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Page last updated
Wed 04 Dec 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Unioninae | Unionini

genus Cuneopsis Simpson, 1900: cladomics
     8 valid species

Cladomics Summary

number of species = 8

analyzed in 55 cladograms in 34 publications
maximum number of terminals in an analysis = 4, mean = 2.04

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 34
maximum number of included terminals = 4, mean = 2.68
proportion of cladograms that support basic monophyly = 0.79


Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ tree


Wu et al., 2022 [Publ 4238], fig. 2d

iT tT bi riS oT roT roS iD
4 4 2 4 35 19 7 0.0

*** Top Monophyly Cladogram ***

  +-------------------------- Aculamprotula        
  |                                                
--+  +----------------------- Unio                 
  |  |                                             
  +--+  +-------------------- Pseudocuneopsis      
     |  |                                          
     +--+  +----------------- Nodularia            
        |  |                                       
        +--+  +-------------- Schistodesmus        
           |  |                                    
           +--+           +-- Tchangsinaia         
              |  +--------+                        
              |  |        +-- Inversiunio          
              +--+                                 
                 |  +-------- Cuneopsis_rufescens **  
                 +--+                              
                    |  +----- Cuneopsis_celtiformis **
                    +--+                           
                       |  +-- Cuneopsis_heudei **     
                       +--+                        
                          +-- Cuneopsis_demangei **   

Chen et al., 2023 [Publ 4302], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 38 15 9 --

*** Top Sister-Group Cladogram ***

  +----------------- Aculamprotula        
  |                                       
  |              +-- Middendorffinaia     
--+  +-----------+                        
  |  |           +-- Nodularia            
  |  |                                    
  |  |           +-- Cuneopsis_celtiformis **
  +--+  +--------+                        
     |  |        +-- Cuneopsis_heudei **     
     |  |                                 
     |  |        +-- Diaurora             
     +--+--------+                        
        |        +-- Schistodesmus        
        |                                 
        |  +-------- Pseudocuneopsis      
        +--+                              
           |  +----- Inversiunio          
           +--+                           
              |  +-- Tchangsinaia         
              +--+                        
                 +-- Unio                 

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iT tT bi riS oT roT roS iD
4 4 2 4 88 18 6 0.0
  +----------------------- Aculamprotula        
  |                                             
--+  +-------------------- Unio                 
  |  |                                          
  +--+  +----------------- Nodularia            
     |  |                                       
     +--+----------------- Pseudocuneopsis      
        |                                       
        |  +-------------- Schistodesmus        
        +--+                                    
           |  +----------- Tchangsinaia         
           +--+                                 
              |  +-------- Cuneopsis_rufescens **  
              +--+                              
                 |  +----- Cuneopsis_celtiformis **
                 +--+                           
                    |  +-- Cuneopsis_demangei **   
                    +--+                        
                       +-- Cuneopsis_heudei **     

Dai et al., 2024 [Publ 4358], fig. 4

iT tT bi riS oT roT roS iD
4 4 2 4 48 13 6 0.0
  +-------------------------- Aculamprotula        
--+                                                
  |  +----------------------- Unio                 
  +--+                                             
     |  +-------------------- Pseudocuneopsis      
     +--+                                          
        |  +----------------- Nodularia            
        +--+                                       
           |  +-------------- Schistodesmus        
           +--+                                    
              |  +----------- Tchangsinaia         
              +--+                                 
                 |  +-------- Cuneopsis_rufescens **  
                 +--+                              
                    |  +----- Cuneopsis_celtiformis **
                    +--+                           
                       |  +-- Cuneopsis_heudei **     
                       +--+                        
                          +-- Cuneopsis_demangei **   

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iT tT bi riS oT roT roS iD
4 4 2 4 92 18 6 0.0
  +----------------------- Aculamprotula        
  |                                             
--+  +-------------------- Unio                 
  |  |                                          
  +--+                 +-- Nodularia            
     |  +--------------+                        
     |  |              +-- Pseudocuneopsis      
     +--+                                       
        |  +-------------- Tchangsinaia         
        +--+                                    
           |  +----------- Schistodesmus        
           +--+                                 
              |  +-------- Cuneopsis_rufescens **  
              +--+                              
                 |  +----- Cuneopsis_celtiformis **
                 +--+                           
                    |  +-- Cuneopsis_demangei **   
                    +--+                        
                       +-- Cuneopsis_heudei **     

