|
Bivalvia | Unionoida superfamily HYRIOIDEA Swainson, 1840: cladomics
1 family
Cladomics Summary
number of species = 129
number of subtaxa (rank: family) = 1
analyzed in 99 cladograms in 51 publications
maximum number of terminals in an analysis = 19, mean = 3.92
maximum number of subtaxa (family) in an analysis = 1, mean = 1.0
Basic Monophyly
number of cladograms that test basic monophyly (i.e., more than 1 included species) = 57
maximum number of included terminals = 19, mean = 6.07
maximum number of subtaxa (family) = 1, mean = 1.0
proportion of cladograms that support basic monophyly = 1.0
Pairwise Cladogram Clustering
The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
78 |
77 |
3 |
-- |
*** Top Monophyly Cladogram *** *** Top Sister-Group Cladogram ***
+-- Hyridella_depressa **
+-----------------------------------+
| +-- Velesunio_ambiguus **
|
| +-- Cumberlandia_monodonta
| +--------------------------------+
| | +-- Margaritifera_margaritifera
| |
| | +----------- Mutela_rostrata
--+ | |
| | +--------------------+ +-- Aspatharia_pfeifferiana
| | | | +-----+
| | | | | +-- Chambardia_wahlbergi
| | | +--+
| | | | +----- Etheria_elliptica
| | | +--+
| | | | +-- Tamsiella_schroeteriana
+--+ | +--+
| | +-- Mycetopoda_siliquosa
| |
| | +-------- Trapezidens_dolichorhynchus
| | +--------------------+
| | | | +----- Leoparreysia_tavoyensis
| | | +--+
| | | | +-- Coelatura_aegyptiaca
| | | +--+
| | | +-- Indochinella_pugio
| | |
| | | +----- Unio_gibbus
+--+ | |
| | +-----------+----- Unio_tumidus
| | | |
| | | | +-- Unio_crassus
| | | +--+
| | +--+ +-- Unio_pictorum
| | | |
| | | | +-------------- Aculamprotula_tortuosa
| | | | |
| | | +--+ +-- Nodularia_nuxpersicae
| | | | +--------+
| | | | | +-- Nodularia_douglasiae
| | | +--+
| | | | +-------- Schistodesmus_lampreyanus
| | | +--+
| | | | +----- Tchangsinaia_piscicula
| | | +--+
| | +-----+ | +-- Cuneopsis_heudei
| | | | +--+
| | | | +-- Cuneopsis_rufescens
+--+ | |
| | | +-------- Lanceolaria_gladiola
| | | +--------+
| | | | | +----- Lanceolaria_grayii__2
| | | | +--+
| | | | | +-- Lanceolaria_lanceolata
| | | | +--+
| | | | +-- Lanceolaria_grayii__1
| | | |
| | +--+ +----------- Cristaria_plicata
| | | +--+
| | | | | +-------- Anemina_arcaeformis
| | | | +--+
| | | | | +----- Pletholophus_tenuis
| | | | +--+
| | | | | +-- Sinanodonta_jourdyi
| | | | +--+
| | +--+ +-- Sinanodonta_lucida
| | |
| | | +----- Pseudanodonta_complanata
| | | |
| | | +-----+ +-- Anodonta_anatina
| | | | | |
| | | | +--+-- Anodonta_cygnea
| | | | |
| | +--+ +-- Anodonta_nuttalliana
| | |
| | | +-- Pyganodon_grandis
+--+ | +-----+
| | | +-- Simpsonaias_ambigua
| +--+
| | +----- Strophitus_undulatus
| +--+
| | +-- Alasmidonta_marginata
| +--+
| +-- Lasmigona_costata
|
| +-- Lens_contradens
| +--+
| | +-- Lens_comptus
| +--+
| | | +-- Physunio_modelli
| | +--+
| | +-- unionoid_species
| |
| +--------------+ +----- Hyriopsis_desowitzi
| | | |
| | | +----- Rectidens_sumatrensis
| | | |
| | | | +-- Hyriopsis_khoratensis
| | +--+--+
| | | +-- Hyriopsis_myersiana
| | |
| | | +-- Ensidens_ingallsianus
| | | |
| | +--+-- Ensidens_telus
| | |
| | +-- Ensidens_jaculus
| |
| | +-------- Amblema_plicata
+--+ +--+
| | | +----- Cambarunio_iris
| | +--+
| | | +-- Lampsilis_cardium
| | +--+
| +--------+ +-- Ortmanniana_ligamentina
| | |
| | | +-- Quadrula_quadrula
| | | +-----+
| | | | +-- Quadrula_verrucosa
| | +--+
| | | +----- Elliptio_complanata
| | +--+
| | | +-- Pleurobema_sintoxia
| | +--+
| | +-- Eurynia_dilatata
+--+
| +-- Chamberlainia_somsakpanhai
| +-----+
| | +-- Sinohyriopsis_cumingii
| |
| +--------+ +-- Lamprotula_caveata
| | | +--+
| | | | +-- Lamprotula_leaii
| | +--+
| | | +-- Potomida_littoralis
| | +--+
| | +-- Pronodularia_japanensis
+--+
| +-- Sinosolenaia_carinata
| +-----+
| | +-- Sinosolenaia_oleivora
| +-----+
| | | +----- Gonidea_angulata
| | +--+
| | | +-- Microcondylaea_bonellii
+--+ +--+
| +-- Leguminaia_wheatleyi
|
| +----------- Parvasolenaia_rivularis
| |
+--+ +-- Pilsbryoconcha_exilis
| +-----+
| | +-- Pilsbryoconcha_acuta
+--+
| +----- Pseudodon_vagulus
+--+
| +-- Bineurus_exilis
+--+
+-- Nyeinchanconcha_nyeinchani
Santos-Neto et al., 2016 [Publ 3739], fig. 2a,b
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 19 |
19 |
20 |
1 |
1 |
0 |
0 |
-- |
+-------- Westralunio_carteri **
|
+--------------+ +-- Hyridella_drapeta **
| | +--+
| | | +-- Hyridella_depressa **
| +--+
| | +-- Diplodon_gratus **
| +--+
| +-- Diplodon_suavidicus__2 **
|
| +-- Hyridella_australis **
| +--------------+
| | +-- Cucumerunio_novaehollandiae **
+-----+
| | +-------------- Echyridella_menziesii **
--+ +--+
| | +----------- Velesunio_ambiguus **
| +--+
| | +-------- Microdontia_anodontaeformis **
| +--+
| | +----- Lortiella **
| +--+
| | +-- Alathyria_jacksoni **
| +--+
| +-- Velesunio_angasi **
|
| +-------------------- Diplodon_suavidicus__1 **
| |
+--+ +-- Triplodon_corrugatus **
| +--------------+
| | +-- Prisodon_syrmatophorus **
+--+
| +-------------- Castalia_stevensi **
| |
+--+ +-- Callonaia_duprei **
| +--------+
| | +-- Castalia_ambigua__1 **
+--+
| +-------- Castalia_ambigua__4 **
+--+
| +----- Castalia_ambigua__2 **
+--+
| +-- Castaliella_quadrata **
+--+
+-- Castalia_ambigua__3 **
Santos-Neto et al., 2016 [Publ 3739], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 19 |
19 |
21 |
1 |
1 |
0 |
0 |
-- |
+----------------------- Diplodon_suavidicus__2 **
|
+--+ +-- Triplodon_corrugatus **
| | +-----------------+
| | | +-- Prisodon_syrmatophorus **
| +--+
| | +----------------- Castalia_ambigua__4 **
| +--+
| | +-------------- Callonaia_duprei **
| +--+
| | +----------- Castalia_ambigua__3 **
| +--+
| | +-------- Castalia_stevensi **
| +--+
--+ | +----- Castalia_ambigua__2 **
| +--+
| | +-- Castaliella_quadrata **
| +--+
| +-- Castalia_ambigua__1 **
|
| +-------- Westralunio_carteri **
| |
| +-----------+ +-- Hyridella_drapeta **
| | | +--+
| | | | +-- Hyridella_depressa **
| | +--+
| | | +-- Diplodon_suavidicus__1 **
+-----+ +--+
| +-- Diplodon_gratus **
|
| +----------------- Echyridella_menziesii **
| |
+--+ +-- Cucumerunio_novaehollandiae **
| +-----------+
| | +-- Hyridella_australis **
+--+
| +----------- Velesunio_ambiguus **
+--+
| +-------- Alathyria_jacksoni **
+--+
| +----- Velesunio_angasi **
+--+
| +-- Lortiella **
+--+
+-- Microdontia_anodontaeformis **
Huang et al., 2019 [Publ 3843], fig. S3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
14 |
12 |
1 |
182 |
180 |
2 |
-- |
+----- Velesunio_ambiguus **
+--+
| | +-- Alathyria_jacksoni **
| +--+
+--------+ +-- Alathyria_profuga **
| |
| | +----- Alathyria_pertexta **
| +--+
| | +-- Lortiella_froggatti **
+-----------------------------------------------+ +--+
| | +-- Microdontia_anodontaeformis **
| |
| | +-------------- Westralunio_carteri **
| | |
| +--+ +----------- Echyridella_menziesii **
| | |
| | | +----- Diplodon_suavidicus **
| +--+ +--+
| | | | +-- Castalia_ambigua **
| | | +--+
| +--+ +-- Triplodon_corrugatus **
| |
| | +----- Hyridella_australis **
| +--+
| | +-- Hyridella_depressa **
| +--+
| +-- Hyridella_drapeta **
|
| +-------- Anodontites_trapesialis
| +--+
| | | +----- Mutela_rostrata
| | +--+
| | | +-- Mutela_dubia
| | +--+
| +--------------------------------------------------+ +-- Mutela_hargeri
| | |
| | | +----- Mycetopoda_siliquosa
--+ | | +--+
| | | | | +-- Tamsiella_schroeteriana
| | | | +--+
| | +--+ +-- Lamproscapha_ensiformis
| | |
| | | +----- Etheria_elliptica
| | +--+
| | | +-- Aspatharia_pfeifferiana
| | +--+
| | +-- Chambardia_wahlbergi
| |
| | +-------------------- Gibbosula_rochechouartii
| | |
| | | +----------------- Gibbosula_laosensis
| | +--------------------------------------+ |
| | | | | +-------- Cumberlandia_monodonta
| | | | | +-----+
| | | +--+ | | +----- Pseudunio_marocanus
| | | | | +--+
| | | | | | +-- Pseudunio_auricularius
| | | | | +--+
| | | +--+ +-- Pseudunio_homsensis
| | | |
| | | | +-- Margaritifera_dahurica
| | | | +--------+
| | | | | +-- Margaritifera_margaritifera
| | | +--+
| | | | +-------- Margaritifera_falcata
+--+ | | |
| | +--+ +-- Margaritifera_hembeli
| | | +--+
| | | | +-- Margaritifera_marrianae
| | +--+
| | | +-- Margaritifera_laevis
| | +--+
| | +-- Margaritifera_middendorffi
| |
| | +----- Trapezidens_exolescens
| | +-----+
| | | | +-- Trapezidens_angustior
| | | +--+
| | | +-- Trapezidens_dolichorhynchus
| | |
| | +--------------+ +-- Lamellidens_corrianus
| | | | +-----+
| | | | | +-- Lamellidens_savadiensis
| | | | |
| | | | | +----- Lamellidens_lamellatus
| | | +--+ |
| | | | +----- Lamellidens_jenkinsianus
| | | | |
| | | | | +-- Lamellidens_marginalis
| | | +--+--+
| | | | +-- Lamellidens_ferrugineus
| | | |
| | +-----------------------------+ | +-- Lamellidens_mainwaringi__1
| | | | | |
| | | | +--+-- Lamellidens_mainwaringi__2
| | | | |
| | | | +-- Lamellidens_generosus
+--+ | |
| | | +----- Parreysia_keralaensis
| | | +-----------------+
| | | | | +-- Parreysia_nagpoorensis
| | | | +--+
| | | | +-- Parreysia_plagiosoma
| | | |
| | | | +----- Leoparreysia_canefrii
| | +--+ +--------------+
| | | | | +-- Leoparreysia_tavoyensis
| | | | +--+
| | | | +-- Leoparreysia_olivacea
| | | |
| | | | +----------- Coelatura_aegyptiaca__2
| | | | |
| | +--+ +-----+ +-- Coelatura_aegyptiaca__1
| | | | | +-----+
| | | | | | +-- Nitia_teretiuscula
| | | | +--+
| | | | | +----- Coelatura_kunenensis
| | | | +--+
| | | | | +-- Coelatura_gabonensis
| | | | +--+
| | +--+ +-- Prisodontopsis_mweruensis
| | |
| | | +-------- Indonaia_shurtleffiana
| | | |
| | | +-----+ +----- Indonaia_bonneaudii
| | | | | |
| | | | +--+----- Indonaia_rugosa
| | | | |
| | | | | +-- Indonaia_andersoniana
| | +--+ +--+
| | | +-- Indonaia_caerulea
| | |
| | | +----------- Indochinella_pugio
| | | |
| | | | +----- Scabies_phaselus
| | +--+ +--+
| | | | | +-- Scabies_humilis
| | | | +--+
| | +--+ +-- Scabies_mandarinus
| | |
| | | +----- Radiatula_mouhoti
| | +--+
| | | +-- Radiatula_myitkyinae
| | +--+
+--+ +-- Radiatula_chaudhurii
|
| +----- Indopseudodon_indawgyiensis
| +--------------+
| | | +-- Indopseudodon_bogani
| | +--+
| | +-- Indopseudodon_salwenianus
| |
| | +-------- Pilsbryoconcha_exilis__3
| +--------------------------------+ +--------+
| | | | | +----- Pilsbryoconcha_exilis__2
| | | | +--+
| | | | | +-- Pilsbryoconcha_carinifera
| | | | +--+
| | | | +-- Pilsbryoconcha_exilis__1
| | | |
| | +--+ +----------- Sundadontina_tumida
| | | +--+
| | | | | +-------- Pseudodon_vagulus
| | | | +--+
| | | | | +----- Pseudodon_vondembuschianus__1
| | | | +--+
| | | | | +-- Pseudodon_mekongi
| | +--+ +--+
| | | +-- Pseudodon_vondembuschianus__2
| | |
| | | +-- Nyeinchanconcha_nyeinchani
| | | +--------+
| | | | +-- Sundadontina_cumingii
| | +--+
| | | +-------- Thaiconcha_callifera
| | +--+
| | | +----- Bineurus_loeiensis
| | +--+
| | | +-- Bineurus_mouhotii
| | +--+
| | +-- Bineurus_exilis
| |
| | +-- Ensidens_telus
| | +-----+
| | | +-- Ensidens_ingallsianus
| | +-----+
| | | | +----- Ensidens_sagittarius
| | | +--+
| | | | +-- Ensidens_spiculus
| | +--+ +--+
| | | | +-- Ensidens_jaculus
| | | |
| | | | +----------- Hyriopsis_desowitzi
| | | | |
| | | +--+ +-------- Rectidens_sumatrensis
| | | | |
| | | +--+ +-- Hyriopsis_panhai
+--+ | | +--+
| | | | +-- Hyriopsis_khoratensis
| +--------------------------------+ +--+
| | | | +-- Hyriopsis_bialata
| | | +--+
| | | +-- Hyriopsis_myersiana
| | |
| | | +----- Physunio
| | | +--------+
| | | | | +-- Physunio_superbus
| | | | +--+
| | | | +-- Physunio_modelli
| | | |
| | +--+ +-------- Yaukthwa_inlenensis
| | | +--+
| | | | | +----- Yaukthwa_panhai
| | | | +--+
| | | | | +-- Yaukthwa_nesemanni
| | | | +--+
| | +--+ +-- Yaukthwa_rectangularis
| | |
| | | +-- Lens_contradens
| | | +-----+
| | | | +-- Lens_micropterus
| | +--+
| | | +----- Lens_eximius
| | +--+
| | | +-- Lens_novoselovi
| | +--+
| | +-- Lens_comptus
| |
| | +----- Chamberlainia_somsakpanhai
| | +--------------------------------------+
| | | | +-- Sinohyriopsis_cumingii
| | | +--+
| | | +-- Sinohyriopsis_schlegelii
| | |
| | | +-- Potomida_littoralis
| | | +-----+
| | | | +-- Pronodularia_japanensis
| | | +-----+
| | | | | +----- Lamprotula_leaii
| | | | +--+
| | | | | +-- Lamprotula_caveata
+--+ | | +--+
| | | +-- Lamprotula_cornuumlunae
| +--+ +--------------------------+
| | | | | +----- Sinosolenaia_recognita
| | | | | +-----+
| | | | | | | +-- Sinosolenaia_oleivora
| | | | | | +--+
| | | | | | +-- Sinosolenaia_iridinea
| | | | +--+
| | | | | +----- Gonidea_angulata
| | | | | +--+
| | | | | | | +-- Leguminaia_wheatleyi
| | | | | | +--+
| | | | +--+ +-- Microcondylaea_bonellii
| | | | |
| | | | | +----- Ptychorhynchus_pfisteri
| | +--+ +--+
| | | | +-- Parvasolenaia_rivularis
| | | +--+
| | | +-- Koreosolenaia_sitgyensis
| | |
| | | +-- Megalonaias_nervosa
| | | +--+
| | | | +-- Uniomerus_carolinianus
| | | +--------------------------------+
| | | | | +-- Quadrula_quadrula
| | | | +--+
| | | | +-- Quadrula_verrucosa
| | | |
| | | | +-------- Elliptoideus_sloatianus
| | +--+ |
| | | +--------------------------+ +-- Elliptio_complanata
| | | | | +--+
| | | | | | +-- Eurynia_dilatata
| | | | +--+
| | | | | +-- Fusconaia_flava
| | +--+ +--+
| | | +-- Pleurobema_sintoxia
| | |
| | | +-------------------------------- Amblema_plicata
| | | |
| | +--+ +----------------------------- Toxolasma_parvum
| | | |
+--+ +--+ +-- Obliquaria_reflexa
| | +-----------------------+
| | | +-- Plectomerus_dombeyanus
| +--+
| | +----------------------- Potamilus_leptodon
| | |
| +--+ +-------------------- Potamilus_alatus
| | |
| | | +-- Medionidus_conradicus
| +--+ +--------------+
| | | +-- Ptychobranchus_subtentus
| | |
| +--+ +-- Ellipsaria_lineolata
| | +-----------+
| | | +-- Truncilla_truncata
| +--+
| | +----------- Leaunio_lienosus
| | |
| +--+ +-- Venustaconcha_ellipsiformis
| | +-----+
| | | +-- Cambarunio_iris
| +--+
| | +----- Ortmanniana_ligamentina
| +--+
| | +-- Lampsilis_cardium
| +--+
| +-- Lampsilis_ornata
|
| +----------------------------------- Acuticosta_chinensis
| |
| | +-------- Aculamprotula_tortuosa
| | |
| | +-----------+ +-- Aculamprotula_coreana
| | | | +--+
| | | | | +-- Aculamprotula_scripta
| | | +--+
| | | | +-- Aculamprotula_fibrosa
| | | +--+
| | | +-- Aculamprotula_tientsinensis
| | +-----------+
| | | | +-------- Unio_tumidus
+-----------+ | | +--------+
| | | | | +----- Unio_crassus
| | | | +--+
| | | | | +-- Unio_delphinus
| | | | +--+
| | +--+ +-- Unio_pictorum
| | |
| | | +----- Nodularia_douglasiae
| | | +--------+
| | | | | +-- Nodularia_huana
| | | | +--+
| | +--+ +-- Nodularia_nuxpersicae
| | |
+--+ | +----------- Schistodesmus_lampreyanus
| +--+
| | +-------- Tchangsinaia_piscicula
| +--+
| | +----- Cuneopsis_rufescens
| +--+
| | +-- Cuneopsis_heudei
| +--+
| +-- Cuneopsis_celtiformis
|
| +----------------------------- Lepidodesma_languilati
| |
| | +-------- Lanceolaria_gladiola
| | +-----------------+
| | | | +----- Lanceolaria_grayii
| | | +--+
+--+ | | +-- Lanceolaria_lanceolata
| | +--+
| | +-- Lanceolaria_triformis
| |
| | +-------------------- Anemina_arcaeformis
| | |
| | +--+ +----------------- Cristaria_plicata
+--+ | | |
| | +--+ +-------------- Pletholophus_tenuis
| | | |
| | +--+ +-- Sinanodonta_angula
| | | +--------+
| | | | +-- Sinanodonta_lucida
| | +--+
| | | +-------- Sinanodonta_woodiana
| | +--+
| | | +----- Sinanodonta_jourdyi
+--+ +--+
| | +-- Sinanodonta_elliptica
| +--+
| +-- Sinanodonta_pacifica
|
| +-- Alasmidonta_marginata
| +--------+
| | +-- Lasmigona_costata
| |
| | +----- Anodonta_anatina
| +-----+
| | | +-- Anodonta_cygnea
+-----------+ +--+
| +-- Anodonta_nuttalliana
|
| +----- Pseudanodonta_complanata
| +--+
| | | +-- Simpsonaias_ambigua
| | +--+
+--+ +-- Strophitus_undulatus
|
| +----- Pyganodon_grandis
+--+
| +-- Utterbackia_imbecillis
+--+
+-- Utterbackia_peninsularis
Huang et al., 2019 [Publ 3843], fig. S4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
14 |
9 |
1 |
184 |
182 |
2 |
-- |
+----------- Etheria_elliptica
|
| +-- Aspatharia_pfeifferiana
+--------+
| +-- Chambardia_wahlbergi
|
+-----------------------+ +----- Mycetopoda_siliquosa
| +-----+
| | | +-- Tamsiella_schroeteriana
| | +--+
| | +-- Lamproscapha_ensiformis
| |
| | +-------- Anodontites_trapesialis
| +--+
| | +----- Mutela_hargeri
| +--+
| | +-- Mutela_dubia
| +--+
| +-- Mutela_rostrata
|
| +----------- Echyridella_menziesii **
| |
| +----------- Westralunio_carteri **
| |
| | +----- Diplodon_suavidicus **
| +-----+
| | | +-- Castalia_ambigua **
| | +--+
| | +-- Triplodon_corrugatus **
| |
| +--------------------+ +----- Hyridella_australis **
| | +-----+
| | | | +-- Hyridella_depressa **
| | | +--+
| | | +-- Hyridella_drapeta **
| | |
| | | +----- Velesunio_ambiguus **
| | | +--+
--+ | | | | +-- Alathyria_jacksoni **
| | | | +--+
| | +--+ +-- Alathyria_profuga **
| | |
| | | +----- Alathyria_pertexta **
| | +--+
| | | +-- Lortiella_froggatti **
| | +--+
| | +-- Microdontia_anodontaeformis **
| |
| | +-------------------- Gibbosula_rochechouartii
| | |
| | +--------+ +----------------- Gibbosula_laosensis
| | | | |
| | | +--+ +-------------- Cumberlandia_monodonta
| | | | |
| | | | | +----- Pseudunio_marocanus
| | | +--+ +-----+
| | | | | | +-- Pseudunio_auricularius
| | | | | +--+
| | | | | +-- Pseudunio_homsensis
| | | +--+
| | | | +-------- Margaritifera_falcata
| | | | |
| | | | | +-- Margaritifera_dahurica
| | | | +-----+
| | | +--+ +-- Margaritifera_margaritifera
| | | |
| | | | +----- Margaritifera_laevis
| | | | |
| | | +--+----- Margaritifera_middendorffi
| | | |
+--+ | | +-- Margaritifera_hembeli
| | +--+
| | +-- Margaritifera_marrianae
| |
| | +-- Parreysia_keralaensis
| | |
| | +--------------------+-- Parreysia_nagpoorensis
| | | |
| | | +-- Parreysia_plagiosoma
| | |
| | | +----- Trapezidens_exolescens
| | | +--------+
| | | | | +-- Trapezidens_angustior
| | | | +--+
| | +--------+ +-- Trapezidens_dolichorhynchus
| | | |
| | | | +----------- Lamellidens_corrianus
| | | | |
| | | +--+ +-------- Lamellidens_savadiensis
| | | | |
| | | | | +----- Lamellidens_marginalis
| | | +--+ |
| | | | +----- Lamellidens_ferrugineus
| | | | |
| | +--+ +--+ +-- Lamellidens_mainwaringi__1
| | | | | |
| | | | | +-- Lamellidens_mainwaringi__2
| | | | | |
| | | | +--+-- Lamellidens_lamellatus
| | | | |
| | | | +-- Lamellidens_generosus
| | | | |
| | | | +-- Lamellidens_jenkinsianus
| | | |
| | | | +----- Leoparreysia_tavoyensis
| | | | +--------------+
| | | | | | +-- Leoparreysia_canefrii
| | | | | +--+
| | | | | +-- Leoparreysia_olivacea
| | | | |
+--+ | | | +-------- Coelatura_aegyptiaca__2
| | | | |
| | | | +-------- Nitia_teretiuscula
| | +--+ +--------+
| | | | | +----- Coelatura_aegyptiaca__1
| | | | | |
| | | | +--+ +-- Coelatura_gabonensis
| | | | | |
| | | | +--+-- Coelatura_kunenensis
| | | | |
| | | | +-- Prisodontopsis_mweruensis
| | +--+
| | | +----- Indonaia_shurtleffiana
| | | |
| | | | +-- Indonaia_bonneaudii
| | | +--------+--+
| | | | | +-- Indonaia_rugosa
| | | | |
| | | | | +-- Indonaia_andersoniana
| | | | +--+
| | +--+ +-- Indonaia_caerulea
| | |
| | | +----------- Indochinella_pugio
| | | |
| | | | +----- Scabies_humilis
| | +--+ +--+
| | | | | +-- Scabies_phaselus
| | | | +--+
| | +--+ +-- Scabies_mandarinus
| | |
| | | +----- Radiatula_mouhoti
| | +--+
| | | +-- Radiatula_myitkyinae
| | +--+
| | +-- Radiatula_chaudhurii
| |
| | +-------------------- Chamberlainia_somsakpanhai
| | |
| | +-------------------- Megalonaias_nervosa
| | |
| | +-------------------- Uniomerus_carolinianus
| | |
| | | +-- Quadrula_quadrula
| | +-----------------+
| | | +-- Quadrula_verrucosa
| | |
| | | +-- Sinohyriopsis_cumingii
| | +-----------------+
| | | +-- Sinohyriopsis_schlegelii
| | |
| | | +-- Potomida_littoralis
| | | +-----+
| | | | +-- Pronodularia_japanensis
| | +-----------+
| | | | +----- Lamprotula_leaii
| | | +--+
| | | | +-- Lamprotula_caveata
| | | +--+
| | | +-- Lamprotula_cornuumlunae
| | |
| | | +-------- Sinosolenaia_carinata
+--+ | |
| | | +----- Gonidea_angulata
| | +--+
| | | | +-- Leguminaia_wheatleyi
| | | +--+
| | | +-- Microcondylaea_bonellii
| +-----------+
| | | +----- Ptychorhynchus_pfisteri
| | +--+
| | | | +-- Parvasolenaia_rivularis
| | | +--+
| | | +-- Koreosolenaia_sitgyensis
| | |
| | | +----- Sinosolenaia_recognita
| | +--+
| | | +-- Sinosolenaia_iridinea
| | +--+
| | +-- Sinosolenaia_oleivora
| |
| | +----- Indopseudodon_salwenianus
| | +-----+
| | | | +-- Indopseudodon_indawgyiensis
| | | +--+
| | | +-- Indopseudodon_bogani
| | |
| | | +-------- Nyeinchanconcha_nyeinchani
| | | |
| | | +-------- Thaiconcha_callifera
| +--------+ |
| | | +-------- Sundadontina_cumingii
| | | |
| | | +-------- Sundadontina_tumida__1
| | | |
| | | +-------- Sundadontina_tumida__2
| | | |
| | | | +----- Pilsbryoconcha_exilis__2
| | +--+--+
| | | | +-- Pilsbryoconcha_carinifera
| | | +--+
| | | +-- Pilsbryoconcha_exilis__1
| | |
| | | +----- Pseudodon_vagulus
| | +--+
| | | | +-- Pseudodon_mekongi
| | | +--+
| | | +-- Pseudodon_vondembuschianus
| | |
| | | +----- Bineurus_loeiensis
| | +--+
| | | +-- Bineurus_mouhotii
| | +--+
| | +-- Bineurus_exilis
| |
| | +-------------- Amblema_plicata
| | |
| | +-------------- Toxolasma_parvum
| | |
| | | +-- Obliquaria_reflexa
| | +-----------+
| | | +-- Plectomerus_dombeyanus
| | |
| | | +----------- Elliptoideus_sloatianus
| | +--+
| | | | +-------- Elliptio_complanata
| | | +--+
| | | | +----- Eurynia_dilatata
| +-----+ +--+
| | | | +-- Fusconaia_flava
| | | +--+
| | | +-- Pleurobema_sintoxia
| | |
| | | +-------- Ellipsaria_lineolata
| | | |
| | | +-------- Potamilus_leptodon
| | | |
| | | +-------- Medionidus_conradicus
| | | |
+-----+ | +-------- Potamilus_alatus
| | |
| | +-------- Ptychobranchus_subtentus
| +-----+
| +-------- Truncilla_truncata
| |
| +-------- Venustaconcha_ellipsiformis
| |
| +-------- Cambarunio_iris
