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Page last updated
Thu 07 Sep 2023

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Gonideinae | Rectidentini

genus Hyriopsis Conrad, 1853: cladomics
     10 valid species

Cladomics Summary

number of species = 10

analyzed in 32 cladograms in 21 publications
maximum number of terminals in an analysis = 9, mean = 3.66

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 21
maximum number of included terminals = 9, mean = 5.05
proportion of cladograms that support basic monophyly = 0.24

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ tree

Konopleva et al., 2023 [Publ 4262], fig. 8

iT tT bi riS oT roT roS iD
9 10 8 9 75 7 2 0.0

*** Top Monophyly Cladogram *** 

                       +-- Ensidens_ingallsianus  
                 +-----+                          
                 |     +-- Ensidens_telus         
                 |                                
  +--------------+     +-- Ensidens_jaculus       
  |              |  +--+                          
  |              |  |  +-- Ensidens_spiculus[1]   
  |              +--+                             
  |                 |  +-- Ensidens_spiculus[2]   
--+                 +--+                          
  |                    +-- Ensidens_dugasti       
  |                                               
  |                    +-- Hyriopsis_desowitzi **    
  |  +-----------------+                          
  |  |                 +-- Hyriopsis_delaportei **   
  +--+                                            
     |  +----------------- Rectidens_sumatrensis  
     |  |                                         
     +--+  +-------------- Hyriopsis_panhai **       
        |  |                                      
        +--+  +----------- Hyriopsis_myersiana **    
           |  |                                   
           +--+        +-- Hyriopsis_khoratensis **  
              |  +-----+                          
              |  |     +-- Hyriopsis_bialata **      
              +--+                                
                 |  +----- Hyriopsis_sakhonensis **  
                 +--+                             
                    |  +-- Hyriopsis_phuphaniensis **
                    +--+                          
                       +-- Hyriopsis_kratiensis **

Konopleva et al., 2019 [Publ 4041], fig. 3

iT tT bi riS oT roT roS iD
5 13 0 5 30 8 2 15.0

*** Top Sister-Group Cladogram ***

  +-- Ensidens_ingallsianus
  |                        
  +-- Ensidens_telus       
  |                        
  +-- Ensidens_dugasti     
  |                        
  +-- Hyriopsis_khoratensis **
  |                        
  +-- Hyriopsis_bialata **    
  |                        
  +-- Hyriopsis_myersiana **  
  |                        
--+-- Hyriopsis_desowitzi **  
  |                        
  +-- Rectidens_sumatrensis
  |                        
  +-- Hyriopsis_panhai **     
  |                        
  +-- Ensidens_jaculus[1]  
  |                        
  +-- Ensidens_jaculus[2]  
  |                        
  +-- Ensidens_spiculus    
  |                        
  +-- Ensidens_sagittarius

Pfeiffer et al., 2021 [Publ 4126], fig. 27

iT tT bi riS oT roT roS iD
9 9 5 9 2 1 1 7.0 
  +-------------- Rectidens_sumatrensis  
  |                                      
  |           +-- Hyriopsis_delaportei **   
--+  +--------+                          
  |  |        +-- Hyriopsis_desowitzi **    
  |  |                                   
  +--+  +-------- Hyriopsis_khoratensis **  
     |  |                                
     |  |     +-- Hyriopsis_kratiensis **   
     |  +-----+                          
     +--+     +-- Hyriopsis_phuphaniensis **
        |                                
        |  +----- Hyriopsis_sakhonensis **  
        |  |                             
        +--+  +-- Hyriopsis_myersiana **    
           |  |                          
           +--+-- Hyriopsis_panhai **       
              |                          
              +-- Hyriopsis_bialata **

