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Page last updated
Thu 04 Jul 2019

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

This page documents data for Watters et al. (2009), such as genera and species attributed to that publication (including known misspellings, misidentifications, and other unavailable names) as well as taxonomic opinions, if available. Taxonomic opinions refer to uses of genera and species as valid and invalid in that work.

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Publication information

Watters, G.T., M.A. Hoggarth & D.H. Stansbery. 2009. The Freshwater Mussels of Ohio. 421 pp.

Attributed Species Names

Anodonta linneana Conrad, 1853
‘Lea’ of Authors.
Conrad, 1853, Proc. Acad. Nat. Sci. 6: 264.
Hanley, 1856, Cat. Rec. Bivalve Shells, Suppl.: 387.
Sowerby, 1870, Conch. Icon. 17: pl. 35, sp. 144.
Simpson, 1900, Proc. U.S. Nat. Mus. 22: 646.
Simpson, 1914, Descr. Cat. Naiades: 431.
Frierson, 1927, Check List N. Amer. Naiades: 14.
Haas, 1969, Das Tierreich 88: 364.
Parmalee & Bogan, 1998, The Freshwater Mussels of Tennessee: 206.
Watters et al., 2009, Freshw. Mussels OH: 260.
(error for Anodonta linnaeana Lea, 1852)

Anodonta arkansasensis Call, 1895
‘Lea’ of Authors.
Call, 1895, Trans. Acad. Sci. St. Louis 7: 60.
Simpson, 1900, Proc. U.S. Nat. Mus. 22: 617.
Simpson, 1914, Descr. Cat. Naiades: 347.
Williams et al., 2008, Freshw. Mussels AL and Mobile Basin: 709.
Watters et al., 2009, Freshw. Mussels OH: 294.
(error for Anodonta arkansensis Lea, 1852)

Unio stewardsonii var. stewardsonii Parmalee & Bogan, 1998
‘De Gregorio’ of Authors.
Parmalee & Bogan, 1998, The Freshwater Mussels of Tennessee: 97.
Watters et al., 2009, Freshw. Mussels OH: 95.
(error for Unio stewardsonii var. stewensoni de Gregorio, 1914)

Unio pertenuis Watters et al., 2009
‘de Gregorio’ Watters et al., 2009, Freshw. Mussels OH: 302.
(error for Unio pertennis de Gregorio, 1914)

Unio lehmanni Watters et al., 2009
‘Wright’ Watters et al., 2009, Freshw. Mussels OH: 82.
(error for Unio lehmanii S.H. Wright, 1897)

Taxonomic Opinions

Actinonaias ligamentina carinata (Barnes, 1823)
+ Actinonaias carinata orbis Morrison, 1942

Actinonaias ligamentina ligamentina (Lamarck, 1819)
+ Unio crassus Say, 1817
+ Lampsilis (sec. Eurynia) ligamentina var. gibbus Simpson, 1900
+ Unio tecomensis de Gregorio, 1914

Alasmidonta marginata Say, 1818
+ Unio rugulosus Wood, 1828
+ Decurambis scriptum Rafinesque, 1831
+ Alasmodonta truncata Lea, 1836
+ Margaritana marginata Küster, 1862
+ Margaritana marginata var. truncata B.H. Wright, 1898
+ Unio calceolus f.a sciotincola de Gregorio, 1914
+ Alasmidonta (Decurambis) marginata susquehannae Ortmann, 1919
+ Alasmidonta (Decurambis) marginata variabilis F.C. Baker, 1928

Alasmidonta viridis (Rafinesque, 1820)
+ Unio calceolus Lea, 1828
+ Lasmigona viridis Rafinesque, 1831
+ Lasmigona viridis var. chloris Rafinesque, 1831
+ Lasmigona viridis var. radiata Rafinesque, 1831
+ Lasmigona viridis var. fuscata Rafinesque, 1831
+ Alasmodonta truncata Conrad, 1834
+ Margaritana deltoidea Lea, 1838
+ Calceola angulata Swainson, 1840
+ Margaritana minor Lea, 1845
+ Alasmidonta (Pressodonta) calceolus magnalacustris F.C. Baker, 1928
+ Alasmidonta (Pressodonta) calceolus danielsi F.C. Baker, 1928

Amblema plicata perplicata (Conrad, 1841)