| |
| +-------- Leaunio_lienosus
| |
| | +----- Ortmanniana_ligamentina
| +--+
| | +-- Lampsilis_cardium
| +--+
| +-- Lampsilis_ornata
|
| +-------------- Hyriopsis_desowitzi
| |
| +-------------- Hyriopsis_myersiana
| |
| +-------------- Hyriopsis_panhai
| |
| +--+ +-- Hyriopsis_bialata
| | +-----------+
| | | +-- Hyriopsis_khoratensis
| | |
| | | +----------- Rectidens_sumatrensis
| | | |
| | +--+ +-- Ensidens_telus
| | | +-----+
| | | | +-- Ensidens_ingallsianus
| | +--+
| | | +----- Ensidens_sagittarius
| | +--+
+--+ | +-- Ensidens_spiculus
| | +--+
| | +-- Ensidens_jaculus
| |
| | +----------- Physunio_modelli
| | |
| | +----------- Lens
| | |
| | +----------- Physunio_superbus
| | |
| | | +-------- Yaukthwa_inlenensis
| | +--+
| +-----+ | +----- Yaukthwa_panhai
| | +--+
| | | +-- Yaukthwa_nesemanni
| | +--+
| | +-- Yaukthwa_rectangularis
| |
| | +-- Lens_comptus
| | +-----+
| | | +-- Lens_novoselovi
| +--+
| | +----- Lens_eximius
| +--+
| | +-- Lens_contradens
| +--+
| +-- Lens_micropterus
|
| +----------------- Acuticosta_chinensis
| |
| +----------------- Alasmidonta_marginata
| |
| +----------------- Anodonta_anatina
| |
| +----------------- Anemina_arcaeformis
| |
| +----------------- Anodonta_cygnea
| |
| +----------------- Cristaria_plicata
| |
| +----------------- Anodonta_nuttalliana
| |
| +----------------- Lasmigona_costata
| |
| +----------------- Lepidodesma_languilati
| |
| +----------------- Pseudanodonta_complanata
| |
| +----------------- Pyganodon_grandis
| |
| +----------------- Simpsonaias_ambigua
| |
| +----------------- Strophitus_undulatus
| |
| | +-- Schistodesmus_lampreyanus
| +--------------+
| | +-- Schistodesmus_spinosus
| |
| | +-- Utterbackia_imbecillis
| +--------------+
| | +-- Utterbackia_peninsularis
| |
| | +-------- Lanceolaria_gladiola
| +--------+
| | | +----- Lanceolaria_grayii
| | +--+
+--+ | +-- Lanceolaria_lanceolata
| +--+
| +-- Lanceolaria_triformis
|
| +-------- Unio_tumidus
+--------+
| | +----- Unio_crassus
| +--+
| | +-- Unio_delphinus
| +--+
| +-- Unio_pictorum
|
| +-------- Aculamprotula_tortuosa
| |
+--------+ +-- Aculamprotula_coreana
| | +--+
| | | +-- Aculamprotula_scripta
| +--+
| | +-- Aculamprotula_fibrosa
| +--+
| +-- Aculamprotula_tientsinensis
|
| +-------- Cuneopsis_heudei
| |
| +-------- Cuneopsis_celtiformis
| |
+--------+-------- Tchangsinaia_piscicula
| |
| +-------- Cuneopsis_rufescens
| |
| | +----- Nodularia_huana
| +--+
| | +-- Nodularia_douglasiae
| +--+
| +-- Nodularia_nuxpersicae
|
| +-------------- Pletholophus_tenuis
| |
+--+ +-- Sinanodonta_angula
| +--------+
| | +-- Sinanodonta_lucida
+--+
| +-------- Sinanodonta_woodiana
+--+
| +----- Sinanodonta_jourdyi
+--+
| +-- Sinanodonta_elliptica
+--+
+-- Sinanodonta_pacifica
Graf et al., 2015 [Publ 3619], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
14 |
12 |
1 |
6 |
5 |
2 |
-- |
+-- Unio_pictorum
+-----+
| +-- Margaritifera_margaritifera
+--------+
| | +----- Chambardia_wahlbergi
| +--+
| | +-- Tamsiella_schroeteriana
| +--+
| +-- Etheria_elliptica
|
| +----------- Westralunio_carteri **
| |
--+ | +----- Alathyria_jacksoni **
| +--+ +--+
| | | | | +-- Velesunio_ambiguus **
| | | | +--+
| | +--+ +-- Alathyria_profuga **
| | |
| | | +----- Alathyria_pertexta **
| | +--+
| | | +-- Microdontia_anodontaeformis **
+--+ +--+
| +-- Lortiella_froggatti **
|
| +----- Diplodon_suavidicus **
| +--+
| | | +-- Triplodon_corrugatus **
| | +--+
| | +-- Castalia_ambigua **
+-----+
| +-- Hyridella_depressa **
| +--+
| | +-- Hyridella_drapeta **
+--+
| +-- Hyridella_australis **
+--+
+-- Cucumerunio_novaehollandiae **
Huang et al., 2019 [Publ 3843], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
14 |
12 |
1 |
184 |
182 |
2 |
-- |
+----- Alathyria_jacksoni **
+--+
| | +-- Alathyria_profuga **
| +--+
+--------+ +-- Velesunio_ambiguus **
| |
| | +----- Alathyria_pertexta **
| +--+
| | +-- Lortiella_froggatti **
+--------------------------------+ +--+
| | +-- Microdontia_anodontaeformis **
| |
| | +-------------- Westralunio_carteri **
| | |
| +--+ +----------- Echyridella_menziesii **
| | |
| | | +----- Diplodon_suavidicus **
| +--+ +--+
| | | | +-- Castalia_ambigua **
| | | +--+
| +--+ +-- Triplodon_corrugatus **
| |
| | +----- Hyridella_australis **
| +--+
| | +-- Hyridella_depressa **
| +--+
| +-- Hyridella_drapeta **
|
| +-------- Anodontites_trapesialis
| +--+
| | | +----- Mutela_rostrata
| | +--+
| | | +-- Mutela_dubia
| | +--+
| +-----------------------------------+ +-- Mutela_hargeri
| | |
| | | +----- Mycetopoda_siliquosa
| | | +--+
--+ | | | | +-- Tamsiella_schroeteriana
| | | | +--+
| | +--+ +-- Lamproscapha_ensiformis
| | |
| | | +----- Etheria_elliptica
| | +--+
| | | +-- Aspatharia_pfeifferiana
| | +--+
| | +-- Chambardia_wahlbergi
| |
| | +----------------- Gibbosula_rochechouartii
| | |
| | | +-------------- Gibbosula_laosensis
| | +--------------------------+ |
| | | | | +----------- Cumberlandia_monodonta
| | | | | |
| | | +--+ | +----- Pseudunio_marocanus
| | | | +-----+
| | | | | | +-- Pseudunio_auricularius
| | | | | +--+
| | | +--+ +-- Pseudunio_homsensis
| | | |
| | | | +-- Margaritifera_dahurica
| | | | +-----+
| | | | | +-- Margaritifera_margaritifera
| | | | |
| | | +--+ +----- Margaritifera_laevis
| | | | |
| | | | | +-- Margaritifera_falcata
| | | +--+--+
+--+ | | +-- Margaritifera_middendorffi
| | |
| | | +-- Margaritifera_hembeli
| | +--+
| | +-- Margaritifera_marrianae
| |
| | +----- Trapezidens_exolescens
| | +-----------------+
| | | | +-- Trapezidens_dolichorhynchus
| | | +--+
| | +--+ +-- Trapezidens_angustior
| | | |
| | | | +-------------------- Lamellidens_corrianus
| | | | |
| | | +--+ +----------------- Lamellidens_savadiensis
| | | | |
| | | +--+ +-- Lamellidens_lamellatus
| | | | +-----------+
| | | | | +-- Lamellidens_jenkinsianus
| | | +--+
| | | | +----------- Lamellidens_generosus
| | | +--+
| | | | +-------- Lamellidens_mainwaringi__2
| | | +--+
| | +--------------+ | +----- Lamellidens_mainwaringi__1
| | | | +--+
| | | | | +-- Lamellidens_ferrugineus
| | | | +--+
| | | | +-- Lamellidens_marginalis
| | | |
| | | | +----- Parreysia_keralaensis
| | | | +-----------------+
| | | | | | +-- Parreysia_nagpoorensis
| | | | | +--+
| | | | | +-- Parreysia_plagiosoma
+--+ | | |
| | | | +-- Leoparreysia_tavoyensis
| | | | |
| | +--+ +-----------------+-- Leoparreysia_canefrii
| | | | |
| | | | +-- Leoparreysia_olivacea
| | | |
| | | | +-------- Coelatura_aegyptiaca__2
| | | | |
| | | | +--------+ +----- Coelatura_aegyptiaca__1
| | +--+ | | |
| | | | | +----- Nitia_teretiuscula
| | | | +--+
| | | | | +-- Coelatura_gabonensis
| | | | | |
| | | | +--+-- Coelatura_kunenensis
| | | | |
| | +--+ +-- Prisodontopsis_mweruensis
| | |
| | | +-------- Indonaia_shurtleffiana
| | | |
| | | +-----+ +----- Indonaia_bonneaudii
| | | | | |
| | | | +--+----- Indonaia_rugosa
| | | | |
| | | | | +-- Indonaia_andersoniana
| | +--+ +--+
| | | +-- Indonaia_caerulea
| | |
| | | +----------- Indochinella_pugio
| | | |
| | | | +----- Scabies_humilis
| | +--+ +--+
| | | | | +-- Scabies_phaselus
| | | | +--+
| | +--+ +-- Scabies_mandarinus
| | |
| | | +----- Radiatula_myitkyinae
| | +--+
| | | +-- Radiatula_chaudhurii
| | +--+
| | +-- Radiatula_mouhoti
| |
| | +-------------------------------- Acuticosta_chinensis
| | |
| | | +----------------------------- Lepidodesma_languilati
| | | |
| | | | +-------- Aculamprotula_tortuosa
| | | | |
| | | | +-----------+ +----- Aculamprotula_coreana
| | | | | | |
| | | | | +--+----- Aculamprotula_scripta
| | | | | |
+--+ | | | | +-- Aculamprotula_fibrosa
| | | | +--+
| | | | +-- Aculamprotula_tientsinensis
| +-----+ +--------+
| | | | | +-------- Unio_tumidus
| | | | | +--------+
| | | | | | | +----- Unio_crassus
| | | | | | +--+
| | | | | | | +-- Unio_delphinus
| | | | | | +--+
| | | | +--+ +-- Unio_pictorum
| | | | |
| | | | | +-- Schistodesmus_lampreyanus
| | | | | +-----------+
| | | | | | +-- Schistodesmus_spinosus
| | | | | |
| | | | +--+ +----- Nodularia_douglasiae
| | +--+ | +-----+
| | | | | | +-- Nodularia_huana
| | | | | +--+
| | | +--+ +-- Nodularia_nuxpersicae
| | | |
| | | | +-------- Tchangsinaia_piscicula
| | | +--+
| | | | +----- Cuneopsis_rufescens
| | | +--+
| | | | +-- Cuneopsis_heudei
| | | +--+
| | | +-- Cuneopsis_celtiformis
| | |
| | | +-------- Lanceolaria_gladiola
| | | |
| | | +-----------------+ +----- Lanceolaria_grayii__1
| | | | | |
| | | | +--+----- Lanceolaria_grayii__2
| | | | |
| | | | | +-- Lanceolaria_lanceolata
| | | | +--+
| | | | +-- Lanceolaria_triformis
| | | |
| | | | +-------------------- Anemina_arcaeformis
| | | | |
| | | | +--+ +----------------- Cristaria_plicata
| | +--+ | | |
| | | | +--+ +-------------- Pletholophus_tenuis
| | | | | |
| | | | +--+ +-- Sinanodonta_angula
| | | | | +--------+
| | | | | | +-- Sinanodonta_lucida
| | | | +--+
| | | | | +-------- Sinanodonta_woodiana
| | | | +--+
| | | | | +----- Sinanodonta_jourdyi
| | | | +--+
| | +--+ | +-- Sinanodonta_elliptica
| | | +--+
| | | +-- Sinanodonta_pacifica
| | |
| | | +-------- Alasmidonta_marginata
| | | |
| | | +-------- Lasmigona_costata
| | | |
| | | +-------- Pseudanodonta_complanata
| | | |
| | | +-------- Simpsonaias_ambigua
+--+ | |
| +--------------+-------- Strophitus_undulatus
| |
| | +----- Anodonta_anatina
| +--+
| | | +-- Anodonta_cygnea
| | +--+
| | +-- Anodonta_nuttalliana
| |
| | +----- Pyganodon_grandis
| +--+
| | +-- Utterbackia_imbecillis
| +--+
| +-- Utterbackia_peninsularis
|
| +----------- Hyriopsis_desowitzi
| |
| +----------- Rectidens_sumatrensis
| |
| | +-- Hyriopsis_bialata
| | |
| | +-- Hyriopsis_myersiana
| +-----+--------+
| | | +-- Hyriopsis_panhai
| | | |
| | | +-- Hyriopsis_khoratensis
| | |
| | | +-- Ensidens_ingallsianus
| | | +-----+
| | | | +-- Ensidens_telus
| | +--+
| | | +----- Ensidens_sagittarius
| +-----------------+ +--+
| | | | +-- Ensidens_spiculus
| | | +--+
| | | +-- Ensidens_jaculus
| | |
| | | +----- Physunio
| | | +--------+
| | | | | +-- Physunio_modelli
| | | | +--+
| | | | +-- Physunio_superbus
| | | |
| | +--+ +-------- Yaukthwa_inlenensis
| | | +--+
| | | | | +----- Yaukthwa_panhai
| | | | +--+
| | | | | +-- Yaukthwa_nesemanni
| | | | +--+
| | +--+ +-- Yaukthwa_rectangularis
| | |
| | | +-- Lens_contradens
| | | +-----+
| | | | +-- Lens_micropterus
| | +--+
| | | +----- Lens_eximius
| | +--+
| | | +-- Lens_comptus
| | +--+
| | +-- Lens_novoselovi
| |
| | +-- Megalonaias_nervosa
| | +--+
+--+ | +-- Uniomerus_carolinianus
| +-----------------------+
| | | +-- Quadrula_quadrula
| | +--+
| | +-- Quadrula_verrucosa
| |
| | +-------- Elliptoideus_sloatianus
| +--+ |
| | | +-----------------+ +-- Elliptio_complanata
| | | | | +--+
| | | | | | +-- Eurynia_dilatata
| | | | +--+
| | | | | +-- Fusconaia_flava
| | +--+ +--+
| | | +-- Pleurobema_sintoxia
| | |
| | | +----------------------- Amblema_plicata
| | | |
| | | | +----- Toxolasma_parvum
| | +--+ +--------------+
| | | | | +-- Obliquaria_reflexa
| | | | +--+
| | | | +-- Plectomerus_dombeyanus
| | +--+
| | | +----------------- Potamilus_leptodon
| | | |
| | | | +-------------- Potamilus_alatus
| | +--+ |
| | | | +----------- Venustaconcha_ellipsiformis
| | | | |
| | +--+ +----------- Leaunio_lienosus
| | | |
+--+ | | +-- Ellipsaria_lineolata
| | +--------+
| +--+ +-- Truncilla_truncata
| |
| | +-- Medionidus_conradicus
| +--------+
| | +-- Ptychobranchus_subtentus
| |
| | +-------- Cambarunio_iris
| +--+
| | +----- Ortmanniana_ligamentina
| +--+
| | +-- Lampsilis_cardium
| +--+
| +-- Lampsilis_ornata
|
| +----- Chamberlainia_somsakpanhai
| +-----------------------+
| | | +-- Sinohyriopsis_cumingii
| | +--+
| | +-- Sinohyriopsis_schlegelii
| |
| | +-- Potomida_littoralis
| | +-----+
| | | +-- Pronodularia_japanensis
| | +-----------------+
+--+ | | +----- Lamprotula_leaii
| | +--+
| | | +-- Lamprotula_caveata
| | +--+
| | +-- Lamprotula_cornuumlunae
| |
| | +-------- Sinosolenaia_carinata
| | +--+
| | | | +----- Sinosolenaia_recognita
| | | +--+
+--+ | | +-- Sinosolenaia_oleivora
| | +--+
| +-----------+ +-- Sinosolenaia_iridinea
| | |
| | | +----- Gonidea_angulata
| | | +--+
| | | | | +-- Leguminaia_wheatleyi
| | | | +--+
| | +--+ +-- Microcondylaea_bonellii
| | |
| | | +-- Ptychorhynchus_pfisteri
| | | |
+--+ +-----+-- Parvasolenaia_rivularis
| |
| +-- Koreosolenaia_sitgyensis
|
| +----- Indopseudodon_salwenianus
| +--------------+
| | | +-- Indopseudodon_indawgyiensis
| | +--+
| | +-- Indopseudodon_bogani
| |
| | +-------- Pilsbryoconcha_exilis__3
+--+ +--------+
| | | +----- Pilsbryoconcha_exilis__2
| | +--+
| | | +-- Pilsbryoconcha_exilis__1
| | +--+
| | +-- Pilsbryoconcha_carinifera
| |
+--+ +----------- Nyeinchanconcha_nyeinchani
| +--+
| | | +-------- Thaiconcha_callifera
| | +--+
| | | +----- Bineurus_loeiensis
| | +--+
| | | +-- Bineurus_mouhotii
+--+ +--+
| +-- Bineurus_exilis
|
| +----- Pseudodon_vagulus
| +--+
| | | +-- Pseudodon_mekongi
| | +--+
+-----+ +-- Pseudodon_vondembuschianus
|
| +-- Sundadontina_cumingii
| |
+-----+-- Sundadontina_tumida__1
|
+-- Sundadontina_tumida__2
Marshall et al., 2014 [Publ 3593], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
1 |
4 |
4 |
1 |
-- |
+----- Cumberlandia_monodonta
+-----------+
| | +-- Pyganodon_grandis
| +--+
| +-- Unio_pictorum
--+
| +-------------- Amblema_plicata
| |
| | +----- Echyridella_aucklandica **
| | +-----+
+--+ | | +-- Echyridella_menziesii **
| | +--+
| | +-- Echyridella_onekaka **
| |
+--+ +----- Velesunio_ambiguus **
| +--+
| | | +-- Velesunio **
| | +--+
| | +-- Lortiella **
+--+
| +-- Cucumerunio_novaehollandiae **
| +--+
| | +-- Hyridella_australis **
+--+
| +-- Diplodon_chilensis **
+--+
+-- Diplodon_gratus **
Walker et al., 2006 [Publ 2666], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
7 |
1 |
26 |
25 |
2 |
-- |
+----------------- Castalia_stevensi **
|
+----------------- Diplodon_gratus **
+--------+
| | +-------------- Hyridella_depressa **
| | |
| | +-------------- Hyridella_australis **
| +--+
| | +----------- Echyridella_menziesii **
| | |
| | | +-- Velesunio_ambiguus **
| +--+ +-----+
| | | +-- Velesunio__3 **
| | |
| +--+ +----- Velesunio__2 **
| | |
| | | +-- Alathyria_jacksoni **
| +--+--+
| | +-- Velesunio__1 **
| |
--+ | +-- Velesunio_angasi **
| +--+
| +-- Lortiella_rugata **
|
| +-- Mutela_dubia
| +--------------+
| | +-- Mutela_rostrata
| +-----+
| | | +-------------- Etheria_elliptica
| | | |
| | +--+ +----------- Chambardia_rubens
| | | |
| | +--+ +-------- Mulleria_rivoli
| | | |
| | +--+ +-- Anodontites_guanarensis
| | | +--+
| | | | +-- Anodontites_trigona
+--+ +--+
| | +-- Tamsiella_tamsiana
| +--+
| +-- Monocondylaea_minuana
|
| +-- Pyganodon_fragilis
| +-----------------+
| | +-- Pyganodon_grandis
| |
| | +----------- Cumberlandia_monodonta
| | |
+--+ +-----+ +-------- Pseudunio_auricularius
| | | |
| | +--+ +-- Margaritifera_margaritifera
| | | +--+
| | | | +-- Margaritifera_dahurica
| | +--+
+--+ | +-- Margaritifera_falcata
| +--+
| +-- Margaritifera_middendorffi
|
| +-- Gonidea_angulata
| +-----------+
| | +-- Sundadontina_tumida
+--+
| +----------- Cyrtonaias_tampicoensis
| |
+--+----------- Fusconaia_flava
|
| +-------- Potamilus_purpuratus
+--+
| +----- Lampsilis_teres
+--+
| +-- Ligumia_recta
+--+
+-- Ortmanniana_ligamentina
Graf & Cummings, 2006 [Publ 1968], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
7 |
1 |
38 |
37 |
2 |
-- |
+----- Velesunio_ambiguus **
+--------+
| | +-- Velesunio_angasi **
| +--+
+--+ +-- Lortiella_rugata **
| |
| | +----------- Echyridella_menziesii **
| | |
| +--+ +-- Hyridella_depressa **
| | +-----+
| | | +-- Hyridella_australis **
| +--+
| | +----- Castalia **
| +--+
+--------------------+ | +-- Diplodon_chilensis **
| | +--+
| | +-- Diplodon_gratus **
| |
| | +----- Etheria_elliptica
| | +-----+
| | | | +-- Mulleria_rivoli
| | | +--+
| | | +-- Pseudomulleria_dalyi
| | |
| +-----+ +----- Monocondylaea_minuana
| | +--+
| | | | +-- Anodontites_trigona
| | | +--+
| | | +-- Anodontites_guanarensis
--+ +--+
| | +-- Chambardia_wahlbergi
| | +--+
| | | +-- Aspatharia_rugifera
| +--+
| | +-- Mutela_dubia
| +--+
| +-- Mutela_rostrata
|
| +-- Margaritifera_margaritifera
| +--------------------------------+
| | +-- Cumberlandia_monodonta
| |
| | +-- Pilsbryoconcha_exilis
| | +-----------------------------+
+--+ | +-- Pseudodon_vondembuschianus
| |
| | +----------------------------- Coelatura_aegyptiaca
| | |
| | +----------------------------- Lens_contradens
+--+ |
| | +-- Unio_pictorum
| | +--------+
| | | +-- Unio_caffer
| +-----------------+
| | | +-------- Lasmigona_costata
+--+ +--+
| | +----- Alasmidonta_marginata
| +--+
| | +-- Pyganodon_grandis
| +--+
| +-- Strophitus_undulatus
|
| +-------------------------- Gonidea_angulata
| |
| | +-- Quadrula_verrucosa
+--+ +--------------------+
| | +-- Quadrula_quadrula
| |
+--+ +-------------------- Amblema_plicata
| |
| | +----- Eurynia_dilatata
+--+ +-----------+
| | | +-- Fusconaia_flava
| | +--+
| | +-- Pleurobema_sintoxia
+--+
| +-------------- Obliquaria_reflexa
| |
| +-------------- Truncilla_truncata
+--+
+-------------- Ptychobranchus_fasciolaris
|
| +----------- Ligumia_recta
+--+
| +-------- Cambarunio_iris
+--+
| +----- Epioblasma_triquetra
+--+
| +-- Lampsilis_cardium
+--+
+-- Ortmanniana_ligamentina
Hoeh et al., 2009 [Publ 2716], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
7 |
1 |
38 |
37 |
2 |
-- |
+----- Velesunio_ambiguus **
+--------+
| | +-- Velesunio_angasi **
| +--+
+-----------------------------+ +-- Lortiella_rugata **
| |
| | +----------- Echyridella_menziesii **
| | |
| +--+ +-- Hyridella_depressa **
| | +-----+
| | | +-- Hyridella_australis **
| +--+
| | +----- Castalia **
| +--+
| | +-- Diplodon_chilensis **
| +--+
| +-- Diplodon_gratus **
--+
| +----- Chambardia_wahlbergi
| |
| +--------+----- Aspatharia_rugifera
| | |
| | | +-- Mutela_rostrata
| +--------------------------+ +--+
| | | +-- Mutela_dubia
| | |
| | | +----------- Etheria_elliptica
| | +--+
| | | +-------- Mulleria_rivoli
| | +--+
| | | +----- Monocondylaea_minuana
+--+ +--+
| | +-- Anodontites_trigona
| +--+
| +-- Anodontites_guanarensis
|
| +-- Margaritifera_margaritifera
| +-----------------------------------+
| | +-- Cumberlandia_monodonta
| |
+--+ +----------------------------------- Coelatura_aegyptiaca
| |
| | +-------------------------------- Pseudomulleria_dalyi
| | |
+--+ | +-- Unio_pictorum
| | +--------+
| | | +-- Unio_caffer
| | +-----------------+
+--+ | | +-------- Lasmigona_costata
| | +--+
| | | +----- Alasmidonta_marginata
| | +--+
| | | +-- Strophitus_undulatus
| | +--+
+--+ +-- Pyganodon_grandis
|
| +-------------------------- Lens_contradens
| |
| | +----- Gonidea_angulata
| | +-----------------+
| | | | +-- Pilsbryoconcha_exilis
+--+ | +--+
| | +-- Pseudodon_vondembuschianus
| |
| | +----------- Amblema_plicata
| | |
+--+ | +----- Obliquaria_reflexa
| +--------+ +--+
| | | | | +-- Quadrula_verrucosa
| | | | +--+
| | +--+ +-- Quadrula_quadrula
| | |
| | | +----- Eurynia_dilatata
+--+ +--+
| | +-- Pleurobema_sintoxia
| +--+
| +-- Fusconaia_flava
|
| +----------------- Ptychobranchus_fasciolaris
+--+
| +-------------- Truncilla_truncata
+--+
| +----------- Ortmanniana_ligamentina
+--+
| +-------- Lampsilis_cardium
+--+
| +----- Cambarunio_iris
+--+
| +-- Ligumia_recta
+--+
+-- Epioblasma_triquetra
Bolotov et al., 2017 [Publ 3761], fig. S2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
7 |
1 |
130 |
128 |
2 |
-- |
+-------------------- Chamberlainia_somsakpanhai
|
+-----------+ +----------------- Cumberlandia_monodonta
| | |
| | | +-------------- Gibbosula_laosensis
| +--+ |
| | | +-- Pseudunio_auricularius
| | +-----------+
| +--+ +-- Pseudunio_marocanus
| |
| | +-- Margaritifera_dahurica
| | +--------+
| | | +-- Margaritifera_margaritifera
| +--+
| | +-------- Margaritifera_falcata
| +--+
| | +----- Margaritifera_middendorffi
| +--+
| | +-- Margaritifera_laevis
| +--+
| +-- Margaritifera_marrianae
|
| +-- Tamsiella_schroeteriana
| +--+
| | +-- Etheria_elliptica
| +--------+
| | | +-- Aspatharia_pfeifferiana
| | +--+
| | +-- Chambardia_wahlbergi
| |
+-----------------+ +----- Diplodon_suavidicus **
| | +-----+
| | | | +-- Castalia_ambigua **
| | | +--+
| | | +-- Triplodon_corrugatus **
| +--+
| | +----- Velesunio_ambiguus **
| | +--+
| | | | +-- Alathyria_jacksoni **
| | | +--+
| +--+ +-- Alathyria_profuga **
| |
| | +----- Microdontia_anodontaeformis **
| +--+
| | +-- Alathyria_pertexta **
| +--+
| +-- Lortiella_froggatti **
|
| +-- Parreysia_nagpoorensis
| |
| +-----------+-- Parreysia_plagiosoma
| | |
| | +-- Parreysia_corrugata
| |
| +--------+ +-- Trapezidens_dolichorhynchus
| | | +--------+
| | | | +-- Trapezidens_exolescens
| | | |
--+ | +--+ +-------- Lamellidens_savadiensis
| | | |
| | | +-------- Lamellidens_corrianus
| | +--+
| | | +-- Lamellidens_marginalis
| | | +--+
| | | | +-- Lamellidens_ferrugineus
| +-----+ +--+
| | | | +-- Lamellidens_jenkinsianus
| | | +--+
| | | +-- Lamellidens_mainwaringi
| | |
| | | +-- Leoparreysia_olivacea
| | | +-----------------+
| | | | +-- Leoparreysia_tavoyensis
| | | |
| | | | +-------------- Nitia_teretiuscula
| | | | +--+
| | | | | | +----------- Coelatura_aegyptiaca__1
| | +--+ | +--+
| | | | | +-------- Coelatura_aegyptiaca__2
| | | | +--+
| | | | | +----- Coelatura_gabonensis
| | | | +--+
| | | | | +-- Coelatura_kunenensis
| | +--+ +--+
| | | +-- Prisodontopsis_mweruensis
| | |
| | | +-------- Indonaia_shurtleffiana
| | | +-----+
| | | | | +----- Indonaia_andersoniana
| | | | +--+
| | | | | +-- Indonaia_rugosa
| | | | +--+
| | +--+ +-- Indonaia_bonneaudii
| | |
| | | +-- Indochinella_pugio
| | | +--------+
| | | | +-- Scabies_crispata
| | | |
| | +--+ +-- Radiatula_myitkyinae
| | | +-----+
| | | | +-- Radiatula_chaudhurii
| | +--+
| | | +----- Scabies_mandarinus
| | +--+
| | | +-- Scabies_humilis
| | +--+
| | +-- Scabies_phaselus
