MUSSELpdb | family Iridinidae

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Page last updated
Tue 01 Jul 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Etherioidea

family IRIDINIDAE Swainson, 1840: cladomics
2 subfamilies

Cladomics Summary

number of species = 53
number of subtaxa (rank: subfamily) = 2

analyzed in 70 cladograms in 38 publications
maximum number of terminals in an analysis = 5, mean = 1.96
maximum number of subtaxa (subfamily) in an analysis = 2, mean = 1.33

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 36
maximum number of included terminals = 5, mean = 2.86
maximum number of subtaxa (subfamily) = 2, mean = 1.64
proportion of cladograms that support basic monophyly = 0.47

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 23
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (subfamily) taxon and at least one rigorous outgroup (family).
maximum number of included terminals = 5, mean = 3.35
maximum number of subtaxa (subfamily) = 2, mean = 2.0
proportion of cladograms that support rigorous monophyly = 0.17

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination

NJ tree

Walker et al., 2006 [Publ 2666], fig. 6

iT	tT	bi	riS	oT	roT	roS	iD
3	9	7	2	33	6	2	0.0

*** Top Monophyly Cladogram *** *** Top Sister-Group Cladogram ***

  +----------- Iridininae **   
--+                         
  |  +-------- Etheria      
  +--+                      
     |  +----- Aspathariinae **
     +--+                   
        |  +-- Mulleria     
        +--+                
           +-- Mycetopodidae

Huang et al., 2019 [Publ 3843], fig. S4

iT	tT	bi	riS	oT	roT	roS	iD
5	10	6	2	194	5	2	14.0

  +-------- Etheriidae      
  |                         
  +-------- Aspathariinae **   
  |                         
--+     +-- Anodontites     
  +-----+                   
  |     +-- Iridininae **      
  |                         
  |  +----- Mycetopodinae   
  +--+                      
     |  +-- Monocondylaeinae
     +--+                   
        +-- Lamproscapha

Huang et al., 2019 [Publ 3843], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
5	10	8	2	194	5	2	16.0

           +-- Anodontites     
  +--------+                   
  |        +-- Iridininae **      
  |                            
--+        +-- Etheriidae      
  |  +-----+                   
  |  |     +-- Aspathariinae **   
  +--+                         
     |  +----- Mycetopodinae   
     +--+                      
        |  +-- Monocondylaeinae
        +--+                   
           +-- Lamproscapha

Huang et al., 2019 [Publ 3843], fig. S3

iT	tT	bi	riS	oT	roT	roS	iD
5	10	8	2	192	5	2	16.0

           +-- Anodontites     
  +--------+                   
  |        +-- Iridininae **      
  |                            
--+        +-- Etheriidae      
  |  +-----+                   
  |  |     +-- Aspathariinae **   
  +--+                         
     |  +----- Mycetopodinae   
     +--+                      
        |  +-- Monocondylaeinae
        +--+                   
           +-- Lamproscapha

Hoeh et al., 2009 [Publ 2716], fig. 4

iT	tT	bi	riS	oT	roT	roS	iD
4	4	2	2	42	6	2	8.0

  +-------- Etheriidae   
--+                      
  |  +----- Mycetopodidae
  +--+                   
     |  +-- Iridininae **   
     +--+                
        +-- Aspathariinae **

Graf & Cummings, 2006 [Publ 1968], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
4	4	2	2	43	6	2	8.0

  +-------- Etheriidae   
--+                      
  |  +----- Mycetopodidae
  +--+                   
     |  +-- Aspathariinae **
     +--+                
        +-- Iridininae **

Hoeh et al., 2009 [Publ 2716], fig. 5

iT	tT	bi	riS	oT	roT	roS	iD
4	4	2	2	42	6	2	8.0

  +-------------- Pseudomulleria
--+                             
  |  +----------- Mulleria      
  +--+                          
     |  +-------- Etheria       
     +--+                       
        |  +----- Mycetopodidae 
        +--+                    
           |  +-- Iridininae **    
           +--+                 
              +-- Aspathariinae **

Hoeh et al., 2009 [Publ 2716], fig. 3

iT	tT	bi	riS	oT	roT	roS	iD
4	4	1	2	43	6	2	8.0

        +-- Chambardia **   
        |                
  +-----+-- Aspatharia **   
  |     |                
--+     +-- Iridininae **   
  |                      
  |  +----- Etheria      
  +--+                   
     |  +-- Mulleria     
     +--+                
        +-- Mycetopodidae

