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Page last updated
Thu 04 Jul 2019

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

This page documents data for Williams et al. (2008), such as genera and species attributed to that publication (including known misspellings, misidentifications, and other unavailable names) as well as taxonomic opinions, if available. Taxonomic opinions refer to uses of genera and species as valid and invalid in that work.

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Publication information

Williams, J.D., A.E. Bogan & J.T. Garner. 2008. Freshwater Mussels of Alabama and the Mobile Basin in Georgia, Mississippi and Tennessee: 908.

Attributed Genus Names

Leucosilla Scudder, 1882
‘Rafinesque’ of Authors.
Scudder, 1882, Nomen. Zool., Suppl.: 187.
Cox et al., 1969, Treatise on Invert. Paleo 2: N862.
Williams et al., 2008, Freshw. Mussels AL and Mobile Basin: 102.
(error for Loncosilla Rafinesque, 1831)

Euanodonata Williams et al., 2008
Williams et al., 2008, Freshw. Mussels AL and Mobile Basin: 136.
(error for Euanodonta Westerlund, 1890)

Attributed Species Names

Anodonta arkansasensis Call, 1895
‘Lea’ of Authors.
Call, 1895, Trans. Acad. Sci. St. Louis 7: 60.
Simpson, 1900, Proc. U.S. Nat. Mus. 22: 617.
Simpson, 1914, Descr. Cat. Naiades: 347.
Williams et al., 2008, Freshw. Mussels AL and Mobile Basin: 709.
Watters et al., 2009, Freshw. Mussels OH: 294.
(error for Anodonta arkansensis Lea, 1852)

Unio triangularis var. longisculus Parmalee & Bogan, 1998
‘de Gregorio’ Parmalee & Bogan, 1998, The Freshwater Mussels of Tennessee: 108.
‘de Gregorio’ Williams et al., 2008, Freshw. Mussels AL and Mobile Basin: 283.
(error for Unio triangularis var. longiusculus de Gregorio, 1914)

Unio propecaelatus Parmalee & Bogan, 1998
‘de Gregorio’ Parmalee & Bogan, 1998, The Freshwater Mussels of Tennessee: 169.
‘de Gregorio’ Williams et al., 2008, Freshw. Mussels AL and Mobile Basin: 483.
(error for Unio propecoelatus de Gregorio, 1914)

Unio subteritus [sic] var. purcheornatus Parmalee & Bogan, 1998
‘de Gregorio’ Parmalee & Bogan, 1998, The Freshwater Mussels of Tennessee: 204.
‘de Gregorio’ Williams et al., 2008, Freshw. Mussels AL and Mobile Basin: 629.
(error for Unio subteritus [sic] var. purchreornatus de Gregorio, 1914)

Unio pectitis Williams et al., 2008
‘Conrad’ Williams et al., 2008, Freshw. Mussels AL and Mobile Basin: 258.
(error for Unio pectitus Conrad, 1854)

Epioblasma capsaeformis Williams et al., 2008
Williams et al., 2008, Freshw. Mussels AL and Mobile Basin: 288.
(misidentified reference to Unio capsaeformis Lea, 1834)

Unio plantus Williams et al., 2008
‘Barnes’ Williams et al., 2008, Freshw. Mussels AL and Mobile Basin: 419.
(error for Unio planus Barnes, 1823)

Unio elegans var. elagantopsis Williams et al., 2008
‘de Gregorio’ Williams et al., 2008, Freshw. Mussels AL and Mobile Basin: 739.
(error for Unio elegans var. elegantopsis de Gregorio, 1914)

Taxonomic Opinions

Actinonaias ligamentina (Lamarck, 1819)
+ Unio crassus Say, 1817
+ Unio (Elliptio) fasciata Rafinesque, 1820
+ Unio (Elliptio) fasciata var. alternata Rafinesque, 1820
+ Unio (Elliptio) fasciata var. cuprea Rafinesque, 1820
+ Unio (Elliptio) fasciata var. nigrofasciata Rafinesque, 1820
+ Unio carinatus Barnes, 1823
+ Unio ellipticus Barnes, 1823
+ Mya gravis Wood, 1828
+ Obliquaria (Sintoxia) calendis Rafinesque, 1831
+ Unio pinguis Lea, 1857
+ Unio upsonii Marsh, 1887
+ Lampsilis (sec. Eurynia) ligamentina var. gibbus Simpson, 1900
+ Lampsilis (Eurynia) ligamentina var. nigrescens Simpson, 1914
+ Actinonaias carinata orbis Morrison, 1942

Actinonaias pectorosus (Conrad, 1834)
+ Unio perdix Lea, 1834
+ Unio biangularis Lea, 1840
+ Unio biangulatus Lea, 1840

Alasmidonta marginata Say, 1818
+ Unio rugulosus Wood, 1828
+ Decurambis scriptum Rafinesque, 1831
+ Alasmodonta truncata Lea, 1836
+ Alasmidonta (Decurambis) marginata susquehannae Ortmann, 1919
+ Alasmidonta (Decurambis) marginata variabilis F.C. Baker, 1928

Alasmidonta mccordi Athearn, 1964

Alasmidonta triangulata (Lea, 1858)

Alasmidonta viridis (Rafinesque, 1820)
+ Unio calceolus Lea, 1828
+ Alasmodonta truncata Conrad, 1834
+ Margaritana deltoidea Lea, 1838
+ Calceola angulata Swainson, 1840
+ Margaritana minor Lea, 1845
+ Unio diversus Conrad, 1856
+ Alasmidonta (Pressodonta) calceolus danielsi F.C. Baker, 1928
+ Alasmidonta (Pressodonta) calceolus magnalacustris F.C. Baker, 1928

