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Page last updated
Sat 03 May 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Unioninae | Unionini

genus Schistodesmus Simpson, 1900: cladomics
     3 valid species

Cladomics Summary

number of species = 3

analyzed in 81 cladograms in 49 publications
maximum number of terminals in an analysis = 2, mean = 1.35

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 28
maximum number of included terminals = 2, mean = 2.0
proportion of cladograms that support basic monophyly = 0.96

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iT tT bi riS oT roT roS iD
2 7 0 3 94 20 6 0.0

*** Top Monophyly Cladogram *** 

  +-------------------- Aculamprotula               
  |                                                 
--+  +----------------- Unio                        
  |  |                                              
  +--+  +-------------- Pseudocuneopsis             
     |  |                                           
     +--+  +----------- Nodularia                   
        |  |                                        
        +--+        +-- Schistodesmus_lampreyanus__1 **
           |  +-----+                               
           |  |     +-- Cuneopsis                   
           +--+                                     
              |  +----- Tchangsinaia                
              +--+                                  
                 |  +-- Schistodesmus_lampreyanus__2 **
                 +--+                               
                    +-- Schistodesmus_spinosus **

Chen et al., 2023 [Publ 4302], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 38 15 9 --

*** Top Sister-Group Cladogram ***

  +----------------- Aculamprotula            
  |                                           
  |              +-- Middendorffinaia         
--+  +-----------+                            
  |  |           +-- Nodularia                
  |  |                                        
  +--+  +----------- Cuneopsis                
     |  |                                     
     |  |     +----- Diaurora                 
     |  +-----+                               
     +--+     |  +-- Schistodesmus_lampreyanus **
        |     +--+                            
        |        +-- Schistodesmus_spinosus **   
        |                                     
        |  +-------- Pseudocuneopsis          
        +--+                                  
           |  +----- Inversiunio              
           +--+                               
              |  +-- Tchangsinaia             
              +--+                            
                 +-- Unio

Wu et al., 2018 [Publ 3836], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 51 12 5 -- 
  +-------------- Aculamprotula            
  |                                        
--+  +----------- Unio                     
  |  |                                     
  +--+        +-- Tchangsinaia             
     |  +-----+                            
     |  |     +-- Cuneopsis                
     +--+                                  
        |  +----- Nodularia                
        +--+                               
           |  +-- Schistodesmus_lampreyanus **
           +--+                            
              +-- Schistodesmus_spinosus **

Liu et al., 2023 [Publ 4310], fig. 5

iT tT bi riS oT roT roS iD
2 2 0 2 22 12 7 -- 
  +----------- Unio                     
  |                                     
  +----------- Aculamprotula            
  |                                     
--+  +-------- Cuneopsis                
  |  |                                  
  |  |  +----- Nodularia                
  |  +--+                               
  |  |  |  +-- Schistodesmus_lampreyanus **
  +--+  +--+                            
     |     +-- Schistodesmus_spinosus **   
     |                                  
     |     +-- Tchangsinaia             
     |     |                            
     +-----+-- Inversiunio              
           |                            
           +-- Pseudocuneopsis

Hou et al., 2025 [Publ 4401], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 2 26 24 5 -- 
  +-------------- Middendorffinaia         
  |                                        
--+  +----------- Pseudocuneopsis          
  |  |                                     
  +--+  +-------- Nodularia                
     |  |                                  
     +--+     +-- Schistodesmus_lampreyanus **
        |  +--+                            
        |  |  +-- Schistodesmus_spinosus **   
        +--+                               
           |  +-- Tchangsinaia             
           +--+                            
              +-- Cuneopsis

Wu et al., 2024 [Publ 4379], fig. 5

iT tT bi riS oT roT roS iD
2 2 0 2 52 27 6 -- 
  +----------------- Aculamprotula            
  |                                           
--+  +-------------- Unio                     
  |  |                                        
  +--+  +----------- Pseudocuneopsis          
     |  |                                     
     +--+  +-------- Nodularia                
        |  |                                  
        +--+     +-- Schistodesmus_spinosus **   
           |  +--+                            
           |  |  +-- Schistodesmus_lampreyanus **
           +--+                               
              |  +-- Tchangsinaia             
              +--+                            
                 +-- Cuneopsis

