Obliquaria (Ellipsaria) verrucosa Rafinesque, 1820
Rafinesque, 1818, Amer. Mon. Mag. 3: 355 [as ‘Potamilus verrucosus’]. [nomen nudum]
Rafinesque, 1819, Jour. Phys. Chim. Hist. Nat. 88: 426 [as ‘U. (Plagiola) verrucosa’]. [nomen nudum]
Rafinesque, 1820, Ann. Gén. Sci. Phys. 5 (13): 304, pl. 81, figs. 10-12 [also as ‘U. verrucosa’].
Rafinesque, 1864, [in] Binney & Tryon, reprint: 23, 29, 48.
Type(s): LECTOTYPE ANSP_20235, Ohio River.

Obliquaria (Ellipsaria) verrucosa — Rafinesque (1820)
Potamilus verrucosus — Rafinesque (1818)
Unio (Plagiola) verrucosa — Rafinesque (1819)
Unio verrucosa — Ferussac (1835), Sowerby (1868)
Tritogonia verrucosa — Agassiz (1852), Vanatta (1915), F.C. Baker (1928), Murray & Leonard (1962), ..., FMCS (2021), Graf & Cummings (2021), Hopper et al. (2023), Hauck et al. (2023)
Unio verrucosus — Conrad (1853), Küster (1862)
Unio (Tritogenia) verrucosus — Mörch (1853)
Unio (Iridea) verrucosa — Adams & Adams (1857)
Unio (Limnium sec. Ellipsaria) verrucosus — Fischer (1887)
Quadrula verrucosa — Utterback (1915), Ortmann (1918), Ortmann & Walker (1922), Ortmann (1924), ..., Marshall et al. (2018), Huang et al. (2019), Jeratthitikul et al. (2022), Neemuchwala et al. (2023)
Quadrula (Tritogonia) verrucosa — Frierson (1927)

Unio verrucosus Say, 1834
‘Rafinesque’ of Authors.
‘Rafinesque’ Say, 1834, Amer. Conch. 6: ??.
‘Rafinesque’ Sowerby, 1868, Conch. Icon. 16: pl. 1, sp. 4.
(reference to Obliquaria (Ellipsaria) verrucosa Rafinesque, 1820)

Unio verrucosus — Say (1834)
Tritogonia tuberculata — Simpson (1900), Simpson (1914)
Quadrula (Cyclonaias) tuberculata — Haas (1930)
Tritogonia verrucosus — Haas (1969)
Tritogonia verrucosa — Howells et al. (1996), Watters et al. (2009)

Unio conjugans B.H. Wright, 1899
B.H. Wright, 1899, Nautilus 13(8): 89.
Simpson, 1900, Proc. Acad. Nat. Sci. 52: 79, pl. 4, fig. 1.
Type(s): HOLOTYPE USNM_159501, Hiawassa River, Polk Co., Tenn.

Unio conjugans — B.H. Wright (1899)
Tritogonia conjugans — Simpson (1900), Simpson (1900), Simpson (1914)
Quadrula verrucosa — Ortmann (1918), Williams et al. (2008)
Quadrula (s.s.) conjugans — Frierson (1927)
Quadrula (s.s.) quadrula conjugans — Haas (1969)
Tritogonia verrucosa — Parmalee & Bogan (1998), Watters et al. (2009), Lopes-Lima et al. (2019)

Unio tuberculatus Barnes, 1823
Barnes, 1823, Amer. J. Sci. 6 (1-2): 125, pl. 7, figs. 8a-b.
(renamed Quadrula tritogonia Ortmann, 1909)

Unio tuberculatus — Barnes (1823), Hildreth (1828), Lea (1829), Ravenel (1834), ..., Singley (1893), Call (1895), Pilsbry & Rhoads (1896), de Gregorio (1914)
Unio verrucosa — Ferussac (1835), Sowerby (1868)
Margarita (Unio) tuberculatus — Lea (1836), Lea (1838)
Tritogonia verrucosa — Agassiz (1852), Vanatta (1915), F.C. Baker (1928), Murray & Leonard (1962), ..., Strayer & Jirka (1997), Parmalee & Bogan (1998), Watters et al. (2009), Lopes-Lima et al. (2019)
Margaron (Unio) tuberculatus — Lea (1852), Lea (1870)
Unio verrucosus — Conrad (1853), Küster (1862)
Unio (Tritogenia) verrucosus — Mörch (1853)
Unio (Limnium sec. Tritogonia) tuberculatus — Fischer (1887)
Tritogonia tuberculata — Simpson (1900), Ortmann (1909), Simpson (1914), Thiele (1934)
Unio (s.s.) tuberculatus — Call (1900)
Quadrula tritogonia — Ortmann (1909), Ortmann (1910)
Quadrula tuberculata — Ortmann (1912)
Quadrula verrucosa — Utterback (1915), Ortmann & Walker (1922), Williams et al. (2008)
Quadrula (Tritogonia) verrucosa — Frierson (1927)

Unio (Theliderma) pustulata Swainson, 1840
Swainson, 1840, Treat. Malacol.: 271, fig. 54d; 273.
(homonym of Unio pustulatus Lea, 1831)

Unio (Theliderma) pustulata — Swainson (1840)
Tritogonia pustulata — Agassiz (1852)
Margaron (Unio) tuberculatus — Lea (1852), Lea (1870)
Unio (Tritogenia) verrucosus — Mörch (1853)
Unio verrucosa — Sowerby (1868)
Tritogonia tuberculata — Simpson (1900), Simpson (1914)
Quadrula tritogonia — Ortmann (1909)
Quadrula (Tritogonia) verrucosa — Frierson (1927)
Tritogonia verrucosa — Haas (1969), Parmalee & Bogan (1998), Watters et al. (2009), Lopes-Lima et al. (2019)
Quadrula verrucosa — Williams et al. (2008)

Unio gigas Sowerby, 1867
‘Swainson’ Sowerby, 1867, Conch. Icon. 16: pl. 56, sp. 287.
(misidentified reference to Unio gigas Swainson, 1824)
Type(s): MANUSCRIPT NAME BMNH_1968861, River Ohio.

