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Page last updated
Wed 04 Dec 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

The nominal species (available and otherwise) in this synonymy are listed by usage rather than strictly by publication date. The primary sorting order is by the most recent year (since 1969) that the species name was treated as valid, and then nominal species are sorted secondarily by publication year.

The taxonomic history of the valid species (since 1900) is listed at the bottom of the page.

Search the database pages:

Bivalvia | Unionoida | Etherioidea | Iridinidae | Iridininae | Mutela

species Mutela dubia (Gmelin, 1791): nominal species
     334 specimens available.

Mytilus dubius Gmelin, 1791
Adanson, 1757, Hist. Nat. Sénégal, Coquillages: 234, pl. 17, fig. 21 [as ‘le mutel’]. [pre-linnaean]
Gmelin, 1791, Syst. Nat. 1(6): 3363, sp. 56.
(renamed Anodonta adansonii Gray, 1825)
Type(s): HOLOTYPE MNHN_UNCATALOGED, “lacs d’eau douce de l’intérieur du Sénégal”.

Mytilus dubiusGmelin (1791), Dillwyn (1817), Wood (1825)
Anodonta dubiaBosc (1801)
Anodonta adansoniiGray (1825)
Margarita (Dipsas) plicatusLea (1836), Lea (1838)
Iridina niloticaLea (1836), Sowerby (1868)
Platiris (Spatha) niloticaLea (1838)
Unio bialataHanley (1843)
Platiris (Spatha) dubiaLea (1852), Lea (1870)
Mutela dubiaMörch (1853), Thiele (1934), Modell (1942), Franc (1949), ..., Graf & Cummings (2021), Wu et al. (2022), Wu et al. (2023), Wu et al. (2024)
Mutela dubiusAdams & Adams (1857), Lévèque (1967), Kenmuir (1980)
Mutela (s.s.) dubiaFischer (1887), Simpson (1900), Kobelt (1910), Simpson (1914), Pilsbry & Bequaert (1927), Haas (1969), Haas (1969)
Iridina dubiusAncey (1888)
Mutela niloticaWesterlund (1890)
Mutela dubia dubiaMandahl-Barth (1988), Daget (1998), Graf & Cummings (2007)
Mutela dubia niloticaIbrahim et al. (1999)

Iridina nilotica Cailliaud, 1823
Cailliaud, 1823, Voyage à Meroe, Atlas 2: pl. 60, fig. 11.
Sowerby, 1824, Zool. Jour. 1: 53, pl. 2.
Type(s): SYNTYPE MNHN_MP_0280, Bahr Yousef, Egypte.

Iridina niloticaCailliaud (1823), Wood (1828), Lea (1834), Lea (1836), Hanley (1843), Sowerby (1868)
Iridina exoticaDeshayes (1832)
Platiris (Spatha) niloticaLea (1838), Sowerby (1839)
Calliscapha niloticaSwainson (1840)
Platiris (Spatha) dubiaLea (1852), Lea (1870)
Mutela (Calliscapha) niloticaAdams & Adams (1857), Martens (1897)
Spatha (Mutela) niloticaMartens (1866)
Mutela niloticaJickeli (1874), Servain (1890), Westerlund (1890), Pallary (1903), Pallary (1909), Haas (1929), Graf (2000)
Mutela exoticaE.A. Smith (1880), E.A. Smith (1881), E.A. Smith (1904)
Mutela (s.s.) niloticaSimpson (1900), Kobelt (1910), Simpson (1914), Pilsbry & Bequaert (1927), Haas (1969)
Mutela (Mutelina) niloticaKobelt (1904)
Mutela dubiaVan Damme (1984), Graf & Cummings (2011)
Mutela dubia niloticaMandahl-Barth (1988), Daget (1998), Ibrahim et al. (1999), Graf & Cummings (2007), IUCN (2010)

Mutela nilotica var. emini Martens, 1897
Martens, 1897, Besch. Weich. Deutsch Ost Afr. 4: 253.
(intro'd as intra-sp. of Iridina nilotica Cailliaud, 1823)
Type(s): SYNTYPES ZMB_108807, Kassenye, Albert Nyanza.