| |
| | +----- Unio_tumidus
| | +-----+
| | | | +-- Unio_crassus
| | | +--+
| | | +-- Unio_pictorum
| | +-----+
+--+ | | +-------- Aculamprotula_tortuosa
| | | |
| | | +-------- Schistodesmus_lampreyanus
| | | |
| | +--+ +-- Nodularia_douglasiae
| | +-----+
| | | +-- Nodularia_nuxpersicae
| | |
| | | +----- Cuneopsis_heudei
| | +--+
| | | +-- Tchangsinaia_piscicula
| | +--+
| +--------+ +-- Cuneopsis_rufescens
| | |
| | | +-- Lanceolaria_grayii
| | | +-----------+
| | | | +-- Lanceolaria_gladiola
| | | |
| | | | +-- Cristaria_plicata
| | | | +--------+
| | | | | +-- Cristaria_radiata
| | | +--+
| | | | | +-------- Pletholophus_tenuis
| | | | +--+
| | | | | +----- Sinanodonta_lucida
| | +--+ +--+
| | | | +-- Sinanodonta_lauta
| | | +--+
| | | +-- Sinanodonta_jourdyi
| | |
| | | +-------- Anodonta_anatina
| | | |
| | | +-------- Pseudanodonta_complanata
| | | |
| | | | +-- Anodonta_cygnea
| | +-----+-----+
| | | +-- Anodonta_nuttalliana
| | |
| | | +----- Pyganodon_grandis
| | | |
| | | +----- Simpsonaias_ambigua
| | +--+
| | +----- Strophitus_undulatus
| | |
| | | +-- Alasmidonta_marginata
| | +--+
| | +-- Lasmigona_costata
| |
| | +-------- Lens_contradens
| | +-----+
+--+ | | +----- Lens_eximius
| | +--+
| | | +-- Lens_comptus
| | +--+
| +--+ +-- Lens_novoselovi
| | |
| | | +----- Physunio
| | | +-----+
| | | | | +-- Physunio_modelli
| | | | +--+
| | +--+ +-- Physunio_superbus
| | |
| | | +-------- Yaukthwa_inlenensis
| | +--+
| | | +----- Yaukthwa_panhai
| +-----+ +--+
| | | | +-- Yaukthwa_rectangularis
| | | +--+
| | | +-- Yaukthwa_nesemanni
| | |
| | | +----- Hyriopsis_panhai
| | | |
| | | +--------+----- Hyriopsis_khoratensis
| | | | |
| | | | | +-- Hyriopsis_myersiana
| | | | +--+
| | +--+ +-- Hyriopsis_bialata
| | |
| | | +----------- Hyriopsis_desowitzi
| | | |
| | | +----------- Rectidens_sumatrensis
| | +--+
| | | +-- Ensidens_telus
| | | +-----+
| | | | +-- Ensidens_ingallsianus
| | +--+
| | | +----- Ensidens_sagittarius
| | +--+
| | | +-- Ensidens_spiculus
+--+ +--+
| +-- Ensidens_jaculus
|
| +-------- Amblema_plicata
| +--+
| | | +----- Cambarunio_iris
| | +--+
| | | +-- Ortmanniana_ligamentina
| | +--+
| +--------+ +-- Lampsilis_cardium
| | |
| | | +-- Quadrula_quadrula
| | | +-----+
| | | | +-- Quadrula_verrucosa
| | +--+
| | | +----- Pleurobema_sintoxia
| | +--+
| | | +-- Elliptio_complanata
| | +--+
| | +-- Eurynia_dilatata
| |
| | +-------- Sinohyriopsis_cumingii
| | |
| | +-----+ +-- Lamprotula_caveata
+--+ | | +--+
| | | | +-- Lamprotula_leaii
| | +--+
| | | +-- Potomida_littoralis
| +--+ +--+
| | | +-- Pronodularia_japanensis
| | |
| | | +----------- Sinosolenaia_carinata
| | | |
| | | | +-- Sinosolenaia_oleivora
| | +--+--------+
| | | +-- Sinosolenaia_recognita
| | |
| | | +-------- Gonidea_angulata
| | +--+
| | | +----- Microcondylaea_bonellii
+--+ +--+
| | +-- Leguminaia_wheatleyi
| +--+
| +-- Leguminaia
|
| +-------- Pilsbryoconcha_exilis__3
| +-----+
| | | +----- Pilsbryoconcha_exilis__2
| | +--+
| | | +-- Pilsbryoconcha_exilis__1
| | +--+
| | +-- Pilsbryoconcha_carinifera
| |
+--+ +----------- Indopseudodon_indawgyiensis
| |
| | +-------- Thaiconcha_callifera
| +--+
| | | +----- Bineurus_exilis
| | +--+
| | | +-- Bineurus_mouhotii
+--+ +--+
| +-- Bineurus_loeiensis
|
| +----- Pseudodon_vagulus
| +--+
| | | +-- Pseudodon_mekongi
| | +--+
| | +-- Pseudodon_vondembuschianus
+--+
| +-- Pseudodon_inoscularis
| +--+
| | +-- Sundadontina_tumida
+--+
| +-- Nyeinchanconcha_nyeinchani
+--+
+-- Sundadontina_cumingii
Bolotov et al., 2016 [Publ 3743], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
7 |
1 |
21 |
18 |
2 |
-- |
+----- Diplodon_suavidicus **
+-----+
| | +-- Triplodon_corrugatus **
| +--+
| +-- Castalia_ambigua **
+--------------+
| | +----- Microdontia_anodontaeformis **
| | +--+
| | | | +-- Lortiella_froggatti **
| | | +--+
| +--+ +-- Alathyria_pertexta **
| |
| | +----- Velesunio_ambiguus **
| +--+
| | +-- Alathyria_jacksoni **
--+ +--+
| +-- Alathyria_profuga **
|
| +-------- Tamsiella_schroeteriana
| +--------------+
| | | +----- Etheria_elliptica
| | +--+
| | | +-- Aspatharia_pfeifferiana
| | +--+
| | +-- Chambardia_wahlbergi
+--+
| +-------- Lampsilis_cardium
| +-----------+
| | | +----- Anodonta_anatina
| | +--+
| | | +-- Unio_pictorum
| | +--+
+--+ +-- Unio_ravoisieri
|
| +-- Pseudunio_auricularius
| +--------------+
| | +-- Pseudunio_marocanus
| |
+--+ +-- Gibbosula_laosensis
| +-----------+
| | +-- Cumberlandia_monodonta
| |
+--+ +-- Margaritifera_dahurica
| +--------+
| | +-- Margaritifera_margaritifera
+--+
| +-------- Margaritifera_falcata
+--+
| +----- Margaritifera_laevis
+--+
| +-- Margaritifera_middendorffi
+--+
+-- Margaritifera_marrianae
Bolotov et al., 2017 [Publ 3761], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
7 |
1 |
130 |
128 |
2 |
-- |
+-------- Tamsiella_schroeteriana
+-----+
| | +----- Etheria_elliptica
| +--+
| | +-- Chambardia_wahlbergi
| +--+
| +-- Aspatharia_pfeifferiana
+-----------------------+
| | +----- Diplodon_suavidicus **
| | +-----+
| | | | +-- Castalia_ambigua **
| | | +--+
| | | +-- Triplodon_corrugatus **
| +--+
| | +----- Alathyria_pertexta **
| | +--+
| | | | +-- Microdontia_anodontaeformis **
| | | +--+
| +--+ +-- Lortiella_froggatti **
| |
| | +----- Velesunio_ambiguus **
| +--+
| | +-- Alathyria_jacksoni **
| +--+
| +-- Alathyria_profuga **
|
| +-- Trapezidens_dolichorhynchus
| +-----------+
| | +-- Trapezidens_exolescens
| +-----------+
| | | +----------- Lamellidens_corrianus
| | | |
| | +--+ +-------- Lamellidens_savadiensis
| | | |
| | +--+ +-- Lamellidens_jenkinsianus
| | | +--+
| | | | +-- Lamellidens_mainwaringi
| | +--+
| | | +-- Lamellidens_ferrugineus
| +--------+ +--+
| | | +-- Lamellidens_marginalis
| | |
| | | +-- Parreysia_nagpoorensis
| | | |
| | | +--------------------+-- Parreysia_plagiosoma
| | | | |
| | | | +-- Parreysia_corrugata
--+ | | |
| | | | +-- Leoparreysia_tavoyensis
| | +--+ +-----------------+
| | | | +-- Leoparreysia_olivacea
| | | |
| | | | +-------------- Nitia_teretiuscula
| | | | +--+
| | | | | | +----------- Coelatura_aegyptiaca__2
| | +--+ | +--+
| | | | | +-------- Coelatura_aegyptiaca__1
| | | | +--+
| | | | | +----- Prisodontopsis_mweruensis
| | | | +--+
| | | | | +-- Coelatura_gabonensis
| | +--+ +--+
| | | +-- Coelatura_kunenensis
| | |
| | | +-------- Indonaia_shurtleffiana
| | | +-----+
| | | | | +----- Indonaia_andersoniana
| | | | +--+
| | | | | +-- Indonaia_rugosa
| | | | +--+
| | +--+ +-- Indonaia_bonneaudii
| | |
| | | +-- Indochinella_pugio
| | | +--------+
| | | | +-- Scabies_crispata
| | | |
| | +--+ +-- Radiatula_myitkyinae
| | | +-----+
| | | | +-- Radiatula_chaudhurii
| | +--+
| | | +----- Scabies_mandarinus
| | +--+
| | | +-- Scabies_humilis
| | +--+
| | +-- Scabies_phaselus
| |
| | +-- Ensidens_telus
+--+ +-----+
| | +-- Ensidens_ingallsianus
| +-----+
| | | +----- Ensidens_sagittarius
| | +--+
| | | +-- Ensidens_spiculus
| | +--+
| +--+ +-- Ensidens_jaculus
| | |
| | | +-- Hyriopsis_bialata
| | | +--------+
| | | | +-- Hyriopsis_khoratensis
| | +--+
| | | +-------- Hyriopsis_panhai
| | +--+
| | | +----- Hyriopsis_myersiana
| | +--+
| +--------------+ | +-- Rectidens_sumatrensis
| | | +--+
| | | +-- Hyriopsis_desowitzi
| | |
| | | +----- Physunio
| | | +--------+
| | | | | +-- Physunio_modelli
| | | | +--+
| | | | +-- Physunio_superbus
| | | |
| | +--+ +-------- Yaukthwa_inlenensis
| | | +--+
| | | | | +----- Yaukthwa_panhai
| | | | +--+
| | | | | +-- Yaukthwa_rectangularis
| | | | +--+
| | +--+ +-- Yaukthwa_nesemanni
| | |
| | | +-- Lens_contradens
| | | +-----+
| | | | +-- Lens_micropterus
| | +--+
| | | +----- Lens_eximius
| | +--+
| | | +-- Lens_comptus
| | +--+
| | +-- Lens_novoselovi
| |
| | +-------------------- Indopseudodon_indawgyiensis
| | |
| | | +-------- Pilsbryoconcha_exilis__1
| | | +--------+
+--+ +--------+ | | +----- Pilsbryoconcha_exilis__3
| | | | +--+
| | | | | +-- Pilsbryoconcha_exilis__2
| | | | +--+
| | | | +-- Pilsbryoconcha_carinifera
| | +--+
| | | +-------- Sundadontina_tumida
| | | +-----+
| | | | | +----- Pseudodon_vagulus
| | | | +--+
| | | | | +-- Pseudodon_mekongi
| | | | +--+
| | +--+ +-- Pseudodon_vondembuschianus
| | |
| | | +-- Nyeinchanconcha_nyeinchani
| | | +--------+
| | | | +-- Sundadontina_cumingii
| | +--+
| | | +-------- Thaiconcha_callifera
| | +--+
| | | +----- Bineurus_exilis
| | +--+
| | | +-- Bineurus_mouhotii
| | +--+
| | +-- Bineurus_loeiensis
| |
| | +-------- Amblema_plicata
| | +--+
| | | | +----- Ortmanniana_ligamentina
| | | +--+
| | | | +-- Lampsilis_cardium
| | | +--+
+--+ +-----+ +-- Cambarunio_iris
| | |
| | | +-- Quadrula_quadrula
| | | +-----+
| | | | +-- Quadrula_verrucosa
| | +--+
| | | +----- Pleurobema_sintoxia
| | +--+
| | | +-- Eurynia_dilatata
| | +--+
| | +-- Elliptio_complanata
| +--------+
| | | +-- Sinohyriopsis_cumingii
| | | +-----+
| | | | +-- Chamberlainia_somsakpanhai
| | | |
| | | +-----+ +-- Potomida_littoralis
| | | | | +--+
| | | | | | +-- Pronodularia_japanensis
| | | | +--+
| | | | | +-- Lamprotula_caveata
| | | | +--+
| | +--+ +-- Lamprotula_leaii
| | |
| | | +----- Sinosolenaia_carinata
| | | +-----+
| | | | | +-- Sinosolenaia_oleivora
| | | | +--+
| | +--+ +-- Sinosolenaia_recognita
| | |
| | | +-------- Gonidea_angulata
| | +--+
+--+ | +----- Microcondylaea_bonellii
| +--+
| | +-- Leguminaia_wheatleyi
| +--+
| +-- Leguminaia
|
| +----- Unio_tumidus
| +--------+
| | | +-- Unio_pictorum
| | +--+
| | +-- Unio_crassus
| +--------+
| | | +----------- Aculamprotula_tortuosa
| | | |
| | | | +-- Nodularia_douglasiae
| | +--+ +-----+
| | | | +-- Nodularia_nuxpersicae
| | | |
| | +--+ +-- Cuneopsis_rufescens
| | | +--+
| | | | +-- Cuneopsis_heudei
| | +--+
+--+ | +-- Tchangsinaia_piscicula
| +--+
| +-- Schistodesmus_lampreyanus
|
| +-- Lanceolaria_grayii
| +-----------------+
| | +-- Lanceolaria_gladiola
| |
| | +-- Cristaria_plicata
| | +--------+
| | | +-- Cristaria_radiata
+--+ +-----+
| | | +-------- Pletholophus_tenuis
| | +--+
| | | +----- Sinanodonta_lucida
| | +--+
| | | +-- Sinanodonta_jourdyi
+--+ +--+
| +-- Sinanodonta_lauta
|
| +-------------- Anodonta_anatina
| |
| | +----- Pseudanodonta_complanata
+--+ +-----+
| | | +-- Anodonta_cygnea
| | +--+
| | +-- Anodonta_nuttalliana
+--+
| +-- Lasmigona_costata
| +-----+
| | +-- Alasmidonta_marginata
+--+
| +----- Simpsonaias_ambigua
+--+
| +-- Pyganodon_grandis
+--+
+-- Strophitus_undulatus
Bolotov et al., 2016 [Publ 3743], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
7 |
1 |
21 |
18 |
2 |
-- |
+----- Diplodon_suavidicus **
+-----+
| | +-- Triplodon_corrugatus **
| +--+
| +-- Castalia_ambigua **
+--------------+
| | +----- Microdontia_anodontaeformis **
| | +--+
| | | | +-- Alathyria_pertexta **
| | | +--+
| +--+ +-- Lortiella_froggatti **
| |
| | +----- Velesunio_ambiguus **
| +--+
| | +-- Alathyria_jacksoni **
--+ +--+
| +-- Alathyria_profuga **
|
| +-------- Tamsiella_schroeteriana
| +--------------+
| | | +----- Etheria_elliptica
| | +--+
| | | +-- Chambardia_wahlbergi
| | +--+
| | +-- Aspatharia_pfeifferiana
+--+
| +-------- Lampsilis_cardium
| +-----------+
| | | +----- Anodonta_anatina
| | +--+
| | | +-- Unio_pictorum
| | +--+
+--+ +-- Unio_ravoisieri
|
| +-- Pseudunio_auricularius
| +--------------+
| | +-- Pseudunio_marocanus
| |
+--+ +-- Gibbosula_laosensis
| +-----------+
| | +-- Cumberlandia_monodonta
| |
+--+ +-- Margaritifera_margaritifera
| +--------+
| | +-- Margaritifera_dahurica
+--+
| +-------- Margaritifera_falcata
+--+
| +----- Margaritifera_laevis
+--+
| +-- Margaritifera_marrianae
+--+
+-- Margaritifera_middendorffi
Hoeh et al., 2009 [Publ 2716], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
7 |
1 |
37 |
36 |
2 |
-- |
+----- Velesunio_ambiguus **
+--------+
| | +-- Velesunio_angasi **
| +--+
+--------------------------------+ +-- Lortiella_rugata **
| |
| | +----------- Echyridella_menziesii **
| | |
| +--+ +-- Hyridella_depressa **
| | +-----+
| | | +-- Hyridella_australis **
| +--+
| | +----- Castalia **
| +--+
| | +-- Diplodon_chilensis **
| +--+
| +-- Diplodon_gratus **
--+
| +-- Etheria_elliptica
| +--------+
| | +-- Mulleria_rivoli
| |
| | +----- Monocondylaea_minuana
| +--------------------------------+ +--+
| | | | | +-- Anodontites_trigona
| | | | +--+
| | | | +-- Anodontites_guanarensis
| | +--+
| | | +-- Mutela_rostrata
| | | +--+
+--+ | | +-- Mutela_dubia
| +--+
| | +-- Chambardia_wahlbergi
| +--+
| +-- Aspatharia_rugifera
|
| +----------------------------------------- Coelatura_aegyptiaca
| |
+--+ +-------------------------------------- Pseudomulleria_dalyi
| |
| | +-- Margaritifera_margaritifera
+--+ +--------------------------------+
| | +-- Cumberlandia_monodonta
| |
| | +-- Pilsbryoconcha_exilis
+--+ +-----------------------------+
| | +-- Pseudodon_vondembuschianus
| |
| | +----------------------------- Lens_contradens
+--+ |
| | +-- Unio_pictorum
| | +--------+
| | | +-- Unio_caffer
| | +--------------+
+--+ | | +-------- Lasmigona_costata
| | +--+
| | | +----- Alasmidonta_marginata
| | +--+
| | | +-- Strophitus_undulatus
| | +--+
+--+ +-- Pyganodon_grandis
|
| +----------------------- Gonidea_angulata
| |
| | +-- Eurynia_dilatata
| | |
| | +-----+-- Pleurobema_sintoxia
+--+ | |
| +-----------+ +-- Fusconaia_flava
| | |
| | | +----- Obliquaria_reflexa
| | +--+
+--+ | +-- Quadrula_verrucosa
| +--+
| +-- Quadrula_quadrula
|
| +----------------- Ptychobranchus_fasciolaris
+--+
| +-------------- Truncilla_truncata
+--+
| +----------- Ortmanniana_ligamentina
+--+
| +-------- Lampsilis_cardium
+--+
| +----- Cambarunio_iris
+--+
| +-- Ligumia_recta
+--+
+-- Epioblasma_triquetra
Hoeh et al., 2009 [Publ 2716], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
7 |
1 |
37 |
36 |
2 |
-- |
+-- Margaritifera_margaritifera
+--------------------+
| +-- Cumberlandia_monodonta
|
| +----- Velesunio_ambiguus **
| +--------+
| | | +-- Velesunio_angasi **
| | +--+
--+ +-----+ +-- Lortiella_rugata **
| | |
| | | +----------- Echyridella_menziesii **
| | | |
| | +--+ +-- Hyridella_depressa **
| | | +-----+
| | | | +-- Hyridella_australis **
| | +--+
+--+ | +----- Castalia **
| +--+
| | +-- Diplodon_chilensis **
| +--+
| +-- Diplodon_gratus **
|
| +----------------- Pseudomulleria_dalyi
| |
+--+ +-------------- Mulleria_rivoli
| |
+--+ +----------- Etheria_elliptica
| |
| | +----- Monocondylaea_minuana
+--+ +--+
| | | +-- Anodontites_trigona
| | +--+
| | +-- Anodontites_guanarensis
+--+
| +-- Mutela_rostrata
| +--+
| | +-- Mutela_dubia
+--+
| +-- Chambardia_wahlbergi
+--+
+-- Aspatharia_rugifera
Baker et al., 2004 [Publ 2685], fig. 2-3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
9 |
1 |
1 |
1 |
1 |
-- |
+-------------------- Epioblasma_triquetra
|
| +----- Hyridella_depressa **
--+ +-----------+
| | | +-- Hyridella_australis **
| | +--+
| | +-- Diplodon_gratus **
+--+
| +-- Echyridella_menziesii **
| +-----------+
| | +-- Castalia_stevensi **
| |
+--+ +-- Velesunio__3 **
| +--------+
| | +-- Velesunio__4 **
+--+
| +-------- Velesunio__2 **
+--+
| +----- Velesunio_angasi **
+--+
| +-- Velesunio__1 **
+--+
+-- Alathyria_jacksoni **
Santos-Neto et al., 2016 [Publ 3739], fig. 3, 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
6 |
1 |
0 |
0 |
0 |
-- |
+-------------- Diplodon_suavidicus **
|
| +----- Triplodon_corrugatus **
--+ +-----+
| | | +-- Prisodon_obliquus **
| | +--+
+--+ +-- Prisodon_syrmatophorus **
|
| +-------- Callonaia_duprei **
+--+
| +----- Castalia_ambigua__2 **
+--+
| +-- Castalia_ambigua__1 **
+--+
+-- Castaliella_quadrata **
Baker et al., 2003 [Publ 2690], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
1 |
1 |
1 |
1 |
-- |
+-------------- Epioblasma_triquetra
|
--+ +-- Hyridella_australis **
| +--------+
| | +-- Diplodon_gratus **
+--+
| +-------- Hyridella_depressa **
| |
+--+ +-- Velesunio_angasi **
| +--+
| | +-- Velesunio_ambiguus **
+--+
| +-- Echyridella_menziesii **
+--+
+-- Castalia_stevensi **
Graf & Ó Foighil, 2000 [Publ 2649], fig. 1, 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
5 |
1 |
5 |
5 |
1 |
-- |
+----------------- Cumberlandia_monodonta
|
| +-- Unio_pictorum
| +--+
| | +-- Pyganodon_grandis
+-----------+
--+ | +-- Amblema_plicata
| +--+
| +-- Lampsilis_cardium
|
| +-------------- Velesunio_ambiguus **
| |
+--+ +-- Diplodon_chilensis **
| +--------+
| | +-- Castalia **
+--+
| +-------- Hyridella_australis__2 **
+--+
| +----- Hyridella_australis__1 **
+--+
| +-- Echyridella_menziesii **
+--+
+-- Hyridella_depressa **
Hoeh et al., 2002 [Publ 2653], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
1 |
26 |
25 |
2 |
-- |
+-- Castalia_stevensi **
+-----+
| +-- Diplodon_gratus **
+-----------------------+
| | +----- Echyridella_menziesii **
| +--+
| | +-- Velesunio_angasi **
| +--+
| +-- Lortiella **
|
--+ +-- Mutela_dubia
| +-----+
| | +-- Mutela_rostrata
| +--------------------+
| | | +----- Monocondylaea_minuana
| | +--+
| | | +-- Anodontites_guanarensis
| | +--+
+--+ +-- Anodontites_trigona
|
| +-------------------------- Coelatura_aegyptiaca
| |
| | +-- Margaritifera_margaritifera
| | +--------------------+
+--+ | +-- Cumberlandia_monodonta
| |
| | +-------------- Gonidea_angulata
| | +-----+
+--+ | | +----------- Unio_pictorum
| | +--+
| | | +-------- Unio_tumidus
| | +--+
| | | +----- Anodonta_cygnea
| | +--+
+--+ | +-- Pyganodon_grandis
| +--+
| +-- Strophitus_undulatus
|
| +-- Glebula_rotundata
| +--------------+
| | +-- Cyrtonaias_tampicoensis
| |
+--+ +-------------- Toxolasma_lividum
| |
| | +----- Potamilus_alatus
| +--------+
+--+ | +-- Ligumia_recta
| +--+
| +-- Ortmanniana_ligamentina
|
| +----------- Amblema_plicata
+--+
| +-------- Quadrula_quadrula
+--+
| +----- Eurynia_dilatata
+--+
| +-- Pleurobema_clava
+--+
+-- Fusconaia_flava
Bogan & Hoeh, 2000 [Publ 2650], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
1 |
29 |
28 |
2 |
-- |
+-- Castalia_stevensi **
+-----+
| +-- Diplodon_gratus **
+--------------------+
| | +----- Echyridella_menziesii **
| +--+
| | +-- Velesunio_angasi **
| +--+
| +-- Lortiella_rugata **
|
| +-- Mutela_dubia
--+ +-----------+
| | +-- Mutela_rostrata
| +-----------+
| | | +----------- Etheria_elliptica
| | +--+
| | | +-------- Mulleria_rivoli
| | +--+
| | | +----- Monocondylaea_minuana
| | +--+
+--+ | +-- Anodontites_trigona
| +--+
| +-- Anodontites_guanarensis
|
| +----------------------- Coelatura_aegyptiaca
| |
| | +-------------------- Pseudomulleria_dalyi
+--+ |
| | +----- Anodonta_cygnea
| | +-----------+
+--+ | | +-- Strophitus_undulatus
| | +--+
| | +-- Pyganodon_grandis
| |
+--+ +-------------- Gonidea_angulata
| |
| | +-- Unio_pictorum
| +-----------+
| | +-- Unio_tumidus
| |
+--+ +-- Cumberlandia_monodonta
| +--------+
| | +-- Margaritifera_margaritifera
| |
| | +-------- Toxolasma_lividum
| | |
+--+ +-------- Amblema_plicata
| |
| | +-- Glebula_rotundata
| +-----+
| | +-- Cyrtonaias_tampicoensis
| |
+--+ +----- Potamilus_alatus
+--+
| | +-- Ortmanniana_ligamentina
| +--+
| +-- Ligumia_recta
|
| +----- Quadrula_quadrula
| |
+--+ +-- Eurynia_dilatata
| |
+--+-- Pleurobema_clava
|
+-- Fusconaia_flava
Roe & Hoeh, 2003 [Publ 2655], fig. 4.3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
1 |
26 |
25 |
2 |
-- |
+-- Mutela_dubia
+-----+
| +-- Mutela_rostrata
+--+
| | +----- Monocondylaea_minuana
| +--+
| | +-- Anodontites_guanarensis
| +--+
+-----------------+ +-- Anodontites_trigona
| |
| | +-- Castalia_stevensi **
| | +-----+
| | | +-- Diplodon_gratus **
| +--+
| | +----- Echyridella_menziesii **
| +--+
--+ | +-- Velesunio_angasi **
| +--+
| +-- Lortiella_rugata **
|
| +-------------------------- Coelatura_aegyptiaca
| |
| | +-- Unio_pictorum
| | +-----+
| | | +-- Unio_tumidus
+--+ +--------------+
| | | +----- Anodonta_cygnea
| | +--+
| | | +-- Pyganodon_grandis
| | +--+
+--+ +-- Strophitus_undulatus
|
| +-------------------- Gonidea_angulata
| |
| | +----------------- Amblema_plicata
+--+ |
| | +-------- Quadrula_quadrula
| | +-----+
+--+ | | +----- Eurynia_dilatata
| | +--+
| | | +-- Fusconaia_flava
| | +--+
+--+ +-- Pleurobema_clava
|
| +-- Cyrtonaias_tampicoensis
| +--------+
| | +-- Glebula_rotundata
+--+
| +-------- Toxolasma_lividum
+--+
| +----- Ortmanniana_ligamentina
+--+
| +-- Ligumia_recta
+--+
+-- Potamilus_alatus
Hoeh et al., 2001 [Publ 2652], fig. 14.4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
1 |
26 |
25 |
2 |
-- |
+-- Castalia_stevensi **
+-----+
| +-- Diplodon_gratus **
+--------------------------+
| | +----- Echyridella_menziesii **
| +--+
| | +-- Lortiella_rugata **
| +--+
| +-- Velesunio_angasi **
|
--+ +-- Mutela_dubia
| +-----+
| | +-- Mutela_rostrata
| +-----------------------+
| | | +----- Monocondylaea_minuana
| | +--+
| | | +-- Anodontites_guanarensis
| | +--+
+--+ +-- Anodontites_trigona
|
| +----------------------------- Coelatura_aegyptiaca
| |
| | +-- Cumberlandia_monodonta
+--+ +-----------------------+
| | +-- Margaritifera_margaritifera
| |
| | +----------------------- Gonidea_angulata
+--+ |
| | +-------- Unio_pictorum
| | |
| | +-----------+-------- Unio_tumidus
+--+ | |
| | | +----- Anodonta_cygnea
| | +--+
| | | +-- Strophitus_undulatus
| | +--+
+--+ +-- Pyganodon_grandis
|
| +----------------- Amblema_plicata
| |
| | +-------- Quadrula_quadrula
| | +-----+
+--+ | | +----- Eurynia_dilatata
| | +--+
| | | +-- Fusconaia_flava
| | +--+
+--+ +-- Pleurobema_clava
|
| +-- Cyrtonaias_tampicoensis
| +--------+
| | +-- Glebula_rotundata
+--+
| +-------- Toxolasma_lividum
+--+
| +----- Potamilus_alatus
+--+
| +-- Ligumia_recta
+--+
+-- Ortmanniana_ligamentina
Roe & Hoeh, 2003 [Publ 2655], fig. 4.2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
2 |
1 |
26 |
25 |
2 |
-- |
+-- Mutela_dubia
+-----+
| +-- Mutela_rostrata
+--+
| | +----- Monocondylaea_minuana
| +--+
| | +-- Anodontites_guanarensis
| +--+
+-----------------------+ +-- Anodontites_trigona
| |
| | +-- Castalia_stevensi **