Amblema elliottii (Lea, 1856)

Amblema neislerii (Lea, 1858)

Amblema plicata (Say, 1817)
+ Unio peruviana Lamarck, 1819
+ Unio rariplicata Lamarck, 1819
+ Amblema costata Rafinesque, 1820
+ Unio undulatus Barnes, 1823
+ Unio perplicatus Conrad, 1841
+ Unio atrocostatus Lea, 1845
+ Unio hippopoeus Lea, 1845
+ Unio latecostatus Lea, 1845
+ Unio pearlensis Conrad, 1855
+ Unio brazosensis Lea, 1868
+ Unio lincecumii Lea, 1868
+ Unio pauciplicatus Lea, 1872
+ Unio quintardii Cragin, 1887
+ Unio pilsbryi Marsh, 1891
+ Cokeria southalli Marshall, 1916

Anodonta heardi Gordon & Hoeh, 1995

Anodonta sp.

Anodonta suborbiculata Say, 1831
+ Anodonta (Nayadina) venusta de Gregorio, 1914

Anodontoides radiatus (Conrad, 1834)
+ Margaritana elliottii Lea, 1858
+ Margaritana elliptica Lea, 1859
+ Anodonta showalterii Lea, 1860

Arcidens confragosus (Say, 1829)
+ Arcidens confragosa jacintoensis Strecker, 1931

Cumberlandia monodonta (Say, 1829)
+ Unio soleniformis Lea, 1831

Cyclonaias tuberculata (Rafinesque, 1820)
+ Unio verrucosus Barnes, 1823
+ Unio verrucosus purpureus Hildreth, 1828
+ Unio graniferus Lea, 1838
+ Quadrula (sec. Rotundaria) granifera var. pusilla Simpson, 1900
+ Quadrula (Cyclonaias) tuberculata utterbackiana Frierson, 1927

Cyprogenia stegaria (Rafinesque, 1820)
+ Obovaria stegaria var. tuberculata Rafinesque, 1820
+ Unio irroratus Lea, 1828
+ Unio verrucosus albus Hildreth, 1828
+ Cyprogenia irrorata var. pusilla Simpson, 1900

Dromus dromas (Lea, 1834)
+ Unio caperatus Lea, 1845
+ Unio abacoides Haldeman, 1846

Ellipsaria lineolata (Rafinesque, 1820)
+ Obliquaria (Plagiola) depressa Rafinesque, 1820
+ Obliquaria (Ellipsaria) ellipsaria Rafinesque, 1820
+ Unio securis Lea, 1829
+ Unio compressus Rafinesque, 1831

Elliptio arca (Conrad, 1834)
+ Unio luridus Lea, 1852
+ Unio subgibbosus Lea, 1857
+ Unio rufus Lea, 1857

Elliptio arctata (Conrad, 1834)
+ Unio strigosus Lea, 1840
+ Unio tortivus Lea, 1840
+ Unio viridans Lea, 1859

Elliptio chipolaensis (Walker, 1905)

Elliptio crassidens (Lamarck, 1819)
+ Unio (Elliptio) nigra Rafinesque, 1820
+ Unio (Elliptio) nigra var. fusca Rafinesque, 1820
+ Unio (Elliptio) nigra var. maculata Rafinesque, 1820
+ Unio cuneatus Barnes, 1823
+ Obliquaria (Aximedia) venus Rafinesque, 1831
+ Unio incrassatus Lea, 1840

Elliptio dilatata (Rafinesque, 1820)
+ Unio nasuta Lamarck, 1819
+ Unio gibbosus Barnes, 1823
+ Unio mucronatus Barnes, 1823
+ Obliquaria violacea Rafinesque, 1831
+ Obliquaria bicolor Rafinesque, 1831
+ Bariosta fulvus Rafinesque, 1831
+ Unio arctior Lea, 1838
+ Unio stonensis Lea, 1840
+ Unio gibbosus var. armathwaitensis B.H. Wright, 1898
+ Unio (sec. Elliptio) gibbosus var. delicatus Simpson, 1900
+ Unio propeverutus de Gregorio, 1914
+ Elliptio dilatata var. sterkii Grier, 1918

Elliptio fraterna (Lea, 1852)

Elliptio fumata (Lea, 1857)
+ Unio subniger Lea, 1857
+ Unio hallenbeckii Lea, 1859
+ Unio quadratus Lea, 1859
+ Unio basalis Lea, 1872
+ Unio gesnerii Lea, 1874
+ Unio invenustus Lea, 1874

Elliptio mcmichaeli Clench & Turner, 1956

Elliptio nigella (Lea, 1852)
+ Unio denigratus Lea, 1857

Elliptio pullata (Lea, 1856)
+ Unio pullatis Lea, 1856
+ Unio sublatus Lea, 1857
+ Unio tetricus Lea, 1857
+ Unio aquilus Lea, 1857
+ Unio roswellensis Lea, 1858
+ Unio viridiradiatus Lea, 1859
+ Unio salebrosus Lea, 1859
+ Unio verutus Lea, 1859
+ Unio mercerii Lea, 1862
+ Unio corneus Lea, 1874
+ Unio dooleyensis Lea, 1874
+ Unio singularis B.H. Wright, 1899

Elliptio purpurella (Lea, 1857)

Elliptoideus sloatianus (Lea, 1840)
+ Unio atromarginatus Lea, 1840
+ Unio plectrophorus Conrad, 1849
+ Unio plectophorus Conrad, 1849
+ Unio aratus Conrad, 1849

