Obliquaria (Quadrula) metanevra Rafinesque, 1820
Rafinesque, 1820, Ann. Gén. Sci. Phys. 5 (13): 305, pl. 81, figs. 15-16 [also as ‘U. metanevra’].
Rafinesque, 1864, [in] Binney & Tryon, reprint: 50.
Type(s): NEOTYPE ANSP_20238, Ohio River.

Obliquaria (Quadrula) metanevra — Rafinesque (1820)
Unio metanever — Ravenel (1834), Reeve (1864), Lewis (1871)
Unio metanevrus — Conrad (1835), Ferussac (1835), Lapham (1860), B.H. Wright (1888), Call (1895)
Margarita (Unio) metanever — Lea (1836), Lea (1838)
Unio (Theliderma) metanevra — Swainson (1840)
Quadrula metanever — Agassiz (1852)
Margaron (Unio) metanevrus — Lea (1852), Lea (1870)
Unio metaneurus — Küster (1852), Conrad (1853)
Unio (Quadrula) metanever — Mörch (1853)
Unio (Quadrula) metanevrus — Adams & Adams (1857)
Unio (Limnium sec. Quadrula) metanever — Fischer (1887)
Quadrula (s.s.) metanevra — Simpson (1900), Simpson (1914)
Unio (s.s.) metanevra — Call (1900)
Quadrula metanevra — Ortmann (1909), Ortmann (1909), Ortmann (1910), Ortmann (1912), ..., McMurray et al. (2012), ITIS (2012), Szumowski et al. (2012), Inoue et al. (2018)
Quadrula (Orthonymus) metanevra — Frierson (1927), Haas (1930)
Quadrula (sec. Orthonymus) metanevra — F.C. Baker (1928)
Quadrula (sec. Theliderma) metaneura — Thiele (1934)
Orthonymus metanevrus metanevrus — Haas (1969)
Theliderma metanevra — Graf & Cummings (2007), Cummings & Graf (2010), Campbell & Lydeard (2012), Williams et al. (2017), ..., Graf & Cummings (2021), FMCS (2021), Neemuchwala et al. (2023), Hopper et al. (2023)

Unio tuberosus Lea, 1840
Lea, 1840, Proc. Amer. Phil. Soc. 1: 286.
Lea, 1842, Trans. Amer. Phil. Soc. 8 [O. 3]: 210 [48], pl. 14, fig. 25.
Type(s): HOLOTYPE USNM_84219, Cumberland R., Middle Tenn.

Unio tuberosus — Lea (1840), Conrad (1853), Lewis (1871), B.H. Wright (1888), de Gregorio (1914)
Margaron (Unio) tuberosus — Lea (1852), Lea (1870)
Unio (Quadrula) tuberosus — Adams & Adams (1857)
Quadrula (s.s.) tuberosa — Simpson (1900), Simpson (1914)
Quadrula intermedia — Ortmann (1918), Ortmann (1924), Ortmann (1925), Burch (1975)
Quadrula (Orthonymus) metanevra-tuberosa — Frierson (1927)
Orthonymus metanevrus tuberosus — Haas (1969)
Quadrula tuberosa — R.I. Johnson (1980), Turgeon et al. (1988), Williams et al. (1993), Turgeon et al. (1998), Cicerello & Schuster (2003), ITIS (2012)
Theliderma tuberosa — IUCN (1996), Graf & Cummings (2007), Cummings & Graf (2010)
Quadrula metanevra — Parmalee & Bogan (1998), Williams et al. (2008)
Theliderma metanevra — Williams et al. (2017), Lopes-Lima et al. (2019), Graf & Cummings (2021)

Unio wardii Lea, 1861
Lea, 1861, Proc. Acad. Nat. Sci. 13: 392.
Lea, 1862, Jour. Acad. Nat. Sci. 5 [O. 9]: 187 [9], pl. 24, fig. 257.
Type(s): HOLOTYPE USNM_84237, Mahoning R., Ohio.