| | +--+
| | | +-- Velesunio_angasi **
| +-----+
| | +-- Echyridella_menziesii **
--+ | |
| +--+-- Lortiella_rugata **
| |
| +-- Diplodon_gratus **
|
| +-- Cumberlandia_monodonta
| +-----------------------------+
| | +-- Margaritifera_margaritifera
+--+
| +----------------------------- Gonidea_angulata
| |
+--+ +-------------------------- Coelatura_aegyptiaca
| |
+--+ +----------------------- Fusconaia_flava
| |
| +----------------------- Amblema_plicata
+--+
| +-------------------- Quadrula_quadrula
| |
| | +----------------- Pleurobema_clava
+--+ |
| | +-------------- Eurynia_dilatata
| | |
+--+ | +----------- Ortmanniana_ligamentina
| +--+
| | | +-------- Glebula_rotundata
| | +--+
| | | +----- Potamilus_alatus
+--+ +--+
| | +-- Ligumia_recta
| +--+
| +-- Toxolasma_lividum
|
| +----------- Cyrtonaias_tampicoensis
| |
+--+ +-- Unio_pictorum
| +-----+
| | +-- Unio_tumidus
+--+
| +----- Pyganodon_grandis
+--+
| +-- Anodonta_cygnea
+--+
+-- Strophitus_undulatus
Santos-Neto et al., 2018 [Publ 3888], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
1 |
0 |
0 |
0 |
-- |
+-- Triplodon_corrugatus **
+-----+
| +-- Prisodon_syrmatophorus **
--+
| +----- Callonaia_duprei **
+--+
| +-- Castalia_ambigua **
+--+
+-- Castalia_stevensi **
Hoeh et al., 2001 [Publ 2652], fig. 14.2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
1 |
1 |
26 |
25 |
2 |
-- |
+----- Pyganodon_grandis
+-----+
| | +-- Anodonta_cygnea
| +--+
| +-- Strophitus_undulatus
|
| +----- Diplodon_gratus **
--+ |
| +----- Echyridella_menziesii **
| +--+
| | +----- Lortiella_rugata **
| | |
| | | +-- Castalia_stevensi **
| | +--+
+--+ +-- Velesunio_angasi **
|
| +-- Mutela_dubia
| +--+
| | +-- Mutela_rostrata
+--+
| +-- Anodontites_guanarensis
| |
+--+-- Anodontites_trigona
|
+-- Monocondylaea_minuana
Guerra et al., 2019 [Publ 4168], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
1 |
36 |
35 |
2 |
-- |
+----------------- Chambardia_rubens
+--------------------+
| | +-------------- Mutela_dubia
| +--+
| | +----------- Anodontites_trapesialis
| +--+
| | +-------- Monocondylaea_parchappii
| +--+
| | +----- Lamproscapha_ensiformis
| +--+
| | +-- Tamsiella_schroeteriana
| +--+
| +-- Fossula_fossiculifera
--+
| +----------- Westralunio_carteri **
| +-----------------------+
| | | +-------- Echyridella_menziesii **
| | +--+
| | | +----- Diplodon_suavidicus **
| | +--+
| | | +-- Castalia_ambigua **
| | +--+
| | +-- Prisodon_obliquus **
| |
| | +-------------- Gibbosula_crassa
+--+ |
| +-----------------+ +----------- Cumberlandia_monodonta
| | | |
| | +--+ +-- Pseudunio_marocanus
| | | +-----+
| | | | +-- Pseudunio_auricularius
| | +--+
| | | +----- Margaritifera_falcata
| | +--+
| | | +-- Margaritifera_dahurica
+--+ +--+
| +-- Margaritifera_margaritifera
|
| +-------- Quadrula_quadrula
| +--------------------+
| | | +----- Toxolasma_parvum
| | +--+
| | | +-- Venustaconcha_ellipsiformis
| | +--+
| | +-- Potamilus_alatus
+--+
| +-------- Sinosolenaia_carinata
| +-----------------+
| | | +----- Pronodularia_japanensis
| | +--+
| | | +-- Lamprotula_leaii
| | +--+
+--+ +-- Potomida_littoralis
|
| +----------------------- Aculamprotula_tortuosa
| |
| | +-------- Schistodesmus_lampreyanus
+--+ +-----------+
| | | +----- Unio_tumidus
| | +--+
| | | +-- Unio_nanus
+--+ +--+
| +-- Unio_delphinus
|
| +----------------- Lanceolaria_lanceolata
| |
+--+ +-------------- Sinanodonta_woodiana
| |
+--+ +----------- Anodonta_anatina
| |
+--+ +-- Platynaias_compressa
| +-----+
| | +-- Platynaias_subviridis
+--+
| +----- Pyganodon_grandis
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Utterbackia_imbecillis
Guerra et al., 2019 [Publ 4168], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
1 |
36 |
35 |
2 |
-- |
+----------------- Mutela_dubia
+--------------+
| | +-------------- Chambardia_rubens
| +--+
| | +----------- Anodontites_trapesialis
| +--+
| | +-------- Monocondylaea_parchappii
| +--+
| | +----- Lamproscapha_ensiformis
| +--+
| | +-- Tamsiella_schroeteriana
| +--+
| +-- Fossula_fossiculifera
--+
| +-- Echyridella_menziesii **
| +-----+
| | +-- Westralunio_carteri **
| +--------------------+
| | | +----- Diplodon_suavidicus **
| | +--+
| | | +-- Castalia_ambigua **
| | +--+
| | +-- Prisodon_obliquus **
| |
| | +-------------- Gibbosula_crassa
| | |
+--+ +-----------+ +----------- Cumberlandia_monodonta
| | | |
| | +--+ +-- Pseudunio_marocanus
| | | +-----+
| | | | +-- Pseudunio_auricularius
| | +--+
| | | +----- Margaritifera_falcata
| | +--+
| | | +-- Margaritifera_dahurica
| | +--+
+--+ +-- Margaritifera_margaritifera
|
| +-------- Quadrula_quadrula
| +--------------+
| | | +----- Toxolasma_parvum
| | +--+
| | | +-- Venustaconcha_ellipsiformis
| | +--+
| | +-- Potamilus_alatus
| |
+--+ +-------- Sinosolenaia_carinata
| +-----------+
| | | +----- Pronodularia_japanensis
| | +--+
| | | +-- Lamprotula_leaii
| | +--+
| | +-- Potomida_littoralis
| |
+--+ +-------------- Lanceolaria_lanceolata
| +--+
| | | +----------- Aculamprotula_tortuosa
| | +--+
| | | +-------- Schistodesmus_lampreyanus
| | +--+
| | | +----- Unio_tumidus
+--+ +--+
| | +-- Unio_nanus
| +--+
| +-- Unio_delphinus
|
| +-------------- Sinanodonta_woodiana
| |
+--+ +----------- Anodonta_anatina
| |
+--+ +-- Platynaias_compressa
| +-----+
| | +-- Platynaias_subviridis
+--+
| +----- Pyganodon_grandis
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Utterbackia_imbecillis
Whelan et al., 2011 [Publ 3380], fig. 1 MP
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
1 |
32 |
31 |
2 |
-- |
+-- Margaritifera_margaritifera
+--------+
| +-- Cumberlandia_monodonta
+-----------------+
| | +-------- Velesunio_ambiguus **
| +--+
| | +----- Hyridella_australis **
| +--+
| | +-- Castalia_ambigua **
| +--+
| +-- Triplodon_corrugatus **
|
| +-- Lamellidens_savadiensis
| +--------------+
--+ | +-- Trapezidens_dolichorhynchus
| +--------+
| | | +-------------- Leoparreysia_tavoyensis
| | | |
| | +--+ +-- Radiatula_chaudhurii
| | | +--------+
| | | | +-- Indochinella_pugio
| | +--+
| | | +-------- Nitia_teretiuscula
| | | |
| | +--+ +----- Coelatura_aegyptiaca__1
+--+ | |
| +--+----- Coelatura_aegyptiaca__2
| |
| | +-- Coelatura_gabonensis
| +--+
| +-- Prisodontopsis_aviculaeformis
|
| +----------------------- Lens_contradens
| |
| | +-------------------- Pseudodon_vondembuschianus
+--+ |
| | +-- Pyganodon_grandis
| | +--+
+--+ | +-- Lasmigona_costata
| +-----------+
| | | +-- Unio_pictorum
| | +--+
| | +-- Unio_caffer
+--+
| +-------------- Gonidea_angulata
| |
| | +----- Pronodularia_japanensis
+--+ +-----+
| | | +-- Potomida_littoralis
| | +--+
+--+ +-- Potomida_semirugata
|
| +-------- Quadrula_quadrula
| |
+--+-------- Pleurobema_sintoxia
|
| +----- Amblema_plicata
+--+
| +-- Lampsilis_cardium
+--+
+-- Truncilla_truncata
Graf, 2002 [Publ 2651], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
1 |
14 |
13 |
1 |
-- |
+-------- Velesunio_ambiguus **
+--------------+
| | +----- Hyridella_depressa **
| +--+
| | +-- Diplodon_chilensis **
| +--+
--+ +-- Castalia **
|
| +-------------------- Cumberlandia_monodonta
| |
| | +-- Pilsbryoconcha_exilis
+--+ +--------------+
| | +-- Pseudodon_vondembuschianus
| |
+--+ +-------------- Lens_contradens
| |
| | +-- Unio_pictorum
| | +--+
+--+ | +-- Unio_caffer
| +-----+
| | | +-- Alasmidonta_marginata
| | +--+
+--+ +-- Pyganodon_grandis
|
| +-------- Gonidea_angulata
| |
+--+ +-- Lampsilis_cardium
| +--+
| | +-- Obliquaria_reflexa
+--+
| +-- Eurynia_dilatata
+--+
+-- Amblema_plicata
Whelan et al., 2011 [Publ 3380], fig. 1 ML
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
1 |
32 |
31 |
2 |
-- |
+-------- Velesunio_ambiguus **
+--------------------------+
| | +----- Hyridella_australis **
| +--+
| | +-- Castalia_ambigua **
| +--+
| +-- Triplodon_corrugatus **
|
| +-- Margaritifera_margaritifera
--+ +-----------------------------+
| | +-- Cumberlandia_monodonta
| |
| | +-- Aspatharia_pfeifferiana
| | +-----+
| | | +-- Chambardia_wahlbergi
| | +--------------------+
| | | | +----- Mutela_rostrata
+--+ | +--+
| | | +-- Mycetopoda_siliquosa
| | +--+
| | +-- Tamsiella_schroeteriana
| |
| | +-- Lamellidens_savadiensis
| | +-----------------+
| | | +-- Trapezidens_dolichorhynchus
+--+ +-----+
| | | +----------------- Leoparreysia_tavoyensis
| | | |
| | +--+ +-- Radiatula_chaudhurii
| | | +-----------+
| | | | +-- Indochinella_pugio
| | +--+
| | | +----------- Nitia_teretiuscula
| | +--+
| | | +-------- Coelatura_aegyptiaca__2
| | +--+
+--+ | +----- Coelatura_aegyptiaca__1
| +--+
| | +-- Coelatura_gabonensis
| +--+
| +-- Prisodontopsis_aviculaeformis
|
| +-- Pseudodon_vondembuschianus
| +--------------------+
| | +-- Lens_contradens
| |
| | +-- Pyganodon_grandis
| | +--+
+--+ | +-- Lasmigona_costata
| +--------------+
| | | +-- Unio_pictorum
| | +--+
| | +-- Unio_caffer
+--+
| +----------------- Gonidea_angulata
| |
| | +----- Pronodularia_japanensis
+--+ +--------+
| | | +-- Potomida_littoralis
| | +--+
+--+ +-- Potomida_semirugata
|
| +----------- Pleurobema_sintoxia
+--+
| +-------- Quadrula_quadrula
+--+
| +----- Amblema_plicata
+--+
| +-- Truncilla_truncata
+--+
+-- Lampsilis_cardium
Klunzinger et al., 2022 [Publ 4259], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
1 |
0 |
0 |
0 |
-- |
+-------- Cucumerunio_novaehollandiae **
--+
| +----- Velesunio_ambiguus **
+--+
| +-- Westralunio_inbisi **
+--+
+-- Westralunio_carteri **
Graf, 2000 [Publ 2647], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
1 |
13 |
12 |
2 |
-- |
+-------------- Grandidieria
|
| +-------- Castalia **
| +--+
--+ | | +----- Diplodon **
| | +--+
| | | +-- Hyridella **
| | +--+
+--+ +-- Velesunio **
|
| +-- Mutela__1
| +-----+
| | +-- Mutela__2
+--+
| +----- Anodontites
| |
+--+----- Mycetopoda
|
| +-- Etheria
+--+
+-- Mulleria
Klunzinger et al., 2021 [Publ 4086], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
1 |
2 |
2 |
1 |
-- |
+-- Margaritifera_margaritifera
+-----+
| +-- Unio_pictorum
|
--+ +-- Westralunio_inbisi **
| +--+
| | +-- Westralunio_carteri **
+--+
| +-- Velesunio_ambiguus **
+--+
+-- Cucumerunio_novaehollandiae **
Pfeiffer & Graf, 2015 [Publ 3620], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
1 |
36 |
35 |
2 |
-- |
+-------- Velesunio_ambiguus **
+--------------------------+
| | +----- Hyridella_australis **
| +--+
| | +-- Triplodon_corrugatus **
| +--+
| +-- Castalia_ambigua **
|
--+ +----- Etheria_elliptica
| +--------------------------+
| | | +-- Lamproscapha_ensiformis
| | +--+
| | +-- Chambardia_wahlbergi
| |
| | +-- Margaritifera_margaritifera
| | +--------------------------+
+--+ | +-- Cumberlandia_monodonta
| |
| | +-- Trapezidens_dolichorhynchus
| | +-----------+
| | | +-- Lamellidens_savadiensis
| | |
| | +-----------+ +-- Leoparreysia_olivacea
+--+ | | +--------+
| | | | +-- Leoparreysia_tavoyensis
| | +--+
| | | +-------- Coelatura_aegyptiaca
| | +--+
| | | +----- Indochinella_pugio
| | +--+
| | | +-- Radiatula_chaudhurii
| | +--+
+--+ +-- Scabies_mandarinus
|
| +----- Unio_pictorum
| +-----------------+
| | | +-- Pyganodon_grandis
| | +--+
| | +-- Lasmigona_costata
| |
| | +----------- Quadrula_quadrula
| | +--------+
+--+ | | +-------- Amblema_plicata
| | +--+
| | | +----- Potamilus_alatus
| | +--+
| | | +-- Lampsilis_cardium
| | +--+
| | +-- Truncilla_truncata
+--+
| +-------- Ensidens_telus
| +--------+
| | | +----- Lens_micropterus
| | +--+
| | | +-- Physunio
| | +--+
| | +-- Physunio_modelli
+--+
| +-------- Gonidea_angulata
| +-----+
| | | +----- Potomida_littoralis
| | +--+
| | | +-- Pronodularia_japanensis
+--+ +--+
| +-- Lamprotula_caveata
|
| +----------- Pilsbryoconcha_carinifera
| |
+--+ +-- Pseudodon_vondembuschianus
| +-----+
| | +-- Pseudodon_vagulus
+--+
| +----- Sundadontina_tumida
+--+
| +-- Nyeinchanconcha_nyeinchani
+--+
+-- Bineurus_exilis
Sano et al., 2020 [Publ 4138], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
66 |
64 |
2 |
-- |
+----- Triplodon_corrugatus **
+-----------------------------------------------+
| | +-- Hyridella_australis **
| +--+
| +-- Velesunio_ambiguus **
|
| +----------- Mutela_hargeri
| |
| +--------------------------------------+ +-- Aspatharia_pfeifferiana
| | | +-----+
| | | | +-- Chambardia_wahlbergi
--+ | +--+
| | | +----- Etheria_elliptica
| | +--+
| | | +-- Tamsiella_schroeteriana
| | +--+
| | +-- Lamproscapha_ensiformis
| |
| | +----------------------- Cumberlandia_monodonta
| | |
| | | +-- Pseudunio_auricularius
| | +-----------------------+ +-----------------+
+--+ | | | +-- Pseudunio_marocanus
| | | |
| | +--+ +-- Gibbosula_laosensis
| | | +--------------+
| | | | +-- Gibbosula_rochechouartii
| | | |
| | +--+ +-- Margaritifera_margaritifera
| | | +-----------+
| | | | +-- Margaritifera_dahurica
| | +--+
| | | +----------- Margaritifera_falcata
| | +--+
| | | +-------- Margaritifera_middendorffi
| | +--+
+--+ | +----- Margaritifera_laevis
| +--+
| | +-- Margaritifera_hembeli
| +--+
| +-- Margaritifera_marrianae
|
| +-------------------------------------------- Scabies_crispata
| |
| | +----- Sinohyriopsis_cumingii__2
| | +--------+
| | | | +-- Sinohyriopsis_schlegelii
| | | +--+
| | | +-- Sinohyriopsis_cumingii__1
| | +--------------------------+
| | | | +----------- Lampsilis_cardium
+--+ | | |
| | | | +----- Lamprotula_leaii
| | +--+ +--+
| | | | | +-- Potomida_littoralis
| | | | +--+
| | +--+ +-- Pronodularia_japanensis
| | |
| | | +----- Inversidens_brandtii
| | +--+
| | | +-- Obovalis_omiensis
+--+ +--+
| +-- Ptychorhynchus_pfisteri
|
| +----- Buldowskia_iwakawai
| +--------------------------------+
| | | +-- Anemina_arcaeformis
| | +--+
| | +-- Anemina_euscaphys
| |
| | +-------- Pletholophus_tenuis
| | +--+
| | | | +----- Beringiana_fukuharai
| | | +--+
+--+ | | +-- Beringiana_japonica
| +-----------------------+ +--+
| | | +-- Beringiana_beringiana
| | |
| | | +-------- Sinanodonta_lucida
| | +--+
| | | +----- Sinanodonta_tumens
| | +--+
| | | +-- Sinanodonta_woodiana
| | +--+
+--+ +-- Sinanodonta_lauta
|
| +-------------------------------- Cristaria_plicata
| |
| | +-- Anodonta_anatina
| | +--------+
| | | +-- Anodonta_cygnea
| | |
| | +-----------------+ +----- Pyganodon_grandis
+--+ | | +--+
| | | | | +-- Utterbackia_imbecillis
| | | | +--+
| | +--+ +-- Utterbackia_peninsularis
| | |
| | | +----- Alasmidonta_varicosa
| | +--+
+--+ | +-- Platynaias_compressa
| +--+
| +-- Prolasmidonta_heterodon
|
| +----- Lanceolaria_grayii__1
| +--------------------+
| | | +-- Lanceolaria_lanceolata
| | +--+
| | +-- Lanceolaria_grayii__2
+--+
| +----------------------- Lepidodesma_languilati
| |
| | +----- Aculamprotula_tortuosa
| | +--------------+
+--+ | | +-- Aculamprotula_tientsinensis
| | +--+
| | +-- Aculamprotula_coreana
| |
+--+ +----- Unio_nanus
| +-----------+
| | | +-- Unio_delphinus
| | +--+
| | +-- Unio_pictorum
| |
+--+ +-------- Inversiunio_jokohamensis
| +-----+
| | | +----- Inversiunio_yanagawensis__2
| | +--+
| | | +-- Inversiunio_reinianus
| | +--+
+--+ +-- Inversiunio_yanagawensis__1
|
| +-- Schistodesmus_lampreyanus
| +--------+
| | +-- Tchangsinaia_piscicula
+--+
| +-------- Nodularia_nipponensis__1
+--+
| +----- Nodularia_nipponensis__2
+--+
| +-- Nodularia_nipponensis__3
+--+
+-- Nodularia_douglasiae
Combosch et al., 2017 [Publ 3806], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
12 |
10 |
2 |
-- |
+-- Chambardia_wahlbergi
+--------+
| +-- Aspatharia_pfeifferiana
+--+
| | +-------- Etheria_elliptica
| +--+
| | +----- Mutela_hargeri
| +--+
| | +-- Tamsiella_schroeteriana
--+ +--+
| +-- Lamproscapha_ensiformis
|
| +----- Velesunio_ambiguus **
| +-----+
| | | +-- Triplodon_corrugatus **
| | +--+
+--+ +-- Hyridella_australis **
|
| +-------- Margaritifera_margaritifera
+--+
| +----- Unio_pictorum
+--+
| +-- Scabies_crispata
+--+
+-- Lampsilis_cardium
Sonowal et al., 2021 [Publ 4221], fig. 3a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
0 |
1 |
12 |
9 |
1 |
-- |
+-- Triplodon_corrugatus **
|
--+-- Hyridella_australis **
|
+-- Velesunio_ambiguus **
Wu et al., 2021 [Publ 4366], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
88 |
86 |
2 |
-- |
+-- Chambardia_wahlbergi
+-----+
| +-- Aspatharia_pfeifferiana
+--------------------------------+
| | +----- Etheria_elliptica
| +--+
| | +-- Tamsiella_schroeteriana
| +--+
| +-- Lamproscapha_ensiformis
|
| +----- Alathyria_jacksoni **
| +--------------------------------+
| | | +-- Diplodon_suavidicus **
| | +--+
--+ | +-- Castalia_ambigua **
| |
| | +----------- Gibbosula_laosensis
| | |
| | | +-- Cumberlandia_monodonta
| | +-----------------------+ +-----+
| | | | | +-- Pseudunio_homsensis
| | | | |
| | | +--+ +-- Margaritifera_middendorffi
| | | | +--+
| | | | | +-- Margaritifera_laevis
| | | +--+
+--+ | | +-- Margaritifera_dahurica
| | +--+
| | +-- Margaritifera_margaritifera
| |
| | +-------- Trapezidens_exolescens
| | +-----------+
| | | | +----- Lamellidens_savadiensis
| | | +--+
| | | | +-- Lamellidens_ferrugineus
| | | +--+
| | +-----------+ +-- Lamellidens_generosus
| | | |
| | | | +----- Leoparreysia_olivacea
| | | | +-----------+
| | | | | | +-- Leoparreysia_canefrii
| | | | | +--+
+--+ | +--+ +-- Leoparreysia_tavoyensis
| | |
| | | +-------------- Coelatura_aegyptiaca
| | +--+
| | | +----------- Indonaia_andersoniana
| | +--+
| | | +-------- Radiatula_myitkyinae
| | +--+
| | | +----- Scabies_crispata
| | +--+
| | | +-- Radiatula_mouhoti
| | +--+
| | +-- Radiatula_chaudhurii
| |
| | +-------------------------- Acuticosta_chinensis
| | |
| | | +----- Aculamprotula_tortuosa
| | | +-----------+
| | | | | +-- Aculamprotula_tientsinensis
| | | | +--+
| | | | +-- Aculamprotula_scripta
| | | +-----+
| | | | | +-- Unio_pictorum
+--+ +--+ | | +-----------+
| | | | | | +-- Unio_tumidus
| | | | +--+
| | | | | +----------- Nodularia_douglasiae
| | | | | |
| | | | +--+ +-- Cuneopsis_heudei
| | | | | +-----+
| | | | | | +-- Cuneopsis_rufescens
| | | | +--+
| | +--+ | +----- Tchangsinaia_piscicula
| | | +--+
| | | | +-- Schistodesmus_spinosus
| | | +--+
| | | +-- Schistodesmus_lampreyanus
| | |
| | | +----- Lanceolaria_gladiola
| | | +--------------+
| | | | | +-- Lanceolaria_grayii
| | | | +--+
| | | | +-- Lanceolaria_lanceolata
| | +--+
| | | +----------------- Lepidodesma_languilati
| | | |
| | | | +-------- Anemina_arcaeformis
| | +--+ +-----+
| | | | | +----- Cristaria_plicata
| | | | +--+
| | | | | +-- Sinanodonta_woodiana
| | +--+ +--+
+--+ | +-- Sinanodonta_lucida
| |
| | +----------- Anodonta_cygnea
| | |
| +--+ +-- Lasmigona_costata
| | +-----+
| | | +-- Anodonta_anatina
| +--+
| | +----- Pseudanodonta_complanata
| +--+
| | +-- Pyganodon_grandis
| +--+
| +-- Strophitus_undulatus
|
| +-------- Physunio_modelli
| +--+
| | | +----- Lens_eximius
| | +--+
| | | +-- Yaukthwa_nesemanni
| +--------------+ +--+
| | | +-- Yaukthwa_panhai
| | |
| | | +-------- Hyriopsis_desowitzi
| | +--+
| | | +----- Ensidens_telus
| | +--+
| | | +-- Ensidens_spiculus
| | +--+
| | +-- Ensidens_sagittarius
| |
+--+ +-- Quadrula_quadrula
| +--------+
| | +-- Elliptio_complanata
| +-----------+
| | | +-------- Obliquaria_reflexa
| | +--+
| | | +----- Potamilus_alatus
| | +--+
| | | +-- Lampsilis_cardium
| | +--+
| | +-- Truncilla_truncata
+--+
| +----- Chamberlainia_somsakpanhai
| +--------------+
| | | +-- Sinohyriopsis_cumingii
| | +--+
| | +-- Sinohyriopsis_schlegelii
| |
| | +-------- Pronodularia_japanensis
| | +--------+
+--+ | | +----- Potomida_littoralis
| | +--+
| | | +-- Lamprotula_leaii
| | +--+
| | +-- Lamprotula_caveata
| |
| | +----- Indopseudodon_indawgyiensis
| | +--+
+--+ | | +-- Indopseudodon_salwenianus
| | +--+
| +-----+ +-- Indopseudodon_bogani
| | |
| | | +----- Pilsbryoconcha_exilis
| | +--+
| | | +-- Pseudodon_vagulus
| | +--+
| | +-- Pilsbryoconcha_carinifera
+--+
| +-------- Inversidens_brandtii
| +--+
| | | +----- Gonidea_angulata
| | +--+
| | | +-- Microcondylaea_bonellii
| | +--+
+--+ +-- Leguminaia_wheatleyi
|
| +-- Sinosolenaia_oleivora
| +--+
| | +-- Sinosolenaia_carinata
+-----+
| +-- Ptychorhynchus_pfisteri
+--+
+-- Parvasolenaia_rivularis
Combosch et al., 2017 [Publ 3806], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
12 |
10 |
2 |
-- |
+----- Triplodon_corrugatus **
+--------------+
| | +-- Velesunio_ambiguus **
| +--+
| +-- Hyridella_australis **
--+
| +----- Unio_pictorum
| +-----------+
| | | +-- Lampsilis_cardium
| | +--+
+--+ +-- Scabies_crispata
|
| +-------------- Margaritifera_margaritifera
| |
+--+ +-- Chambardia_wahlbergi
| +--------+
| | +-- Aspatharia_pfeifferiana
+--+
| +-------- Etheria_elliptica
+--+
| +----- Mutela_hargeri
+--+
| +-- Tamsiella_schroeteriana
+--+
+-- Lamproscapha_ensiformis
Wu et al., 2020 [Publ 4121], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
31 |
29 |
2 |
-- |
+----- Triplodon_corrugatus **
+--------------------+
| | +-- Hyridella_australis **
| +--+
| +-- Velesunio_ambiguus **
|
| +----------- Mutela_hargeri
| |
| +-----------+ +-- Aspatharia_pfeifferiana
| | | +-----+
--+ | | | +-- Chambardia_wahlbergi
| | +--+
| | | +----- Etheria_elliptica
| | +--+
| | | +-- Tamsiella_schroeteriana
| | +--+
| | +-- Lamproscapha_ensiformis
| |
| | +-- Lampsilis_cardium
| | +--------------+
+--+ | +-- Scabies_crispata
| |
| | +-------- Sinosolenaia_carinata
| +--+ +-----+
| | | | | +----- Potomida_littoralis
| | | | +--+
| | | | | +-- Lamprotula_caveata
| | +--+ +--+
| | | +-- Lamprotula_leaii
| | |
| | | +----------- Unio_pictorum
| | +--+
| | | +-------- Aculamprotula_tortuosa
+--+ +--+
| | +----- Aculamprotula_polysticta
| +--+
| | +-- Aculamprotula_fibrosa
| +--+
| +-- Aculamprotula_tientsinensis
|
| +----------------- Cumberlandia_monodonta
| |
| | +-- Gibbosula_laosensis
| | +--+
+--+ | +-- Gibbosula_rochechouartii
| +--------+
| | | +-- Pseudunio_auricularius
| | +--+
| | +-- Pseudunio_marocanus
+--+
| +-- Margaritifera_dahurica
| +--------+
| | +-- Margaritifera_margaritifera
+--+
| +-------- Margaritifera_falcata