Mutela nilotica var. eminiMartens (1897)
Mutela (Calliscapha) nilotica var. eminiMartens (1897)
Mutela (s.s.) nilotica var. eminiSimpson (1900), Kobelt (1910), Simpson (1914)
Mutela (s.s.) eminiPilsbry & Bequaert (1927)
Mutela dubia var. eminiFranc (1949)
Mutela (s.s.) niloticaHaas (1969)
Mutela dubiaVan Damme (1984), Graf & Cummings (2011)
Mutela dubia eminiMandahl-Barth (1988), Daget (1998), Graf & Cummings (2007), IUCN (2010)

Mutela (s.s.) garambae Pilsbry & Bequaert, 1927
Pilsbry & Bequaert, 1927, Bull. AMNH 53: 431, pl. 40, figs. 1-2.
Type(s): HOLOTYPE (ORIG. DESIG.) AMNH_72067, Middle Garambe River, Belgian Congo (Zaire), Africa.

Mutela (s.s.) garambaePilsbry & Bequaert (1927)
Mutela niloticaHaas (1929)
Mutela (s.s.) dubiaHaas (1969)
Mutela dubiaVan Damme (1984), Graf & Cummings (2011)
Mutela dubia garambaeMandahl-Barth (1988), Daget (1998), Graf & Cummings (2007), IUCN (2010)

Iridina exotica Lamarck, 1819
Bruguière, 1797, Encycl. Méth. 2: pl. 204, figs. 1a-b [as Anodontites sp.].
Lamarck, 1819, Hist. Natur. Anim. sans Vert. 6: 89, sp. 1.

Iridina exoticaLamarck (1819), Children (1823), Bory de St. Vincent (1827), Deshayes (1832), ..., Bourguignat (1886), Ancey (1888), Modell (1942), Graf (2000)
Anodonta exoticaBlainville (1825)
Anodontes exoticaGriffith & Pidgeon (1834)
Platiris (Iridina) exoticaLea (1838), Sowerby (1839), Lea (1852), Lea (1870)
Iridina striataSwainson (1840)
Mutela dubiaMörch (1853), Van Damme (1984)
Mutela exoticaE.A. Smith (1880), E.A. Smith (1881), Fischer (1887), E.A. Smith (1904)
Mutela (s.s.) exoticaSimpson (1900), Kobelt (1904), Kobelt (1910), Simpson (1914)
Iridina (s.s.) exoticaPilsbry & Bequaert (1927), Thiele (1934), Haas (1969), Haas (1969)
Mutela dubia var. exoticaFranc (1949)
Mutela dubia dubiaMandahl-Barth (1988), Daget (1998)

Mutelina singularis Pallary, 1924
Pallary, 1924, Mém. Inst. Égypte 7 (n.s.): 53, pl. 4, fig. 17.
Type(s): UNKNOWN UMMZ_111999, Canal Mahmoudich, Egypt.

Mutelina singularisPallary (1924)
Mutela (s.s.) singularisPilsbry & Bequaert (1927)
Mutela (s.s.) niloticaHaas (1969)
Mutela dubiaVan Damme (1984), Graf & Cummings (2011)
Mutela dubia niloticaMandahl-Barth (1988), Daget (1998), Ibrahim et al. (1999), Graf & Cummings (2007)
Mutela singularisIbrahim et al. (1999)

Scapha elongata Humphrey, 1797
Humphrey, 1797, Mus. Col.: 59. [nomen nudum]

Scapha elongataHumphrey (1797)

Iridina elongata Sowerby & Sowerby, 1821
Sowerby & Sowerby, 1821, Genera of Shells 2: 40, pl. 90.
Sowerby, 1839, Conch. Man.: 54, fig. 150.

Iridina elongataSowerby & Sowerby (1821), Wood (1828), Sowerby (1839), Swainson (1840), Hanley (1843), Sowerby (1868)
Iridina exoticaLea (1836)
Platiris (Spatha) elongataLea (1838)
Platiris (Iridina) exoticaLea (1852), Lea (1870)
Mutela exoticaE.A. Smith (1881), E.A. Smith (1904)
Mutela (s.s.) exoticaSimpson (1900), Simpson (1914)
Mutela (s.s.) exotica var. elongataKobelt (1910)
Iridina (s.s.) exoticaPilsbry & Bequaert (1927), Haas (1969)
Mutela dubiaVan Damme (1984)
Mutela dubia dubiaMandahl-Barth (1988), Daget (1998)

Iridina striata Swainson, 1823
Swainson, 1823, Philos. Mag. & J. 61: 112.
Oken, 1834, Isis (Oken) 27(5): 458.