Wu et al., 2020 [Publ 4121], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
3	6	4	2	31	3	2	11.0

  +-------- Iridininae **   
--+                      
  |  +----- Aspathariinae **
  +--+                   
     |  +-- Etheriidae   
     +--+                
        +-- Mycetopodidae

Sano et al., 2020 [Publ 4138], fig. 4

iT	tT	bi	riS	oT	roT	roS	iD
3	6	4	2	65	3	2	11.0

  +-------- Iridininae **   
--+                      
  |  +----- Aspathariinae **
  +--+                   
     |  +-- Etheriidae   
     +--+                
        +-- Mycetopodidae

Combosch et al., 2017 [Publ 3806], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
3	6	4	2	12	3	2	13.0

  +-------- Aspathariinae **
--+                      
  |  +----- Etheriidae   
  +--+                   
     |  +-- Iridininae **   
     +--+                
        +-- Mycetopodidae

Combosch et al., 2017 [Publ 3806], fig. 1

iT	tT	bi	riS	oT	roT	roS	iD
3	6	4	2	12	3	2	13.0

  +-------- Aspathariinae **
--+                      
  |  +----- Etheriidae   
  +--+                   
     |  +-- Iridininae **   
     +--+                
        +-- Mycetopodidae

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iT	tT	bi	riS	oT	roT	roS	iD
3	6	4	2	77	3	2	11.0

  +-------- Iridininae **   
--+                      
  |  +----- Aspathariinae **
  +--+                   
     |  +-- Etheriidae   
     +--+                
        +-- Mycetopodidae

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iT	tT	bi	riS	oT	roT	roS	iD
3	5	3	2	33	2	1	12.0

  +----- Aspathariinae **
--+                   
  |  +-- Iridininae **   
  +--+                
     +-- Mycetopodidae

Sano et al., 2020 [Publ 4138], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
3	6	4	2	65	3	2	11.0

  +-------- Iridininae **   
--+                      
  |  +----- Aspathariinae **
  +--+                   
     |  +-- Etheriidae   
     +--+                
        +-- Mycetopodidae

Sano et al., 2020 [Publ 4138], fig. 3

iT	tT	bi	riS	oT	roT	roS	iD
3	6	4	2	65	3	2	11.0

  +-------- Iridininae **   
--+                      
  |  +----- Aspathariinae **
  +--+                   
     |  +-- Etheriidae   
     +--+                
        +-- Mycetopodidae

Huang et al., 2018 [Publ 3828], fig. 3_5

iT	tT	bi	riS	oT	roT	roS	iD
3	6	4	2	26	3	2	11.0

  +-------- Iridininae **   
--+                      
  |  +----- Aspathariinae **
  +--+                   
     |  +-- Etheriidae   
     +--+                
        +-- Mycetopodidae

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iT	tT	bi	riS	oT	roT	roS	iD
3	5	3	2	33	2	1	12.0

  +----- Aspathariinae **
--+                   
  |  +-- Iridininae **   
  +--+                
     +-- Mycetopodidae

Wu et al., 2020 [Publ 4121], fig. S1

iT	tT	bi	riS	oT	roT	roS	iD
3	6	4	2	31	3	2	11.0

  +-------- Iridininae **   
--+                      
  |  +----- Aspathariinae **
  +--+                   
     |  +-- Etheriidae   
     +--+                
        +-- Mycetopodidae

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

iT	tT	bi	riS	oT	roT	roS	iD
3	6	4	2	25	3	2	11.0

  +-------- Iridininae **   
--+                      
  |  +----- Aspathariinae **
  +--+                   
     |  +-- Etheriidae   
     +--+                
        +-- Mycetopodidae

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

iT	tT	bi	riS	oT	roT	roS	iD
3	6	4	2	25	3	2	11.0

  +-------- Iridininae **   
--+                      
  |  +----- Aspathariinae **
  +--+                   
     |  +-- Etheriidae   
     +--+                
        +-- Mycetopodidae