| |
+--+ +-- Margaritifera_hembeli
| +--+
| | +-- Margaritifera_marrianae
+--+
| +-- Margaritifera_laevis
+--+
+-- Margaritifera_middendorffi
Sano et al., 2020 [Publ 4138], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
66 |
64 |
2 |
-- |
+----- Triplodon_corrugatus **
+--------------------------------------------+
| | +-- Hyridella_australis **
| +--+
| +-- Velesunio_ambiguus **
|
| +----------- Mutela_hargeri
| |
| +-----------------------------------+ +-- Aspatharia_pfeifferiana
| | | +-----+
| | | | +-- Chambardia_wahlbergi
--+ | +--+
| | | +----- Etheria_elliptica
| | +--+
| | | +-- Tamsiella_schroeteriana
| | +--+
| | +-- Lamproscapha_ensiformis
| |
| | +-------------------- Cumberlandia_monodonta
| | |
| | | +-- Gibbosula_rochechouartii
| | | +--+
+--+ +-----------------------+ | +-- Gibbosula_laosensis
| | | +-----------+
| | | | | +-- Pseudunio_auricularius
| | | | +--+
| | | | +-- Pseudunio_marocanus
| | +--+
| | | +-- Margaritifera_dahurica
| | | +-----------+
| | | | +-- Margaritifera_margaritifera
| | +--+
| | | +----------- Margaritifera_falcata
| | +--+
| | | +-------- Margaritifera_middendorffi
| | +--+
+--+ | +----- Margaritifera_laevis
| +--+
| | +-- Margaritifera_hembeli
| +--+
| +-- Margaritifera_marrianae
|
| +----------------------------------------- Scabies_crispata
| |
| | +----------------- Lampsilis_cardium
| | |
| | | +----- Sinohyriopsis_cumingii__2
| | +--------------------+ +--------+
| | | | | | +-- Sinohyriopsis_schlegelii
| | | | | +--+
| | | +--+ +-- Sinohyriopsis_cumingii__1
+--+ | |
| | | +----------- Inversidens_brandtii
| | | |
| | +--+ +-- Ptychorhynchus_pfisteri
| | | +-----+
| | | | +-- Obovalis_omiensis
| | +--+
| | | +----- Lamprotula_leaii
| | +--+
+--+ | +-- Pronodularia_japanensis
| +--+
| +-- Potomida_littoralis
|
| +----- Buldowskia_iwakawai
| +-----------------------------+
| | | +-- Anemina_arcaeformis
| | +--+
| | +-- Anemina_euscaphys
| |
| | +-------------- Pletholophus_tenuis
| | |
| | | +----- Beringiana_fukuharai
| | +-----------------+ +-----+
+--+ | | | | +-- Beringiana_japonica
| | | | +--+
| | +--+ +-- Beringiana_beringiana
| | |
| | | +-------- Sinanodonta_lucida
| | +--+
| | | +----- Sinanodonta_tumens
| | +--+
| | | +-- Sinanodonta_lauta
+--+ +--+
| +-- Sinanodonta_woodiana
|
| +----------------------------- Cristaria_plicata
| |
| | +-- Anodonta_cygnea
| | +--------+
| | | +-- Anodonta_anatina
| | |
| | +--------------+ +----- Alasmidonta_varicosa
+--+ | | +--+
| | | | | +-- Prolasmidonta_heterodon
| | | | +--+
| | +--+ +-- Platynaias_compressa
| | |
| | | +----- Pyganodon_grandis
| | +--+
+--+ | +-- Utterbackia_imbecillis
| +--+
| +-- Utterbackia_peninsularis
|
| +----- Lanceolaria_grayii__1
| +-----------------+
| | | +-- Lanceolaria_lanceolata
| | +--+
| | +-- Lanceolaria_grayii__2
+--+
| +-------------------- Lepidodesma_languilati
| |
| | +----- Aculamprotula_tortuosa
| | +-----------+
+--+ | | +-- Aculamprotula_tientsinensis
| | +--+
| | +-- Aculamprotula_coreana
| |
+--+ +----- Unio_nanus
| +--------+
| | | +-- Unio_delphinus
| | +--+
| | +-- Unio_pictorum
| |
+--+ +-------- Inversiunio_jokohamensis
| +--+
| | | +----- Inversiunio_yanagawensis__2
| | +--+
| | | +-- Inversiunio_reinianus
| | +--+
+--+ +-- Inversiunio_yanagawensis__1
|
| +-- Tchangsinaia_piscicula
| +-----+
| | +-- Schistodesmus_lampreyanus
+--+
| +----- Nodularia_nipponensis__1
+--+
| +-- Nodularia_nipponensis__2
+--+
+-- Nodularia_douglasiae
Lopes-Lima et al., 2018 [Publ 3829], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
25 |
23 |
2 |
-- |
+----- Triplodon_corrugatus **
+--------------------+
| | +-- Velesunio_ambiguus **
| +--+
| +-- Hyridella_australis **
|
| +----------- Mutela_hargeri
| |
--+ +-----------+ +-- Aspatharia_pfeifferiana
| | | +-----+
| | | | +-- Chambardia_wahlbergi
| | +--+
| | | +----- Etheria_elliptica
| | +--+
| | | +-- Lamproscapha_ensiformis
| | +--+
+--+ +-- Tamsiella_schroeteriana
|
| +----- Potomida_littoralis
| +--------------+
| | | +-- Lampsilis_cardium
| | +--+
| | +-- Unio_pictorum
| |
+--+ +----- Gibbosula_rochechouartii
| +-----------+
| | | +-- Gibbosula_crassa
| | +--+
| | +-- Gibbosula_laosensis
| |
+--+ +-------- Cumberlandia_monodonta
| +-----+
| | | +----- Pseudunio_marocanus
| | +--+
| | | +-- Pseudunio_auricularius
| | +--+
+--+ +-- Pseudunio_homsensis
|
| +-- Margaritifera_dahurica
| +--------+
| | +-- Margaritifera_margaritifera
+--+
| +-------- Margaritifera_falcata
| |
+--+ +-- Margaritifera_middendorffi
| +--+
| | +-- Margaritifera_laevis
+--+
| +-- Margaritifera_hembeli
+--+
+-- Margaritifera_marrianae
Konopleva et al., 2017 [Publ 3807], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
28 |
26 |
2 |
-- |
+----- Diplodon_suavidicus **
+--------------------+
| | +-- Triplodon_corrugatus **
| +--+
| +-- Castalia_ambigua **
|
| +----- Tamsiella_schroeteriana
--+ +-----------------+
| | | +-- Etheria_elliptica
| | +--+
| | +-- Chambardia_wahlbergi
| |
| | +-- Margaritifera_dahurica
| | +-----------------+
+--+ | +-- Margaritifera_laevis
| |
| | +-------- Ensidens_sagittarius
| | +-----+
| | | | +----- Lens_micropterus
| | | +--+
| | | | +-- Physunio
| | +--+ +--+
+--+ | | +-- Physunio_modelli
| | |
| | | +----------- Sundadontina_tumida
| | | |
| | +--+ +-- Nyeinchanconcha_nyeinchani
| | | +-----+
| | | | +-- Bineurus_exilis
| | +--+
| | | +----- Pilsbryoconcha_carinifera
| | +--+
+--+ | +-- Pseudodon_vagulus
| +--+
| +-- Pseudodon_vondembuschianus
|
| +-- Trapezidens_exolescens
| +-----+
| | +-- Trapezidens_dolichorhynchus
| +-----+
| | | +----- Lamellidens_savadiensis
| | +--+
| | | +-- Lamellidens_jenkinsianus
| | +--+
+--+ +-- Lamellidens_mainwaringi
|
| +-- Leoparreysia_tavoyensis
| +--------+
| | +-- Leoparreysia_olivacea
+--+
| +-------- Coelatura_aegyptiaca
+--+
| +----- Indochinella_pugio
+--+
| +-- Radiatula_chaudhurii
+--+
+-- Scabies_mandarinus
Wu et al., 2021 [Publ 4366], fig. 2, I
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
88 |
86 |
2 |
-- |
+----- Alathyria_jacksoni **
+-----------------------------------+
| | +-- Castalia_ambigua **
| +--+
| +-- Diplodon_suavidicus **
|
| +-- Aspatharia_pfeifferiana
| +-----+
| | +-- Chambardia_wahlbergi
| +-----------------------------+
| | | +----- Etheria_elliptica
| | +--+
| | | +-- Tamsiella_schroeteriana
--+ | +--+
| | +-- Lamproscapha_ensiformis
| |
| | +----------- Gibbosula_laosensis
| | |
| | | +-- Pseudunio_homsensis
| | +-----------------------+ +-----+
| | | | | +-- Cumberlandia_monodonta
| | | | |
| | | +--+ +-- Margaritifera_middendorffi
| | | | +--+
| | | | | +-- Margaritifera_laevis
+--+ | +--+
| | | +-- Margaritifera_dahurica
| | +--+
| | +-- Margaritifera_margaritifera
| |
| | +-------- Trapezidens_exolescens
| | +-----------+
| | | | +----- Lamellidens_savadiensis
| | | +--+
| | | | +-- Lamellidens_generosus
| | | +--+
| | +-----------+ +-- Lamellidens_ferrugineus
| | | |
| | | | +----- Leoparreysia_olivacea
| | | | +-----------+
| | | | | | +-- Leoparreysia_canefrii
| | | | | +--+
+--+ | +--+ +-- Leoparreysia_tavoyensis
| | |
| | | +-------------- Coelatura_aegyptiaca
| | +--+
| | | +----------- Indonaia_andersoniana
| | +--+
| | | +-------- Scabies_crispata
| | +--+
| | | +----- Radiatula_myitkyinae
| | +--+
| | | +-- Radiatula_mouhoti
| | +--+
| | +-- Radiatula_chaudhurii
| |
| | +-------------------------- Acuticosta_chinensis
| | |
| | | +-------------------- Lepidodesma_languilati
| | | |
| | | | +----- Aculamprotula_tortuosa
| | | +--+ +-----------+
| | | | | | | +-- Aculamprotula_scripta
| | | | | | +--+
| | | | | | +-- Aculamprotula_tientsinensis
+--+ +--+ | +--+
| | | | | +-- Unio_pictorum
| | | | | +-----------+
| | | | | | +-- Unio_tumidus
| | | | +--+
| | | | | +----------- Nodularia_douglasiae
| | | | | |
| | | | +--+ +-- Cuneopsis_heudei
| | | | | +-----+
| | +--+ | | +-- Cuneopsis_rufescens
| | | +--+
| | | | +----- Tchangsinaia_piscicula
| | | +--+
| | | | +-- Schistodesmus_lampreyanus
| | | +--+
| | | +-- Schistodesmus_spinosus
| | |
| | | +----- Lanceolaria_gladiola
| | | +-----------+
| | | | | +-- Lanceolaria_grayii
| | | | +--+
| | | | +-- Lanceolaria_lanceolata
| | | |
| | +-----+ +-------- Anemina_arcaeformis
| | | +-----+
| | | | | +----- Cristaria_plicata
| | | | +--+
| | | | | +-- Sinanodonta_lucida
| | +--+ +--+
+--+ | +-- Sinanodonta_woodiana
| |
| | +----------- Lasmigona_costata
| | |
| +--+ +-- Anodonta_anatina
| | +-----+
| | | +-- Anodonta_cygnea
| +--+
| | +----- Pseudanodonta_complanata
| +--+
| | +-- Pyganodon_grandis
| +--+
| +-- Strophitus_undulatus
|
| +-------- Hyriopsis_desowitzi
| +--+
| | | +----- Ensidens_telus
| | +--+
| | | +-- Ensidens_spiculus
| +--------------+ +--+
| | | +-- Ensidens_sagittarius
| | |
| | | +-------- Physunio_modelli
| | +--+
| | | +----- Lens_eximius
| | +--+
| | | +-- Yaukthwa_nesemanni
| | +--+
| | +-- Yaukthwa_panhai
+--+
| +-------------- Quadrula_quadrula
| +--------+
| | | +----------- Elliptio_complanata
| | +--+
| | | +-------- Obliquaria_reflexa
| | +--+
| | | +----- Potamilus_alatus
| | +--+
| | | +-- Lampsilis_cardium
| | +--+
+--+ +-- Truncilla_truncata
|
| +----- Chamberlainia_somsakpanhai
| +--------------+
| | | +-- Sinohyriopsis_cumingii
| | +--+
| | +-- Sinohyriopsis_schlegelii
| |
| | +-------- Potomida_littoralis
| | +--------+
+--+ | | +----- Pronodularia_japanensis
| | +--+
| | | +-- Lamprotula_caveata
| | +--+
| | +-- Lamprotula_leaii
| |
| | +----- Indopseudodon_indawgyiensis
+--+ +--+
| | | +-- Indopseudodon_bogani
| | +--+
| +-----+ +-- Indopseudodon_salwenianus
| | |
| | | +----- Pilsbryoconcha_exilis
| | +--+
| | | +-- Pseudodon_vagulus
| | +--+
+--+ +-- Pilsbryoconcha_carinifera
|
| +----- Gonidea_angulata
| +-----+
| | | +-- Leguminaia_wheatleyi
| | +--+
+--+ +-- Microcondylaea_bonellii
|
| +-------- Inversidens_brandtii
| |
+--+ +-- Sinosolenaia_carinata
| +--+
| | +-- Sinosolenaia_oleivora
+--+
| +-- Parvasolenaia_rivularis
+--+
+-- Ptychorhynchus_pfisteri
Sano et al., 2020 [Publ 4138], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
66 |
64 |
2 |
-- |
+----- Triplodon_corrugatus **
+-----------------------------------------------+
| | +-- Hyridella_australis **
| +--+
| +-- Velesunio_ambiguus **
|
| +----------- Mutela_hargeri
| |
| +--------------------------------------+ +-- Aspatharia_pfeifferiana
| | | +-----+
| | | | +-- Chambardia_wahlbergi
--+ | +--+
| | | +----- Etheria_elliptica
| | +--+
| | | +-- Tamsiella_schroeteriana
| | +--+
| | +-- Lamproscapha_ensiformis
| |
| | +-------------------- Cumberlandia_monodonta
| | |
| | | +-- Pseudunio_auricularius
| | +--------------------------+ +--------------+
+--+ | | | +-- Pseudunio_marocanus
| | | |
| | +--+ +-- Gibbosula_rochechouartii
| | | +-----------+
| | | | +-- Gibbosula_laosensis
| | | |
| | +--+ +-- Margaritifera_margaritifera
| | | +--------+
| | | | +-- Margaritifera_dahurica
| | +--+
| | | +-------- Margaritifera_falcata
| | | |
| | +--+ +-- Margaritifera_laevis
| | | +--+
+--+ | | +-- Margaritifera_middendorffi
| +--+
| | +-- Margaritifera_hembeli
| +--+
| +-- Margaritifera_marrianae
|
| +-------------------------------------------- Scabies_crispata
| |
| | +----------------- Lampsilis_cardium
| | |
| | | +----- Sinohyriopsis_cumingii__2
| | +-----------------------+ +--------+
| | | | | | +-- Sinohyriopsis_schlegelii
| | | | | +--+
| | | +--+ +-- Sinohyriopsis_cumingii__1
+--+ | |
| | | +----------- Inversidens_brandtii
| | | |
| | +--+ +-- Obovalis_omiensis
| | | +-----+
| | | | +-- Ptychorhynchus_pfisteri
| | +--+
| | | +----- Lamprotula_leaii
| | +--+
+--+ | +-- Pronodularia_japanensis
| +--+
| +-- Potomida_littoralis
|
| +----- Buldowskia_iwakawai
| +--------------------------------+
| | | +-- Anemina_arcaeformis
| | +--+
| | +-- Anemina_euscaphys
| |
| | +-------- Sinanodonta_lucida
| | +--+
| | | | +----- Sinanodonta_tumens
| | | +--+
+--+ | | +-- Sinanodonta_woodiana
| +-----------------------+ +--+
| | | +-- Sinanodonta_lauta
| | |
| | | +-------- Pletholophus_tenuis
| | +--+
| | | +----- Beringiana_fukuharai
| | +--+
| | | +-- Beringiana_japonica
| | +--+
+--+ +-- Beringiana_beringiana
|
| +-------------------------------- Cristaria_plicata
| |
| | +-- Anodonta_cygnea
| | +--------+
| | | +-- Anodonta_anatina
| | |
| | +-----------------+ +----- Alasmidonta_varicosa
+--+ | | +--+
| | | | | +-- Prolasmidonta_heterodon
| | | | +--+
| | +--+ +-- Platynaias_compressa
| | |
| | | +----- Pyganodon_grandis
| | +--+
+--+ | +-- Utterbackia_imbecillis
| +--+
| +-- Utterbackia_peninsularis
|
| +-------------------------- Lepidodesma_languilati
| |
| | +----- Lanceolaria_grayii__1
| | +-----------------+
+--+ | | +-- Lanceolaria_lanceolata
| | +--+
| | +-- Lanceolaria_grayii__2
| |
+--+ +----- Aculamprotula_tortuosa
| +--------------+
| | | +-- Aculamprotula_tientsinensis
| | +--+
| | +-- Aculamprotula_coreana
| |
+--+ +----- Unio_nanus
| +-----------+
| | | +-- Unio_delphinus
| | +--+
| | +-- Unio_pictorum
| |
+--+ +-------- Nodularia_nipponensis__3
| +-----+
| | | +----- Nodularia_nipponensis__1
| | +--+
| | | +-- Nodularia_douglasiae
| | +--+
+--+ +-- Nodularia_nipponensis__2
|
| +-- Schistodesmus_lampreyanus
| +--------+
| | +-- Tchangsinaia_piscicula
+--+
| +-------- Inversiunio_jokohamensis
+--+
| +----- Inversiunio_yanagawensis__2
+--+
| +-- Inversiunio_reinianus
+--+
+-- Inversiunio_yanagawensis__1
Sonowal et al., 2021 [Publ 4221], fig. 3b
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
0 |
1 |
12 |
9 |
1 |
-- |
+-- Triplodon_corrugatus **
|
+--+-- Velesunio_ambiguus **
| |
| +-- Hyridella_australis **
|
| +-- Aculamprotula_tientsinensis
| |
--+ +-- Lamprotula_leaii
| |
| +-- Lampsilis_cardium
| |
| +-- Potomida_littoralis
| |
+--+-- Unio_pictorum
|
+-- Anodonta_cygnea
|
+-- Unio_ravoisieri
|
+-- Lamellidens_mainwaringi
|
+-- Elliptio_complanata
Sonowal et al., 2021 [Publ 4221], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
13 |
10 |
1 |
-- |
+----- Triplodon_corrugatus **
+-----------------------+
| | +-- Velesunio_ambiguus **
| +--+
--+ +-- Hyridella_australis **
|
| +-------------------------- Lamellidens_mainwaringi
+--+
| +----------------------- Lamprotula_leaii
+--+
| +-------------------- Potomida_littoralis
+--+
| +----------------- Lampsilis_cardium
+--+
| +-------------- Elliptio_complanata
+--+
| +----------- Aculamprotula_tientsinensis
+--+
| +-------- Unio_pictorum
+--+
| +----- Anodonta_cygnea
+--+
| +-- Unio_ravoisieri
+--+
+-- Lamellidens_corrianus
Lopes-Lima et al., 2018 [Publ 3829], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
25 |
23 |
2 |
-- |
+----- Triplodon_corrugatus **
+-----------------------+
| | +-- Velesunio_ambiguus **
| +--+
| +-- Hyridella_australis **
|
| +----------- Mutela_hargeri
| |
--+ +--------------+ +-- Chambardia_wahlbergi
| | | +-----+
| | | | +-- Aspatharia_pfeifferiana
| | +--+
| | | +----- Etheria_elliptica
| | +--+
| | | +-- Tamsiella_schroeteriana
| | +--+
+--+ +-- Lamproscapha_ensiformis
|
| +----- Potomida_littoralis
| +-----------------+
| | | +-- Lampsilis_cardium
| | +--+
| | +-- Unio_pictorum
| |
+--+ +----- Gibbosula_rochechouartii
| +--------------+
| | | +-- Gibbosula_laosensis
| | +--+
| | +-- Gibbosula_crassa
| |
+--+ +-------- Cumberlandia_monodonta
| +--------+
| | | +----- Pseudunio_marocanus
| | +--+
| | | +-- Pseudunio_auricularius
| | +--+
+--+ +-- Pseudunio_homsensis
|
| +-- Margaritifera_margaritifera
| +-----------+
| | +-- Margaritifera_dahurica
+--+
| +----------- Margaritifera_falcata
+--+
| +-------- Margaritifera_laevis
+--+
| +----- Margaritifera_middendorffi
+--+
| +-- Margaritifera_hembeli
+--+
+-- Margaritifera_marrianae
Olivera-Hyde et al., 2020 [Publ 4249], fig. 10
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
0 |
0 |
0 |
-- |
+----- Callonaia_duprei **
--+
| +-- Castalia_ambigua **
+--+
+-- Castalia_stevensi **
Huang et al., 2018 [Publ 3828], fig. 3, 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
26 |
24 |
2 |
-- |
+----- Triplodon_corrugatus **
+-----------------------+
| | +-- Velesunio_ambiguus **
| +--+
| +-- Hyridella_australis **
|
| +----------- Mutela_hargeri
| |
| +--------------+ +-- Aspatharia_pfeifferiana
--+ | | +-----+
| | | | +-- Chambardia_wahlbergi
| | +--+
| | | +----- Etheria_elliptica
| | +--+
| | | +-- Tamsiella_schroeteriana
| | +--+
| | +-- Lamproscapha_ensiformis
| |
+--+ +-- Potomida_littoralis
| +-----+
| | +-- Lamprotula_leaii
| |
| +--------------+ +-- Lampsilis_cardium
| | | +--+
| | | | +-- Scabies_crispata
| | +--+
| | | +-- Unio_pictorum
| | +--+
+--+ +-- Aculamprotula_tientsinensis
|
| +-- Gibbosula_rochechouartii
| +-----------------+
| | +-- Gibbosula_laosensis
| |
| | +----- Cumberlandia_monodonta
+--+ +-----------+
| | | +-- Pseudunio_auricularius
| | +--+
| | +-- Pseudunio_marocanus
+--+
| +-- Margaritifera_margaritifera
| +-----------+
| | +-- Margaritifera_dahurica
+--+
| +----------- Margaritifera_falcata
+--+
| +-------- Margaritifera_laevis
+--+
| +----- Margaritifera_hembeli
+--+
| +-- Margaritifera_middendorffi
+--+
+-- Margaritifera_marrianae
Wu et al., 2020 [Publ 4121], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
1 |
31 |
29 |
2 |
-- |
+----- Triplodon_corrugatus **
+-----------------------+
| | +-- Hyridella_australis **
| +--+
| +-- Velesunio_ambiguus **
|
| +----------- Mutela_hargeri
| |
| +--------------+ +-- Aspatharia_pfeifferiana
| | | +-----+
--+ | | | +-- Chambardia_wahlbergi
| | +--+
| | | +----- Etheria_elliptica
| | +--+
| | | +-- Tamsiella_schroeteriana
| | +--+
| | +-- Lamproscapha_ensiformis
| |
| | +-- Lampsilis_cardium
| | +--------------+
+--+ | +-- Scabies_crispata
| |
| | +-------- Sinosolenaia_carinata
| +-----+ +-----+
| | | | | +----- Potomida_littoralis
| | | | +--+
| | | | | +-- Lamprotula_caveata
| | +--+ +--+
| | | +-- Lamprotula_leaii
| | |
| | | +----------- Unio_pictorum
| | +--+
+--+ | +-------- Aculamprotula_tortuosa
| +--+
| | +----- Aculamprotula_polysticta
| +--+
| | +-- Aculamprotula_tientsinensis
| +--+
| +-- Aculamprotula_fibrosa
|
| +-------------------- Gibbosula_rochechouartii
| |
+--+ +----------------- Gibbosula_laosensis
| |
| | +----- Cumberlandia_monodonta
+--+ +--------+
| | | +-- Pseudunio_auricularius
| | +--+
| | +-- Pseudunio_marocanus
+--+
| +-- Margaritifera_dahurica
| +--------+
| | +-- Margaritifera_margaritifera
+--+
| +-------- Margaritifera_falcata
| |
+--+ +-- Margaritifera_hembeli
| +--+
| | +-- Margaritifera_marrianae
+--+
| +-- Margaritifera_laevis
+--+
+-- Margaritifera_middendorffi
Guerra et al., 2019 [Publ 4168], fig. 2-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
20 |
20 |
1 |
-- |
+-- Echyridella_menziesii **
+--------------------------+
| +-- Westralunio_carteri **
|
| +----- Pseudunio_marocanus
| +--------------------+
--+ | | +-- Margaritifera_margaritifera
| | +--+
| | +-- Cumberlandia_monodonta
| |
| | +----- Quadrula_quadrula
+--+ +-----------------+
| | | +-- Venustaconcha_ellipsiformis
| | +--+
| | +-- Potamilus_alatus
| |
| | +-------- Sinosolenaia_carinata
+--+ +-----------+
| | | +----- Potomida_littoralis
| | +--+
| | | +-- Lamprotula_leaii
| | +--+
| | +-- Pronodularia_japanensis
+--+
| +-- Utterbackia_peninsularis
| +--+
| | +-- Pyganodon_grandis
| +-----------+
| | | +-- Anodonta_anatina
| | +--+
+--+ +-- Sinanodonta_woodiana
|
| +-------------- Lanceolaria_lanceolata
+--+
| +----------- Aculamprotula_tortuosa
+--+
| +-------- Schistodesmus_lampreyanus
+--+
| +----- Unio_tumidus
+--+
| +-- Unio_nanus
+--+
+-- Unio_delphinus
Guerra et al., 2019 [Publ 4168], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
20 |
20 |
1 |
-- |
+-- Echyridella_menziesii **
+--------------------------+
| +-- Westralunio_carteri **
|
| +----- Cumberlandia_monodonta
| +--------------------+
--+ | | +-- Margaritifera_margaritifera
| | +--+
| | +-- Pseudunio_marocanus
| |
| | +----- Quadrula_quadrula
+--+ +-----------------+
| | | +-- Venustaconcha_ellipsiformis
| | +--+
| | +-- Potamilus_alatus
| |
| | +-------- Sinosolenaia_carinata
+--+ +-----------+
| | | +----- Potomida_littoralis
| | +--+
| | | +-- Lamprotula_leaii
| | +--+
| | +-- Pronodularia_japanensis
+--+
| +-------- Sinanodonta_woodiana
| +--------+
| | | +----- Anodonta_anatina
| | +--+
| | | +-- Utterbackia_peninsularis
+--+ +--+
| +-- Pyganodon_grandis
|
| +-------------- Lanceolaria_lanceolata
+--+
| +----------- Schistodesmus_lampreyanus
+--+
| +-------- Aculamprotula_tortuosa
+--+
| +----- Unio_tumidus
+--+
| +-- Unio_nanus
+--+
+-- Unio_delphinus
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
118 |
116 |
2 |
-- |
+-- Hyridella_depressa **
+--------------------------------------------+
| +-- Velesunio_ambiguus **
|
| +-- Margaritifera_laevis
| +-----------------------------------------+
| | +-- Margaritifera_middendorffi
| |
| | +-- Lamproscapha_ensiformis
--+ | +-----+
| | | +-- Mycetopoda_siliquosa
| | +--------------------------------+
| | | | +----- Etheria_elliptica
| | | +--+
| | | | +-- Aspatharia_pfeifferiana
| | | +--+
| | | +-- Chambardia_wahlbergi
| | |
+--+ | +-- Coelatura_aegyptiaca
| | |
| | +-----------------------------------+-- Leoparreysia_tavoyensis
| | | |
| | | +-- Radiatula_mouhoti
| | |
| | | +-- Lens_contradens
| | | |
| | | +-- Ensidens_sagittarius
| | | |
| | | +-----------------------+-- Physunio_modelli
| | | | |
| | | | +-- Rectidens_sumatrensis
| | | | |
+--+ | | +-- Yaukthwa_nesemanni
| | |
| | | +-- Amblema_plicata
| | | |
| | | +-- Elliptio_complanata
| | | +--------------------+
| | +--------+ | +-- Lampsilis_cardium
| | | | | |
| | | | | +-- Quadrula_quadrula
| | | | |
| | | | | +----- Chamberlainia_somsakpanhai
| | | | | +-----------+
| | | | | | | +-- Sinohyriopsis_cumingii
| | | | | | +--+
| | | | | +--+ +-- Sinohyriopsis_schlegelii
| | | | | | |
| | | | | | | +-------------- Pronodularia_japanensis
| | | +--+ | | |
| | | | | +--+ +----------- Potomida_littoralis
