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Page last updated
Wed 04 Dec 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida

superfamily HYRIOIDEA Swainson, 1840: cladomics
     1 family

Cladomics Summary

number of species = 129
number of subtaxa (rank: family) = 1

analyzed in 99 cladograms in 51 publications
maximum number of terminals in an analysis = 19, mean = 3.92
maximum number of subtaxa (family) in an analysis = 1, mean = 1.0

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 57
maximum number of included terminals = 19, mean = 6.07
maximum number of subtaxa (family) = 1, mean = 1.0
proportion of cladograms that support basic monophyly = 1.0


Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ tree


Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iT tT bi riS oT roT roS iD
2 2 0 1 78 77 3 --

*** Top Monophyly Cladogram *** *** Top Sister-Group Cladogram ***

  +----------------------------------- Hyriidae **        
  |                                                    
--+  +-------------------------------- Margaritiferidae
  |  |                                                 
  +--+  +----------------------------- Etherioidea     
     |  |                                              
     +--+  +-------------------------- Parreysiinae    
        |  |                                           
        |  |  +----------------------- Unioninae       
        +--+  |                                        
           |  |              +-------- Rectidentini    
           |  |  +-----------+                         
           +--+  |           |  +----- Lens            
              |  |           +--+                      
              |  |              |  +-- Physunio        
              |  |              +--+                   
              +--+                 +-- unionoid        
                 |                                     
                 |  +----------------- Ambleminae      
                 |  |                                  
                 |  |              +-- Chamberlainiini 
                 +--+  +-----------+                   
                    |  |           +-- Lamprotulini    
                    |  |                               
                    +--+           +-- Parvasolenaia   
                       |  +--------+                   
                       |  |        +-- Pseudodontini   
                       +--+                            
                          |  +-------- Sinosolenaia    
                          +--+                         
                             |  +----- Gonidea         
                             +--+                      
                                |  +-- Microcondylaea  
                                +--+                   
                                   +-- Leguminaia      

Santos-Neto et al., 2016 [Publ 3739], fig. 2a_b

iT tT bi riS oT roT roS iD
19 19 20 1 1 0 0 --
-- (Hyriidae

Santos-Neto et al., 2016 [Publ 3739], fig. 4

iT tT bi riS oT roT roS iD
19 19 21 1 1 0 0 --
-- (Hyriidae

Huang et al., 2019 [Publ 3843], fig. S3

iT tT bi riS oT roT roS iD
14 14 12 1 182 180 2 --
  +----- Hyriidae **   
--+                 
  |  +-- Etherioidea
  +--+              
     +-- Unionoidea 

Graf et al., 2015 [Publ 3619], fig. 1

iT tT bi riS oT roT roS iD
14 14 12 1 6 5 2 --
  +----- Hyriidae **   
--+                 
  |  +-- Unionoidea 
  +--+              
     +-- Etherioidea

Huang et al., 2019 [Publ 3843], fig. S4

iT tT bi riS oT roT roS iD
14 14 9 1 184 182 2 --
  +----- Etherioidea
--+                 
  |  +-- Hyriidae **   
  +--+              
     +-- Unionoidea 

Huang et al., 2019 [Publ 3843], fig. 2

iT tT bi riS oT roT roS iD
14 14 12 1 184 182 2 --
  +----- Hyriidae **   
--+                 
  |  +-- Etherioidea
  +--+              
     +-- Unionoidea 

Walker et al., 2006 [Publ 2666], fig. 6

iT tT bi riS oT roT roS iD
10 10 7 1 26 25 2 --
  +----- Hyriidae **   
--+                 
  |  +-- Etherioidea
  +--+              
     +-- Unionoidea 

Marshall et al., 2014 [Publ 3593], fig. 1

iT tT bi riS oT roT roS iD
10 10 8 1 4 4 1 --
     +-- Margaritiferidae
  +--+                   
  |  +-- Unioninae       
--+                      
  |  +-- Ambleminae      
  +--+                   
     +-- Hyriidae **        

Graf & Cummings, 2006 [Publ 1968], fig. 2

iT tT bi riS oT roT roS iD
9 9 7 1 38 37 2 --
  +----- Unionoidea 
--+                 
  |  +-- Hyriidae **   
  +--+              
     +-- Etherioidea