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iT tT bi riS oT roT roS iD
4 4 2 4 88 18 6 0.0
  +----------------------- Aculamprotula        
  |                                             
--+  +-------------------- Unio                 
  |  |                                          
  +--+  +----------------- Nodularia            
     |  |                                       
     +--+----------------- Pseudocuneopsis      
        |                                       
        |  +-------------- Schistodesmus        
        +--+                                    
           |  +----------- Tchangsinaia         
           +--+                                 
              |  +-------- Cuneopsis_rufescens **  
              +--+                              
                 |  +----- Cuneopsis_celtiformis **
                 +--+                           
                    |  +-- Cuneopsis_demangei **   
                    +--+                        
                       +-- Cuneopsis_heudei **     

Wu et al., 2022 [Publ 4238], fig. 3

iT tT bi riS oT roT roS iD
4 4 2 4 70 16 6 0.0
  +-------------------------- Aculamprotula        
--+                                                
  |  +----------------------- Unio                 
  +--+                                             
     |  +-------------------- Pseudocuneopsis      
     +--+                                          
        |  +----------------- Nodularia            
        +--+                                       
           |  +-------------- Schistodesmus        
           +--+                                    
              |  +----------- Tchangsinaia         
              +--+                                 
                 |  +-------- Cuneopsis_rufescens **  
                 +--+                              
                    |  +----- Cuneopsis_celtiformis **
                    +--+                           
                       |  +-- Cuneopsis_heudei **     
                       +--+                        
                          +-- Cuneopsis_demangei **   

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iT tT bi riS oT roT roS iD
4 4 2 4 92 18 6 0.0
  +-------------------------- Aculamprotula               
  |                                                       
--+  +----------------------- Unio                        
  |  |                                                    
  +--+  +-------------------- Pseudocuneopsis             
     |  |                                                 
     +--+  +----------------- Nodularia                   
        |  |                                              
        |  |           +----- Tchangsinaia                
        +--+  +--------+                                  
           |  |        |  +-- Schistodesmus_lampreyanus__2
           |  |        +--+                               
           +--+           +-- Schistodesmus_spinosus      
              |                                           
              |  +----------- Schistodesmus_lampreyanus__1
              +--+                                        
                 |  +-------- Cuneopsis_rufescens **         
                 +--+                                     
                    |  +----- Cuneopsis_celtiformis **       
                    +--+                                  
                       |  +-- Cuneopsis_demangei **          
                       +--+                               
                          +-- Cuneopsis_heudei **            

Wu et al., 2024 [Publ 4379], fig. 5

iT tT bi riS oT roT roS iD
4 4 2 4 50 25 6 0.0
  +-------------------------- Aculamprotula        
--+                                                
  |  +----------------------- Unio                 
  +--+                                             
     |  +-------------------- Pseudocuneopsis      
     +--+                                          
        |  +----------------- Nodularia            
        +--+                                       
           |  +-------------- Schistodesmus        
           +--+                                    
              |  +----------- Tchangsinaia         
              +--+                                 
                 |  +-------- Cuneopsis_rufescens **  
                 +--+                              
                    |  +----- Cuneopsis_celtiformis **
                    +--+                           
                       |  +-- Cuneopsis_demangei **   
                       +--+                        
                          +-- Cuneopsis_heudei **     

Huang et al., 2019 [Publ 3843], fig. S4

iT tT bi riS oT roT roS iD
3 7 2 3 195 15 5 6.0
  +-- Cuneopsis_heudei **     
  |                        
  +-- Cuneopsis_celtiformis **
  |                        
--+-- Tchangsinaia         
  |                        
  +-- Cuneopsis_rufescens **  
  |                        
  +-- Nodularia            

Huang et al., 2019 [Publ 3843], fig. 1

iT tT bi riS oT roT roS iD
3 4 2 3 39 11 4 2.0
  +-------------- Schistodesmus        
--+                                    
  |  +----------- Nodularia            
  +--+                                 
     |  +-------- Cuneopsis_rufescens **  
     +--+                              
        |  +----- Tchangsinaia         
        +--+                           
           |  +-- Cuneopsis_heudei **     
           +--+                        
              +-- Cuneopsis_celtiformis **