Amblema plicata plicata (Say, 1817)
+ Unio peruviana Lamarck, 1819
+ Unio rariplicata Lamarck, 1819
+ Amblema costata Rafinesque, 1820
+ Unio crassus Barnes, 1823
+ Unio gigantea Lea, 1834
+ Unio hippopoeus Lea, 1845
+ Unio hippopaeus Lea, 1852
+ Unio pearlensis Conrad, 1855
+ Unio heros Küster, 1855
+ Unio brazosensis Lea, 1868
+ Unio lincecumii Lea, 1868
+ Unio pauciplicatus Lea, 1872
+ Unio quintardii Cragin, 1887
+ Unio lincecurei Paetel, 1890
+ Unio pilsbryi Marsh, 1891
+ Cokeria southalli Marshall, 1916

Anodonta suborbiculata Say, 1831

Anodontoides ferussacianus (Lea, 1834)
+ Anodonta buchanensis Lea, 1838
+ Anodonta subcylindracea Lea, 1838
+ Anodonta argentea Lea, 1840
+ Anodonta ferruginea Lea, 1840
+ Anodon plicatus Haldeman, 1842
+ Anodonta oblita Lea, 1852
+ Anodonta modesta Lea, 1857
+ Anodon subcylindracea Sowerby, 1867
+ Anodonta subcilindracea de Gregorio, 1914
+ Anodonta ferussaceana de Gregorio, 1914
+ Anodontoides birgei F.C. Baker, 1923

Arcidens confragosus (Say, 1829)
+ Arcidens confragosa jacintoensis Strecker, 1931

Cumberlandia monodonta (Say, 1829)
+ Unio soleniformis Lea, 1831

Cyclonaias tuberculata (Rafinesque, 1820)
+ Unio verrucosus Barnes, 1823
+ Unio tuberculosus Valenciennes, 1827
+ Unio verrucosus purpureus Hildreth, 1828
+ Unio graniferus Lea, 1838
+ Quadrula (sec. Rotundaria) granifera var. pusilla Simpson, 1900
+ Plethobasus cooperianus Utterback, 1915
+ Quadrula (Cyclonaias) tuberculata utterbackiana Frierson, 1927

Cyprogenia stegaria (Rafinesque, 1820)
+ Obovaria stegaria var. tuberculata Rafinesque, 1820
+ Unio irroratus Lea, 1828
+ Unio stegarius Say, 1834
+ Cyprogenia irrorata var. pusilla Simpson, 1900

Ellipsaria lineolata (Rafinesque, 1820)
+ Obliquaria (Plagiola) depressa Rafinesque, 1820
+ Obliquaria (Ellipsaria) ellipsaria Rafinesque, 1820
+ Unio securis Lea, 1829
+ Unio compressus Rafinesque, 1831
+ Unio lineolatus Say, 1834

Elliptio complanata (Lightfoot, 1786)

Elliptio crassidens (Lamarck, 1819)
+ Unio (Elliptio) nigra Rafinesque, 1820
+ Unio (Elliptio) nigra var. fusca Rafinesque, 1820
+ Unio (Elliptio) nigra var. maculata Rafinesque, 1820
+ Unio cuneatus Barnes, 1823
+ Obliquaria (Aximedia) venus Rafinesque, 1831
+ Unio discus Sowerby, 1868
+ Unio crassus Sowerby, 1868
+ Unio lehmanii S.H. Wright, 1897
+ Unio danielsii B.H. Wright, 1899
+ Unio polymorphus B.H. Wright, 1899
+ Elliptio pachyodon Pilsbry, 1953

Elliptio dilatata (Rafinesque, 1820)
+ Unio nasuta Lamarck, 1819
+ Obliquaria (Ellipsaria) sinuata Rafinesque, 1820
+ Unio mucronatus Barnes, 1823
+ Unio gibbosus Barnes, 1823
+ Obliquaria violacea Rafinesque, 1831
+ Bariosta fulvus Rafinesque, 1831
+ Obliquaria bicolor Rafinesque, 1831
+ Unio euliratus Rafinesque, 1831
+ Unio dilatatus Say, 1834
+ Unio arctior Lea, 1838
+ Unio stonensis Lea, 1840
+ Unio gibbosus var. armathwaitensis B.H. Wright, 1898
+ Unio (sec. Elliptio) gibbosus var. delicatus Simpson, 1900
+ Unio propeverutus de Gregorio, 1914
+ Elliptio dilatata var. subgibbosa Utterback, 1915
+ Elliptio dilatata var. sterkii Grier, 1918