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iT tT bi riS oT roT roS iD
2 2 0 2 90 20 6 -- 
  +-------------- Aculamprotula            
  |                                        
--+  +----------- Unio                     
  |  |                                     
  +--+  +-------- Nodularia                
     |  |                                  
     |  +-------- Pseudocuneopsis          
     +--+                                  
        |     +-- Schistodesmus_lampreyanus **
        |  +--+                            
        |  |  +-- Schistodesmus_spinosus **   
        +--+                               
           |  +-- Tchangsinaia             
           +--+                            
              +-- Cuneopsis

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iT tT bi riS oT roT roS iD
2 2 0 2 90 20 6 -- 
  +-------------- Aculamprotula            
  |                                        
--+  +----------- Unio                     
  |  |                                     
  +--+  +-------- Nodularia                
     |  |                                  
     |  +-------- Pseudocuneopsis          
     +--+                                  
        |     +-- Schistodesmus_lampreyanus **
        |  +--+                            
        |  |  +-- Schistodesmus_spinosus **   
        +--+                               
           |  +-- Tchangsinaia             
           +--+                            
              +-- Cuneopsis

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iT tT bi riS oT roT roS iD
2 2 0 2 119 25 7 -- 
  +----------------- Aculamprotula            
  |                                           
--+  +-------------- Middendorffinaia         
  |  |                                        
  +--+  +----------- Unio                     
     |  |                                     
     +--+  +-------- Nodularia                
        |  |                                  
        +--+  +----- Cuneopsis                
           |  |                               
           |  |  +-- Schistodesmus_lampreyanus **
           +--+--+                            
              |  +-- Schistodesmus_spinosus **   
              |                               
              |  +-- Tchangsinaia             
              +--+                            
                 +-- Inversiunio

Chen et al., 2023 [Publ 4302], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 38 15 9 -- 
  +----------------- Aculamprotula            
  |                                           
  |              +-- Middendorffinaia         
--+  +-----------+                            
  |  |           +-- Nodularia                
  |  |                                        
  +--+  +----------- Cuneopsis                
     |  |                                     
     |  |     +----- Diaurora                 
     |  +-----+                               
     +--+     |  +-- Schistodesmus_lampreyanus **
        |     +--+                            
        |        +-- Schistodesmus_spinosus **   
        |                                     
        |  +-------- Pseudocuneopsis          
        +--+                                  
           |  +----- Inversiunio              
           +--+                               
              |  +-- Tchangsinaia             
              +--+                            
                 +-- Unio

Wu et al., 2022 [Publ 4238], fig. 2e

iT tT bi riS oT roT roS iD
2 2 0 2 15 10 6 -- 
  +-------------------- Aculamprotula            
--+                                              
  |  +----------------- Unio                     
  +--+                                           
     |  +-------------- Pseudocuneopsis          
     +--+                                        
        |  +----------- Nodularia                
        +--+                                     
           |  +-------- Tchangsinaia             
           +--+                                  
              |  +----- Cuneopsis                
              +--+                               
                 |  +-- Schistodesmus_lampreyanus **
                 +--+                            
                    +-- Schistodesmus_spinosus **

Liu et al., 2023 [Publ 4310], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 2 22 12 7 -- 
  +----------------- Unio                     
  |                                           
--+  +-------------- Aculamprotula            
  |  |                                        
  +--+  +----------- Cuneopsis                
     |  |                                     
     |  |     +----- Nodularia                
     +--+  +--+                               
        |  |  |  +-- Schistodesmus_lampreyanus **
        |  |  +--+                            
        +--+     +-- Schistodesmus_spinosus **   
           |                                  
           |  +----- Tchangsinaia             
           +--+                               
              |  +-- Inversiunio              
              +--+                            
                 +-- Pseudocuneopsis

Hou et al., 2024 [Publ 4400], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 2 75 47 7 -- 
  +-------------------- Aculamprotula            
  |                                              
--+  +----------------- Unio                     
  |  |                                           
  +--+  +-------------- Middendorffinaia         
     |  |                                        
     +--+  +----------- Pseudocuneopsis          
        |  |                                     
        +--+  +-------- Nodularia                
           |  |                                  
           +--+     +-- Schistodesmus_lampreyanus **
              |  +--+                            
              |  |  +-- Schistodesmus_spinosus **   
              +--+                               
                 |  +-- Tchangsinaia             
                 +--+                            
                    +-- Cuneopsis