Wu et al., 2021 [Publ 4366], fig. 2_I

iT	tT	bi	riS	oT	roT	roS	iD
2	2	0	1	89	3	2	--

  +----- Aspathariinae **
--+                   
  |  +-- Etheriidae   
  +--+                
     +-- Mycetopodidae

Bolotov et al., 2017 [Publ 3761], fig. S1

iT	tT	bi	riS	oT	roT	roS	iD
2	2	0	1	137	2	2	--

  +----- Mycetopodidae
--+                   
  |  +-- Etheriidae   
  +--+                
     +-- Aspathariinae **

Wu et al., 2021 [Publ 4366], fig. 6

iT	tT	bi	riS	oT	roT	roS	iD
2	2	0	1	89	3	2	--

  +----- Aspathariinae **
--+                   
  |  +-- Etheriidae   
  +--+                
     +-- Mycetopodidae

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

iT	tT	bi	riS	oT	roT	roS	iD
2	2	0	1	32	6	2	--

  +-------- Iridininae **   
--+                      
  |  +----- Etheria      
  +--+                   
     |  +-- Mulleria     
     +--+                
        +-- Mycetopodidae

Bolotov et al., 2016 [Publ 3743], fig. 4

iT	tT	bi	riS	oT	roT	roS	iD
2	2	0	1	28	2	2	--

  +----- Mycetopodidae
--+                   
  |  +-- Etheriidae   
  +--+                
     +-- Aspathariinae **

Bolotov et al., 2016 [Publ 3743], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
2	2	0	1	28	2	2	--

  +----- Mycetopodidae
--+                   
  |  +-- Etheriidae   
  +--+                
     +-- Aspathariinae **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iT	tT	bi	riS	oT	roT	roS	iD
2	2	0	1	119	3	2	--

  +----- Mycetopodidae
--+                   
  |  +-- Etheriidae   
  +--+                
     +-- Aspathariinae **

Hoeh et al., 2001 [Publ 2652], fig. 14f2

iT	tT	bi	riS	oT	roT	roS	iD
2	2	0	1	29	3	1	--

  +-- Iridininae **   
--+                
  +-- Mycetopodidae

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

iT	tT	bi	riS	oT	roT	roS	iD
2	2	0	1	29	3	1	--

  +-- Iridininae **   
--+                
  +-- Mycetopodidae

Bolotov et al., 2017 [Publ 3761], fig. S2

iT	tT	bi	riS	oT	roT	roS	iD
2	2	0	1	137	2	2	--

  +----- Aspathariinae **
--+                   
  |  +-- Mycetopodidae
  +--+                
     +-- Etheriidae

Guerra et al., 2019 [Publ 4168], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
2	7	0	2	39	5	1	11.0

  +----- Aspathariinae **
--+                   
  |  +-- Iridininae **   
  +--+                
     +-- Mycetopodidae

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

iT	tT	bi	riS	oT	roT	roS	iD
2	2	0	1	29	3	1	--

  +-- Iridininae **   
--+                
  +-- Mycetopodidae

Guerra et al., 2019 [Publ 4168], fig. 1

iT	tT	bi	riS	oT	roT	roS	iD
2	7	0	2	39	5	1	9.0

  +----- Iridininae **   
--+                   
  |  +-- Aspathariinae **
  +--+                
     +-- Mycetopodidae

Hoeh et al., 2001 [Publ 2652], fig. 14f4

iT	tT	bi	riS	oT	roT	roS	iD
2	2	0	1	29	3	1	--

  +-- Iridininae **   
--+                
  +-- Mycetopodidae

Hoeh et al., 2002 [Publ 2653], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
2	2	0	1	29	3	1	--

  +-- Iridininae **   
--+                
  +-- Mycetopodidae

Wu et al., 2021 [Publ 4366], fig. 3-BI

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	72	1	1	--

  +-- Mycetopodidae
--+                
  +-- Iridininae **

Wu et al., 2020 [Publ 4121], fig. 3

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	48	1	1	--

  +-- Mycetopodidae
--+                
  +-- Iridininae **

Dai et al., 2025 [Publ 4440], fig. 3

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	87	1	1	--

  +-- Mycetopodidae
--+                
  +-- Iridininae **

González & Giribet, 2015 [Publ 3632], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	5	0	0	--