Epioblasma flexuosa (Rafinesque, 1820)
+ Obliquaria (Quadrula) flexuosa var. bullata Rafinesque, 1820
+ Unio foliatus Hildreth, 1828

Epioblasma lewisi (Walker, 1910)

Epioblasma obliquata obliquata (Rafinesque, 1820)
+ Unio ridibundus Say, 1829
+ Unio pectitus Conrad, 1854
+ Unio stewardsonii var. stewensoni de Gregorio, 1914
+ Unio propesulcatus de Gregorio, 1914
+ Unio propeculcatus Parmalee & Bogan, 1998

Epioblasma obliquata perobliqua (Conrad, 1836)

Epioblasma personata (Say, 1829)
+ Unio pileus Lea, 1831
+ Unio capillaris Lea, 1834

Epioblasma phillipsii (Conrad, 1835)
+ Unio cincinnatiensis Lea, 1840

Epioblasma torulosa rangiana (Lea, 1838)
+ Epioblasma biloba Rafinesque, 1831
+ Truncilla (Scalenaria) sulcata var. delicata Simpson, 1900

Epioblasma torulosa torulosa (Rafinesque, 1820)
+ Amblema torulosa var. angulata Rafinesque, 1820
+ Amblema gibbosa Rafinesque, 1820
+ Amblema gibbosa var. olivacea Rafinesque, 1820
+ Amblema gibbosa var. radiata Rafinesque, 1820
+ Amblema gibbosa var. difformis Rafinesque, 1820

Epioblasma triquetra (Rafinesque, 1820)
+ Unio triangularis Barnes, 1823
+ Unio cuneatus Swainson, 1823
+ Unio formosus Lea, 1831
+ Unio triangularis var. pergibosus de Gregorio, 1914
+ Unio triangularis var. longiusculus de Gregorio, 1914

Fusconaia ebena (Lea, 1831)
+ Amblema antrosa Rafinesque, 1820
+ Obovaria pachostea Rafinesque, 1820
+ Obovaria obovalis Rafinesque, 1820
+ Unio obovalis Say, 1834
+ Unio obliqua Ferussac, 1835
+ Unio (s.s.) mytiloides Swainson, 1840

Fusconaia flava (Rafinesque, 1820)
+ Obliquaria (Sintoxia) lateralis Rafinesque, 1820
+ Unio undatus Barnes, 1823
+ Unio rubiginosus Lea, 1829
+ Unio trigonus Lea, 1831
+ Unio undatus Conrad, 1834
+ Unio flavus Conrad, 1837
+ Unio triangularis Küster, 1852
+ Unio pilaris Reeve, 1865
+ Fusconaia selecta Wheeler, 1914
+ Unio validus var. continuus de Gregorio, 1914
+ Fusconaia undata trigonoides Utterback, 1915
+ Fusconaia flava var. parvula Grier, 1918
+ Fusconaia undata wagneri F.C. Baker, 1928

Fusconaia subrotunda lesueuriana (Lea, 1840)

Fusconaia subrotunda subrotunda (Lea, 1831)
+ Obliquaria (Rotundaria) subrotunda var. maculata Rafinesque, 1820
+ Unio kirtlandianus Lea, 1834
+ Unio personatus Conrad, 1834
+ Unio politus Conrad, 1837
+ Unio kirklandianus Hanley, 1843
+ Fusconaja [sic] subrotunda leucogona Ortmann, 1913

Glebula rotundata (Lamarck, 1819)
+ Unio suborbiculata Lamarck, 1819
+ Unio glebulus Say, 1831
+ Unio subglobosus Lea, 1834
+ Unio grandensis Conrad, 1855

Hemistena lata (Rafinesque, 1820)
+ Unio dehiscens Say, 1829
+ Unio oriens Lea, 1831
+ Odatelia radiata Rafinesque, 1832
+ Unio (Sayunio) dehiscens var. oriensopsis de Gregorio, 1914

Lampsilis abrupta (Say, 1831)
+ Unio orbiculatus Hildreth, 1828
+ Toxolasma cyclips Rafinesque, 1831
+ Toxolasma cyclips var. fuscata Rafinesque, 1831
+ Toxolasma cyclips var. lutescens Rafinesque, 1831
+ Unio cyclips Ferussac, 1835
+ Unio crassus Conrad, 1836

