Bivalvia | Unionoida | Unionoidea | Unionidae | Ambleminae | Quadrulini genus Uniomerus Conrad, 1853: cladomics 5 valid species
Cladomics Summary
number of species = 5
analyzed in 46 cladograms in 31 publications
maximum number of terminals in an analysis = 4, mean = 1.17
Basic Monophyly
number of cladograms that test basic monophyly (i.e., more than 1 included species) = 5
maximum number of included terminals = 4, mean = 2.6
proportion of cladograms that support basic monophyly = 1.0
Neemuchwala et al., 2023 [Publ 4292], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
4 |
23 |
18 |
2 |
0.0 |
*** Top Monophyly Cladogram *** *** Top Sister-Group Cladogram ***
+-------------- Megalonaias
|
| +-------- Uniomerus_carolinianus **
--+ +--+
| | | +----- Uniomerus_columbensis **
| | +--+
| | | +-- Uniomerus_tetralasmus **
+--+ +--+
| +-- Uniomerus_declivis **
|
| +-------- Theliderma
+--+
| +----- Cyclonaias
+--+
| +-- Quadrula
+--+
+-- Pustulosa
Lopes-Lima et al., 2019 [Publ 3901], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
0 |
3 |
25 |
15 |
2 |
2.0 |
+----- Megalonaias
|
+--------------------+ +-- Uniomerus_carolinianus **
| | |
| +--+-- Uniomerus_declivis **
| |
--+ +-- Uniomerus_tetralasmus **
|
| +----------------------- Quadrula_quadrula
| |
| | +-- Quadrula_verrucosa
+--+ +-----------------+
| | +-- Quadrula_nobilis
| |
+--+ +----------------- Theliderma
| |
| | +----- Pustulosa_kieneriana
+--+ +--------+
| | | +-- Pustulosa_infucata
| | +--+
+--+ +-- Pustulosa_kleiniana
|
| +----------- Cyclonaias
| |
+--+ +-- Pustulosa_pustulosa
| +-----+
| | +-- Pustulosa_succissa
+--+
| +----- Pustulosa_nodulata
+--+
| +-- Pustulosa_necki
+--+
+-- Pustulosa_petrina
Inoue et al., 2018 [Publ 3858], fig. 1_3a-c
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
102 |
5 |
1 |
-- |
+-- Uniomerus_declivis **
+--+
| +-- Uniomerus_tetralasmus **
|
| +-- Cyclonaias
--+ |
| +-- Pustulosa_pustulosa__1
| |
| +-- Theliderma
+--+
+-- Pustulosa_pustulosa__2
|
+-- Quadrula_quadrula
|
+-- Quadrula_verrucosa
Campbell & Lydeard, 2012 [Publ 3309], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
71 |
9 |
1 |
-- |
+-- Uniomerus_declivis **
+--------------+
| +-- Uniomerus_tetralasmus **
|
--+ +-- Quadrula_verrucosa
| +-----------+
| | +-- Quadrula_nobilis
| |
+--+ +----------- Quadrula_quadrula
| |
| +----------- Theliderma
+--+
| +-- Pustulosa_infucata
| +-----+
| | +-- Pustulosa_kleiniana
+--+
| +----- Cyclonaias
+--+
| +-- Pustulosa_pustulosa
+--+
+-- Pustulosa_succissa
Campbell & Lydeard, 2012 [Publ 3309], fig. 1-2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
71 |
9 |
1 |
-- |
+-- Uniomerus_declivis **
+-----------------+
| +-- Uniomerus_tetralasmus **
--+
| +----------------- Theliderma
| |
+--+ +-------------- Quadrula_verrucosa
| |
| | +-- Quadrula_quadrula
+--+ +--------+
| | +-- Quadrula_nobilis
| |
+--+ +-- Pustulosa_infucata
| +-----+
| | +-- Pustulosa_kleiniana
+--+
| +----- Cyclonaias
+--+
| +-- Pustulosa_pustulosa
+--+
+-- Pustulosa_succissa
Campbell et al., 2005 [Publ 2660], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
101 |
10 |
2 |
-- |
+-------------------- Megalonaias
|
--+ +----------------- Uniomerus_declivis **
| |
+--+ +-------------- Theliderma
| |
+--+ +----------- Quadrula_quadrula
| |
| | +-- Quadrula_nobilis
+--+ +-----+
| | +-- Quadrula_verrucosa
| |
+--+ +----- Pustulosa_kieneriana
| |
| | +-- Cyclonaias
+--+--+
| +-- Pustulosa_succissa
|
| +-- Pustulosa_infucata
+--+
+-- Pustulosa_kleiniana
Wu et al., 2022 [Publ 4238], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
73 |
1 |
1 |
-- |
+-- Uniomerus_tetralasmus **
--+
+-- Quadrula
Mejia-Trujillo & Smith, 2022 [Publ 4274], fig. 2-F
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
14 |
1 |
1 |
-- |
+-- Quadrula
--+
+-- Uniomerus_tetralasmus **