Wu et al., 2018 [Publ 3836], fig. 2

iT tT bi riS oT roT roS iD
3 3 1 3 50 11 5 1.0
  +----------------- Aculamprotula        
  |                                       
--+  +-------------- Unio                 
  |  |                                    
  +--+           +-- Nodularia            
     |  +--------+                        
     |  |        +-- Schistodesmus        
     +--+                                 
        |  +-------- Tchangsinaia         
        +--+                              
           |  +----- Cuneopsis_rufescens **  
           +--+                           
              |  +-- Cuneopsis_celtiformis **
              +--+                        
                 +-- Cuneopsis_heudei **     

Huang et al., 2019 [Publ 3843], fig. S3

iT tT bi riS oT roT roS iD
3 3 1 3 193 14 5 1.0
  +-------------------- Aculamprotula        
--+                                          
  |  +----------------- Unio                 
  +--+                                       
     |  +-------------- Nodularia            
     +--+                                    
        |  +----------- Schistodesmus        
        +--+                                 
           |  +-------- Tchangsinaia         
           +--+                              
              |  +----- Cuneopsis_rufescens **  
              +--+                           
                 |  +-- Cuneopsis_heudei **     
                 +--+                        
                    +-- Cuneopsis_celtiformis **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iT tT bi riS oT roT roS iD
3 3 1 3 117 23 7 1.0
  +-------------------- Aculamprotula        
  |                                          
--+  +----------------- Middendorffinaia     
  |  |                                       
  +--+  +-------------- Unio                 
     |  |                                    
     +--+  +----------- Nodularia            
        |  |                                 
        +--+  +-------- Schistodesmus        
           |  |                              
           |  |     +-- Tchangsinaia         
           +--+-----+                        
              |     +-- Inversiunio          
              |                              
              |  +----- Cuneopsis_rufescens **  
              +--+                           
                 |  +-- Cuneopsis_heudei **     
                 +--+                        
                    +-- Cuneopsis_celtiformis **

Huang et al., 2019 [Publ 3843], fig. S1

iT tT bi riS oT roT roS iD
3 3 1 3 42 11 4 1.0
  +----------------- Aculamprotula        
--+                                       
  |  +-------------- Nodularia            
  +--+                                    
     |  +----------- Tchangsinaia         
     +--+                                 
        |  +-------- Schistodesmus        
        +--+                              
           |  +----- Cuneopsis_rufescens **  
           +--+                           
              |  +-- Cuneopsis_heudei **     
              +--+                        
                 +-- Cuneopsis_celtiformis **

Huang et al., 2019 [Publ 3843], fig. 2

iT tT bi riS oT roT roS iD
3 3 1 3 195 15 5 1.0
  +-------------------- Aculamprotula        
--+                                          
  |  +----------------- Unio                 
  +--+                                       
     |  +-------------- Schistodesmus        
     +--+                                    
        |  +----------- Nodularia            
        +--+                                 
           |  +-------- Tchangsinaia         
           +--+                              
              |  +----- Cuneopsis_rufescens **  
              +--+                           
                 |  +-- Cuneopsis_heudei **     
                 +--+                        
                    +-- Cuneopsis_celtiformis **

Bolotov et al., 2017 [Publ 3761], fig. S1

iT tT bi riS oT roT roS iD
2 2 0 2 137 8 5 --
  +-------------- Unio               
  |                                  
--+  +----------- Aculamprotula      
  |  |                               
  +--+  +-------- Nodularia          
     |  |                            
     +--+     +-- Cuneopsis_rufescens **
        |  +--+                      
        |  |  +-- Cuneopsis_heudei **   
        +--+                         
           |  +-- Tchangsinaia       
           +--+                      
              +-- Schistodesmus      

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iT tT bi riS oT roT roS iD
2 2 0 2 54 10 5 --
  +----------------- Middendorffinaia   
--+                                     
  |  +-------------- Aculamprotula      
  +--+                                  
     |  +----------- Unio               
     +--+                               
        |  +-------- Nodularia          
        +--+                            
           |  +----- Tchangsinaia       
           +--+                         
              |  +-- Cuneopsis_rufescens **
              +--+                      
                 +-- Cuneopsis_heudei **   

