Mya margaritifera Linnaeus, 1758
Lister, 1678, Hist. Anim. Anglic.: [app.] 15, pl. 1, fig. 1. [pre-linnaean]
Lister, 1685, Conch.: pl. 149, fig. 4. [pre-linnaean]
Linnaeus, 1758, Syst. Nat. 1: 671, sp. 20.
Linnaeus, 1767, Syst. Nat. 1(2): 1112, sp. 29.
Type(s): LECTOTYPE LS_29, “Habitat in totius orbis arctici cataractis.”.

Mya margaritifera — Linnaeus (1758), Linnaeus (1767), Müller (1774), Poiret (1789), ..., Montagu (1803), Turton (1807), Dillwyn (1817), Wood (1825)
Unio margaritiferus — Retzius (1788), Spengler (1793), Hanley (1843), Sowerby (1868)
Unio margaritifera — Cuvier (1798), Poiret (1801), Bosc (1801), Draparnaud (1806), ..., Lea (1834), Griffith & Pidgeon (1834), Held (1836), Kira (1954)
Margartifera margaritifera — Schumacher (1816)
Margaritana fluviatilis — Schumacher (1817)
Unio sinuata — Children (1823)
Unio sinuatus — Menke (1828), Menke (1830), Gassies (1849)
Unio elongatus — Wood (1828)
Alasmodonta margaritifera — Ferussac (1835), C.B. Adams (1847)
Unio margaritifer — Rossmässler (1835), Schlüter (1838), Küster (1855), Malzine (1867)
Margarita (Margaritana) margaritifera — Lea (1836), Lea (1838)
Margaritana margaritifera — Sowerby (1839), Swainson (1840), Dupuy (1852), Lea (1856), ..., Frierson (1927), Germain (1931), Zhadin (1938), Zhadin (1965)
Alasmodon arcuata — De Kay (1843)
Damalis margaritifera — Leach (1847)
Margaron (Margaritana) margaritifera — Lea (1852), Lea (1870)
Damaris elongata — Leach (1852)
Unio (Baphia) margaritifera — Mörch (1853)
Unio (Margaritana) margaritifer — Moquin-Tandon (1855), Drouët (1857), Schrenck (1867)
Baphia margaritifera — Adams & Adams (1857)
Unio (Margaritana s.s.) margaritifera — Fischer (1887)
Margaritana (s.s.) margaritifera — Thiele (1934)
Margaritifera margaritifera — Mandahl-Barth (1938), Modell (1942), Kira (1959), Heard & Guckert (1970), ..., Prié et al. (2023), Hopper et al. (2023), Wu et al. (2023), Wu et al. (2024)
Margaritifera margaritifera margaritifera — Haas (1940), Bogatov (2022)
Margaritifera (s.s.) margaritifera margaritifera — Haas (1969), Falkner et al. (2001), Falkner et al. (2002)
Margaritifera (s.s.) margaritifera — Haas (1969), Bolotov et al. (2016)

Unio elongata Lamarck, 1819
Pennant, 1777, British Zool. 4: pl. 43, fig. 18.
Lamarck, 1819, Hist. Natur. Anim. sans Vert. 6: 70, sp. 2.
Type(s): POSSIBLE SYNTYPE MNHN_MPL_0028, Habite dans les rivières de l’Angleterre, et probablement du nord de l’Europe [Lives in the rivers of England and Northern Europe probably].

