MUSSELpdb | subfamily Hyriinae

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Page last updated
Thu 29 May 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Hyrioidea | Hyriidae

subfamily HYRIINAE Swainson, 1840: cladomics
4 tribes, 1 genus

Cladomics Summary

number of species = 86
number of subtaxa (rank: tribe) = 5

analyzed in 90 cladograms in 46 publications
maximum number of terminals in an analysis = 13, mean = 2.72
maximum number of subtaxa (tribe) in an analysis = 5, mean = 2.17

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 51
maximum number of included terminals = 13, mean = 4.04
maximum number of subtaxa (tribe) = 5, mean = 3.06
proportion of cladograms that support basic monophyly = 0.47

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 47
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (tribe) taxon and at least one rigorous outgroup (subfamily).
maximum number of included terminals = 13, mean = 4.0
maximum number of subtaxa (tribe) = 5, mean = 3.13
proportion of cladograms that support rigorous monophyly = 0.43

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination

NJ tree

Santos-Neto et al., 2016 [Publ 3739], fig. 4

iT	tT	bi	riS	oT	roT	roS	iD
13	19	21	5	7	6	1	0.0

*** Top Monophyly Cladogram ***

                    +----- Diplodon_suavidicus__2 **
  +-----------------+                            
  |                 |  +-- Hyriini **               
  |                 +--+                         
  |                    +-- Castaliini **            
  |                                              
  |              +-------- Westralunio           
--+              |                               
  |  +-----------+     +-- Hyridella_drapeta **     
  |  |           |  +--+                         
  |  |           |  |  +-- Hyridella_depressa **    
  |  |           +--+                            
  |  |              |  +-- Diplodon_suavidicus__1 **
  +--+              +--+                         
     |                 +-- Diplodon_gratus **       
     |                                           
     |  +----------------- Echyridella **           
     |  |                                        
     +--+              +-- Cucumerunio **           
        |  +-----------+                         
        |  |           +-- Hyridella_australis **   
        +--+                                     
           |  +----------- Velesunio_ambiguus    
           +--+                                  
              |  +-------- Alathyria             
              +--+                               
                 |  +----- Velesunio_angasi      
                 +--+                            
                    |  +-- Lortiella             
                    +--+                         
                       +-- Microdontia

Huang et al., 2019 [Publ 3843], fig. S4

iT	tT	bi	riS	oT	roT	roS	iD
7	14	9	5	192	7	1	22.5

*** Top Sister-Group Cladogram ***

  +----------- Echyridella **       
  |                              
  +----------- Westralunio       
  |                              
  +----------- Hyridellini **       
  |                              
  |     +----- Rhipidodontini **    
  +-----+                        
--+     |  +-- Castaliini **        
  |     +--+                     
  |        +-- Hyriini **           
  |                              
  |     +----- Velesunio         
  |  +--+                        
  |  |  |  +-- Alathyria_jacksoni
  |  |  +--+                     
  +--+     +-- Alathyria_profuga 
     |                           
     |  +----- Alathyria_pertexta
     +--+                        
        |  +-- Lortiella         
        +--+                     
           +-- Microdontia

Santos-Neto et al., 2016 [Publ 3739], fig. 2a_b

iT	tT	bi	riS	oT	roT	roS	iD
13	19	20	5	7	6	1	11.0

                 +----- Diplodon_suavidicus__1 **
  +--------------+                            
  |              |  +-- Hyriini **               
  |              +--+                         
  |                 +-- Castaliini **            
  |                                           
  |           +-------- Westralunio           
  |           |                               
  +-----------+     +-- Hyridella_drapeta **     
  |           |  +--+                         
--+           |  |  +-- Hyridella_depressa **    
  |           +--+                            
  |              |  +-- Diplodon_gratus **       
  |              +--+                         
  |                 +-- Diplodon_suavidicus__2 **
  |                                           
  |                 +-- Hyridella_australis **   
  |  +--------------+                         
  |  |              +-- Cucumerunio **           
  +--+                                        
     |  +-------------- Echyridella **           
     +--+                                     
        |  +----------- Velesunio_ambiguus    
        +--+                                  
           |  +-------- Microdontia           
           +--+                               
              |  +----- Lortiella             
              +--+                            
                 |  +-- Alathyria             
                 +--+                         
                    +-- Velesunio_angasi