Hoeh et al., 2009 [Publ 2716], fig. 5

iT tT bi riS oT roT roS iD
9 9 7 1 37 36 2 --
  +----------------- Parreysiinae    
  |                                  
  |           +----- Margaritiferidae
--+  +--------+                      
  |  |        |  +-- Hyriidae **        
  |  |        +--+                   
  +--+           +-- Etherioidea     
     |                               
     |  +----------- Pseudodontini   
     +--+                            
        |  +-------- Contradentini   
        +--+                         
           |  +----- Unioninae       
           +--+                      
              |  +-- Gonideini       
              +--+                   
                 +-- Ambleminae      

Bolotov et al., 2017 [Publ 3761], fig. S1

iT tT bi riS oT roT roS iD
9 9 7 1 130 128 2 --
  +-------- Margaritiferidae
--+                         
  |  +----- Unionidae       
  +--+                      
     |  +-- Etherioidea     
     +--+                   
        +-- Hyriidae **        

Hoeh et al., 2009 [Publ 2716], fig. 3

iT tT bi riS oT roT roS iD
9 9 7 1 38 37 2 --
  +-------------------------- Hyriidae **        
  |                                           
  |                 +-------- Iridinidae      
--+  +--------------+                         
  |  |              |  +----- Etheria         
  |  |              +--+                      
  |  |                 |  +-- Mulleria        
  +--+                 +--+                   
     |                    +-- Mycetopodidae   
     |                                        
     |  +-------------------- Margaritiferidae
     +--+                                     
        |  +----------------- Parreysiinae    
        +--+                                  
           |  +-------------- Pseudomulleria  
           +--+                               
              |  +----------- Unioninae       
              +--+                            
                 |  +-------- Contradentini   
                 +--+                         
                    |  +----- Ambleminae      
                    +--+                      
                       |  +-- Gonideini       
                       +--+                   
                          +-- Pseudodontini   

Bolotov et al., 2016 [Publ 3743], fig. 4

iT tT bi riS oT roT roS iD
9 9 7 1 21 18 2 --
  +----- Hyriidae **   
--+                 
  |  +-- Etherioidea
  +--+              
     +-- Unionoidea 

Bolotov et al., 2016 [Publ 3743], fig. 2

iT tT bi riS oT roT roS iD
9 9 7 1 21 18 2 --
  +----- Hyriidae **   
--+                 
  |  +-- Etherioidea
  +--+              
     +-- Unionoidea 

Bolotov et al., 2017 [Publ 3761], fig. S2

iT tT bi riS oT roT roS iD
9 9 7 1 130 128 2 --
                       +-- Chamberlainia   
  +--------------------+                   
  |                    +-- Margaritiferidae
  |                                        
  |                    +-- Etherioidea     
--+--------------------+                   
  |                    +-- Hyriidae **        
  |                                        
  |  +-------------------- Parreysiinae    
  |  |                                     
  +--+  +----------------- Unioninae       
     |  |                                  
     +--+              +-- Contradentini   
        |  +-----------+                   
        |  |           +-- Rectidentini    
        +--+                               
           |  +----------- Ambleminae      
           +--+                            
              |  +-------- Pseudodontini   
              +--+                         
                 |  +----- Gonideini       
                 +--+                      
                    |  +-- Sinohyriopsis   
                    +--+                   
                       +-- Lamprotulini    

Hoeh et al., 2009 [Publ 2716], fig. 4

iT tT bi riS oT roT roS iD
9 9 7 1 37 36 2 --
  +-------------------------- Hyriidae **        
  |                                           
  |                       +-- Etheria         
  |                    +--+                   
--+                    |  +-- Mulleria        
  |  +-----------------+                      
  |  |                 |  +-- Mycetopodidae   
  |  |                 +--+                   
  +--+                    +-- Iridinidae      
     |                                        
     |  +-------------------- Parreysiinae    
     +--+                                     
        |  +----------------- Pseudomulleria  
        +--+                                  
           |  +-------------- Margaritiferidae
           +--+                               
              |  +----------- Pseudodontini   
              +--+                            
                 |  +-------- Contradentini   
                 +--+                         
                    |  +----- Unioninae       
                    +--+                      
                       |  +-- Gonideini       
                       +--+                   
                          +-- Ambleminae      

Baker et al., 2004 [Publ 2685], fig. 2-3

iT tT bi riS oT roT roS iD
8 8 9 1 1 1 1 --
  +-- Unionoidea
--+             
  +-- Hyriidae **  