Unio elongata — Lamarck (1819), Nilsson (1822), Jay (1835)
Unio margaritifera — Pfeiffer (1821), Lea (1834), Held (1836)
Unio margaritiferus — Menke (1828)
Unio elongatus — Wood (1828), Sowerby (1868)
Alasmodonta margaritifera — Ferussac (1835)
Unio margaritifer — Rossmässler (1835), Küster (1855)
Margarita (Margaritana) margaritifera — Lea (1836), Lea (1838)
Margaritana elongata — Swainson (1840), Locard (1889), Fagot (1892), Locard (1893), Dumas (1895), Locard (1899)
Margaron (Margaritana) margaritifera — Lea (1852), Lea (1870)
Margaritana margaritifera — Dupuy (1852), Drouët (1898), Simpson (1900), Preston (1912), Simpson (1914), Germain (1931)
Damaris elongata — Leach (1852)
Unio (Baphia) margaritifera — Mörch (1853)
Unio (Margaritana) margaritifer var. C elongatus — Moquin-Tandon (1855)
Unio (Margaritana) margaritifer — Drouët (1857)
Margaritana margaritifera var. elongata — Colbeau (1868)
Margaritana margaritifera forma elongata — Westerlund (1890)
Margaritana (Margaritiferana) elongata — Fagot (1893)
Margaritifera margaritifera margaritifera — Haas (1940)
Margaritifera (s.s.) margaritifera margaritifera — Haas (1969)
Margaritifera margaritifera — Ziuganov et al. (1994), Graf (2007), Araujo et al. (2009), Watters et al. (2009), Kantor et al. (2010), Fauna Europaea (2012), Bolotov et al. (2013), Vinarski & Kantor (2016)
Margaritifera elongata — Bogatov et al. (2003), Starobogatov et al. (2004), Kantor & Sysoev (2005)
Margaritifera margaritifera elongata — Bogatov (2022)

Margaritana borealis Westerlund, 1871
Westerlund, 1871, Nova Reg. Soc. Scient. 8: 168.

Margaritana borealis — Westerlund (1871)
Margaritana margaritifera forma borealis — Westerlund (1890)
Margaritifera borealis — Bogatov et al. (2003), Starobogatov et al. (2004), Kantor & Sysoev (2005)
Margaritifera margaritifera — Graf (2007), Kantor et al. (2010), Bolotov et al. (2013), Vinarski & Kantor (2016)
Margaritifera margaritifera borealis — Bogatov (2022)

Margaritana durrovensis Phillips, 1928
Phillips, 1928, Proc. Malac. Soc. London 18: 71, pls. 3-5.
Bloomer, 1928, Proc. Malac. Soc. London 18: 74, text fig.
Type(s): SYNTYPES BMNH_1928-10-23-1-2, River Nore, Durrow, Queen’s Co., Ireland.

Margaritana durrovensis — Phillips (1928)
Margaritifera margaritifera margaritifera — Haas (1940)
Margaritifera (s.s.) margaritifera margaritifera — Haas (1969)
Margaritifera margaritifera — Ziuganov et al. (1994), Araujo et al. (2009), Vinarski & Kantor (2016), Lopes-Lima et al. (2017)
Margaritifera margaritifera durrovensis — IUCN (1996), Machordom et al. (2003), Walker et al. (2006)
Margaritifera (s.s.) margaritifera durrovensis — Falkner et al. (2001)

Unio brunneus Bonhomme, 1840
Bonhomme, 1840, Mém. Aveyron 2: 460.

Unio brunneus — Bonhomme (1840)
Margaron (Unio) litoralis — Lea (1852), Lea (1870)
Margaritana margaritifera — Dupuy (1852), Westerlund (1890), Germain (1931)
Unio (Margaritana) margaritifer var. D minor — Moquin-Tandon (1855)
Unio (Margaritana) margaritifer var. minor — Drouët (1857)
Unio litoralis — Reeve (1865)
Margaritana brunnea — Locard (1889), Locard (1893)
Margaritifera margaritifera margaritifera — Haas (1940)
Margaritifera (s.s.) margaritifera margaritifera — Haas (1969)
Margaritifera (s.s.) margaritifera brunnea — Falkner et al. (2001), Falkner et al. (2002)
Margaritifera margaritifera — Araujo et al. (2009), Vinarski & Kantor (2016), Lopes-Lima et al. (2017)

Margaritana margaritifera parvula Haas, 1908
Haas, 1908, Beitr. Kenntn. Mittleleurop. Najadeen, Beilage zu... 1908 (1): 15.
(intro'd as intra-sp. of Mya margaritifera Linnaeus, 1758)
Type(s): LECTOTYPE SMF_4982, Ulfenbach bei Affolterbach.

