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Page last updated
Wed 04 Dec 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Unioninae

tribe LEPIDODESMINI Huang & Wu in Huang et al. 2019: cladomics
     1 genus

Cladomics Summary

number of species = 3
number of subtaxa (rank: genus) = 1

analyzed in 53 cladograms in 30 publications
maximum number of terminals in an analysis = 2, mean = 1.09
maximum number of subtaxa (genus) in an analysis = 1, mean = 1.0

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 5
maximum number of included terminals = 2, mean = 2.0
maximum number of subtaxa (genus) = 1, mean = 1.0
proportion of cladograms that support basic monophyly = 1.0


Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iT tT bi riS oT roT roS iD
1 1 0 1 119 68 4 --

*** Top Monophyly Cladogram *** *** Top Sister-Group Cladogram ***

  +----- Acuticostini 
  |                   
  +----- Lepidodesma **  
--+                   
  +----- Unionini     
  |                   
  |  +-- Lanceolariini
  +--+                
     +-- Anodontini   

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iT tT bi riS oT roT roS iD
2 2 0 1 90 45 4 --
  +----------- Lepidodesma **  
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolariini

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iT tT bi riS oT roT roS iD
2 2 0 1 94 45 4 --
  +----------- Lepidodesma **  
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolariini

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iT tT bi riS oT roT roS iD
2 2 0 1 90 45 4 --
  +----------- Lepidodesma **  
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolariini

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iT tT bi riS oT roT roS iD
2 2 0 1 94 45 4 --
  +----------- Lepidodesma **  
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolariini

Wu et al., 2023 [Publ 4339], fig. 5

iT tT bi riS oT roT roS iD
2 2 0 1 17 9 1 --
  +-- Lepidodesma **
--+              
  +-- Anodontini 

Dai et al., 2024 [Publ 4358], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 51 26 4 --
  +----------- Lepidodesma **  
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lanceolariini
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Anodontini   

Sano et al., 2020 [Publ 4138], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 68 35 3 --
                 +-- Buldowskia   
  +--------------+                
  |              +-- Anemina      
  |                               
--+           +----- Sinanodonta  
  |  +--------+                   
  |  |        |  +-- Pletholophus 
  |  |        +--+                
  +--+           +-- Beringiana   
     |                            
     |  +----------- Cristaria    
     |  |                         
     +--+        +-- Anodontina   
        |  +-----+                
        |  |     +-- Alasmidontina
        +--+                      
           |  +----- Lepidodesma **  
           +--+                   
              |  +-- Lanceolariini
              +--+                
                 +-- Unionini     

Sano et al., 2020 [Publ 4138], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 68 35 3 --
                 +-- Buldowskia   
  +--------------+                
  |              +-- Anemina      
  |                               
--+           +----- Pletholophus 
  |  +--------+                   
  |  |        |  +-- Beringiana   
  |  |        +--+                
  +--+           +-- Sinanodonta  
     |                            
     |  +----------- Cristaria    
     |  |                         
     +--+        +-- Anodontina   
        |  +-----+                
        |  |     +-- Alasmidontina
        +--+                      
           |  +----- Lanceolariini
           +--+                   
              |  +-- Lepidodesma **  
              +--+                
                 +-- Unionini     

Sano et al., 2017 [Publ 3799], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 32 20 4 --
  +-------------- Anodontini   
  |                            
  |           +-- Tchangsinaia 
--+  +--------+                
  |  |        +-- Nodularia    
  |  |                         
  +--+        +-- Schistodesmus
     |  +-----+                
     |  |     +-- Acuticostini 
     +--+                      
        |  +----- Inversiunio  
        +--+                   
           |  +-- Lepidodesma **  
           +--+                
              +-- Lanceolariini

Wu et al., 2021 [Publ 4366], fig. 3-ML

iT tT bi riS oT roT roS iD
1 1 0 1 73 33 4 --
  +----------- Lepidodesma **  
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolariini

Huang et al., 2019 [Publ 3843], fig. S4

iT tT bi riS oT roT roS iD
1 1 0 1 197 43 4 --
  +----- Acuticostini         
  |                           
  +----- Alasmidonta          
  |                           
  +----- Anodonta_anatina     
  |                           
  +----- Anemina              
  |                           
  +----- Anodonta_cygnea      
  |                           
  +----- Cristaria            
  |                           
  +----- Anodonta_nuttalliana 
  |                           
  +----- Lasmigona            
  |                           
  +----- Lepidodesma **          
  |                           
  +----- Pseudanodonta        
  |                           
  +----- Pyganodon            
  |                           
--+----- Simpsonaias          
  |                           
  +----- Strophitus           
  |                           
  +----- Schistodesmus        
  |                           
  +----- Utterbackia          
  |                           
  +----- Lanceolariini        
  |                           
  +----- Unio                 
  |                           
  +----- Aculamprotula        
  |                           
  |  +-- Pletholophus         
  +--+                        
  |  +-- Sinanodonta          
  |                           
  |  +-- Cuneopsis_heudei     
  |  |                        
  |  +-- Cuneopsis_celtiformis
  |  |                        
  +--+-- Tchangsinaia         
     |                        
     +-- Cuneopsis_rufescens  
     |                        
     +-- Nodularia            

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 23 23 3 --
  +-------- Anodontini   
--+                      
  |  +----- Lanceolariini
  +--+                   
     |  +-- Lepidodesma **  
     +--+                
        +-- Unionini     

Sano et al., 2020 [Publ 4138], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 68 35 3 --
                 +-- Buldowskia   
  +--------------+                
  |              +-- Anemina      
  |                               
--+           +----- Sinanodonta  
  |  +--------+                   
  |  |        |  +-- Pletholophus 
  |  |        +--+                
  +--+           +-- Beringiana   
     |                            
     |  +----------- Cristaria    
     |  |                         
     +--+        +-- Anodontina   
        |  +-----+                
        |  |     +-- Alasmidontina
        +--+                      
           |  +----- Lanceolariini
           +--+                   
              |  +-- Lepidodesma **  
              +--+                
                 +-- Unionini     

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iT tT bi riS oT roT roS iD
1 1 0 1 55 32 4 --
  +-------- Unionini     
  |                      
--+     +-- Lepidodesma **  
  |  +--+                
  |  |  +-- Lanceolariini
  +--+                   
     |  +-- Acuticostini 
     +--+                
        +-- Anodontini   

Cha et al., 2018 [Publ 3916], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 38 21 3 --
  +-------- Lepidodesma **  
--+                      
  |  +----- Unionini     
  +--+                   
     |  +-- Lanceolariini
     +--+                
        +-- Anodontini   

Dai et al., 2024 [Publ 4386], fig. 5-BI

iT tT bi riS oT roT roS iD
1 1 0 1 20 15 4 --
  +----------------- Acuticostini    
  |                                  
  +----------------- Cristariina     
  |                                  
  +----------------- Anodontina      
--+                                  
  |  +-------------- Lanceolariini   
  |  |                               
  |  |           +-- Aculamprotula   
  |  +-----------+                   
  +--+           +-- Lepidodesma **     
     |                               
     |  +----------- Unio            
     |  |                            
     +--+  +-------- Middendorffinaia
        |  |                         
        +--+  +----- Cuneopsis       
           |  |                      
           +--+----- Nodularia       
              |                      
              |  +-- Inversiunio     
              +--+                   
                 +-- Schistodesmus   

Huang et al., 2002 [Publ 2657], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 33 13 4 --
  +----------- Anodontini   
  |                         
  |     +----- Acuticostini 
--+  +--+                   
  |  |  |  +-- Nodularia    
  |  |  +--+                
  +--+     +-- Tchangsinaia 
     |                      
     |  +----- Lanceolariini
     +--+                   
        |  +-- Schistodesmus
        +--+                
           +-- Lepidodesma **  