Huang et al., 2019 [Publ 3843], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 197 17 5 -- 
  +-------------- Aculamprotula            
  |                                        
--+  +----------- Unio                     
  |  |                                     
  +--+        +-- Schistodesmus_lampreyanus **
     |  +-----+                            
     |  |     +-- Schistodesmus_spinosus **   
     +--+                                  
        |  +----- Nodularia                
        +--+                               
           |  +-- Tchangsinaia             
           +--+                            
              +-- Cuneopsis

Wu et al., 2021 [Publ 4366], fig. 6

iT tT bi riS oT roT roS iD
2 2 0 2 89 9 5 -- 
  +----------------- Aculamprotula            
--+                                           
  |  +-------------- Unio                     
  +--+                                        
     |  +----------- Nodularia                
     +--+                                     
        |  +-------- Cuneopsis                
        +--+                                  
           |  +----- Tchangsinaia             
           +--+                               
              |  +-- Schistodesmus_spinosus **   
              +--+                            
                 +-- Schistodesmus_lampreyanus **

Zhou et al., 2007 [Publ 2658], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 19 5 3 -- 
  +-- Schistodesmus_lampreyanus **
--+                            
  +-- Schistodesmus_spinosus **

Hou et al., 2025 [Publ 4432], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 2 26 24 5 -- 
  +-------------- Middendorffinaia         
  |                                        
--+  +----------- Pseudocuneopsis          
  |  |                                     
  +--+  +-------- Nodularia                
     |  |                                  
     +--+     +-- Schistodesmus_lampreyanus **
        |  +--+                            
        |  |  +-- Schistodesmus_spinosus **   
        +--+                               
           |  +-- Tchangsinaia             
           +--+                            
              +-- Cuneopsis

Wu et al., 2024 [Publ 4379], fig. 4_6

iT tT bi riS oT roT roS iD
2 2 0 2 20 12 2 -- 
  +----- Nodularia                
--+                               
  |  +-- Schistodesmus_lampreyanus **
  +--+                            
     +-- Schistodesmus_spinosus **

Huang et al., 2019 [Publ 3843], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 41 13 4 -- 
                 +-- Schistodesmus_lampreyanus **
  +--------------+                            
  |              +-- Schistodesmus_spinosus **   
--+                                           
  |  +-------------- Nodularia                
  +--+                                        
     |  +----------- Cuneopsis_rufescens      
     +--+                                     
        |  +-------- Tchangsinaia             
        +--+                                  
           |  +----- Cuneopsis_celtiformis    
           +--+                               
              |  +-- Cuneopsis_szechenyii     
              +--+                            
                 +-- Cuneopsis_heudei

Wu et al., 2022 [Publ 4238], fig. 2d

iT tT bi riS oT roT roS iD
2 2 0 2 37 21 7 -- 
  +-------------------- Aculamprotula            
  |                                              
--+  +----------------- Unio                     
  |  |                                           
  +--+  +-------------- Pseudocuneopsis          
     |  |                                        
     +--+  +----------- Nodularia                
        |  |                                     
        +--+        +-- Schistodesmus_lampreyanus **
           |  +-----+                            
           |  |     +-- Schistodesmus_spinosus **   
           +--+                                  
              |  +----- Cuneopsis                
              +--+                               
                 |  +-- Tchangsinaia             
                 +--+                            
                    +-- Inversiunio

Huang et al., 2019 [Publ 3843], fig. S4

iT tT bi riS oT roT roS iD
2 2 0 2 197 17 5 -- 
  +-- Schistodesmus_lampreyanus **
--+                            
  +-- Schistodesmus_spinosus **

Dai et al., 2024 [Publ 4358], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 2 50 15 6 -- 
  +----------------- Aculamprotula            
  |                                           
--+  +-------------- Unio                     
  |  |                                        
  +--+  +----------- Pseudocuneopsis          
     |  |                                     
     +--+  +-------- Nodularia                
        |  |                                  
        +--+     +-- Schistodesmus_lampreyanus **
           |  +--+                            
           |  |  +-- Schistodesmus_spinosus **   
           +--+                               
              |  +-- Tchangsinaia             
              +--+                            
                 +-- Cuneopsis

Wu et al., 2021 [Publ 4366], fig. 2_I

iT tT bi riS oT roT roS iD
2 2 0 2 89 9 5 -- 
  +----------------- Aculamprotula            
--+                                           
  |  +-------------- Unio                     
  +--+                                        
     |  +----------- Nodularia                
     +--+                                     
        |  +-------- Cuneopsis                
        +--+                                  
           |  +----- Tchangsinaia             
           +--+                               
              |  +-- Schistodesmus_lampreyanus **
              +--+                            
                 +-- Schistodesmus_spinosus **