Iridina striataSwainson (1823), Swainson (1840)
Iridina exoticaLea (1836)
Platiris (Iridina) exoticaLea (1838), Lea (1852), Lea (1870)
Mutela (s.s.) exoticaSimpson (1900), Simpson (1914)
Iridina (s.s.) exoticaPilsbry & Bequaert (1927), Haas (1969)
Mutela dubiaVan Damme (1984)
Mutela dubia dubiaMandahl-Barth (1988), Daget (1998)

Anodonta adansonii Gray, 1825
Gray, 1825, Ann. Phil. 9: 138.
(new name for Mytilus dubius Gmelin, 1791)

Anodonta adansoniiGray (1825)
Mutela dubiaMörch (1853)

Anodonta purpurea Deshayes, 1826
Deshayes, 1826, Nouv. Bull. Sci. 1826: 61.
Férussac, 1826, Bull. Sci. Nat. et Géol. 9: 250.

Anodonta purpureaDeshayes (1826)

Mutela oudnoei Koenig, 1826
Koenig, 1826, [in] Denham & Clapperton, Narr. Travels Africa, Appendix: 254.

Mutela oudnoeiKoenig (1826)
Platiris (Spatha) niloticaLea (1838)
Platiris (Spatha) dubiaLea (1852), Lea (1870)
Iridina niloticaSowerby (1868)
Mutela (s.s.) oudnoeiPilsbry & Bequaert (1927)
Mutela (s.s.) dubiaHaas (1969)

Iridina mutel Rang, 1835
Adanson, 1757, Hist. Nat. Sénégal, Coquillages: 234, pl. 17, fig. 21 [as ‘le mutel’]. [pre-linnaean]
Rang, 1835, Nouv. Ann. Mus. 4(3): 314.

Iridina mutelRang (1835)
Iridina exoticaDeshayes (1832)
Platiris (Spatha) niloticaLea (1838)
Platiris (Spatha) dubiaLea (1852)
Mutela mutelJousseaume (1886)
Mutela (s.s.) dubiaPilsbry & Bequaert (1927), Haas (1969)
Mutela dubia dubiaDaget (1998)

Spatha cailliaudi Martens, 1866
‘Parr.’ Martens, 1866, Malak. Blatt. 13: 10.
(intro'd as syn of Iridina nilotica Cailliaud, 1823)

Spatha cailliaudiMartens (1866)
Spatha (Mutela) niloticaMartens (1866)
Mutela niloticaJickeli (1874)

Spatha (Mutela) plicata Martens, 1866
‘Parreyss in collect.’ Martens, 1866, Malak. Blatt. 13: 10.

Spatha (Mutela) plicataMartens (1866)
Mutela plicataJickeli (1874), Servain (1890), Westerlund (1890), Pallary (1909)
Mutela (s.s.) plicataPilsbry & Bequaert (1927)
Mutela (s.s.) niloticaHaas (1969)
Mutela dubiaVan Damme (1984)
Mutela dubia niloticaMandahl-Barth (1988), Daget (1998), Ibrahim et al. (1999)

Iridina angustata Sowerby, 1868
Sowerby, 1868, Conch. Icon. 16: pl. 2, sp. 5.

Iridina angustataSowerby (1868)
Platiris (Spatha) dubiaLea (1870)
Mutela angustataJickeli (1874), Servain (1890), Westerlund (1890)
Mutela (s.s.) angustataSimpson (1900), Kobelt (1910), Pilsbry & Bequaert (1927)
Mutela nilotica var. angustataPallary (1909)
Mutela (s.s.) dubiaSimpson (1914), Haas (1969)
Mutela niloticaHaas (1929)
Mutela dubia var. angustataFranc (1949)
Mutela dubiaVan Damme (1984)
Mutela dubia dubiaMandahl-Barth (1988), Daget (1998)
Mutela dubia niloticaIbrahim et al. (1999)

Mycetopus plicatus Sowerby, 1868
‘Gray?, MS” Sowerby, 1868, Conch. Icon. 16: pl. 2, sp. 3.
(homonym of Spatha (Mutela) plicata Martens, 1866)
Type(s): MANUSCRIPT NAME BMNH_46-7-24-18, Syene, Egypt.

