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Page last updated
Thu 04 Apr 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Ambleminae

tribe PLEUROBEMINI Hannibal, 1912: cladomics
     9 genera

Cladomics Summary

number of species = 105
number of subtaxa (rank: genus) = 9

analyzed in 136 cladograms in 84 publications
maximum number of terminals in an analysis = 157, mean = 8.3
maximum number of subtaxa (genus) in an analysis = 9, mean = 3.12

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 95
maximum number of included terminals = 157, mean = 11.45
maximum number of subtaxa (genus) = 9, mean = 4.04
proportion of cladograms that support basic monophyly = 0.77

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 88
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (tribe).
maximum number of included terminals = 157, mean = 11.64
maximum number of subtaxa (genus) = 9, mean = 4.17
proportion of cladograms that support rigorous monophyly = 0.75


Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ tree


Campbell & Lydeard, 2012 [Publ 3308], fig. 3

iT tT bi riS oT roT roS iD
32 44 35 9 19 19 4 0.0

*** Top Monophyly Cladogram ***

                                   +-------------- Uniomerus_declivis      
                                   |                                       
  +--------------------------------+           +-- Pustulosa_infucata      
  |                                |  +--------+                           
  |                                |  |        +-- Pustulosa_kleiniana     
  |                                +--+                                    
  |                                   |  +-------- Quadrula_quadrula       
  |                                   +--+                                 
  |                                      |  +----- Cyclonaias_tuberculata  
  |                                      +--+                              
  |                                         |  +-- Pustulosa_succissa      
  |                                         +--+                           
  |                                            +-- Pustulosa_pustulosa     
--+                                                                        
  |                                +-------------- Elliptoideus_sloatianus ** 
  |  +-----------------------------+                                       
  |  |                             |  +----------- Plectomerus_dombeyanus  
  |  |                             +--+                                    
  |  |                                |  +-------- Obovaria_unicolor       
  |  |                                +--+                                 
  |  |                                   |  +----- Obovaria_olivaria       
  |  |                                   +--+                              
  |  |                                      |  +-- Lampsilis_ovata         
  |  |                                      +--+                           
  |  |                                         +-- Lemiox_rimosus          
  +--+                                                                     
     |                                +----------- Cyrtonaias_tampicoensis 
     |                                |                                    
     |                                |     +----- Popenaias_popeii        
     |  +-----------------------------+  +--+                              
     |  |                             |  |  |  +-- Amblema_elliottii       
     |  |                             |  |  +--+                           
     |  |                             +--+     +-- Amblema_plicata         
     |  |                                |                                 
     |  |                                |     +-- Reginaia_ebenus[1]      
     |  |                                |     |                           
     +--+                                +-----+-- Reginaia_ebenus[2]      
        |                                      |                           
        |                                      +-- Reginaia_rotulata       
        |                                                                  
        |                                      +-- Elliptio_arca **           
        |  +-----------------------------------+                           
        |  |                                   +-- Elliptio_crassidens **     
        |  |                                                               
        +--+  +----------------------------------- Eurynia_dilatata **        
           |  |                                                            
           |  |  +-------------------------------- Parvaspina_collina **      
           |  |  |                                                         
           +--+  |                             +-- Hemistena_lata **          
              |  +-----------------------------+                           
              |  |                             +-- Plethobasus_cyphyus **     
              |  |                                                         
              +--+                          +----- Pleuronaia_barnesiana **   
                 |  +-----------------------+                              
                 |  |                       |  +-- Pleuronaia_dolabelloides **
                 |  |                       +--+                           
                 |  |                          +-- Pleuronaia_gibber **       
                 |  |                                                      
                 |  |                          +-- Pleurobema_stabile **      
                 +--+                    +-----+                           
                    |                    |     +-- Pleurobema_sintoxia **     
                    |  +-----------------+                                 
                    |  |                 |  +----- Pleurobema_decisum **      
                    |  |                 +--+                              
                    |  |                    |  +-- Pleurobema_clava **        
                    +--+                    +--+                           
                       |                       +-- Pleurobema_pyriforme **    
                       |                                                   
                       |  +----------------------- Pleurobema_cordatum **     
                       |  |                                                
                       +--+  +-------------------- Fusconaia_subrotunda **    
                          |  |                                             
                          |  |                 +-- Fusconaia_cor **           
                          +--+  +--------------+                           
                             |  |              +-- Fusconaia_cuneolus **      
                             |  |                                          
                             +--+  +-------------- Fusconaia_ozarkensis **    
                                |  |                                       
                                |  |           +-- Fusconaia_flava[6] **      
                                |  |  +--------+                           
                                +--+  |        +-- Fusconaia_masoni **        
                                   |  |                                    
                                   |  |        +-- Fusconaia_escambia **      
                                   |  |        |                           
                                   +--+        +-- Fusconaia_burkei[1] **     
                                      |  +-----+                           
                                      |  |     +-- Fusconaia_burkei[2] **     
                                      |  |     |                           
                                      |  |     +-- Fusconaia_burkei[3] **     
                                      +--+                                 
                                         |  +----- Fusconaia_flava[5] **      
                                         |  |                              
                                         |  |  +-- Fusconaia_cerina **        
                                         +--+  |                           
                                            |  +-- Fusconaia_flava[1] **      
                                            |  |                           
                                            +--+-- Fusconaia_flava[2] **      
                                               |                           
                                               +-- Fusconaia_flava[3] **      
                                               |                           
                                               +-- Fusconaia_flava[4] **      

Campbell et al., 2005 [Publ 2660], fig. 2

iT tT bi riS oT roT roS iD
36 36 28 9 79 67 4 39.0

*** Top Sister-Group Cladogram ***

                    +-------------------- Megalonaias_nervosa           
                    |                                                   
  +-----------------+  +----------------- Uniomerus_declivis            
  |                 |  |                                                
  |                 +--+  +-------------- Theliderma_metanevra          
  |                    |  |                                             
  |                    +--+  +----------- Quadrula_quadrula             
  |                       |  |                                          
  |                       |  |        +-- Quadrula_nobilis              
  |                       +--+  +-----+                                 
  |                          |  |     +-- Quadrula_verrucosa            
  |                          |  |                                       
  |                          +--+  +----- Pustulosa_kieneriana          
  |                             |  |                                    
  |                             |  |  +-- Cyclonaias_tuberculata        
  |                             +--+--+                                 
  |                                |  +-- Pustulosa_succissa            
  |                                |                                    
  |                                |  +-- Pustulosa_infucata            
  |                                +--+                                 
  |                                   +-- Pustulosa_kleiniana           
  |                                                                     
  |     +-------------------------------- Elliptoideus_sloatianus **       
  |     |                                                               
  |     |  +----------------------------- Plethobasus_cyphyus **           
  |     |  |                                                            
  |     |  +----------------------------- Parvaspina_collina **            
  |  +--+  |                                                            
  |  |  |  |                          +-- Elliptio_arca **                 
  |  |  |  +--------------------------+                                 
  |  |  |  |                          +-- Elliptio_crassidens **           
  |  |  |  |                                                            
  |  |  |  |  +-------------------------- Eurynia_dilatata **              
  |  |  +--+  |                                                         
--+  |     |  +-------------------------- Hemistena_lata **                
  |  |     |  |                                                         
  |  |     |  |                       +-- Pleurobema_rubrum[1] **          
  |  |     |  |                    +--+                                 
  |  |     |  |                    |  +-- Pleurobema_sintoxia **           
  |  |     |  |  +-----------------+                                    
  |  |     |  |  |                 |  +-- Pleurobema_rubrum[2] **          
  |  |     |  |  |                 +--+                                 
  |  |     +--+  |                    +-- Pleurobema_cordatum **           
  |  |        |  |                                                      
  |  |        |  |        +-------------- Fusconaia_subrotunda **          
  |  |        |  |        |                                             
  |  |        |  |        |           +-- Fusconaia_cor **                 
  |  |        |  +--------+  +--------+                                 
  |  |        |  |        |  |        +-- Fusconaia_cuneolus **            
  |  |        |  |        |  |                                          
  |  |        |  |        +--+        +-- Fusconaia_cerina **              
  |  |        |  |           |  +-----+                                 
  |  |        |  |           |  |     +-- Fusconaia_flava **               
  |  |        +--+           +--+                                       
  |  |           |              |  +----- Fusconaia_burkei[2] **           
  |  |           |              +--+                                    
  |  |           |                 |  +-- Fusconaia_escambia **            
  |  |           |                 +--+                                 
  |  |           |                    +-- Fusconaia_burkei[1] **           
  |  |           |                                                      
  |  |           |                 +----- Pleuronaia_gibber **             
  |  |           |  +--------------+                                    
  |  |           |  |              |  +-- Pleuronaia_barnesiana **         
  |  |           |  |              +--+                                 
  |  |           |  |                 +-- Pleuronaia_dolabelloides **      
  |  |           |  |                                                   
  |  |           |  |           +-------- Pleurobema_rubellum **           
  +--+           +--+  +--------+                                       
     |              |  |        |  +----- Pleurobema_strodeanum **         
     |              |  |        +--+                                    
     |              |  |           |  +-- Pleurobema_perovatum **          
     |              |  |           +--+                                 
     |              |  |              +-- Pleurobema_taitianum **          
     |              +--+                                                
     |                 |              +-- Pleurobema_oviforme[1] **        
     |                 |              |                                 
     |                 |  +-----------+-- Pleurobema_oviforme[2] **        
     |                 |  |           |                                 
     |                 |  |           +-- Pleurobema_clava **              
     |                 +--+                                             
     |                    |  +----------- Pleurobema_pyriforme **          
     |                    |  |                                          
     |                    +--+        +-- Pleurobema_athearni **           
     |                       |  +-----+                                 
     |                       |  |     +-- Pleurobema_georgianum[2] **      
     |                       +--+                                       
     |                          |  +----- Pleurobema_decisum **            
     |                          |  |                                    
     |                          +--+  +-- Pleurobema_hanleyianum[1] **     
     |                             |  |                                 
     |                             +--+-- Pleurobema_hanleyianum[2] **     
     |                                |                                 
     |                                +-- Pleurobema_georgianum[1] **      
     |                                                                  
     |                          +-------- Popenaias_popeii              
     |                          |                                       
     |           +--------------+     +-- Amblema_elliottii             
     |           |              |  +--+                                 
     |           |              |  |  +-- Amblema_plicata               
     |           |              +--+                                    
     |           |                 |  +-- Reginaia_ebenus               
     |           |                 |  |                                 
     |           |                 +--+-- Reginaia_rotulata[1]          
     |           |                    |                                 
     |           |                    +-- Reginaia_rotulata[2]          
     |           |                                                      
     |           |                 +----- Plectomerus_dombeyanus        
     |           |  +--------------+                                    
     +-----------+  |              |  +-- Toxolasma_parvum              
                 |  |              +--+                                 
                 |  |                 +-- Toxolasma_texasiense          
                 |  |                                                   
                 |  |              +----- Obliquaria_reflexa            
                 |  |  +-----------+                                    
                 |  |  |           |  +-- Cyrtonaias_tampicoensis       
                 |  |  |           +--+                                 
                 |  |  |              +-- Glebula_rotundata             
                 |  |  |                                                
                 |  |  |  +-------------- Sagittunio_nasutus            
                 |  |  |  |                                             
                 +--+  |  +-------------- Leaunio_vanuxemensis          
                    |  |  |                                             
                    |  |  |           +-- Ellipsaria_lineolata          
                    |  |  +-----------+                                 
                    |  |  |           +-- Truncilla_truncata            
                    |  |  |                                             
                    |  |  |        +----- Lampsilis_teres               
                    |  |  +--------+                                    
                    |  |  |        |  +-- Ortmanniana_ligamentina       
                    |  |  |        +--+                                 
                    |  |  |           +-- Lampsilis_siliquoidea         
                    |  |  |                                             
                    |  |  |        +----- Villosa_villosa               
                    |  |  +--------+                                    
                    +--+  |        |  +-- Ligumia_recta                 
                       |  |        +--+                                 
                       |  |           +-- Cambarunio_iris               
                       |  |                                             
                       |  |     +-------- Ortmanniana_pectorosa         
                       |  +-----+                                       
                       |  |     |  +----- Lampsilis_ornata              
                       |  |     +--+                                    
                       |  |        |  +-- Lampsilis_cardium             
                       |  |        +--+                                 
                       |  |           +-- Lampsilis_ovata               
                       |  |                                             
                       |  |           +-- Potamilus_fragilis            
                       |  |        +--+                                 
                       |  |        |  +-- Potamilus_leptodon            
                       |  +--------+                                    
                       |  |        |  +-- Potamilus_alatus              
                       |  |        +--+                                 
                       |  |           +-- Potamilus_purpuratus          
                       |  |                                             
                       |  |           +-- Ptychobranchus_fasciolaris    
                       +--+  +--------+                                 
                          |  |        +-- Venustaconcha_ellipsiformis[1]
                          |  |                                          
                          +--+  +-------- Medionidus_conradicus         
                          |  |  |                                       
                          |  |  |     +-- Lemiox_rimosus                
                          |  +--+-----+                                 
                          |     |     +-- Medionidus_acutissimus        
                          |     |                                       
                          |     |  +----- Dromus_dromas                 
                          |     +--+                                    
                          |        |  +-- Cyprogenia_aberti             
                          |        +--+                                 
                          |           +-- Cyprogenia_stegaria           
                          |                                             
                          |        +----- Hamiota_australis[2]          
                          |  +-----+                                    
                          |  |     |  +-- Hamiota_australis[1]          
                          |  |     +--+                                 
                          |  |        +-- Hamiota_subangulata           
                          +--+                                          
                          |  |        +-- Hamiota_altilis[3]            
                          |  |  +-----+                                 
                          |  |  |     +-- Hamiota_perovalis[2]          
                          |  +--+                                       
                          |     |  +----- Hamiota_altilis[2]            
                          |     +--+                                    
                          |        |  +-- Hamiota_altilis[1]            
                          |        +--+                                 
                          |           +-- Hamiota_perovalis[1]          
                          |                                             
                          |  +----------- Obovaria_olivaria             
                          |  |                                          
                          |  +----------- Venustaconcha_ellipsiformis[2]
                          |  |                                          
                          |  |     +----- Obovaria_unicolor             
                          |  +-----+                                    
                          +--+     |  +-- Obovaria_arkansasensis        
                             |     +--+                                 
                             |        +-- Obovaria_subrotunda           
                             |                                          
                             |        +-- Epioblasma_brevidens          
                             |  +-----+                                 
                             |  |     +-- Epioblasma_triquetra          
                             +--+                                       
                                |  +----- Epioblasma_walkeri[1]         
                                +--+                                    
                                   |  +-- Epioblasma_walkeri[2]         
                                   +--+                                 
                                      +-- Epioblasma_capsaeformis       