Marshall et al., 2014 [Publ 3593], fig. 1

iT	tT	bi	riS	oT	roT	roS	iD
7	10	8	3	7	3	1	25.0

  +-------- Echyridella **   
--+                       
  |  +----- Velesunioninae
  +--+                    
     |  +-- Hyridellini **   
     +--+                 
        +-- Rhipidodontini **

Huang et al., 2019 [Publ 3843], fig. S3

iT	tT	bi	riS	oT	roT	roS	iD
7	7	5	5	190	7	1	22.0

     +-------------- Westralunio       
  +--+                                 
  |  |  +----------- Echyridella **       
  |  +--+                              
  |     |  +-------- Hyridellini **       
  |     +--+                           
  |        |  +----- Rhipidodontini **    
  |        +--+                        
--+           |  +-- Castaliini **        
  |           +--+                     
  |              +-- Hyriini **           
  |                                    
  |           +----- Velesunio         
  |        +--+                        
  |        |  |  +-- Alathyria_jacksoni
  |        |  +--+                     
  +--------+     +-- Alathyria_profuga 
           |                           
           |  +----- Alathyria_pertexta
           +--+                        
              |  +-- Lortiella         
              +--+                     
                 +-- Microdontia

Graf et al., 2015 [Publ 3619], fig. 1

iT	tT	bi	riS	oT	roT	roS	iD
7	7	5	4	13	7	1	20.0

  +----------- Velesunioninae
--+                          
  |  +-------- Hyridellini **   
  +--+                       
     |  +----- Rhipidodontini **
     +--+                    
        |  +-- Hyriini **       
        +--+                 
           +-- Castaliini **

Huang et al., 2019 [Publ 3843], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
7	7	5	5	192	7	1	22.0

     +-------------- Westralunio       
  +--+                                 
  |  |  +----------- Echyridella **       
  |  +--+                              
  |     |  +-------- Hyridellini **       
  |     +--+                           
  |        |  +----- Rhipidodontini **    
  |        +--+                        
--+           |  +-- Castaliini **        
  |           +--+                     
  |              +-- Hyriini **           
  |                                    
  |           +----- Alathyria_jacksoni
  |        +--+                        
  |        |  |  +-- Alathyria_profuga 
  |        |  +--+                     
  +--------+     +-- Velesunio         
           |                           
           |  +----- Alathyria_pertexta
           +--+                        
              |  +-- Lortiella         
              +--+                     
                 +-- Microdontia

Santos-Neto et al., 2016 [Publ 3739], fig. 3_5

iT	tT	bi	riS	oT	roT	roS	iD
7	7	6	3	0	0	0	18.0

  +----- Rhipidodontini **
--+                    
  |  +-- Hyriini **       
  +--+                 
     +-- Castaliini **

Hoeh et al., 2009 [Publ 2716], fig. 3

iT	tT	bi	riS	oT	roT	roS	iD
6	6	4	4	41	3	1	25.5

  +----------- Velesunioninae
--+                          
  |  +-------- Echyridella **   
  +--+                       
     |  +----- Hyridellini **   
     +--+                    
        |  +-- Castaliini **    
        +--+                 
           +-- Rhipidodontini **

Graf & Cummings, 2006 [Publ 1968], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
6	6	4	4	41	3	1	25.5

  +----------- Velesunioninae
--+                          
  |  +-------- Echyridella **   
  +--+                       
     |  +----- Hyridellini **   
     +--+                    
        |  +-- Castaliini **    
        +--+                 
           +-- Rhipidodontini **

Hoeh et al., 2009 [Publ 2716], fig. 5

iT	tT	bi	riS	oT	roT	roS	iD
6	6	4	4	40	3	1	25.5

  +----------- Velesunioninae
--+                          
  |  +-------- Echyridella **   
  +--+                       
     |  +----- Hyridellini **   
     +--+                    
        |  +-- Castaliini **    
        +--+                 
           +-- Rhipidodontini **