Margaritana margaritifera parvula — Haas (1908), Haas (1924)
Margaritifera margaritifera margaritifera — Haas (1940)
Margaritifera (s.s.) margaritifera margaritifera — Haas (1969)
Margaritifera (s.s.) margaritifera parvula — Falkner et al. (2001)
Margaritifera margaritifera — Vinarski & Kantor (2016), Lopes-Lima et al. (2017)

Mya margaritifera Da Costa, 1778
Da Costa, 1778, British Conch.: 225, pl. 15, fig. 3.
(reference to Mya margaritifera Linnaeus, 1758)

Mya margaritifera — Da Costa (1778), Turton (1807), Dillwyn (1817)
Unio elongata — Lamarck (1819)
Unio margaritiferus — Menke (1828)
Margaritana elongata — Swainson (1840), Locard (1889), Locard (1899)
Margaron (Margaritana) margaritifera — Lea (1852), Lea (1870)

Margaritana fluviatilis Schumacher, 1817
Schumacher, 1816, Overs. Dansk. Vid. Selsk. Forhandl. Kjobehhaven 1816: 7. [nomen nudum]
Schumacher, 1817, Essai. Nouv. Syst.: 124, pl. 10, fig. 4.

Margaritana fluviatilis — Schumacher (1817)
Margartifera fluviatilis — Schumacher (1816)
Margarita (Margaritana) margaritifera — Lea (1836), Lea (1838)
Margaron (Margaritana) margaritifera — Lea (1852), Lea (1870)
Unio (Margaritana) margaritifer — Drouët (1857)
Margaritana margaritifera — Locard (1882), Westerlund (1890)
Margaritifera margaritifera — Habe (1977), Ziuganov et al. (1994), Watters et al. (2009), Vinarski & Kantor (2016)

Alasmodonta arcuata Barnes, 1823
Barnes, 1823, Amer. J. Sci. 6 (1-2): 277, pl. 12, figs. 20a-d.

Alasmodonta arcuata — Barnes (1823), Ravenel (1834), Linsley (1845)
Unio margaritifera — Lea (1834)
Alasmodonta margaritifera — Ferussac (1835)
Margarita (Margaritana) margaritifera — Lea (1836), Lea (1838)
Margaritana arcuata — Swainson (1840), Stimpson (1851), Agassiz (1852), Conrad (1853), Küster (1862)
Alasmodon arcuata — De Kay (1843)
Margaron (Margaritana) margaritifera — Lea (1852), Lea (1870)
Margaritana margaritifera — Dupuy (1852), Simpson (1900), Hannibal (1912), Preston (1912), Simpson (1914), Frierson (1927)
Unio margaritifer — Küster (1855)
Margaritifera (s.s.) margaritifera margaritifera — Haas (1969)
Margaritifera margaritifera — Ziuganov et al. (1994), Strayer & Jirka (1997), Araujo et al. (2009), Watters et al. (2009), Vinarski & Kantor (2016)

Unio sinuata C. Pfeiffer, 1825
C. Pfeiffer, 1825, Naturgesch. Dtsch. Land-Süßw. Moll. 2: .
(misidentified reference to Unio sinuata Lamarck, 1819)

Unio sinuata — C. Pfeiffer (1825)
Unio margaritifer — Rossmässler (1835), Küster (1855)
Margarita (Margaritana) margaritifera — Lea (1836), Lea (1838)
Unio margaritifer var. minor — Schlüter (1838)
Unio (Margaritana) margaritifer — Drouët (1857)
Margaritana margaritifera — Clessin (1877), Westerlund (1890), Simpson (1900), Preston (1912), Simpson (1914)

Unio elongata Michaud, 1831
Michaud, 1831, Compl. Hist. Nat. Moll., Draparnaud: 113, pl. 16, fig. 29.
(reference to Unio elongata Lamarck, 1819)

Unio elongata — Michaud (1831)
Unio margaritifer var. minor — Rossmässler (1835)
Unio margaritifer — Küster (1855)
Unio sinuata — Sowerby (1868)
Margaritana michaudi — Locard (1889)
Margaritana margaritifera — Westerlund (1890), Germain (1931)
Unio (sec. Lymnium) batavus — Simpson (1900), Simpson (1914)

Unio roissyi Michaud, 1831
Michaud, 1831, Compl. Hist. Nat. Moll., Draparnaud: 112, pl. 16, figs. 27-28.