Epioblasma arcaeformis (Lea, 1831)
+ Unio nexus Say, 1831

Epioblasma biemarginata (Lea, 1857)

Epioblasma brevidens (Lea, 1831)

Epioblasma capsaeformis (Lea, 1834)

Epioblasma cincinnatiensis (Lea, 1840)

Epioblasma flexuosa (Rafinesque, 1820)
+ Unio foliatus Hildreth, 1828

Epioblasma florentina (Lea, 1857)
+ Unio saccatus Küster, 1862
+ Unio sacculus Reeve, 1864
+ Truncilla walkeri Wilson & Clark, 1914
+ Truncilla curtisii Utterback, 1916

Epioblasma haysiana (Lea, 1834)
+ Unio sowerbianus Lea, 1834

Epioblasma lenior (Lea, 1842)
+ Unio lenis Lea, 1840

Epioblasma lewisii (Walker, 1910)

Epioblasma metastriata (Conrad, 1838)
+ Unio compactus Lea, 1859

Epioblasma obliquata (Rafinesque, 1820)
+ Unio ridibundus Say, 1829
+ Unio perplexus Say, 1829
+ Unio sulcatus Lea, 1829
+ Unio flagellatus Say, 1830
+ Unio gibbosus var. perobliquus Conrad, 1836
+ Unio pectitus Conrad, 1854
+ Truncilla (Scalenaria) sulcata var. delicata Simpson, 1900
+ Unio propesulcatus de Gregorio, 1914

Epioblasma othcaloogensis (Lea, 1857)
+ Unio modicellus Lea, 1859

Epioblasma penita (Conrad, 1834)

Epioblasma personata (Say, 1829)
+ Unio pileus Lea, 1831
+ Unio capillaris Lea, 1834

Epioblasma propinqua (Lea, 1857)

Epioblasma sp. cf. capaeformis

Epioblasma stewardsonii (Lea, 1852)

Epioblasma torulosa (Rafinesque, 1820)
+ Amblema gibbosa Rafinesque, 1820
+ Amblema gibbosa var. difformis Rafinesque, 1820
+ Amblema gibbosa var. olivacea Rafinesque, 1820
+ Amblema gibbosa var. radiata Rafinesque, 1820
+ Amblema torulosa var. angulata Rafinesque, 1820
+ Epioblasma biloba Rafinesque, 1831
+ Unio perplexus Lea, 1831
+ Unio rangianus Lea, 1838
+ Unio gubernaculum Reeve, 1865

Epioblasma triquetra (Rafinesque, 1820)
+ Unio cuneatus Swainson, 1823
+ Unio triangularis Barnes, 1823
+ Unio formosus Lea, 1831
+ Unio triangularis var. longiusculus de Gregorio, 1914
+ Unio triangularis var. pergibosus de Gregorio, 1914

Epioblasma turgidula (Lea, 1858)
+ Unio deviatus Reeve, 1864
+ Truncilla lefevrei Utterback, 1916

Fusconaia apalachicola Williams & Fradkin, 1999

Fusconaia burkei (Walker in Ortmann & Walker, 1922)

Fusconaia cerina (Conrad, 1838)
+ Unio glandaceus Lea, 1861
+ Unio negatus Lea, 1861
+ Unio rubidus Lea, 1861
+ Quadrula rubidula Frierson, 1905
+ Quadrula (sec. Fusconaia) castanea Simpson, 1914

Fusconaia cor (Conrad, 1834)
+ Unio edgarianus Lea, 1840
+ Unio obuncus Lea, 1871
+ Unio andersonensis Lea, 1872
+ Fusconaia cor var. analoga Ortmann, 1918

Fusconaia cuneolus (Lea, 1840)
+ Unio appressus Lea, 1871
+ Unio tuscumbiensis Lea, 1871
+ Unio flavidus Lea, 1872

Fusconaia ebena (Lea, 1831)
+ Obovaria obovalis Rafinesque, 1820
+ Obovaria pachostea Rafinesque, 1820
+ Amblema antrosa Rafinesque, 1820

Fusconaia escambia Clench & Turner, 1956

Fusconaia flava (Rafinesque, 1820)

Fusconaia masoni
+ Unio subplanus Conrad, 1837

Fusconaia rotulata (Wright, 1899)

Fusconaia subrotunda (Lea, 1831)
+ Obliquaria (Rotundaria) subrotunda var. maculata Rafinesque, 1820
+ Unio kirtlandianus Lea, 1834
+ Unio personatus Conrad, 1834
+ Unio pilaris Lea, 1840
+ Unio lesueurianus Lea, 1840
+ Unio globatus Lea, 1871
+ Unio bursapastoris B.H. Wright, 1896
+ Quadrula flexuosa Simpson, 1900
+ Quadrula (sec. Fusconaia) kirtlandiana var. minor Simpson, 1900
+ Quadrula andrewsii Marsh, 1902
+ Quadrula andrewsae Marsh, 1902
+ Quadrula beauchampii Marsh, 1902
+ Fusconaja [sic] subrotunda leucogona Ortmann, 1913

Glebula rotundata (Lamarck, 1819)
+ Unio suborbiculata Lamarck, 1819
+ Unio glebulus Say, 1831
+ Unio subglobosus Lea, 1834

Hamiota altilis (Conrad, 1834)
+ Unio clarkianus Lea, 1852
+ Unio gerhardtii Lea, 1862

Hamiota australis (Simpson, 1900)

Hamiota perovalis (Conrad, 1834)
+ Unio placitus Lea, 1852
+ Unio spillmanii Lea, 1861