Wu et al., 2023 [Publ 4296], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 2 21 11 7 -- 
  +----------------- Unio                     
  |                                           
--+  +-------------- Aculamprotula            
  |  |                                        
  +--+  +----------- Cuneopsis                
     |  |                                     
     |  |     +----- Nodularia                
     +--+  +--+                               
        |  |  |  +-- Schistodesmus_lampreyanus **
        |  |  +--+                            
        +--+     +-- Schistodesmus_spinosus **   
           |                                  
           |  +----- Tchangsinaia             
           +--+                               
              |  +-- Inversiunio              
              +--+                            
                 +-- Pseudocuneopsis

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iT tT bi riS oT roT roS iD
2 2 0 2 94 20 6 -- 
  +----------------- Aculamprotula            
  |                                           
--+  +-------------- Unio                     
  |  |                                        
  +--+           +-- Nodularia                
     |  +--------+                            
     |  |        +-- Pseudocuneopsis          
     +--+                                     
        |  +-------- Tchangsinaia             
        +--+                                  
           |  +----- Cuneopsis                
           +--+                               
              |  +-- Schistodesmus_lampreyanus **
              +--+                            
                 +-- Schistodesmus_spinosus **

Zhang et al., 2004 [Publ 2735], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 12 5 4 -- 
  +----------------- Tchangsinaia             
--+                                           
  |  +-------------- Cuneopsis_heudei         
  +--+                                        
     |  +----------- Nodularia                
     +--+                                     
        |  +-------- Pseudocuneopsis          
        +--+                                  
           |  +----- Cuneopsis_rufescens      
           +--+                               
              |  +-- Schistodesmus_lampreyanus **
              +--+                            
                 +-- Schistodesmus_spinosus **

Wu et al., 2022 [Publ 4238], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 72 18 6 -- 
  +----------------- Aculamprotula            
  |                                           
--+  +-------------- Unio                     
  |  |                                        
  +--+  +----------- Pseudocuneopsis          
     |  |                                     
     +--+  +-------- Nodularia                
        |  |                                  
        +--+     +-- Schistodesmus_lampreyanus **
           |  +--+                            
           |  |  +-- Schistodesmus_spinosus **   
           +--+                               
              |  +-- Tchangsinaia             
              +--+                            
                 +-- Cuneopsis

Huang et al., 2019 [Publ 3843], fig. S1

iT tT bi riS oT roT roS iD
2 2 0 2 44 13 4 -- 
  +-------------- Aculamprotula            
--+                                        
  |  +----------- Nodularia                
  +--+                                     
     |  +-------- Tchangsinaia             
     +--+                                  
        |  +----- Cuneopsis                
        +--+                               
           |  +-- Schistodesmus_spinosus **   
           +--+                            
              +-- Schistodesmus_lampreyanus **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iT tT bi riS oT roT roS iD
1 1 0 1 75 9 5 -- 
  +----------------- Unio                     
--+                                           
  |  +-------------- Aculamprotula            
  +--+                                        
     |  +----------- Nodularia                
     +--+                                     
        |  +-------- Schistodesmus_lampreyanus **
        +--+                                  
           |  +----- Cuneopsis_szechenyii     
           +--+                               
              |  +-- Tchangsinaia             
              +--+                            
                 +-- Cuneopsis_rufescens

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iT tT bi riS oT roT roS iD
1 1 0 1 75 9 5 -- 
  +----------------- Unio                     
--+                                           
  |  +-------------- Aculamprotula            
  +--+                                        
     |  +----------- Nodularia                
     +--+                                     
        |  +-------- Schistodesmus_lampreyanus **
        +--+                                  
           |  +----- Cuneopsis_szechenyii     
           +--+                               
              |  +-- Tchangsinaia             
              +--+                            
                 +-- Cuneopsis_rufescens

Sano et al., 2017 [Publ 3799], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 32 7 3 -- 
-- Schistodesmus_lampreyanus

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 53 11 6 -- 
  +-------------- Aculamprotula            
  |                                        
--+  +----------- Unio                     
  |  |                                     
  +--+        +-- Schistodesmus_lampreyanus **
     |  +-----+                            
     |  |     +-- Cuneopsis                
     +--+                                  
        |  +----- Pseudocuneopsis          
        +--+                               
           |  +-- Nodularia                
           +--+                            
              +-- Tchangsinaia