Johnson et al., 2018 [Publ 3844], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
15 |
13 |
2 |
-- |
+-------------------------- Megalonaias
|
--+ +----------------------- Uniomerus_tetralasmus **
| |
+--+ +-------------------- Theliderma
| |
+--+ +----------------- Quadrula
| |
| | +----- Pustulosa_kieneriana
+--+ +--------+
| | | +-- Pustulosa_infucata
| | +--+
| | +-- Pustulosa_kleiniana
+--+
| +-- Pustulosa_pustulosa
| +--------+
| | +-- Pustulosa_succissa
+--+
| +-------- Cyclonaias
+--+
| +----- Pustulosa_necki
+--+
| +-- Pustulosa_nodulata
+--+
+-- Pustulosa_petrina
Dai et al., 2023 [Publ 4285], fig. 5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
52 |
1 |
1 |
-- |
+-- Quadrula
--+
+-- Uniomerus_tetralasmus **
Pfeiffer et al., 2018 [Publ 3835], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
3 |
3 |
2 |
-- |
+----- Quadrula
--+
| +-- Uniomerus_declivis **
+--+
+-- Megalonaias
Pfeiffer et al., 2019 [Publ 3937], fig. 6
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
56 |
1 |
1 |
-- |
+-- Quadrula
--+
+-- Uniomerus_tetralasmus **
Wu et al., 2023 [Publ 4339], fig. 4_S5-aa
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
95 |
1 |
1 |
-- |
+-- Quadrula
--+
+-- Uniomerus_tetralasmus **
Lydeard et al., 1996 [Publ 2656], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
27 |
3 |
2 |
-- |
-- Uniomerus_carolinianus
Rosenberg et al., 1997 [Publ 2720], fig. 2A
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
21 |
3 |
2 |
-- |
-- Uniomerus_tetralasmus
Huang et al., 2019 [Publ 3843], fig. S4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
197 |
3 |
2 |
-- |
-- Uniomerus_carolinianus
Lopes-Lima et al., 2019 [Publ 3901], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
28 |
18 |
2 |
-- |
+-------------------------- Uniomerus_tetralasmus **
|
--+ +----------------------- Theliderma
| |
+--+ +-------------------- Quadrula
| |
+--+ +----------------- Pustulosa_kieneriana
| |
| | +-- Pustulosa_infucata
+--+ +-----------+
| | +-- Pustulosa_kleiniana
| |
+--+ +-- Pustulosa_pustulosa
| +--------+
| | +-- Pustulosa_succissa
+--+
| +-------- Cyclonaias
+--+
| +----- Pustulosa_necki
+--+
| +-- Pustulosa_petrina
+--+
+-- Pustulosa_nodulata
Huang et al., 2002 [Publ 2657], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
33 |
1 |
1 |
-- |
-- Uniomerus_carolinianus
Wu et al., 2024 [Publ 4350], fig. 8b_S2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
91 |
1 |
1 |
-- |
+-- Quadrula
--+
+-- Uniomerus_tetralasmus **
Rosenberg et al., 1994 [Publ 2719], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
18 |
3 |
2 |
-- |
-- Uniomerus_tetralasmus
Huang et al., 2002 [Publ 2657], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
33 |
1 |
1 |
-- |
-- Uniomerus_carolinianus
Huang et al., 2019 [Publ 3843], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
197 |
3 |
2 |
-- |
+----- Quadrula
--+
| +-- Megalonaias
+--+
+-- Uniomerus_carolinianus **
Huang et al., 2019 [Publ 3843], fig. S3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
195 |
3 |
2 |
-- |
+----- Quadrula
--+
| +-- Megalonaias
+--+
+-- Uniomerus_carolinianus **
Huang et al., 2019 [Publ 3843], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
56 |
1 |
1 |
-- |
+-- Quadrula
--+
+-- Uniomerus_tetralasmus **
Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
18 |
1 |
1 |
-- |
+-- Uniomerus_tetralasmus **
--+
+-- Quadrula
Campbell et al., 2008 [Publ 2675], fig. 5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
24 |
0 |
0 |
-- |
-- Uniomerus_declivis
Kandl et al., 2001 [Publ 3384], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
5 |
0 |
0 |
-- |
-- Uniomerus_carolinianus
Perkins et al., 2017 [Publ 3777], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
23 |
0 |
0 |
-- |
-- Uniomerus_declivis
Pfeiffer et al., 2016 [Publ 3660], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