Liu et al., 2023 [Publ 4310], fig. 5

iT tT bi riS oT roT roS iD
2 2 0 2 22 12 7 --
  +-------- Unio               
  |                            
  +-------- Aculamprotula      
  |                            
--+     +-- Cuneopsis_rufescens **
  |  +--+                      
  |  |  +-- Cuneopsis_heudei **   
  |  |                         
  |  |  +-- Nodularia          
  +--+--+                      
     |  +-- Schistodesmus      
     |                         
     |  +-- Tchangsinaia       
     |  |                      
     +--+-- Inversiunio        
        |                      
        +-- Pseudocuneopsis    

Wu et al., 2023 [Publ 4296], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 2 21 11 7 --
  +----------------- Unio               
  |                                     
--+  +-------------- Aculamprotula      
  |  |                                  
  |  |           +-- Cuneopsis_heudei **   
  +--+  +--------+                      
     |  |        +-- Cuneopsis_rufescens **
     |  |                               
     +--+        +-- Nodularia          
        |  +-----+                      
        |  |     +-- Schistodesmus      
        +--+                            
           |  +----- Tchangsinaia       
           +--+                         
              |  +-- Inversiunio        
              +--+                      
                 +-- Pseudocuneopsis    

Wu et al., 2021 [Publ 4366], fig. 2_I

iT tT bi riS oT roT roS iD
2 2 0 2 89 9 5 --
  +-------------- Aculamprotula      
  |                                  
--+  +----------- Unio               
  |  |                               
  +--+  +-------- Nodularia          
     |  |                            
     +--+     +-- Cuneopsis_heudei **   
        |  +--+                      
        |  |  +-- Cuneopsis_rufescens **
        +--+                         
           |  +-- Tchangsinaia       
           +--+                      
              +-- Schistodesmus      

Chen et al., 2023 [Publ 4302], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 38 15 9 --
  +----------------- Aculamprotula        
  |                                       
  |              +-- Middendorffinaia     
--+  +-----------+                        
  |  |           +-- Nodularia            
  |  |                                    
  |  |           +-- Cuneopsis_celtiformis **
  +--+  +--------+                        
     |  |        +-- Cuneopsis_heudei **     
     |  |                                 
     |  |        +-- Diaurora             
     +--+--------+                        
        |        +-- Schistodesmus        
        |                                 
        |  +-------- Pseudocuneopsis      
        +--+                              
           |  +----- Inversiunio          
           +--+                           
              |  +-- Tchangsinaia         
              +--+                        
                 +-- Unio                 

Liu et al., 2017 [Publ 3939], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 13 9 3 --
  +-------- Unio               
  |                            
--+  +----- Schistodesmus      
  |  |                         
  +--+----- Nodularia          
     |                         
     |  +-- Cuneopsis_heudei **   
     +--+                      
        +-- Cuneopsis_rufescens **

Wu et al., 2022 [Publ 4238], fig. 2e

iT tT bi riS oT roT roS iD
2 2 0 2 15 10 6 --
  +-------------------- Aculamprotula        
--+                                          
  |  +----------------- Unio                 
  +--+                                       
     |  +-------------- Pseudocuneopsis      
     +--+                                    
        |  +----------- Nodularia            
        +--+                                 
           |  +-------- Tchangsinaia         
           +--+                              
              |  +----- Schistodesmus        
              +--+                           
                 |  +-- Cuneopsis_celtiformis **
                 +--+                        
                    +-- Cuneopsis_heudei **     

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iT tT bi riS oT roT roS iD
2 2 0 2 78 9 5 --
  +----------------- Unio               
--+                                     
  |  +-------------- Aculamprotula      
  +--+                                  
     |  +----------- Nodularia          
     +--+                               
        |  +-------- Schistodesmus      
        +--+                            
           |  +----- Tchangsinaia       
           +--+                         
              |  +-- Cuneopsis_heudei **   
              +--+                      
                 +-- Cuneopsis_rufescens **