Baker et al., 2003 [Publ 2690], fig. 5

iT tT bi riS oT roT roS iD
7 7 5 1 1 1 1 --
  +-- Unionoidea
--+             
  +-- Hyriidae **  

Santos-Neto et al., 2016 [Publ 3739], fig. 3_5

iT tT bi riS oT roT roS iD
7 7 6 1 0 0 0 --
-- (Hyriinae

Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2

iT tT bi riS oT roT roS iD
6 6 5 1 5 5 1 --
  +-- Margaritiferidae
  |                   
--+-- Unionidae       
  |                   
  +-- Hyriidae **        

Hoeh et al., 2002 [Publ 2653], fig. 2

iT tT bi riS oT roT roS iD
5 5 3 1 26 25 2 --
  +----- Hyriidae **   
--+                 
  |  +-- Etherioidea
  +--+              
     +-- Unionoidea 

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

iT tT bi riS oT roT roS iD
5 5 3 1 26 25 2 --
  +-------- Margaritiferidae
--+                         
  |  +----- Unionidae       
  +--+                      
     |  +-- Etherioidea     
     +--+                   
        +-- Hyriidae **        

Hoeh et al., 2001 [Publ 2652], fig. 14f2

iT tT bi riS oT roT roS iD
5 5 1 1 26 25 2 --
  +----------------- Margaritiferidae
  |                                  
  |  +-------------- Amblemini       
  |  |                               
  |  +-------------- Parreysiinae    
  |  |                               
  |  +-------------- Eurynia         
--+  |                               
  |  +-------------- Fusconaia       
  |  |                               
  |  +-------------- Gonideinae      
  |  |                               
  |  +-------------- Pleurobema      
  |  |                               
  +--+-------------- Quadrulini      
     |                               
     |  +----------- Ortmanniana     
     +--+                            
     |  |  +-------- Glebula         
     |  +--+                         
     |     |  +----- Potamilus       
     |     +--+                      
     |        |  +-- Ligumia         
     |        +--+                   
     |           +-- Toxolasma       
     |                               
     |  +----------- Cyrtonaias      
     +--+                            
        |  +-------- Unionini        
        +--+                         
           |  +----- Anodontini      
           +--+                      
              |  +-- Hyriidae **        
              +--+                   
                 +-- Etherioidea     

Guerra et al., 2019 [Publ 4168], fig. 1

iT tT bi riS oT roT roS iD
5 5 3 1 36 35 2 --
  +----- Etherioidea
--+                 
  |  +-- Hyriidae **   
  +--+              
     +-- Unionoidea 

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

iT tT bi riS oT roT roS iD
5 5 2 1 26 25 2 --
  +----- Unionoidea 
--+                 
  |  +-- Etherioidea
  +--+              
     +-- Hyriidae **   

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

iT tT bi riS oT roT roS iD
5 5 3 1 29 28 2 --
  +-------------------- Hyriidae **        
  |                                     
  |           +-------- Iridinidae      
--+  +--------+                         
  |  |        |  +----- Etheria         
  |  |        +--+                      
  |  |           |  +-- Mulleria        
  +--+           +--+                   
     |              +-- Mycetopodidae   
     |                                  
     |  +-------------- Parreysiinae    
     |  |                               
     +--+  +----------- Pseudomulleria  
        |  |                            
        +--+  +-------- Anodontini      
           |  |                         
           +--+  +----- Gonideinae      
              |  |                      
              +--+----- Unionini        
                 |                      
                 |  +-- Margaritiferidae
                 +--+                   
                    +-- Ambleminae      

Guerra et al., 2019 [Publ 4168], fig. 2

iT tT bi riS oT roT roS iD
5 5 3 1 36 35 2 --
  +----- Etherioidea
--+                 
  |  +-- Hyriidae **   
  +--+              
     +-- Unionoidea 

Santos-Neto et al., 2018 [Publ 3888], fig. 5

iT tT bi riS oT roT roS iD
5 5 3 1 0 0 0 --
-- (Hyriinae

Hoeh et al., 2001 [Publ 2652], fig. 14f4

iT tT bi riS oT roT roS iD
5 5 3 1 26 25 2 --
  +----- Hyriidae **   
--+                 
  |  +-- Etherioidea
  +--+              
     +-- Unionoidea 

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iT tT bi riS oT roT roS iD
4 4 2 1 36 35 2 --
  +----- Hyriidae **   
--+                 
  |  +-- Etherioidea
  +--+              
     +-- Unionoidea 