Xue et al., 2016 [Publ 3659], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 15 3 3 -- 
  +-------- Aculamprotula            
--+                                  
  |  +----- Schistodesmus_lampreyanus **
  +--+                               
     |  +-- Nodularia                
     +--+                            
        +-- Unio

Bolotov et al., 2017 [Publ 3761], fig. S2

iT tT bi riS oT roT roS iD
1 1 0 1 138 9 5 -- 
  +----------- Unio                     
  |                                     
--+  +-------- Aculamprotula            
  |  |                                  
  |  +-------- Schistodesmus_lampreyanus **
  +--+                                  
     +-------- Nodularia                
     |                                  
     |  +----- Cuneopsis_szechenyii     
     +--+                               
        |  +-- Tchangsinaia             
        +--+                            
           +-- Cuneopsis_rufescens

Guerra et al., 2019 [Publ 4168], fig. 2-M

iT tT bi riS oT roT roS iD
1 1 0 1 21 4 2 -- 
  +----- Aculamprotula            
--+                               
  |  +-- Schistodesmus_lampreyanus **
  +--+                            
     +-- Unio

Froufe et al., 2019 [Publ 3936], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 44 9 4 -- 
  +----------- Aculamprotula            
--+                                     
  |  +-------- Unio                     
  +--+                                  
     |  +----- Nodularia                
     +--+                               
        |  +-- Tchangsinaia             
        +--+                            
           +-- Schistodesmus_lampreyanus **

Cha et al., 2018 [Publ 3916], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 38 9 4 -- 
  +----------- Aculamprotula            
--+                                     
  |  +-------- Unio                     
  +--+                                  
     |  +----- Schistodesmus_lampreyanus **
     +--+                               
        |  +-- Nodularia                
        +--+                            
           +-- Tchangsinaia

Aunins et al., 2018 [Publ 3863], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 40 10 5 -- 
  +-------------- Aculamprotula            
--+                                        
  |  +----------- Unio                     
  +--+                                     
     |  +-------- Nodularia                
     +--+                                  
        |  +----- Tchangsinaia             
        +--+                               
           |  +-- Cuneopsis                
           +--+                            
              +-- Schistodesmus_lampreyanus **

Huang et al., 2019 [Publ 3843], fig. S3

iT tT bi riS oT roT roS iD
1 1 0 1 196 17 5 -- 
  +-------------- Aculamprotula            
--+                                        
  |  +----------- Unio                     
  +--+                                     
     |  +-------- Nodularia                
     +--+                                  
        |  +----- Schistodesmus_lampreyanus **
        +--+                               
           |  +-- Tchangsinaia             
           +--+                            
              +-- Cuneopsis

Dai et al., 2024 [Publ 4386], fig. 5-ML

iT tT bi riS oT roT roS iD
1 1 0 1 20 10 6 -- 
  +----------------- Aculamprotula            
--+                                           
  |  +-------------- Unio                     
  +--+                                        
     |  +----------- Middendorffinaia         
     +--+                                     
        |  +-------- Nodularia                
        +--+                                  
           |  +----- Cuneopsis                
           +--+                               
              |  +-- Inversiunio              
              +--+                            
                 +-- Schistodesmus_lampreyanus **

Dai et al., 2024 [Publ 4386], fig. 5-BI

iT tT bi riS oT roT roS iD
1 1 0 1 20 10 6 -- 
  +----------- Unio                     
  |                                     
--+  +-------- Middendorffinaia         
  |  |                                  
  +--+  +----- Cuneopsis                
     |  |                               
     +--+----- Nodularia                
        |                               
        |  +-- Inversiunio              
        +--+                            
           +-- Schistodesmus_lampreyanus **

Kim et al., 2021 [Publ 4197], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 16 10 4 -- 
  +-------- Unio                     
--+                                  
  |  +----- Nodularia                
  +--+                               
     |  +-- Schistodesmus_lampreyanus **
     +--+                            
        +-- Cuneopsis

Sano et al., 2020 [Publ 4138], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 68 12 5 -- 
  +-------------- Aculamprotula            
--+                                        
  |  +----------- Unio                     
  +--+                                     
     |  +-------- Inversiunio              
     +--+                                  
        |  +----- Nodularia                
        +--+                               
           |  +-- Tchangsinaia             
           +--+                            
              +-- Schistodesmus_lampreyanus **