Bolotov et al., 2017 [Publ 3764], fig. S1

iT tT bi riS oT roT roS iD
2 10 0 2 123 8 5 10.0
  +-- Unio_crassus         
  |                        
  +-- Unio_pictorum        
  |                        
  +-- Unio_tumidus         
  |                        
  +-- Aculamprotula        
  |                        
  +-- Cuneopsis_heudei **     
--+                        
  +-- Tchangsinaia         
  |                        
  +-- Cuneopsis_rufescens **  
  |                        
  +-- Nodularia_douglasiae 
  |                        
  +-- Nodularia_nuxpersicae
  |                        
  +-- Schistodesmus        

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iT tT bi riS oT roT roS iD
2 3 0 2 74 8 5 3.0
  +----------------- Unio               
--+                                     
  |  +-------------- Aculamprotula      
  +--+                                  
     |  +----------- Nodularia          
     +--+                               
        |  +-------- Schistodesmus      
        +--+                            
           |  +----- Cuneopsis_heudei **   
           +--+                         
              |  +-- Tchangsinaia       
              +--+                      
                 +-- Cuneopsis_rufescens **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iT tT bi riS oT roT roS iD
2 3 0 2 74 8 5 3.0
  +----------------- Unio               
--+                                     
  |  +-------------- Aculamprotula      
  +--+                                  
     |  +----------- Nodularia          
     +--+                               
        |  +-------- Schistodesmus      
        +--+                            
           |  +----- Cuneopsis_heudei **   
           +--+                         
              |  +-- Tchangsinaia       
              +--+                      
                 +-- Cuneopsis_rufescens **

Bolotov et al., 2017 [Publ 3761], fig. S2

iT tT bi riS oT roT roS iD
2 3 0 2 137 8 5 3.0
  +----------- Unio               
  |                               
--+  +-------- Aculamprotula      
  |  |                            
  |  +-------- Schistodesmus      
  +--+                            
     +-------- Nodularia          
     |                            
     |  +----- Cuneopsis_heudei **   
     +--+                         
        |  +-- Tchangsinaia       
        +--+                      
           +-- Cuneopsis_rufescens **

Choi et al., 2020 [Publ 4169], fig. 2a

iT tT bi riS oT roT roS iD
2 2 0 2 10 10 4 --
  +----------- Nodularia          
  |                               
--+  +-------- Schistodesmus      
  |  |                            
  +--+     +-- Unio               
     |  +--+                      
     |  |  +-- Tchangsinaia       
     +--+                         
        |  +-- Cuneopsis_heudei **   
        +--+                      
           +-- Cuneopsis_rufescens **

Dai et al., 2024 [Publ 4386], fig. 6

iT tT bi riS oT roT roS iD
2 2 0 2 38 18 6 --
  +-------------------- Aculamprotula        
--+                                          
  |  +----------------- Unio                 
  +--+                                       
     |  +-------------- Pseudocuneopsis      
     +--+                                    
        |  +----------- Nodularia            
        +--+                                 
           |  +-------- Schistodesmus        
           +--+                              
              |  +----- Tchangsinaia         
              +--+                           
                 |  +-- Cuneopsis_heudei **     
                 +--+                        
                    +-- Cuneopsis_celtiformis **

Wu et al., 2021 [Publ 4366], fig. 6

iT tT bi riS oT roT roS iD
2 2 0 2 89 9 5 --
  +-------------- Aculamprotula      
  |                                  
--+  +----------- Unio               
  |  |                               
  +--+  +-------- Nodularia          
     |  |                            
     +--+     +-- Cuneopsis_heudei **   
        |  +--+                      
        |  |  +-- Cuneopsis_rufescens **
        +--+                         
           |  +-- Tchangsinaia       
           +--+                      
              +-- Schistodesmus      

Zhang et al., 2004 [Publ 2735], fig. 1

iT tT bi riS oT roT roS iD
2 6 0 2 12 5 4 6.0
  +-------------- Tchangsinaia       
--+                                  
  |  +----------- Cuneopsis_heudei **   
  +--+                               
     |  +-------- Nodularia          
     +--+                            
        |  +----- Pseudocuneopsis    
        +--+                         
           |  +-- Cuneopsis_rufescens **
           +--+                      
              +-- Schistodesmus      

