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The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)
FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!
The nominal species (available and otherwise) in this synonymy are listed by usage rather than strictly by publication date. The primary sorting order is by the most recent year (since 1969) that the species name was treated as valid, and then nominal species are sorted secondarily by publication year. The taxonomic history of the valid species (since 1900) is listed at the bottom of the page.
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Unionoida | Unionoidea | Unionidae | Ambleminae | Lampsilini | Epioblasma species Epioblasma torulosa (Rafinesque, 1820): nominal species 5 specimens available.
Amblema torulosa Rafinesque, 1820
Rafinesque, 1820, Ann. Gén. Sci. Phys. 5 (13): 314, pl. 82, fig. 11-12 [also as ‘Unio torulosa’].
Rafinesque, 1864, [in] Binney & Tryon, reprint: 57.
Type(s): HOLOTYPE ANSP_20218, [no locality available - North America].
Amblema torulosa — Rafinesque (1820)
Epioblasma torulosa — Rafinesque (1831), Cummings & Mayer (1992), Cummings & Mayer (1997), Parmalee & Bogan (1998), ..., Williams et al. (2017), Graf & Cummings (2021), FMCS (2021), Hopper et al. (2023)
Unio torulosus — Ferussac (1835), Conrad (1853)
Unio gibbosus — Conrad (1836)
Truncilla torulosa — Vanatta (1915), Ortmann (1918)
Dysnomia (Pilea) torulosa — Ortmann & Walker (1922), Ortmann (1925)
Dysnomia (s.s.) torulosa — Ortmann (1924)
Dysnomia (Torulosa) torulosa — Frierson (1927), Haas (1930), Haas (1969)
Epioblasma (sec. Torulosa) torulosum — Thiele (1934)
Dysnomia (Torulosa) torulosa torulosa — Haas (1969)
Dysnomia torulosa — Stansbery (1970)
Dysnomia torulosa torulosa — Burch (1975)
Plagiola (Torulosa) torulosa — R.I. Johnson (1978)
Plagiola torulosa — R.I. Johnson (1980)
Epioblasma torulosa torulosa — Turgeon et al. (1988), Williams et al. (1993), Bogan (1993), Turgeon et al. (1998), Lydeard et al. (1999), Cicerello & Schuster (2003), Watters et al. (2009), ITIS (2012), IUCN (2012)
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Amblema gibbosa Rafinesque, 1820
Rafinesque, 1818, Amer. Mon. Mag. 3: 355 [as ‘Potamilus gibbosus’]. [nomen nudum]
Rafinesque, 1820, Ann. Gén. Sci. Phys. 5 (13): 315 [also as ‘Unio gibbosa’].
Rafinesque, 1864, [in] Binney & Tryon, reprint: 23, 58.
Type(s): LECTOTYPE ANSP_20232, Riv. Ohio.
Amblema gibbosa — Rafinesque (1820)
Potamilus gibbosus — Rafinesque (1818)
Unio gibbosus — Ferussac (1835), Conrad (1836), Küster (1848), Reeve (1864)
Margarita (Unio) perplexus — Lea (1836), Lea (1838)
Dysnomia gibbosa — Agassiz (1852)
Margaron (Unio) perplexus — Lea (1852), Lea (1870)
Unio torulosus — Conrad (1853)
Unio (Dysnomia) gibbosus — Mörch (1853)
Truncilla torulosa — Vanatta (1915), Ortmann (1918)
Dysnomia (Pilea) torulosa — Ortmann & Walker (1922)
Dysnomia (Torulosa) torulosa — Frierson (1927)
Dysnomia (Torulosa) torulosa torulosa — Haas (1969)
Dysnomia torulosa torulosa — Burch (1975)
Plagiola (Torulosa) torulosa — R.I. Johnson (1978)
Epioblasma torulosa — Parmalee & Bogan (1998), Williams et al. (2008)
Epioblasma torulosa torulosa — Watters et al. (2009)
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Amblema gibbosa var. difformis Rafinesque, 1820
Rafinesque, 1820, Ann. Gén. Sci. Phys. 5 (13): 315.
Rafinesque, 1864, [in] Binney & Tryon, reprint: 58.