Hamiota subangulata (Lea, 1840)
+ Unio kirklandianus S.H. Wright, 1897

Hemistena lata (Rafinesque, 1820)
+ Unio dehiscens Say, 1829
+ Unio oriens Lea, 1831
+ Odatelia radiata Rafinesque, 1832
+ Unio hildrethi Delessert, 1841
+ Unio (Sayunio) dehiscens var. oriensopsis de Gregorio, 1914

Lampsilis abrupta (Say, 1831)
+ Toxolasma cyclips Rafinesque, 1831

Lampsilis binominata Simpson, 1900
+ Unio lineatus Lea, 1840

Lampsilis fasciola Rafinesque, 1820
+ Unio multiradiatus Lea, 1829
+ Unio perradiatus Lea, 1858

Lampsilis floridensis (Lea, 1852)

Lampsilis ornata (Conrad, 1835)
+ Unio excavatus Lea, 1857
+ Unio doliaris Lea, 1865

Lampsilis ovata (Say, 1817)

Lampsilis straminea (Conrad, 1834)
+ Unio claibornensis Lea, 1838
+ Unio obtusus Lea, 1840
+ Unio pallescens Lea, 1845
+ Unio contrarius Conrad, 1849
+ Unio perpastus Lea, 1861

Lampsilis teres (Rafinesque, 1820)
+ Unio anodontoides Lea, 1831
+ Lampsilis fallaciosus Smith, 1899

Lampsilis virescens (Lea, 1858)

Lasmigona alabamensis Clarke, 1985

Lasmigona complanata (Barnes, 1823)
+ Unio gigas Swainson, 1824
+ Unio katherinae Lea, 1838

Lasmigona costata (Rafinesque, 1820)
+ Alasmodonta rugosa Barnes, 1823
+ Amblasmodon hians Rafinesque, 1831
+ Lasmigona costata var. ereganensis Grier, 1918
+ Lasmigona costata pepinensis F.C. Baker, 1928
+ Lasmigona costata nuda F.C. Baker, 1928

Lasmigona etowaensis (Conrad, 1849)
+ Margaritana etowahensis Lea, 1858
+ Margaritana georgiana Lea, 1859

Lasmigona holstonia (Lea, 1838)
+ Sulcularia badia Rafinesque, 1831
+ Alasmodonta impressa J.G. Anthony, 1865

Lasmigona subviridis (Conrad, 1835)
+ Unio tappanianus Lea, 1838
+ Unio neglectus Lea, 1843
+ Unio hyalinus Lea, 1845
+ Margaritana quadrata Lea, 1861
+ Unio pertenuis Lea, 1863

Lemiox rimosus (Rafinesque, 1831)
+ Unio caelatus Conrad, 1834
+ Unio coelatus Conrad, 1834

Leptodea fragilis (Rafinesque, 1820)
+ Unio planus Barnes, 1823
+ Unio gracilis Barnes, 1823
+ Unio (Niaa) atrata Swainson, 1823
+ Lasmonos fragilis Rafinesque, 1831
+ Unio permiscens Lea, 1859
+ Lampsilis simpsoni Ferriss, 1900
+ Lampsilis gracilis lacustris F.C. Baker, 1922
+ Medionidus mcglameriae van der Schalie, 1939

Leptodea leptodon (Rafinesque, 1820)
+ Anodon purpurascens Swainson, 1823
+ Symphynota tenuissima Lea, 1829
+ Unio velum Say, 1829
+ Lampsilis blatchleyi Daniels, 1902

Ligumia recta (Lamarck, 1819)
+ Unio (Eurynia) latissima Rafinesque, 1820
+ Unio praelongus Barnes, 1823
+ Unio sageri Conrad, 1836
+ Unio arquatus Conrad, 1854
+ Unio leprosus Miles, 1861

Ligumia subrostrata (Say, 1831)
+ Unio nashvillianus Lea, 1834
+ Unio mississippiensis Conrad, 1850
+ Unio rutersvillensis Lea, 1859
+ Unio cocoduensis Reeve, 1865
+ Unio topekaensis Lea, 1868
+ Lampsilis (Eurynia) subrostrata var. furva Simpson, 1914

Margaritifera marrianae Johnson, 1983

Medionidus acutissimus (Lea, 1831)
+ Unio rubellinus Lea, 1857
+ Unio semiplicatus Küster, 1862

Medionidus conradicus (Lea, 1834)
+ Obliquaria plateolus Rafinesque, 1831

Medionidus parvulus (Lea, 1860)

Medionidus penicillatus (Lea, 1857)
+ Unio kingii B.H. Wright, 1900

Megalonaias nervosa (Rafinesque, 1820)
+ Unio (crassus var.) giganteus Barnes, 1823
+ Unio undulatus Barnes, 1823
+ Unio heros Say, 1829
+ Unio multiplicatus Lea, 1831
+ Unio boykinianus Lea, 1840
+ Unio eightsii Lea, 1860
+ Unio triumphans B.H. Wright, 1898

Obliquaria reflexa Rafinesque, 1820
+ Unio cornutus Barnes, 1823
+ Unio phillipsii Conrad, 1835
+ Obliquaria reflexa conradi Frierson, 1927

Obovaria haddletoni (Athearn, 1964)

Obovaria jacksoniana (Frierson, 1912)
+ Unio castaneus Lea, 1831

Obovaria olivaria (Rafinesque, 1820)
+ Unio ellipsis Lea, 1828
+ Unio pealei Lea, 1871