Hoeh et al., 2009 [Publ 2716], fig. 4

iT	tT	bi	riS	oT	roT	roS	iD
6	6	4	4	40	3	1	25.5

  +----------- Velesunioninae
--+                          
  |  +-------- Echyridella **   
  +--+                       
     |  +----- Hyridellini **   
     +--+                    
        |  +-- Castaliini **    
        +--+                 
           +-- Rhipidodontini **

Walker et al., 2006 [Publ 2666], fig. 6

iT	tT	bi	riS	oT	roT	roS	iD
5	10	7	4	31	5	1	21.5

  +-------- Castaliini **         
  |                            
  +-------- Rhipidodontini **     
--+                            
  |  +----- Hyridella_depressa ** 
  |  |                         
  +--+----- Hyridella_australis **
     |                         
     |  +-- Echyridella **        
     +--+                      
        +-- Velesunioninae

Baker et al., 2004 [Publ 2685], fig. 2-3

iT	tT	bi	riS	oT	roT	roS	iD
5	8	9	4	4	3	1	25.0

     +----- Hyridella_depressa ** 
  +--+                         
  |  |  +-- Hyridella_australis **
  |  +--+                      
--+     +-- Rhipidodontini **     
  |                            
  |  +----- Velesunioninae     
  +--+                         
     |  +-- Echyridella **        
     +--+                      
        +-- Castaliini **

Baker et al., 2003 [Publ 2690], fig. 5

iT	tT	bi	riS	oT	roT	roS	iD
5	7	5	4	3	2	1	24.0

           +-- Hyridella_australis **
  +--------+                      
  |        +-- Rhipidodontini **     
--+                               
  |  +-------- Hyridella_depressa ** 
  +--+                            
     |  +----- Velesunioninae     
     +--+                         
        |  +-- Echyridella **        
        +--+                      
           +-- Castaliini **

Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2

iT	tT	bi	riS	oT	roT	roS	iD
5	5	4	4	6	1	1	26.5

  +-------------- Velesunioninae        
  |                                     
--+           +-- Rhipidodontini **        
  |  +--------+                         
  |  |        +-- Castaliini **            
  +--+                                  
     |  +-------- Hyridella_australis__2 **
     +--+                               
        |  +----- Hyridella_australis__1 **
        +--+                            
           |  +-- Echyridella **           
           +--+                         
              +-- Hyridella_depressa **

Santos-Neto et al., 2018 [Publ 3888], fig. 5

iT	tT	bi	riS	oT	roT	roS	iD
5	5	3	2	0	0	0	18.0

  +-- Hyriini **   
--+             
  +-- Castaliini **

Guerra et al., 2019 [Publ 4168], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
4	4	2	4	37	1	1	22.0

  +----------- Velesunioninae
--+                          
  |  +-------- Echyridella **   
  +--+                       
     |  +----- Rhipidodontini **
     +--+                    
        |  +-- Castaliini **    
        +--+                 
           +-- Hyriini **

Guerra et al., 2019 [Publ 4168], fig. 1

iT	tT	bi	riS	oT	roT	roS	iD
4	5	3	4	37	1	1	23.0

        +-- Echyridella **   
  +-----+                 
  |     +-- Velesunioninae
--+                       
  |  +----- Rhipidodontini **
  +--+                    
     |  +-- Castaliini **    
     +--+                 
        +-- Hyriini **

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

iT	tT	bi	riS	oT	roT	roS	iD
3	5	3	3	31	2	1	20.5

     +-- Castaliini **    
  +--+                 
  |  +-- Rhipidodontini **
--+                    
  |  +-- Echyridella **   
  +--+                 
     +-- Velesunioninae

Bolotov et al., 2017 [Publ 3761], fig. S2

iT	tT	bi	riS	oT	roT	roS	iD
3	3	1	3	136	6	1	20.0

  +-------- Velesunioninae
--+                       
  |  +----- Rhipidodontini **
  +--+                    
     |  +-- Castaliini **    
     +--+                 
        +-- Hyriini **

Hoeh et al., 2001 [Publ 2652], fig. 14f4

iT	tT	bi	riS	oT	roT	roS	iD
3	5	3	3	28	2	1	20.5

     +-- Castaliini **    
  +--+                 
  |  +-- Rhipidodontini **
--+                    
  |  +-- Echyridella **   
  +--+                 
     +-- Velesunioninae