Inoue et al., 2018 [Publ 3858], fig. 1, 3a-c

iT tT bi riS oT roT roS iD
157 157 53 9 48 25 4 144.0
                                               +-- Reginaia_ebenus          
  +--------------------------------------------+                            
  |                                            +-- Reginaia_rotulata        
  |                                                                         
  |                                            +-- Amblema_plicata          
  +--------------------------------------------+                            
  |                                            +-- Amblema_elliottii        
  |                                                                         
  |                                         +----- Popenaias_popeii         
  |                                         |                               
  |                                         |  +-- Dromus_dromas            
  |                                         |  |                            
  |                                         |  +-- Epioblasma_rangiana      
  |                                         |  |                            
  +-----------------------------------------+  +-- Glebula_rotundata        
  |                                         |  |                            
  |                                         |  +-- Lampsilis_ovata          
  |                                         |  |                            
  |                                         |  +-- Lampsilis_virescens      
  |                                         |  |                            
  |                                         |  +-- Lemiox_rimosus           
  |                                         +--+                            
  |                                            +-- Obliquaria_reflexa       
  |                                            |                            
  |                                            +-- Obovaria_olivaria        
  |                                            |                            
--+                                            +-- Obovaria_unicolor        
  |                                            |                            
  |                                            +-- Plectomerus_dombeyanus   
  |                                            |                            
  |                                            +-- Potamilus_alatus         
  |                                            |                            
  |                                            +-- Potamilus_capax          
  |                                                                         
  |                                            +-- Uniomerus_declivis       
  |                                         +--+                            
  |                                         |  +-- Uniomerus_tetralasmus    
  |                                         |                               
  |                                         |  +-- Cyclonaias_tuberculata   
  |  +--------------------------------------+  |                            
  |  |                                      |  +-- Pustulosa_pustulosa[1]   
  |  |                                      |  |                            
  |  |                                      |  +-- Theliderma_metanevra     
  |  |                                      +--+                            
  |  |                                         +-- Pustulosa_pustulosa[2]   
  |  |                                         |                            
  |  |                                         +-- Quadrula_quadrula        
  |  |                                         |                            
  |  |                                         +-- Quadrula_verrucosa       
  |  |                                                                      
  |  |  +----------------------------------------- Elliptoideus_sloatianus **  
  +--+  |                                                                   
     |  |                                      +-- Elliptio_lanceolata **      
     |  |  +-----------------------------------+                            
     |  |  |                                   +-- Parvaspina_collina **       
     |  |  |                                                                
     |  |  |  +----------------------------------- Plethobasus_cyphyus **      
     |  |  |  |                                                             
     |  |  |  +----------------------------------- Eurynia_dilatata **         
     |  |  |  |                                                             
     |  |  |  +----------------------------------- Hemistena_lata **           
     |  |  |  |                                                             
     |  |  |  |                                +-- Elliptio_complanata **      
     +--+  |  |                                |                            
        |  |  |                                +-- Elliptio_hopetonensis **    
        |  |  |                                |                            
        |  |  |                                +-- Elliptio_producta[1] **     
        |  |  |                                |                            
        |  |  |                                +-- Elliptio_shepardiana **     
        |  |  |                                |                            
        |  |  |                                +-- Elliptio_angustata **       
        |  |  |                                |                            
        |  |  |                                +-- Elliptio_arca **            
        |  |  |                                |                            
        |  |  |                                +-- Elliptio_arctata **         
        |  |  |                                |                            
        +--+  |                                +-- Elliptio_jayensis **        
           |  |                                |                            
           |  |                                +-- Elliptio_crassidens **      
           |  |                                |                            
           |  +--------------------------------+-- Elliptio_dariensis **       
           |  |                                |                            
           |  |                                +-- Elliptio_fisheriana **      
           |  |                                |                            
           |  |                                +-- Elliptio_folliculata **     
           |  |                                |                            
           |  |                                +-- Elliptio_fumata **          
           |  |                                |                            
           |  |                                +-- Elliptio_icterina **        
           |  |                                |                            
           |  |                                +-- Elliptio_mcmichaeli **      
           |  |                                |                            
           |  |                                +-- Elliptio_producta[2] **     
           |  |                                |                            
           |  |                                +-- Elliptio_pullata **         
           |  |                                |                            
           |  |                                +-- Elliptio_purpurella **      
           |  |                                |                            
           |  |                                +-- Elliptio_congaraea **       
           |  |                                                             
           +--+                                +-- Pleuronaia_barnesiana **    
              |                                |                            
              |  +-----------------------------+-- Pleuronaia_dolabelloides ** 
              |  |                             |                            
              |  |                             +-- Pleuronaia_gibber **        
              |  |                                                          
              |  |     +-------------------------- Pleurobema_plenum **        
              |  |     |                                                    
              |  |     |  +----------------------- Pleurobema_stabile **       
              |  |     |  |                                                 
              |  |     |  |                 +----- Pleurobema_pyriforme **     
              |  |  +--+  |              +--+                               
              |  |  |  |  |              |  |  +-- Pleurobema_georgianum[2] ** 
              |  |  |  |  |              |  +--+                            
              |  |  |  |  |  +-----------+     +-- Pleurobema_athearni **      
              |  |  |  |  |  |           |                                  
              |  |  |  +--+  |           |  +----- Pleurobema_decisum **       
              |  |  |     |  |           |  |                               
              |  |  |     |  |           +--+  +-- Pleurobema_georgianum[1] ** 
              |  |  |     |  |              |  |                            
              |  |  |     |  |              +--+-- Pleurobema_hanleyianum[1] **
              |  |  |     |  |                 |                            
              |  |  |     |  |                 +-- Pleurobema_hanleyianum[2] **
              |  |  |     |  |                                              
              |  |  |     +--+     +-------------- Pleurobema_beadleianum[2] **
              |  |  |        |     |                                        
              |  |  |        |  +--+  +----------- Pleurobema_strodeanum[2] ** 
              |  |  |        |  |  |  |                                     
              |  |  |        |  |  +--+----------- Pleurobema_rubellum **      
              |  |  |        |  |     |                                     
              |  |  |        |  |     |  +-------- Pleurobema_beadleianum[1] **
              |  |  |        |  |     +--+                                  
              |  |  |        |  |        |  +----- Pleurobema_taitianum **     
              |  |  |        |  |        +--+                               
              |  |  |        |  |           |  +-- Pleurobema_perovatum **     
              |  |  |        +--+           +--+                            
              |  |  |           |              +-- Pleurobema_strodeanum[1] ** 
              |  |  |           |                                           
              |  |  |           |           +----- Pleurobema_oviforme[6] **   
              |  |  |           |           |                               
              |  |  |           |        +--+----- Pleurobema_clava[3] **      
              |  |  |           |        |  |                               
              |  |  |           |        |  |  +-- Pleurobema_oviforme[5] **   
              |  |  |           |        |  +--+                            
              |  |  |           +--------+     +-- Pleurobema_clava[2] **      
              +--+  |                    |                                  
                 |  |                    |  +----- Pleurobema_oviforme[2] **   
                 |  |                    |  |                               
                 |  |                    |  +----- Pleurobema_oviforme[3] **   
                 |  |                    +--+                               
                 |  |                       +----- Pleurobema_oviforme[4] **   
                 |  |                       |                               
                 |  |                       |  +-- Pleurobema_clava[1] **      
                 |  |                       +--+                            
                 |  |                          +-- Pleurobema_oviforme[1] **   
                 |  |                                                       
                 |  |  +-------------------------- Pleurobema_cordatum **      
                 |  |  |                                                    
                 |  |  |                    +----- Fusconaia_subrotunda **     
                 |  |  |  +-----------------+                               
                 |  +--+  |                 |  +-- Fusconaia_cor **            
                 |  |  |  |                 +--+                            
                 |  |  |  |                    +-- Fusconaia_cuneolus **       
                 |  |  |  |                                                 
                 |  |  |  |  +-------------------- Fusconaia_ozarkensis **     
                 |  |  +--+  |                                              
                 |  |     |  |                 +-- Fusconaia_escambia **       
                 |  |     |  |  +--------------+                            
                 |  |     |  |  |              +-- Fusconaia_burkei **         
                 |  |     |  |  |                                           
                 |  |     |  |  |           +----- Fusconaia_flava[27] **      
                 |  |     |  |  |           |                               
                 |  |     +--+  |           |  +-- Fusconaia_iheringi **       
                 |  |        |  |           +--+                            
                 |  |        |  |           |  +-- Fusconaia_mitchelli **      
                 |  |        |  +-----------+                               
                 |  |        |  |           |  +-- Fusconaia_askewi[1] **      
                 |  |        |  |           |  |                            
                 |  |        |  |           |  +-- Fusconaia_askewi[2] **      
                 |  |        |  |           |  |                            
                 |  |        +--+           |  +-- Fusconaia_askewi[3] **      