Obovaria retusa (Lamarck, 1819)
+ Obovaria torsa Rafinesque, 1820

Obovaria subrotunda (Rafinesque, 1820)
+ Unio (Aximedia) levigata Rafinesque, 1820
+ Obovaria striata Rafinesque, 1820
+ Unio orbiculatus Hildreth, 1828
+ Unio circulus Lea, 1829
+ Unio lens Lea, 1831
+ Unio leibii Lea, 1862
+ Unio depygis Conrad, 1866
+ Obovaria lens var. parva Simpson, 1914
+ Obovaria lens var. elongata Simpson, 1914
+ Obovaria subrotunda globula Morrison, 1942

Obovaria unicolor (Lea, 1845)
+ Unio uber Conrad, 1866
+ Unio tinkeri B.H. Wright, 1899
+ Obovaria (s.s.) nux Simpson, 1914

Pegias fabula (Lea, 1838)
+ Margaritana curreyiana Lea, 1840
+ Unio propecoelatus de Gregorio, 1914

Plectomerus dombeyanus (Valenciennes, 1827)
+ Unio crassidens Lamarck, 1819
+ Unio interruptus Say, 1831
+ Unio trapezoides Lea, 1831

Plethobasus cicatricosus (Say, 1829)
+ Unio varicosus Lea, 1831
+ Unio detectus Frierson, 1911
+ Unio cicatricoides Frierson, 1911

Plethobasus cooperianus (Lea, 1834)

Plethobasus cyphyus (Rafinesque, 1820)
+ Unio aesopus Green, 1827
+ Unio compertus Frierson, 1911

Pleurobema athearni Gangloff Williams and Feminella, 2006

Pleurobema clava (Lamarck, 1819)
+ Unio (Aximedia) elliptica Rafinesque, 1820
+ Obliquaria (Scalenaria) scalenia Rafinesque, 1820
+ Pleurobema cuneata Rafinesque, 1820
+ Pleurobema mytiloides Rafinesque, 1820
+ Unio patulus Lea, 1829
+ Unio bournianus Lea, 1840
+ Unio consanguineus de Gregorio, 1914

Pleurobema cordatum (Rafinesque, 1820)
+ Unio obliqua Lamarck, 1819
+ Obovaria cordata var. rosea Rafinesque, 1820

Pleurobema curtum (Lea, 1859)
+ Unio anaticulus Lea, 1861

Pleurobema decisum (Lea, 1831)
+ Unio chattanoogaensis Lea, 1858
+ Unio consanguineus Lea, 1861
+ Unio crebrivittatus Lea, 1861

Pleurobema fibuloides (Lea, 1859)
+ Unio altus Conrad, 1854
+ Unio crapulus Lea, 1861
+ Unio lewisii Lea, 1861

Pleurobema georgianum (Lea, 1841)
+ Unio nucleopsis Conrad, 1849
+ Unio troschelianus Lea, 1852
+ Unio favosus Lea, 1856
+ Unio irrasus Lea, 1861
+ Unio medius Lea, 1861

Pleurobema hanleyianum (Lea, 1852)
+ Pleurobema aldrichianum Goodrich, 1931

Pleurobema hartmanianus (Lea, 1860)
+ Unio showalterii Lea, 1860

Pleurobema marshalli Frierson, 1927

Pleurobema oviforme (Conrad, 1834)
+ Unio ravenelianus Lea, 1834
+ Unio rudis Conrad, 1837
+ Unio holstonensis Lea, 1840
+ Unio mundus Lea, 1857
+ Unio lesleyi Lea, 1860
+ Unio tesserulae Lea, 1862
+ Unio striatissimus J.G. Anthony, 1865
+ Unio clinchensis Lea, 1867
+ Unio planior Lea, 1868
+ Unio lawi Lea, 1871
+ Unio acuens Lea, 1871
+ Unio pattinoides Lea, 1871
+ Unio conasaugaensis Lea, 1872
+ Unio brevis Lea, 1872
+ Unio bellulus Lea, 1872
+ Unio swordianus S.H. Wright, 1897

Pleurobema perovatum (Conrad, 1834)
+ Unio nux Lea, 1852
+ Unio johannis Lea, 1859
+ Unio cinnamomicus Lea, 1861
+ Unio flavidulus Lea, 1861
+ Unio concolor Lea, 1861
+ Unio interventus Lea, 1861
+ Unio pallidofulvus Lea, 1861
+ Unio litus Lea, 1871
+ Unio simulans Lea, 1871
+ Unio pinkstoni S.H. Wright, 1897

Pleurobema plenum (Lea, 1840)

Pleurobema pyriforme (Lea, 1857)
+ Unio striatus Lea, 1840
+ Unio modicus Lea, 1857
+ Pleurobema simpsoni Vanatta, 1915

Pleurobema rubellum (Conrad, 1834)
+ Unio furvus Conrad, 1834
+ Unio pulvinulus Lea, 1845
+ Pleurobema avellana Simpson, 1900
+ Unio (Pleurobema) hagleri Frierson, 1900
+ Pleurobema (s.s.) fictum Frierson, 1927

Pleurobema rubrum (Rafinesque, 1820)
+ Unio cardiacea Guérin, 1823
+ Unio pyramidatus Lea, 1831

Pleurobema sintoxia (Rafinesque, 1820)
+ Unio rubens Menke, 1828
+ Unio coccineus Conrad, 1836
+ Unio catillus Conrad, 1836
+ Unio solidus Lea, 1838
+ Unio cuneus Conrad, 1838
+ Unio gouldianus Jay, 1839
+ Unio fulgidus Lea, 1845
+ Quadrula (sec. Fusconaia) coccinea var. paupercula Simpson, 1900
+ Pleurobema missouriensis Marsh, 1901
+ Quadrula (sec. Fusconaia) coccinea var. magnalacustris Simpson, 1914
+ Pleurobema coccineum var. mississippiensis F.C. Baker, 1928