Lampsilis cardium Rafinesque, 1820
+ Unio ventricosus Barnes, 1823
+ Unio occidens Lea, 1829
+ Unio subovatus Lea, 1831
+ Unio cardium Conrad, 1834
+ Unio lenis Conrad, 1838
+ Unio ovatus Deshayes, 1839
+ Unio canadensis Lea, 1857
+ Unio dolabraeformis Sowerby, 1867
+ Unio latissimus Sowerby, 1868
+ Lampsilis ventricosa cohongoronta Ortmann, 1912
+ Lampsilis (s.s.) ventricosa var. lurida Simpson, 1914
+ Lampsilis ventricosa perglobosa F.C. Baker, 1928
+ Lampsilis ventricosa winnebagoensis F.C. Baker, 1928

Lampsilis fasciola Rafinesque, 1820
+ Unio multiradiatus Lea, 1829
+ Unio fasciolus Say, 1834
+ Unio perradiatus Lea, 1858
+ Unio altilis Reeve, 1865
+ Unio perovalis Sowerby, 1866

Lampsilis ovata (Say, 1817)

Lampsilis radiata luteola (Lamarck, 1819)
+ Unio siliquoideus Barnes, 1823
+ Unio inflatus Barnes, 1823
+ Unio childreni Hanley, 1843
+ Unio siliquoides Küster, 1848
+ Unio hydianus Küster, 1861
+ Unio distans J.G. Anthony, 1865
+ Unio affinis Sowerby, 1868
+ Unio multiradiatus Sowerby, 1868
+ Unio superiorensis Marsh, 1897
+ Lampsilis siliquoidea pepinensis F.C. Baker, 1927
+ Lampsilis siliquoidea chadwicki F.C. Baker, 1928

Lampsilis rafinesqueana Frierson, 1927
+ Actinonaias carinata carinata Murray & Leonard, 1962

Lampsilis teres (Rafinesque, 1820)
+ Unio anodontoides Lea, 1831
+ Unio floridensis Lea, 1852
+ Unio oriens Sowerby, 1868
+ Lampsilis anodontoides F.C. Baker, 1898
+ Lampsilis fallaciosus Smith, 1899

Lasmigona complanata (Barnes, 1823)
+ Unio gigas Swainson, 1824
+ Unio katherinae Lea, 1838
+ Alasmodon complanatus Sowerby, 1839
+ Complanaria gigas Swainson, 1840

Lasmigona complanata alabamensis Clarke, 1985

Lasmigona compressa (Lea, 1829)
+ Unio alasmodontina Lea, 1829
+ Unio compressus var. plebius C.B. Adams, 1842
+ Complanaria alasmodontina Stimpson, 1851
+ Margaron (Unio) pressus Lea, 1852
+ Unio compressa var. lindus de Gregorio, 1914

Lasmigona costata (Rafinesque, 1820)
+ Alasmodonta rugosa Barnes, 1823
+ Lasmigona rugosa Rafinesque, 1831
+ Amblasmodon hians Rafinesque, 1831
+ Alasmodonta hians Ferussac, 1835
+ Lasmigona costata var. eriganensis Grier, 1918
+ Lasmigona costata nuda F.C. Baker, 1928
+ Lasmigona costata pepinensis F.C. Baker, 1928

Lasmigona subviridis (Conrad, 1835)
+ Unio viridis Conrad, 1835
+ Unio tappanianus Lea, 1838
+ Unio hyalinus Lea, 1845
+ Unio tappianus Catlow & Reeve, 1845
+ Unio pertenuis Lea, 1863

Leptodea fragilis (Rafinesque, 1820)
+ Unio gracilis Barnes, 1823
+ Unio planus Barnes, 1823
+ Unio (Niaa) atrata Swainson, 1823
+ Mya plana Eaton, 1826
+ Lasmonos fragilis Rafinesque, 1831
+ Unio leptodea var. planus Jay, 1850
+ Lampsilis simpsoni Ferriss, 1900
+ Lampsilis gracilis lacustris F.C. Baker, 1922

Leptodea leptodon (Rafinesque, 1820)
+ Anodon purpurascens Swainson, 1823
+ Unio velum Say, 1829
+ Symphynota tenuissima Lea, 1829
+ Unio leptodon Conrad, 1836
+ Lampsilis blatchleyi Daniels, 1902

Ligumia nasuta (Say, 1817)
+ Obliquaria attenuata Rafinesque, 1820
+ Unio rostrata Valenciennes, 1827
+ Unio vaughanianus Sowerby, 1868