Liu et al., 2023 [Publ 4310], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 2 22 12 7 --
  +----------------- Unio               
  |                                     
--+  +-------------- Aculamprotula      
  |  |                                  
  |  |           +-- Cuneopsis_heudei **   
  +--+  +--------+                      
     |  |        +-- Cuneopsis_rufescens **
     |  |                               
     +--+        +-- Nodularia          
        |  +-----+                      
        |  |     +-- Schistodesmus      
        +--+                            
           |  +----- Tchangsinaia       
           +--+                         
              |  +-- Inversiunio        
              +--+                      
                 +-- Pseudocuneopsis    

Wu et al., 2021 [Publ 4366], fig. 3-ML

iT tT bi riS oT roT roS iD
1 1 0 1 73 14 6 --
  +----------------- Aculamprotula   
--+                                  
  |  +-------------- Unio            
  +--+                               
     |  +----------- Pseudocuneopsis 
     +--+                            
        |  +-------- Nodularia       
        +--+                         
           |  +----- Tchangsinaia    
           +--+                      
              |  +-- Schistodesmus   
              +--+                   
                 +-- Cuneopsis_heudei **

Wu et al., 2020 [Publ 4147], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 28 9 6 --
  +----------------- Aculamprotula   
--+                                  
  |  +-------------- Unio            
  +--+                               
     |  +----------- Pseudocuneopsis 
     +--+                            
        |  +-------- Nodularia       
        +--+                         
           |  +----- Schistodesmus   
           +--+                      
              |  +-- Cuneopsis_heudei **
              +--+                   
                 +-- Tchangsinaia    

Ouyang et al., 2011 [Publ 3408], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 12 2 2 --
  +-- Nodularia       
--+                   
  +-- Cuneopsis_heudei **

Dai et al., 2024 [Publ 4386], fig. 5-BI

iT tT bi riS oT roT roS iD
1 1 0 1 20 10 6 --
  +----------- Unio                 
  |                                 
--+  +-------- Middendorffinaia     
  |  |                              
  +--+  +----- Cuneopsis_celtiformis **
     |  |                           
     +--+----- Nodularia            
        |                           
        |  +-- Inversiunio          
        +--+                        
           +-- Schistodesmus        

Wu et al., 2021 [Publ 4366], fig. 3-BI

iT tT bi riS oT roT roS iD
1 1 0 1 73 14 6 --
  +----------------- Aculamprotula   
--+                                  
  |  +-------------- Unio            
  +--+                               
     |  +----------- Pseudocuneopsis 
     +--+                            
        |  +-------- Nodularia       
        +--+                         
           |  +----- Schistodesmus   
           +--+                      
              |  +-- Cuneopsis_heudei **
              +--+                   
                 +-- Tchangsinaia    

Wu et al., 2021 [Publ 4366], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 68 14 6 --
  +----------------- Aculamprotula   
--+                                  
  |  +-------------- Unio            
  +--+                               
     |  +----------- Pseudocuneopsis 
     +--+                            
        |  +-------- Nodularia       
        +--+                         
           |  +----- Schistodesmus   
           +--+                      
              |  +-- Tchangsinaia    
              +--+                   
                 +-- Cuneopsis_heudei **

Aunins et al., 2018 [Publ 3863], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 40 10 5 --
  +-------------- Aculamprotula   
--+                               
  |  +----------- Unio            
  +--+                            
     |  +-------- Nodularia       
     +--+                         
        |  +----- Tchangsinaia    
        +--+                      
           |  +-- Cuneopsis_heudei **
           +--+                   
              +-- Schistodesmus   

Sano et al., 2020 [Publ 4138], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 33 6 4 --
  +-- Cuneopsis_celtiformis **
--+                        
  +-- Nodularia            

Dai et al., 2024 [Publ 4386], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 12 9 5 --
  +-------- Middendorffinaia     
--+                              
  |  +----- Inversiunio          
  +--+                           
     |  +-- Cuneopsis_celtiformis **
     +--+                        
        +-- Nodularia            

Zhou et al., 2007 [Publ 2658], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 20 6 2 --
  +-- Nodularia            
--+                        
  +-- Cuneopsis_celtiformis **

Choi et al., 2020 [Publ 4169], fig. 3a

iT tT bi riS oT roT roS iD
1 1 0 1 5 5 2 --
  +----- Nodularia       
--+                      
  |  +-- Cuneopsis_heudei **
  +--+                   
     +-- Unio            