Klunzinger et al., 2022 [Publ 4259], fig. 1

iT tT bi riS oT roT roS iD
4 4 2 1 0 0 0 --
-- (Hyriidae

Graf, 2002 [Publ 2651], fig. 2

iT tT bi riS oT roT roS iD
4 4 2 1 14 13 1 --
  +-- Hyriidae **  
--+             
  +-- Unionoidea

Graf, 2000 [Publ 2647], fig. 1

iT tT bi riS oT roT roS iD
4 4 2 1 13 12 2 --
  +-------------- Margaritiferidae
  |                               
  |     +-------- Unionini        
--+  +--+                         
  |  |  |  +----- Pleurobemini    
  |  |  +--+                      
  |  |     |  +-- Anodontini      
  +--+     +--+                   
     |        +-- Lampsilini      
     |                            
     |  +-------- Parreysiini     
     +--+                         
        |  +----- Coelaturini     
        +--+                      
           |  +-- Hyriidae **        
           +--+                   
              +-- Etherioidea     

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iT tT bi riS oT roT roS iD
4 4 2 1 32 31 2 --
  +-------- Hyriidae **        
--+                         
  |  +----- Margaritiferidae
  +--+                      
     |  +-- Etherioidea     
     +--+                   
        +-- Unionidae       

Klunzinger et al., 2021 [Publ 4086], fig. 2

iT tT bi riS oT roT roS iD
4 4 2 1 2 2 1 --
  +-- Unionoidea
--+             
  +-- Hyriidae **  

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iT tT bi riS oT roT roS iD
4 4 2 1 32 31 2 --
  +-------- Etherioidea     
--+                         
  |  +----- Unionidae       
  +--+                      
     |  +-- Margaritiferidae
     +--+                   
        +-- Hyriidae **        

Combosch et al., 2017 [Publ 3806], fig. 1

iT tT bi riS oT roT roS iD
3 3 1 1 12 10 2 --
  +-------- Hyriidae **        
--+                         
  |  +----- Unionidae       
  +--+                      
     |  +-- Margaritiferidae
     +--+                   
        +-- Etherioidea     

Combosch et al., 2017 [Publ 3806], fig. 2

iT tT bi riS oT roT roS iD
3 3 1 1 12 10 2 --
  +----- Etherioidea
--+                 
  |  +-- Hyriidae **   
  +--+              
     +-- Unionoidea 

Wu et al., 2021 [Publ 4366], fig. 6

iT tT bi riS oT roT roS iD
3 3 1 1 88 86 2 --
  +----- Etherioidea
--+                 
  |  +-- Hyriidae **   
  +--+              
     +-- Unionoidea 

Olivera-Hyde et al., 2020 [Publ 4249], fig. 10

iT tT bi riS oT roT roS iD
3 3 1 1 0 0 0 --
-- (Castaliini

Sano et al., 2020 [Publ 4138], fig. 3

iT tT bi riS oT roT roS iD
3 3 1 1 66 64 2 --
  +----- Hyriidae **   
--+                 
  |  +-- Etherioidea
  +--+              
     +-- Unionoidea 

Wu et al., 2020 [Publ 4121], fig. 2

iT tT bi riS oT roT roS iD
3 3 1 1 31 29 2 --
  +----- Hyriidae **   
--+                 
  |  +-- Etherioidea
  +--+              
     +-- Unionoidea 

Sonowal et al., 2021 [Publ 4221], fig. 3b

iT tT bi riS oT roT roS iD
3 3 0 1 12 9 1 --
  +-- Hyriidae **  
--+             
  +-- Unionoidea

Konopleva et al., 2017 [Publ 3807], fig. 6

iT tT bi riS oT roT roS iD
3 3 1 1 28 26 2 --
  +----- Hyriidae **   
--+                 
  |  +-- Etherioidea
  +--+              
     +-- Unionoidea 

Sonowal et al., 2021 [Publ 4221], fig. 3a

iT tT bi riS oT roT roS iD
3 3 0 1 12 9 1 --
-- (Hyriidae

Sano et al., 2020 [Publ 4138], fig. 4

iT tT bi riS oT roT roS iD
3 3 1 1 66 64 2 --
  +----- Hyriidae **   
--+                 
  |  +-- Etherioidea
  +--+              
     +-- Unionoidea 