Pleurobema stabilis (Lea, 1861)
+ Unio murrayensis Lea, 1868

Pleurobema strodeanum (Wright, 1898)
+ Pleurobema patsaligensis Simpson, 1900

Pleurobema taitianum (Lea, 1834)
+ Pleurobema tombigbeanum Frierson, 1908
+ Pleurobema (s.s.) aldrichi Frierson, 1927

Pleurobema verum (Lea, 1861)
+ Unio instructus Lea, 1861

Pleuronaia barnesiana (Lea, 1838)
+ Unio bigbyensis Lea, 1841
+ Unio estabrookianus Lea, 1845
+ Unio tumescens Lea, 1845
+ Unio meredithii Lea, 1858
+ Unio pudicus Lea, 1860
+ Unio ornatus Lea, 1861
+ Unio lyonii Lea, 1865
+ Unio striatissimus J.G. Anthony, 1865
+ Unio fassinans Lea, 1868
+ Unio crudus Lea, 1871
+ Unio radiosus Lea, 1871
+ Unio lenticularis Lea, 1872
+ Unio tellicoensis Lea, 1872
+ Pleurobema fassinans var. rhomboidea Simpson, 1900

Pleuronaia dolabelloides (Lea, 1840)
+ Unio maculatus Conrad, 1834
+ Unio mooresianus Lea, 1857
+ Unio thorntonii Lea, 1857
+ Unio subglobatus Lea, 1871
+ Unio circumactus Lea, 1871
+ Unio recurvatus Lea, 1871
+ Pleurobema conradi Vanatta, 1915

Potamilus alatus (Say, 1817)
+ Metaptera megaptera Rafinesque, 1820

Potamilus inflatus (Lea, 1831)
+ Unio alabamensis Conrad, 1834

Potamilus ohiensis (Rafinesque, 1820)
+ Symphynota laevissima Lea, 1829

Potamilus purpuratus (Lamarck, 1819)
+ Unio ater Lea, 1829
+ Unio atra Deshayes, 1830
+ Unio lugubris Say, 1832
+ Unio poulsoni Conrad, 1834
+ Unio coloradoensis Lea, 1856
+ Unio dolosus Lea, 1860

Ptychobranchus fasciolaris (Rafinesque, 1820)
+ Obliquaria (Plagiola) interrupta Rafinesque, 1820
+ Obliquaria (Ellipsaria) fasciolaris var. interrupta Rafinesque, 1820
+ Obliquaria (Ellipsaria) fasciolaris var. fuscata Rafinesque, 1820
+ Obliquaria (Ellipsaria) fasciolaris var. longa Rafinesque, 1820
+ Obliquaria (Ellipsaria) fasciolaris var. obliterata Rafinesque, 1820
+ Unio phaseolus Hildreth, 1828
+ Unio planulatus Lea, 1829
+ Unio camelus Lea, 1834
+ Unio compressissimus Lea, 1845
+ Unio imperitus de Gregorio, 1914
+ Unio lanceolatus var. blandus de Gregorio, 1914
+ Unio compressissimus f.a performosus de Gregorio, 1914
+ Ptychobranchus fasciolaris var. lacustris F.C. Baker, 1928

Ptychobranchus foremanianus (Lea, 1842)
+ Unio velatus Conrad, 1854
+ Unio woodwardius Lea, 1857
+ Unio trinacrus Lea, 1861

Ptychobranchus greenii (Conrad, 1834)
+ Unio brumleyanus Lea, 1841
+ Unio brumbyanus Lea, 1841
+ Unio flavescens Lea, 1845
+ Unio simplex Lea, 1845

Ptychobranchus jonesi (van der Schalie, 1934)

Ptychobranchus subtentum (Say, 1825)
+ Unio subteritus [sic] var. purchreornatus de Gregorio, 1914

Pyganodon cataracta (Say, 1817)
+ Anodonta teres Conrad, 1834
+ Anodonta excurvata DeKay, 1843
+ Anodonta virgulata Lea, 1857
+ Anodonta dariensis Lea, 1858
+ Anodonta williamsii Lea, 1862
+ Anodonta tryonii Lea, 1862
+ Anodonta dolearis Lea, 1863
+ Anodonta doliaris Lea, 1863

Pyganodon grandis (Say, 1829)
+ Anodonta lugubris Say, 1829
+ Anodonta corpulenta Cooper, 1834
+ Anodonta declivis Conrad, 1834
+ Anodonta stewartiana Lea, 1834
+ Anodonta palna Lea, 1834
+ Anodonta plana Lea, 1834
+ Symphynota benedictensis Lea, 1834
+ Anodonta gigantea Lea, 1838
+ Anodonta decora Lea, 1838
+ Anodonta salmonia Lea, 1838
+ Anodonta ovata Lea, 1838
+ Anodonta pepinianus Lea, 1838
+ Anodonta footiana Lea, 1840
+ Anodonta harpethensis Lea, 1840
+ Anodonta maryattana Lea, 1840
+ Anodonta maryattiana Lea, 1840
+ Anodonta nilssonii Küster, 1842
+ Anodonta sulcata Küster, 1842
+ Anodonta opaca Lea, 1852
+ Anodonta linnaeana Lea, 1852
+ Anodonta virens Lea, 1852
+ Anodonta lewisii Lea, 1857
+ Anodonta danielsii Lea, 1858
+ Anodonta gesnerii Lea, 1858
+ Anodonta texasensis Lea, 1859
+ Anodonta simpsoniana Lea, 1861
+ Anodonta leonensis Lea, 1862
+ Anodonta dallasiana Lea, 1863
+ Anodonta bealei Lea, 1863
+ Anodon subangulata J.G. Anthony, 1865
+ Anodon imbricata J.G. Anthony, 1865
+ Anodon opalina J.G. Anthony, 1865
+ Anodon subinflata J.G. Anthony, 1865
+ Anodon micans J.G. Anthony, 1865
+ Anodon mcnielii J.G. Anthony, 1866
+ Anodon subgibbosa J.G. Anthony, 1866
+ Anodon inornata J.G. Anthony, 1866
+ Margaron (Anodonta) benedictii Lea, 1870
+ Anodonta hockingensis Call, 1880
+ Anodonta somersii Call, 1880
+ Anodonta houghtonensis DeCamp, 1881
+ Anodonta dakota Frierson, 1910

