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Page last updated
Mon 13 Mar 2023

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Gonideinae | Rectidentini

genus Hyriopsis Conrad, 1853: cladomics
     10 valid species

Cladomics Summary

number of species = 10

analyzed in 29 cladograms in 20 publications
maximum number of terminal species in an analysis = 9, mean = 3.93

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 21
maximum number of included species = 9, mean = 5.05
proportion of cladograms that support basic monophyly = 0.24

Konopleva et al., 2023, fig. 8

iT tT bi riS oT roT roS iD
9 10 7 9 74 7 2 0.0

*** Top Monophyly Cladogram *** 

                       +-- Ensidens_ingallsianus  
                 +-----+                          
                 |     +-- Ensidens_telus         
                 |                                
  +--------------+     +-- Ensidens_jaculus       
  |              |  +--+                          
  |              |  |  +-- Ensidens               
  |              +--+                             
  |                 |  +-- Ensidens_spiculus      
--+                 +--+                          
  |                    +-- Ensidens_dugasti       
  |                                               
  |                    +-- Hyriopsis_desowitzi **    
  |  +-----------------+                          
  |  |                 +-- Hyriopsis_delaportei **   
  +--+                                            
     |  +----------------- Rectidens_sumatrensis  
     |  |                                         
     +--+  +-------------- Hyriopsis_panhai **       
        |  |                                      
        +--+  +----------- Hyriopsis_myersiana **    
           |  |                                   
           +--+        +-- Hyriopsis_khoratensis **  
              |  +-----+                          
              |  |     +-- Hyriopsis_bialata **      
              +--+                                
                 |  +----- Hyriopsis_sakhonensis **  
                 +--+                             
                    |  +-- Hyriopsis_phuphaniensis **
                    +--+                          
                       +-- Hyriopsis_kratiensis **

Zieritz et al., 2021, fig. 5

iT tT bi riS oT roT roS iD
2 2 0 2 29 5 4 --

*** Top Sister-Group Cladogram ***

  +-------- Ensidens_sagittarius 
--+                              
  |  +----- Rectidens_sumatrensis
  +--+                           
     |  +-- Hyriopsis_bialata **    
     +--+                        
        +-- Hyriopsis_desowitzi **

Pfeiffer et al., 2021, fig. 27

iT tT bi riS oT roT roS iD
9 9 4 9 2 2 2 7.0 
  +----------------- Ctenodesma_borneensis  
  |                                         
--+  +-------------- Rectidens_sumatrensis  
  |  |                                      
  |  |           +-- Hyriopsis_delaportei **   
  +--+  +--------+                          
     |  |        +-- Hyriopsis_desowitzi **    
     |  |                                   
     +--+  +-------- Hyriopsis_khoratensis **  
        |  |                                
        |  |     +-- Hyriopsis_kratiensis **   
        |  +-----+                          
        +--+     +-- Hyriopsis_phuphaniensis **
           |                                
           |  +----- Hyriopsis_sakhonensis **  
           |  |                             
           +--+  +-- Hyriopsis_myersiana **    
              |  |                          
              +--+-- Hyriopsis_panhai **       
                 |                          
                 +-- Hyriopsis_bialata **

Jeratthitkul et al., 2021, fig. 4

iT tT bi riS oT roT roS iD
8 9 6 8 13 3 2 7.0 
                 +-- Ensidens_ingallsianus  
  +--------------+                          
  |              +-- Ensidens_jaculus       
  |                                         
--+              +-- Hyriopsis_desowitzi **    
  |  +-----------+                          
  |  |           +-- Hyriopsis_delaportei **   
  |  |                                      
  +--+  +----------- Rectidens_sumatrensis  
     |  |                                   
     |  |     +----- Hyriopsis_myersiana **    
     +--+  +--+                             
        |  |  |  +-- Hyriopsis_bialata **      
        |  |  +--+                          
        +--+     +-- Hyriopsis_panhai **       
           |                                
           |  +----- Hyriopsis_khoratensis **  
           +--+                             
              |  +-- Hyriopsis_phuphaniensis **
              +--+                          
                 +-- Hyriopsis_sakhonensis **