Zhou et al., 2016 [Publ 3710], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 19 1 1 -- 
  +-- Schistodesmus_lampreyanus **
--+                            
  +-- Unio

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 35 5 4 -- 
  +----------- Aculamprotula            
--+                                     
  |  +-------- Unio                     
  +--+                                  
     |  +----- Schistodesmus_lampreyanus **
     +--+                               
        |  +-- Nodularia                
        +--+                            
           +-- Tchangsinaia

Chen et al., 2020 [Publ 4170], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 33 2 1 -- 
-- Schistodesmus_lampreyanus

Wu et al., 2022 [Publ 4238], fig. 2a_S1

iT tT bi riS oT roT roS iD
1 1 0 1 11 11 6 -- 
  +----------------- Aculamprotula  
--+                                 
  |  +-------------- Nodularia      
  +--+                              
     |  +----------- Pseudocuneopsis
     +--+                           
        |  +-------- Schistodesmus **  
        +--+                        
           |  +----- Cuneopsis      
           +--+                     
              |  +-- Tchangsinaia   
              +--+                  
                 +-- Inversiunio

Fukata & Iigo, 2019 [Publ 4108], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 15 11 6 -- 
  +----------- Unio                     
  |                                     
--+  +-------- Pseudocuneopsis          
  |  |                                  
  +--+     +-- Schistodesmus_lampreyanus **
     |  +--+                            
     |  |  +-- Cuneopsis                
     +--+                               
        |  +-- Tchangsinaia             
        +--+                            
           +-- Nodularia

Choi et al., 2020 [Publ 4169], fig. 2a

iT tT bi riS oT roT roS iD
1 1 0 1 11 11 4 -- 
  +----------- Nodularia                
--+                                     
  |  +-------- Schistodesmus_lampreyanus **
  +--+                                  
     |  +----- Cuneopsis                
     +--+                               
        |  +-- Unio                     
        +--+                            
           +-- Tchangsinaia

Guerra et al., 2019 [Publ 4168], fig. 1-M

iT tT bi riS oT roT roS iD
1 1 0 1 21 4 2 -- 
  +----- Schistodesmus_lampreyanus **
--+                               
  |  +-- Aculamprotula            
  +--+                            
     +-- Unio

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iT tT bi riS oT roT roS iD
1 1 0 1 79 10 5 -- 
  +-------------- Unio                     
--+                                        
  |  +----------- Aculamprotula            
  +--+                                     
     |  +-------- Nodularia                
     +--+                                  
        |  +----- Schistodesmus_lampreyanus **
        +--+                               
           |  +-- Tchangsinaia             
           +--+                            
              +-- Cuneopsis

Guerra et al., 2019 [Publ 4168], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 40 4 2 -- 
  +-- Schistodesmus_lampreyanus **
--+                            
  +-- Unio

Dai et al., 2023 [Publ 4285], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 52 6 6 -- 
  +-------------- Aculamprotula            
  |                                        
--+  +----------- Unio                     
  |  |                                     
  +--+        +-- Pseudocuneopsis          
     |  +-----+                            
     |  |     +-- Nodularia                
     +--+                                  
        |  +----- Schistodesmus_lampreyanus **
        +--+                               
           |  +-- Cuneopsis                
           +--+                            
              +-- Tchangsinaia

Sano et al., 2020 [Publ 4138], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 33 6 4 -- 
  +-- Schistodesmus_lampreyanus **
--+                            
  +-- Inversiunio

Sano et al., 2020 [Publ 4138], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 68 12 5 -- 
  +-------------- Aculamprotula            
--+                                        
  |  +----------- Unio                     
  +--+                                     
     |  +-------- Nodularia                
     +--+                                  
        |  +----- Inversiunio              
        +--+                               
           |  +-- Schistodesmus_lampreyanus **
           +--+                            
              +-- Tchangsinaia

Huang et al., 2019 [Publ 3843], fig. 3-M

iT tT bi riS oT roT roS iD
1 1 0 1 22 4 2 -- 
  +----- Schistodesmus_lampreyanus **
--+                               
  |  +-- Aculamprotula            
  +--+                            
     +-- Unio

Araujo et al., 2017 [Publ 3801], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 13 2 2 -- 
  +-- Schistodesmus_lampreyanus **
--+                            
  +-- Aculamprotula