-- Aspatharia_pfeifferiana

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	56	2	2	--

  +----- Mycetopodidae
--+                   
  |  +-- Etheriidae   
  +--+                
     +-- Aspathariinae **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	56	2	2	--

  +----- Mycetopodidae
--+                   
  |  +-- Etheriidae   
  +--+                
     +-- Aspathariinae **

González & Giribet, 2015 [Publ 3632], fig. 3

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	5	0	0	--

-- Aspatharia_pfeifferiana

Konopleva et al., 2017 [Publ 3807], fig. 6

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	30	2	2	--

  +----- Mycetopodidae
--+                   
  |  +-- Etheriidae   
  +--+                
     +-- Aspathariinae **

Graf et al., 2015 [Publ 3619], fig. 1

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	19	2	2	--

  +----- Aspathariinae **
--+                   
  |  +-- Mycetopodidae
  +--+                
     +-- Etheriidae

Huang et al., 2019 [Publ 3843], fig. 3

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	55	1	1	--

  +-- Mycetopodidae
--+                
  +-- Iridininae **

Guerra et al., 2017 [Publ 3772], fig. 2b

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	24	1	1	--

  +-- Iridininae **   
--+                
  +-- Mycetopodidae

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	94	1	1	--

  +-- Mycetopodidae
--+                
  +-- Iridininae **

Wu et al., 2021 [Publ 4366], fig. 4

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	67	1	1	--

  +-- Iridininae **   
--+                
  +-- Mycetopodidae

Elderkin et al., 2016 [Publ 3693], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	2	1	1	--

-- Aspatharia_pfeifferiana

Bieler et al., 2014 [Publ 3622], fig. 28

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	4	0	0	--

-- Aspatharia_pfeifferiana

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	90	1	1	--

  +-- Mycetopodidae
--+                
  +-- Iridininae **

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	29	0	0	--

-- Mutela_rostrata

Wu et al., 2021 [Publ 4366], fig. 3-ML

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	72	1	1	--

  +-- Mycetopodidae
--+                
  +-- Iridininae **

Bieler et al., 2014 [Publ 3622], fig. 26_27_29_30

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	4	0	0	--

-- Aspatharia_pfeifferiana

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	90	1	1	--

  +-- Mycetopodidae
--+                
  +-- Iridininae **

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	29	0	0	--

-- Mutela_rostrata

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	39	2	2	--

  +----- Etheriidae   
--+                   
  |  +-- Mycetopodidae
  +--+                
     +-- Aspathariinae **

Hoeh et al., 1998 [Publ 2695], fig. 4

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	4	0	0	--

-- Mutela_rostrata

Huang et al., 2018 [Publ 3828], fig. 6

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	41	1	1	--

  +-- Mycetopodidae
--+                
  +-- Iridininae **

González & Giribet, 2015 [Publ 3632], fig. 6

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	5	0	0	--

-- Aspatharia_pfeifferiana

Wu et al., 2019 [Publ 3864], fig. 3

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	45	0	0	--

-- Mutela_dubia

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	94	1	1	--

  +-- Mycetopodidae
--+                
  +-- Iridininae **

Guerra et al., 2017 [Publ 3772], fig. 2a

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	24	1	1	--

  +-- Iridininae **   
--+                
  +-- Mycetopodidae

Smith, 2021 [Publ 4331], fig. 1

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	19	1	1	--

  +-- Etheriidae   
--+                
  +-- Aspathariinae **

Lemer et al., 2019 [Publ 4093], fig. 1

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	5	1	1	--

  +-- Etheriidae   
--+                
  +-- Aspathariinae **

Wu et al., 2022 [Publ 4238], fig. 3

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	72	1	1	--

  +-- Iridininae **   
--+                
  +-- Mycetopodidae

González & Giribet, 2015 [Publ 3632], fig. 4

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	5	0	0	--

-- Aspatharia_pfeifferiana

Hoeh et al., 1998 [Publ 2695], fig. 3

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	4	0	0	--

-- Mutela_rostrata

Graf, 2000 [Publ 2647], fig. 1

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	16	4	2	--

  +----- Iridininae **   
  |                   
--+  +-- Anodontitinae
  |  |                
  +--+-- Mycetopodinae
     |                
     +-- Etheriidae

Page updated: 09:55:40 Tue 01 Jul 2025

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