Pfeiffer et al., 2021, fig. 9

iT tT bi riS oT roT roS iD
6 6 4 6 23 7 2 6.0 
                    +-- Ensidens_ingallsianus
           +--------+                        
           |        +-- Ensidens_telus       
  +--------+                                 
  |        |  +-------- Ensidens_spiculus    
  |        +--+                              
  |           |  +----- Ensidens_dugasti     
  |           +--+                           
--+              |  +-- Ensidens_jaculus     
  |              +--+                        
  |                 +-- Ensidens_sagittarius 
  |                                          
  |  +----------------- Rectidens_sumatrensis
  +--+                                       
     |  +-------------- Hyriopsis_delaportei ** 
     +--+                                    
        |  +----------- Hyriopsis_myersiana **  
        +--+                                 
           |  +-------- Hyriopsis_bialata **    
           +--+                              
              |  +----- Hyriopsis_khoratensis **
              +--+                           
                 |  +-- Hyriopsis_kratiensis ** 
                 +--+                        
                    +-- Hyriopsis_sakhonensis **

Bolotov et al., 2017, fig. S1

iT tT bi riS oT roT roS iD
5 6 3 5 133 6 2 7.0 
              +-- Ensidens_telus       
        +-----+                        
        |     +-- Ensidens_ingallsianus
  +-----+                              
  |     |  +----- Ensidens_sagittarius 
  |     +--+                           
  |        |  +-- Ensidens_spiculus    
  |        +--+                        
--+           +-- Ensidens_jaculus     
  |                                    
  |           +-- Hyriopsis_bialata **    
  |  +--------+                        
  |  |        +-- Hyriopsis_khoratensis **
  +--+                                 
     |  +-------- Hyriopsis_panhai **     
     +--+                              
        |  +----- Hyriopsis_myersiana **  
        +--+                           
           |  +-- Rectidens_sumatrensis
           +--+                        
              +-- Hyriopsis_desowitzi **

Bolotov et al., 2017, fig. S2

iT tT bi riS oT roT roS iD
5 11 6 5 133 6 2 13.0 
           +----- Hyriopsis_panhai **     
           |                           
  +--------+----- Hyriopsis_khoratensis **
  |        |                           
  |        |  +-- Hyriopsis_myersiana **  
  |        +--+                        
--+           +-- Hyriopsis_bialata **    
  |                                    
  |  +----------- Hyriopsis_desowitzi **  
  |  |                                 
  |  +----------- Rectidens_sumatrensis
  +--+                                 
     |        +-- Ensidens_telus       
     |  +-----+                        
     |  |     +-- Ensidens_ingallsianus
     +--+                              
        |  +----- Ensidens_sagittarius 
        +--+                           
           |  +-- Ensidens_spiculus    
           +--+                        
              +-- Ensidens_jaculus

Bolotov et al., 2017, fig. S1

iT tT bi riS oT roT roS iD
5 6 2 5 120 6 2 6.0 
           +-- Ensidens_telus       
     +-----+                        
     |     +-- Ensidens_ingallsianus
  +--+                              
  |  |  +----- Ensidens_jaculus     
  |  +--+                           
  |     |  +-- Ensidens_spiculus    
  |     +--+                        
--+        +-- Ensidens_sagittarius 
  |                                 
  |  +-------- Hyriopsis_desowitzi **  
  |  |                              
  |  +-------- Rectidens_sumatrensis
  +--+                              
     |  +----- Hyriopsis_panhai **     
     |  |                           
     +--+  +-- Hyriopsis_myersiana **  
        |  |                        
        +--+-- Hyriopsis_bialata **    
           |                        
           +-- Hyriopsis_khoratensis **