Huang et al., 2019 [Publ 3843], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 41 27 4 --
  +-------------------------------- Anemina              
  |                                                      
--+  +----------------------------- Acuticostini         
  |  |                                                   
  +--+  +-------------------------- Sinanodonta          
     |  |                                                
     +--+  +----------------------- Aculamprotula        
        |  |                                             
        +--+                    +-- Cristaria            
           |  +-----------------+                        
           |  |                 +-- Lanceolariini        
           +--+                                          
              |  +----------------- Lepidodesma **          
              +--+                                       
                 |  +-------------- Schistodesmus        
                 +--+                                    
                    |  +----------- Nodularia            
                    +--+                                 
                       |  +-------- Cuneopsis_rufescens  
                       +--+                              
                          |  +----- Tchangsinaia         
                          +--+                           
                             |  +-- Cuneopsis_heudei     
                             +--+                        
                                +-- Cuneopsis_celtiformis

Wu et al., 2022 [Publ 4238], fig. 2d

iT tT bi riS oT roT roS iD
1 1 0 1 38 38 4 --
  +----------- Lepidodesma **  
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lanceolariini
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Anodontini   

Froufe et al., 2019 [Publ 3936], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 44 22 3 --
  +-------- Lepidodesma **  
--+                      
  |  +----- Unionini     
  +--+                   
     |  +-- Lanceolariini
     +--+                
        +-- Anodontini   

Wu et al., 2018 [Publ 3836], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 52 32 4 --
  +-------- Lepidodesma **  
  |                      
--+-------- Unionini     
  |                      
  |  +----- Anodontini   
  +--+                   
     |  +-- Acuticostini 
     +--+                
        +-- Lanceolariini

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 35 14 4 --
  +----------- Lepidodesma **  
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lanceolariini
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Anodontini   

Huang et al., 2019 [Publ 3843], fig. S3

iT tT bi riS oT roT roS iD
1 1 0 1 195 42 4 --
  +----------- Acuticostini 
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lepidodesma **  
     +--+                   
        |  +-- Lanceolariini
        +--+                
           +-- Anodontini   

Wu et al., 2021 [Publ 4366], fig. 2_I

iT tT bi riS oT roT roS iD
1 1 0 1 90 25 4 --
  +-------- Acuticostini 
  |                      
--+     +-- Lepidodesma **  
  |  +--+                
  |  |  +-- Unionini     
  +--+                   
     |  +-- Lanceolariini
     +--+                
        +-- Anodontini   

Wu et al., 2021 [Publ 4366], fig. 3-BI

iT tT bi riS oT roT roS iD
1 1 0 1 73 33 4 --
  +----------- Lepidodesma **  
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolariini

Wu et al., 2021 [Publ 4366], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 68 32 4 --
  +----------- Lepidodesma **  
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolariini

Aunins et al., 2018 [Publ 3863], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 40 26 4 --
  +----------- Lepidodesma **  
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolariini

Sano et al., 2020 [Publ 4138], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 33 21 4 --
  +----- Lanceolaria_grayii__1  
  |                             
  +----- Lanceolaria_grayii__2  
  |                             
  +----- Cristaria_plicata__1   
  |                             
  +----- Cristaria_plicata__2   
  |                             
  +----- Sinanodonta_lauta      
  |                             
  +----- Beringiana_japonica    
  |                             
  +----- Beringiana_fukuharai__1
  |                             
  +----- Beringiana_beringiana  
  |                             
  +----- Beringiana_fukuharai__2
  |                             
  +----- Beringiana_fukuharai__3
  |                             
  +----- Buldowskia_iwakawai__1 
  |                             
  +----- Buldowskia_iwakawai__2 
  |                             
  +----- Sinanodonta_tumens__1  
--+                             
  +----- Sinanodonta_tumens__2  
  |                             
  +----- Pletholophus           
  |                             
  +----- Anemina                
  |                             
  +----- Lanceolaria_lanceolata 
  |                             
  +----- Cristaria_plicata__3   
  |                             
  +----- Lanceolaria_gladiola   
  |                             
  +----- Sinanodonta_angula     
  |                             
  +----- Lepidodesma **            
  |                             
  |  +-- Aculamprotula          
  +--+                          
  |  +-- Acuticostini           
  |                             
  |  +-- Cuneopsis              
  +--+                          
  |  +-- Nodularia              
  |                             
  |  +-- Schistodesmus          
  +--+                          
     +-- Inversiunio            