Quadrula apiculata (Say, 1829)
+ Unio asper Lea, 1831
+ Unio forsheyi Lea, 1859

Quadrula asperata (Lea, 1861)
+ Unio pauperculus Lea, 1861
+ Unio vallatus Lea, 1868
+ Unio cahabensis Lea, 1871
+ Quadrula archeri Frierson, 1905

Quadrula cylindrica (Say, 1817)
+ Unio naviformis Lamarck, 1819
+ Unio cylindricus var. strigillatus B.H. Wright, 1898

Quadrula infucata (Conrad, 1834)
+ Unio securiformis Conrad, 1849

Quadrula intermedia (Conrad, 1836)
+ Unio tuberosus var. perlobatus de Gregorio, 1914
+ Quadrula biangulata Morrison, 1942

Quadrula kieneriana (Lea, 1852)

Quadrula metanevra (Rafinesque, 1820)
+ Unio nodosus Barnes, 1823
+ Unio tuberosus Lea, 1840
+ Unio wardii Lea, 1862

Quadrula nobilis (Conrad, 1853)

Quadrula nodulata (Rafinesque, 1820)

Quadrula pustulosa (Lea, 1831)
+ Obliquaria (Quadrula) retusa Rafinesque, 1820
+ Obliquaria (Quadrula) bullata Rafinesque, 1820
+ Unio premorsus Rafinesque, 1831
+ Unio nodulosus Say, 1834
+ Unio prasinus Conrad, 1834
+ Unio schoolcraftensis Lea, 1834
+ Unio mortoni Conrad, 1836
+ Unio turgidus Lea, 1838
+ Unio dorfeuillianus Lea, 1838
+ Unio pernodosus Lea, 1845
+ Unio schoolcraftii Lea, 1870

Quadrula quadrula (Rafinesque, 1820)
+ Unio rugosus Barnes, 1823
+ Unio lacrymosus Lea, 1828
+ Unio asperrimus Lea, 1831
+ Unio lunulatus Pratt, 1876
+ Quadrula quadrula contraryensis Utterback, 1915
+ Quadrula (Quadrula) quadrula bullocki F.C. Baker, 1928

Quadrula refulgens (Lea, 1868)

Quadrula rumphiana (Lea, 1852)
+ Unio blandianus Lea, 1856

Quadrula sparsa (Lea, 1841)

Quadrula stapes (Lea, 1831)

Quadrula succissa (Lea, 1852)
+ Unio cacao Lea, 1859

Quadrula verrucosa (Rafinesque, 1820)
+ Unio tuberculatus Barnes, 1823
+ Unio (Theliderma) pustulata Swainson, 1840
+ Unio conjugans B.H. Wright, 1899
+ Quadrula tritogonia Ortmann, 1909
+ Quadrula parkeri Geiser, 1911

Strophitus connasaugaensis (Lea, 1858)
+ Margaritana gesnerii Lea, 1858
+ Anodonta hallenbeckii Lea, 1858
+ Margaritana alabamensis Lea, 1861

Strophitus subvexus (Conrad, 1834)
+ Margaritana spillmanii Lea, 1858
+ Margaritana tombecbeensis Lea, 1858
+ Margaritana tombigbeensis Lea, 1858
+ Margaritana columbensis Lea, 1867

Strophitus undulatus (Say, 1817)
+ Anodonta pensylvanica Lamarck, 1819
+ Anodon rugosus Swainson, 1822
+ Alasmodonta edentula Say, 1829
+ Anodon areolatus Swainson, 1829
+ Anodonta virgata Conrad, 1836
+ Anodonta wardiana Lea, 1838
+ Anodonta pavonia Lea, 1838
+ Anodon unadilla DeKay, 1843
+ Anodonta tetragona Lea, 1845
+ Anodonta arkansensis Lea, 1852
+ Anodonta shaefferiana Lea, 1852
+ Alasmodon rhombica J.G. Anthony, 1865
+ Anodon papyracea J.G. Anthony, 1865
+ Anodon annulatus Sowerby, 1867
+ Anodon quadriplicatus Sowerby, 1870
+ Strophitus undulatus ovatus Frierson, 1927
+ Strophitus undulatus tennesseensis Frierson, 1927
+ Strophitus rugosus pepinensis F.C. Baker, 1928
+ Strophitus rugosus winnebagoensis F.C. Baker, 1928
+ Strophitus rugosus lacustris F.C. Baker, 1928

Toxolasma corvunculus of Authors, non Lea, 1868

Toxolasma cylindrellus (Lea, 1868)

Toxolasma germanus (Lea, 1861)

Toxolasma granulatus (Lea, 1861)

Toxolasma lividum (Rafinesque, 1831)
+ Unio glans Lea, 1831
+ Unio moestus Lea, 1841

Toxolasma parvum (Barnes, 1823)
+ Carunculina parva cahni F.C. Baker, 1927

Toxolasma paulum (Lea, 1840)

Toxolasma sp.