                 |  |           |           +--+                            
                 |  |           |              +-- Fusconaia_askewi[4] **      
                 |  |           |              |                            
                 |  |           |              +-- Fusconaia_askewi[5] **      
                 |  |           |              |                            
                 |  |           |              +-- Fusconaia_chunii **         
                 |  |           |                                           
                 +--+           |  +-------------- Fusconaia_masoni **         
                    |           |  |                                        
                    |           |  |  +----------- Fusconaia_flava[26] **      
                    |           |  |  |                                     
                    |           |  |  |        +-- Fusconaia_cerina[8] **      
                    |           +--+  |  +-----+                            
                    |              |  |  |     +-- Fusconaia_flava[25] **      
                    |              |  |  |                                  
                    |              |  |  |  +----- Fusconaia_flava[5] **       
                    |              |  |  |  |                               
                    |              |  |  |  +----- Fusconaia_flava[6] **       
                    |              +--+  |  |                               
                    |                 |  |  +----- Fusconaia_flava[7] **       
                    |                 |  |  |                               
                    |                 |  |  +----- Fusconaia_flava[8] **       
                    |                 |  |  |                               
                    |                 |  |  +----- Fusconaia_cerina[4] **      
                    |                 |  |  |                               
                    |                 |  |  +----- Fusconaia_flava[9] **       
                    |                 |  |  |                               
                    |                 +--+  +----- Fusconaia_flava[10] **      
                    |                    |  |                               
                    |                    |  +----- Fusconaia_flava[11] **      
                    |                    |  |                               
                    |                    |  +----- Fusconaia_flava[12] **      
                    |                    |  |                               
                    |                    |  +----- Fusconaia_flava[13] **      
                    |                    |  |                               
                    |                    |  +----- Fusconaia_flava[14] **      
                    |                    |  |                               
                    |                    |  +----- Fusconaia_flava[15] **      
                    |                    |  |                               
                    |                    |  +----- Fusconaia_flava[16] **      
                    |                    |  |                               
                    |                    |  +----- Fusconaia_flava[17] **      
                    |                    +--+                               
                    |                       +----- Fusconaia_flava[18] **      
                    |                       |                               
                    |                       +----- Fusconaia_flava[19] **      
                    |                       |                               
                    |                       +----- Fusconaia_flava[20] **      
                    |                       |                               
                    |                       +----- Fusconaia_flava[21] **      
                    |                       |                               
                    |                       +----- Fusconaia_flava[22] **      
                    |                       |                               
                    |                       +----- Fusconaia_flava[23] **      
                    |                       |                               
                    |                       +----- Fusconaia_flava[24] **      
                    |                       |                               
                    |                       +----- Fusconaia_cerina[5] **      
                    |                       |                               
                    |                       +----- Fusconaia_cerina[6] **      
                    |                       |                               
                    |                       +----- Fusconaia_cerina[7] **      
                    |                       |                               
                    |                       |  +-- Fusconaia_flava[1] **       
                    |                       |  |                            
                    |                       |  +-- Fusconaia_flava[2] **       
                    |                       |  |                            
                    |                       |  +-- Fusconaia_flava[3] **       
                    |                       |  |                            
                    |                       +--+-- Fusconaia_flava[4] **       
                    |                          |                            
                    |                          +-- Fusconaia_cerina[1] **      
                    |                          |                            
                    |                          +-- Fusconaia_cerina[2] **      
                    |                          |                            
                    |                          +-- Fusconaia_cerina[3] **      
                    |                                                       
                    |           +----------------- Pleurobema_riddellii **     
                    |           |                                           
                    |           |  +-------------- Pleurobema_sintoxia[21] **  
                    |           |  |                                        
                    |           |  |  +----------- Pleurobema_sintoxia[19] **  
                    |           |  |  |                                     
                    |           |  |  +----------- Pleurobema_sintoxia[20] **  
                    |           |  |  |                                     
                    +-----------+  |  |     +----- Pleurobema_sintoxia[18] **  
                                |  |  |     |                               
                                |  |  |     +----- Pleurobema_rubrum[19] **    
                                |  |  |     |                               
                                |  |  |     +----- Pleurobema_rubrum[20] **    
                                |  |  +-----+                               
                                |  |  |     +----- Pleurobema_rubrum[21] **    
                                |  |  |     |                               
                                |  |  |     |  +-- Pleurobema_rubrum[17] **    
                                +--+  |     |  |                            
                                   |  |     |  +-- Pleurobema_sintoxia[16] **  
                                   |  |     +--+                            
                                   |  |        +-- Pleurobema_sintoxia[17] **  
                                   |  |        |                            
                                   |  |        +-- Pleurobema_rubrum[18] **    
                                   |  |                                     
                                   |  |  +-------- Pleurobema[4] **            
                                   |  |  |                                  
                                   |  |  +-------- Pleurobema[5] **            
                                   |  |  |                                  
                                   |  |  +-------- Pleurobema_sintoxia[1] **   
                                   |  |  |                                  
                                   |  |  +-------- Pleurobema[8] **            
                                   |  |  |                                  
                                   |  |  +-------- Pleurobema[9] **            
                                   +--+  |                                  
                                      |  +-------- Pleurobema[10] **           
                                      |  |                                  
                                      |  +-------- Pleurobema_sintoxia[3] **   
                                      |  |                                  
                                      |  +-------- Pleurobema_sintoxia[4] **   
                                      |  |                                  
                                      |  +-------- Pleurobema_sintoxia[5] **   
                                      |  |                                  
                                      |  +-------- Pleurobema_sintoxia[6] **   
                                      |  |                                  
                                      |  +-------- Pleurobema_sintoxia[7] **   
                                      |  |                                  
                                      |  +-------- Pleurobema_sintoxia[8] **   
                                      |  |                                  
                                      |  +-------- Pleurobema_sintoxia[9] **   
                                      |  |                                  
                                      |  +-------- Pleurobema_sintoxia[10] **  
                                      |  |                                  
                                      |  +-------- Pleurobema_sintoxia[11] **  
                                      |  |                                  
                                      |  +-------- Pleurobema_sintoxia[12] **  
                                      |  |                                  
                                      |  +-------- Pleurobema_sintoxia[13] **  
                                      |  |                                  
                                      |  +-------- Pleurobema_sintoxia[14] **  
                                      |  |                                  
                                      |  +-------- Pleurobema_sintoxia[15] **  
                                      |  |                                  
                                      +--+-------- Pleurobema_rubrum[4] **     
                                         |                                  
                                         +-------- Pleurobema_rubrum[5] **     
                                         |                                  
                                         +-------- Pleurobema_rubrum[6] **     
                                         |                                  
                                         +-------- Pleurobema_rubrum[7] **     
                                         |                                  
                                         +-------- Pleurobema_rubrum[8] **     
                                         |                                  
                                         +-------- Pleurobema_rubrum[9] **     
                                         |                                  
                                         +-------- Pleurobema_rubrum[10] **    
                                         |                                  
                                         +-------- Pleurobema_rubrum[11] **    
                                         |                                  
                                         +-------- Pleurobema_rubrum[12] **    
                                         |                                  
                                         +-------- Pleurobema_rubrum[13] **    
                                         |                                  
                                         +-------- Pleurobema_rubrum[14] **    
                                         |                                  
                                         +-------- Pleurobema_rubrum[15] **    
                                         |                                  
                                         +-------- Pleurobema_rubrum[16] **    
                                         |                                  
                                         |     +-- Pleurobema[6] **            
                                         +-----+                            
                                         |     +-- Pleurobema_rubrum[1] **     
                                         |                                  
                                         |     +-- Pleurobema_sintoxia[2] **   
                                         +-----+                            
                                         |     +-- Pleurobema_rubrum[2] **     
                                         |                                  
                                         |     +-- Pleurobema[7] **            
                                         +-----+                            
                                         |     +-- Pleurobema_rubrum[3] **     
                                         |                                  
                                         |  +----- Pleurobema[3] **            
                                         +--+                               
                                            |  +-- Pleurobema[1] **            
                                            +--+                            
                                               +-- Pleurobema[2] **            