37 |
5 |
1 |
-- |
+-------------- Uniomerus_declivis **
--+
| +----------- Quadrula_quadrula
+--+
| +-------- Quadrula_verrucosa
+--+
| +----- Pustulosa_pustulosa
+--+
| +-- Pustulosa_petrina
+--+
+-- Cyclonaias
Campbell et al., 2008 [Publ 2675], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
24 |
0 |
0 |
-- |
-- Uniomerus_declivis
Wu et al., 2024 [Publ 4350], fig. 8a_S1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
91 |
1 |
1 |
-- |
+-- Quadrula
--+
+-- Uniomerus_tetralasmus **
Lopes-Lima et al., 2019 [Publ 3901], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
25 |
15 |
2 |
-- |
+----------------------------- Megalonaias
|
--+ +-------------------------- Uniomerus_tetralasmus **
| |
+--+ +----------------------- Theliderma
| |
+--+ +-------------------- Quadrula_quadrula
| |
| | +-- Quadrula_verrucosa
+--+ +--------------+
| | +-- Quadrula_nobilis
| |
| | +----- Pustulosa_kieneriana
+--+ +--------+
| | | +-- Pustulosa_infucata
| | +--+
| | +-- Pustulosa_kleiniana
+--+
| +-- Pustulosa_pustulosa
| +--------+
| | +-- Pustulosa_succissa
+--+
| +-------- Cyclonaias
+--+
| +----- Pustulosa_necki
+--+
| +-- Pustulosa_petrina
+--+
+-- Pustulosa_nodulata
Wu et al., 2023 [Publ 4339], fig. 4_S5-nt
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
95 |
1 |
1 |
-- |
+-- Quadrula
--+
+-- Uniomerus_tetralasmus **
Campbell & Lydeard, 2012 [Publ 3308], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
42 |
6 |
1 |
-- |
+-------------- Uniomerus_declivis **
|
--+ +-- Pustulosa_infucata
| +--------+
| | +-- Pustulosa_kleiniana
+--+
| +-------- Quadrula
+--+
| +----- Cyclonaias
+--+
| +-- Pustulosa_succissa
+--+
+-- Pustulosa_pustulosa
Pfeiffer et al., 2019 [Publ 3937], fig. 4-5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
56 |
1 |
1 |
-- |
+-- Quadrula
--+
+-- Uniomerus_tetralasmus **
Wu et al., 2021 [Publ 4366], fig. 3-BI
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
73 |
1 |
1 |
-- |
+-- Uniomerus_tetralasmus **
--+
+-- Quadrula
Smith, 2021 [Publ 4331], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
19 |
1 |
1 |
-- |
+-- Uniomerus_tetralasmus **
--+
+-- Megalonaias
Pfeiffer et al., 2019 [Publ 3866], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
34 |
4 |
2 |
-- |
+----------- Megalonaias
--+
| +-------- Uniomerus_declivis **
+--+
| +----- Quadrula_verrucosa
+--+
| +-- Cyclonaias
+--+
+-- Quadrula_quadrula
Lydeard et al., 1996 [Publ 2656], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
26 |
3 |
2 |
-- |
-- Uniomerus_carolinianus
Krebs et al., 2003 [Publ 2691], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
32 |
3 |
2 |
-- |
-- Uniomerus_carolinianus
Dai et al., 2024 [Publ 4358], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
51 |
1 |
1 |
-- |
+-- Quadrula
--+
+-- Uniomerus_tetralasmus **
Wu et al., 2021 [Publ 4366], fig. 3-ML
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
73 |
1 |
1 |
-- |
+-- Uniomerus_tetralasmus **
--+
+-- Quadrula
Burlakova et al., 2019 [Publ 3899], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
32 |
1 |
1 |
-- |
+-- Quadrula
--+
+-- Uniomerus_declivis **
Fukata & Iigo, 2020 [Publ 4106], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
53 |
1 |
1 |
-- |
+-- Quadrula
--+
+-- Uniomerus_tetralasmus **
Campbell et al., 2005 [Publ 2660], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
101 |
10 |
2 |
-- |
+-- Megalonaias
+-----------+
| +-- Uniomerus_declivis **
--+
| +----------- Theliderma
| |
+--+ +-------- Quadrula
| |
+--+ +----- Pustulosa_kieneriana
| |
| | +-- Cyclonaias
+--+--+
| +-- Pustulosa_succissa
|
| +-- Pustulosa_infucata
+--+
+-- Pustulosa_kleiniana
Rosenberg et al., 1997 [Publ 2720], fig. 2B
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
21 |
3 |
2 |
-- |
-- Uniomerus_tetralasmus
|