Zieritz et al., 2016, fig. S2-1

iT tT bi riS oT roT roS iD
5 5 2 5 1 1 1 8.0 
  +----------- Rectidens_sumatrensis
  |                                 
--+  +-------- Hyriopsis_desowitzi **  
  |  |                              
  +--+  +----- Hyriopsis_khoratensis **
     |  |                           
     +--+  +-- Hyriopsis_myersiana **  
        |  |                        
        +--+-- Hyriopsis_panhai **     
           |                        
           +-- Hyriopsis_bialata **

Huang et al., 2019, fig. 2

iT tT bi riS oT roT roS iD
5 11 5 5 194 6 2 12.0 
  +----------- Hyriopsis_desowitzi **  
  |                                 
  +----------- Rectidens_sumatrensis
  |                                 
  |        +-- Hyriopsis_bialata **    
  |        |                        
  |        +-- Hyriopsis_myersiana **  
--+--------+                        
  |        +-- Hyriopsis_panhai **     
  |        |                        
  |        +-- Hyriopsis_khoratensis **
  |                                 
  |        +-- Ensidens_ingallsianus
  |  +-----+                        
  |  |     +-- Ensidens_telus       
  +--+                              
     |  +----- Ensidens_sagittarius 
     +--+                           
        |  +-- Ensidens_spiculus    
        +--+                        
           +-- Ensidens_jaculus

Huang et al., 2019, fig. S3

iT tT bi riS oT roT roS iD
5 6 4 5 192 6 2 8.0 
              +-- Ensidens_telus       
        +-----+                        
        |     +-- Ensidens_ingallsianus
  +-----+                              
  |     |  +----- Ensidens_sagittarius 
  |     +--+                           
  |        |  +-- Ensidens_spiculus    
--+        +--+                        
  |           +-- Ensidens_jaculus     
  |                                    
  |  +----------- Hyriopsis_desowitzi **  
  |  |                                 
  +--+  +-------- Rectidens_sumatrensis
     |  |                              
     +--+     +-- Hyriopsis_panhai **     
        |  +--+                        
        |  |  +-- Hyriopsis_khoratensis **
        +--+                           
           |  +-- Hyriopsis_bialata **    
           +--+                        
              +-- Hyriopsis_myersiana **

Huang et al., 2019, fig. S4

iT tT bi riS oT roT roS iD
5 11 6 5 194 6 2 12.0 
  +-------------- Hyriopsis_desowitzi **  
  |                                    
  +-------------- Hyriopsis_myersiana **  
  |                                    
  +-------------- Hyriopsis_panhai **     
  |                                    
--+           +-- Hyriopsis_bialata **    
  +-----------+                        
  |           +-- Hyriopsis_khoratensis **
  |                                    
  |  +----------- Rectidens_sumatrensis
  |  |                                 
  +--+        +-- Ensidens_telus       
     |  +-----+                        
     |  |     +-- Ensidens_ingallsianus
     +--+                              
        |  +----- Ensidens_sagittarius 
        +--+                           
           |  +-- Ensidens_spiculus    
           +--+                        
              +-- Ensidens_jaculus

Konopleva et al., 2019, fig. 1

iT tT bi riS oT roT roS iD
5 6 3 5 47 6 2 5.0 
              +-- Ensidens_ingallsianus
        +-----+                        
        |     +-- Ensidens_telus       
  +-----+                              
  |     |  +----- Ensidens_sagittarius 
  |     +--+                           
  |        |  +-- Ensidens_jaculus     
--+        +--+                        
  |           +-- Ensidens_spiculus    
  |                                    
  |  +----------- Hyriopsis_desowitzi **  
  |  |                                 
  +--+----------- Rectidens_sumatrensis
     |                                 
     |  +-------- Hyriopsis_panhai **     
     +--+                              
        |  +----- Hyriopsis_myersiana **  
        +--+                           
           |  +-- Hyriopsis_khoratensis **
           +--+                        
              +-- Hyriopsis_bialata **

