |  +-------------------------- Indochinella_pugio      
  |  |                                                   
  |  |                 +-------- Radiatula_crispisulcata 
  |  |  +--------------+                                 
  +--+  |              |  +----- Radiatula_mouhoti       
     |  |              +--+                              
     |  |                 |  +-- Radiatula_chaudhurii    
     |  |                 +--+                           
     +--+                    +-- Radiatula_myitkyinae    
        |                                                
        |  +-------------------- Scabiellus_songkramensis
        |  |                                             
        |  |                 +-- Unionetta_fabagina      
        +--+  +--------------+                           
           |  |              +-- Harmandia_somboriensis  
           |  |                                          
           +--+  +-------------- Scabies_nucleus         
              |  |                                       
              |  |           +-- Scabies_crispata        
              +--+  +--------+                           
                 |  |        +-- Scabies_mandarinus      
                 |  |                                    
                 +--+        +-- Scabies_humilis         
                    |  +-----+                           
                    |  |     +-- Scabies_pilata          
                    +--+                                 
                       |  +----- Scabies_anceps          
                       +--+                              
                          |  +-- Scabies_phaselus        
                          +--+                           
                             +-- Scabies_scobinatus

Bolotov et al., 2022 [Publ 4250], fig. S2

iT tT bi riS oT roT roS iD
5 5 3 5 10 7 2 7.0 
        +----------- Indonaia_rugosa **        
  +-----+                                   
  |     |  +-------- Indonaia_shurtleffiana ** 
  |     +--+                                
  |        |  +----- Indonaia_andersoniana **  
  |        +--+                             
--+           |  +-- Indonaia_subclathrata **  
  |           +--+                          
  |              +-- Indonaia_caerulea **      
  |                                         
  |  +-------------- Indochinella_pugio     
  |  |                                      
  +--+  +----------- Radiatula_crispisulcata
     |  |                                   
     +--+        +-- Radiatula_chaudhurii   
        |  +-----+                          
        |  |     +-- Radiatula_myitkyinae   
        +--+                                
           |  +----- Radiatula_chindwinensis
           +--+                             
              |  +-- Radiatula_mouhoti      
              +--+                          
                 +-- Radiatula_myitthanensis

Bolotov et al., 2017 [Publ 3761], fig. S1

iT tT bi riS oT roT roS iD
4 4 2 4 137 7 3 8.0 
        +-------- Indonaia_shurtleffiana **
  +-----+                               
  |     |  +----- Indonaia_andersoniana ** 
  |     +--+                            
  |        |  +-- Indonaia_rugosa **       
  |        +--+                         
--+           +-- Indonaia_bonneaudii **   
  |                                     
  |           +-- Indochinella_pugio    
  |  +--------+                         
  |  |        +-- Scabies_crispata      
  |  |                                  
  +--+        +-- Radiatula_myitkyinae  
     |  +-----+                         
     |  |     +-- Radiatula_chaudhurii  
     +--+                               
        |  +----- Scabies_mandarinus    
        +--+                            
           |  +-- Scabies_humilis       
           +--+                         
              +-- Scabies_phaselus

Bolotov et al., 2017 [Publ 3761], fig. S2

iT tT bi riS oT roT roS iD
4 4 2 4 137 7 3 8.0 
        +-------- Indonaia_shurtleffiana **
  +-----+                               
  |     |  +----- Indonaia_andersoniana ** 
  |     +--+                            
  |        |  +-- Indonaia_rugosa **       
  |        +--+                         
--+           +-- Indonaia_bonneaudii **   
  |                                     
  |           +-- Indochinella_pugio    
  |  +--------+                         
  |  |        +-- Scabies_crispata      
  |  |                                  
  +--+        +-- Radiatula_myitkyinae  
     |  +-----+                         
     |  |     +-- Radiatula_chaudhurii  
     +--+                               
        |  +----- Scabies_mandarinus    
        +--+                            
           |  +-- Scabies_humilis       
           +--+                         
              +-- Scabies_phaselus

Bolotov et al., 2018 [Publ 3826], fig. 2

iT tT bi riS oT roT roS iD
4 4 2 4 81 7 3 8.0 
        +-------- Indonaia_shurtleffiana **
  +-----+                               
  |     |  +----- Indonaia_rugosa **       
  |     +--+                            
  |        |  +-- Indonaia_caerulea **     
  |        +--+                         
--+           +-- Indonaia_andersoniana ** 
  |                                     
  |  +----------- Indochinella_pugio    
  |  |                                  
  |  |     +----- Radiatula_myitkyinae  
  +--+  +--+                            
     |  |  |  +-- Radiatula_mouhoti     
     |  |  +--+                         
     +--+     +-- Radiatula_chaudhurii  
        |                               
        |  +----- Scabies_mandarinus    
        +--+                            
           |  +-- Scabies_humilis       
           +--+                         
              +-- Scabies_phaselus