Bolotov et al., 2017 [Publ 3761], fig. S1

iT tT bi riS oT roT roS iD
1 1 0 1 138 9 5 -- 
  +-------------- Unio                     
--+                                        
  |  +----------- Aculamprotula            
  +--+                                     
     |  +-------- Nodularia                
     +--+                                  
        |  +----- Cuneopsis                
        +--+                               
           |  +-- Tchangsinaia             
           +--+                            
              +-- Schistodesmus_lampreyanus **

Guerra et al., 2019 [Publ 4168], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 40 4 2 -- 
  +----- Aculamprotula            
--+                               
  |  +-- Schistodesmus_lampreyanus **
  +--+                            
     +-- Unio

Wu et al., 2021 [Publ 4366], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 68 14 6 -- 
  +----------------- Aculamprotula            
--+                                           
  |  +-------------- Unio                     
  +--+                                        
     |  +----------- Pseudocuneopsis          
     +--+                                     
        |  +-------- Nodularia                
        +--+                                  
           |  +----- Schistodesmus_lampreyanus **
           +--+                               
              |  +-- Tchangsinaia             
              +--+                            
                 +-- Cuneopsis

Bogan et al., 2023 [Publ 4295], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 30 0 0 -- 
-- Schistodesmus_lampreyanus

Liu et al., 2017 [Publ 3939], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 14 10 3 -- 
  +----- Unio                     
  |                               
--+  +-- Schistodesmus_lampreyanus **
  |  |                            
  +--+-- Cuneopsis                
     |                            
     +-- Nodularia

Wu et al., 2021 [Publ 4366], fig. 3-BI

iT tT bi riS oT roT roS iD
1 1 0 1 73 14 6 -- 
  +----------------- Aculamprotula            
--+                                           
  |  +-------------- Unio                     
  +--+                                        
     |  +----------- Pseudocuneopsis          
     +--+                                     
        |  +-------- Nodularia                
        +--+                                  
           |  +----- Schistodesmus_lampreyanus **
           +--+                               
              |  +-- Cuneopsis                
              +--+                            
                 +-- Tchangsinaia

Wu et al., 2020 [Publ 4121], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 49 13 6 -- 
  +----------------- Aculamprotula            
--+                                           
  |  +-------------- Unio                     
  +--+                                        
     |  +----------- Pseudocuneopsis          
     +--+                                     
        |  +-------- Nodularia                
        +--+                                  
           |  +----- Tchangsinaia             
           +--+                               
              |  +-- Cuneopsis                
              +--+                            
                 +-- Schistodesmus_lampreyanus **

Froufe et al., 2019 [Publ 3936], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 44 9 4 -- 
  +----------- Aculamprotula            
--+                                     
  |  +-------- Unio                     
  +--+                                  
     |  +----- Nodularia                
     +--+                               
        |  +-- Tchangsinaia             
        +--+                            
           +-- Schistodesmus_lampreyanus **

Sano et al., 2020 [Publ 4138], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 68 12 5 -- 
  +-------------- Aculamprotula            
--+                                        
  |  +----------- Unio                     
  +--+                                     
     |  +-------- Inversiunio              
     +--+                                  
        |  +----- Nodularia                
        +--+                               
           |  +-- Schistodesmus_lampreyanus **
           +--+                            
              +-- Tchangsinaia

Huang et al., 2002 [Publ 2657], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 33 2 2 -- 
-- Schistodesmus_lampreyanus

Dai et al., 2024 [Publ 4386], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 39 19 6 -- 
  +----------------- Aculamprotula            
--+                                           
  |  +-------------- Unio                     
  +--+                                        
     |  +----------- Pseudocuneopsis          
     +--+                                     
        |  +-------- Nodularia                
        +--+                                  
           |  +----- Schistodesmus_lampreyanus **
           +--+                               
              |  +-- Tchangsinaia             
              +--+                            
                 +-- Cuneopsis

Wu et al., 2019 [Publ 3864], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 46 12 6 -- 
  +----------------- Aculamprotula            
--+                                           
  |  +-------------- Unio                     
  +--+                                        
     |  +----------- Pseudocuneopsis          
     +--+                                     
        |  +-------- Nodularia                
        +--+                                  
           |  +----- Tchangsinaia             
           +--+                               
              |  +-- Cuneopsis                
              +--+                            
                 +-- Schistodesmus_lampreyanus **