Konopleva et al., 2019, fig. 3

iT tT bi riS oT roT roS iD
5 13 0 5 30 8 2 15.0 
  +-- Ensidens_ingallsianus
  |                        
  +-- Ensidens_telus       
  |                        
  +-- Ensidens_dugasti     
  |                        
  +-- Hyriopsis_khoratensis **
  |                        
  +-- Hyriopsis_bialata **    
  |                        
  +-- Hyriopsis_myersiana **  
  |                        
--+-- Hyriopsis_desowitzi **  
  |                        
  +-- Rectidens_sumatrensis
  |                        
  +-- Hyriopsis_panhai **     
  |                        
  +-- Ensidens_jaculus[1]  
  |                        
  +-- Ensidens_jaculus[2]  
  |                        
  +-- Ensidens_spiculus    
  |                        
  +-- Ensidens_sagittarius

Bolotov et al., 2020, fig. 1

iT tT bi riS oT roT roS iD
5 6 4 5 116 7 2 6.0 
     +----------- Hyriopsis_desowitzi **  
     |                                 
  +--+  +-------- Rectidens_sumatrensis
  |  |  |                              
  |  +--+     +-- Hyriopsis_khoratensis **
  |     |  +--+                        
  |     |  |  +-- Hyriopsis_bialata **    
  |     +--+                           
  |        |  +-- Hyriopsis_myersiana **  
--+        +--+                        
  |           +-- Hyriopsis_panhai **     
  |                                    
  |           +-- Ensidens_telus       
  |     +-----+                        
  |     |     +-- Ensidens_ingallsianus
  |     |                              
  +-----+     +-- Ensidens_jaculus     
        |  +--+                        
        |  |  +-- Ensidens_spiculus    
        +--+                           
           |  +-- Ensidens_dugasti     
           +--+                        
              +-- Ensidens_sagittarius

Bolotov et al., 2020, fig. S1

iT tT bi riS oT roT roS iD
5 6 4 5 110 7 2 4.0 
     +-------------- Hyriopsis_desowitzi **  
  +--+                                    
  |  |  +----------- Rectidens_sumatrensis
  |  +--+                                 
  |     |  +-------- Hyriopsis_panhai **     
  |     +--+                              
  |        |  +----- Hyriopsis_myersiana **  
  |        +--+                           
--+           |  +-- Hyriopsis_khoratensis **
  |           +--+                        
  |              +-- Hyriopsis_bialata **    
  |                                       
  |              +-- Ensidens_telus       
  |        +-----+                        
  |        |     +-- Ensidens_ingallsianus
  |        |                              
  +--------+     +-- Ensidens_dugasti     
           |  +--+                        
           |  |  +-- Ensidens_sagittarius 
           +--+                           
              |  +-- Ensidens_spiculus    
              +--+                        
                 +-- Ensidens_jaculus

Zieritz et al., 2020, fig. 4

iT tT bi riS oT roT roS iD
5 6 2 5 26 7 3 6.0 
  +-------------- Ctenodesma_borneensis
  |                                    
  |           +-- Ensidens_telus       
  |     +-----+                        
  |     |     +-- Ensidens_ingallsianus
--+  +--+                              
  |  |  |  +----- Ensidens_sagittarius 
  |  |  +--+                           
  |  |     |  +-- Ensidens_spiculus    
  |  |     +--+                        
  +--+        +-- Ensidens_jaculus     
     |                                 
     |  +-------- Hyriopsis_desowitzi **  
     |  |                              
     |  +-------- Rectidens_sumatrensis
     +--+                              
        |  +----- Hyriopsis_panhai **     
        |  |                           
        +--+  +-- Hyriopsis_khoratensis **
           |  |                        
           +--+-- Hyriopsis_bialata **    
              |                        
              +-- Hyriopsis_myersiana **

Muanta et al., 2019, fig. 2

iT tT bi riS oT roT roS iD
4 4 2 4 6 6 1 8.0 
        +-------- Hyriopsis_desowitzi **    
  +-----+                                
  |     |  +----- Hyriopsis_myersiana **    
  |     +--+                             
  |        |  +-- Hyriopsis_khoratensis **  
  |        +--+                          
--+           +-- Hyriopsis_panhai **       
  |                                      
  |           +-- Ensidens_ingallsianus  
  |  +--------+                          
  |  |        +-- Ensidens_telus         
  +--+                                   
     |  +-------- Ensidens_sagittarius[2]
     +--+                                
        |  +----- Ensidens_jaculus       
        +--+                             
           |  +-- Ensidens_sagittarius[1]
           +--+                          
              +-- Ensidens_dugasti