Graf, 2002 [Publ 2651], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
3	3	1	3	15	1	1	23.5

  +-------- Velesunioninae
--+                       
  |  +----- Hyridellini **   
  +--+                    
     |  +-- Rhipidodontini **
     +--+                 
        +-- Castaliini **

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

iT	tT	bi	riS	oT	roT	roS	iD
3	5	3	3	28	2	1	20.5

     +-- Castaliini **    
  +--+                 
  |  +-- Rhipidodontini **
--+                    
  |  +-- Echyridella **   
  +--+                 
     +-- Velesunioninae

Hoeh et al., 2001 [Publ 2652], fig. 14f2

iT	tT	bi	riS	oT	roT	roS	iD
3	5	1	3	28	2	1	22.5

  +----- Rhipidodontini **
  |                    
  +----- Echyridella **   
--+                    
  +----- Lortiella     
  |                    
  |  +-- Castaliini **    
  +--+                 
     +-- Velesunio

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

iT	tT	bi	riS	oT	roT	roS	iD
3	5	2	3	28	2	1	23.5

     +-- Castaliini **    
  +--+                 
  |  +-- Velesunio     
--+                    
  |  +-- Echyridella **   
  |  |                 
  +--+-- Lortiella     
     |                 
     +-- Rhipidodontini **

Bolotov et al., 2017 [Publ 3761], fig. S1

iT	tT	bi	riS	oT	roT	roS	iD
3	3	1	3	136	6	1	20.0

  +-------- Velesunioninae
--+                       
  |  +----- Rhipidodontini **
  +--+                    
     |  +-- Castaliini **    
     +--+                 
        +-- Hyriini **

Bolotov et al., 2016 [Publ 3743], fig. 4

iT	tT	bi	riS	oT	roT	roS	iD
3	3	1	3	27	6	1	20.0

  +-------- Velesunioninae
--+                       
  |  +----- Rhipidodontini **
  +--+                    
     |  +-- Hyriini **       
     +--+                 
        +-- Castaliini **

Graf, 2000 [Publ 2647], fig. 1

iT	tT	bi	riS	oT	roT	roS	iD
3	4	2	3	14	1	1	23.0

  +-------- Castaliini **    
--+                       
  |  +----- Rhipidodontini **
  +--+                    
     |  +-- Hyridellini **   
     +--+                 
        +-- Velesunioninae

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iT	tT	bi	riS	oT	roT	roS	iD
3	3	1	3	33	1	1	20.0

  +-------- Velesunioninae
--+                       
  |  +----- Hyridellini **   
  +--+                    
     |  +-- Castaliini **    
     +--+                 
        +-- Hyriini **

Bolotov et al., 2016 [Publ 3743], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
3	3	1	3	27	6	1	20.0

  +-------- Velesunioninae
--+                       
  |  +----- Rhipidodontini **
  +--+                    
     |  +-- Hyriini **       
     +--+                 
        +-- Castaliini **

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iT	tT	bi	riS	oT	roT	roS	iD
3	3	1	3	33	1	1	20.0

  +-------- Velesunioninae
--+                       
  |  +----- Hyridellini **   
  +--+                    
     |  +-- Castaliini **    
     +--+                 
        +-- Hyriini **

Olivera-Hyde et al., 2020 [Publ 4249], fig. 10

iT	tT	bi	riS	oT	roT	roS	iD
3	3	1	1	0	0	0	20.0

-- (Castaliini

Konopleva et al., 2017 [Publ 3807], fig. 6

iT	tT	bi	riS	oT	roT	roS	iD
3	3	1	3	28	0	0	20.0

  +----- Rhipidodontini **
--+                    
  |  +-- Hyriini **       
  +--+                 
     +-- Castaliini **

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
3	3	1	3	37	1	1	20.0

  +-------- Velesunioninae
--+                       
  |  +----- Hyridellini **   
  +--+                    
     |  +-- Hyriini **       
     +--+                 
        +-- Castaliini **

Hoeh et al., 2002 [Publ 2653], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
3	5	3	3	28	2	1	20.0

     +-- Castaliini **    
  +--+                 
  |  +-- Rhipidodontini **
--+                    
  |  +-- Echyridella **   
  +--+                 
     +-- Velesunioninae