Campbell & Lydeard, 2012 [Publ 3309], fig. 1-2

iT tT bi riS oT roT roS iD
42 42 36 9 32 32 4 51.0
                                         +-- Uniomerus_declivis         
                       +-----------------+                              
                       |                 +-- Uniomerus_tetralasmus      
  +--------------------+                                                
  |                    |  +----------------- Theliderma_metanevra       
  |                    |  |                                             
  |                    +--+  +-------------- Quadrula_verrucosa         
  |                       |  |                                          
  |                       |  |           +-- Quadrula_quadrula          
  |                       +--+  +--------+                              
  |                          |  |        +-- Quadrula_nobilis           
  |                          |  |                                       
  |                          +--+        +-- Pustulosa_infucata         
  |                             |  +-----+                              
  |                             |  |     +-- Pustulosa_kleiniana        
  |                             +--+                                    
  |                                |  +----- Cyclonaias_tuberculata     
  |                                +--+                                 
  |                                   |  +-- Pustulosa_pustulosa        
  |                                   +--+                              
  |                                      +-- Pustulosa_succissa         
  |                                                                     
  |                                   +----- Plectomerus_dombeyanus     
  |                 +-----------------+                                 
  |                 |                 |  +-- Reginaia_ebenus            
  |                 |                 +--+                              
--+                 |                    +-- Reginaia_rotulata          
  |                 |                                                   
  |                 |                 +----- Popenaias_popeii           
  |  +--------------+  +--------------+                                 
  |  |              |  |              |  +-- Amblema_elliottii          
  |  |              |  |              +--+                              
  |  |              |  |                 +-- Amblema_plicata            
  |  |              |  |                                                
  |  |              |  |  +----------------- Potamilus_alatus           
  |  |              +--+  |                                             
  |  |                 |  +----------------- Potamilus_capax            
  |  |                 |  |                                             
  |  |                 |  +----------------- Glebula_rotundata          
  |  |                 |  |                                             
  |  |                 |  |              +-- Toxolasma_lividum          
  |  |                 |  +--------------+                              
  |  |                 +--+              +-- Paetulunio_fabalis         
  |  |                    |                                             
  |  |                    |              +-- Dromus_dromas              
  |  |                    +--------------+                              
  |  |                    |              +-- Lemiox_rimosus             
  |  |                    |                                             
  |  |                    |              +-- Lampsilis_ovata            
  +--+                    |  +-----------+                              
     |                    |  |           +-- Obliquaria_reflexa         
     |                    +--+                                          
     |                       |  +----------- Lampsilis_virescens        
     |                       |  |                                       
     |                       +--+        +-- Epioblasma_rangiana        
     |                          |  +-----+                              
     |                          |  |     +-- Obovaria_arkansasensis     
     |                          +--+                                    
     |                             |  +----- Obovaria_olivaria          
     |                             +--+                                 
     |                                |  +-- Venustaconcha_ellipsiformis
     |                                +--+                              
     |                                   +-- Venustaconcha_pleasii      
     |                                                                  
     |  +----------------------------------- Elliptoideus_sloatianus **    
     |  |                                                               
     |  |  +-------------------------------- Pleurobema_stabile **         
     |  |  |                                                            
     |  |  |                             +-- Pleurobema_clava **           
     |  |  |                 +-----------+                              
     +--+  |                 |           +-- Pleurobema_oviforme **        
        |  |                 |                                          
        |  |  +--------------+  +----------- Pleurobema_pyriforme **       
        |  |  |              |  |                                       
        |  |  |              |  |        +-- Pleurobema_athearni **        
        |  |  |              |  |     +--+                              
        +--+  |              +--+     |  +-- Pleurobema_georgianum **      
           |  |                 |  +--+                                 
           |  |                 |  |  |  +-- Pleurobema_decisum **         
           |  |                 |  |  +--+                              
           |  |                 |  |     +-- Pleurobema_hanleyianum **     
           |  |                 +--+                                    
           |  |                    |  +----- Pleurobema_beadleianum **     
           |  |                    |  |                                 
           |  |                    |  +----- Pleurobema_strodeanum **      
           |  |                    +--+                                 
           |  |                       +----- Pleurobema_rubellum **        
           +--+                       |                                 
              |                       |  +-- Pleurobema_perovatum **       
              |                       +--+                              
              |                          +-- Pleurobema_taitianum **       
              |                                                         
              |                    +-------- Pleurobema_riddellii **       
              |  +-----------------+                                    
              |  |                 |  +----- Pleurobema_cordatum **        
              |  |                 +--+                                 
              |  |                    |  +-- Pleurobema_rubrum **          
              |  |                    +--+                              
              |  |                       +-- Pleurobema_sintoxia **        
              |  |                                                      
              |  |        +----------------- Fusconaia_subrotunda **       
              |  |        |                                             
              +--+  +-----+              +-- Fusconaia_cor **              
                 |  |     |  +-----------+                              
                 |  |     |  |           +-- Fusconaia_cuneolus **         
                 |  |     +--+                                          
                 |  |        |  +----------- Fusconaia_ozarkensis **       
                 |  |        |  |                                       
                 |  |        +--+        +-- Fusconaia_escambia **         
                 |  |           |  +-----+                              
                 |  |           |  |     +-- Fusconaia_burkei **           
                 |  |           +--+                                    
                 +--+              |  +----- Fusconaia_masoni **           
                    |              +--+                                 
                    |                 |  +-- Fusconaia_flava **            
                    |                 +--+                              
                    |                    +-- Fusconaia_cerina **           
                    |                                                   
                    |                 +----- Pleuronaia_gibber **          
                    |  +--------------+                                 
                    |  |              |  +-- Pleuronaia_barnesiana **      
                    |  |              +--+                              
                    |  |                 +-- Pleuronaia_dolabelloides **   
                    +--+                                                
                       |                 +-- Eurynia_dilatata **           
                       |  +--------------+                              
                       |  |              +-- Parvaspina_collina **         
                       |  |                                             
                       +--+              +-- Hemistena_lata **             
                          |  +-----------+                              
                          |  |           +-- Plethobasus_cyphyus **        
                          |  |                                          
                          +--+  +----------- Elliptio_shepardiana **       
                             |  |                                       
                             |  |  +-------- Elliptio_complanata **        
                             +--+  |                                    
                                |  |  +----- Elliptio_purpurella **        
                                |  +--+                                 
                                |  |  |  +-- Elliptio_arca **              
                                +--+  +--+                              
                                   |     +-- Elliptio_arctata **           
                                   |                                    
                                   |     +-- Elliptio_mcmichaeli **        
                                   |     |                              
                                   +-----+-- Elliptio_pullata **           
                                         |                              
                                         +-- Elliptio_crassidens **        

Campbell & Lydeard, 2012 [Publ 3309], fig. 3

iT tT bi riS oT roT roS iD
42 42 37 9 32 32 4 49.0
                                      +-- Uniomerus_declivis         
                       +--------------+                              
                       |              +-- Uniomerus_tetralasmus      
                       |                                             
  +--------------------+              +-- Quadrula_verrucosa         
  |                    |  +-----------+                              
  |                    |  |           +-- Quadrula_nobilis           
  |                    |  |                                          
  |                    +--+  +----------- Quadrula_quadrula          
  |                       |  |                                       
  |                       |  +----------- Theliderma_metanevra       
  |                       +--+                                       
  |                          |        +-- Pustulosa_infucata         
  |                          |  +-----+                              
  |                          |  |     +-- Pustulosa_kleiniana        
  |                          +--+                                    
  |                             |  +----- Cyclonaias_tuberculata     
  |                             +--+                                 
  |                                |  +-- Pustulosa_pustulosa        
  |                                +--+                              
  |                                   +-- Pustulosa_succissa         
  |                                                                  
  |                                   +-- Amblema_elliottii          
  |                                +--+                              
  |                                |  +-- Amblema_plicata            
  |     +--------------------------+                                 
  |     |                          |  +-- Reginaia_ebenus            
--+     |                          +--+                              
  |     |                             +-- Reginaia_rotulata          
  |  +--+                                                            
  |  |  |  +----------------------------- Popenaias_popeii           
  |  |  |  |                                                         
  |  |  |  |                          +-- Glebula_rotundata          
  |  |  +--+  +-----------------------+                              
  |  |     |  |                       +-- Paetulunio_fabalis         
  |  |     |  |                                                      
  |  |     +--+                       +-- Toxolasma_lividum          
  |  |        |  +--------------------+                              
  |  |        |  |                    +-- Plectomerus_dombeyanus     
  |  |        |  |                                                   
  |  |        +--+  +-------------------- Obliquaria_reflexa         
  |  |           |  |                                                
  |  |           |  |                 +-- Dromus_dromas              
  |  |           |  |              +--+                              
  |  |           +--+              |  +-- Lemiox_rimosus             
  |  |              |  +-----------+                                 
  |  |              |  |           |  +-- Potamilus_alatus           
  |  |              |  |           +--+                              
  |  |              +--+              +-- Potamilus_capax            
  |  |                 |                                             
  |  |                 |  +-------------- Lampsilis_ovata            
  +--+                 |  |                                          
     |                 +--+  +----------- Lampsilis_virescens        
     |                    |  |                                       
     |                    +--+        +-- Epioblasma_rangiana        
     |                       |  +-----+                              
     |                       |  |     +-- Obovaria_arkansasensis     
     |                       +--+                                    
     |                          |  +----- Obovaria_olivaria          
     |                          +--+                                 
     |                             |  +-- Venustaconcha_ellipsiformis
     |                             +--+                              
     |                                +-- Venustaconcha_pleasii      
     |                                                               
     |  +-------------------------------- Elliptoideus_sloatianus **    
     |  |                                                            
     |  |                             +-- Eurynia_dilatata **           
     |  |  +--------------------------+                              
     |  |  |                          +-- Parvaspina_collina **         
     |  |  |                                                         
     |  |  |                          +-- Hemistena_lata **             
     |  |  +--------------------------+                              
     |  |  |                          +-- Plethobasus_cyphyus **        
     +--+  |                                                         
        |  |                 +----------- Elliptio_shepardiana **       
        |  |                 |                                       
        |  |                 |     +----- Elliptio_crassidens **        
        |  +-----------------+  +--+                                 
        |  |                 |  |  |  +-- Elliptio_mcmichaeli **        
        |  |                 |  |  +--+                              
        |  |                 +--+     +-- Elliptio_pullata **           
        |  |                    |                                    
        |  |                    |  +----- Elliptio_complanata **        
        +--+                    |  |                                 
           |                    +--+  +-- Elliptio_purpurella **        
           |                       |  |                              
           |                       +--+-- Elliptio_arca **              
           |                          |                              
           |                          +-- Elliptio_arctata **           
           |                                                         
           |                       +----- Pleuronaia_gibber **          
           |  +--------------------+                                 
           |  |                    |  +-- Pleuronaia_barnesiana **      
           |  |                    +--+                              
           |  |                       +-- Pleuronaia_dolabelloides **   
           |  |                                                      
           |  |           +-------------- Fusconaia_ozarkensis **       
           |  |           |                                          
           |  |           |        +----- Fusconaia_subrotunda **       
           |  |  +--------+  +-----+                                 
           +--+  |        |  |     |  +-- Fusconaia_cor **              
              |  |        |  |     +--+                              
              |  |        |  |        +-- Fusconaia_cuneolus **         
              |  |        +--+                                       
              |  |           |        +-- Fusconaia_escambia **         
              |  |           |  +-----+                              
              |  |           |  |     +-- Fusconaia_burkei **           
              |  |           +--+                                    
              |  |              |  +----- Fusconaia_masoni **           
              +--+              +--+                                 
                 |                 |  +-- Fusconaia_cerina **           
                 |                 +--+                              
                 |                    +-- Fusconaia_flava **            
                 |                                                   
                 |              +-------- Pleurobema_riddellii **       
                 |  +-----------+                                    
                 |  |           |  +----- Pleurobema_cordatum **        
                 |  |           +--+                                 
                 |  |              |  +-- Pleurobema_rubrum **          
                 |  |              +--+                              
                 +--+                 +-- Pleurobema_sintoxia **        
                    |                                                
                    |  +----------------- Pleurobema_stabile **         
                    |  |                                             
                    |  |     +----------- Pleurobema_strodeanum **      
                    |  |  +--+                                       
                    +--+  |  |  +-------- Pleurobema_rubellum **        
                       |  |  +--+                                    
                       |  |     |  +----- Pleurobema_beadleianum **     
                       |  |     +--+                                 
                       |  |        |  +-- Pleurobema_perovatum **       
                       +--+        +--+                              
                          |           +-- Pleurobema_taitianum **       
                          |                                          
                          |           +-- Pleurobema_clava **           
                          |  +--------+                              
                          |  |        +-- Pleurobema_oviforme **        
                          +--+                                       
                             |  +-------- Pleurobema_pyriforme **       
                             |  |                                    
                             +--+     +-- Pleurobema_athearni **        
                                |  +--+                              
                                |  |  +-- Pleurobema_georgianum **      
                                +--+                                 
                                   |  +-- Pleurobema_decisum **         
                                   +--+                              
                                      +-- Pleurobema_hanleyianum **     

