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Page last updated
Sun 07 Sep 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

The MUSSEL Project Database has moved to a new server, with a new URL! The new address for this page is: https://musselpdb.org/db.php?ty=validfam_cld&id=Lanceolariini

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Unioninae

tribe LANCEOLARIINI Froufe, Lopes-Lima, & Bogan in Lopes-Lima et al., 2017: cladomics
     1 genus

Cladomics Summary

number of species = 15
number of subtaxa (rank: genus) = 1

analyzed in 94 cladograms in 58 publications
maximum number of terminals in an analysis = 8, mean = 2.26
maximum number of subtaxa (genus) in an analysis = 1, mean = 1.0

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 68
maximum number of included terminals = 8, mean = 2.74
maximum number of subtaxa (genus) = 1, mean = 1.0
proportion of cladograms that support basic monophyly = 0.9

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ tree

Sano et al., 2020 [Publ 4138], fig. 1

iT tT bi riS oT roT roS iD
3 22 5 1 31 19 4 0.0

*** Top Monophyly Cladogram *** 

  +----- Lanceolaria_grayii__1 **  
  |                             
  +----- Lanceolaria_grayii__2 **  
  |                             
  +----- Cristaria_plicata__1   
  |                             
  +----- Cristaria_plicata__2   
  |                             
  +----- Sinanodonta_lauta      
  |                             
  +----- Beringiana_japonica    
  |                             
  +----- Beringiana_fukuharai__1
  |                             
  +----- Beringiana_beringiana  
  |                             
  +----- Beringiana_fukuharai__2
  |                             
  +----- Beringiana_fukuharai__3
  |                             
  +----- Buldowskia_iwakawai__1 
  |                             
  +----- Buldowskia_iwakawai__2 
  |                             
  +----- Sinanodonta_tumens__1  
--+                             
  +----- Sinanodonta_tumens__2  
  |                             
  +----- Pletholophus           
  |                             
  +----- Anemina                
  |                             
  +----- Lanceolaria_lanceolata ** 
  |                             
  +----- Cristaria_plicata__3   
  |                             
  +----- Lanceolaria_gladiola **   
  |                             
  +----- Sinanodonta_angula     
  |                             
  +----- Lepidodesmini          
  |                             
  |  +-- Aculamprotula          
  +--+                          
  |  +-- Acuticostini           
  |                             
  |  +-- Cuneopsis              
  +--+                          
  |  +-- Nodularia              
  |                             
  |  +-- Schistodesmus          
  +--+                          
     +-- Inversiunio

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iT tT bi riS oT roT roS iD
8 8 6 1 113 62 4 36.0

*** Top Sister-Group Cladogram ***

  +----- Acuticostini 
  |                   
  +----- Lepidodesmini
--+                   
  +----- Unionini     
  |                   
  |  +-- Lanceolaria **  
  +--+                
     +-- Anodontini

Lopes-Lima et al., 2020 [Publ 4080], fig. 11

iT tT bi riS oT roT roS iD
8 8 6 1 4 2 1 36.0 
  +-- Unionini   
--+              
  +-- Lanceolaria **

Huang et al., 2019 [Publ 3843], fig. S1

iT tT bi riS oT roT roS iD
5 5 4 1 40 26 4 30.0 
  +-------------- Anemina      
  |                            
--+  +----------- Sinanodonta  
  |  |                         
  +--+  +-------- Acuticostini 
     |  |                      
     +--+     +-- Lepidodesmini
        |  +--+                
        |  |  +-- Cristaria    
        +--+                   
           |  +-- Lanceolaria **  
           +--+                
              +-- Unionini

