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Page last updated
Wed 04 Dec 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Ambleminae | Quadrulini

genus Uniomerus Conrad, 1853: cladomics
     5 valid species

Cladomics Summary

number of species = 5

analyzed in 47 cladograms in 32 publications
maximum number of terminals in an analysis = 4, mean = 1.17

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 5
maximum number of included terminals = 4, mean = 2.6
proportion of cladograms that support basic monophyly = 1.0


Neemuchwala et al., 2023 [Publ 4292], fig. 2

iT tT bi riS oT roT roS iD
4 4 2 4 23 18 2 0.0

*** Top Monophyly Cladogram *** *** Top Sister-Group Cladogram ***

  +-------------- Megalonaias           
  |                                     
  |     +-------- Uniomerus_carolinianus **
--+  +--+                               
  |  |  |  +----- Uniomerus_columbensis ** 
  |  |  +--+                            
  |  |     |  +-- Uniomerus_tetralasmus ** 
  +--+     +--+                         
     |        +-- Uniomerus_declivis **    
     |                                  
     |  +-------- Theliderma            
     +--+                               
        |  +----- Cyclonaias            
        +--+                            
           |  +-- Quadrula              
           +--+                         
              +-- Pustulosa             

Lopes-Lima et al., 2019 [Publ 3901], fig. 2

iT tT bi riS oT roT roS iD
3 3 0 3 25 15 2 2.0
                       +----- Megalonaias           
                       |                            
  +--------------------+  +-- Uniomerus_carolinianus **
  |                    |  |                         
  |                    +--+-- Uniomerus_declivis **    
  |                       |                         
--+                       +-- Uniomerus_tetralasmus ** 
  |                                                 
  |  +----------------------- Quadrula_quadrula     
  |  |                                              
  |  |                    +-- Quadrula_verrucosa    
  +--+  +-----------------+                         
     |  |                 +-- Quadrula_nobilis      
     |  |                                           
     +--+  +----------------- Theliderma            
        |  |                                        
        |  |           +----- Pustulosa_kieneriana  
        +--+  +--------+                            
           |  |        |  +-- Pustulosa_infucata    
           |  |        +--+                         
           +--+           +-- Pustulosa_kleiniana   
              |                                     
              |  +----------- Cyclonaias            
              |  |                                  
              +--+        +-- Pustulosa_pustulosa   
                 |  +-----+                         
                 |  |     +-- Pustulosa_succissa    
                 +--+                               
                    |  +----- Pustulosa_nodulata    
                    +--+                            
                       |  +-- Pustulosa_necki       
                       +--+                         
                          +-- Pustulosa_petrina     

Inoue et al., 2018 [Publ 3858], fig. 1_3a-c

iT tT bi riS oT roT roS iD
2 2 0 2 102 5 1 --
     +-- Uniomerus_declivis **    
  +--+                         
  |  +-- Uniomerus_tetralasmus ** 
  |                            
  |  +-- Cyclonaias            
--+  |                         
  |  +-- Pustulosa_pustulosa__1
  |  |                         
  |  +-- Theliderma            
  +--+                         
     +-- Pustulosa_pustulosa__2
     |                         
     +-- Quadrula_quadrula     
     |                         
     +-- Quadrula_verrucosa    

Campbell & Lydeard, 2012 [Publ 3309], fig. 1-2

iT tT bi riS oT roT roS iD
2 2 0 2 71 9 1 --
                    +-- Uniomerus_declivis **   
  +-----------------+                        
  |                 +-- Uniomerus_tetralasmus **
--+                                          
  |  +----------------- Theliderma           
  |  |                                       
  +--+  +-------------- Quadrula_verrucosa   
     |  |                                    
     |  |           +-- Quadrula_quadrula    
     +--+  +--------+                        
        |  |        +-- Quadrula_nobilis     
        |  |                                 
        +--+        +-- Pustulosa_infucata   
           |  +-----+                        
           |  |     +-- Pustulosa_kleiniana  
           +--+                              
              |  +----- Cyclonaias           
              +--+                           
                 |  +-- Pustulosa_pustulosa  
                 +--+                        
                    +-- Pustulosa_succissa   