Burlakova et al., 2012 [Publ 3486], fig. 3

iT tT bi riS oT roT roS iD
42 42 25 8 1 1 1 44.0
  +----------------------------------- Plectomerus_dombeyanus  
  |                                                            
  |  +-------------------------------- Hemistena_lata **          
  |  |                                                         
--+  +-------------------------------- Pleurobema_stabile **      
  |  |                                                         
  |  |  +----------------------------- Eurynia_dilatata **        
  |  |  |                                                      
  +--+  |                    +-------- Pleurobema_riddellii **    
     |  |                    |                                 
     |  |  +-----------------+  +----- Pleurobema_sintoxia **     
     |  |  |                 |  |                              
     |  |  |                 +--+----- Pleurobema_rubrum[2] **    
     |  |  |                    |                              
     +--+  |                    |  +-- Pleurobema_cordatum **     
        |  |                    +--+                           
        |  |                       +-- Pleurobema_rubrum[1] **    
        |  |                                                   
        |  |  +----------------------- Parvaspina_collina **      
        |  |  |                                                
        |  |  |                 +----- Pleuronaia_gibber **       
        |  |  +-----------------+                              
        |  |  |                 |  +-- Pleuronaia_dolabelloides **
        +--+  |                 +--+                           
           |  |                    +-- Pleuronaia_barnesiana **   
           |  |                                                
           |  |                 +----- Plethobasus_cyphyus **     
           |  +-----------------+                              
           |  |                 |  +-- Elliptio_arca **           
           |  |                 +--+                           
           |  |                    +-- Elliptio_crassidens **     
           |  |                                                
           |  |                    +-- Pleurobema_clava **        
           |  |           +--------+                           
           |  |           |        +-- Pleurobema_oviforme **     
           |  |           |                                    
           |  |           |  +-------- Pleurobema_decisum **      
           +--+-----------+  |                                 
              |           |  +-------- Pleurobema_pyriforme **    
              |           |  |                                 
              |           |  +-------- Pleurobema_hanleyianum **  
              |           +--+                                 
              |              +-------- Pleurobema_georgianum **   
              |              |                                 
              |              |  +----- Pleurobema_rubellum **     
              |              |  |                              
              |              |  +----- Pleurobema_beadleianum **  
              |              +--+                              
              |                 +----- Pleurobema_strodeanum **   
              |                 |                              
              |                 |  +-- Pleurobema_perovatum **    
              |                 +--+                           
              |                    +-- Pleurobema_taitianum **    
              |                                                
              |  +-------------------- Fusconaia_subrotunda **    
              |  |                                             
              |  |                 +-- Fusconaia_cor **           
              +--+  +--------------+                           
                 |  |              +-- Fusconaia_cuneolus **      
                 |  |                                          
                 |  |  +-------------- Fusconaia_ozarkensis **    
                 +--+  |                                       
                    |  |     +-------- Fusconaia_masoni **        
                    |  |     |                                 
                    |  |     |     +-- Fusconaia_burkei **        
                    |  |  +--+-----+                           
                    +--+  |  |     +-- Fusconaia_escambia **      
                       |  |  |                                 
                       |  |  |  +----- Fusconaia_cerina **        
                       |  |  +--+                              
                       |  |     |  +-- Fusconaia_askewi[1] **     
                       |  |     +--+                           
                       +--+        +-- Fusconaia_flava **         
                          |                                    
                          |        +-- Fusconaia_askewi[2] **     
                          |        |                           
                          |        +-- Fusconaia_askewi[3] **     
                          |        |                           
                          |        +-- Fusconaia_askewi[4] **     
                          +--------+                           
                                   +-- Fusconaia_lananensis[1] ** 
                                   |                           
                                   +-- Fusconaia_lananensis[2] ** 
                                   |                           
                                   +-- Fusconaia_askewi[5] **     

Burlakova et al., 2012 [Publ 3486], fig. 2

iT tT bi riS oT roT roS iD
40 40 30 8 0 0 0 46.0
  +----------------------------------- Parvaspina_collina **      
  |                                                            
  |  +-------------------------------- Pleurobema_plenum **       
  |  |                                                         
--+  |                             +-- Plethobasus_cyphyus **     
  |  |                       +-----+                           
  |  |                       |     +-- Hemistena_lata **          
  |  |  +--------------------+                                 
  +--+  |                    |  +----- Eurynia_dilatata **        
     |  |                    +--+                              
     |  |                       |  +-- Elliptio_arca **           
     |  |                       +--+                           
     |  |                          +-- Elliptio_crassidens **     
     |  |                                                      
     |  |  +-------------------------- Pleurobema_cordatum[2] **  
     +--+  |                                                   
        |  |           +-------------- Fusconaia_subrotunda **    
        |  |           |                                       
        |  |           |           +-- Fusconaia_cor **           
        |  |           |  +--------+                           
        |  |  +--------+  |        +-- Fusconaia_cuneolus **      
        |  |  |        |  |                                    
        |  |  |        |  |  +-------- Fusconaia_flava[2] **      
        |  |  |        |  |  |                                 
        +--+  |        +--+  +-------- Fusconaia_ozarkensis **    
           |  |           |  |                                 
           |  |           |  +-------- Fusconaia_masoni **        
           |  |           |  |                                 
           |  |           |  |  +----- Fusconaia_askewi **        
           |  |           +--+--+                              
           |  |              |  |  +-- Fusconaia_burkei **        
           |  |              |  +--+                           
           |  |              |     +-- Fusconaia_escambia **      
           |  |              |                                 
           +--+              |     +-- Fusconaia_flava[1] **      
              |              |     |                           
              |              +-----+-- Fusconaia_cerina[1] **     
              |                    |                           
              |                    +-- Fusconaia_cerina[2] **     
              |                                                
              |              +-------- Pleurobema_stabile **      
              |              |                                 
              |  +-----------+  +----- Pleurobema_riddellii **    
              |  |           |  |                              
              |  |           +--+  +-- Pleurobema_cordatum[1] **  
              |  |              |  |                           
              |  |              |  +-- Pleurobema_rubrum **       
              |  |              +--+                           
              |  |                 +-- Pleurobema_sintoxia[1] **  
              +--+                 |                           
                 |                 +-- Pleurobema_sintoxia[2] **  
                 |                                             
                 |              +----- Pleuronaia_barnesiana **   
                 |  +-----------+                              
                 |  |           |  +-- Pleuronaia_dolabelloides **
                 |  |           +--+                           
                 |  |              +-- Pleuronaia_gibber **       
                 +--+                                          
                    |              +-- Pleurobema_georgianum **   
                    |  +-----------+                           
                    |  |           +-- Pleurobema_pyriforme **    
                    |  |                                       
                    +--+           +-- Pleurobema_decisum **      
                       |  +--------+                           
                       |  |        +-- Pleurobema_hanleyianum **  
                       |  |                                    
                       +--+        +-- Pleurobema_clava **        
                          |  +-----+                           
                          |  |     +-- Pleurobema_oviforme **     
                          |  |                                 
                          +--+  +----- Pleurobema_rubellum **     
                             |  |                              
                             |  +----- Pleurobema_beadleianum **  
                             +--+                              
                                +----- Pleurobema_strodeanum **   
                                |                              
                                |  +-- Pleurobema_perovatum **    
                                +--+                           
                                   +-- Pleurobema_taitianum **    