Jeratthitkul et al., 2021 [Publ 4212], fig. 4

iT tT bi riS oT roT roS iD
8 9 7 8 13 3 2 7.0 
                 +-- Ensidens_ingallsianus  
  +--------------+                          
  |              +-- Ensidens_jaculus       
  |                                         
--+              +-- Hyriopsis_desowitzi **    
  |  +-----------+                          
  |  |           +-- Hyriopsis_delaportei **   
  |  |                                      
  +--+  +----------- Rectidens_sumatrensis  
     |  |                                   
     |  |     +----- Hyriopsis_myersiana **    
     +--+  +--+                             
        |  |  |  +-- Hyriopsis_bialata **      
        |  |  +--+                          
        +--+     +-- Hyriopsis_panhai **       
           |                                
           |  +----- Hyriopsis_khoratensis **  
           +--+                             
              |  +-- Hyriopsis_phuphaniensis **
              +--+                          
                 +-- Hyriopsis_sakhonensis **

Pfeiffer et al., 2021 [Publ 4126], fig. 9

iT tT bi riS oT roT roS iD
6 6 4 6 23 7 2 6.0 
                    +-- Ensidens_ingallsianus
           +--------+                        
           |        +-- Ensidens_telus       
  +--------+                                 
  |        |  +-------- Ensidens_spiculus    
  |        +--+                              
  |           |  +----- Ensidens_dugasti     
  |           +--+                           
--+              |  +-- Ensidens_jaculus     
  |              +--+                        
  |                 +-- Ensidens_sagittarius 
  |                                          
  |  +----------------- Rectidens_sumatrensis
  +--+                                       
     |  +-------------- Hyriopsis_delaportei ** 
     +--+                                    
        |  +----------- Hyriopsis_myersiana **  
        +--+                                 
           |  +-------- Hyriopsis_bialata **    
           +--+                              
              |  +----- Hyriopsis_khoratensis **
              +--+                           
                 |  +-- Hyriopsis_kratiensis ** 
                 +--+                        
                    +-- Hyriopsis_sakhonensis **

Bolotov et al., 2017 [Publ 3761], fig. S1

iT tT bi riS oT roT roS iD
5 6 4 5 133 6 2 10.0 
              +-- Ensidens_telus       
        +-----+                        
        |     +-- Ensidens_ingallsianus
  +-----+                              
  |     |  +----- Ensidens_sagittarius 
  |     +--+                           
  |        |  +-- Ensidens_spiculus    
  |        +--+                        
--+           +-- Ensidens_jaculus     
  |                                    
  |           +-- Hyriopsis_bialata **    
  |  +--------+                        
  |  |        +-- Hyriopsis_khoratensis **
  +--+                                 
     |  +-------- Hyriopsis_panhai **     
     +--+                              
        |  +----- Hyriopsis_myersiana **  
        +--+                           
           |  +-- Rectidens_sumatrensis
           +--+                        
              +-- Hyriopsis_desowitzi **

Bolotov et al., 2017 [Publ 3761], fig. S2

iT tT bi riS oT roT roS iD
5 11 7 5 133 6 2 14.0 
           +----- Hyriopsis_panhai **     
           |                           
  +--------+----- Hyriopsis_khoratensis **
  |        |                           
  |        |  +-- Hyriopsis_myersiana **  
  |        +--+                        
--+           +-- Hyriopsis_bialata **    
  |                                    
  |  +----------- Hyriopsis_desowitzi **  
  |  |                                 
  |  +----------- Rectidens_sumatrensis
  +--+                                 
     |        +-- Ensidens_telus       
     |  +-----+                        
     |  |     +-- Ensidens_ingallsianus
     +--+                              
        |  +----- Ensidens_sagittarius 
        +--+                           
           |  +-- Ensidens_spiculus    
           +--+                        
              +-- Ensidens_jaculus

Bolotov et al., 2017 [Publ 3764], fig. S1

iT tT bi riS oT roT roS iD
5 6 2 5 123 6 2 6.0 
           +-- Ensidens_telus       
     +-----+                        
     |     +-- Ensidens_ingallsianus
  +--+                              
  |  |  +----- Ensidens_jaculus     
  |  +--+                           
  |     |  +-- Ensidens_spiculus    
  |     +--+                        
--+        +-- Ensidens_sagittarius 
  |                                 
  |  +-------- Hyriopsis_desowitzi **  
  |  |                              
  |  +-------- Rectidens_sumatrensis
  +--+                              
     |  +----- Hyriopsis_panhai **     
     |  |                           
     +--+  +-- Hyriopsis_myersiana **  
        |  |                        
        +--+-- Hyriopsis_bialata **    
           |                        
           +-- Hyriopsis_khoratensis **