Sano et al., 2020 [Publ 4138], fig. 2

iT tT bi riS oT roT roS iD
3 3 1 1 66 64 2 --
  +----- Hyriidae **   
--+                 
  |  +-- Etherioidea
  +--+              
     +-- Unionoidea 

Sonowal et al., 2021 [Publ 4221], fig. 4

iT tT bi riS oT roT roS iD
3 3 1 1 13 10 1 --
  +-- Hyriidae **  
--+             
  +-- Unionoidea

Wu et al., 2021 [Publ 4366], fig. 2_I

iT tT bi riS oT roT roS iD
3 3 1 1 88 86 2 --
  +----- Hyriidae **   
--+                 
  |  +-- Etherioidea
  +--+              
     +-- Unionoidea 

Huang et al., 2018 [Publ 3828], fig. 3_5

iT tT bi riS oT roT roS iD
3 3 1 1 26 24 2 --
  +----- Hyriidae **   
--+                 
  |  +-- Etherioidea
  +--+              
     +-- Unionoidea 

Wu et al., 2020 [Publ 4121], fig. S1

iT tT bi riS oT roT roS iD
3 3 1 1 31 29 2 --
  +----- Hyriidae **   
--+                 
  |  +-- Etherioidea
  +--+              
     +-- Unionoidea 

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

iT tT bi riS oT roT roS iD
3 3 1 1 25 23 2 --
  +----- Hyriidae **   
--+                 
  |  +-- Etherioidea
  +--+              
     +-- Unionoidea 

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

iT tT bi riS oT roT roS iD
3 3 1 1 25 23 2 --
  +----- Hyriidae **   
--+                 
  |  +-- Etherioidea
  +--+              
     +-- Unionoidea 

Guerra et al., 2019 [Publ 4168], fig. 1-M

iT tT bi riS oT roT roS iD
2 2 0 1 20 20 1 --
  +-- Hyriidae **  
--+             
  +-- Unionoidea

Guerra et al., 2019 [Publ 4168], fig. 2-M

iT tT bi riS oT roT roS iD
2 2 0 1 20 20 1 --
  +-- Hyriidae **  
--+             
  +-- Unionoidea

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iT tT bi riS oT roT roS iD
2 2 0 1 118 116 2 --
  +-------- Hyriidae **        
--+                         
  |  +----- Margaritiferidae
  +--+                      
     |  +-- Etherioidea     
     +--+                   
        +-- Unionidae       

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iT tT bi riS oT roT roS iD
1 1 0 1 91 90 2 --
  +----- Hyriidae **   
--+                 
  |  +-- Etherioidea
  +--+              
     +-- Unionoidea 

Elderkin et al., 2016 [Publ 3693], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 2 2 1 --
  +-- Etheriidae
  |             
--+-- Iridinidae
  |             
  +-- Hyriidae **  

Bieler et al., 2014 [Publ 3622], fig. 26_27_29_30

iT tT bi riS oT roT roS iD
1 1 0 1 4 3 2 --
  +-------- Hyriidae **        
--+                         
  |  +----- Unionidae       
  +--+                      
     |  +-- Etherioidea     
     +--+                   
        +-- Margaritiferidae

Chase et al., 2018 [Publ 3861], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 10 10 1 --
  +-- Hyriidae **  
--+             
  +-- Unionoidea

Breton et al., 2011 [Publ 3376], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 36 36 1 --
  +-- Hyriidae **  
--+             
  +-- Unionoidea

Huang et al., 2018 [Publ 3828], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 42 41 2 --
  +----- Etherioidea
--+                 
  |  +-- Hyriidae **   
  +--+              
     +-- Unionoidea 

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iT tT bi riS oT roT roS iD
1 1 0 1 56 55 2 --
  +----- Etherioidea
--+                 
  |  +-- Hyriidae **   
  +--+              
     +-- Unionoidea 

González & Giribet, 2015 [Publ 3632], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 5 3 2 --
  +-------- Hyriidae **        
--+                         
  |  +----- Unionidae       
  +--+                      
     |  +-- Etherioidea     
     +--+                   
        +-- Margaritiferidae

Wu et al., 2021 [Publ 4366], fig. 3-BI

iT tT bi riS oT roT roS iD
1 1 0 1 73 72 2 --
  +----- Hyriidae **   
--+                 
  |  +-- Etherioidea
  +--+              
     +-- Unionoidea 