Combosch et al., 2017 [Publ 3806], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
2	2	0	2	13	1	1	--

  +----- Velesunioninae
--+                    
  |  +-- Hyriini **       
  +--+                 
     +-- Hyridellini **

Combosch et al., 2017 [Publ 3806], fig. 1

iT	tT	bi	riS	oT	roT	roS	iD
2	3	0	2	13	1	1	20.0

  +----- Hyriini **       
--+                    
  |  +-- Velesunioninae
  +--+                 
     +-- Hyridellini **

Wu et al., 2020 [Publ 4121], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
2	3	0	2	32	1	1	20.0

  +----- Hyriini **       
--+                    
  |  +-- Hyridellini **   
  +--+                 
     +-- Velesunioninae

Sano et al., 2020 [Publ 4138], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
2	3	0	2	67	1	1	20.0

  +----- Hyriini **       
--+                    
  |  +-- Hyridellini **   
  +--+                 
     +-- Velesunioninae

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

iT	tT	bi	riS	oT	roT	roS	iD
2	3	0	2	26	1	1	20.0

  +----- Hyriini **       
--+                    
  |  +-- Velesunioninae
  +--+                 
     +-- Hyridellini **

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

iT	tT	bi	riS	oT	roT	roS	iD
2	3	0	2	26	1	1	20.0

  +----- Hyriini **       
--+                    
  |  +-- Velesunioninae
  +--+                 
     +-- Hyridellini **

Sonowal et al., 2021 [Publ 4221], fig. 3a

iT	tT	bi	riS	oT	roT	roS	iD
2	3	0	2	13	1	1	21.0

  +-- Hyriini **       
  |                 
--+-- Hyridellini **   
  |                 
  +-- Velesunioninae

Sonowal et al., 2021 [Publ 4221], fig. 3b

iT	tT	bi	riS	oT	roT	roS	iD
2	3	0	2	13	1	1	21.0

  +-- Hyriini **       
  |                 
--+-- Velesunioninae
  |                 
  +-- Hyridellini **

Sano et al., 2020 [Publ 4138], fig. 4

iT	tT	bi	riS	oT	roT	roS	iD
2	3	0	2	67	1	1	20.0

  +----- Hyriini **       
--+                    
  |  +-- Hyridellini **   
  +--+                 
     +-- Velesunioninae

Wu et al., 2021 [Publ 4366], fig. 6

iT	tT	bi	riS	oT	roT	roS	iD
2	2	0	2	89	1	1	--

  +----- Velesunioninae
--+                    
  |  +-- Rhipidodontini **
  +--+                 
     +-- Castaliini **

Sonowal et al., 2021 [Publ 4221], fig. 4

iT	tT	bi	riS	oT	roT	roS	iD
2	3	0	2	14	1	1	20.0

  +----- Hyriini **       
--+                    
  |  +-- Velesunioninae
  +--+                 
     +-- Hyridellini **

Sano et al., 2020 [Publ 4138], fig. 3

iT	tT	bi	riS	oT	roT	roS	iD
2	3	0	2	67	1	1	20.0

  +----- Hyriini **       
--+                    
  |  +-- Hyridellini **   
  +--+                 
     +-- Velesunioninae

Wu et al., 2021 [Publ 4366], fig. 2_I

iT	tT	bi	riS	oT	roT	roS	iD
2	2	0	2	89	1	1	--

  +----- Velesunioninae
--+                    
  |  +-- Castaliini **    
  +--+                 
     +-- Rhipidodontini **

Huang et al., 2018 [Publ 3828], fig. 3_5

iT	tT	bi	riS	oT	roT	roS	iD
2	3	0	2	27	1	1	20.0

  +----- Hyriini **       
--+                    
  |  +-- Velesunioninae
  +--+                 
     +-- Hyridellini **

Wu et al., 2020 [Publ 4121], fig. S1

iT	tT	bi	riS	oT	roT	roS	iD
2	3	0	2	32	1	1	20.0

  +----- Hyriini **       
--+                    
  |  +-- Hyridellini **   
  +--+                 
     +-- Velesunioninae