| | | | | | |
| | | | | +--+ +-------- Lamprotula_caveata
| | | | | | |
+--+ | | | +--+ +----- Lamprotula_leaii__2
| | | | | |
| | | | +--+----- Lamprotula_leaii__3
| | | | |
| | | | | +-- Lamprotula_seomjinensis
| | | | +--+
| | | | +-- Lamprotula_leaii__1
| | +--+
| | | +-- Indopseudodon_indawgyiensis
| | | +--------+
| | | | +-- Indopseudodon_salwenianus
| | | |
| | | +-----+ +-- Pilsbryoconcha_exilis
| | | | | +-----+
| | | | | | +-- Pilsbryoconcha_carinifera
| | | | +--+
| | | | | +----- Pseudodon_mekongi
| | | | | |
| | | | +--+ +-- Nyeinchanconcha_nyeinchani
| | | | | |
| | | | +--+-- Bineurus_mouhotii
| | | | |
| | +--+ +-- Sundadontina_tumida
| | |
| | | +-------- Inversidens_brandtii
| | | |
| | | +-----+ +-- Obovalis_omiensis
| | | | | +--+
| | | | | | +-- Ptychorhynchus_pfisteri
| | | | +--+
| | | | | +-- Parvasolenaia_rivularis
| | | | +--+
| | +--+ +-- Koreosolenaia_sitgyensis
| | |
+--+ | +----- Gonidea_angulata
| | +-----+
| | | | +-- Leguminaia_wheatleyi
| | | +--+
| +--+ +-- Microcondylaea_bonellii
| |
| | +-------- Sinosolenaia_carinata
| +--+
| | +----- Sinosolenaia_recognita
| +--+
| | +-- Sinosolenaia_iridinea
| +--+
| +-- Sinosolenaia_oleivora
|
| +-------------------------------- Acuticosta_chinensis
| |
| +-------------------------------- Lepidodesma_languilati
| |
| | +----------- Aculamprotula_tortuosa
| | +--------------+
| | | | +-------- Aculamprotula_scripta
| | | +--+
| | | | +----- Aculamprotula_coreana
| | | +--+
| | | | +-- Aculamprotula_fibrosa
| | | +--+
| +-----+ +-- Aculamprotula_tientsinensis
| | |
| | | +----------------------- Middendorffinaia_mongolica
| | | |
| | | | +----- Unio_gibbus
| | | | |
| | | | +--------------+----- Unio_tumidus
| | +--+ | |
| | | | | +-- Unio_crassus
| | | | +--+
| | | | +-- Unio_pictorum
| | | |
| | | | +-------------- Nodularia_breviconcha
| | +--+ +--+
| | | | | +----------- Nodularia_huana
| | | | +--+
| | | | | +-------- Nodularia_nuxpersicae
| | | | +--+
| | | | | +----- Nodularia_nipponensis
| | | | +--+
| | | | | +-- Nodularia_douglasiae
| | +--+ +--+
| | | +-- Nodularia_dualobtusa
| | |
+--+ | +-- Schistodesmus_lampreyanus
| | +-----------+
| | | +-- Schistodesmus_spinosus
| | |
| | | +----- Cuneopsis_rufescens
| | +--------+
| +--+ | +-- Cuneopsis_heudei
| | +--+
| | +-- Cuneopsis_celtiformis
| |
| | +----------- Tchangsinaia_piscicula
| +--+
| | +-------- Inversiunio_jokohamensis
| +--+
| | +----- Inversiunio_verrucosus
| +--+
| | +-- Inversiunio_reinianus
| +--+
| +-- Inversiunio_yanagawensis
|
| +----------------- Lanceolaria_oxyrhyncha
| |
| +-----------+ +-------------- Lanceolaria_gladiola
| | | |
| | +--+ +-- Lanceolaria_bilirata
| | | +--------+
| | | | +-- Lanceolaria_grayii
| | +--+
| | | +-------- Lanceolaria_acrorrhyncha
| | +--+
| | | +----- Lanceolaria_kihirai
| | +--+
| | | +-- Lanceolaria_lanceolata
| | +--+
| | +-- Lanceolaria_triformis
| |
| | +-------- Pseudanodonta_complanata
+--+ |
| | +----- Anodonta_cygnea
| +--+
| | | +-- Anodonta_anatina
| +-----------------+ +--+
| | | +-- Anodonta_nuttalliana
| | |
| | | +----- Pyganodon_grandis
| | | |
| | | +----- Simpsonaias_ambigua
| | +--+
| | +----- Strophitus_undulatus
| | |
| | | +-- Alasmidonta_marginata
| | +--+
| | +-- Lasmigona_costata
+--+
| +-- Cristaria_plicata
| +--------------------+
| | +-- Cristaria_clessini
| |
| | +-------------- Anemina_arcaeformis
| | |
| +--------+ +----------- Amuranodonta_kijaensis
| | | |
| | +--+ +-------- Buldowskia_suifunica
| | | |
| | +--+ +-- Buldowskia_flavotincta
+--+ | +--+
| | | +-- Buldowskia_shadini
| +--+
| | +-- Buldowskia_iwakawai
| +--+
| +-- Buldowskia_kamiyai
|
| +-------------------- Pletholophus_tenuis
| |
| | +-------- Beringiana_japonica
| | +--------+
+--+ | | +----- Beringiana_beringiana
| | +--+
| | | +-- Beringiana_fukuharai
| | +--+
+--+ +-- Beringiana_gosannensis
|
| +-- Sinanodonta_angula
| +-----------+
| | +-- Sinanodonta_lucida
| |
+--+ +----------- Sinanodonta_schrenkii
| |
| | +-- Sinanodonta_calipygos
| +--------+
+--+ +-- Sinanodonta_tumens
|
| +-------- Sinanodonta_woodiana
| |
+--+ +----- Sinanodonta_elliptica
| |
+--+----- Sinanodonta_jourdyi
|
| +-- Sinanodonta_lauta
+--+
+-- Sinanodonta_pacifica
Walker et al., 2006 [Publ 2666], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
14 |
13 |
1 |
-- |
+-------------------------- Hyridella **
|
| +----- Anodonta
--+ +-----------------+
| | | +-- Pyganodon_fragilis
| | +--+
| | +-- Pyganodon_grandis
+--+
| +-------------------- Pronodularia_japanensis
| |
| | +-- Gonidea_angulata
+--+ +--------------+
| | +-- Pseudodon_vondembuschianus
| |
+--+ +-- Fusconaia_flava
| +-----------+
| | +-- Cumberlandia_monodonta
+--+
| +----------- Cyrtonaias_tampicoensis
+--+
| +-------- Potamilus_purpuratus
+--+
| +----- Ligumia_recta
+--+
| +-- Ortmanniana_ligamentina
+--+
+-- Lampsilis_teres
Wu et al., 2019 [Publ 3864], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
46 |
46 |
2 |
-- |
+-- Echyridella_menziesii **
+--------+
| +-- Mutela_dubia
--+
| +-------- Gibbosula_rochechouartii
+--+
| +----- Cumberlandia_monodonta
+--+
| +-- Margaritifera_dahurica
+--+
+-- Margaritifera_falcata
Plazzi et al., 2011 [Publ 3676], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
17 |
17 |
1 |
-- |
+----- Alathyria_jacksoni **
|
+-----+----- Pronodularia_japanensis
| |
| | +-- Sinohyriopsis_cumingii
| +--+
--+ +-- Sinohyriopsis_schlegelii
|
| +-------- Quadrula_quadrula
| |
+--+-------- Parvaspina_collina
|
| +----- Lampsilis_ornata
+--+
| +-- Epioblasma_rangiana
+--+
+-- Venustaconcha_ellipsiformis
Chase et al., 2018 [Publ 3861], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
10 |
10 |
1 |
-- |
+-- Cumberlandia_monodonta
+-----------+
| +-- Echyridella_menziesii **
|
| +-------- Lamprotula_leaii
--+ +--+
| | | +----- Quadrula_quadrula
| | +--+
| | | +-- Lampsilis_powellii
| | +--+
+--+ +-- Lampsilis_siliquoidea
|
| +-- Unio_delphinus
| +-----+
| | +-- Unio_tumidus
+--+
| +----- Anodonta_anatina
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Pyganodon_grandis
Wu et al., 2022 [Publ 4238], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
73 |
73 |
2 |
-- |
+-- Mutela_dubia
+--------------------------------------------+
| +-- Anodontites_trapesialis
|
| +-------------------------------------------- Echyridella_menziesii **
| |
| | +-------------- Gibbosula_rochechouartii
| | +--------------------------+
--+ | | | +----------- Cumberlandia_monodonta
| | | +--+
| | | | +-------- Pseudunio_marocanus
| | | +--+
| | | | +----- Margaritifera_falcata
| | | +--+
| | | | +-- Margaritifera_dahurica
+--+ | +--+
| | +-- Margaritifera_margaritifera
| |
| | +-- Uniomerus_tetralasmus
| | +--------------------------+
| | | +-- Quadrula_quadrula
| | |
| | +--------+ +-- Pleurobema_oviforme
| | | | +-----------------------+
| | | | | +-- Elliptio_complanata
| | | +--+
+--+ | | +----------------------- Amblema_plicata
| | | |
| | +--+ +-------------------- Popenaias_popeii
| | | |
| | +--+ +----------------- Toxolasma_parvum
| | | |
| | +--+ +-- Potamilus_alatus
| | | +-----------+
| | | | +-- Potamilus_leptodon
| | +--+
| | | +----------- Leaunio_lienosus
| | | |
| | +--+ +-- Lampsilis_cardium
| | | +-----+
| | | | +-- Lampsilis_ornata
| | +--+
| | | +----- Venustaconcha_ellipsiformis
| | +--+
| | | +-- Lampsilis_siliquoidea
| | +--+
+--+ +-- Lampsilis_powellii
|
| +----- Chamberlainia_somsakpanhai
| +-----------+
| | | +-- Sinohyriopsis_schlegelii
| | +--+
| | +-- Sinohyriopsis_cumingii
| |
| +-----------------+ +-------- Potomida_littoralis
| | | +-----+
| | | | | +----- Pronodularia_japanensis
| | | | +--+
| | | | | +-- Lamprotula_caveata
| | | | +--+
| | +--+ +-- Lamprotula_leaii
| | |
| | | +-- Pilsbryoconcha_exilis
| | | +--------+
| | | | +-- Pseudodon_vondembuschianus
| | | |
| | +--+ +-- Ptychorhynchus_pfisteri
| | | +-----+
| | | | +-- Parvasolenaia_rivularis
| | +--+
| | | +----- Microcondylaea_bonellii
+--+ +--+
| | +-- Sinosolenaia_carinata
| +--+
| +-- Sinosolenaia_recognita
|
| +-------------------------------- Lepidodesma_languilati
| |
| | +-------- Acuticosta_chinensis
| | +--------+
| | | | +----- Lanceolaria_gladiola
| | | +--+
| | | | +-- Lanceolaria_grayii
| | | +--+
| | | +-- Lanceolaria_lanceolata
| | |
| | +-----------+ +-------- Cristaria_plicata
| | | | |
+--+ | | +-----+ +-- Sinanodonta_woodiana
| | | | | +--+
| | | | | | +-- Sinanodonta_lucida
| | | | +--+
| | | | | +-- Anemina_arcaeformis
| | +--+ +--+
| | | +-- Anemina_euscaphys
| | |
| | | +-- Anodonta_cygnea
| | | +--------+
| | | | +-- Anodonta_anatina
| | +--+
| | | +-------- Platynaias_compressa
+--+ +--+
| | +----- Pyganodon_grandis
| +--+
| | +-- Utterbackia_imbecillis
| +--+
| +-- Utterbackia_peninsularis
|
| +-------- Aculamprotula_tortuosa
| |
| +-----------------+ +-- Aculamprotula_tientsinensis
| | | +--+
| | | | +-- Aculamprotula_coreana
| | +--+
| | | +-- Aculamprotula_polysticta
| | +--+
| | +-- Aculamprotula_scripta
+--+
| +-------- Unio_tumidus
| +--------------+
| | | +----- Unio_nanus
| | +--+
| | | +-- Unio_pictorum
| | +--+
+--+ +-- Unio_delphinus
|
| +-- Pseudocuneopsis_capitata
| +-----------------+
| | +-- Pseudocuneopsis_sichuanensis
| |
+--+ +-- Nodularia_douglasiae
| +--------------+
| | +-- Nodularia_breviconcha
| |
+--+ +-- Schistodesmus_lampreyanus
| +-----------+
| | +-- Schistodesmus_spinosus
+--+
| +----------- Tchangsinaia_piscicula
+--+
| +-------- Cuneopsis_rufescens
+--+
| +----- Cuneopsis_celtiformis
+--+
| +-- Cuneopsis_heudei
+--+
+-- Cuneopsis_demangei
Elderkin et al., 2016 [Publ 3693], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
2 |
2 |
1 |
-- |
+-- Etheria_elliptica
|
--+-- Aspatharia_pfeifferiana
|
+-- Velesunio_ambiguus **
González & Giribet, 2015 [Publ 3632], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
5 |
3 |
2 |
-- |
+----- Unio_pictorum
|
--+----- Velesunio_ambiguus **
|
| +-- Aspatharia_pfeifferiana
+--+
+-- Margaritifera_margaritifera
Huang et al., 2019 [Publ 3843], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
56 |
55 |
2 |
-- |
+-- Anodontites_trapesialis
+--------------------------------------+
| +-- Mutela_dubia
|
| +-------------------------------------- Echyridella_menziesii **
| |
| | +-------------- Gibbosula_rochechouartii
| | +--------------------+
--+ | | | +----------- Cumberlandia_monodonta
| | | +--+
| | | | +-------- Pseudunio_marocanus
| | | +--+
| | | | +----- Margaritifera_falcata
| | | +--+
| | | | +-- Margaritifera_margaritifera
| | | +--+
+--+ | +-- Margaritifera_dahurica
| |
| | +-- Sinohyriopsis_cumingii
| | +-----------+
| | | +-- Sinohyriopsis_schlegelii
| | |
| | | +-------- Potomida_littoralis
| | +--------+ +--+
| | | | | | +----- Pronodularia_japanensis
| | | | | +--+
| | | | | | +-- Lamprotula_caveata
| | | | | +--+
+--+ | +--+ +-- Lamprotula_leaii
| | |
| | | +-- Ptychorhynchus_pfisteri
| | | +--+
| +--------+ | | +-- Parvasolenaia_rivularis
| | | +-----+
| | | | +-- Sinosolenaia_oleivora
| | | +--+
| | | +-- Sinosolenaia_carinata
| | |
| | | +-- Quadrula_quadrula
| | | +-----------------+
| | | | +-- Uniomerus_tetralasmus
| | +--+
| | | +----------------- Elliptio_complanata
| | | |
| | +--+ +-------------- Toxolasma_parvum
| | | |
| | +--+ +-- Potamilus_leptodon
| | | +--------+
| | | | +-- Potamilus_alatus
+--+ +--+
| | +-------- Leaunio_lienosus
| +--+
| | +----- Venustaconcha_ellipsiformis
| +--+
| | +-- Lampsilis_cardium
| +--+
| +-- Lampsilis_ornata
|
| +----------------------------- Lepidodesma_languilati
| |
| | +-------- Aculamprotula_tortuosa
| | +-----+
| | | | +----- Aculamprotula_scripta
| | | +--+
| | | | +-- Aculamprotula_tientsinensis
| | | +--+
| | +-----------+ +-- Aculamprotula_coreana
+--+ | |
| | | +----- Nodularia_douglasiae
| | | +-----+
| | | | | +-- Tchangsinaia_piscicula
| | | | +--+
| | +--+ +-- Schistodesmus_lampreyanus
| | |
| | | +-------- Unio_tumidus
| | +--+
+--+ | +----- Unio_nanus
| +--+
| | +-- Unio_delphinus
| +--+
| +-- Unio_pictorum
|
| +----- Lanceolaria_gladiola
| +-----------------+
| | | +-- Lanceolaria_lanceolata
| | +--+
| | +-- Lanceolaria_grayii
+--+
| +-------------------- Acuticosta_chinensis
| |
| | +-------- Cristaria_plicata
| | |
+--+ +--------+ +-- Sinanodonta_lucida
| | | +--+
| | | | +-- Sinanodonta_woodiana
| | +--+
+--+ | +-- Anemina_arcaeformis
| +--+
| +-- Anemina_euscaphys
|
| +-------------- Anodonta_cygnea
+--+
| +----------- Anodonta_anatina
+--+
| +-------- Lasmigona_costata
+--+
| +----- Pyganodon_grandis
+--+
| +-- Utterbackia_imbecillis
+--+
+-- Utterbackia_peninsularis
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
29 |
28 |
2 |
-- |
+----------------------------------------- Mutela_rostrata
|
| +-------------------------------------- Hyridella_depressa **
--+ |
| | +-- Margaritifera_margaritifera
| | +--------------------------------+
+--+ | +-- Cumberlandia_monodonta
| |
| | +-- Unio_pictorum
| | +-----------------------------+
+--+ | +-- Unio_caffer
| |
| | +----------- Anodonta_cygnea
| | |
| | +-----------------+ +-- Alasmidonta_marginata
+--+ | | +-----+
| | | | +-- Lasmigona_costata
| | +--+
| | | +----- Strophitus_undulatus
| | +--+
| | | +-- Pyganodon_grandis
+--+ +--+
| +-- Pyganodon_fragilis
|
| +-------- Quadrula_quadrula
| +-----------------+
| | | +----- Eurynia_dilatata
| | +--+
| | | +-- Pleurobema_sintoxia
+--+ +--+
| +-- Fusconaia_flava
|
| +----------------------- Amblema_plicata
| |
+--+ +-------------------- Truncilla_truncata
| |
+--+ +----------------- Ligumia_recta
| |
| | +-- Ptychobranchus_fasciolaris
+--+ +-----------+
| | +-- Sagittunio_nasutus
| |
+--+ +----------- Lampsilis_siliquoidea
| |
| | +----- Ortmanniana_ligamentina
+--+ +--+
| | | +-- Lampsilis_cardium
| | +--+
| | +-- Lampsilis_fasciola
+--+
| +-- Epioblasma_brevidens
| +--+
| | +-- Epioblasma_triquetra
+--+
| +-- Cambarunio_iris
+--+
+-- Leaunio_vanuxemensis
Chase et al., 2018 [Publ 3861], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
10 |
10 |
1 |
-- |
+-------------------- Echyridella_menziesii **
|
--+ +----------------- Cumberlandia_monodonta
| |
| | +----- Quadrula_quadrula
+--+ +--------+
| | | +-- Lampsilis_powellii
| | +--+
+--+ +-- Lampsilis_siliquoidea
|
| +----------- Lamprotula_leaii
| |
+--+ +-- Unio_delphinus
| +-----+
| | +-- Unio_tumidus
+--+
| +----- Anodonta_anatina
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Pyganodon_grandis
Wu et al., 2021 [Publ 4366], fig. 3-ML
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
73 |
72 |
2 |
-- |
+-------------------------------------------- Echyridella_menziesii **
|
| +-- Anodontites_trapesialis
| +--------------------------------------+
| | +-- Mutela_dubia
| |
| | +-- Gibbosula_rochechouartii
--+ | +-----------+
| | | +-- Gibbosula_crassa
| | +-----------------------+
| | | | +----------- Cumberlandia_monodonta
| | | +--+
| | | | +-------- Pseudunio_marocanus
| | | +--+
| | | | +----- Margaritifera_falcata
+--+ | +--+
| | | +-- Margaritifera_margaritifera
| | +--+
| | +-- Margaritifera_dahurica
| |
| | +-- Uniomerus_tetralasmus
| | +--------------------------+
| | | +-- Quadrula_quadrula
| | |
| | +-----+ +-- Pleurobema_oviforme
| | | | +-----------------------+
| | | | | +-- Elliptio_complanata
| | | +--+
+--+ | | +----------------------- Amblema_plicata
| | | |
| | +--+ +-------------------- Popenaias_popeii
| | | |
| | +--+ +----------------- Toxolasma_parvum
| | | |
| | +--+ +-- Potamilus_alatus
| | | +-----------+
| | | | +-- Potamilus_leptodon
| | +--+
| | | +----------- Leaunio_lienosus
| | | |
| | +--+ +-- Lampsilis_ornata
| | | +-----+
| | | | +-- Lampsilis_cardium
| | +--+
| | | +----- Venustaconcha_ellipsiformis
| | +--+
| | | +-- Lampsilis_siliquoidea
| | +--+
+--+ +-- Lampsilis_powellii
|
| +----- Chamberlainia_somsakpanhai
| +-----------+
| | | +-- Sinohyriopsis_schlegelii
| | +--+
| | +-- Sinohyriopsis_cumingii
| |
| +--------------+ +-------- Potomida_littoralis
| | | +-----+
| | | | | +----- Pronodularia_japanensis
| | | | +--+
| | | | | +-- Lamprotula_caveata
| | | | +--+
| | +--+ +-- Lamprotula_leaii
| | |
| | | +-- Pilsbryoconcha_exilis
| | | +--------+
| | | | +-- Pseudodon_vondembuschianus
| | | |
| | +--+ +-- Parvasolenaia_rivularis
| | | +-----+
| | | | +-- Ptychorhynchus_pfisteri
| | +--+
| | | +----- Microcondylaea_bonellii
+--+ +--+
| | +-- Sinosolenaia_oleivora
| +--+
| +-- Sinosolenaia_carinata
|
| +----------------------------- Lepidodesma_languilati
| |
| | +-------- Aculamprotula_tortuosa
| | |
| | +--------+ +-- Aculamprotula_scripta
| | | | +--+
| | | | | +-- Aculamprotula_polysticta
| | | +--+
| | | | +-- Aculamprotula_coreana
| | | +--+
| | +--------+ +-- Aculamprotula_tientsinensis
| | | |
+--+ | | +-------- Unio_tumidus
| | | +-----+
| | | | | +----- Unio_nanus
| | | | +--+
| | | | | +-- Unio_pictorum
| | +--+ +--+
| | | +-- Unio_delphinus
| | |
| | | +----------- Pseudocuneopsis_capitata
| | | |
| | +--+ +-- Nodularia_douglasiae
| | | +-----+
| | | | +-- Nodularia_breviconcha
+--+ +--+
| | +----- Tchangsinaia_piscicula
| +--+
| | +-- Schistodesmus_lampreyanus
| +--+
| +-- Cuneopsis_heudei
|
| +-------- Acuticosta_chinensis
| +--------------+
| | | +----- Lanceolaria_gladiola
| | +--+
| | | +-- Lanceolaria_grayii
| | +--+
| | +-- Lanceolaria_lanceolata
| |
+--+ +-------- Cristaria_plicata
| |
| +-----------+ +-- Sinanodonta_woodiana
| | | +--+
| | | | +-- Sinanodonta_lucida
| | +--+
| | | +-- Anemina_euscaphys
+--+ +--+
| +-- Anemina_arcaeformis
|
| +----------------- Anodonta_cygnea
| |
+--+ +-------------- Anodonta_anatina
| |
| | +----- Pyganodon_grandis
+--+ +-----+
| | | +-- Utterbackia_peninsularis
| | +--+
+--+ +-- Utterbackia_imbecillis
|
| +-------- Prolasmidonta_heterodon
+--+
| +----- Alasmidonta_varicosa
+--+
| +-- Platynaias_subviridis
+--+
+-- Platynaias_compressa
Walker et al., 2006 [Publ 2666], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
14 |
13 |
1 |
-- |
+-------------------------------- Hyridella **
|
--+ +----------------------------- Cumberlandia_monodonta
| |
+--+ +-------------------------- Pseudodon_vondembuschianus
| |
+--+ +----------------------- Gonidea_angulata
| |
+--+ +-------------------- Inversidens
| |
| | +----- Anodonta
+--+ +-----------+
| | | +-- Pyganodon_fragilis
| | +--+
+--+ +-- Pyganodon_grandis
|
| +-------------- Fusconaia_flava
+--+
| +----------- Cyrtonaias_tampicoensis
+--+
| +-------- Potamilus_purpuratus
+--+
| +----- Ligumia_recta
+--+
| +-- Ortmanniana_ligamentina
+--+
+-- Lampsilis_teres
Huang et al., 2018 [Publ 3828], fig. 6-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
16 |
16 |
1 |
-- |
+----------------------- Echyridella_menziesii **
|
| +-------------------- Cumberlandia_monodonta
--+ |
| | +----- Quadrula_quadrula
| | +-----------+
+--+ | | +-- Venustaconcha_ellipsiformis
| | +--+
| | +-- Potamilus_alatus
| |
| | +-------- Sinosolenaia_carinata
+--+ +-----+
| | | +----- Potomida_littoralis
| | +--+
| | | +-- Lamprotula_leaii
| | +--+
| | +-- Pronodularia_japanensis
+--+
| +-------- Sinanodonta_woodiana
| +--+
| | | +----- Anodonta_anatina
| | +--+
| | | +-- Pyganodon_grandis
+--+ +--+
| +-- Utterbackia_peninsularis
|
| +-------- Aculamprotula_tortuosa
+--+
| +----- Unio_tumidus
+--+
| +-- Unio_crassus
+--+
+-- Unio_delphinus
Bieler et al., 2014 [Publ 3622], fig. 26, 27, 29, 30
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
4 |
3 |
2 |
-- |
+-------- Velesunio_ambiguus **
--+
| +----- Unio_pictorum
+--+
| +-- Aspatharia_pfeifferiana
+--+
+-- Margaritifera_margaritifera
Huang et al., 2018 [Publ 3828], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
42 |
41 |
2 |
-- |
+-- Anodontites_trapesialis
+--------------------------+
| +-- Mutela_dubia
|
| +-------------------------- Echyridella_menziesii **
| |
| | +-------------- Gibbosula_rochechouartii
--+ | +--------+
| | | | +----------- Cumberlandia_monodonta
| | | +--+
| | | | +-------- Pseudunio_marocanus
| | | +--+
| | | | +----- Margaritifera_falcata
| | | +--+
+--+ | | +-- Margaritifera_margaritifera
| | +--+
| | +-- Margaritifera_dahurica
| |
| | +----------- Quadrula_quadrula
| | |
| | +-----+ +-------- Toxolasma_parvum
| | | | |
| | | +--+ +-- Lampsilis_ornata
| | | | +--+
| | | | | +-- Venustaconcha_ellipsiformis
+--+ | +--+
| | | +-- Potamilus_leptodon
| +--+ +--+
| | | +-- Potamilus_alatus
| | |
| | | +-- Sinohyriopsis_cumingii
| | | +-----------+
| | | | +-- Sinohyriopsis_schlegelii
| | | |
| | +--+ +-- Sinosolenaia_oleivora
| | | +--------+
| | | | +-- Sinosolenaia_carinata
| | +--+
| | | +-------- Potomida_littoralis
| | +--+
| | | +----- Pronodularia_japanensis
| | +--+
| | | +-- Lamprotula_caveata
+--+ +--+
| +-- Lamprotula_leaii
|
| +----- Aculamprotula_tortuosa
| +--------+
| | | +-- Aculamprotula_coreana
| | +--+
| | +-- Aculamprotula_tientsinensis
| +--+
| | | +-- Tchangsinaia_piscicula
| | | +--------+
| | | | +-- Nodularia_douglasiae
| | +--+
| | | +-------- Unio_tumidus
| | +--+
| | | +----- Unio_crassus
| | +--+
+--+ | +-- Unio_delphinus
| +--+
| +-- Unio_pictorum
|
| +----------- Anodonta_anatina
| +--+
| | | +-------- Platynaias_compressa
| | +--+
| | | +----- Pyganodon_grandis
| | +--+
+--+ | +-- Utterbackia_imbecillis
| +--+
| +-- Utterbackia_peninsularis
|
| +-------- Cristaria_plicata
| |
+-----+ +-- Anemina_arcaeformis
| +--+
| | +-- Anemina_euscaphys
+--+
| +-- Sinanodonta_woodiana
+--+
+-- Sinanodonta_lucida
Wu et al., 2023 [Publ 4339], fig. 4, S5-aa
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
95 |
94 |
2 |
-- |
+----------------------------------------------- Echyridella_menziesii **
|
| +-- Anodontites_trapesialis
| +-----------------------------------------+
| | +-- Mutela_dubia
| |
| | +-- Gibbosula_crassa
| | +-----------+
--+ | | +-- Gibbosula_rochechouartii
| | +--------------------------+
| | | | +----------- Cumberlandia_monodonta
| | | +--+
| | | | +-------- Pseudunio_marocanus