Huang et al., 2019 [Publ 3843], fig. S4

iT tT bi riS oT roT roS iD
4 4 2 1 195 41 4 33.0 
  +-------- Acuticostini         
  |                              
  +-------- Alasmidonta          
  |                              
  +-------- Anodonta_anatina     
  |                              
  +-------- Anemina              
  |                              
  +-------- Anodonta_cygnea      
  |                              
  +-------- Cristaria            
  |                              
  +-------- Anodonta_nuttalliana 
  |                              
  +-------- Lasmigona            
  |                              
  +-------- Lepidodesmini        
  |                              
  +-------- Pseudanodonta        
  |                              
  +-------- Pyganodon            
  |                              
--+-------- Simpsonaias          
  |                              
  +-------- Strophitus           
  |                              
  +-------- Schistodesmus        
  |                              
  +-------- Utterbackia          
  |                              
  +-------- Lanceolaria **          
  |                              
  +-------- Unio                 
  |                              
  +-------- Aculamprotula        
  |                              
  |     +-- Pletholophus         
  +-----+                        
  |     +-- Sinanodonta          
  |                              
  |  +----- Cuneopsis_celtiformis
  |  |                           
  |  +----- Tchangsinaia         
  |  |                           
  +--+----- Cuneopsis_rufescens  
     |                           
     +----- Nodularia            
     |                           
     |  +-- Cuneopsis_szechenyii 
     +--+                        
        +-- Cuneopsis_heudei

Huang et al., 2019 [Publ 3843], fig. 2

iT tT bi riS oT roT roS iD
4 4 2 1 195 41 4 33.0 
  +-------- Acuticostini 
  |                      
--+  +----- Lepidodesmini
  |  |                   
  +--+----- Unionini     
     |                   
     |  +-- Lanceolaria **  
     +--+                
        +-- Anodontini

Huang et al., 2019 [Publ 3843], fig. 1

iT tT bi riS oT roT roS iD
4 4 2 1 39 25 4 33.0 
  +----------------------------------- Anemina              
  |                                                         
--+  +-------------------------------- Acuticostini         
  |  |                                                      
  +--+  +----------------------------- Sinanodonta          
     |  |                                                   
     +--+  +-------------------------- Aculamprotula        
        |  |                                                
        +--+                       +-- Cristaria            
           |  +--------------------+                        
           |  |                    +-- Lanceolaria **          
           +--+                                             
              |  +-------------------- Lepidodesmini        
              +--+                                          
                 |  +----------------- Schistodesmus        
                 +--+                                       
                    |  +-------------- Nodularia            
                    +--+                                    
                       |  +----------- Cuneopsis_rufescens  
                       +--+                                 
                          |  +-------- Tchangsinaia         
                          +--+                              
                             |  +----- Cuneopsis_celtiformis
                             +--+                           
                                |  +-- Cuneopsis_szechenyii 
                                +--+                        
                                   +-- Cuneopsis_heudei

Huang et al., 2019 [Publ 3843], fig. S3

iT tT bi riS oT roT roS iD
4 4 2 1 193 40 4 33.0 
  +----------- Acuticostini 
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lepidodesmini
     +--+                   
        |  +-- Lanceolaria **  
        +--+                
           +-- Anodontini

Wu et al., 2018 [Publ 3836], fig. 2

iT tT bi riS oT roT roS iD
4 4 2 1 49 29 4 33.0 
  +-------- Lepidodesmini
  |                      
--+-------- Unionini     
  |                      
  |  +----- Anodontini   
  +--+                   
     |  +-- Acuticostini 
     +--+                
        +-- Lanceolaria **

Wu et al., 2023 [Publ 4296], fig. 4

iT tT bi riS oT roT roS iD
3 3 1 1 20 18 3 32.0 
  +-------- Acuticostini
--+                     
  |  +----- Lanceolaria ** 
  +--+                  
     |  +-- Anodontini  
     +--+               
        +-- Unionini

Kim et al., 2021 [Publ 4197], fig. 1

iT tT bi riS oT roT roS iD
3 3 1 1 14 13 3 32.0 
  +-------------- Aculamprotula
  |                            
--+           +-- Acuticostini 
  |  +--------+                
  |  |        +-- Lanceolaria **  
  +--+                         
     |  +-------- Unio         
     +--+                      
        |  +----- Nodularia    
        +--+                   
           |  +-- Schistodesmus
           +--+                
              +-- Cuneopsis