Zieritz et al., 2016 [Publ 3804], fig. S2-1

iT tT bi riS oT roT roS iD
5 5 2 5 1 1 1 8.0 
  +----------- Rectidens_sumatrensis
  |                                 
--+  +-------- Hyriopsis_desowitzi **  
  |  |                              
  +--+  +----- Hyriopsis_khoratensis **
     |  |                           
     +--+  +-- Hyriopsis_myersiana **  
        |  |                        
        +--+-- Hyriopsis_panhai **     
           |                        
           +-- Hyriopsis_bialata **

Huang et al., 2019 [Publ 3843], fig. 2

iT tT bi riS oT roT roS iD
5 11 5 5 196 6 2 12.0 
  +----------- Hyriopsis_desowitzi **  
  |                                 
  +----------- Rectidens_sumatrensis
  |                                 
  |        +-- Hyriopsis_bialata **    
  |        |                        
  |        +-- Hyriopsis_myersiana **  
--+--------+                        
  |        +-- Hyriopsis_panhai **     
  |        |                        
  |        +-- Hyriopsis_khoratensis **
  |                                 
  |        +-- Ensidens_ingallsianus
  |  +-----+                        
  |  |     +-- Ensidens_telus       
  +--+                              
     |  +----- Ensidens_sagittarius 
     +--+                           
        |  +-- Ensidens_spiculus    
        +--+                        
           +-- Ensidens_jaculus

Huang et al., 2019 [Publ 3843], fig. S3

iT tT bi riS oT roT roS iD
5 6 4 5 193 6 2 8.0 
              +-- Ensidens_telus       
        +-----+                        
        |     +-- Ensidens_ingallsianus
  +-----+                              
  |     |  +----- Ensidens_sagittarius 
  |     +--+                           
  |        |  +-- Ensidens_spiculus    
--+        +--+                        
  |           +-- Ensidens_jaculus     
  |                                    
  |  +----------- Hyriopsis_desowitzi **  
  |  |                                 
  +--+  +-------- Rectidens_sumatrensis
     |  |                              
     +--+     +-- Hyriopsis_panhai **     
        |  +--+                        
        |  |  +-- Hyriopsis_khoratensis **
        +--+                           
           |  +-- Hyriopsis_bialata **    
           +--+                        
              +-- Hyriopsis_myersiana **

Huang et al., 2019 [Publ 3843], fig. S4

iT tT bi riS oT roT roS iD
5 11 6 5 195 6 2 12.0 
  +-------------- Hyriopsis_desowitzi **  
  |                                    
  +-------------- Hyriopsis_myersiana **  
  |                                    
  +-------------- Hyriopsis_panhai **     
  |                                    
--+           +-- Hyriopsis_bialata **    
  +-----------+                        
  |           +-- Hyriopsis_khoratensis **
  |                                    
  |  +----------- Rectidens_sumatrensis
  |  |                                 
  +--+        +-- Ensidens_telus       
     |  +-----+                        
     |  |     +-- Ensidens_ingallsianus
     +--+                              
        |  +----- Ensidens_sagittarius 
        +--+                           
           |  +-- Ensidens_spiculus    
           +--+                        
              +-- Ensidens_jaculus

Konopleva et al., 2019 [Publ 3900], fig. 1

iT tT bi riS oT roT roS iD
5 6 3 5 47 6 2 5.0 
              +-- Ensidens_ingallsianus
        +-----+                        
        |     +-- Ensidens_telus       
  +-----+                              
  |     |  +----- Ensidens_sagittarius 
  |     +--+                           
  |        |  +-- Ensidens_jaculus     
--+        +--+                        
  |           +-- Ensidens_spiculus    
  |                                    
  |  +----------- Hyriopsis_desowitzi **  
  |  |                                 
  +--+----------- Rectidens_sumatrensis
     |                                 
     |  +-------- Hyriopsis_panhai **     
     +--+                              
        |  +----- Hyriopsis_myersiana **  
        +--+                           
           |  +-- Hyriopsis_khoratensis **
           +--+                        
              +-- Hyriopsis_bialata **