| | | +--+
| | | | +----- Margaritifera_falcata
| | | +--+
| | | | +-- Margaritifera_dahurica
+--+ | +--+
| | +-- Margaritifera_margaritifera
| |
| | +-- Quadrula_quadrula
| | +--------------------------+
| | | +-- Uniomerus_tetralasmus
| | |
| | +--------+ +-- Elliptio_complanata
| | | | +-----------------------+
| | | | | +-- Pleurobema_oviforme
| | | +--+
| | | | +----------------------- Amblema_plicata
| | | | |
| | | +--+ +-------------------- Popenaias_popeii
+--+ | | |
| | +--+ +----------------- Toxolasma_parvum
| | | |
| | | | +----- Potamilus_leptodon
| | +--+ +--------+
| | | | | +-- Potamilus_alatus
| | | | +--+
| | | | +-- Potamilus_streckersoni
| | +--+
| | | +----------- Leaunio_lienosus
| | | |
| | | | +----- Lampsilis_ornata
| | +--+ +--+
| | | | | +-- Lampsilis_cardium
| | | | +--+
| | +--+ +-- Lampsilis_cariosa
| | |
| | | +----- Venustaconcha_ellipsiformis
| | +--+
| | | +-- Lampsilis_powellii
| | +--+
| | +-- Lampsilis_siliquoidea
+--+
| +----- Chamberlainia_somsakpanhai
| +-----------------+
| | | +-- Sinohyriopsis_cumingii
| | +--+
| | +-- Sinohyriopsis_schlegelii
| |
| | +-- Hyriopsis_bialata
| | +--+
| +-----------+ | +-- Rectidens_sumatrensis
| | | +--------------+
| | | | | +-- Lens_contradens
| | | | +--+
| | | | +-- Physunio_superbus
| | | |
| | | | +----------- Potomida_littoralis
| | +--+ +-----+
| | | | | +-------- Pronodularia_japanensis
| | | | +--+
| | | | | +----- Lamprotula_leaii
| | | | +--+
| | | | | +-- Lamprotula_caveata
| | +--+ +--+
| | | +-- Lamprotula_cornuumlunae
| | |
| | | +-- Pseudodon_vondembuschianus
| | | +-----------+
| | | | +-- Pilsbryoconcha_exilis
| | +--+
| | | +----------- Microcondylaea_bonellii
+--+ | |
| +--+ +-- Parvasolenaia_rivularis
| | +-----+
| | | +-- Ptychorhynchus_pfisteri
| +--+
| | +----- Sinosolenaia_carinata
| +--+
| | +-- Sinosolenaia_iridinea
| +--+
| +-- Sinosolenaia_oleivora
|
| +-------------------------------- Lamellidens_jenkinsianus
| |
| | +-- Lepidodesma_aligera
| | +--------------------------+
| | | +-- Lepidodesma_languilati
| | |
| | | +-------- Aculamprotula_tortuosa
| | | |
| | | +--------------+ +-- Aculamprotula_coreana
| | | | | +--+
+--+ | | | | +-- Aculamprotula_tientsinensis
| | | +--+
| | | | +-- Aculamprotula_polysticta
| | | +--+
| | | +-- Aculamprotula_scripta
| | |
| | +--+ +-------------- Unio_tumidus
| | | | +-----+
| | | | | | +----------- Unio_nanus
| | | | | +--+
+--+ | | | | +-------- Unio_elongatulus
| | | | +--+
| | | | | +----- Unio_pictorum
| | | | +--+
| | | | | +-- Unio_delphinus
| | +--+ +--+
| | | +-- Unio_mancus
| | |
| | | +-- Nodularia_breviconcha
| | | +--+
| | | | +-- Nodularia_nipponensis
| | | +-----------+
| | | | | +-- Pseudocuneopsis_capitata
| | | | +--+
| | +--+ +-- Pseudocuneopsis_sichuanensis
| | |
| | | +-------------- Tchangsinaia_piscicula
| | | |
+--+ +--+ +-- Schistodesmus_lampreyanus
| | +--------+
| | | +-- Schistodesmus_spinosus
| +--+
| | +-------- Cuneopsis_rufescens
| +--+
| | +----- Cuneopsis_celtiformis
| +--+
| | +-- Cuneopsis_demangei
| +--+
| +-- Cuneopsis_heudei
|
| +-------- Acuticosta_chinensis
| +--------+
| | | +----- Lanceolaria_gladiola
| | +--+
| | | +-- Lanceolaria_grayii
| | +--+
| | +-- Lanceolaria_lanceolata
| |
| | +----- Cristaria_plicata
| | +-----+
| | | | +-- Anemina_arcaeformis
+--------+ | +--+
| +--+ +-- Anemina_euscaphys
| | |
| | | +-------- Beringiana_fukuharai
| | +--+
| | | +----- Sinanodonta_lucida
| | +--+
| | | +-- Sinanodonta_tumens
| | +--+
| | +-- Sinanodonta_woodiana
+--+
| +-- Anodonta_nuttalliana
| +-----+
| | +-- Pseudanodonta_complanata
| +--+
| | | +----- Anodonta_anatina
| | +--+
| | | +-- Anodonta_cygnea
| | +--+
| | +-- Anodonta_exulcerata
+--+
| +----- Pyganodon_grandis
| +--+
| | | +-- Utterbackia_peninsularis
| | +--+
| | +-- Utterbackia_imbecillis
+--+
| +-- Alasmidonta_varicosa
| +--+
| | +-- Platynaias_compressa
+--+
| +-- Prolasmidonta_heterodon
+--+
+-- Lasmigona_complanata
González & Giribet, 2015 [Publ 3632], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
5 |
3 |
2 |
-- |
+-------- Velesunio_ambiguus **
--+
| +----- Unio_pictorum
+--+
| +-- Aspatharia_pfeifferiana
+--+
+-- Margaritifera_margaritifera
Guerra et al., 2017 [Publ 3772], fig. 2a-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
7 |
7 |
1 |
-- |
+----------- Echyridella_menziesii **
|
--+ +-------- Cumberlandia_monodonta
| |
| | +-- Sinosolenaia_carinata
+--+ +--+
| | +-- Pronodularia_japanensis
| |
| | +-- Utterbackia_peninsularis
+--+--+
| +-- Pyganodon_grandis
|
| +-- Venustaconcha_ellipsiformis
+--+
+-- Quadrula_quadrula
Guerra et al., 2017 [Publ 3772], fig. 2b
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
24 |
23 |
2 |
-- |
+-- Mutela_dubia
+--------------------------------+
| +-- Anodontites_trapesialis
|
--+ +-------------------------------- Echyridella_menziesii **
| |
| | +-- Margaritifera_falcata
| | +--------------------------+
+--+ | +-- Cumberlandia_monodonta
| |
| | +-------- Quadrula_quadrula
| | +-----------------+
+--+ | | +----- Toxolasma_parvum
| | +--+
| | | +-- Venustaconcha_ellipsiformis
| | +--+
| | +-- Lampsilis_ornata
| |
+--+ +-- Sinohyriopsis_schlegelii
| +-----+
| | +-- Sinohyriopsis_cumingii
| +--------------+
| | | +----- Pronodularia_japanensis
| | +--+
| | | +-- Sinosolenaia_oleivora
+--+ +--+
| +-- Sinosolenaia_carinata
|
| +-------------------- Unio_pictorum
| |
+--+ +----------------- Aculamprotula_tortuosa
| |
+--+ +-- Cristaria_plicata
| +-----------+
| | +-- Sinanodonta_woodiana
+--+
| +----------- Anodonta_anatina
| |
+--+ +-- Platynaias_subviridis
| +-----+
| | +-- Platynaias_compressa
+--+
| +----- Pyganodon_grandis
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Utterbackia_imbecillis
Huang et al., 2019 [Publ 3843], fig. 3-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
22 |
22 |
1 |
-- |
+----------------------------- Echyridella_menziesii **
|
| +-- Pseudunio_marocanus
| +-----------------------+
--+ | +-- Margaritifera_margaritifera
| |
| | +----- Quadrula_quadrula
| | +-----------------+
| | | | +-- Venustaconcha_ellipsiformis
+--+ | +--+
| | +-- Potamilus_alatus
| |
| | +----------- Sinohyriopsis_cumingii
| | |
| | | +----- Potomida_littoralis
+--+ +--------+ +--+
| | | | | +-- Lamprotula_leaii
| | | | +--+
| | +--+ +-- Pronodularia_japanensis
| | |
| | | +----- Parvasolenaia_rivularis
| | +--+
| | | +-- Sinosolenaia_oleivora
+--+ +--+
| +-- Sinosolenaia_carinata
|
| +-------- Sinanodonta_woodiana
| +--------+
| | | +----- Anodonta_anatina
| | +--+
| | | +-- Pyganodon_grandis
+--+ +--+
| +-- Utterbackia_peninsularis
|
| +-------------- Lanceolaria_lanceolata
+--+
| +----------- Schistodesmus_lampreyanus
+--+
| +-------- Aculamprotula_tortuosa
+--+
| +----- Unio_tumidus
+--+
| +-- Unio_nanus
+--+
+-- Unio_delphinus
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
29 |
28 |
2 |
-- |
+-------------------------------- Mutela_rostrata
|
| +----------------------------- Hyridella_depressa **
--+ |
| | +-- Margaritifera_margaritifera
| | +-----------------------+
+--+ | +-- Cumberlandia_monodonta
| |
| | +-- Unio_pictorum
| | +--------------------+
+--+ | +-- Unio_caffer
| |
| | +----------- Anodonta_cygnea
| | |
| | +--------+ +-- Alasmidonta_marginata
+--+ | | +-----+
| | | | +-- Lasmigona_costata
| | +--+
| | | +----- Strophitus_undulatus
| | +--+
| | | +-- Pyganodon_grandis
| | +--+
+--+ +-- Pyganodon_fragilis
|
| +-------- Quadrula_quadrula
| +--------+
| | | +----- Eurynia_dilatata
| | +--+
| | | +-- Pleurobema_sintoxia
| | +--+
+--+ +-- Fusconaia_flava
|
| +-------------- Amblema_plicata
| |
| | +----------- Ligumia_recta
| | |
+--+ +----------- Truncilla_truncata
| |
| | +-- Ptychobranchus_fasciolaris
| +--------+
+--+ +-- Sagittunio_nasutus
|
| +----- Lampsilis_siliquoidea
| +--+
| | | +-- Epioblasma_triquetra
| | +--+
| | +-- Epioblasma_brevidens
+--+
| +----- Ortmanniana_ligamentina
| |
| +----- Cambarunio_iris
+--+
+----- Leaunio_vanuxemensis
|
| +-- Lampsilis_cardium
+--+
+-- Lampsilis_fasciola
Breton et al., 2011 [Publ 3376], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
36 |
36 |
1 |
-- |
+-------------------------------------- Echyridella_menziesii **
|
| +-------- Cumberlandia_monodonta
| +--------------------------+
| | | +----- Margaritifera_margaritifera
| | +--+
--+ | | +-- Margaritifera_falcata
| | +--+
| | +-- Margaritifera_marrianae
| |
| | +----- Quadrula_quadrula
| | +--------------------------+
+--+ | | +-- Cyclonaias_tuberculata
| | +--+
| | +-- Pustulosa_pustulosa
| |
| | +-------------------------- Pronodularia_japanensis
| | |
| | +--+ +----------------------- Sinohyriopsis_cumingii
| | | | |
| | | +--+ +-------------------- Cristaria_plicata
| | | | |
+--+ | | | +-- Lasmigona_complanata
| | +--+ +--------------+
| | | | +-- Lasmigona_costata
| | | |
| | +--+ +-------------- Strophitus_undulatus
| | | |
| | | | +-- Platynaias_compressa
| | | | +--+
| | +--+ | +-- Platynaias_subviridis
| | | +-----+
| | | | | +-- Alasmidonta_marginata
+--+ | | +--+
| | | +-- Alasmidonta_undulata
| +--+
| | +-- Pyganodon_lacustris
| | +-----+
| | | +-- Pyganodon_grandis
| +--+
| | +----- Utterbackia_peninsularis__2
| +--+
| | +-- Utterbackia_peninsularis__1
| +--+
| +-- Utterbackia_imbecillis
|
| +-------------------------- Megalonaias_nervosa
| |
+--+ +----------------------- Fusconaia_flava
| |
+--+ +-------------------- Reginaia_ebenus
| |
| | +-------- Toxolasma_paulum__2
+--+ +--------+
| | | +----- Toxolasma_lividum
| | +--+
| | | +-- Toxolasma_parvum
+--+ +--+
| +-- Toxolasma_paulum__1
|
| +-------------- Ellipsaria_lineolata
| |
+--+ +----------- Truncilla_macrodon
| |
| | +-- Lemiox_rimosus
+--+ +-----+
| | +-- Potamilus_metnecktayi
| |
+--+ +-- Cambarunio_iris
| +--+
| | +-- Venustaconcha_ellipsiformis
+--+
| +-- Lampsilis_powellii
| |
+--+-- Lampsilis_ornata
|
+-- Lampsilis_siliquoidea
Chase et al., 2018 [Publ 3847], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
37 |
37 |
1 |
-- |
+-- Echyridella_menziesii **
+--------+
| +-- Reginaia_ebenus
--+
| +-------- Fusconaia_flava
+--+
| +----- Megalonaias_nervosa
+--+
| +-- Cyclonaias_tuberculata
+--+
+-- Pustulosa_pustulosa
González & Giribet, 2015 [Publ 3632], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
5 |
3 |
2 |
-- |
+-- Aspatharia_pfeifferiana
+--+
| +-- Margaritifera_margaritifera
--+
| +-- Unio_pictorum
+--+
+-- Velesunio_ambiguus **
Hoeh et al., 2002 [Publ 2673], fig. 1-F
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
8 |
8 |
1 |
-- |
+-------------------- Echyridella_menziesii **
|
--+ +-- Pyganodon_fragilis
| +--------------+
| | +-- Pyganodon_grandis
+--+
| +-------------- Fusconaia_flava
+--+
| +----------- Cyrtonaias_tampicoensis
+--+
| +-------- Ortmanniana_ligamentina
+--+
| +----- Lampsilis_teres
+--+
| +-- Potamilus_purpuratus
+--+
+-- Ligumia_recta
Hoeh et al., 2002 [Publ 2673], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
8 |
8 |
1 |
-- |
+-------------------- Echyridella_menziesii **
|
--+ +-- Pyganodon_fragilis
| +--------------+
| | +-- Pyganodon_grandis
+--+
| +-------------- Fusconaia_flava
+--+
| +----------- Potamilus_purpuratus
+--+
| +-------- Cyrtonaias_tampicoensis
+--+
| +----- Ligumia_recta
+--+
| +-- Lampsilis_teres
+--+
+-- Ortmanniana_ligamentina
Wu et al., 2021 [Publ 4366], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
68 |
67 |
2 |
-- |
+----------------------------------------- Echyridella_menziesii **
|
| +-- Mutela_dubia
| +-----------------------------------+
| | +-- Anodontites_trapesialis
| |
| | +-- Gibbosula_rochechouartii
--+ | +-----------+
| | | +-- Gibbosula_crassa
| | +--------------------+
| | | | +----------- Cumberlandia_monodonta
| | | +--+
| | | | +-------- Pseudunio_marocanus
| | | +--+
+--+ | | +----- Margaritifera_falcata
| | +--+
| | | +-- Margaritifera_margaritifera
| | +--+
| | +-- Margaritifera_dahurica
| |
| | +-------------------------- Quadrula_quadrula
| | |
| | +-----+ +----------------------- Pleurobema_oviforme
| | | | |
| | | +--+ +-------------------- Amblema_plicata
| | | | |
+--+ | +--+ +----------------- Popenaias_popeii
| | | |
| | +--+ +-------------- Toxolasma_parvum
| | | |
| | | | +-- Potamilus_alatus
| | +--+ +--------+
| | | | +-- Potamilus_leptodon
| | | |
| | +--+ +-- Lampsilis_ornata
| | | +-----+
| | | | +-- Lampsilis_cardium
| | +--+
| | | +----- Venustaconcha_ellipsiformis
| | +--+
| | | +-- Lampsilis_siliquoidea
| | +--+
| | +-- Lampsilis_powellii
| |
+--+ +----- Chamberlainia_somsakpanhai
| +-----------+
| | | +-- Sinohyriopsis_schlegelii
| | +--+
| | +-- Sinohyriopsis_cumingii
| |
| +-----------+ +----- Potomida_littoralis
| | | +--------+
| | | | | +-- Lamprotula_caveata
| | | | +--+
| | | | +-- Lamprotula_leaii
| | +--+
| | | +-- Pilsbryoconcha_exilis
| | | +--------+
| | | | +-- Pseudodon_vondembuschianus
| | | |
| | +--+ +-- Parvasolenaia_rivularis
| | | +-----+
| | | | +-- Ptychorhynchus_pfisteri
| | +--+
| | | +----- Microcondylaea_bonellii
| | +--+
+--+ | +-- Sinosolenaia_oleivora
| +--+
| +-- Sinosolenaia_carinata
|
| +-------------------------- Lepidodesma_languilati
| |
| | +-------- Aculamprotula_tortuosa
| | |
| | +--------+ +-- Aculamprotula_scripta
| | | | +--+
| | | | | +-- Aculamprotula_polysticta
| | | +--+
| | | | +-- Aculamprotula_coreana
| | | +--+
| | +-----+ +-- Aculamprotula_tientsinensis
| | | |
+--+ | | +-------- Unio_tumidus
| | | +-----+
| | | | | +----- Unio_nanus
| | | | +--+
| | | | | +-- Unio_pictorum
| | +--+ +--+
| | | +-- Unio_delphinus
| | |
| | | +----------- Pseudocuneopsis_capitata
| | | |
| | +--+ +-- Nodularia_douglasiae
| | | +-----+
| | | | +-- Nodularia_breviconcha
+--+ +--+
| | +----- Schistodesmus_lampreyanus
| +--+
| | +-- Tchangsinaia_piscicula
| +--+
| +-- Cuneopsis_heudei
|
| +-------- Acuticosta_chinensis
| +-----------+
| | | +----- Lanceolaria_gladiola
| | +--+
| | | +-- Lanceolaria_grayii
| | +--+
| | +-- Lanceolaria_lanceolata
| |
+--+ +-------- Cristaria_plicata
| |
| +--------+ +-- Sinanodonta_woodiana
| | | +--+
| | | | +-- Sinanodonta_lucida
| | +--+
| | | +-- Anemina_euscaphys
+--+ +--+
| +-- Anemina_arcaeformis
|
| +-------------- Anodonta_cygnea
| |
+--+ +----------- Anodonta_anatina
| |
| | +----- Pyganodon_grandis
+--+ +--+
| | | +-- Utterbackia_imbecillis
| | +--+
+--+ +-- Utterbackia_peninsularis
|
| +----- Prolasmidonta_heterodon
+--+
| +-- Platynaias_compressa
+--+
+-- Alasmidonta_varicosa
Bieler et al., 2014 [Publ 3622], fig. 28
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
4 |
3 |
2 |
-- |
+-- Aspatharia_pfeifferiana
+--+
| +-- Margaritifera_margaritifera
--+
| +-- Unio_pictorum
+--+
+-- Velesunio_ambiguus **
Pfeiffer et al., 2019 [Publ 3937], fig. 4-5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
56 |
55 |
2 |
-- |
+----- Tamsiella_schroeteriana
+--------------------------------+
| | +-- Etheria_elliptica
| +--+
| +-- Aspatharia_pfeifferiana
|
| +----------------------------------- Hyridella_australis **
| |
--+ | +-- Margaritifera_margaritifera
| | +-----------------------------+
| | | +-- Margaritifera_hembeli
| | |
| | | +-- Quadrula_quadrula
| | | +--------------------+
| | | | +-- Uniomerus_tetralasmus
+--+ | |
| | | +----- Elliptoideus_sloatianus
| | +-----+ +--------------+
| | | | | | +-- Elliptio_complanata
| | | | | +--+
| | | | | +-- Pleurobema_strodeanum
| | | | |
| | | +--+ +----- Reginaia_ebenus
| | | | +-----------+
| | | | | | +-- Plectomerus_dombeyanus
+--+ | | | +--+
| | | | +-- Amblema_plicata
| | +--+
| | | +-- Psoronaias_semigranosa
| | | +-----------+
| | | | +-- Popenaias_popeii
| | +--+
| | | +----------- Obliquaria_reflexa
| | +--+
| | | +-------- Cyrtonaias_tampicoensis
| | +--+
| | | +----- Leaunio_lienosus
| | +--+
| | | +-- Lampsilis_cardium
| | +--+
| | +-- Pachynaias_spheniopsis
| |
+--+ +-- Chamberlainia_hainesiana
| +--------------------+
| | +-- Sinohyriopsis_cumingii
| |
| | +----------------- Lamprotula_cornuumlunae
| | +--+
| | | | +-------------- Gonidea_angulata
| +--+ | +--+
| | | | | +----------- Indopseudodon_kayinensis
| | | | +--+
| | | | | +-------- Bineurus_anodontinum
| | | | +--+
| | | | | +----- Pilsbryoconcha_exilis
| | | | +--+
| | +--+ | +-- Pseudodon_vondembuschianus
| | | +--+
| | | +-- Sundadontina_tanintharyiensis
| | |
| | | +----- Ensidens_ingallsianus
| | | +-----+
| | | | | +-- Hyriopsis_bialata
| | | | +--+
| | | | +-- Rectidens_sumatrensis
| | +--------+
| | | +-- Trapezoideus_foliaceus
| | | +-----+
+--+ | | +-- Physunio_superbus
| +--+
| | +----- Yaukthwa_paiensis
| +--+
| | +-- Lens_contradens
| +--+
| +-- Solenaia_emarginata
|
| +----------- Lanceolaria_lanceolata
| |
| | +----- Unio_gibbus
| +--------+ +--+
| | | | | +-- Nodularia_douglasiae
| | | | +--+
| | | | +-- Tchangsinaia_piscicula
| | +--+
| | | +-- Cristaria_plicata
| | | +--+
| | | | +-- Sinanodonta_woodiana
+-----+ +--+
| | +-- Anodonta_cygnea
| +--+
| +-- Pyganodon_grandis
|
| +-- Trapezidens_dolichorhynchus
| +--------------+
| | +-- Lamellidens_savadiensis
| |
+--+ +-- Leoparreysia_tavoyensis
| +-----------+
| | +-- Leoparreysia_olivacea
| |
+--+ +-- Coelatura_choziensis
| +--------+
| | +-- Prisodontopsis_aviculaeformis
+--+
| +-------- Indonaia_caerulea
+--+
| +----- Indochinella_pugio
+--+
| +-- Scabies_phaselus
+--+
+-- Harmandia_somboriensis
Chase et al., 2018 [Publ 3847], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
9 |
9 |
1 |
-- |
+-- Cumberlandia_monodonta
+-----------+
| +-- Echyridella_menziesii **
|
--+ +----------- Sinosolenaia_carinata
| |
| | +----- Potamilus_alatus
+--+ +--+
| | | +-- Lampsilis_siliquoidea
| | +--+
| | +-- Lampsilis_powellii
+--+
| +-- Anodonta_anatina
| +--+
| | +-- Utterbackia_peninsularis
+--+
| +-- Unio_delphinus
+--+
+-- Unio_tumidus
Walker et al., 2006 [Publ 2666], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
14 |
13 |
1 |
-- |
+-------------------------------- Hyridella **
|
--+ +----------------------------- Cumberlandia_monodonta
| |
+--+ +-------------------------- Pseudodon_vondembuschianus
| |
+--+ +----------------------- Gonidea_angulata
| |
+--+ +-------------------- Inversidens
| |
| | +----- Anodonta
+--+ +-----------+
| | | +-- Pyganodon_fragilis
| | +--+
+--+ +-- Pyganodon_grandis
|
| +-------------- Fusconaia_flava
+--+
| +----------- Potamilus_purpuratus
+--+
| +-------- Cyrtonaias_tampicoensis
+--+
| +----- Ligumia_recta
+--+
| +-- Ortmanniana_ligamentina
+--+
+-- Lampsilis_teres
Wu et al., 2023 [Publ 4339], fig. 4, S5-nt
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
95 |
94 |
2 |
-- |
+-------------------------------------------------- Echyridella_menziesii **
|
| +-- Anodontites_trapesialis
| +--------------------------------------------+
| | +-- Mutela_dubia
| |
| | +-- Gibbosula_crassa
| | +-----------+
--+ | | +-- Gibbosula_rochechouartii
| | +-----------------------------+
| | | | +----------- Cumberlandia_monodonta
| | | +--+
| | | | +-------- Pseudunio_marocanus
| | | +--+
| | | | +----- Margaritifera_falcata
| | | +--+
| | | | +-- Margaritifera_dahurica
+--+ | +--+
| | +-- Margaritifera_margaritifera
| |
| | +-- Quadrula_quadrula
| | +--------------------------+
| | | +-- Uniomerus_tetralasmus
| | |
| | +-----------+ +-- Elliptio_complanata
| | | | +-----------------------+
| | | | | +-- Pleurobema_oviforme
| | | +--+
| | | | +----------------------- Amblema_plicata
| | | | |
| | | +--+ +-------------------- Popenaias_popeii
+--+ | | |
| | +--+ +----------------- Toxolasma_parvum
| | | |
| | | | +----- Potamilus_leptodon
| | +--+ +--------+
| | | | | +-- Potamilus_alatus
| | | | +--+
| | | | +-- Potamilus_streckersoni
| | +--+
| | | +----------- Leaunio_lienosus
| | | |
| | | | +----- Lampsilis_ornata
| | +--+ +--+
| | | | | +-- Lampsilis_cardium
| | | | +--+
| | +--+ +-- Lampsilis_cariosa
| | |
| | | +----- Venustaconcha_ellipsiformis
| | +--+
| | | +-- Lampsilis_powellii
| | +--+
| | +-- Lampsilis_siliquoidea
+--+
| +----- Chamberlainia_somsakpanhai
| +-----------------+
| | | +-- Sinohyriopsis_cumingii
| | +--+
| | +-- Sinohyriopsis_schlegelii
| |
| | +-- Hyriopsis_bialata
| | +--+
| +--------------+ | +-- Rectidens_sumatrensis
| | | +--------------+
| | | | | +-- Lens_contradens
| | | | +--+
| | | | +-- Physunio_superbus
| | | |
| | | | +----------- Potomida_littoralis
| | +--+ +-----+
| | | | | +-------- Pronodularia_japanensis
| | | | +--+
| | | | | +----- Lamprotula_leaii
| | | | +--+
| | | | | +-- Lamprotula_caveata
| | +--+ +--+
| | | +-- Lamprotula_cornuumlunae
| | |
| | | +-- Pseudodon_vondembuschianus
| | | +-----------+
| | | | +-- Pilsbryoconcha_exilis
| | | |
| | +--+ +-- Ptychorhynchus_pfisteri
+--+ | +--------+
| | | +-- Microcondylaea_bonellii
| +--+
| | +-------- Parvasolenaia_rivularis
| +--+
| | +----- Sinosolenaia_carinata
| +--+
| | +-- Sinosolenaia_iridinea
| +--+
| +-- Sinosolenaia_oleivora
|
| +----------------------------------- Lamellidens_jenkinsianus
| |
| | +-- Lepidodesma_aligera
| | +-----------------------------+
| | | +-- Lepidodesma_languilati
| | |
| | | +-------- Acuticosta_chinensis
| | | +--------------+
| | | | | +----- Lanceolaria_gladiola
| | | | +--+
+--+ | | | +-- Lanceolaria_grayii
| | | +--+
| | | +-- Lanceolaria_lanceolata
| | |
| | | +-------------- Cristaria_plicata
| | +-----+ |
| | | | +-----+ +-- Anemina_arcaeformis
| | | | | | +--------+
| | | | | | | +-- Anemina_euscaphys
| | | | | +--+
+--+ | | | | +-------- Beringiana_fukuharai
| | | | +--+
| | | | | +----- Sinanodonta_lucida
| | | | +--+
| | +--+ | +-- Sinanodonta_tumens
| | | +--+
| | | +-- Sinanodonta_woodiana
| | |
| | | +----------------- Anodonta_nuttalliana
| | | |
| | | +----------------- Pseudanodonta_complanata
| | | |
| | +--+ +-------------- Anodonta_cygnea
| | | |
| | | | +-- Anodonta_exulcerata
| | | +-----------+
| | | | +-- Anodonta_anatina
| | +--+
+--+ | +----- Pyganodon_grandis
| | +-----+
| | | | +-- Utterbackia_imbecillis