Campbell & Lydeard, 2012 [Publ 3309], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 71 9 1 --
                 +-- Uniomerus_declivis **   
  +--------------+                        
  |              +-- Uniomerus_tetralasmus **
  |                                       
--+              +-- Quadrula_verrucosa   
  |  +-----------+                        
  |  |           +-- Quadrula_nobilis     
  |  |                                    
  +--+  +----------- Quadrula_quadrula    
     |  |                                 
     |  +----------- Theliderma           
     +--+                                 
        |        +-- Pustulosa_infucata   
        |  +-----+                        
        |  |     +-- Pustulosa_kleiniana  
        +--+                              
           |  +----- Cyclonaias           
           +--+                           
              |  +-- Pustulosa_pustulosa  
              +--+                        
                 +-- Pustulosa_succissa   

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iT tT bi riS oT roT roS iD
1 1 0 1 91 1 1 --
  +-- Quadrula             
--+                        
  +-- Uniomerus_tetralasmus **

Wu et al., 2021 [Publ 4366], fig. 3-ML

iT tT bi riS oT roT roS iD
1 1 0 1 73 1 1 --
  +-- Uniomerus_tetralasmus **
--+                        
  +-- Quadrula             

Dai et al., 2024 [Publ 4358], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 51 1 1 --
  +-- Quadrula             
--+                        
  +-- Uniomerus_tetralasmus **

Huang et al., 2019 [Publ 3843], fig. S4

iT tT bi riS oT roT roS iD
1 1 0 1 197 3 2 --
-- Uniomerus_carolinianus

Perkins et al., 2017 [Publ 3777], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 23 0 0 --
-- Uniomerus_declivis

Pfeiffer et al., 2016 [Publ 3660], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 37 5 1 --
  +-------------- Uniomerus_declivis ** 
--+                                  
  |  +----------- Quadrula_quadrula  
  +--+                               
     |  +-------- Quadrula_verrucosa 
     +--+                            
        |  +----- Pustulosa_pustulosa
        +--+                         
           |  +-- Pustulosa_petrina  
           +--+                      
              +-- Cyclonaias         

Huang et al., 2002 [Publ 2657], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 33 1 1 --
-- Uniomerus_carolinianus

Mejia-Trujillo & Smith, 2022 [Publ 4274], fig. 2-F

iT tT bi riS oT roT roS iD
1 1 0 1 14 1 1 --
  +-- Quadrula             
--+                        
  +-- Uniomerus_tetralasmus **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iT tT bi riS oT roT roS iD
1 1 0 1 56 1 1 --
  +-- Quadrula             
--+                        
  +-- Uniomerus_tetralasmus **

Huang et al., 2019 [Publ 3843], fig. S3

iT tT bi riS oT roT roS iD
1 1 0 1 195 3 2 --
  +----- Quadrula              
--+                            
  |  +-- Megalonaias           
  +--+                         
     +-- Uniomerus_carolinianus **

Wu et al., 2021 [Publ 4366], fig. 3-BI

iT tT bi riS oT roT roS iD
1 1 0 1 73 1 1 --
  +-- Uniomerus_tetralasmus **
--+                        
  +-- Quadrula             

Campbell & Lydeard, 2012 [Publ 3308], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 42 6 1 --
  +-------------- Uniomerus_declivis ** 
  |                                  
--+           +-- Pustulosa_infucata 
  |  +--------+                      
  |  |        +-- Pustulosa_kleiniana
  +--+                               
     |  +-------- Quadrula           
     +--+                            
        |  +----- Cyclonaias         
        +--+                         
           |  +-- Pustulosa_succissa 
           +--+                      
              +-- Pustulosa_pustulosa

Lydeard et al., 1996 [Publ 2656], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 27 3 2 --
-- Uniomerus_carolinianus

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iT tT bi riS oT roT roS iD
1 1 0 1 95 1 1 --
  +-- Quadrula             
--+                        
  +-- Uniomerus_tetralasmus **

Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 18 1 1 --
  +-- Uniomerus_tetralasmus **
--+                        
  +-- Quadrula             

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iT tT bi riS oT roT roS iD
1 1 0 1 91 1 1 --
  +-- Quadrula             
--+                        
  +-- Uniomerus_tetralasmus **

Rosenberg et al., 1997 [Publ 2720], fig. 2A

iT tT bi riS oT roT roS iD
1 1 0 1 21 3 2 --
-- Uniomerus_tetralasmus

Lydeard et al., 1996 [Publ 2656], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 26 3 2 --
-- Uniomerus_carolinianus

Krebs et al., 2003 [Publ 2691], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 32 3 2 --
-- Uniomerus_carolinianus

Dai et al., 2023 [Publ 4285], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 52 1 1 --
  +-- Quadrula             
--+                        
  +-- Uniomerus_tetralasmus **

Campbell et al., 2005 [Publ 2660], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 101 10 2 --
  +-------------------- Megalonaias         
  |                                         
--+  +----------------- Uniomerus_declivis **  
  |  |                                      
  +--+  +-------------- Theliderma          
     |  |                                   
     +--+  +----------- Quadrula_quadrula   
        |  |                                
        |  |        +-- Quadrula_nobilis    
        +--+  +-----+                       
           |  |     +-- Quadrula_verrucosa  
           |  |                             
           +--+  +----- Pustulosa_kieneriana
              |  |                          
              |  |  +-- Cyclonaias          
              +--+--+                       
                 |  +-- Pustulosa_succissa  
                 |                          
                 |  +-- Pustulosa_infucata  
                 +--+                       
                    +-- Pustulosa_kleiniana 

Johnson et al., 2018 [Publ 3844], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 15 13 2 --
  +-------------------------- Megalonaias          
  |                                                
--+  +----------------------- Uniomerus_tetralasmus **
  |  |                                             
  +--+  +-------------------- Theliderma           
     |  |                                          
     +--+  +----------------- Quadrula             
        |  |                                       
        |  |           +----- Pustulosa_kieneriana 
        +--+  +--------+                           
           |  |        |  +-- Pustulosa_infucata   
           |  |        +--+                        
           |  |           +-- Pustulosa_kleiniana  
           +--+                                    
              |           +-- Pustulosa_pustulosa  
              |  +--------+                        
              |  |        +-- Pustulosa_succissa   
              +--+                                 
                 |  +-------- Cyclonaias           
                 +--+                              
                    |  +----- Pustulosa_necki      
                    +--+                           
                       |  +-- Pustulosa_nodulata   
                       +--+                        
                          +-- Pustulosa_petrina    

Kandl et al., 2001 [Publ 3384], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 5 0 0 --
-- Uniomerus_carolinianus

Pfeiffer et al., 2018 [Publ 3835], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 3 3 2 --
  +----- Quadrula          
--+                        
  |  +-- Uniomerus_declivis **
  +--+                     
     +-- Megalonaias       

Campbell et al., 2008 [Publ 2675], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 24 0 0 --
-- Uniomerus_declivis

Wu et al., 2022 [Publ 4238], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 73 1 1 --
  +-- Uniomerus_tetralasmus **
--+                        
  +-- Quadrula             

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iT tT bi riS oT roT roS iD
1 1 0 1 95 1 1 --
  +-- Quadrula             
--+                        
  +-- Uniomerus_tetralasmus **

Lopes-Lima et al., 2019 [Publ 3901], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 25 15 2 --
  +----------------------------- Megalonaias          
  |                                                   
--+  +-------------------------- Uniomerus_tetralasmus **
  |  |                                                
  +--+  +----------------------- Theliderma           
     |  |                                             
     +--+  +-------------------- Quadrula_quadrula    
        |  |                                          
        |  |                 +-- Quadrula_verrucosa   
        +--+  +--------------+                        
           |  |              +-- Quadrula_nobilis     
           |  |                                       
           |  |           +----- Pustulosa_kieneriana 
           +--+  +--------+                           
              |  |        |  +-- Pustulosa_infucata   
              |  |        +--+                        
              |  |           +-- Pustulosa_kleiniana  
              +--+                                    
                 |           +-- Pustulosa_pustulosa  
                 |  +--------+                        
                 |  |        +-- Pustulosa_succissa   
                 +--+                                 
                    |  +-------- Cyclonaias           
                    +--+                              
                       |  +----- Pustulosa_necki      
                       +--+                           
                          |  +-- Pustulosa_petrina    
                          +--+                        
                             +-- Pustulosa_nodulata   