Chase et al., 2018 [Publ 3847], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 9 9 1 --
  +----- Unionidae       
--+                      
  |  +-- Margaritiferidae
  +--+                   
     +-- Hyriidae **        

Guerra et al., 2017 [Publ 3772], fig. 2b-M

iT tT bi riS oT roT roS iD
1 1 0 1 7 7 1 --
  +-- Hyriidae **  
--+             
  +-- Unionoidea

Walker et al., 2006 [Publ 2666], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 14 13 1 --
  +-- Hyriidae **  
--+             
  +-- Unionoidea

Walker et al., 2006 [Publ 2666], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 14 13 1 --
  +-- Hyriidae **  
--+             
  +-- Unionoidea

Wu et al., 2020 [Publ 4121], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 49 48 2 --
  +----- Hyriidae **   
--+                 
  |  +-- Etherioidea
  +--+              
     +-- Unionoidea 

Chase et al., 2018 [Publ 3861], fig. 1-M

iT tT bi riS oT roT roS iD
1 1 0 1 10 10 1 --
  +----- Unionidae       
--+                      
  |  +-- Margaritiferidae
  +--+                   
     +-- Hyriidae **        

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

iT tT bi riS oT roT roS iD
1 1 0 1 29 28 2 --
  +----- Etherioidea
--+                 
  |  +-- Hyriidae **   
  +--+              
     +-- Unionoidea 

Guerra et al., 2017 [Publ 3772], fig. 2b

iT tT bi riS oT roT roS iD
1 1 0 1 24 23 2 --
  +----- Etherioidea
--+                 
  |  +-- Hyriidae **   
  +--+              
     +-- Unionoidea 

Wu et al., 2019 [Publ 3864], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 46 46 2 --
  +-------- Unionidae       
--+                         
  |  +----- Margaritiferidae
  +--+                      
     |  +-- Hyriidae **        
     +--+                   
        +-- Etherioidea     

Wu et al., 2022 [Publ 4238], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 73 73 2 --
  +----- Etherioidea
--+                 
  |  +-- Hyriidae **   
  +--+              
     +-- Unionoidea 

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iT tT bi riS oT roT roS iD
1 1 0 1 95 94 2 --
  +----- Hyriidae **   
--+                 
  |  +-- Etherioidea
  +--+              
     +-- Unionoidea 

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

iT tT bi riS oT roT roS iD
1 1 0 1 29 28 2 --
  +----- Etherioidea
--+                 
  |  +-- Hyriidae **   
  +--+              
     +-- Unionoidea 

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 56 55 2 --
  +-------- Etherioidea     
--+                         
  |  +----- Unionidae       
  +--+                      
     |  +-- Hyriidae **        
     +--+                   
        +-- Margaritiferidae

González & Giribet, 2015 [Publ 3632], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 5 3 2 --
  +----- Unionidae       
  |                      
--+----- Hyriidae **        
  |                      
  |  +-- Etherioidea     
  +--+                   
     +-- Margaritiferidae

Guerra et al., 2017 [Publ 3772], fig. 2a-M

iT tT bi riS oT roT roS iD
1 1 0 1 7 7 1 --
  +-- Hyriidae **  
--+             
  +-- Unionoidea

Huang et al., 2019 [Publ 3843], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 56 55 2 --
  +----- Etherioidea
--+                 
  |  +-- Hyriidae **   
  +--+              
     +-- Unionoidea 

Plazzi et al., 2011 [Publ 3676], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 17 17 1 --
  +----------- Cristariina    
  |                           
  |  +-------- Lanceolariini  
  |  |                        
--+  +-------- Anodonta       
  |  |                        
  |  +-------- Unionini       
  |  |                        
  +--+     +-- Alasmidontina  
     +-----+                  
     |     +-- Pseudanodonta  
     |                        
     |  +----- Ambleminae     
     |  |                     
     +--+  +-- Hyriidae **       
        |  |                  
        +--+-- Lamprotulini   
           |                  
           +-- Chamberlainiini

Chase et al., 2018 [Publ 3847], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 37 37 1 --
  +-------------- Margaritiferidae
  |                               
--+  +----------- Unioninae       
  |  |                            
  +--+  +-------- Lampsilini      
     |  |                         
     +--+     +-- Hyriidae **        
        |  +--+                   
        |  |  +-- Amblemini       
        +--+                      
           |  +-- Pleurobemini    
           +--+                   
              +-- Quadrulini      