Unio roissyi — Michaud (1831)
Alasmodonta margaritifera — Ferussac (1835)
Margarita (Margaritana) margaritifera — Lea (1836), Lea (1838)
Margaritana margaritifera — Dupuy (1852), Westerlund (1890), Simpson (1900), Preston (1912), Simpson (1914), Germain (1931)
Unio (Margaritana) margaritifer var. B roissyi — Moquin-Tandon (1855)
Unio (Margaritana) margaritifer var. roissyi — Drouët (1857)
Unio sinuata — Sowerby (1868)
Margaron (Margaritana) margaritifera — Lea (1870)
Margaritana roissyi — Locard (1889), Locard (1893), Rochebrune (1903)
Margaritifera margaritifera margaritifera — Haas (1940)
Margaritifera (s.s.) margaritifera margaritifera — Haas (1969)
Margaritifera margaritifera — Ziuganov et al. (1994), Watters et al. (2009), Araujo et al. (2009), Vinarski & Kantor (2016)

Unio margaritifer var. minor Rossmässler, 1835
Rossmässler, 1835, Icon. Land- und Süssw.-Moll. 1 (2): 21, pl. 9, fig. 129.
(intro'd as intra-sp. of Mya margaritifera Linnaeus, 1758)

Unio margaritifer var. minor — Rossmässler (1835), Schlüter (1838)
Unio (Margaritana) margaritifer var. D minor — Moquin-Tandon (1855)
Unio (Margaritana) margaritifer var. minor — Drouët (1857)
Margaritana margaritifera — Drouët (1881), Simpson (1900), Preston (1912), Simpson (1914), Germain (1931)
Margaritana brunnea — Locard (1889)
Margaritifera margaritifera margaritifera — Haas (1940)
Margaritifera margaritifera — Watters et al. (2009), Vinarski & Kantor (2016)

Unio rivalis Lea, 1836
‘Zeig.’ Lea, 1836, Synopsis Fam. Naiades: 46.
‘Zeig.’ Lea, 1838, Synopsis Fam. Naiades: 28.
(intro'd as syn of Mya margaritifera Linnaeus, 1758)

Unio rivalis — Lea (1836)
Margarita (Margaritana) margaritifera — Lea (1836), Lea (1838)
Margaron (Margaritana) margaritifera — Lea (1852), Lea (1870)

Unio tristis Morelet, 1845
Morelet, 1845, Moll. Terr. Fluv. Portugal: 107, pl. 31, fig. 2.
Type(s): HOLOTYPE BMNH_93-2-4-1961, la Tamega, pres a Amarante, Portugal.

Unio tristis — Morelet (1845)
Margaron (Unio) batavus — Lea (1852), Lea (1870)
Margaritana tristis — Bourguignat (1876)
Margaritana elongata — Locard (1899)
Margaritana margaritifera — Simpson (1900), Preston (1912), Simpson (1914)
Margaritifera margaritifera margaritifera — Haas (1940)
Margaritifera (s.s.) margaritifera margaritifera — Haas (1969)
Margaritifera margaritifera — Ziuganov et al. (1994), Watters et al. (2009), Araujo et al. (2009), Vinarski & Kantor (2016)

Unio roisii Lea, 1852
‘Mich.’ Lea, 1852, Synopsis Fam. Naiades: 40.
(error for Unio roissyi Michaud, 1831)

Unio roisii — Lea (1852)

Unio raveneliana Chenu, 1859
‘Lea’ Chenu, 1859, Manuel Conchyl. 2: 144, fig. 714.
(misidentified reference to Unio ravenelianus Lea, 1834)

Unio raveneliana — Chenu (1859)
Margaritana margaritifera — Simpson (1900), Preston (1912), Simpson (1914)

Unio reyniesii Lea, 1870
‘Mich.’ Lea, 1870, Synopsis Fam. Naiades: 64.
(error for Unio roissyi Michaud, 1831)

Unio reyniesii — Lea (1870)

Margaritana margaritifera var. compressa Westerlund, 1873
Westerlund, 1873, Fauna Moll. Terr. Fluv. Suec. Norv. Dan. (2): 578.
(intro'd as intra-sp. of Mya margaritifera Linnaeus, 1758)