Ligumia recta (Lamarck, 1819)
+ Unio (Eurynia) latissima Rafinesque, 1820
+ Unio praelongus Barnes, 1823
+ Unio rectus Conrad, 1836
+ Unio sageri Conrad, 1836
+ Unio arquatus Conrad, 1854
+ Unio leprosus Miles, 1861
+ Unio angustatus var. cuniculus de Gregorio, 1914

Ligumia subrostrata (Say, 1831)
+ Unio nashvillianus Lea, 1834
+ Unio mississippiensis Conrad, 1850
+ Unio rutersvillensis Lea, 1859
+ Unio cocoduensis Reeve, 1865
+ Unio topekaensis Lea, 1868
+ Lampsilis (Eurynia) subrostrata var. furva Simpson, 1914

Margaritifera margaritifera (Linnaeus, 1758)
+ Dichonca crassissima Klein, 1753
+ Margaritana fluviatilis Schumacher, 1817
+ Unio elongata Lamarck, 1819
+ Alasmodonta arcuata Barnes, 1823
+ Unio roissyi Michaud, 1831
+ Unio margaritifer var. minor Rossmässler, 1835
+ Unio tristis Morelet, 1845

Megalonaias nervosa (Rafinesque, 1820)
+ Unio (crassus var.) giganteus Barnes, 1823
+ Unio undulatus Barnes, 1823
+ Unio undulata Valenciennes, 1827
+ Unio heros Say, 1829
+ Unio multiplicatus Lea, 1831
+ Unio atrocostatus Lea, 1845
+ Unio eightsii Lea, 1860
+ Unio atrocostatus Reeve, 1864
+ Unio eighti Paetel, 1890
+ Unio triumphans B.H. Wright, 1898

Obliquaria reflexa Rafinesque, 1820
+ Unio cornutus Barnes, 1823
+ Unio torulosus Short & Eaton, 1831
+ Obliquaria reflexa conradi Frierson, 1927

Obovaria olivaria (Rafinesque, 1820)
+ Unio brevialis Sowerby & Sowerby, 1823
+ Unio ellipsis Lea, 1828
+ Unio olivarius Conrad, 1834
+ Diplasma vitrea Potiez & Michaud, 1844
+ Unio taitanus Sowerby, 1868
+ Unio pealei Lea, 1871
+ Unio pearlii Scammon, 1906

Obovaria retusa (Lamarck, 1819)
+ Obovaria torsa Rafinesque, 1820
+ Unio torsus Potiez & Michaud, 1844

Obovaria subrotunda (Rafinesque, 1820)
+ Unio (Aximedia) levigata Rafinesque, 1820
+ Obovaria striata Rafinesque, 1820
+ Unio orbiculatus Hildreth, 1828
+ Unio rotundus Wood, 1828
+ Unio circulus Lea, 1829
+ Unio lens Lea, 1831
+ Unio laevigatus Say, 1834
+ Unio leibii Lea, 1862
+ Unio depygis Conrad, 1866
+ Obovaria lens var. parva Simpson, 1914
+ Obovaria lens var. elongata Simpson, 1914
+ Obovaria subrotunda globula Morrison, 1942

Plethobasus cicatricosus (Say, 1829)
+ Unio varicosus Lea, 1831
+ Unio detectus Frierson, 1911
+ Unio cicatricoides Frierson, 1911
+ Plethobasus pachosteus Morrison, 1969

Plethobasus cooperianus (Lea, 1834)
+ Unio striatus Reeve, 1864

Plethobasus cyphyus (Rafinesque, 1820)
+ Unio cyphia Rafinesque, 1820
+ Unio aesopus Green, 1827
+ Unio scyphius Küster, 1861
+ Unio compertus Frierson, 1911

Pleurobema clava (Lamarck, 1819)
+ Unio (Aximedia) elliptica Rafinesque, 1820
+ Pleurobema mytiloides Rafinesque, 1820
+ Pleurobema cuneata Rafinesque, 1820
+ Obliquaria (Scalenaria) scalenia Rafinesque, 1820
+ Unio patulus Lea, 1829
+ Unio cuneatus Say, 1834
+ Unio bournianus Lea, 1840
+ Unio consanguineus de Gregorio, 1914
+ Unio anaticulus var. ohiensis de Gregorio, 1914

Pleurobema cordatum (Rafinesque, 1820)
+ Unio obliqua Lamarck, 1819
+ Obovaria cordata var. rosea Rafinesque, 1820
+ Unio mytiloides Short & Eaton, 1831
+ Unio obliquus Call, 1900