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 56 1 1 --
  +-- Quadrula             
--+                        
  +-- Uniomerus_tetralasmus **

Smith, 2021 [Publ 4331], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 19 1 1 --
  +-- Uniomerus_tetralasmus **
--+                        
  +-- Megalonaias          

Rosenberg et al., 1997 [Publ 2720], fig. 2B

iT tT bi riS oT roT roS iD
1 1 0 1 21 3 2 --
-- Uniomerus_tetralasmus

Lopes-Lima et al., 2019 [Publ 3901], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 28 18 2 --
  +-------------------------- Uniomerus_tetralasmus **
  |                                                
--+  +----------------------- Theliderma           
  |  |                                             
  +--+  +-------------------- Quadrula             
     |  |                                          
     +--+  +----------------- Pustulosa_kieneriana 
        |  |                                       
        |  |              +-- Pustulosa_infucata   
        +--+  +-----------+                        
           |  |           +-- Pustulosa_kleiniana  
           |  |                                    
           +--+           +-- Pustulosa_pustulosa  
              |  +--------+                        
              |  |        +-- Pustulosa_succissa   
              +--+                                 
                 |  +-------- Cyclonaias           
                 +--+                              
                    |  +----- Pustulosa_necki      
                    +--+                           
                       |  +-- Pustulosa_petrina    
                       +--+                        
                          +-- Pustulosa_nodulata   

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 53 1 1 --
  +-- Quadrula             
--+                        
  +-- Uniomerus_tetralasmus **

Huang et al., 2002 [Publ 2657], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 33 1 1 --
-- Uniomerus_carolinianus

Rosenberg et al., 1994 [Publ 2719], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 18 3 2 --
-- Uniomerus_tetralasmus

Campbell et al., 2005 [Publ 2660], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 101 10 2 --
              +-- Megalonaias         
  +-----------+                       
  |           +-- Uniomerus_declivis **  
--+                                   
  |  +----------- Theliderma          
  |  |                                
  +--+  +-------- Quadrula            
     |  |                             
     +--+  +----- Pustulosa_kieneriana
        |  |                          
        |  |  +-- Cyclonaias          
        +--+--+                       
           |  +-- Pustulosa_succissa  
           |                          
           |  +-- Pustulosa_infucata  
           +--+                       
              +-- Pustulosa_kleiniana 

Burlakova et al., 2019 [Publ 3899], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 32 1 1 --
  +-- Quadrula          
--+                     
  +-- Uniomerus_declivis **

Campbell et al., 2008 [Publ 2675], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 24 0 0 --
-- Uniomerus_declivis

Huang et al., 2019 [Publ 3843], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 197 3 2 --
  +----- Quadrula              
--+                            
  |  +-- Megalonaias           
  +--+                         
     +-- Uniomerus_carolinianus **

Pfeiffer et al., 2019 [Publ 3866], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 34 4 2 --
  +----------- Megalonaias       
--+                              
  |  +-------- Uniomerus_declivis **
  +--+                           
     |  +----- Quadrula_verrucosa
     +--+                        
        |  +-- Cyclonaias        
        +--+                     
           +-- Quadrula_quadrula 

Wu et al., 2024 [Publ 4388], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 53 1 1 --
  +-- Quadrula             
--+                        
  +-- Uniomerus_tetralasmus **

Huang et al., 2019 [Publ 3843], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 56 1 1 --
  +-- Quadrula             
--+                        
  +-- Uniomerus_tetralasmus **

Page updated: 14:56:46 Wed 04 Dec 2024

 
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