Chen et al., 2023 [Publ 4302], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 39 30 4 --
  +-------- Acuticostini 
  |                      
--+  +----- Lepidodesma **  
  |  |                   
  +--+----- Unionini     
     |                   
     |  +-- Lanceolariini
     +--+                
        +-- Anodontini   

Dai et al., 2024 [Publ 4386], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 12 11 2 --
  +-------------- Unio            
  |                               
  |        +----- Aculamprotula   
--+  +-----+                      
  |  |     |  +-- Anodontini      
  |  |     +--+                   
  +--+        +-- Lepidodesma **     
     |                            
     |  +-------- Middendorffinaia
     +--+                         
        |  +----- Inversiunio     
        +--+                      
           |  +-- Cuneopsis       
           +--+                   
              +-- Nodularia       

Wu et al., 2019 [Publ 3864], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 46 26 4 --
  +----------- Lepidodesma **  
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolariini

Dai et al., 2023 [Publ 4285], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 52 16 4 --
  +----------- Lepidodesma **  
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolariini

Kim et al., 2021 [Publ 4197], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 16 15 3 --
-- Lepidodesma_languilati

Dai et al., 2024 [Publ 4386], fig. 5-ML

iT tT bi riS oT roT roS iD
1 1 0 1 20 15 4 --
  +----------- Anodontina   
  |                         
--+        +-- Acuticostini 
  |  +-----+                
  |  |     +-- Cristariina  
  +--+                      
     |  +----- Lepidodesma **  
     +--+                   
        |  +-- Lanceolariini
        +--+                
           +-- Unionini     

Huang et al., 2019 [Publ 3843], fig. S1

iT tT bi riS oT roT roS iD
1 1 0 1 44 30 4 --
  +-------------- Anemina      
  |                            
--+  +----------- Sinanodonta  
  |  |                         
  +--+  +-------- Acuticostini 
     |  |                      
     +--+     +-- Lepidodesma **  
        |  +--+                
        |  |  +-- Cristaria    
        +--+                   
           |  +-- Lanceolariini
           +--+                
              +-- Unionini     

Wu et al., 2022 [Publ 4238], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 73 35 4 --
  +----------- Lepidodesma **  
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolariini

Chen et al., 2023 [Publ 4302], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 39 30 4 --
           +-- Anemina               
  +--------+                         
  |        +-- Amuranodonta          
  |                                  
  |     +----- Pletholophus          
--+  +--+                            
  |  |  |  +-- Sinanodonta           
  |  |  +--+                         
  |  |     +-- Beringiana            
  |  |                               
  +--+  +----- Lanceolaria_gladiola  
     |  |                            
     |  +----- Acuticostini          
     |  |                            
     |  +----- Lepidodesma **           
     |  |                            
     +--+----- Lanceolaria_oxyrhyncha
        |                            
        +----- Cristaria             
        |                            
        +----- Unionini              
        |                            
        |  +-- Alasmidontina         
        +--+                         
           +-- Anodontina            

Zhou et al., 2016 [Publ 3710], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 19 8 2 --
  +----- Anodontini 
--+                 
  |  +-- Lepidodesma **
  +--+              
     +-- Unionini   