Wu et al., 2019 [Publ 3864], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 46 12 6 --
  +----------------- Aculamprotula   
--+                                  
  |  +-------------- Unio            
  +--+                               
     |  +----------- Pseudocuneopsis 
     +--+                            
        |  +-------- Nodularia       
        +--+                         
           |  +----- Tchangsinaia    
           +--+                      
              |  +-- Cuneopsis_heudei **
              +--+                   
                 +-- Schistodesmus   

Dai et al., 2023 [Publ 4285], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 52 6 6 --
  +-------------- Aculamprotula        
  |                                    
--+  +----------- Unio                 
  |  |                                 
  +--+        +-- Pseudocuneopsis      
     |  +-----+                        
     |  |     +-- Nodularia            
     +--+                              
        |  +----- Schistodesmus        
        +--+                           
           |  +-- Cuneopsis_celtiformis **
           +--+                        
              +-- Tchangsinaia         

Wu et al., 2024 [Publ 4379], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 11 11 3 --
  +----- Nodularia            
  |                           
--+  +-- Cuneopsis_celtiformis **
  |  |                        
  +--+-- Tchangsinaia         
     |                        
     +-- Pseudocuneopsis      

Kim et al., 2021 [Publ 4197], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 16 10 4 --
  +-------- Unio            
--+                         
  |  +----- Nodularia       
  +--+                      
     |  +-- Schistodesmus   
     +--+                   
        +-- Cuneopsis_heudei **

Dai et al., 2024 [Publ 4386], fig. 5-ML

iT tT bi riS oT roT roS iD
1 1 0 1 20 10 6 --
  +----------------- Aculamprotula        
--+                                       
  |  +-------------- Unio                 
  +--+                                    
     |  +----------- Middendorffinaia     
     +--+                                 
        |  +-------- Nodularia            
        +--+                              
           |  +----- Cuneopsis_celtiformis **
           +--+                           
              |  +-- Inversiunio          
              +--+                        
                 +-- Schistodesmus        

Wang et al., 2018 [Publ 3846], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 31 7 4 --
  +----------- Aculamprotula   
--+                            
  |  +-------- Unio            
  +--+                         
     |  +----- Nodularia       
     +--+                      
        |  +-- Tchangsinaia    
        +--+                   
           +-- Cuneopsis_heudei **

Wu et al., 2020 [Publ 4121], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 49 13 6 --
  +----------------- Aculamprotula   
--+                                  
  |  +-------------- Unio            
  +--+                               
     |  +----------- Pseudocuneopsis 
     +--+                            
        |  +-------- Nodularia       
        +--+                         
           |  +----- Tchangsinaia    
           +--+                      
              |  +-- Cuneopsis_heudei **
              +--+                   
                 +-- Schistodesmus   

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 53 11 6 --
  +-------------- Aculamprotula   
  |                               
--+  +----------- Unio            
  |  |                            
  +--+        +-- Schistodesmus   
     |  +-----+                   
     |  |     +-- Cuneopsis_heudei **
     +--+                         
        |  +----- Pseudocuneopsis 
        +--+                      
           |  +-- Nodularia       
           +--+                   
              +-- Tchangsinaia    

Fukata & Iigo, 2019 [Publ 4108], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 15 11 6 --
  +----------- Unio            
  |                            
--+  +-------- Pseudocuneopsis 
  |  |                         
  +--+     +-- Schistodesmus   
     |  +--+                   
     |  |  +-- Cuneopsis_heudei **
     +--+                      
        |  +-- Tchangsinaia    
        +--+                   
           +-- Nodularia       

Wu et al., 2024 [Publ 4388], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 53 8 6 --
  +----------------- Aculamprotula   
--+                                  
  |  +-------------- Unio            
  +--+                               
     |  +----------- Pseudocuneopsis 
     +--+                            
        |  +-------- Nodularia       
        +--+                         
           |  +----- Tchangsinaia    
           +--+                      
              |  +-- Cuneopsis_heudei **
              +--+                   
                 +-- Schistodesmus   

Page updated: 14:57:32 Wed 04 Dec 2024

 
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