Bolotov et al., 2020 [Publ 4100], fig. S1

iT tT bi riS oT roT roS iD
4 4 2 4 110 16 6 6.0 
                             +-------- Indonaia_shurtleffiana **  
  +--------------------------+                                 
  |                          |  +----- Indonaia_andersoniana **   
  |                          +--+                              
  |                             |  +-- Indonaia_caerulea **       
  |                             +--+                           
--+                                +-- Indonaia_subclathrata **   
  |                                                            
  |  +-------------------------------- Indochinella_pugio      
  |  |                                                         
  |  |                          +----- Radiatula_mouhoti       
  +--+  +-----------------------+                              
     |  |                       |  +-- Radiatula_myitkyinae    
     |  |                       +--+                           
     +--+                          +-- Radiatula_chaudhurii    
        |                                                      
        |  +-------------------------- Radiatula_crispisulcata 
        |  |                                                   
        +--+  +----------------------- Scabiellus_songkramensis
           |  |                                                
           +--+  +-------------------- Harmandia_somboriensis  
              |  |                                             
              +--+  +----------------- Unionetta_fabagina      
                 |  |                                          
                 +--+  +-------------- Scabies_mandarinus      
                    |  |                                       
                    +--+  +----------- Scabies_crispata        
                       |  |                                    
                       +--+  +-------- Scabies_nucleus         
                          |  |                                 
                          +--+     +-- Scabies_humilis         
                             |  +--+                           
                             |  |  +-- Scabies_pilata          
                             +--+                              
                                |  +-- Scabies_phaselus        
                                |  |                           
                                +--+-- Scabies_scobinatus      
                                   |                           
                                   +-- Scabies_anceps

Pfeiffer et al., 2018 [Publ 3827], fig. 2

iT tT bi riS oT roT roS iD
3 3 1 3 26 15 6 7.0 
                             +----- Indonaia_bonneaudii **     
  +--------------------------+                              
  |                          |  +-- Indonaia_andersoniana **   
  |                          +--+                           
--+                             +-- Indonaia_caerulea **       
  |                                                         
  |  +----------------------------- Indochinella_pugio      
  |  |                                                      
  +--+                          +-- Radiatula_chaudhurii    
     |  +-----------------------+                           
     |  |                       +-- Radiatula_myitkyinae    
     +--+                                                   
        |  +----------------------- Scabiellus_songkramensis
        |  |                                                
        +--+  +-------------------- Unionetta_fabagina[2]   
           |  |                                             
           +--+  +----------------- Harmandia_somboriensis  
              |  |                                          
              +--+  +-------------- Unionetta_fabagina[1]   
                 |  |                                       
                 +--+           +-- Scabies_crispata        
                    |  +--------+                           
                    |  |        +-- Scabies_mandarinus      
                    +--+                                    
                       |  +-------- Scabies_nucleus         
                       |  |                                 
                       +--+     +-- Scabies_pilata          
                          |  +--+                           
                          |  |  +-- Scabies_humilis         
                          +--+                              
                             |  +-- Scabies_phaselus        
                             |  |                           
                             +--+-- Scabies_anceps          
                                |                           
                                +-- Scabies_scobinatus

Jeratthitikul et al., 2019 [Publ 3922], fig. 1

iT tT bi riS oT roT roS iD
3 3 1 3 20 15 6 7.0 
                                   +----- Indonaia_rugosa **         
  +--------------------------------+                              
  |                                |  +-- Indonaia_caerulea **       
  |                                +--+                           
  |                                   +-- Indonaia_andersoniana **   
--+                                                               
  |  +----------------------------------- Indochinella_pugio      
  |  |                                                            
  |  |                             +----- Radiatula_mouhoti       
  +--+  +--------------------------+                              
     |  |                          |  +-- Radiatula_myitkyinae    
     |  |                          +--+                           
     +--+                             +-- Radiatula_chaudhurii    
        |                                                         
        |  +----------------------------- Scabiellus_songkramensis
        +--+                                                      
           |  +-------------------------- Harmandia_somboriensis  
           +--+                                                   
              |  +----------------------- Unionetta_fabagina      
              +--+                                                
                 |  +-------------------- Scabies_crispata        
                 +--+                                             
                    |  +----------------- Scabies_mandarinus      
                    +--+                                          
                       |  +-------------- Scabies_pilata          
                       +--+                                       
                          |  +----------- Scabies_nucleus         
                          +--+                                    
                             |  +-------- Scabies_humilis         
                             +--+                                 
                                |  +----- Scabies_scobinatus      
                                +--+                              
                                   |  +-- Scabies_phaselus        
                                   +--+                           
                                      +-- Scabies_anceps