Mycetopus plicatusSowerby (1868)
Platiris (Mycetopus) plicatusLea (1870)
Mutela plicataJickeli (1874), Pallary (1909), Pallary (1909)
Mutela (Pseudomutela) plicataSimpson (1900), Kobelt (1910), Simpson (1914)
Mutela niloticaPallary (1903)
Pseudomutela plicataPallary (1924)
Mutela (s.s.) plicataPilsbry & Bequaert (1927)
Mutela dubia niloticaDaget (1998)

Mutela adansoni Jousseaume, 1886
‘Bourguignat’ Jousseaume, 1886, Bull. Soc. Zool. France 11: 488.
(reference to Anodonta adansonii Gray, 1825)

Mutela adansoniJousseaume (1886)
Mutela (s.s.) dubiaPilsbry & Bequaert (1927)

Mutela chambardi Servain, 1890
Servain, 1890, Bull. Soc. Malac. France 7: 303. [nomen nudum]

Mutela chambardiServain (1890)

Mutela conica Servain, 1890
Servain, 1890, Bull. Soc. Malac. France 7: 303. [nomen nudum]

Mutela conicaServain (1890)

Mutela letourneuxi Servain, 1890
Servain, 1890, Bull. Soc. Malac. France 7: 303. [nomen nudum]

Mutela letourneuxiServain (1890)

Mutela saulcyi Servain, 1890
Servain, 1890, Bull. Soc. Malac. France 7: 303. [nomen nudum]

Mutela saulcyiServain (1890)

Mutela elongata Morgan, 1897
‘Sowerby’ Morgan, 1897, Recherch. origin. Egypte II: 99.
(reference to Iridina elongata Sowerby & Sowerby, 1821)

Mutela elongataMorgan (1897)
Mutela dubia niloticaDaget (1998)

Mutela chevalieri Germain, 1904
Germain, 1904, Bull. Mus. Hist. Nat. 10: 469.
Germain, 1907, L’Afrique Centrale Française: 566, lithograph, fig. 1.
Type(s): HOLOTYPE MNHN_MP_0203, Haut Oubangui.

Mutela chevalieriGermain (1904)
Mutela (s.s.) chevalieriKobelt (1910), Pilsbry & Bequaert (1927)
Mutela (s.s.) dubiaHaas (1969)
Mutela dubiaVan Damme (1984), IUCN (2010), Graf & Cummings (2011)
Mutela dubia dubiaMandahl-Barth (1988), Daget (1998)

Mutela angustata var. curta Germain, 1906
Germain, 1906, Bull. Mus. Hist. Nat. 12: 174.
(intro'd as intra-sp. of Iridina angustata Sowerby, 1868)
Type(s): SYNTYPES MNHN_MP_0197, Lac Tchad.

Mutela angustata var. curtaGermain (1906)
Mutela (s.s.) dubia var. curtaSimpson (1914)
Mutela (s.s.) angustata var. curtaPilsbry & Bequaert (1927)
Mutela (s.s.) dubiaHaas (1969)
Mutela dubiaVan Damme (1984)
Mutela dubia dubiaDaget (1998)
Mutela dubia niloticaIbrahim et al. (1999)

Mutela angustata var. ponderosa Germain, 1906
Germain, 1906, Bull. Mus. Hist. Nat. 12: 56, Abb. 1.
(intro'd as intra-sp. of Iridina angustata Sowerby, 1868)
Type(s): SYNTYPES MNHN_MP_0292, Lac Chad.

Mutela angustata var. ponderosaGermain (1906)
Mutela (s.s.) dubia var. ponderosaSimpson (1914)
Mutela (s.s.) angustata var. ponderosaPilsbry & Bequaert (1927)
Mutela (s.s.) dubiaHaas (1969)
Mutela dubiaVan Damme (1984)
Mutela dubia dubiaDaget (1998)

Pliodon (Cameronia) hardeleti Germain, 1906
Germain, 1906, Bull. Mus. Hist. Nat. 12: 57, fig. 2.

Pliodon (Cameronia) hardeletiGermain (1906)
Pleiodon (Cameronia) hardeletiSimpson (1914)
Iridina (s.s.) hardeletiPilsbry & Bequaert (1927)
Iridina (s.s.) exoticaHaas (1969)
Mutela dubiaVan Damme (1984)
Mutela dubia dubiaMandahl-Barth (1988), Daget (1998)

Pliodon (Cameronia) tchadiensis Germain, 1906
Germain, 1906, Bull. Mus. Hist. Nat. 12: 60, fig. 4.
Type(s): HOLOTYPE MNHN_MP_0301, Lac Tchad.