Wu et al., 2021 [Publ 4366], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 68 67 2 --
  +----- Hyriidae **   
--+                 
  |  +-- Etherioidea
  +--+              
     +-- Unionoidea 

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iT tT bi riS oT roT roS iD
1 1 0 1 95 94 2 --
  +----- Hyriidae **   
--+                 
  |  +-- Etherioidea
  +--+              
     +-- Unionoidea 

Walker et al., 2006 [Publ 2666], fig. 6-M

iT tT bi riS oT roT roS iD
1 1 0 1 13 13 1 --
  +-- Hyriidae **  
--+             
  +-- Unionoidea

González & Giribet, 2015 [Publ 3632], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 5 3 2 --
  +----- Etherioidea
--+                 
  |  +-- Hyriidae **   
  +--+              
     +-- Unionoidea 

González & Giribet, 2015 [Publ 3632], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 5 3 2 --
     +-- Etherioidea     
  +--+                   
  |  +-- Margaritiferidae
--+                      
  |  +-- Unionidae       
  +--+                   
     +-- Hyriidae **        

Hoeh et al., 2002 [Publ 2673], fig. 1-M

iT tT bi riS oT roT roS iD
1 1 0 1 8 8 1 --
  +-- Hyriidae **  
--+             
  +-- Unionoidea

Wu et al., 2021 [Publ 4366], fig. 3-ML

iT tT bi riS oT roT roS iD
1 1 0 1 73 72 2 --
  +----- Hyriidae **   
--+                 
  |  +-- Etherioidea
  +--+              
     +-- Unionoidea 

Hoeh et al., 2002 [Publ 2673], fig. 1-F

iT tT bi riS oT roT roS iD
1 1 0 1 8 8 1 --
  +-- Hyriidae **  
--+             
  +-- Unionoidea

Bieler et al., 2014 [Publ 3622], fig. 28

iT tT bi riS oT roT roS iD
1 1 0 1 4 3 2 --
     +-- Etherioidea     
  +--+                   
  |  +-- Margaritiferidae
--+                      
  |  +-- Unionidae       
  +--+                   
     +-- Hyriidae **        

Guerra et al., 2017 [Publ 3772], fig. 2a

iT tT bi riS oT roT roS iD
1 1 0 1 24 23 2 --
  +----- Etherioidea
--+                 
  |  +-- Hyriidae **   
  +--+              
     +-- Unionoidea 

Huang et al., 2019 [Publ 3843], fig. 3-M

iT tT bi riS oT roT roS iD
1 1 0 1 22 22 1 --
  +-- Hyriidae **  
--+             
  +-- Unionoidea

Huang et al., 2018 [Publ 3828], fig. 6-M

iT tT bi riS oT roT roS iD
1 1 0 1 16 16 1 --
  +-- Hyriidae **  
--+             
  +-- Unionoidea

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iT tT bi riS oT roT roS iD
1 1 0 1 91 90 2 --
  +----- Hyriidae **   
--+                 
  |  +-- Etherioidea
  +--+              
     +-- Unionoidea 

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 33 33 1 --
  +----------------------------------- Margaritiferinae       
  |                                                           
  +----------------------------------- Hyriidae **               
--+                                                           
  |  +-------------------------------- Parreysiinae           
  |  |                                                        
  +--+  +----------------------------- Unioninae              
     |  |                                                     
     +--+  +-------------------------- Ambleminae             
        |  |                                                  
        +--+  +----------------------- Chamberlainiini        
           |  |                                               
           +--+                    +-- Gonideini              
              |  +-----------------+                          
              |  |                 +-- Pseudodontini          
              +--+                                            
                 |  +----------------- Contradentini          
                 |  |                                         
                 +--+  +-------------- Pronodularia           
                    |  |                                      
                    +--+  +----------- Potomida               
                       |  |                                   
                       +--+  +-------- Lamprotula_leaii       
                          |  |                                
                          +--+  +----- Lamprotula_caveata__2  
                             |  |                             
                             +--+  +-- Lamprotula_cornuumlunae
                                |  |                          
                                +--+-- Gibbosulinae           
                                   |                          
                                   +-- Lamprotula_caveata__1  

Walker et al., 2006 [Publ 2666], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 14 13 1 --
  +-- Hyriidae **  
--+             
  +-- Unionoidea

Page updated: 14:53:32 Wed 04 Dec 2024

 
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