Pleurobema plenum (Lea, 1840)
+ Quadrula cordata Vanatta, 1915

Pleurobema rubrum (Rafinesque, 1820)
+ Unio cardiacea Guérin, 1823
+ Unio obliquus Wood, 1828
+ Unio mytiloides Deshayes, 1830
+ Unio pyramidatus Lea, 1831
+ Unio ruber Conrad, 1853
+ Unio mytilloides Frierson, 1927

Pleurobema sintoxia (Rafinesque, 1820)
+ Unio coccineus Conrad, 1836
+ Unio catillus Conrad, 1836
+ Unio cuneus Conrad, 1838
+ Unio solidus Lea, 1838
+ Unio cardiacea Deshayes, 1839
+ Unio gouldianus Jay, 1839
+ Unio fulgidus Lea, 1845
+ Unio obovalis Conrad, 1853
+ Unio catilus B.H. Wright, 1888
+ Quadrula (sec. Fusconaia) coccinea var. paupercula Simpson, 1900
+ Pleurobema missouriensis Marsh, 1901
+ Quadrula (sec. Fusconaia) coccinea var. magnalacustris Simpson, 1914
+ Pleurobema coccineum var. mississippiensis F.C. Baker, 1928

Potamilus alatus (Say, 1817)
+ Metaptera megaptera Rafinesque, 1820

Potamilus capax (Green, 1832)
+ Symphynota globosa Lea, 1834

Potamilus ohiensis (Rafinesque, 1820)
+ Symphynota laevissima Lea, 1829
+ Unio ohiensis Say, 1834
+ Unio discoideus Sowerby, 1866

Ptychobranchus fasciolaris (Rafinesque, 1820)
+ Obliquaria (Plagiola) interrupta Rafinesque, 1820
+ Obliquaria (Ellipsaria) fasciolaris var. interrupta Rafinesque, 1820
+ Obliquaria (Ellipsaria) fasciolaris var. fuscata Rafinesque, 1820
+ Obliquaria (Ellipsaria) fasciolaris var. obliterata Rafinesque, 1820
+ Obliquaria (Ellipsaria) fasciolaris var. longa Rafinesque, 1820
+ Unio phaseolus Hildreth, 1828
+ Unio planulatus Lea, 1829
+ Unio camelus Lea, 1834
+ Unio fasciolaris Conrad, 1834
+ Unio compressissimus Lea, 1845
+ Unio arquatus Conrad, 1854
+ Unio compressissimus f.a performosus de Gregorio, 1914
+ Unio lanceolatus var. blandus de Gregorio, 1914
+ Unio imperitus de Gregorio, 1914
+ Ptychobranchus fasciolaris var. lacustris F.C. Baker, 1928

Pyganodon cataracta marginata (Say, 1817)

Pyganodon grandis grandis (Say, 1829)
+ Anodonta lugubris Say, 1829
+ Anodonta palna Lea, 1834
+ Anodonta stewartiana Lea, 1834
+ Symphynota benedictensis Lea, 1834
+ Anodonta declivis Conrad, 1834
+ Anodonta corpulenta Cooper, 1834
+ Anodonta gigantea Lea, 1838
+ Anodonta ovata Lea, 1838
+ Anodonta salmonia Lea, 1838
+ Anodonta decora Lea, 1838
+ Anodonta pepinianus Lea, 1838
+ Anodonta harpethensis Lea, 1840
+ Anodonta maryattiana Lea, 1840
+ Anodonta footiana Lea, 1840
+ Anodonta maryattana Lea, 1840
+ Anodonta nilssonii Küster, 1842
+ Anodonta sulcata Küster, 1842
+ Anodonta virens Lea, 1852
+ Anodonta linnaeana Lea, 1852
+ Anodonta opaca Lea, 1852
+ Anodonta lewisii Lea, 1857
+ Anodonta danielsii Lea, 1858
+ Anodonta texasensis Lea, 1859
+ Anodonta simpsoniana Lea, 1861
+ Anodonta leonensis Lea, 1862
+ Anodonta dallasiana Lea, 1863
+ Anodonta bealei Lea, 1863
+ Anodon subangulata J.G. Anthony, 1865
+ Anodon subinflata J.G. Anthony, 1865
+ Anodon imbricata J.G. Anthony, 1865
+ Anodon micans J.G. Anthony, 1865
+ Anodon opalina J.G. Anthony, 1865
+ Anodon mcnielii J.G. Anthony, 1866
+ Anodon subgibbosa J.G. Anthony, 1866
+ Anodon inornata J.G. Anthony, 1866
+ Anodon edentulus Sowerby, 1867
+ Margaron (Anodonta) benedictii Lea, 1870
+ Margaron (Anodonta) subglobosa Lea, 1870
+ Anodonta somersii Call, 1880
+ Anodonta hockingensis Call, 1880
+ Anodonta houghtonensis DeCamp, 1881
+ Anodonta dakota Frierson, 1910
+ Anodonta (Nayadina) venusta de Gregorio, 1914
+ Anodonta dakotana Utterback, 1915