Campbell et al., 2005 [Publ 2660], fig. 1

iT tT bi riS oT roT roS iD
39 39 29 9 78 66 4 44.0
                                            +-- Megalonaias_nervosa           
                                +-----------+                                 
                                |           +-- Uniomerus_declivis            
                                |                                             
  +-----------------------------+  +----------- Theliderma_metanevra          
  |                             |  |                                          
  |                             |  |     +----- Quadrula_verrucosa            
  |                             +--+  +--+                                    
  |                                |  |  |  +-- Quadrula_quadrula             
  |                                |  |  +--+                                 
  |                                |  |     +-- Quadrula_nobilis              
  |                                +--+                                       
  |                                   |  +----- Pustulosa_kieneriana          
  |                                   |  |                                    
  |                                   |  |  +-- Cyclonaias_tuberculata        
  |                                   +--+--+                                 
  |                                      |  +-- Pustulosa_succissa            
  |                                      |                                    
  |                                      |  +-- Pustulosa_infucata            
  |                                      +--+                                 
  |                                         +-- Pustulosa_kleiniana           
  |                                                                           
  |        +----------------------------------- Elliptoideus_sloatianus **       
  |        |                                                                  
  |        |  +-------------------------------- Hemistena_lata **                
  |        |  |                                                               
  |  +-----+  |                          +----- Plethobasus_cyphyus **           
  |  |     |  |  +-----------------------+                                    
  |  |     |  |  |                       |  +-- Elliptio_arca **                 
  |  |     |  |  |                       +--+                                 
  |  |     |  |  |                          +-- Elliptio_crassidens **           
  |  |     +--+  |                                                            
  |  |        |  |                          +-- Eurynia_dilatata **              
  |  |        |  |  +-----------------------+                                 
  |  |        |  |  |                       +-- Parvaspina_collina **            
--+  |        |  |  |                                                         
  |  |        |  |  |                    +----- Pleuronaia_gibber **             
  |  |        |  |  +--------------------+                                    
  |  |        |  |  |                    |  +-- Pleuronaia_barnesiana **         
  |  |        +--+  |                    +--+                                 
  |  |           |  |                       +-- Pleuronaia_dolabelloides **      
  |  |           |  |                                                         
  |  |           |  |              +----------- Fusconaia_subrotunda **          
  |  |           |  |              |                                          
  |  |           |  |              |        +-- Fusconaia_cor **                 
  |  |           |  +--------------+  +-----+                                 
  |  |           |  |              |  |     +-- Fusconaia_cuneolus **            
  |  |           |  |              |  |                                       
  |  |           |  |              |  |     +-- Fusconaia_cerina[1] **           
  |  |           |  |              +--+     |                                 
  |  |           |  |                 |  +--+-- Fusconaia_cerina[2] **           
  |  |           +--+                 |  |  |                                 
  |  |              |                 |  |  +-- Fusconaia_flava **               
  |  |              |                 +--+                                    
  |  |              |                    |  +-- Fusconaia_burkei[1] **           
  |  |              |                    |  |                                 
  |  |              |                    |  +-- Fusconaia_burkei[2] **           
  |  |              |                    +--+                                 
  |  |              |                       +-- Fusconaia_burkei[3] **           
  |  |              |                       |                                 
  |  |              |                       +-- Fusconaia_escambia **            
  |  |              |                                                         
  |  |              |                       +-- Pleurobema_rubrum[2] **          
  |  |              |                    +--+                                 
  |  |              |                    |  +-- Pleurobema_cordatum **           
  |  |              |  +-----------------+                                    
  |  |              |  |                 |  +-- Pleurobema_rubrum[1] **          
  +--+              |  |                 +--+                                 
     |              |  |                    +-- Pleurobema_sintoxia **           
     |              +--+                                                      
     |                 |  +-------------------- Pleurobema_pyriforme **          
     |                 |  |                                                   
     |                 |  |                 +-- Pleurobema_georgianum[2] **      
     |                 |  |                 |                                 
     |                 +--+  +--------------+-- Pleurobema_georgianum[3] **      
     |                    |  |              |                                 
     |                    |  |              +-- Pleurobema_athearni **           
     |                    |  |                                                
     |                    |  |           +----- Pleurobema_decisum **            
     |                    +--+           |                                    
     |                       |  +--------+  +-- Pleurobema_hanleyianum[1] **     
     |                       |  |        |  |                                 
     |                       |  |        +--+-- Pleurobema_hanleyianum[2] **     
     |                       |  |           |                                 
     |                       |  |           +-- Pleurobema_georgianum[1] **      
     |                       +--+                                             
     |                          |           +-- Pleurobema_oviforme[1] **        
     |                          |           |                                 
     |                          |  +--------+-- Pleurobema_oviforme[2] **        
     |                          |  |        |                                 
     |                          +--+        +-- Pleurobema_clava **              
     |                             |                                          
     |                             |  +-------- Pleurobema_strodeanum **         
     |                             +--+                                       
     |                                |  +----- Pleurobema_rubellum **           
     |                                +--+                                    
     |                                   |  +-- Pleurobema_perovatum **          
     |                                   +--+                                 
     |                                      +-- Pleurobema_taitianum **          
     |                                                                        
     |                                   +----- Plectomerus_dombeyanus        
     |  +--------------------------------+                                    
     |  |                                |  +-- Reginaia_ebenus               
     |  |                                +--+                                 
     |  |                                   +-- Reginaia_rotulata             
     |  |                                                                     
     |  |                                +----- Popenaias_popeii              
     |  |  +-----------------------------+                                    
     |  |  |                             |  +-- Amblema_elliottii             
     +--+  |                             +--+                                 
        |  |                                +-- Amblema_plicata               
        |  |                                                                  
        |  |                                +-- Toxolasma_parvum              
        |  |                          +-----+                                 
        |  |                          |     +-- Toxolasma_texasiense          
        |  |  +-----------------------+                                       
        |  |  |                       |  +----- Obliquaria_reflexa            
        +--+  |                       +--+                                    
           |  |                          |  +-- Cyrtonaias_tampicoensis       
           |  |                          +--+                                 
           |  |                             +-- Glebula_rotundata             
           |  |                                                               
           |  |                             +-- Potamilus_fragilis            
           |  |                          +--+                                 
           |  |                          |  +-- Potamilus_leptodon            
           |  |              +-----------+                                    
           |  |              |           |  +-- Potamilus_alatus              
           |  |              |           +--+                                 
           +--+              |              +-- Potamilus_purpuratus          
              |              |                                                
              |  +-----------+              +-- Sagittunio_nasutus            
              |  |           |           +--+                                 
              |  |           |           |  +-- Medionidus_conradicus         
              |  |           |  +--------+                                    
              |  |           |  |        |  +-- Ptychobranchus_fasciolaris    
              |  |           |  |        +--+                                 
              |  |           +--+           +-- Venustaconcha_ellipsiformis[2]
              |  |              |                                             
              |  |              |  +----------- Medionidus_acutissimus        
              |  |              +--+                                          
              |  |                 |  +-------- Lemiox_rimosus                
              |  |                 +--+                                       
              +--+                    |  +----- Dromus_dromas                 
                 |                    +--+                                    
                 |                       |  +-- Cyprogenia_aberti             
                 |                       +--+                                 
                 |                          +-- Cyprogenia_stegaria           
                 |                                                            
                 |                          +-- Ellipsaria_lineolata          
                 |  +-----------------------+                                 
                 |  |                       +-- Truncilla_truncata            
                 |  |                                                         
                 |  |  +----------------------- Villosa_villosa               
                 |  |  |                                                      
                 |  |  |                    +-- Obovaria_olivaria             
                 +--+  |     +--------------+                                 
                    |  |     |              +-- Venustaconcha_ellipsiformis[1]
                    |  |     |                                                
                    |  |  +--+           +----- Obovaria_unicolor             
                    |  |  |  |  +--------+                                    
                    |  |  |  |  |        |  +-- Obovaria_arkansasensis        
                    +--+  |  |  |        +--+                                 
                       |  |  +--+           +-- Obovaria_subrotunda           
                       |  |     |                                             
                       |  |     |  +----------- Epioblasma_brevidens          
                       |  |     +--+                                          
                       |  |        |  +-------- Epioblasma_triquetra          
                       |  |        +--+                                       
                       |  |           |  +----- Epioblasma_walkeri[2]         
                       |  |           +--+                                    
                       |  |              |  +-- Epioblasma_capsaeformis       
                       +--+              +--+                                 
                          |                 +-- Epioblasma_walkeri[1]         
                          |                                                   
                          |                 +-- Ligumia_recta                 
                          |        +--------+                                 
                          |        |        +-- Cambarunio_iris               
                          |     +--+                                          
                          |     |  |  +-------- Lampsilis_teres               
                          |     |  +--+                                       
                          |     |     |  +----- Ortmanniana_ligamentina       
                          |     |     +--+                                    
                          |     |        |  +-- Lampsilis_siliquoidea         
                          |     |        +--+                                 
                          |     |           +-- Leaunio_vanuxemensis          
                          +-----+                                             
                                |           +-- Lampsilis_cardium             
                                |        +--+                                 
                                |        |  +-- Lampsilis_ovata               
                                |  +-----+                                    
                                |  |     |  +-- Ortmanniana_pectorosa         
                                |  |     +--+                                 
                                |  |        +-- Lampsilis_ornata              
                                |  |                                          
                                +--+     +----- Hamiota_australis[2]          
                                   |  +--+                                    
                                   |  |  |  +-- Hamiota_australis[1]          
                                   |  |  +--+                                 
                                   |  |     +-- Hamiota_subangulata           
                                   +--+                                       
                                      |  +----- Hamiota_altilis[2]            
                                      |  |                                    
                                      |  |  +-- Hamiota_altilis[1]            
                                      +--+--+                                 
                                         |  +-- Hamiota_perovalis[1]          
                                         |                                    
                                         |  +-- Hamiota_altilis[3]            
                                         +--+                                 
                                            +-- Hamiota_perovalis[2]          

Burlakova et al., 2012 [Publ 3486], fig. 1

iT tT bi riS oT roT roS iD
38 38 36 8 6 6 3 50.0
  +----------------------------------------- Popenaias_popeii        
  |                                                                  
  |  +-------------------------------------- Cyrtonaias_tampicoensis 
--+  |                                                               
  |  |  +----------------------------------- Parvaspina_collina **      
  |  |  |                                                            
  +--+  |                             +----- Plethobasus_cyphyus **     
     |  |  +--------------------------+                              
     |  |  |                          |  +-- Elliptio_arca **           
     |  |  |                          +--+                           
     |  |  |                             +-- Elliptio_crassidens **     
     +--+  |                                                         
        |  |           +-------------------- Fusconaia_subrotunda **    
        |  |           |                                             
        |  |           |                 +-- Fusconaia_cor **           
        |  |  +--------+  +--------------+                           
        |  |  |        |  |              +-- Fusconaia_cuneolus **      
        |  |  |        |  |                                          
        +--+  |        +--+  +-------------- Fusconaia_ozarkensis **    
           |  |           |  |                                       
           |  |           |  |     +-------- Fusconaia_askewi[3] **     
           |  |           |  |  +--+                                 
           |  |           +--+  |  |  +----- Fusconaia_askewi[2] **     
           |  |              |  |  +--+                              
           |  |              |  |     |  +-- Fusconaia_lananensis **    
           |  |              |  |     +--+                           
           |  |              +--+        +-- Fusconaia_askewi[1] **     
           |  |                 |                                    
           |  |                 |        +-- Fusconaia_burkei **        
           |  |                 |  +-----+                           
           +--+                 |  |     +-- Fusconaia_escambia **      
              |                 +--+                                 
              |                    |  +----- Fusconaia_masoni **        
              |                    +--+                              
              |                       |  +-- Fusconaia_flava **         
              |                       +--+                           
              |                          +-- Fusconaia_cerina **        
              |                                                      
              |                          +-- Eurynia_dilatata **        
              |  +-----------------------+                           
              |  |                       +-- Hemistena_lata **          
              |  |                                                   
              |  |                    +----- Pleuronaia_gibber **       
              |  |              +-----+                              
              |  |              |     |  +-- Pleuronaia_dolabelloides **
              +--+              |     +--+                           
                 |  +-----------+        +-- Pleuronaia_barnesiana **   
                 |  |           |                                    
                 |  |           |  +-------- Pleurobema_riddellii **    
                 |  |           +--+                                 
                 |  |              |  +----- Pleurobema_sintoxia **     
                 |  |              +--+                              
                 +--+                 |  +-- Pleurobema_rubrum **       
                    |                 +--+                           
                    |                    +-- Pleurobema_cordatum **     
                    |                                                
                    |  +-------------------- Pleurobema_stabile **      
                    |  |                                             
                    +--+  +----------------- Pleurobema_pyriforme **    
                       |  |                                          
                       |  |           +----- Pleurobema_georgianum **   
                       +--+  +--------+                              
                          |  |        |  +-- Pleurobema_hanleyianum **  
                          |  |        +--+                           
                          |  |           +-- Pleurobema_decisum **      
                          +--+                                       
                             |           +-- Pleurobema_clava **        
                             |  +--------+                           
                             |  |        +-- Pleurobema_oviforme **     
                             +--+                                    
                                |  +-------- Pleurobema_beadleianum **  
                                |  |                                 
                                +--+     +-- Pleurobema_rubellum **     
                                   |  +--+                           
                                   |  |  +-- Pleurobema_strodeanum **   
                                   +--+                              
                                      |  +-- Pleurobema_perovatum **    
                                      +--+                           
                                         +-- Pleurobema_taitianum **    