Chase et al., 2018 [Publ 3861], fig. 1-M

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	10	0	0	--

-- Echyridella_menziesii

Guerra et al., 2017 [Publ 3772], fig. 2b

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	24	0	0	--

-- Echyridella_menziesii

Guerra et al., 2019 [Publ 4168], fig. 2-M

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	21	1	1	--

  +-- Echyridella **   
--+                 
  +-- Velesunioninae

Chase et al., 2018 [Publ 3847], fig. 5

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	9	0	0	--

-- Echyridella_menziesii

Chase et al., 2018 [Publ 3861], fig. 1

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	10	0	0	--

-- Echyridella_menziesii

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	56	0	0	--

-- Hyridella_australis

Walker et al., 2006 [Publ 2666], fig. 4

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	14	0	0	--

-- Hyridella

Walker et al., 2006 [Publ 2666], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	14	0	0	--

-- Hyridella

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	91	0	0	--

-- Echyridella_menziesii

Guerra et al., 2017 [Publ 3772], fig. 2b-M

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	7	0	0	--

-- Echyridella_menziesii

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	29	0	0	--

-- Hyridella_depressa

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	91	0	0	--

-- Echyridella_menziesii

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	120	1	1	--

  +-- Hyridellini **   
--+                 
  +-- Velesunioninae

Guerra et al., 2019 [Publ 4168], fig. 1-M

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	21	1	1	--

  +-- Echyridella **   
--+                 
  +-- Velesunioninae

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	79	1	1	--

  +-- Hyridellini **   
--+                 
  +-- Velesunioninae

Walker et al., 2006 [Publ 2666], fig. 3

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	14	0	0	--

-- Hyridella

Klunzinger et al., 2022 [Publ 4259], fig. 1

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	3	3	1	--

  +-- Hyridellini **   
--+                 
  +-- Velesunioninae

Guerra et al., 2017 [Publ 3772], fig. 2a

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	24	0	0	--

-- Echyridella_menziesii

Walker et al., 2006 [Publ 2666], fig. 6-M

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	13	0	0	--

-- Echyridella_menziesii

Huang et al., 2019 [Publ 3843], fig. 3-M

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	22	0	0	--

-- Echyridella_menziesii

Breton et al., 2011 [Publ 3376], fig. 3

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	36	0	0	--

-- Echyridella_menziesii

Chase et al., 2018 [Publ 3847], fig. 4

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	37	0	0	--

-- Echyridella_menziesii

Wu et al., 2021 [Publ 4366], fig. 4

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	68	0	0	--

-- Echyridella_menziesii

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	29	0	0	--

-- Hyridella_depressa

Wu et al., 2021 [Publ 4366], fig. 3-BI

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	73	0	0	--

-- Echyridella_menziesii

Wu et al., 2020 [Publ 4121], fig. 3

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	49	0	0	--

-- Echyridella_menziesii

Huang et al., 2018 [Publ 3828], fig. 6

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	42	0	0	--

-- Echyridella_menziesii

Hoeh et al., 2002 [Publ 2673], fig. 1-F

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	8	0	0	--

-- Echyridella_menziesii

Klunzinger et al., 2021 [Publ 4086], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	5	3	1	--

  +----- Westralunio
--+                 
  |  +-- Velesunio  
  +--+              
     +-- Hyridellini **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	95	0	0	--

-- Echyridella_menziesii

Hoeh et al., 2002 [Publ 2673], fig. 1-M

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	8	0	0	--

-- Echyridella_menziesii

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	56	0	0	--

-- Hyridella_australis

Wu et al., 2019 [Publ 3864], fig. 3

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	46	0	0	--

-- Echyridella_menziesii

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	95	0	0	--

-- Echyridella_menziesii

Wu et al., 2021 [Publ 4366], fig. 3-ML

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	73	0	0	--

-- Echyridella_menziesii

Wu et al., 2022 [Publ 4238], fig. 3

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	73	0	0	--

-- Echyridella_menziesii

Huang et al., 2019 [Publ 3843], fig. 3

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	56	0	0	--

-- Echyridella_menziesii

Guerra et al., 2017 [Publ 3772], fig. 2a-M

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	7	0	0	--

-- Echyridella_menziesii

Huang et al., 2018 [Publ 3828], fig. 6-M

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	16	0	0	--

-- Echyridella_menziesii

Page updated: 09:20:44 Thu 29 May 2025

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