Fukata & Iigo, 2019 [Publ 4108], fig. 1

iT tT bi riS oT roT roS iD
3 3 1 1 13 13 2 32.0 
           +----- Aculamprotula  
  +--------+                     
  |        |  +-- Acuticostini   
  |        +--+                  
--+           +-- Lanceolaria **    
  |                              
  |  +----------- Unio           
  |  |                           
  +--+  +-------- Pseudocuneopsis
     |  |                        
     +--+     +-- Schistodesmus  
        |  +--+                  
        |  |  +-- Cuneopsis      
        +--+                     
           |  +-- Tchangsinaia   
           +--+                  
              +-- Nodularia

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iT tT bi riS oT roT roS iD
3 3 1 1 92 43 4 32.0 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Wu et al., 2022 [Publ 4238], fig. 2d

iT tT bi riS oT roT roS iD
3 3 1 1 35 35 4 32.0 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lanceolaria **  
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Anodontini

Wu et al., 2022 [Publ 4238], fig. 3

iT tT bi riS oT roT roS iD
3 3 1 1 70 32 4 32.0 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Huang et al., 2019 [Publ 3843], fig. 3

iT tT bi riS oT roT roS iD
3 3 1 1 53 23 4 32.0 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lanceolaria **  
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Anodontini

Wu et al., 2021 [Publ 4366], fig. 3-BI

iT tT bi riS oT roT roS iD
3 3 1 1 70 30 4 32.0 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iT tT bi riS oT roT roS iD
3 3 2 1 77 25 2 32.0 
  +----- Unionini   
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Anodontini

Liu et al., 2023 [Publ 4310], fig. 5

iT tT bi riS oT roT roS iD
3 21 14 1 21 19 3 43.0 
  +-------- Acuticostini          
  |                               
--+  +----- Lanceolaria_grayii **    
  |  |                            
  |  +----- Lanceolaria_gladiola **  
  +--+                            
     +----- Lanceolaria_lanceolata **
     |                            
     |  +-- Anodontini            
     +--+                         
        +-- Unionini

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iT tT bi riS oT roT roS iD
3 3 1 1 92 43 4 32.0 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Liu et al., 2023 [Publ 4310], fig. 4

iT tT bi riS oT roT roS iD
3 21 19 1 21 19 3 46.0 
  +----------- Acuticostini          
  |                                  
--+        +-- Lanceolaria_lanceolata **
  |  +-----+                         
  |  |     +-- Lanceolaria_grayii **    
  +--+                               
     |  +----- Lanceolaria_gladiola **  
     +--+                            
        |  +-- Anodontini            
        +--+                         
           +-- Unionini

Hou et al., 2024 [Publ 4400], fig. 4

iT tT bi riS oT roT roS iD
3 3 1 1 73 69 3 32.0 
  +-------- Lepidodesmini
--+                      
  |  +----- Unionini     
  +--+                   
     |  +-- Lanceolaria **  
     +--+                
        +-- Anodontini

Wu et al., 2021 [Publ 4366], fig. 2_I

iT tT bi riS oT roT roS iD
3 3 1 1 88 23 4 32.0 
  +-------- Acuticostini 
  |                      
--+     +-- Lepidodesmini
  |  +--+                
  |  |  +-- Unionini     
  +--+                   
     |  +-- Lanceolaria **  
     +--+                
        +-- Anodontini

Dai et al., 2025 [Publ 4440], fig. 3

iT tT bi riS oT roT roS iD
3 3 1 1 85 41 4 32.0 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Xiang et al., 2025 [Publ 4433], fig. 7

iT tT bi riS oT roT roS iD
3 3 1 1 10 9 1 32.0 
  +-- Lanceolaria **
--+              
  +-- Anodontini