Burdick & White, 2007 [Publ 2679], fig. 3

iT tT bi riS oT roT roS iD
36 37 30 7 2 2 2 46.0
  +----------------------------------------- Lampsilis_ornata      
  |                                                                
--+  +-------------------------------------- Parvaspina_collina **    
  |  |                                                             
  |  |  +----------------------------------- Reginaia_ebenus       
  +--+  |                                                          
     |  |                             +----- Eurynia_dilatata **      
     |  |  +--------------------------+                            
     |  |  |                          |  +-- Plethobasus_cyphyus **   
     +--+  |                          +--+                         
        |  |                             +-- Elliptio_crassidens **   
        |  |                                                       
        |  |  +----------------------------- Pleurobema_plenum **     
        |  |  |                                                    
        |  |  |                          +-- Pleuronaia_barnesiana ** 
        +--+  +--------------------------+                         
           |  |                          +-- Pleuronaia_gibber **     
           |  |                                                    
           |  |              +-------------- Pleurobema_stabile **    
           |  |              |                                     
           |  |              |           +-- Pleurobema_rubellum **   
           |  |              |     +-----+                         
           |  |              |     |     +-- Pleurobema_strodeanum ** 
           +--+  +-----------+  +--+                               
              |  |           |  |  |  +----- Pleurobema_beadleianum **
              |  |           |  |  +--+                            
              |  |           |  |     |  +-- Pleurobema_perovatum **  
              |  |           |  |     +--+                         
              |  |           +--+        +-- Pleurobema_taitianum **  
              |  |              |                                  
              |  |              |        +-- Pleurobema_clava **      
              |  |              |  +-----+                         
              |  |              |  |     +-- Pleurobema_oviforme **   
              |  |              |  |                               
              |  |              +--+     +-- Pleurobema_decisum **    
              +--+                 |  +--+                         
                 |                 |  |  +-- Pleurobema_hanleyianum **
                 |                 +--+                            
                 |                    |  +-- Pleurobema_pyriforme **  
                 |                    +--+                         
                 |                       +-- Pleurobema_georgianum ** 
                 |                                                 
                 |                    +----- Pleurobema_sintoxia[1] **
                 |                    |                            
                 |  +-----------------+  +-- Pleurobema_cordatum[2] **
                 |  |                 |  |                         
                 |  |                 +--+-- Pleurobema_sintoxia[2] **
                 |  |                    |                         
                 +--+                    +-- Pleurobema_rubrum **     
                    |                                              
                    |  +-------------------- Pleurobema_cordatum[1] **
                    |  |                                           
                    +--+  +----------------- Fusconaia_subrotunda **  
                       |  |                                        
                       |  |              +-- Fusconaia_cor **         
                       +--+  +-----------+                         
                          |  |           +-- Fusconaia_cuneolus **    
                          |  |                                     
                          +--+  +----------- Fusconaia_ozarkensis **  
                             |  |                                  
                             |  |        +-- Fusconaia_masoni **      
                             +--+  +-----+                         
                                |  |     +-- Fusconaia_flava[3] **    
                                |  |                               
                                +--+  +----- Fusconaia_flava[1] **    
                                   |  |                            
                                   |  +----- Fusconaia_cerina[4] **   
                                   +--+                            
                                      |  +-- Fusconaia_cerina[1] **   
                                      |  |                         
                                      |  +-- Fusconaia_cerina[2] **   
                                      +--+                         
                                         +-- Fusconaia_cerina[3] **   
                                         |                         
                                         +-- Fusconaia_flava[2] **    

Perkins et al., 2017 [Publ 3777], fig. 2

iT tT bi riS oT roT roS iD
35 35 33 7 2 2 1 57.0
  +----------------------------------------- Amblema_plicata         
  |                                                                  
  +----------------------------------------- Plectomerus_dombeyanus  
  |                                                                  
  |                                      +-- Pleurobema_clava **        
  |                          +-----------+                           
  |                          |           +-- Pleurobema_oviforme **     
  |                          |                                       
  |                          |     +-------- Pleurobema_pyriforme **    
  |  +-----------------------+  +--+                                 
--+  |                       |  |  |  +----- Pleurobema_georgianum **   
  |  |                       |  |  +--+                              
  |  |                       |  |     |  +-- Pleurobema_hanleyianum **  
  |  |                       +--+     +--+                           
  |  |                          |        +-- Pleurobema_decisum **      
  |  |                          |                                    
  |  |                          |  +-------- Pleurobema_rubellum **     
  |  |                          +--+                                 
  |  |                             |  +----- Pleurobema_perovatum **    
  |  |                             +--+                              
  +--+                                |  +-- Pleurobema_beadleianum **  
     |                                +--+                           
     |                                   +-- Pleurobema_strodeanum **   
     |                                                               
     |                                +----- Pleuronaia_gibber **       
     |  +-----------------------------+                              
     |  |                             |  +-- Pleuronaia_barnesiana **   
     |  |                             +--+                           
     |  |                                +-- Pleuronaia_dolabelloides **
     |  |                                                            
     |  |                                +-- Fusconaia_masoni **        
     +--+                             +--+                           
        |                             |  +-- Fusconaia_flava **         
        |  +--------------------------+                              
        |  |                          |  +-- Fusconaia_subrotunda **    
        |  |                          +--+                           
        |  |                             +-- Fusconaia_cor **           
        +--+                                                         
           |                             +-- Parvaspina_collina **      
           |  +--------------------------+                           
           |  |                          +-- Parvaspina_steinstansana **
           |  |                                                      
           +--+                          +-- Eurynia_dilatata **        
              |  +-----------------------+                           
              |  |                       +-- Hemistena_lata **          
              +--+                                                   
                 |  +----------------------- Elliptio_producta[2] **    
                 |  |                                                
                 +--+  +-------------------- Elliptio_producta[1] **    
                    |  |                                             
                    +--+  +----------------- Elliptio_spinosa **        
                       |  |                                          
                       |  |  +-------------- Elliptio_icterina **       
                       +--+  |                                       
                          |  |     +-------- Elliptio_roanokensis **    
                          |  |  +--+                                 
                          +--+  |  |  +----- Elliptio_complanata **     
                             |  |  +--+                              
                             |  |     |  +-- Elliptio_crassidens **     
                             |  |     +--+                           
                             +--+        +-- Elliptio_pullata **        
                                |                                    
                                |        +-- Elliptio_congaraea **      
                                |  +-----+                           
                                |  |     +-- Elliptio_hopetonensis **   
                                |  |                                 
                                +--+     +-- Elliptio_folliculata **    
                                   |  +--+                           
                                   |  |  +-- Elliptio_jayensis **       
                                   +--+                              
                                      |  +-- Elliptio_arca **           
                                      +--+                           
                                         +-- Elliptio_arctata **        

Pieri et al., 2018 [Publ 3867], fig. 3

iT tT bi riS oT roT roS iD
34 34 20 3 3 2 1 38.0
                                      +-- Quadrula_quadrula      
  +-----------------------------------+                          
  |                                   +-- Pustulosa_petrina      
  |                                                              
  |  +----------------------------------- Eurynia_dilatata **       
  |  |                                                           
--+  |                             +----- Pleurobema_clava **       
  |  +-----------------------------+                             
  |  |                             |  +-- Pleurobema_riddellii **   
  |  |                             +--+                          
  |  |                                +-- Pleurobema_sintoxia **    
  +--+                                                           
     |                             +----- Fusconaia_subrotunda **   
     |  +--------------------------+                             
     |  |                          |  +-- Fusconaia_cor **          
     |  |                          +--+                          
     |  |                             +-- Fusconaia_cuneolus **     
     |  |                                                        
     +--+  +----------------------------- Fusconaia_ozarkensis **   
        |  |                                                     
        |  |                 +----------- Fusconaia_cerina[3] **    
        |  |                 |                                   
        |  |  +--------------+  +-------- Fusconaia_flava[1] **     
        +--+  |              |  |                                
           |  |              +--+  +----- Fusconaia_flava[3] **     
           |  |                 |  |                             
           |  |                 +--+  +-- Fusconaia_cerina[1] **    
           |  |                    |  |                          
           |  |                    +--+-- Fusconaia_cerina[2] **    
           +--+                       |                          
              |                       +-- Fusconaia_flava[2] **     
              |                                                  
              |  +----------------------- Fusconaia_masoni **       
              |  |                                               
              |  |                    +-- Fusconaia_burkei **       
              |  +--------------------+                          
              +--+                    +-- Fusconaia_escambia **     
                 |                                               
                 |  +-------------------- Fusconaia_mitchelli **    
                 |  |                                            
                 +--+  +----------------- Fusconaia_chunii **       
                    |  |                                         
                    |  |  +-------------- Fusconaia_askewi[9] **    
                    +--+  |                                      
                       |  |  +----------- Fusconaia_askewi[6] **    
                       |  |  |                                   
                       +--+  +----------- Fusconaia_askewi[7] **    
                          |  |                                   
                          |  +----------- Fusconaia_askewi[8] **    
                          |  |                                   
                          |  +----------- Fusconaia_lananensis[5] **
                          +--+                                   
                             +----------- Fusconaia_lananensis[6] **
                             |                                   
                             |  +-------- Fusconaia_askewi[5] **    
                             |  |                                
                             |  +-------- Fusconaia_lananensis[4] **
                             |  |                                
                             +--+  +----- Fusconaia_askewi[2] **    
                                |  |                             
                                |  +----- Fusconaia_askewi[3] **    
                                |  |                             
                                +--+----- Fusconaia_askewi[4] **    
                                   |                             
                                   |  +-- Fusconaia_askewi[1] **    
                                   |  |                          
                                   |  +-- Fusconaia_lananensis[1] **
                                   +--+                          
                                      +-- Fusconaia_lananensis[2] **
                                      |                          
                                      +-- Fusconaia_lananensis[3] **