Unio wardii — Lea (1861), Sowerby (1868), B.H. Wright (1888)
Margaron (Unio) wardii — Lea (1870)
Quadrula (s.s.) metanevra var. wardii — Simpson (1900), Simpson (1914)
Quadrula metanevra wardii — Utterback (1915)
Quadrula (Orthonymus) metanevra-wardii — Frierson (1927)
Orthonymus metanevrus wardii — Haas (1969)
Quadrula metanevra — Burch (1975), R.I. Johnson (1980), Parmalee & Bogan (1998), Williams et al. (2008), Watters et al. (2009)
Theliderma metanevra — Lopes-Lima et al. (2019)

Unio nodosus Barnes, 1823
Barnes, 1823, Amer. J. Sci. 6 (1-2): 124, pl. 6, figs. 7a-b.
(homonym of Unio (Eurynia) solenoides var. nodosa Rafinesque, 1820)

Unio nodosus — Barnes (1823), Hildreth (1828)
Unio metanevrus — Ferussac (1835), Conrad (1835), Call (1895)
Margarita (Unio) metanever — Lea (1836), Lea (1838)
Quadrula nodosus — Agassiz (1852)
Margaron (Unio) metanevrus — Lea (1852), Lea (1870)
Unio metaneurus — Küster (1852), Conrad (1853)
Unio (Quadrula) nodosus — Mörch (1853)
Unio metanever — Reeve (1864)
Quadrula (s.s.) metanevra — Simpson (1900), Simpson (1914)
Quadrula metanevra — Ortmann & Walker (1922), Burch (1975), Parmalee & Bogan (1998), Williams et al. (2008), Watters et al. (2009)
Quadrula (Orthonymus) metanevra — Frierson (1927), Haas (1930)
Quadrula (sec. Orthonymus) metanevra — F.C. Baker (1928)
Orthonymus metanevrus metanevrus — Haas (1969)
Theliderma metanevra — Lopes-Lima et al. (2019)

Unio metanever Lea, 1829
‘Rafinesque’ of Authors.
Lea, 1829, Trans. Amer. Phil. Soc. 3 [O. 1]: 418.
Jay, 1835, Cat. Rec. Shells: 17.
Hanley, 1843, Illustr. Descr. Cat. Rec. Bivalve Shells: 179.
Agassiz, 1852, Arch. Naturg. 18(1): 48.
Conrad, 1853, Proc. Acad. Nat. Sci. 6: 252.
Mörch, 1853, Cat. Conch. Yoldi 2: 44.
Lewis, 1871, Amer. J. Conch. 6: 218.
Lea, 1871, Proc. Acad. Nat. Sci. 23: 15 [as ‘Margarita (Unio) metanever’].
(reference to Obliquaria (Quadrula) metanevra Rafinesque, 1820)

Unio metanever — Lea (1829)
Margaron (Unio) metanevrus — Lea (1852), Lea (1870)
Quadrula metanevra — Watters et al. (2009)

Unio metaniver Catlow & Reeve, 1845
‘Lea’ Catlow & Reeve, 1845, Conch. Nomen.: 61.
(error for Obliquaria (Quadrula) metanevra Rafinesque, 1820)

Unio metaniver — Catlow & Reeve (1845)
Quadrula (s.s.) metanevra — Simpson (1900)
Quadrula metanevra — Parmalee & Bogan (1998), Watters et al. (2009)

Unio metanera Gray, 1847
Gray, 1847, Proc. Zool. Soc. London 15: 196.
(error for Obliquaria (Quadrula) metanevra Rafinesque, 1820)

Unio metanera — Gray (1847)