Pliodon (Cameronia) tchadiensisGermain (1906)
Pleiodon (Cameronia) tchadiensisSimpson (1914)
Iridina (s.s.) tchadiensisPilsbry & Bequaert (1927)
Iridina (s.s.) exoticaHaas (1969)
Mutela dubiaVan Damme (1984), Graf & Cummings (2011)
Mutela dubia dubiaMandahl-Barth (1988), Daget (1998)

Pliodon (Cameronia) hardeleti var. molli Germain, 1906
Germain, 1906, Bull. Mus. Hist. Nat. 12: 58, Abb. 3.
(intro'd as intra-sp. of Pliodon (Cameronia) hardeleti Germain, 1906)
Type(s): HOLOTYPE MNHN_MP_0225, Lac Tchad.

Pliodon (Cameronia) hardeleti var. molliGermain (1906)
Pleiodon (Cameronia) hardeleti var. molliSimpson (1914)
Iridina (s.s.) hardeleti var. molliPilsbry & Bequaert (1927)
Iridina (s.s.) exoticaHaas (1969)
Mutela dubia dubiaDaget (1998)

Mutelina falemeensis Germain, 1907
Germain, 1907, Bull. Mus. Hist. Nat. 13: 67.
Germain, 1907, L’Afrique Centrale Française: 569, fig. 96.
Type(s): HOLOTYPE MNHN_MP_0245, Fleuve Faleme, Senegal.

Mutelina falemeensisGermain (1907)
Mutela (s.s.) falemeensisSimpson (1914), Pilsbry & Bequaert (1927)
Mutela (s.s.) dubiaHaas (1969)
Mutela rostrataVan Damme (1984), Mandahl-Barth (1988), Ibrahim et al. (1999)
Mutela dubia dubiaDaget (1998)
Mutela dubiaGraf & Cummings (2011)

Mutelina mabillei [sic] var. frasi Germain, 1907
Germain, 1907, Bull. Mus. Hist. Nat. 13: 68.
(intro'd as intra-sp. of Mutelina mabilli Rochebrune, 1886)

Mutelina mabillei var. frasiGermain (1907)
Mutelina mabillei variété frasiGermain (1907)
Mutela (s.s.) mabilli var. frasiSimpson (1914), Pilsbry & Bequaert (1927)
Mutela (s.s.) dubiaHaas (1969)
Mutela rostrataVan Damme (1984)
Mutela dubia dubiaDaget (1998)

Mutela angustata mut. grisea Germain, 1917
Germain, 1917, Bull. Mus. Hist. Nat. 23: 520.
(intro'd as intra-sp. of Iridina angustata Sowerby, 1868)

Mutela angustata mut. griseaGermain (1917)
Mutela dubia dubiaDaget (1998)

Mutela (s.s.) garambae praetenuis Pilsbry & Bequaert, 1927
Pilsbry & Bequaert, 1927, Bull. AMNH 53: 432, pl. 40, fig. 3 [as ‘prætenuis’].
(intro'd as intra-sp. of Mutela (s.s.) garambae Pilsbry & Bequaert, 1927)
Type(s): HOLOTYPE (ORIG. DESIG.) AMNH_72066, Aba River, Africa.

Mutela (s.s.) garambae praetenuisPilsbry & Bequaert (1927)
Mutela niloticaHaas (1929)
Mutela (s.s.) dubiaHaas (1969)
Mutela dubia garambaeMandahl-Barth (1988), Daget (1998)

Iridina tschadiensis Jaeckel, 1972
‘Germain’ Jaeckel, 1972, Zeitschr. f. Geomorphol. 16: 225.
(error for Pliodon (Cameronia) tchadiensis Germain, 1906)

Iridina tschadiensisJaeckel (1972)
Mutela dubia dubiaDaget (1998)

Mutela voltae Odei, 1974
Odei, 1974, Rev. Zool. Afr. 88(2): 446, figs. 1-4.
Type(s): PARATYPES MRAC_799163, Ghana: lac Volta, Zikpo.