Margaritana margaritifera var. compressa — Westerlund (1873)
Margaritana margaritifera forma compressa — Westerlund (1890)
Margaritifera margaritifera — Vinarski & Kantor (2016)

Margaritana margaritifera var. ventricosa Westerlund, 1873
Westerlund, 1873, Fauna Moll. Terr. Fluv. Suec. Norv. Dan. (2): 578.
(intro'd as intra-sp. of Mya margaritifera Linnaeus, 1758)

Margaritana margaritifera var. ventricosa — Westerlund (1873)
Margaritana margaritifera forma ventricosa — Westerlund (1890)
Margaritifera margaritifera — Vinarski & Kantor (2016)

Margaritana freytagi Kobelt, 1886
Kobelt, 1886, Jb. Nassau. Ver. Naturk. 39: 98, pl. 8, figs. 1-4.
Kobelt, 1886, Icon. Land- und Süssw.-Moll. 2: 45, pl. 55, figs. 291-294.
Type(s): LECTOTYPE SMF_4984, Nister b. Hachenburg.

Margaritana freytagi — Kobelt (1886), Kobelt (1886), Westerlund (1890)
Margaritifera margaritifera margaritifera — Haas (1940)
Margaritifera (s.s.) margaritifera margaritifera — Haas (1969)
Margaritifera margaritifera — Ziuganov et al. (1994), Araujo et al. (2009), Vinarski & Kantor (2016)

Margaritana michaudi Locard, 1889
Michaud, 1831, Compl. Hist. Nat. Moll., Draparnaud: 113, pl. 16, fig. 29 [as ‘Unio elongata’].
Locard, 1889, Contr. Faune Malacol. Franc. 13: 123 [17], 180 [74].
Locard, 1893, Coqu. Eau Douce Saumâtre France: 150.
Type(s): PARALECTOTYPE MNHN_22368, Vire [Calvados].

Margaritana michaudi — Locard (1889), Locard (1893), Dumas (1895)
Margaritana margaritifera — Westerlund (1890), Germain (1931)
Margaritifera margaritifera margaritifera — Haas (1940)
Margaritifera (s.s.) margaritifera margaritifera — Haas (1969)
Margaritifera margaritifera — Ziuganov et al. (1994), Araujo et al. (2009), Vinarski & Kantor (2016)

Margaritana pyrenaica Locard, 1889
Bourguignat in Locard, 1889, Contr. Faune Malacol. Franc. 13: 123 [17], 181 [75].
‘Bourguignat’ Locard, 1893, Coqu. Eau Douce Saumâtre France: 150.
Type(s): SYNTYPE MNHN_22478, Carcenac (Aveyron).

Margaritana pyrenaica — Locard (1889), Locard (1893)
Margaritana margaritifera — Westerlund (1890), Germain (1931)
Margaritifera margaritifera margaritifera — Haas (1940)
Margaritifera (s.s.) margaritifera margaritifera — Haas (1969)
Margaritifera margaritifera — Ziuganov et al. (1994), Araujo et al. (2009), Vinarski & Kantor (2016)

Margaritana margaritifera forma jetschini Westerlund, 1890
Westerlund, 1890, Fauna Paläarct. Binnenconch. 2(7): 185. [infra-subsp.]
(intro'd as intra-sp. of Mya margaritifera Linnaeus, 1758)

Margaritana margaritifera forma jetschini — Westerlund (1890)

Margaritana alleni Locard, 1899
Castro in Locard, 1899, Conchyliol. Portug.: 212.