Bolotov et al., 2020 [Publ 4100], fig. 1

iT tT bi riS oT roT roS iD
5 6 4 5 116 7 2 6.0 
     +----------- Hyriopsis_desowitzi **  
     |                                 
  +--+  +-------- Rectidens_sumatrensis
  |  |  |                              
  |  +--+     +-- Hyriopsis_khoratensis **
  |     |  +--+                        
  |     |  |  +-- Hyriopsis_bialata **    
  |     +--+                           
  |        |  +-- Hyriopsis_myersiana **  
--+        +--+                        
  |           +-- Hyriopsis_panhai **     
  |                                    
  |           +-- Ensidens_telus       
  |     +-----+                        
  |     |     +-- Ensidens_ingallsianus
  |     |                              
  +-----+     +-- Ensidens_jaculus     
        |  +--+                        
        |  |  +-- Ensidens_spiculus    
        +--+                           
           |  +-- Ensidens_dugasti     
           +--+                        
              +-- Ensidens_sagittarius

Bolotov et al., 2020 [Publ 4100], fig. S1

iT tT bi riS oT roT roS iD
5 6 4 5 110 7 2 4.0 
     +-------------- Hyriopsis_desowitzi **  
  +--+                                    
  |  |  +----------- Rectidens_sumatrensis
  |  +--+                                 
  |     |  +-------- Hyriopsis_panhai **     
  |     +--+                              
  |        |  +----- Hyriopsis_myersiana **  
  |        +--+                           
--+           |  +-- Hyriopsis_khoratensis **
  |           +--+                        
  |              +-- Hyriopsis_bialata **    
  |                                       
  |              +-- Ensidens_telus       
  |        +-----+                        
  |        |     +-- Ensidens_ingallsianus
  |        |                              
  +--------+     +-- Ensidens_dugasti     
           |  +--+                        
           |  |  +-- Ensidens_sagittarius 
           +--+                           
              |  +-- Ensidens_spiculus    
              +--+                        
                 +-- Ensidens_jaculus

Zieritz et al., 2020 [Publ 4102], fig. 4

iT tT bi riS oT roT roS iD
5 6 2 5 26 6 2 6.0 
           +-- Ensidens_telus       
     +-----+                        
     |     +-- Ensidens_ingallsianus
  +--+                              
  |  |  +----- Ensidens_sagittarius 
  |  +--+                           
  |     |  +-- Ensidens_spiculus    
  |     +--+                        
--+        +-- Ensidens_jaculus     
  |                                 
  |  +-------- Hyriopsis_desowitzi **  
  |  |                              
  |  +-------- Rectidens_sumatrensis
  +--+                              
     |  +----- Hyriopsis_panhai **     
     |  |                           
     +--+  +-- Hyriopsis_khoratensis **
        |  |                        
        +--+-- Hyriopsis_bialata **    
           |                        
           +-- Hyriopsis_myersiana **

Muanta et al., 2019 [Publ 3921], fig. 2

iT tT bi riS oT roT roS iD
4 4 2 4 6 6 1 8.0 
        +-------- Hyriopsis_desowitzi **    
  +-----+                                
  |     |  +----- Hyriopsis_myersiana **    
  |     +--+                             
  |        |  +-- Hyriopsis_khoratensis **  
  |        +--+                          
--+           +-- Hyriopsis_panhai **       
  |                                      
  |           +-- Ensidens_ingallsianus  
  |  +--------+                          
  |  |        +-- Ensidens_telus         
  +--+                                   
     |  +-------- Ensidens_sagittarius[2]
     +--+                                
        |  +----- Ensidens_jaculus       
        +--+                             
           |  +-- Ensidens_sagittarius[1]
           +--+                          
              +-- Ensidens_dugasti

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iT tT bi riS oT roT roS iD
3 7 2 3 78 4 2 10.0 
  +----- Hyriopsis_desowitzi **  
  |                           
  +----- Rectidens_sumatrensis
  |                           
  |  +-- Hyriopsis_khoratensis **
--+--+                        
  |  +-- Hyriopsis_myersiana **  
  |                           
  |  +-- Ensidens_ingallsianus
  |  |                        
  +--+-- Ensidens_telus       
     |                        
     +-- Ensidens_jaculus