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iT tT bi riS oT roT roS iD
3 3 1 1 53 30 4 33.0 
  +-------- Unionini     
  |                      
--+     +-- Lepidodesmini
  |  +--+                
  |  |  +-- Lanceolaria **  
  +--+                   
     |  +-- Acuticostini 
     +--+                
        +-- Anodontini

Wu et al., 2021 [Publ 4366], fig. 6

iT tT bi riS oT roT roS iD
3 3 1 1 88 23 4 32.0 
  +----------- Acuticostini 
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lanceolaria **  
     +--+                   
        |  +-- Lepidodesmini
        +--+                
           +-- Anodontini

Wu et al., 2021 [Publ 4366], fig. 4

iT tT bi riS oT roT roS iD
3 3 1 1 65 29 4 32.0 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Wu et al., 2024 [Publ 4379], fig. 5

iT tT bi riS oT roT roS iD
3 3 1 1 50 48 3 32.0 
  +-------- Lepidodesmini
--+                      
  |  +----- Unionini     
  +--+                   
     |  +-- Lanceolaria **  
     +--+                
        +-- Anodontini

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iT tT bi riS oT roT roS iD
3 3 1 1 50 24 4 32.0 
  +----------- Lepidodesmini
--+                         
  |  +-------- Lanceolaria **  
  +--+                      
     |  +----- Unionini     
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Anodontini

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iT tT bi riS oT roT roS iD
3 3 1 1 88 43 4 32.0 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iT tT bi riS oT roT roS iD
3 3 1 1 88 43 4 32.0 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Wu et al., 2021 [Publ 4366], fig. 3-ML

iT tT bi riS oT roT roS iD
3 3 1 1 70 30 4 32.0 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Wu et al., 2020 [Publ 4121], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 1 47 25 4 -- 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Froufe et al., 2019 [Publ 3936], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 1 42 20 3 -- 
  +-------- Lepidodesmini
--+                      
  |  +----- Unionini     
  +--+                   
     |  +-- Lanceolaria **  
     +--+                
        +-- Anodontini

Sano et al., 2020 [Publ 4138], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 1 66 33 3 -- 
                 +-- Anemina      
  +--------------+                
  |              +-- Buldowskia   
  |                               
--+           +----- Sinanodonta  
  |  +--------+                   
  |  |        |  +-- Pletholophus 
  |  |        +--+                
  +--+           +-- Beringiana   
     |                            
     |  +----------- Cristaria    
     |  |                         
     +--+        +-- Anodontina   
        |  +-----+                
        |  |     +-- Alasmidontina
        +--+                      
           |  +----- Lanceolaria **  
           +--+                   
              |  +-- Lepidodesmini
              +--+                
                 +-- Unionini

Chen et al., 2023 [Publ 4302], fig. 1

iT tT bi riS oT roT roS iD
2 26 0 1 38 29 4 42.0 
           +-- Anemina               
  +--------+                         
  |        +-- Amuranodonta          
  |                                  
  |     +----- Pletholophus          
--+  +--+                            
  |  |  |  +-- Sinanodonta           
  |  |  +--+                         
  |  |     +-- Beringiana            
  |  |                               
  +--+  +----- Lanceolaria_gladiola **  
     |  |                            
     |  +----- Acuticostini          
     |  |                            
     |  +----- Lepidodesmini         
     |  |                            
     +--+----- Lanceolaria_oxyrhyncha **
        |                            
        +----- Cristaria             
        |                            
        +----- Unionini              
        |                            
        |  +-- Alasmidontina         
        +--+                         
           +-- Anodontina

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 1 30 16 2 -- 
  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Bolotov et al., 2017 [Publ 3761], fig. S2

iT tT bi riS oT roT roS iD
2 2 0 1 137 25 2 -- 
  +----------- Unionini            
  |                                
--+  +-------- Lanceolaria **         
  |  |                             
  |  +-------- Cristariina         
  +--+                             
     |  +----- Anodonta_anatina    
     |  |                          
     |  +----- Pseudanodonta       
     +--+                          
        +----- Alasmidontina       
        |                          
        |  +-- Anodonta_cygnea     
        +--+                       
           +-- Anodonta_nuttalliana