Pyganodon lacustris (Lea, 1857)

Quadrula cylindrica cylindrica (Say, 1817)
+ Unio naviformis Lamarck, 1819
+ Unio (Eurynia) solenoides Rafinesque, 1820
+ Unio rugosus Chenu, 1862
+ Unio cilindricus de Gregorio, 1914
+ Unio cilindricus [sic] var. propetipicus de Gregorio, 1914
+ Unio cilindricus [sic] var. acrispatus de Gregorio, 1914

Quadrula fragosa (Conrad, 1835)
+ Unio rugosus Barnes, 1823
+ Margarita (Unio) fragrosus Lea, 1836
+ Unio tragosus Hanley, 1843

Quadrula metanevra (Rafinesque, 1820)
+ Unio nodosus Barnes, 1823
+ Unio metanever Lea, 1829
+ Unio metaniver Catlow & Reeve, 1845
+ Unio metaneurus Herrmannsen, 1849
+ Unio wardii Lea, 1862

Quadrula nodulata (Rafinesque, 1820)
+ Unio pustulatus Lea, 1831
+ Unio nodulatus Say, 1834

Quadrula pustulosa pustulosa (Lea, 1831)
+ Obliquaria (Quadrula) bullata Rafinesque, 1820
+ Obliquaria (Quadrula) retusa Rafinesque, 1820
+ Unio verrucosa Valenciennes, 1827
+ Unio premorsus Rafinesque, 1831
+ Unio nodulosus Say, 1834
+ Unio prasinus Conrad, 1834
+ Unio schoolcraftensis Lea, 1834
+ Unio bullatus Conrad, 1838
+ Unio dorfeuillianus Lea, 1838
+ Unio pernodosus Lea, 1845
+ Unio schoolcraftii Lea, 1870
+ Quadrula pustulosa asperatus Utterback, 1915

Quadrula quadrula (Rafinesque, 1820)
+ Unio lacrymosus Lea, 1828
+ Unio asperrimus Lea, 1831
+ Unio quadrulus Say, 1834
+ Quadrula lachrymosa Swainson, 1840
+ Unio quadratus Reeve, 1864
+ Unio lunulatus Pratt, 1876
+ Quadrula quadrula contraryensis Utterback, 1915
+ Quadrula (Quadrula) quadrula bullocki F.C. Baker, 1928

Simpsonaias ambigua (Say, 1825)
+ Unio hildrethianus Lea, 1834
+ Alasmidonta dubia Ferussac, 1835

Strophitus undulatus (Say, 1817)
+ Anodonta pensylvanica Lamarck, 1819
+ Anodon rugosus Swainson, 1822
+ Anodonta undulata Hildreth, 1828
+ Anodonta pennsylvanica Stark, 1828
+ Alasmodonta edentula Say, 1829
+ Anodon areolatus Swainson, 1829
+ Anodonta marginata Ferussac, 1835
+ Anodonta ferussaciana Ferussac, 1835
+ Anodonta virgata Conrad, 1836
+ Anodonta wardiana Lea, 1838
+ Anodonta pavonia Lea, 1838
+ Anodon unadilla DeKay, 1843
+ Anodonta tetragona Lea, 1845
+ Anodonta arkansensis Lea, 1852
+ Anodonta shaefferiana Lea, 1852
+ Anodonta schaefferiana Adams & Adams, 1857
+ Alasmodon rhombica J.G. Anthony, 1865
+ Anodon papyracea J.G. Anthony, 1865
+ Anodon annulatus Sowerby, 1867
+ Anodonta shafferiana Lea, 1870
+ Anodon quadriplicatus Sowerby, 1870
+ Anodonta salmonea Clessin, 1873
+ Anodonta shefferiana Simpson, 1900
+ Strophitus undulatus tennesseensis Frierson, 1927
+ Strophitus undulatus ovatus Frierson, 1927
+ Strophitus rugosus pepinensis F.C. Baker, 1928
+ Strophitus rugosus winnebagoensis F.C. Baker, 1928
+ Strophitus rugosus lacustris F.C. Baker, 1928