Bolotov et al., 2022 [Publ 4250], fig. 8

iT tT bi riS oT roT roS iD
2 2 0 2 47 7 2 -- 
                 +-- Indonaia_andersoniana **  
  +--------------+                          
  |              +-- Indonaia_subclathrata **  
--+                                         
  |  +-------------- Indochinella_pugio     
  |  |                                      
  +--+  +----------- Radiatula_crispisulcata
     |  |                                   
     +--+        +-- Radiatula_chaudhurii   
        |  +-----+                          
        |  |     +-- Radiatula_myitkyinae   
        +--+                                
           |  +----- Radiatula_mouhoti      
           +--+                             
              |  +-- Radiatula_chindwinensis
              +--+                          
                 +-- Radiatula_myitthanensis

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iT tT bi riS oT roT roS iD
1 1 0 1 56 3 3 -- 
  +-------- Indonaia_caerulea **     
--+                               
  |  +----- Indochinella_pugio    
  +--+                            
     |  +-- Scabies_phaselus      
     +--+                         
        +-- Harmandia_somboriensis

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 56 3 3 -- 
  +-------- Indonaia_caerulea **     
--+                               
  |  +----- Indochinella_pugio    
  +--+                            
     |  +-- Scabies_phaselus      
     +--+                         
        +-- Harmandia_somboriensis

Ortiz-Sepulveda et al., 2020 [Publ 4101], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 35 7 3 -- 
  +----------- Indonaia_andersoniana **
  |                                 
--+  +-------- Indochinella_pugio   
  |  |                              
  |  |  +----- Radiatula_mouhoti    
  +--+  |                           
     |  +----- Radiatula_myitkyinae 
     |  |                           
     +--+----- Radiatula_chaudhurii 
        |                           
        |  +-- Scabies_humilis      
        |  |                        
        +--+-- Scabies_phaselus     
           |                        
           +-- Scabies_mandarinus

Ortiz-Sepulveda et al., 2020 [Publ 4101], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 35 7 3 -- 
  +----- Radiatula_mouhoti    
  |                           
  +----- Radiatula_myitkyinae 
  |                           
  +----- Radiatula_chaudhurii 
  |                           
--+----- Indochinella_pugio   
  |                           
  +----- Indonaia_andersoniana **
  |                           
  |  +-- Scabies_humilis      
  |  |                        
  +--+-- Scabies_phaselus     
     |                        
     +-- Scabies_mandarinus

Wu et al., 2021 [Publ 4215], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 23 0 0 -- 
-- Indonaia_andersoniana

Konopleva et al., 2023 [Publ 4262], fig. 8

iT tT bi riS oT roT roS iD
1 1 0 1 83 13 6 -- 
  +----------------------- Indonaia_andersoniana **   
  |                                                
--+  +-------------------- Indochinella_pugio      
  |  |                                             
  +--+  +----------------- Radiatula_mouhoti       
     |  |                                          
     +--+  +-------------- Scabiellus_songkramensis
        |  |                                       
        |  |           +-- Unionetta_fabagina      
        +--+  +--------+                           
           |  |        +-- Harmandia_somboriensis  
           |  |                                    
           |  |        +-- Scabies_humilis         
           +--+  +-----+                           
              |  |     +-- Scabies_pilata          
              |  |                                 
              |  |  +----- Scabies_crispata        
              |  +--+                              
              +--+  |  +-- Scabies_nucleus         
                 |  +--+                           
                 |     +-- Scabies_mandarinus      
                 |                                 
                 |     +-- Scabies_phaselus        
                 |     |                           
                 +-----+-- Scabies_anceps          
                       |                           
                       +-- Scabies_scobinatus

Konopleva et al., 2023 [Publ 4269], fig. S1, 4

iT tT bi riS oT roT roS iD
1 1 0 1 55 2 2 -- 
  +----- Indonaia_andersoniana **
--+                           
  |  +-- Indochinella_pugio   
  +--+                        
     +-- Radiatula_mouhoti

Chen et al., 2023 [Publ 4302], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 39 0 0 -- 
-- Indonaia_andersoniana

Chen et al., 2023 [Publ 4302], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 39 0 0 -- 
-- Indonaia_andersoniana

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