Wu et al., 2019 [Publ 3864], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 1 44 24 4 -- 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Huang et al., 2002 [Publ 2657], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 1 32 12 4 -- 
  +-------- Strophitus   
  |                      
  +-------- Schistodesmus
  |                      
  +-------- Lepidodesmini
  |                      
  +-------- Acuticostini 
  |                      
  +-------- Cristariina  
--+                      
  +-------- Lanceolaria **  
  |                      
  |     +-- Nodularia    
  +-----+                
  |     +-- Tchangsinaia 
  |                      
  |  +----- Utterbackia  
  |  |                   
  +--+----- Utterbackiana
     |                   
     |  +-- Alasmidonta  
     +--+                
        +-- Lasmigona

Sano et al., 2017 [Publ 3799], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 1 31 19 4 -- 
  +-------------- Anodontini   
  |                            
  |           +-- Tchangsinaia 
--+  +--------+                
  |  |        +-- Nodularia    
  |  |                         
  +--+        +-- Schistodesmus
     |  +-----+                
     |  |     +-- Acuticostini 
     +--+                      
        |  +----- Inversiunio  
        +--+                   
           |  +-- Lepidodesmini
           +--+                
              +-- Lanceolaria **

Zhou et al., 2007 [Publ 2658], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 1 19 11 3 -- 
     +----- Aculamprotula
  +--+                   
  |  |  +-- Tchangsinaia 
  |  +--+                
--+     +-- Nodularia    
  |                      
  |  +----- Schistodesmus
  +--+                   
     |  +-- Lanceolaria **  
     +--+                
        +-- Acuticostini

Wang et al., 2018 [Publ 3865], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 1 29 15 3 -- 
  +-------- Unionini    
--+                     
  |  +----- Anodontini  
  +--+                  
     |  +-- Acuticostini
     +--+               
        +-- Lanceolaria **

Bolotov et al., 2017 [Publ 3761], fig. S1

iT tT bi riS oT roT roS iD
2 2 0 1 137 25 2 -- 
  +----- Unionini   
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Anodontini

Fonseca et al., 2016 [Publ 3775], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 1 29 16 2 -- 
  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Wang et al., 2018 [Publ 3846], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 1 29 15 2 -- 
  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iT tT bi riS oT roT roS iD
2 2 0 1 36 14 4 -- 
  +----------------- Anodontini   
  |                               
  |              +-- Acuticostini 
--+  +-----------+                
  |  |           +-- Lanceolaria **  
  |  |                            
  +--+           +-- Lepidodesmini
     |  +--------+                
     |  |        +-- Aculamprotula
     +--+                         
        |  +-------- Unio         
        +--+                      
           |  +----- Schistodesmus
           +--+                   
              |  +-- Nodularia    
              +--+                
                 +-- Tchangsinaia

Dai et al., 2025 [Publ 4440], fig. 2

iT tT bi riS oT roT roS iD
2 27 0 1 43 34 4 42.0 
              +-- Anemina               
  +-----------+                         
  |           +-- Amuranodonta          
  |                                     
  |        +----- Pletholophus          
--+  +-----+                            
  |  |     |  +-- Sinanodonta           
  |  |     +--+                         
  |  |        +-- Beringiana            
  +--+                                  
     |        +-- Alasmidontina         
     |  +-----+                         
     |  |     +-- Anodontina            
     |  |                               
     +--+  +----- Lanceolaria_gladiola **  
        |  |                            
        |  +----- Lanceolaria_oxyrhyncha **
        |  |                            
        +--+----- Cristaria             
           |                            
           |  +-- Acuticostini          
           |  |                         
           +--+-- Lepidodesmini         
              |                         
              +-- Unionini