Toxolasma lividum (Rafinesque, 1831)
+ Unio glans Lea, 1831
+ Unio moestus Lea, 1841
+ Unio maestus Lea, 1842

Toxolasma parvum (Barnes, 1823)
+ Unio pertennis de Gregorio, 1914
+ Carunculina parva cahni F.C. Baker, 1927

Tritogonia verrucosa (Rafinesque, 1820)
+ Unio tuberculatus Barnes, 1823
+ Unio verrucosus Say, 1834
+ Unio (Theliderma) pustulata Swainson, 1840
+ Unio gigas Sowerby, 1867
+ Unio conjugans B.H. Wright, 1899
+ Tritogonia tuberculata var. obesa Simpson, 1900
+ Quadrula tritogonia Ortmann, 1909
+ Quadrula parkeri Geiser, 1911

Truncilla donaciformis (Lea, 1828)
+ Unio zigzag Lea, 1829
+ Unio nervosa Conrad, 1834
+ Unio zig-zag var. illius de Gregorio, 1914

Truncilla truncata Rafinesque, 1820
+ Truncilla truncata var. fusca Rafinesque, 1820
+ Truncilla truncata var. vermiculata Rafinesque, 1820
+ Unio elegans Lea, 1831
+ Unio truncatus Say, 1834
+ Unio elegans var. elegantopsis de Gregorio, 1914
+ Unio elegans var. magnelegans de Gregorio, 1914
+ Truncilla truncata var. lacustris F.C. Baker, 1928

Uniomerus declivis (Say, 1831)
+ Unio sayanus Conrad, 1838
+ Unio symmetricus Lea, 1845

Uniomerus tetralasmus (Say, 1831)
+ Unio camptodon Say, 1832
+ Unio declivis Conrad, 1836
+ Unio sayii Tappan, 1839
+ Unio parallelus Conrad, 1841
+ Unio rivularis Conrad, 1853
+ Unio porrectus Conrad, 1854
+ Unio subcroceus Conrad, 1854
+ Unio manubius Gould, 1855
+ Unio jamesianus Lea, 1857
+ Unio electrinus Reeve, 1865
+ Unio manubrius Paetel, 1890
+ Uniomerus tetralasma comptodon Utterback, 1915

Utterbackia imbecillis (Say, 1829)
+ Anodonta incerta Lea, 1834
+ Anodonta imbecilis Lea, 1836
+ Anodon horda Gould, 1855
+ Anodonta henryiana Lea, 1857
+ Anodonta hordeum Paetel, 1890
+ Anodonta phalena de Gregorio, 1914
+ Anodonta ohiensis Utterback, 1915
+ Utterbackia imbecillis fusca F.C. Baker, 1927

Venustaconcha ellipsiformis (Conrad, 1836)
+ Unio venustus Lea, 1838
+ Unio spatulatus Lea, 1845

Villosa fabalis (Lea, 1831)
+ Unio lapillus Say, 1831
+ Unio capillus Say, 1831
+ Unio donacopsis de Gregorio, 1914

Villosa iris (Lea, 1829)
+ Unio novieboraci Lea, 1838
+ Unio subrostratus Küster, 1861
+ Unio opalinus J.G. Anthony, 1866
+ Lampsilis (Ligumia) nervosa Frierson, 1927

Villosa lienosa (Conrad, 1834)
+ Unio saxeus Conrad, 1838
+ Unio gouldii Lea, 1845
+ Unio caliginosus Lea, 1845
+ Unio nigerrimus Lea, 1852
+ Unio concestator Lea, 1857
+ Unio intercedens Lea, 1857
+ Unio apicinus Lea, 1857
+ Unio fallax Lea, 1857
+ Unio prattii Lea, 1858
+ Unio porphyreus Lea, 1861
+ Unio contiguus Lea, 1862
+ Unio fuligo Reeve, 1865
+ Unio bicaelatus Reeve, 1865
+ Unio fontanus Conrad, 1866
+ Unio unicostatus B.H. Wright, 1899

Page updated: 10:26:25 Thu 04 Jul 2019

 
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