Mutela voltaeOdei (1974)
Mutela dubia dubiaDaget (1998)

Taxonomic History (since 1900)

Mutela (s.s.) dubiaSimpson (1900), Kobelt (1910), Simpson (1914), Pilsbry & Bequaert (1927), Haas (1969), Haas (1969)
Mutela (s.s.) exoticaSimpson (1900), Kobelt (1904), Kobelt (1910), Simpson (1914)
Mutela (s.s.) niloticaSimpson (1900), Kobelt (1910), Simpson (1914), Pilsbry & Bequaert (1927), Haas (1969)
Mutela (s.s.) angustataSimpson (1900), Kobelt (1910), Pilsbry & Bequaert (1927)
Mutela (Pseudomutela) plicataSimpson (1900), Kobelt (1910), Simpson (1914)
Mutela (s.s.) nilotica var. eminiSimpson (1900), Kobelt (1910), Simpson (1914)
Mutela niloticaPallary (1903), Pallary (1909), Haas (1929), Graf (2000)
Mutela exoticaE.A. Smith (1904)
Mutela (Mutelina) niloticaKobelt (1904)
Mutela chevalieriGermain (1904)
Mutela angustata var. curtaGermain (1906)
Mutela angustata var. ponderosaGermain (1906)
Pliodon (Cameronia) hardeletiGermain (1906)
Pliodon (Cameronia) tchadiensisGermain (1906)
Pliodon (Cameronia) hardeleti var. molliGermain (1906)
Mutelina falemeensisGermain (1907)
Mutelina mabillei var. frasiGermain (1907)
Mutelina mabillei variété frasiGermain (1907)
Mutela plicataPallary (1909), Pallary (1909)
Mutela nilotica var. angustataPallary (1909)
Mutela (s.s.) exotica var. elongataKobelt (1910)
Mutela (s.s.) chevalieriKobelt (1910), Pilsbry & Bequaert (1927)
Mutela (s.s.) dubia var. curtaSimpson (1914)
Mutela (s.s.) dubia var. ponderosaSimpson (1914)
Pleiodon (Cameronia) hardeletiSimpson (1914)
Pleiodon (Cameronia) tchadiensisSimpson (1914)
Pleiodon (Cameronia) hardeleti var. molliSimpson (1914)
Mutela (s.s.) falemeensisSimpson (1914), Pilsbry & Bequaert (1927)
Mutela (s.s.) mabilli var. frasiSimpson (1914), Pilsbry & Bequaert (1927)
Mutela angustata mut. griseaGermain (1917)
Pseudomutela plicataPallary (1924)
Mutelina singularisPallary (1924)
Iridina (s.s.) exoticaPilsbry & Bequaert (1927), Thiele (1934), Haas (1969), Haas (1969)
Mutela (s.s.) oudnoeiPilsbry & Bequaert (1927)
Mutela (s.s.) plicataPilsbry & Bequaert (1927)
Mutela (s.s.) eminiPilsbry & Bequaert (1927)
Mutela (s.s.) angustata var. curtaPilsbry & Bequaert (1927)
Mutela (s.s.) angustata var. ponderosaPilsbry & Bequaert (1927)
Iridina (s.s.) hardeletiPilsbry & Bequaert (1927)
Iridina (s.s.) tchadiensisPilsbry & Bequaert (1927)
Iridina (s.s.) hardeleti var. molliPilsbry & Bequaert (1927)
Mutela (s.s.) singularisPilsbry & Bequaert (1927)
Mutela (s.s.) garambaePilsbry & Bequaert (1927)
Mutela (s.s.) garambae praetenuisPilsbry & Bequaert (1927)
Mutela dubiaThiele (1934), Modell (1942), Franc (1949), Appleton (1979), ..., Graf & Cummings (2021), Wu et al. (2022), Wu et al. (2023), Wu et al. (2024)
Iridina exoticaModell (1942), Graf (2000)
Mutela dubia var. exoticaFranc (1949)
Mutela dubia var. angustataFranc (1949)
Mutela dubia var. eminiFranc (1949)
Mutela dubiusLévèque (1967), Kenmuir (1980)
Iridina tschadiensisJaeckel (1972)
Mutela voltaeOdei (1974)
Mutela rostrataVan Damme (1984), Mandahl-Barth (1988), Ibrahim et al. (1999)
Mutela dubia dubiaMandahl-Barth (1988), Daget (1998), Graf & Cummings (2007)
Mutela dubia niloticaMandahl-Barth (1988), Daget (1998), Ibrahim et al. (1999), Graf & Cummings (2007), IUCN (2010)
Mutela dubia eminiMandahl-Barth (1988), Daget (1998), Graf & Cummings (2007), IUCN (2010)
Mutela dubia garambaeMandahl-Barth (1988), Daget (1998), Graf & Cummings (2007), IUCN (2010)
Mutela singularisIbrahim et al. (1999)

Page updated: 15:05:38 Wed 04 Dec 2024

 
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