Unio gigas — Sowerby (1867)
Tritogonia tuberculata — Simpson (1900), Simpson (1914)
Unio (s.s.) multiplicatus — Call (1900)
Quadrula tritogonia — Ortmann (1909)
Hyriopsis cumingii — Haas (1924)
Quadrula (Tritogonia) verrucosa — Frierson (1927)
Tritogonia verrucosa — Haas (1969), Parmalee & Bogan (1998), Watters et al. (2009)

Tritogonia tuberculata var. obesa Simpson, 1900
Simpson, 1900, Proc. U.S. Nat. Mus. 22: 609.
Simpson, 1914, Descr. Cat. Naiades: 320.
(intro'd as intra-sp. of Unio tuberculatus Barnes, 1823)
Type(s): LECTOTYPE USNM_125669, Neches River, Big Eddy, Texas.

Tritogonia tuberculata var. obesa — Simpson (1900), Simpson (1914)
Quadrula (s.s.) quadrula-aspera — Frierson (1927)
Tritogonia verrucosa — Haas (1969), Parmalee & Bogan (1998), Watters et al. (2009), Lopes-Lima et al. (2019)

Quadrula tritogonia Ortmann, 1909
Ortmann, 1909, Nautilus 22(10): 101.
Ortmann, 1909, Ann. Carnegie Mus. 5: 193.
(new name for Unio tuberculatus Barnes, 1823)

Quadrula tritogonia — Ortmann (1909), Ortmann (1909), Ortmann (1910)
Tritogonia tuberculata — Simpson (1914)
Tritogonia verrucosa — Parmalee & Bogan (1998), Watters et al. (2009), Lopes-Lima et al. (2019)
Quadrula verrucosa — Williams et al. (2008)

Quadrula parkeri Geiser, 1911
Geiser, 1911, The Academician 1: 15.

Quadrula parkeri — Geiser (1911)
Tritogonia tuberculata — Simpson (1914)
Quadrula (Tritogonia) verrucosa — Frierson (1927)
Tritogonia verrucosa — Haas (1969), Parmalee & Bogan (1998), Watters et al. (2009), Lopes-Lima et al. (2019)
Quadrula verrucosa — Williams et al. (2008)

Unio tuberculatus var. supraspeciosus de Gregorio, 1914
de Gregorio, 1914, Il Nat. Siciliano 2: 42, pl. 6, figs. 2a-d [as ‘supra speciosus’].
(intro'd as intra-sp. of Unio tuberculatus Barnes, 1823)

Unio tuberculatus var. supraspeciosus — de Gregorio (1914)
Unio tuberculatus var. supra speciosus — de Gregorio (1914)

Unio (Theliderma) pustululata Lopes-Lima et al., 2019
‘Swainson’ Lopes-Lima et al., 2019, Zool. Scripta 48(3): 7 [Suppl. Appendix 2].
(error for Unio (Theliderma) pustulata Swainson, 1840)

Unio (Theliderma) pustululata — Lopes-Lima et al. (2019)

Taxonomic History (since 1900)

Tritogonia tuberculata — Simpson (1900), Ortmann (1909), Simpson (1914), Thiele (1934)
Unio (s.s.) tuberculatus — Call (1900)
Unio (s.s.) multiplicatus — Call (1900)
Tritogonia conjugans — Simpson (1900), Simpson (1900), Simpson (1914)
Tritogonia tuberculata var. obesa — Simpson (1900), Simpson (1914)
Quadrula tritogonia — Ortmann (1909), Ortmann (1909), Ortmann (1910)
Quadrula parkeri — Geiser (1911)
Quadrula tuberculata — Ortmann (1912)
Unio tuberculatus — de Gregorio (1914)
Unio tuberculatus var. supraspeciosus — de Gregorio (1914)
Unio tuberculatus var. supra speciosus — de Gregorio (1914)
Quadrula verrucosa — Utterback (1915), Ortmann (1918), Ortmann & Walker (1922), Ortmann (1924), ..., Marshall et al. (2018), Huang et al. (2019), Jeratthitikul et al. (2022), Neemuchwala et al. (2023)
Tritogonia verrucosa — Vanatta (1915), F.C. Baker (1928), Murray & Leonard (1962), Haas (1969), ..., FMCS (2021), Graf & Cummings (2021), Hopper et al. (2023), Hauck et al. (2023)
Hyriopsis cumingii — Haas (1924)
Quadrula (Tritogonia) verrucosa — Frierson (1927)
Quadrula (s.s.) conjugans — Frierson (1927)
Quadrula (s.s.) quadrula-aspera — Frierson (1927)
Quadrula (Cyclonaias) tuberculata — Haas (1930)
Tritogonia verrucosus — Haas (1969)
Quadrula (s.s.) quadrula conjugans — Haas (1969)
Unio (Theliderma) pustululata — Lopes-Lima et al. (2019)