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 1 34 13 4 -- 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lanceolaria **  
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Anodontini

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iT tT bi riS oT roT roS iD
2 2 0 1 74 24 2 -- 
  +----- Unionini   
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Anodontini

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iT tT bi riS oT roT roS iD
2 2 0 1 46 20 3 -- 
  +-------- Lepidodesmini
--+                      
  |  +----- Unionini     
  +--+                   
     |  +-- Lanceolaria **  
     +--+                
        +-- Anodontini

Riccardi et al., 2020 [Publ 4089], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 1 17 15 2 -- 
  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Zhang et al., 2004 [Publ 2735], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 1 12 10 3 -- 
  +-------- Anodontini  
--+                     
  |  +----- Unionini    
  +--+                  
     |  +-- Acuticostini
     +--+               
        +-- Lanceolaria **

Xue et al., 2016 [Publ 3659], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 1 13 9 2 -- 
  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Froufe et al., 2019 [Publ 3936], fig. 6

iT tT bi riS oT roT roS iD
2 2 0 1 42 20 3 -- 
  +-------- Lepidodesmini
--+                      
  |  +----- Unionini     
  +--+                   
     |  +-- Lanceolaria **  
     +--+                
        +-- Anodontini

Chen et al., 2023 [Publ 4302], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 1 38 29 4 -- 
  +-------- Acuticostini 
  |                      
--+  +----- Lepidodesmini
  |  |                   
  +--+----- Unionini     
     |                   
     |  +-- Lanceolaria **  
     +--+                
        +-- Anodontini

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 1 21 21 3 -- 
  +-------- Anodontini   
--+                      
  |  +----- Lanceolaria **  
  +--+                   
     |  +-- Lepidodesmini
     +--+                
        +-- Unionini

Huang et al., 2002 [Publ 2657], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 1 32 12 4 -- 
  +----------- Anodontini   
  |                         
  |     +----- Acuticostini 
--+  +--+                   
  |  |  |  +-- Nodularia    
  |  |  +--+                
  +--+     +-- Tchangsinaia 
     |                      
     |  +----- Lanceolaria **  
     +--+                   
        |  +-- Schistodesmus
        +--+                
           +-- Lepidodesmini

Sano et al., 2020 [Publ 4138], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 1 66 33 3 -- 
                 +-- Anemina      
  +--------------+                
  |              +-- Buldowskia   
  |                               
--+           +----- Sinanodonta  
  |  +--------+                   
  |  |        |  +-- Pletholophus 
  |  |        +--+                
  +--+           +-- Beringiana   
     |                            
     |  +----------- Cristaria    
     |  |                         
     +--+        +-- Anodontina   
        |  +-----+                
        |  |     +-- Alasmidontina
        +--+                      
           |  +----- Lepidodesmini
           +--+                   
              |  +-- Lanceolaria **  
              +--+                
                 +-- Unionini

Bolotov et al., 2017 [Publ 3764], fig. S1

iT tT bi riS oT roT roS iD
2 17 0 1 123 25 2 37.0 
  +----- Unionini             
  |                           
  |  +-- Alasmidonta          
  |  |                        
  |  +-- Anodonta_anatina     
  |  |                        
  |  +-- Anodonta_cygnea      
  |  |                        
  |  +-- Anodonta_nuttalliana 
--+  |                        
  |  +-- Lasmigona            
  |  |                        
  |  +-- Pseudanodonta        
  |  |                        
  |  +-- Pyganodon            
  |  |                        
  |  +-- Simpsonaias          
  |  |                        
  |  +-- Strophitus           
  +--+                        
     +-- Sinanodonta_lauta    
     |                        
     +-- Cristaria_plicata    
     |                        
     +-- Cristaria_radiata    
     |                        
     +-- Pletholophus         
     |                        
     +-- Sinanodonta_lucida   
     |                        
     +-- Sinanodonta_jourdyi  
     |                        
     +-- Lanceolaria_gladiola ** 
     |                        
     +-- Lanceolaria_grayii__1 **
     |                        
     +-- Lanceolaria_grayii__2 **