Margaritana alleni — Locard (1899)
Margaritifera margaritifera margaritifera — Haas (1940)
Margaritifera (s.s.) margaritifera margaritifera — Haas (1969)
Margaritifera margaritifera — Ziuganov et al. (1994), Araujo et al. (2009), Vinarski & Kantor (2016)

Margaritana margaritifera var. siluriana Bloomer, 1927
Bloomer, 1927, Proc. Malac. Soc. London 17: 211.
(intro'd as intra-sp. of Mya margaritifera Linnaeus, 1758)

Margaritana margaritifera var. siluriana — Bloomer (1927)
Margaritifera (s.s.) margaritifera margaritifera — Haas (1969)
Margaritifera margaritifera — Vinarski & Kantor (2016)

Margaritana durrovensis Bloomer, 1928
Bloomer, 1928, Proc. Malac. Soc. London 18: 74, text fig.
(reference to Margaritana durrovensis Phillips, 1928)

Margaritana durrovensis — Bloomer (1928)
Margaritifera (s.s.) margaritifera margaritifera — Haas (1969)
Margaritifera margaritifera — Fauna Europaea (2012)

Margaritana durrowensis Haas, 1940
‘Phillips’ Haas, 1940, Zool. Ser. Field Mus. Nat. Hist. 24: 120.
(error for Margaritana durrovensis Phillips, 1928)

Margaritana durrowensis — Haas (1940)

Margaritana tenerus Ziuganov et al., 1994
Ziuganov et al., 1994, Freshw. Pearl Muss. & Fish: 59.
(intro'd as syn of Mya margaritifera Linnaeus, 1758)

Margaritana tenerus — Ziuganov et al. (1994)
Margaritifera margaritifera — Ziuganov et al. (1994)

Margaritana vulgaris Ziuganov et al., 1994
Ziuganov et al., 1994, Freshw. Pearl Muss. & Fish: 59.
(intro'd as syn of Mya margaritifera Linnaeus, 1758)

Margaritana vulgaris — Ziuganov et al. (1994)
Margaritifera margaritifera — Ziuganov et al. (1994)

Margaritana pirenaica Vinarski & Kantor, 2016
‘Boruguignat in Locard’ Vinarski & Kantor, 2016, Analyt. Cat. Fr. Brack. Moll. Russia: 59.
(error for Margaritana pyrenaica Locard, 1889)

Margaritana pirenaica — Vinarski & Kantor (2016)

Taxonomic History (since 1900)

Margaritana margaritifera — Simpson (1900), Kobelt (1904), Ortmann (1909), Ortmann (1910), ..., Frierson (1927), Germain (1931), Zhadin (1938), Zhadin (1965)
Unio (sec. Lymnium) batavus — Simpson (1900), Simpson (1914)
Margaritana roissyi — Rochebrune (1903)
Margaritana margaritifera parvula — Haas (1908), Haas (1924)
Margaritana margaritifera var. siluriana — Bloomer (1927)
Margaritana durrovensis — Bloomer (1928), Phillips (1928)
Margaritana (s.s.) margaritifera — Thiele (1934)
Margaritifera margaritifera — Mandahl-Barth (1938), Modell (1942), Kira (1959), Heard & Guckert (1970), ..., Prié et al. (2023), Hopper et al. (2023), Wu et al. (2023), Wu et al. (2024)
Margaritifera margaritifera margaritifera — Haas (1940), Bogatov (2022)
Margaritana durrowensis — Haas (1940)
Unio margaritifera — Kira (1954)
Margaritifera (s.s.) margaritifera margaritifera — Haas (1969), Falkner et al. (2001), Falkner et al. (2002)
Margaritifera (s.s.) margaritifera — Haas (1969), Bolotov et al. (2016)
Margaritana tenerus — Ziuganov et al. (1994)
Margaritana vulgaris — Ziuganov et al. (1994)
Margaritifera margaritifera durrovensis — IUCN (1996), Machordom et al. (2003), Walker et al. (2006)
Margaritifera (s.s.) margaritifera brunnea — Falkner et al. (2001), Falkner et al. (2002)
Margaritifera (s.s.) margaritifera parvula — Falkner et al. (2001)
Margaritifera (s.s.) margaritifera durrovensis — Falkner et al. (2001)
Margaritifera elongata — Bogatov et al. (2003), Starobogatov et al. (2004), Kantor & Sysoev (2005)
Margaritifera borealis — Bogatov et al. (2003), Starobogatov et al. (2004), Kantor & Sysoev (2005)
Margaritana pirenaica — Vinarski & Kantor (2016)
Margaritifera margaritifera elongata — Bogatov (2022)
Margaritifera margaritifera borealis — Bogatov (2022)