Unio metaneurus Herrmannsen, 1849
‘Rafinesque’ of Authors.
Herrmannsen, 1849, Ind. Gener. Malacol. 2: 571.
Küster, 1852, Conchyl. Cab. 9(2): 50.
Conrad, 1853, Proc. Acad. Nat. Sci. 6: 252.
Thiele, 1934, Handb. Syst. Weichtierk. 3: 826 [1260] [as ‘Quadrula (Theliderma) metaneura’].
(error for Obliquaria (Quadrula) metanevra Rafinesque, 1820)

Unio metaneurus — Herrmannsen (1849)
Margaron (Unio) metanevrus — Lea (1870)
Quadrula (s.s.) metanevra — Simpson (1900), Simpson (1914)
Quadrula metanevra — Parmalee & Bogan (1998), Watters et al. (2009)

Unio rugosus Lea, 1852
Lea, 1852, Synopsis Fam. Naiades: 21, footnote.
(misidentified reference to Unio rugosus Barnes, 1823)

Unio rugosus — Lea (1852)
Quadrula (Orthonymus) metanevra-wardii — Frierson (1927)

Quadrula quadrula Lydeard et al., 2000
‘Rafinesque’ of Authors.
Lydeard et al., 2000, Geol. Soc. London Spec. Publ. 177: 148. [in part]
(misidentified reference to Obliquaria (Quadrula) quadrula Rafinesque, 1820)

Quadrula quadrula — Lydeard et al. (2000)
Theliderma metanevra — Lopes-Lima et al. (2019)

Quadrula nobilis Campbell et al., 2005
‘Conrad’ of Authors.
Campbell et al., 2005, Invert. Biol. 124(2): 155. [in part]
Campbell & Lydeard, 2012, Amer. Malac. Bull. 30(1): 23. [in part]
(misidentified reference to Unio nobilis Conrad, 1854)

Quadrula nobilis — Campbell et al. (2005)
Theliderma metanevra — Lopes-Lima et al. (2019)

Taxonomic History (since 1900)

Quadrula (s.s.) metanevra — Simpson (1900), Simpson (1914)
Unio (s.s.) metanevra — Call (1900)
Quadrula (s.s.) tuberosa — Simpson (1900), Simpson (1914)
Quadrula (s.s.) metanevra var. wardii — Simpson (1900), Simpson (1914)
Quadrula metanevra — Ortmann (1909), Ortmann (1909), Ortmann (1910), Ortmann (1912), ..., McMurray et al. (2012), ITIS (2012), Szumowski et al. (2012), Inoue et al. (2018)
Unio tuberosus — de Gregorio (1914)
Quadrula metanevra wardii — Utterback (1915)
Quadrula intermedia — Ortmann (1918), Ortmann (1924), Ortmann (1925), Burch (1975)
Quadrula (Orthonymus) metanevra — Frierson (1927), Haas (1930)
Quadrula (Orthonymus) metanevra-tuberosa — Frierson (1927)
Quadrula (Orthonymus) metanevra-wardii — Frierson (1927)
Quadrula (sec. Orthonymus) metanevra — F.C. Baker (1928)
Quadrula (sec. Theliderma) metaneura — Thiele (1934)
Orthonymus metanevrus metanevrus — Haas (1969)
Orthonymus metanevrus tuberosus — Haas (1969)
Orthonymus metanevrus wardii — Haas (1969)
Quadrula tuberosa — R.I. Johnson (1980), Turgeon et al. (1988), Williams et al. (1993), Turgeon et al. (1998), Cicerello & Schuster (2003), ITIS (2012)
Theliderma tuberosa — IUCN (1996), Graf & Cummings (2007), Cummings & Graf (2010)
Quadrula quadrula — Lydeard et al. (2000)
Quadrula nobilis — Campbell et al. (2005)
Theliderma metanevra — Graf & Cummings (2007), Cummings & Graf (2010), Campbell & Lydeard (2012), Williams et al. (2017), ..., Graf & Cummings (2021), FMCS (2021), Neemuchwala et al. (2023), Hopper et al. (2023)