| | | +--+
| +--+ +-- Utterbackia_peninsularis
| |
| | +-------- Lasmigona_complanata
| +--+
| | +----- Prolasmidonta_heterodon
| +--+
| | +-- Alasmidonta_varicosa
| +--+
| +-- Platynaias_compressa
|
| +-------- Aculamprotula_tortuosa
| |
| +-----------------+ +-- Aculamprotula_coreana
| | | +--+
| | | | +-- Aculamprotula_tientsinensis
| | +--+
| | | +-- Aculamprotula_polysticta
| | +--+
| | +-- Aculamprotula_scripta
| |
+--+ +----------- Unio_tumidus
| |
| +-----------+ +-------- Unio_nanus
| | | |
| | +--+ +----- Unio_elongatulus
| | | |
| | +--+ +-- Unio_delphinus
| | | |
| | +--+-- Unio_mancus
+--+ |
| +-- Unio_pictorum
|
| +-- Pseudocuneopsis_capitata
| +-----------------+
| | +-- Pseudocuneopsis_sichuanensis
| |
+--+ +-- Nodularia_breviconcha
| +--------------+
| | +-- Nodularia_nipponensis
| |
+--+ +----- Tchangsinaia_piscicula
| +--------+
| | | +-- Schistodesmus_lampreyanus__2
| | +--+
+--+ +-- Schistodesmus_spinosus
|
| +----------- Schistodesmus_lampreyanus__1
+--+
| +-------- Cuneopsis_rufescens
+--+
| +----- Cuneopsis_celtiformis
+--+
| +-- Cuneopsis_demangei
+--+
+-- Cuneopsis_heudei
Pfeiffer et al., 2019 [Publ 3937], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
56 |
55 |
2 |
-- |
+----- Hyridella_australis **
+--------------------+
| | +-- Margaritifera_margaritifera
| +--+
| +-- Margaritifera_hembeli
|
| +-- Quadrula_quadrula
| +-----------------+
| | +-- Uniomerus_tetralasmus
| |
| | +----- Elliptoideus_sloatianus
| | +--+
| | | | +-- Elliptio_complanata
| +--+ | +--+
| | | +--------+ +-- Pleurobema_strodeanum
| | | | |
| | | | | +----- Reginaia_ebenus
| | | | +--+
| | | | | +-- Plectomerus_dombeyanus
| | +--+ +--+
| | | +-- Amblema_plicata
| | |
| | | +-- Psoronaias_semigranosa
| | | +-----------+
--+ | | | +-- Popenaias_popeii
| | +--+
| | | +----------- Obliquaria_reflexa
| | +--+
| | | +-------- Cyrtonaias_tampicoensis
| | +--+
| | | +----- Leaunio_lienosus
| | +--+
| | | +-- Lampsilis_cardium
| | +--+
| | +-- Pachynaias_spheniopsis
| |
| | +-- Chamberlainia_hainesiana
| | +-----------------+
| | | +-- Sinohyriopsis_cumingii
| | |
| | | +-- Gonidea_angulata
| | | +-----------+
| | | | +-- Lamprotula_cornuumlunae
| | | +--+
| +--+ | | +----------- Indopseudodon_kayinensis
| | | | +--+
| | | | | +-------- Pilsbryoconcha_exilis
| | | | +--+
+--+ | | | +----- Bineurus_anodontinum
| | | +--+
| | | | +-- Pseudodon_vondembuschianus
| +--+ +--+
| | +-- Sundadontina_tanintharyiensis
| |
| | +----- Ensidens_ingallsianus
| | +-----+
| | | | +-- Hyriopsis_bialata
| | | +--+
| | | +-- Rectidens_sumatrensis
| +-----+
| | +-- Trapezoideus_foliaceus
| | +-----+
| | | +-- Physunio_superbus
| +--+
| | +----- Yaukthwa_paiensis
| +--+
| | +-- Lens_contradens
| +--+
| +-- Solenaia_emarginata
|
| +-------- Lanceolaria_lanceolata
| +--+
| | | +----- Unio_gibbus
| | +--+
| | | +-- Tchangsinaia_piscicula
| | +--+
| +--------+ +-- Nodularia_douglasiae
| | |
| | | +-- Cristaria_plicata
| | | +--+
| | | | +-- Sinanodonta_woodiana
| | +-----+
| | | +-- Anodonta_cygnea
+--+ +--+
| +-- Pyganodon_grandis
|
| +-- Trapezidens_dolichorhynchus
| +--------------+
| | +-- Lamellidens_savadiensis
| |
+--+ +-- Leoparreysia_tavoyensis
| +-----------+
| | +-- Leoparreysia_olivacea
| |
+--+ +-- Coelatura_choziensis
| +--------+
| | +-- Prisodontopsis_aviculaeformis
+--+
| +-------- Indonaia_caerulea
+--+
| +----- Indochinella_pugio
+--+
| +-- Scabies_phaselus
+--+
+-- Harmandia_somboriensis
Guerra et al., 2017 [Publ 3772], fig. 2b-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
7 |
7 |
1 |
-- |
+----------- Echyridella_menziesii **
|
--+ +-------- Cumberlandia_monodonta
| |
| | +-- Sinosolenaia_carinata
+--+ +--+
| | +-- Pronodularia_japanensis
| |
| | +-- Utterbackia_peninsularis
+--+--+
| +-- Pyganodon_grandis
|
| +-- Venustaconcha_ellipsiformis
+--+
+-- Quadrula_quadrula
Walker et al., 2006 [Publ 2666], fig. 6-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
13 |
13 |
1 |
-- |
+-------------------------- Echyridella_menziesii **
|
| +----- Cumberlandia_monodonta
| |
--+ +-----------------+ +-- Margaritifera_margaritifera
| | | |
| | +--+-- Margaritifera_dahurica__1
| | |
| | +-- Margaritifera_dahurica__2
+--+
| +-- Gonidea_angulata
| +-----------------+
| | +-- Sundadontina_tumida
| |
+--+ +-- Pyganodon_fragilis
| +--------------+
| | +-- Pyganodon_grandis
+--+
| +-------------- Fusconaia_flava
+--+
| +----------- Potamilus_purpuratus
+--+
| +-------- Cyrtonaias_tampicoensis
+--+
| +----- Ligumia_recta
+--+
| +-- Lampsilis_teres
+--+
+-- Ortmanniana_ligamentina
Wu et al., 2024 [Publ 4350], fig. 8a, S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
91 |
90 |
2 |
-- |
+----------------------------------------------- Echyridella_menziesii **
|
| +-- Anodontites_trapesialis
| +-----------------------------------------+
| | +-- Mutela_dubia
| |
| | +-- Margaritifera_dahurica
--+ | +--------------------------------------+
| | | +-- Margaritifera_margaritifera
| | |
| | | +-- Quadrula_quadrula
| | | +--------------------------+
| | | | +-- Uniomerus_tetralasmus
| | | |
+--+ | +--------+ +-- Elliptio_complanata
| | | | +-----------------------+
| | | | | +-- Pleurobema_oviforme
| | | +--+
| | | | +----------------------- Amblema_plicata
| | | | |
| | | +--+ +-------------------- Popenaias_popeii
| | | | |
| | | +--+ +----------------- Toxolasma_parvum
| | | | |
| | | | | +----- Potamilus_leptodon
+--+ | +--+ +--------+
| | | | | +-- Potamilus_alatus
| | | | +--+
| | | | +-- Potamilus_streckersoni
| | +--+
| | | +----------- Leaunio_lienosus
| | | |
| | | | +----- Lampsilis_ornata
| | +--+ +--+
| | | | | +-- Lampsilis_cardium
| | | | +--+
| | +--+ +-- Lampsilis_cariosa
| | |
| | | +----- Venustaconcha_ellipsiformis
| | +--+
| | | +-- Lampsilis_powellii
| | +--+
| | +-- Lampsilis_siliquoidea
| |
+--+ +----- Chamberlainia_somsakpanhai
| +--------------------+
| | | +-- Sinohyriopsis_cumingii
| | +--+
| | +-- Sinohyriopsis_schlegelii
| |
| | +-- Hyriopsis_bialata
| | +--+
| +--------+ | +-- Rectidens_sumatrensis
| | | +-----------------+
| | | | | +-- Lens_contradens
| | | | +--+
| | | | +-- Physunio_superbus
| | | |
| | | | +----------- Potomida_littoralis
| | +--+ +--------+
| | | | | +-------- Pronodularia_japanensis
| | | | +--+
| | | | | +----- Lamprotula_leaii
| | | | +--+
| | | | | +-- Lamprotula_caveata
| | +--+ +--+
| | | +-- Lamprotula_cornuumlunae
| | |
| | | +-- Pseudodon_vondembuschianus
| | | +--------------+
| | | | +-- Pilsbryoconcha_exilis
| | +--+
| | | +-------------- Inversidens_rentianensis
| | | |
+--+ +--+ +-- Parvasolenaia_rivularis
| | +--------+
| | | +-- Ptychorhynchus_pfisteri
| +--+
| | +-------- Microcondylaea_bonellii
| +--+
| | +----- Sinosolenaia_carinata
| +--+
| | +-- Sinosolenaia_iridinea
| +--+
| +-- Sinosolenaia_oleivora
|
| +-------------------------------- Lamellidens_jenkinsianus
| |
| | +-- Lepidodesma_aligera
| | +--------------------------+
| | | +-- Lepidodesma_languilati
| | |
| | | +-------- Aculamprotula_tortuosa
| | | |
| | | +--------------+ +-- Aculamprotula_coreana
| | | | | +--+
+--+ | | | | +-- Aculamprotula_tientsinensis
| | | +--+
| | | | +-- Aculamprotula_polysticta
| | | +--+
| | | +-- Aculamprotula_scripta
| | |
| | +--+ +-------------- Unio_tumidus
| | | | +-----+
| | | | | | +----------- Unio_nanus
| | | | | +--+
+--+ | | | | +-------- Unio_elongatulus
| | | | +--+
| | | | | +----- Unio_pictorum
| | | | +--+
| | | | | +-- Unio_delphinus
| | +--+ +--+
| | | +-- Unio_mancus
| | |
| | | +-- Nodularia_breviconcha
| | | +--------------+
| | | | +-- Nodularia_douglasiae
| | | |
| | | | +-- Pseudocuneopsis_capitata
| | | +--------------+
| | +--+ +-- Pseudocuneopsis_sichuanensis
| | |
| | | +-- Schistodesmus_lampreyanus
| | | +-----------+
+--+ | | +-- Schistodesmus_spinosus
| +--+
| | +----------- Tchangsinaia_piscicula
| +--+
| | +-------- Cuneopsis_rufescens
| +--+
| | +----- Cuneopsis_celtiformis
| +--+
| | +-- Cuneopsis_demangei
| +--+
| +-- Cuneopsis_heudei
|
| +-------- Acuticosta_chinensis
| +--------------+
| | | +----- Lanceolaria_gladiola
| | +--+
| | | +-- Lanceolaria_grayii
| | +--+
| | +-- Lanceolaria_lanceolata
| |
| | +-------------- Cristaria_plicata
| | |
+--+ +-----+ +-- Anemina_arcaeformis
| | | +--------+
| | | | +-- Anemina_euscaphys
| | +--+
| | | +-------- Beringiana_fukuharai
| | +--+
| | | +----- Sinanodonta_lucida
| | +--+
| | | +-- Sinanodonta_tumens
+--+ +--+
| +-- Sinanodonta_woodiana
|
| +-------- Pseudanodonta_complanata
| +--------+
| | | +----- Anodonta_anatina
| | +--+
| | | +-- Anodonta_cygnea
| | +--+
+--+ +-- Anodonta_exulcerata
|
| +-------------- Anodonta_nuttalliana
| |
| | +----- Pyganodon_grandis
+--+ +-----+
| | | +-- Utterbackia_imbecillis
| | +--+
+--+ +-- Utterbackia_peninsularis
|
| +-------- Lasmigona_complanata
+--+
| +----- Prolasmidonta_heterodon
+--+
| +-- Alasmidonta_varicosa
+--+
+-- Platynaias_compressa
González & Giribet, 2015 [Publ 3632], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
5 |
3 |
2 |
-- |
+-------- Aspatharia_pfeifferiana
--+
| +----- Velesunio_ambiguus **
+--+
| +-- Unio_pictorum
+--+
+-- Margaritifera_margaritifera
Guerra et al., 2017 [Publ 3772], fig. 2a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
24 |
23 |
2 |
-- |
+-- Mutela_dubia
+-----------------------------+
| +-- Anodontites_trapesialis
|
| +----------------------------- Echyridella_menziesii **
--+ |
| | +-- Margaritifera_falcata
| | +-----------------------+
| | | +-- Cumberlandia_monodonta
+--+ |
| | +-------- Quadrula_quadrula
| | +--+
| | | | +----- Toxolasma_parvum
| | | +--+
| | | | +-- Venustaconcha_ellipsiformis
+--+ | +--+
| +-----------+ +-- Lampsilis_ornata
| | |
| | | +-- Sinohyriopsis_schlegelii
| | | +-----+
| | | | +-- Sinohyriopsis_cumingii
| | +--+
| | | +----- Pronodularia_japanensis
+--+ +--+
| | +-- Sinosolenaia_oleivora
| +--+
| +-- Sinosolenaia_carinata
|
| +-- Aculamprotula_tortuosa
| +-----------------+
| | +-- Unio_pictorum
+--+
| +----------------- Sinanodonta_woodiana
| |
+--+ +-------------- Cristaria_plicata
| |
+--+ +----------- Anodonta_anatina
| |
+--+ +-- Platynaias_subviridis
| +-----+
| | +-- Platynaias_compressa
+--+
| +----- Pyganodon_grandis
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Utterbackia_imbecillis
Pfeiffer & Graf, 2013 [Publ 3531], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
33 |
33 |
1 |
-- |
+----------------------------------- Margaritifera_margaritifera
|
+----------------------------------- Velesunio_ambiguus **
|
| +-- Lamellidens_savadiensis
| +-----------------------------+
| | +-- Coelatura_aegyptiaca
| |
--+ | +-- Cristaria_plicata
| | +-----+
| | | +-- Sinanodonta_woodiana
| | +--------+
| | | | +----- Strophitus_undulatus
| | | +--+
| | | | +-- Platynaias_compressa
| | +-----------+ +--+
+--+ | | +-- Pyganodon_grandis
| | |
| | | +-------------- Acuticosta_chinensis
| | | |
| | +--+ +-- Unio_pictorum
| | | +--------+
| | | | +-- Nodularia_douglasiae
| | +--+
| | | +-------- Aculamprotula_tortuosa
| | | |
| | +--+ +----- Aculamprotula_scripta
| | | |
+--+ +--+ +-- Aculamprotula_tientsinensis
| | |
| +--+-- Aculamprotula_fibrosa
| |
| +-- Aculamprotula_zonata
|
| +----------- Quadrula_quadrula
| +--------------+
| | | +-------- Eurynia_dilatata
| | +--+
| | | +----- Amblema_plicata
| | +--+
| | | +-- Lampsilis_ornata
+--+ +--+
| +-- Truncilla_truncata
|
| +-- Sinohyriopsis_cumingii
| +--------------------+
| | +-- Sinohyriopsis_schlegelii
| |
+--+ +----- Gonidea_angulata
| +--------------+
| | | +-- Pseudodon_vagulus
| | +--+
+--+ +-- Sundadontina_tumida
|
| +----------------- Physunio
| |
+--+ +-------------- Pronodularia_japanensis
| |
+--+ +----------- Potomida_littoralis
| |
+--+ +-------- Lamprotula_leaii
| |
+--+ +----- Lamprotula_caveata__2
| |
+--+ +-- Lamprotula_cornuumlunae
| |
+--+-- Gibbosula_rochechouartii
|
+-- Lamprotula_caveata__1
Wu et al., 2021 [Publ 4366], fig. 3-BI
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
73 |
72 |
2 |
-- |
+-------------------------------------------- Echyridella_menziesii **
|
| +-- Anodontites_trapesialis
| +--------------------------------------+
| | +-- Mutela_dubia
| |
| | +-- Gibbosula_rochechouartii
--+ | +-----------+
| | | +-- Gibbosula_crassa
| | +-----------------------+
| | | | +----------- Cumberlandia_monodonta
| | | +--+
| | | | +-------- Pseudunio_marocanus
| | | +--+
| | | | +----- Margaritifera_falcata
+--+ | +--+
| | | +-- Margaritifera_margaritifera
| | +--+
| | +-- Margaritifera_dahurica
| |
| | +-- Uniomerus_tetralasmus
| | +--------------------------+
| | | +-- Quadrula_quadrula
| | |
| | +-----+ +-- Pleurobema_oviforme
| | | | +-----------------------+
| | | | | +-- Elliptio_complanata
| | | +--+
+--+ | | +----------------------- Amblema_plicata
| | | |
| | +--+ +-------------------- Popenaias_popeii
| | | |
| | +--+ +----------------- Toxolasma_parvum
| | | |
| | +--+ +-- Potamilus_alatus
| | | +-----------+
| | | | +-- Potamilus_leptodon
| | +--+
| | | +----------- Leaunio_lienosus
| | | |
| | +--+ +-- Lampsilis_ornata
| | | +-----+
| | | | +-- Lampsilis_cardium
| | +--+
| | | +----- Venustaconcha_ellipsiformis
| | +--+
| | | +-- Lampsilis_siliquoidea
| | +--+
+--+ +-- Lampsilis_powellii
|
| +----- Chamberlainia_somsakpanhai
| +-----------+
| | | +-- Sinohyriopsis_schlegelii
| | +--+
| | +-- Sinohyriopsis_cumingii
| |
| +--------------+ +-------- Potomida_littoralis
| | | +-----+
| | | | | +----- Pronodularia_japanensis
| | | | +--+
| | | | | +-- Lamprotula_caveata
| | | | +--+
| | +--+ +-- Lamprotula_leaii
| | |
| | | +-- Pilsbryoconcha_exilis
| | | +--------+
| | | | +-- Pseudodon_vondembuschianus
| | | |
| | +--+ +-- Parvasolenaia_rivularis
| | | +-----+
| | | | +-- Ptychorhynchus_pfisteri
| | +--+
| | | +----- Microcondylaea_bonellii
+--+ +--+
| | +-- Sinosolenaia_oleivora
| +--+
| +-- Sinosolenaia_carinata
|
| +----------------------------- Lepidodesma_languilati
| |
| | +-------- Aculamprotula_tortuosa
| | |
| | +--------+ +-- Aculamprotula_scripta
| | | | +--+
| | | | | +-- Aculamprotula_polysticta
| | | +--+
| | | | +-- Aculamprotula_coreana
| | | +--+
| | +--------+ +-- Aculamprotula_tientsinensis
| | | |
+--+ | | +-------- Unio_tumidus
| | | +-----+
| | | | | +----- Unio_nanus
| | | | +--+
| | | | | +-- Unio_pictorum
| | +--+ +--+
| | | +-- Unio_delphinus
| | |
| | | +----------- Pseudocuneopsis_capitata
| | | |
| | +--+ +-- Nodularia_douglasiae
| | | +-----+
| | | | +-- Nodularia_breviconcha
+--+ +--+
| | +----- Schistodesmus_lampreyanus
| +--+
| | +-- Cuneopsis_heudei
| +--+
| +-- Tchangsinaia_piscicula
|
| +-------- Acuticosta_chinensis
| +--------------+
| | | +----- Lanceolaria_gladiola
| | +--+
| | | +-- Lanceolaria_grayii
| | +--+
| | +-- Lanceolaria_lanceolata
| |
+--+ +-------- Cristaria_plicata
| |
| +-----------+ +-- Sinanodonta_woodiana
| | | +--+
| | | | +-- Sinanodonta_lucida
| | +--+
| | | +-- Anemina_euscaphys
+--+ +--+
| +-- Anemina_arcaeformis
|
| +----------------- Anodonta_cygnea
| |
+--+ +-------------- Anodonta_anatina
| |
| | +----- Pyganodon_grandis
+--+ +-----+
| | | +-- Utterbackia_peninsularis
| | +--+
+--+ +-- Utterbackia_imbecillis
|
| +-------- Prolasmidonta_heterodon
+--+
| +----- Alasmidonta_varicosa
+--+
| +-- Platynaias_subviridis
+--+
+-- Platynaias_compressa
Wu et al., 2024 [Publ 4350], fig. 8b, S2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
91 |
90 |
2 |
-- |
+----------------------------------------------- Echyridella_menziesii **
|
| +-- Anodontites_trapesialis
| +-----------------------------------------+
| | +-- Mutela_dubia
| |
| | +-- Margaritifera_dahurica
--+ | +--------------------------------------+
| | | +-- Margaritifera_margaritifera
| | |
| | | +-- Quadrula_quadrula
| | | +--------------------------+
| | | | +-- Uniomerus_tetralasmus
| | | |
+--+ | +--------+ +-- Elliptio_complanata
| | | | +-----------------------+
| | | | | +-- Pleurobema_oviforme
| | | +--+
| | | | +----------------------- Amblema_plicata
| | | | |
| | | +--+ +-------------------- Popenaias_popeii
| | | | |
| | | +--+ +----------------- Toxolasma_parvum
| | | | |
| | | | | +----- Potamilus_leptodon
+--+ | +--+ +--------+
| | | | | +-- Potamilus_alatus
| | | | +--+
| | +--+ +-- Potamilus_streckersoni
| | |
| | | +----------- Leaunio_lienosus
| | | |
| | +--+ +-------- Venustaconcha_ellipsiformis
| | | |
| | | | +-- Lampsilis_powellii
| | +--+-----+
| | | +-- Lampsilis_siliquoidea
| | |
| | | +----- Lampsilis_ornata
| | +--+
| | | +-- Lampsilis_cardium
| | +--+
| | +-- Lampsilis_cariosa
| |
+--+ +----- Chamberlainia_somsakpanhai
| +--------------------+
| | | +-- Sinohyriopsis_cumingii
| | +--+
| | +-- Sinohyriopsis_schlegelii
| |
| | +-- Hyriopsis_bialata
| | +--+
| +--------+ | +-- Rectidens_sumatrensis
| | | +-----------------+
| | | | | +-- Lens_contradens
| | | | +--+
| | | | +-- Physunio_superbus
| | | |
| | | | +----------- Potomida_littoralis
| | +--+ +--------+
| | | | | +-------- Pronodularia_japanensis
| | | | +--+
| | | | | +----- Lamprotula_leaii
| | | | +--+
| | | | | +-- Lamprotula_caveata
| | +--+ +--+
| | | +-- Lamprotula_cornuumlunae
| | |
| | | +-- Pseudodon_vondembuschianus
| | | +--------------+
| | | | +-- Pilsbryoconcha_exilis
| | +--+
| | | +-------------- Inversidens_rentianensis
| | | |
+--+ +--+ +-- Parvasolenaia_rivularis
| | +--------+
| | | +-- Ptychorhynchus_pfisteri
| +--+
| | +-------- Microcondylaea_bonellii
| +--+
| | +----- Sinosolenaia_carinata
| +--+
| | +-- Sinosolenaia_iridinea
| +--+
| +-- Sinosolenaia_oleivora
|
| +-------------------------------- Lamellidens_jenkinsianus
| |
| | +-- Lepidodesma_aligera
| | +--------------------------+
| | | +-- Lepidodesma_languilati
| | |
| | | +-------- Aculamprotula_tortuosa
| | | |
| | | +--------------+ +-- Aculamprotula_coreana
| | | | | +--+
+--+ | | | | +-- Aculamprotula_tientsinensis
| | | +--+
| | | | +-- Aculamprotula_polysticta
| | | +--+
| | | +-- Aculamprotula_scripta
| | |
| | +--+ +-------------- Unio_tumidus
| | | | +-----+
| | | | | | +----------- Unio_nanus
| | | | | +--+
+--+ | | | | +-------- Unio_elongatulus
| | | | +--+
| | | | | +----- Unio_pictorum
| | | | +--+
| | | | | +-- Unio_delphinus
| | +--+ +--+
| | | +-- Unio_mancus
| | |
| | | +-- Nodularia_breviconcha
| | | +--------------+
| | | | +-- Nodularia_douglasiae
| | | |
| | | | +-- Pseudocuneopsis_capitata
| | | +--------------+
| | +--+ +-- Pseudocuneopsis_sichuanensis
| | |
| | | +-- Schistodesmus_lampreyanus
+--+ | +-----------+
| | | +-- Schistodesmus_spinosus
| +--+
| | +----------- Tchangsinaia_piscicula
| +--+
| | +-------- Cuneopsis_rufescens
| +--+
| | +----- Cuneopsis_celtiformis
| +--+
| | +-- Cuneopsis_demangei
| +--+
| +-- Cuneopsis_heudei
|
| +-------- Acuticosta_chinensis
| +--------------+
| | | +----- Lanceolaria_gladiola
| | +--+
| | | +-- Lanceolaria_grayii
| | +--+
| | +-- Lanceolaria_lanceolata
| |
| | +-------------- Cristaria_plicata
+--+ |
| +-----+ +-- Anemina_arcaeformis
| | | +--------+
| | | | +-- Anemina_euscaphys
| | +--+
| | | +-------- Beringiana_fukuharai
| | +--+
| | | +----- Sinanodonta_lucida
| | +--+
+--+ | +-- Sinanodonta_tumens
| +--+
| +-- Sinanodonta_woodiana
|
| +----------------- Anodonta_nuttalliana
| |
| | +-------- Pseudanodonta_complanata
| | +-----+
+--+ | | +----- Anodonta_anatina
| | +--+
| | | +-- Anodonta_cygnea
| | +--+
+--+ +-- Anodonta_exulcerata
|
| +----- Pyganodon_grandis
| +-----+
| | | +-- Utterbackia_imbecillis
| | +--+
+--+ +-- Utterbackia_peninsularis
|
| +-------- Lasmigona_complanata
+--+
| +----- Prolasmidonta_heterodon
+--+
| +-- Alasmidonta_varicosa
+--+
+-- Platynaias_compressa
Wu et al., 2020 [Publ 4121], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
49 |
48 |
2 |
-- |
+-------------------------------------- Echyridella_menziesii **
|
| +-- Anodontites_trapesialis
| +--------------------------------+
| | +-- Mutela_dubia
--+ |
| | +-------- Gibbosula_rochechouartii
| | +-----------------------+
| | | | +----- Cumberlandia_monodonta
| | | +--+
+--+ | | +-- Margaritifera_falcata
| | +--+
| | +-- Margaritifera_dahurica
| |
| | +----------- Quadrula_quadrula
| | |
| | +-----------------+ +-------- Toxolasma_parvum
+--+ | | |
| | +--+ +-- Venustaconcha_ellipsiformis
| | | +--+
| | | | +-- Lampsilis_ornata
| | +--+
| | | +-- Potamilus_alatus
| | +--+
| | +-- Potamilus_leptodon
| |
| | +-- Sinohyriopsis_schlegelii
| | +--------+
+--+ | +-- Sinohyriopsis_cumingii
| |
| +--------------+ +----- Potomida_littoralis
| | | +--+
| | | | | +-- Pronodularia_japanensis
| | | | +--+
| | +--+ +-- Lamprotula_leaii
| | |
| | | +----- Ptychorhynchus_pfisteri
| | +--+
| | | +-- Sinosolenaia_oleivora
| | +--+
| | +-- Sinosolenaia_carinata
+--+
| +----------------------- Lepidodesma_languilati
| |
| | +----- Acuticosta_chinensis
| | +-----------+
| | | | +-- Lanceolaria_lanceolata
| | | +--+
| | | +-- Lanceolaria_grayii
| | |
| | +--+ +----------- Anodonta_anatina
| | | | +--+
| | | | | | +-------- Platynaias_compressa
+--+ | | | +--+
| | | | | +----- Pyganodon_grandis
| | | | +--+
| | +--+ | +-- Utterbackia_peninsularis
| | | +--+
| | | +-- Utterbackia_imbecillis
| | |
| | | +-------- Cristaria_plicata
| | | |
| | +-----+ +-- Sinanodonta_woodiana
| | | +--+
+--+ | | +-- Sinanodonta_lauta
| +--+
| | +-- Anemina_arcaeformis
| +--+
| +-- Anemina_euscaphys
|
| +-------- Aculamprotula_tortuosa
| |
| +--------+ +-- Aculamprotula_coreana
| | | +--+
| | | | +-- Aculamprotula_tientsinensis
| | +--+
| | | +-- Aculamprotula_polysticta
| | +--+
+--+ +-- Aculamprotula_scripta
|
| +-------- Unio_tumidus
| +-----+
| | | +----- Unio_nanus
| | +--+
| | | +-- Unio_delphinus
+--+ +--+
| +-- Unio_pictorum
|
| +----------- Pseudocuneopsis_capitata
+--+
| +-------- Nodularia_douglasiae
+--+
| +----- Tchangsinaia_piscicula
+--+
| +-- Cuneopsis_heudei
+--+
+-- Schistodesmus_lampreyanus
| 