Sano et al., 2020 [Publ 4138], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 1 66 33 3 -- 
                 +-- Anemina      
  +--------------+                
  |              +-- Buldowskia   
  |                               
--+           +----- Pletholophus 
  |  +--------+                   
  |  |        |  +-- Beringiana   
  |  |        +--+                
  +--+           +-- Sinanodonta  
     |                            
     |  +----------- Cristaria    
     |  |                         
     +--+        +-- Anodontina   
        |  +-----+                
        |  |     +-- Alasmidontina
        +--+                      
           |  +----- Lanceolaria **  
           +--+                   
              |  +-- Lepidodesmini
              +--+                
                 +-- Unionini

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iT tT bi riS oT roT roS iD
2 2 0 1 74 24 2 -- 
  +----- Unionini   
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Anodontini

Bolotov et al., 2020 [Publ 4084], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 1 34 17 2 -- 
  +----- Unionini   
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Anodontini

Ouyang et al., 2011 [Publ 3408], fig. 1

iT tT bi riS oT roT roS iD
2 4 0 1 11 7 3 32.0 
  +----------- Anodontini          
  |                                
--+        +-- Nodularia           
  |  +-----+                       
  |  |     +-- Cuneopsis           
  +--+                             
     |  +----- Lanceolaria_grayii **  
     |  |                          
     +--+----- Acuticostini        
        |                          
        |  +-- Lanceolaria_gladiola **
        +--+                       
           +-- Aculamprotula

Dai et al., 2025 [Publ 4440], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 1 43 34 4 -- 
           +----- Pletholophus 
  +--------+                   
  |        |  +-- Sinanodonta  
  |        +--+                
  |           +-- Beringiana   
--+                            
  |  +----------- Cristaria    
  |  |                         
  |  |        +-- Alasmidontina
  +--+--------+                
     |        +-- Anodontina   
     |                         
     |  +-------- Lanceolaria **  
     +--+                      
        |  +----- Lepidodesmini
        +--+                   
           |  +-- Acuticostini 
           +--+                
              +-- Unionini

Bolotov et al., 2018 [Publ 3826], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 1 81 15 2 -- 
  +----- Unionini   
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Anodontini

Aunins et al., 2018 [Publ 3863], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 1 38 24 4 -- 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Dai et al., 2023 [Publ 4285], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 52 16 4 -- 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Anodontini   
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Lanceolaria **

Guerra et al., 2019 [Publ 4168], fig. 2-M

iT tT bi riS oT roT roS iD
1 1 0 1 21 9 2 -- 
  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_AHE

iT tT bi riS oT roT roS iD
1 1 0 1 21 21 2 -- 
  +----- Unionini   
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Anodontini

Dai et al., 2024 [Publ 4386], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 39 25 4 -- 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lanceolaria **  
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Anodontini

Dai et al., 2024 [Publ 4386], fig. 5-BI

iT tT bi riS oT roT roS iD
1 1 0 1 20 15 4 -- 
  +----------------- Acuticostini    
  |                                  
  +----------------- Cristariina     
  |                                  
  +----------------- Anodontina      
--+                                  
  |  +-------------- Lanceolaria **     
  |  |                               
  |  |           +-- Aculamprotula   
  |  +-----------+                   
  +--+           +-- Lepidodesmini   
     |                               
     |  +----------- Unio            
     |  |                            
     +--+  +-------- Middendorffinaia
        |  |                         
        +--+  +----- Cuneopsis       
           |  |                      
           +--+----- Nodularia       
              |                      
              |  +-- Inversiunio     
              +--+                   
                 +-- Schistodesmus