(intro'd as intra-sp. of Amblema gibbosa Rafinesque, 1820)
Amblema gibbosa var. difformis — Rafinesque (1820)
Dysnomia (Torulosa) torulosa torulosa — Haas (1969)
Epioblasma torulosa — Parmalee & Bogan (1998), Williams et al. (2008)
Epioblasma torulosa torulosa — Watters et al. (2009)
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Amblema gibbosa var. olivacea Rafinesque, 1820
Rafinesque, 1820, Ann. Gén. Sci. Phys. 5 (13): 315.
Rafinesque, 1864, [in] Binney & Tryon, reprint: 58.
(intro'd as intra-sp. of Amblema gibbosa Rafinesque, 1820)
Amblema gibbosa var. olivacea — Rafinesque (1820)
Dysnomia (Torulosa) torulosa torulosa — Haas (1969)
Epioblasma torulosa — Parmalee & Bogan (1998), Williams et al. (2008)
Epioblasma torulosa torulosa — Watters et al. (2009)
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Amblema gibbosa var. radiata Rafinesque, 1820
Rafinesque, 1820, Ann. Gén. Sci. Phys. 5 (13): 315.
Rafinesque, 1864, [in] Binney & Tryon, reprint: 58.
(intro'd as intra-sp. of Amblema gibbosa Rafinesque, 1820)
Amblema gibbosa var. radiata — Rafinesque (1820)
Dysnomia (Torulosa) torulosa torulosa — Haas (1969)
Epioblasma torulosa — Parmalee & Bogan (1998), Williams et al. (2008)
Epioblasma torulosa torulosa — Watters et al. (2009)
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Amblema torulosa var. angulata Rafinesque, 1820
Rafinesque, 1820, Ann. Gén. Sci. Phys. 5 (13): 315.
Rafinesque, 1864, [in] Binney & Tryon, reprint: 57.
(intro'd as intra-sp. of Amblema torulosa Rafinesque, 1820)
Amblema torulosa var. angulata — Rafinesque (1820)
Dysnomia (Torulosa) torulosa torulosa — Haas (1969)
Plagiola (Torulosa) torulosa — R.I. Johnson (1978)
Epioblasma torulosa — Parmalee & Bogan (1998), Williams et al. (2008)
Epioblasma torulosa torulosa — Watters et al. (2009)
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Unio perplexus Lea, 1831
Lea, 1831, Trans. Amer. Phil. Soc. 4 [O. 1]: 112, pl. 17, fig. 42.
(homonym of Unio perplexus Say, 1829)
Type(s): SYNTYPE USNM_84324, Cincinnati, Ohio.
Unio perplexus — Lea (1831), Ravenel (1834), Jay (1835), Hanley (1843), Lewis (1871), B.H. Wright (1888), de Gregorio (1914)
Unio gibbosus — Ferussac (1835), Conrad (1836), Küster (1848), Reeve (1864)
Margarita (Unio) perplexus — Lea (1836), Lea (1838)
Dysnomia gibbosa — Agassiz (1852)
Margaron (Unio) perplexus — Lea (1852), Lea (1870)
Unio torulosus — Conrad (1853)
Unio (Dysnomia) gibbosus — Mörch (1853)
Unio (Rotundaria) perplexus — Adams & Adams (1857)
Truncilla (Pilea) perplexa — Simpson (1900), Simpson (1914)
Unio (s.s.) perplexus — Call (1900)
Truncilla perplexa — Ortmann (1909)
Truncilla torulosa — Vanatta (1915), Ortmann (1918)
Dysnomia (Pilea) torulosa — Ortmann & Walker (1922)
Dysnomia (Torulosa) torulosa — Frierson (1927)
Dysnomia (Torulosa) torulosa torulosa — Haas (1969)
Dysnomia torulosa torulosa — Burch (1975)
Plagiola (Torulosa) torulosa — R.I. Johnson (1978)
Epioblasma torulosa — Strayer & Jirka (1997), Parmalee & Bogan (1998), Williams et al. (2008)
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Unio torulosus Conrad, 1834
‘Rafinesque’ Conrad, 1834, New Fresh-Water Shells: 72.
‘Raf.’ Conrad, 1853, Proc. Acad. Nat. Sci. 6: 259.
(reference to Amblema torulosa Rafinesque, 1820)
Unio torulosus — Conrad (1834)
Truncilla (Pilea) perplexa — Simpson (1900), Simpson (1914)
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Unio gibbosus Ferussac, 1835
‘Rafinesque’ of Authors.