Wu et al., 2024 [Publ 4379], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 53 51 3 --
  +-------- Lepidodesma **  
--+                      
  |  +----- Unionini     
  +--+                   
     |  +-- Lanceolariini
     +--+                
        +-- Anodontini   

Dai et al., 2024 [Publ 4386], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 39 25 4 --
  +----------- Lepidodesma **  
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lanceolariini
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Anodontini   

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iT tT bi riS oT roT roS iD
1 1 0 1 48 22 3 --
  +-------- Lepidodesma **  
--+                      
  |  +----- Unionini     
  +--+                   
     |  +-- Lanceolariini
     +--+                
        +-- Anodontini   

Chen et al., 2020 [Publ 4170], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 33 14 2 --
  +-------- Anodontini   
--+                      
  |  +----- Schistodesmus
  +--+                   
     |  +-- Lepidodesma **  
     +--+                
        +-- Aculamprotula

Wu et al., 2021 [Publ 4366], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 90 25 4 --
  +----------- Acuticostini 
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lanceolariini
     +--+                   
        |  +-- Lepidodesma **  
        +--+                
           +-- Anodontini   

Wu et al., 2020 [Publ 4121], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 49 27 4 --
  +----------- Lepidodesma **  
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolariini

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 53 27 4 --
  +----------- Lepidodesma **  
--+                         
  |  +-------- Lanceolariini
  +--+                      
     |  +----- Unionini     
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Anodontini   

Huang et al., 2002 [Publ 2657], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 33 13 4 --
  +-------- Strophitus   
  |                      
  +-------- Schistodesmus
  |                      
  +-------- Lepidodesma **  
  |                      
  +-------- Acuticostini 
  |                      
  +-------- Cristariina  
--+                      
  +-------- Lanceolariini
  |                      
  |     +-- Nodularia    
  +-----+                
  |     +-- Tchangsinaia 
  |                      
  |  +----- Utterbackia  
  |  |                   
  +--+----- Utterbackiana
     |                   
     |  +-- Alasmidonta  
     +--+                
        +-- Lasmigona    

Froufe et al., 2019 [Publ 3936], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 44 22 3 --
  +-------- Lepidodesma **  
--+                      
  |  +----- Unionini     
  +--+                   
     |  +-- Lanceolariini
     +--+                
        +-- Anodontini   

Wen et al., 2017 [Publ 3795], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 15 6 2 --
  +-------- Anodontini   
--+                      
  |  +----- Aculamprotula
  +--+                   
     |  +-- Lepidodesma **  
     +--+                
        +-- Unio         

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iT tT bi riS oT roT roS iD
1 1 0 1 38 16 4 --
  +----------------- Anodontini   
  |                               
  |              +-- Acuticostini 
--+  +-----------+                
  |  |           +-- Lanceolariini
  |  |                            
  +--+           +-- Lepidodesma **  
     |  +--------+                
     |  |        +-- Aculamprotula
     +--+                         
        |  +-------- Unio         
        +--+                      
           |  +----- Schistodesmus
           +--+                   
              |  +-- Nodularia    
              +--+                
                 +-- Tchangsinaia 

Huang et al., 2019 [Publ 3843], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 197 43 4 --
  +-------- Acuticostini 
  |                      
--+  +----- Lepidodesma **  
  |  |                   
  +--+----- Unionini     
     |                   
     |  +-- Lanceolariini
     +--+                
        +-- Anodontini   

Wu et al., 2024 [Publ 4388], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 53 17 3 --
  +-------- Lepidodesma **  
--+                      
  |  +----- Unionini     
  +--+                   
     |  +-- Lanceolariini
     +--+                
        +-- Anodontini   

Huang et al., 2019 [Publ 3843], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 56 26 4 --
  +----------- Lepidodesma **  
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lanceolariini
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Anodontini   

Page updated: 14:54:26 Wed 04 Dec 2024

 
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