Dai et al., 2024 [Publ 4386], fig. 5-ML

iT tT bi riS oT roT roS iD
1 1 0 1 20 15 4 -- 
  +----------- Anodontina   
  |                         
--+        +-- Acuticostini 
  |  +-----+                
  |  |     +-- Cristariina  
  +--+                      
     |  +----- Lepidodesmini
     +--+                   
        |  +-- Lanceolaria **  
        +--+                
           +-- Unionini

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 56 7 2 -- 
  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Zhou et al., 2017 [Publ 3788], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 15 5 2 -- 
  +-- Lanceolaria **
--+              
  +-- Unionini

Wu et al., 2024 [Publ 4388], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 53 17 3 -- 
  +-------- Lepidodesmini
--+                      
  |  +----- Unionini     
  +--+                   
     |  +-- Lanceolaria **  
     +--+                
        +-- Anodontini

Guerra et al., 2019 [Publ 4168], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 40 12 2 -- 
  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Plazzi et al., 2011 [Publ 3676], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 17 8 2 -- 
-- Lanceolaria_grayii

Huang et al., 2019 [Publ 3843], fig. 3-M

iT tT bi riS oT roT roS iD
1 1 0 1 22 9 2 -- 
  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iT tT bi riS oT roT roS iD
1 1 0 1 56 7 2 -- 
  +----- Lanceolaria **
--+                 
  |  +-- Unionini   
  +--+              
     +-- Anodontini

Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 18 0 0 -- 
-- Lanceolaria_grayii

Guerra et al., 2019 [Publ 4168], fig. 1-M

iT tT bi riS oT roT roS iD
1 1 0 1 21 9 2 -- 
  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Cha et al., 2018 [Publ 3916], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 37 20 3 -- 
  +-------- Lepidodesmini
--+                      
  |  +----- Unionini     
  +--+                   
     |  +-- Lanceolaria **  
     +--+                
        +-- Anodontini

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_MtG

iT tT bi riS oT roT roS iD
1 1 0 1 21 21 2 -- 
  +----- Unionini   
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Anodontini

Sano et al., 2017 [Publ 3799], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 20 12 2 -- 
  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Mabuchi & Nishida, 2020 [Publ 4120], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 11 7 2 -- 
  +-------- Anodontini 
--+                    
  |  +----- Nodularia  
  +--+                 
     |  +-- Lanceolaria **
     +--+              
        +-- Inversiunio

Burzynski & Soroka, 2018 [Publ 3862], fig. 5b

iT tT bi riS oT roT roS iD
1 1 0 1 9 9 2 -- 
  +----- Lanceolaria **
--+                 
  |  +-- Anodontini 
  +--+              
     +-- Unionini

Kano et al., 2019 [Publ 4194], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 9 5 2 -- 
  +-- Lanceolaria **
--+              
  +-- Unionini

Liu et al., 2017 [Publ 3939], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 14 13 3 -- 
  +-- Lanceolaria **
--+              
  +-- Unionini

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

iT tT bi riS oT roT roS iD
1 1 0 1 19 9 2 -- 
  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Guerra et al., 2019 [Publ 4168], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 40 12 2 -- 
  +-------- Aculamprotula
  |                      
--+     +-- Schistodesmus
  |  +--+                
  |  |  +-- Unio         
  +--+                   
     |  +-- Lanceolaria **  
     +--+                
        +-- Anodontini

Dai et al., 2024 [Publ 4358], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 51 26 4 -- 
  +----------- Lepidodesmini
--+                         
  |  +-------- Unionini     
  +--+                      
     |  +----- Lanceolaria **  
     +--+                   
        |  +-- Acuticostini 
        +--+                
           +-- Anodontini

Wu et al., 2022 [Publ 4238], fig. 2e

iT tT bi riS oT roT roS iD
1 1 0 1 16 16 2 -- 
  +----- Anodontini 
--+                 
  |  +-- Lanceolaria **
  +--+              
     +-- Unionini

Page updated: 08:15:36 Sun 07 Sep 2025
 
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