Truncilla donaciformis (Lea, 1828)
+ Unio zigzag Lea, 1829

Truncilla truncata Rafinesque, 1820
+ Truncilla truncata var. fusca Rafinesque, 1820
+ Truncilla truncata var. vermiculata Rafinesque, 1820
+ Unio elegans Lea, 1831
+ Unio elegans var. elegantopsis de Gregorio, 1914
+ Unio elegans var. magnelegans de Gregorio, 1914
+ Truncilla truncata var. lacustris F.C. Baker, 1928

Uniomerus columbensis (Lea, 1857)
+ Unio plantii Lea, 1857

Uniomerus tetralasmus (Say, 1830)
+ Unio camptodon Say, 1832
+ Unio geometricus Lea, 1834
+ Unio excultus Conrad, 1838
+ Unio sayanus Conrad, 1838
+ Unio sayii Tappan, 1839
+ Unio parallelus Conrad, 1841
+ Unio symmetricus Lea, 1845
+ Unio rivularis Conrad, 1853
+ Unio porrectus Conrad, 1854
+ Unio subcroceus Conrad, 1854
+ Unio manubius Gould, 1855
+ Unio jamesianus Lea, 1857
+ Unio electrinus Reeve, 1865

Utterbackia imbecillis (Say, 1829)
+ Anodonta incerta Lea, 1834
+ Anodon horda Gould, 1855
+ Anodonta phalena de Gregorio, 1914
+ Utterbackia imbecillis fusca F.C. Baker, 1927

Utterbackia peggyae (Johnson, 1965)

Villosa choctawensis Athearn, 1964

Villosa fabalis (Lea, 1831)
+ Unio capillus Say, 1831
+ Unio lapillus Say, 1831
+ Unio donacopsis de Gregorio, 1914

Villosa iris (Lea, 1829)
+ Unio notatus Lea, 1838
+ Unio obscurus Lea, 1838
+ Unio zeiglerianus Lea, 1838
+ Unio muhlfeldianus Lea, 1838
+ Unio creperus Lea, 1838
+ Unio glaber Lea, 1838
+ Unio simus Lea, 1838
+ Unio novieboraci Lea, 1838
+ Unio amoenus Lea, 1840
+ Unio tener Lea, 1840
+ Unio dactylus Lea, 1840
+ Unio fatuus Lea, 1840
+ Unio regularis Lea, 1841
+ Unio puniceus Haldeman, 1842
+ Margaritana (Unio) cumberlandianus Lea, 1852
+ Unio discrepans Lea, 1860
+ Unio planicostatus Lea, 1860
+ Unio scitulus Lea, 1860
+ Unio perpictus Lea, 1860
+ Unio opalinus J.G. Anthony, 1866
+ Unio dispansus Lea, 1871

Villosa lienosa (Conrad, 1834)
+ Unio saxeus Conrad, 1838
+ Unio caliginosus Lea, 1845
+ Unio gouldii Lea, 1845
+ Unio nigerrimus Lea, 1852
+ Unio proximus Lea, 1852
+ Unio subellipsis Lea, 1856
+ Unio concestator Lea, 1857
+ Unio apicinus Lea, 1857
+ Unio fallax Lea, 1857
+ Unio intercedens Lea, 1857
+ Unio obfuscus Lea, 1857
+ Unio prattii Lea, 1858
+ Unio dispar Lea, 1860
+ Unio linguaeformis Lea, 1860
+ Unio contiguus Lea, 1862
+ Unio fuligo Reeve, 1865
+ Unio bicaelatus Reeve, 1865
+ Unio fontanus Conrad, 1866
+ Unio unicostatus B.H. Wright, 1899

Villosa nebulosa (Conrad, 1834)
+ Unio radians Lea, 1857
+ Unio jonesii Lea, 1859
+ Unio plancus Lea, 1860
+ Unio sparus Lea, 1868
+ Unio difficilis Lea, 1868

Villosa taeniata (Conrad, 1834)
+ Unio pictus Lea, 1834
+ Unio interruptus Lea, 1838
+ Unio latiradiatus Conrad, 1838
+ Unio pulcher Lea, 1838
+ Unio menkianus Lea, 1838
+ Unio tennesseensis Lea, 1840
+ Unio camelopardilis Lea, 1860
+ Unio fucatus Lea, 1860
+ Unio lindsleyi Lea, 1860
+ Unio punctatus Lea, 1865

Villosa trabalis (Conrad, 1834)
+ Unio troostensis Lea, 1834
+ Unio troostii Lea, 1870

Villosa umbrans (Lea, 1857)
+ Unio umbrosus Lea, 1857
+ Unio tenebricus Lea, 1857
+ Unio fabaceus Lea, 1861
+ Unio porphyreus Lea, 1861
+ Unio proprius Lea, 1865

Villosa vanuxemensis (Lea, 1838)
+ Unio nitens Lea, 1840
+ Unio pybasii Lea, 1858
+ Unio copei Lea, 1868
+ Margaron (Unio) vanuxemii Lea, 1870

Villosa vibex (Conrad, 1834)
+ Unio exiguus Lea, 1840
+ Unio prevostianus Lea, 1852
+ Unio nigrinus Lea, 1852
+ Unio rutilans Lea, 1856
+ Unio sudus Lea, 1857

Villosa villosa (Wright, 1898)
+ Lampsilis (Villosa) wrightiana Frierson, 1927

Page updated: 10:25:37 Thu 04 Jul 2019

 
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