Lopes-Lima et al., 2017, fig. 1-2

iT tT bi riS oT roT roS iD
3 7 2 3 78 4 2 10.0 
  +----- Hyriopsis_desowitzi **  
  |                           
  +----- Rectidens_sumatrensis
  |                           
  |  +-- Hyriopsis_khoratensis **
--+--+                        
  |  +-- Hyriopsis_myersiana **  
  |                           
  |  +-- Ensidens_ingallsianus
  |  |                        
  +--+-- Ensidens_telus       
     |                        
     +-- Ensidens_jaculus

Bolotov et al., 2018, fig. 2

iT tT bi riS oT roT roS iD
3 8 0 3 82 5 1 13.0 
  +-- Ensidens_telus       
  |                        
  +-- Ensidens_ingallsianus
  |                        
  +-- Ensidens_jaculus     
  |                        
  +-- Hyriopsis_desowitzi **  
--+                        
  +-- Hyriopsis_myersiana **  
  |                        
  +-- Hyriopsis_khoratensis **
  |                        
  +-- Ensidens_spiculus    
  |                        
  +-- Ensidens_sagittarius

Pfeiffer et al., 2018, fig. 1

iT tT bi riS oT roT roS iD
2 4 0 2 6 2 2 11.0 
  +-- Ensidens    
  |               
  +-- Hyriopsis[1] **
--+               
  +-- Hyriopsis[2] **
  |               
  +-- Rectidens

Pfeiffer et al., 2019, fig. 4-5

iT tT bi riS oT roT roS iD
1 1 0 1 56 2 2 -- 
  +----- Ensidens_ingallsianus
--+                           
  |  +-- Hyriopsis_bialata **    
  +--+                        
     +-- Rectidens_sumatrensis

Pfeiffer et al., 2019, fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 56 2 2 -- 
  +----- Ensidens_ingallsianus
--+                           
  |  +-- Hyriopsis_bialata **    
  +--+                        
     +-- Rectidens_sumatrensis

Jeratthitikul et al., 2021, fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 36 2 2 -- 
  +----- Ensidens_ingallsianus
--+                           
  |  +-- Hyriopsis_bialata **    
  +--+                        
     +-- Rectidens_sumatrensis

Jeratthitikul et al., 2021, fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 35 2 2 -- 
  +----- Ensidens_ingallsianus
--+                           
  |  +-- Hyriopsis_bialata **    
  +--+                        
     +-- Rectidens_sumatrensis

Zieritz et al., 2021, fig. 3-F, 5

iT tT bi riS oT roT roS iD
1 1 0 1 48 1 1 -- 
  +-- Hyriopsis_bialata **    
--+                        
  +-- Rectidens_sumatrensis

Zieritz et al., 2021, fig. 3-M, 4

iT tT bi riS oT roT roS iD
1 1 0 1 22 1 1 -- 
  +-- Hyriopsis_bialata **    
--+                        
  +-- Rectidens_sumatrensis

Jeratthitikul et al., 2022, fig. S1, 1

iT tT bi riS oT roT roS iD
1 1 0 1 75 2 2 -- 
  +-- Ensidens_ingallsianus
  |                        
--+-- Hyriopsis_bialata **    
  |                        
  +-- Rectidens_sumatrensis

Jeratthitikul et al., 2022, fig. S2, 2

iT tT bi riS oT roT roS iD
1 1 0 1 75 2 2 -- 
  +----- Ensidens_ingallsianus
--+                           
  |  +-- Rectidens_sumatrensis
  +--+                        
     +-- Hyriopsis_bialata **

Page updated: 09:30:54 Mon 13 Mar 2023
 
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