Wu et al., 2024 [Publ 4388], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 53 8 6 -- 
  +----------------- Aculamprotula            
--+                                           
  |  +-------------- Unio                     
  +--+                                        
     |  +----------- Pseudocuneopsis          
     +--+                                     
        |  +-------- Nodularia                
        +--+                                  
           |  +----- Tchangsinaia             
           +--+                               
              |  +-- Cuneopsis                
              +--+                            
                 +-- Schistodesmus_lampreyanus **

Wu et al., 2021 [Publ 4366], fig. 3-ML

iT tT bi riS oT roT roS iD
1 1 0 1 73 14 6 -- 
  +----------------- Aculamprotula            
--+                                           
  |  +-------------- Unio                     
  +--+                                        
     |  +----------- Pseudocuneopsis          
     +--+                                     
        |  +-------- Nodularia                
        +--+                                  
           |  +----- Tchangsinaia             
           +--+                               
              |  +-- Schistodesmus_lampreyanus **
              +--+                            
                 +-- Cuneopsis

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iT tT bi riS oT roT roS iD
1 1 0 1 48 9 4 -- 
  +----------- Aculamprotula            
--+                                     
  |  +-------- Unio                     
  +--+                                  
     |  +----- Nodularia                
     +--+                               
        |  +-- Schistodesmus_lampreyanus **
        +--+                            
           +-- Tchangsinaia

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 32 6 4 -- 
  +----------- Aculamprotula            
--+                                     
  |  +-------- Unio                     
  +--+                                  
     |  +----- Schistodesmus_lampreyanus **
     +--+                               
        |  +-- Tchangsinaia             
        +--+                            
           +-- Nodularia

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 23 9 4 -- 
  +----------- Aculamprotula            
--+                                     
  |  +-------- Unio                     
  +--+                                  
     |  +----- Schistodesmus_lampreyanus **
     +--+                               
        |  +-- Tchangsinaia             
        +--+                            
           +-- Nodularia

Huang et al., 2002 [Publ 2657], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 33 2 2 -- 
-- Schistodesmus_lampreyanus

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iT tT bi riS oT roT roS iD
1 1 0 1 38 5 4 -- 
  +-------- Unio                     
--+                                  
  |  +----- Schistodesmus_lampreyanus **
  +--+                               
     |  +-- Nodularia                
     +--+                            
        +-- Tchangsinaia

Huang et al., 2019 [Publ 3843], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 56 10 4 -- 
  +----------- Aculamprotula            
--+                                     
  |  +-------- Unio                     
  +--+                                  
     |  +----- Nodularia                
     +--+                               
        |  +-- Tchangsinaia             
        +--+                            
           +-- Schistodesmus_lampreyanus **

Bolotov et al., 2017 [Publ 3764], fig. S1

iT tT bi riS oT roT roS iD
1 1 0 1 124 9 5 -- 
  +-- Unio_crassus             
  |                            
  +-- Unio_pictorum            
  |                            
  +-- Unio_tumidus             
  |                            
  +-- Aculamprotula            
  |                            
  +-- Cuneopsis_szechenyii     
--+                            
  +-- Tchangsinaia             
  |                            
  +-- Cuneopsis_rufescens      
  |                            
  +-- Nodularia_douglasiae     
  |                            
  +-- Nodularia_nuxpersicae    
  |                            
  +-- Schistodesmus_lampreyanus **

Fonseca et al., 2016 [Publ 3775], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 31 6 4 -- 
  +----------- Aculamprotula            
--+                                     
  |  +-------- Unio                     
  +--+                                  
     |  +----- Schistodesmus_lampreyanus **
     +--+                               
        |  +-- Nodularia                
        +--+                            
           +-- Tchangsinaia

Wu et al., 2020 [Publ 4147], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 28 9 6 -- 
  +----------------- Aculamprotula            
--+                                           
  |  +-------------- Unio                     
  +--+                                        
     |  +----------- Pseudocuneopsis          
     +--+                                     
        |  +-------- Nodularia                
        +--+                                  
           |  +----- Schistodesmus_lampreyanus **
           +--+                               
              |  +-- Cuneopsis                
              +--+                            
                 +-- Tchangsinaia

Dai et al., 2024 [Publ 4358], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 5 5 2 -- 
  +----- Pseudocuneopsis          
--+                               
  |  +-- Schistodesmus_lampreyanus **
  +--+                            
     +-- Nodularia

Page updated: 10:46:50 Sat 03 May 2025
 
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