Ferussac, 1835, Mag. Zool. 5: 28.
Conrad, 1836, Monogr. Fam. Unionidae 1(6): 50, pl. 27, fig. 1.
Reeve, 1864, Conch. Icon. 16: pl. 11, sp. 41, figs. 41a-b.
(reference to Amblema gibbosa Rafinesque, 1820)
Unio gibbosus — Ferussac (1835), Küster (1848)
Unio torulosus — Ferussac (1835)
Margarita (Unio) perplexus — Lea (1836), Lea (1838)
Unio perplexus — Hanley (1843)
Margaron (Unio) perplexus — Lea (1852), Lea (1870)
Truncilla (Pilea) perplexa — Simpson (1900), Simpson (1914)
Dysnomia torulosa torulosa — Burch (1975)
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Unio obliquus Potiez & Michaud, 1844
‘Lamarck’ Potiez & Michaud, 1844, Galérie Moll. at. Mus. Douai 2: 153, pl. 58, figs. 3-4.
(misidentified reference to Unio obliqua Lamarck, 1819)
Unio obliquus — Potiez & Michaud (1844)
Unio (Dysnomia) gibbosus — Mörch (1853)
Truncilla (Pilea) perplexa — Simpson (1900), Simpson (1914)
Dysnomia (Torulosa) torulosa — Frierson (1927)
Dysnomia (Torulosa) torulosa torulosa — Haas (1969)
Dysnomia torulosa torulosa — Burch (1975)
Plagiola (Torulosa) torulosa — R.I. Johnson (1978)
Epioblasma torulosa — Parmalee & Bogan (1998)
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Unio phillipsii Reeve, 1864
‘Conrad’ Reeve, 1864, Conch. Icon. 16: pl. 4, sp. 15.
(misidentified reference to Unio phillipsii Conrad, 1835)
Unio phillipsii — Reeve (1864)
Truncilla (Pilea) perplexa var. cincinnatiensis — Simpson (1900), Simpson (1914)
Plagiola (Torulosa) torulosa — R.I. Johnson (1978)
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Unio abnormis Haas, 1930
‘Say’ Haas, 1930, Senckenbergiana 12(6): 329. [MS]
(intro'd as syn of Amblema torulosa Rafinesque, 1820)
Unio abnormis — Haas (1930)
Dysnomia (Torulosa) torulosa — Haas (1930)
Epioblasma torulosa — Parmalee & Bogan (1998)
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Taxonomic History (since 1900)
Truncilla (Pilea) perplexa — Simpson (1900), Simpson (1914)
Unio (s.s.) perplexus — Call (1900)
Truncilla (Pilea) perplexa var. cincinnatiensis — Simpson (1900), Simpson (1914)
Truncilla perplexa — Ortmann (1909)
Unio perplexus — de Gregorio (1914)
Truncilla torulosa — Vanatta (1915), Ortmann (1918)
Dysnomia (Pilea) torulosa — Ortmann & Walker (1922), Ortmann (1925)
Dysnomia (s.s.) torulosa — Ortmann (1924)
Dysnomia (Torulosa) torulosa — Frierson (1927), Haas (1930), Haas (1969)
Unio abnormis — Haas (1930)
Epioblasma (sec. Torulosa) torulosum — Thiele (1934)
Dysnomia (Torulosa) torulosa torulosa — Haas (1969)
Dysnomia torulosa — Stansbery (1970)
Dysnomia torulosa torulosa — Burch (1975)
Plagiola (Torulosa) torulosa — R.I. Johnson (1978)
Plagiola torulosa — R.I. Johnson (1980)
Epioblasma torulosa torulosa — Turgeon et al. (1988), Williams et al. (1993), Bogan (1993), Turgeon et al. (1998), Lydeard et al. (1999), Cicerello & Schuster (2003), Watters et al. (2009), ITIS (2012), IUCN (2012)
Epioblasma torulosa — Cummings & Mayer (1992), Strayer & Jirka (1997), Cummings & Mayer (1997), Parmalee & Bogan (1998), ..., Williams et al. (2017), Graf & Cummings (2021), FMCS (2021), Hopper et al. (2023)
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Page updated: 15:02:11 Wed 04 Dec 2024 |
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