Bolotov et al., 2018 [Publ 3826], fig. 2

iT tT bi riS oT roT roS iD
3 8 0 3 82 5 1 13.0 
  +-- Ensidens_telus       
  |                        
  +-- Ensidens_ingallsianus
  |                        
  +-- Ensidens_jaculus     
  |                        
  +-- Hyriopsis_desowitzi **  
--+                        
  +-- Hyriopsis_myersiana **  
  |                        
  +-- Hyriopsis_khoratensis **
  |                        
  +-- Ensidens_spiculus    
  |                        
  +-- Ensidens_sagittarius

Pfeiffer et al., 2018 [Publ 3827], fig. 1

iT tT bi riS oT roT roS iD
2 4 0 2 6 2 2 11.0 
  +-- Ensidens    
  |               
  +-- Hyriopsis[1] **
--+               
  +-- Hyriopsis[2] **
  |               
  +-- Rectidens

Zieritz et al., 2021 [Publ 4218], fig. 5

iT tT bi riS oT roT roS iD
2 2 0 2 30 2 2 -- 
  +-------- Ensidens_sagittarius 
--+                              
  |  +----- Rectidens_sumatrensis
  +--+                           
     |  +-- Hyriopsis_bialata **    
     +--+                        
        +-- Hyriopsis_desowitzi **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iT tT bi riS oT roT roS iD
1 1 0 1 56 2 2 -- 
  +----- Ensidens_ingallsianus
--+                           
  |  +-- Hyriopsis_bialata **    
  +--+                        
     +-- Rectidens_sumatrensis

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 56 2 2 -- 
  +----- Ensidens_ingallsianus
--+                           
  |  +-- Hyriopsis_bialata **    
  +--+                        
     +-- Rectidens_sumatrensis

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 36 2 2 -- 
  +----- Ensidens_ingallsianus
--+                           
  |  +-- Hyriopsis_bialata **    
  +--+                        
     +-- Rectidens_sumatrensis

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 36 2 2 -- 
  +----- Ensidens_ingallsianus
--+                           
  |  +-- Hyriopsis_bialata **    
  +--+                        
     +-- Rectidens_sumatrensis

Zieritz et al., 2021 [Publ 4231], fig. 3-F, 5

iT tT bi riS oT roT roS iD
1 1 0 1 48 1 1 -- 
  +-- Hyriopsis_bialata **    
--+                        
  +-- Rectidens_sumatrensis

Zieritz et al., 2021 [Publ 4231], fig. 3-M, 4

iT tT bi riS oT roT roS iD
1 1 0 1 22 1 1 -- 
  +-- Hyriopsis_bialata **    
--+                        
  +-- Rectidens_sumatrensis

Jeratthitikul et al., 2022 [Publ 4260], fig. S1, 1

iT tT bi riS oT roT roS iD
1 1 0 1 75 2 2 -- 
  +-- Ensidens_ingallsianus
  |                        
--+-- Hyriopsis_bialata **    
  |                        
  +-- Rectidens_sumatrensis

Jeratthitikul et al., 2022 [Publ 4260], fig. S2, 2

iT tT bi riS oT roT roS iD
1 1 0 1 75 2 2 -- 
  +----- Ensidens_ingallsianus
--+                           
  |  +-- Rectidens_sumatrensis
  +--+                        
     +-- Hyriopsis_bialata **

Dai et al., 2023 [Publ 4285], fig. 3-BI

iT tT bi riS oT roT roS iD
1 1 0 1 43 1 1 -- 
  +-- Hyriopsis_bialata **    
--+                        
  +-- Ensidens_ingallsianus

Dai et al., 2023 [Publ 4285], fig. 3-ML

iT tT bi riS oT roT roS iD
1 1 0 1 43 1 1 -- 
  +-- Hyriopsis_bialata **    
--+                        
  +-- Ensidens_ingallsianus

Dai et al., 2023 [Publ 4285], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 52 1 1 -- 
  +-- Hyriopsis_bialata **    
--+                        
  +-- Rectidens_sumatrensis

Page updated: 10:01:26 Thu 07 Sep 2023
 
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