Campbell et al., 2008 [Publ 2675], fig. 5

iT tT bi riS oT roT roS iD
27 28 6 8 3 3 3 46.0
  +-------------- Uniomerus_declivis       
  |                                        
  +-------------- Plectomerus_dombeyanus   
  |                                        
  |           +-- Elliptio_arctata **         
  |  +--------+                            
  |  |        +-- Elliptio_crassidens **      
  |  |                                     
  |  |  +-------- Pleurobema_decisum **       
--+  |  |                                  
  |  |  +-------- Pleurobema_rubellum **      
  |  |  |                                  
  |  |  +-------- Pleurobema_georgianum[2] ** 
  |  |  |                                  
  |  |  +-------- Pleurobema_stabile **       
  |  |  |                                  
  |  |  +-------- Pleurobema_pyriforme **     
  |  |  |                                  
  +--+  +-------- Pleurobema_oviforme[2] **   
     |  |                                  
     |  +-------- Pleurobema_hanleyianum[2] **
     |  |                                  
     |  +-------- Pleuronaia_gibber **        
     |  |                                  
     |  +-------- Pleurobema_georgianum[3] ** 
     |  |                                  
     |  +-------- Pleurobema_cordatum **      
     |  |                                  
     |  +-------- Pleurobema_beadleianum **   
     |  |                                  
     |  +-------- Hemistena_lata **           
     +--+                                  
        +-------- Fusconaia_flava **          
        |                                  
        +-------- Fusconaia_cerina **         
        |                                  
        +-------- Eurynia_dilatata **         
        |                                  
        +-------- Elliptio_arca **            
        |                                  
        +-------- Cyrtonaias_tampicoensis  
        |                                  
        |     +-- Plethobasus_cyphyus **      
        +-----+                            
        |     +-- Parvaspina_collina **       
        |                                  
        |     +-- Pleurobema_athearni **      
        |     |                            
        +-----+-- Pleurobema_georgianum[1] ** 
        |     |                            
        |     +-- Pleurobema_hanleyianum[1] **
        |                                  
        |  +----- Pleurobema_oviforme[1] **   
        |  |                               
        +--+----- Pleurobema_clava **         
           |                               
           |  +-- Pleurobema_rubrum **        
           +--+                            
              +-- Pleurobema_sintoxia **      

Campbell et al., 2008 [Publ 2675], fig. 6

iT tT bi riS oT roT roS iD
27 27 23 8 2 2 2 52.0
  +----------------------------------------- Cyrtonaias_tampicoensis 
  |                                                                  
  +----------------------------------------- Plectomerus_dombeyanus  
--+                                                                  
  |  +-------------------------------------- Parvaspina_collina **      
  |  |                                                               
  |  |  +----------------------------------- Hemistena_lata **          
  +--+  |                                                            
     |  |                             +----- Plethobasus_cyphyus **     
     |  |  +--------------------------+                              
     +--+  |                          |  +-- Elliptio_arca **           
        |  |                          +--+                           
        |  |                             +-- Elliptio_crassidens **     
        |  |                                                         
        +--+                             +-- Fusconaia_cerina **        
           |  +--------------------------+                           
           |  |                          +-- Fusconaia_flava **         
           |  |                                                      
           |  |                          +-- Eurynia_dilatata **        
           +--+  +-----------------------+                           
              |  |                       +-- Pleuronaia_gibber **       
              |  |                                                   
              |  |                       +-- Pleurobema_cordatum **     
              +--+                    +--+                           
                 |                    |  +-- Pleurobema_rubrum[1] **    
                 |  +-----------------+                              
                 |  |                 |  +-- Pleurobema_rubrum[2] **    
                 |  |                 +--+                           
                 +--+                    +-- Pleurobema_sintoxia **     
                    |                                                
                    |  +-------------------- Pleurobema_stabile **      
                    |  |                                             
                    +--+  +----------------- Pleurobema_pyriforme **    
                       |  |                                          
                       |  |           +----- Pleurobema_decisum **      
                       +--+  +--------+                              
                          |  |        |  +-- Pleurobema_hanleyianum **  
                          |  |        +--+                           
                          |  |           +-- Pleurobema_georgianum[2] **
                          +--+                                       
                             |           +-- Pleurobema_georgianum[1] **
                             |  +--------+                           
                             |  |        +-- Pleurobema_athearni **     
                             |  |                                    
                             +--+        +-- Pleurobema_oviforme **     
                                |  +-----+                           
                                |  |     +-- Pleurobema_clava **        
                                |  |                                 
                                +--+  +----- Pleurobema_beadleianum **  
                                   |  |                              
                                   |  +----- Pleurobema_rubellum **     
                                   +--+                              
                                      +----- Pleurobema_strodeanum **   
                                      |                              
                                      |  +-- Pleurobema_perovatum **    
                                      +--+                           
                                         +-- Pleurobema_taitianum **    

Campbell et al., 2008 [Publ 2675], fig. 2

iT tT bi riS oT roT roS iD
26 26 17 8 2 2 2 48.0
  +----------------------- Cyrtonaias_tampicoensis 
  |                                                
  +----------------------- Plectomerus_dombeyanus  
  |                                                
  |                    +-- Pleurobema_cordatum **     
  |                    |                           
  |        +-----------+-- Pleurobema_rubrum **       
  |        |           |                           
  |        |           +-- Pleurobema_sintoxia **     
  |        |                                       
--+  +-----+           +-- Fusconaia_cerina **        
  |  |     |  +--------+                           
  |  |     |  |        +-- Fusconaia_flava **         
  |  |     |  |                                    
  |  |     +--+  +-------- Pleuronaia_gibber **       
  |  |        |  |                                 
  |  |        |  |  +----- Eurynia_dilatata **        
  |  |        +--+  |                              
  |  |           |  +----- Hemistena_lata **          
  |  |           |  |                              
  |  |           +--+----- Plethobasus_cyphyus **     
  +--+              |                              
     |              +----- Parvaspina_collina **      
     |              |                              
     |              |  +-- Elliptio_arca **           
     |              +--+                           
     |                 +-- Elliptio_crassidens **     
     |                                             
     |  +----------------- Pleurobema_stabile **      
     |  |                                          
     |  |  +-------------- Pleurobema_pyriforme **    
     |  |  |                                       
     +--+  |           +-- Pleurobema_georgianum[2] **
        |  +-----------+                           
        |  |           +-- Pleurobema_athearni **     
        |  |                                       
        |  |        +----- Pleurobema_decisum **      
        +--+--------+                              
           |        |  +-- Pleurobema_hanleyianum **  
           |        +--+                           
           |           +-- Pleurobema_georgianum[1] **
           |                                       
           |           +-- Pleurobema_clava **        
           |  +--------+                           
           |  |        +-- Pleurobema_oviforme **     
           +--+                                    
              |  +-------- Pleurobema_rubellum **     
              |  |                                 
              +--+-------- Pleurobema_strodeanum **   
                 |                                 
                 |  +----- Pleurobema_beadleianum **  
                 +--+                              
                    |  +-- Pleurobema_taitianum **    
                    +--+                           
                       +-- Pleurobema_perovatum **    

Campbell et al., 2008 [Publ 2675], fig. 4

iT tT bi riS oT roT roS iD
26 27 9 8 3 3 3 43.0
  +----------------------- Uniomerus_declivis       
  |                                                 
  +----------------------- Plectomerus_dombeyanus   
  |                                                 
  |  +-------------------- Cyrtonaias_tampicoensis  
  |  |                                              
  |  +-------------------- Eurynia_dilatata **         
  |  |                                              
  |  +-------------------- Fusconaia_cerina **         
  |  |                                              
--+  +-------------------- Fusconaia_flava **          
  |  |                                              
  |  +-------------------- Hemistena_lata **           
  |  |                                              
  |  +-------------------- Plethobasus_cyphyus **      
  |  |                                              
  |  +-------------------- Parvaspina_collina **       
  |  |                                              
  |  +-------------------- Pleurobema_cordatum **      
  |  |                                              
  |  +-------------------- Pleuronaia_gibber **        
  +--+                                              
     +-------------------- Pleurobema_oviforme[2] **   
     |                                              
     +-------------------- Pleurobema_stabile **       
     |                                              
     |              +----- Elliptio_arca **            
     +--------------+                               
     |              |  +-- Elliptio_crassidens **      
     |              +--+                            
     |                 +-- Elliptio_arctata **         
     |                                              
     |  +----------------- Pleurobema_decisum **       
     |  |                                           
     |  +----------------- Pleurobema_clava **         
     |  |                                           
     |  +----------------- Pleurobema_oviforme[1] **   
     |  |                                           
     +--+              +-- Pleurobema_rubrum **        
        +--------------+                            
        |              +-- Pleurobema_sintoxia **      
        |                                           
        |  +-------------- Pleurobema_beadleianum **   
        |  |                                        
        +--+  +----------- Pleurobema_pyriforme **     
           |  |                                     
           +--+  +-------- Pleurobema_rubellum **      
              |  |                                  
              |  +-------- Pleurobema_hanleyianum[2] **
              +--+                                  
                 +-------- Pleurobema_georgianum[2] ** 
                 |                                  
                 |  +----- Pleurobema_hanleyianum[1] **
                 +--+                               
                    |  +-- Pleurobema_georgianum[1] ** 
                    +--+                            
                       +-- Pleurobema_athearni **      

Campbell & Lydeard, 2012 [Publ 3308], fig. 1

iT tT bi riS oT roT roS iD
26 26 24 9 17 17 4 43.0
                                      +----------- Quadrula_quadrula       
                                      |                                    
  +-----------------------------------+        +-- Pustulosa_infucata      
  |                                   |  +-----+                           
  |                                   |  |     +-- Pustulosa_kleiniana     
  |                                   +--+                                 
  |                                      |  +----- Cyclonaias_tuberculata  
  |                                      +--+                              
  |                                         |  +-- Pustulosa_succissa      
  |                                         +--+                           
  |                                            +-- Pustulosa_pustulosa     
  |                                                                        
  |                                         +----- Popenaias_popeii        
  |                             +-----------+                              
--+                             |           |  +-- Amblema_elliottii       
  |                             |           +--+                           
  |                             |              +-- Amblema_plicata         
  |  +--------------------------+                                          
  |  |                          |           +----- Cyrtonaias_tampicoensis 
  |  |                          |  +--------+                              
  |  |                          |  |        |  +-- Reginaia_ebenus         
  |  |                          |  |        +--+                           
  |  |                          +--+           +-- Reginaia_rotulata       
  |  |                             |                                       
  |  |                             |  +----------- Plectomerus_dombeyanus  
  |  |                             +--+                                    
  |  |                                |  +-------- Lemiox_rimosus          
  +--+                                +--+                                 
     |                                   |  +----- Lampsilis_ovata         
     |                                   +--+                              
     |                                      |  +-- Obovaria_unicolor       
     |                                      +--+                           
     |                                         +-- Obovaria_olivaria       
     |                                                                     
     |  +----------------------------------------- Elliptoideus_sloatianus ** 
     |  |                                                                  
     |  |  +-------------------------------------- Pleurobema_stabile **      
     +--+  |                                                               
        |  |                                +----- Pleurobema_pyriforme **    
        |  |  +-----------------------------+                              
        +--+  |                             |  +-- Pleurobema_clava **        
           |  |                             +--+                           
           |  |                                +-- Pleurobema_decisum **      
           |  |                                                            
           +--+                                +-- Pleurobema_cordatum **     
              |  +-----------------------------+                           
              |  |                             +-- Pleurobema_sintoxia **     
              |  |                                                         
              |  |                             +-- Eurynia_dilatata **        
              |  |                       +-----+                           
              +--+                       |     +-- Hemistena_lata **          
                 |  +--------------------+                                 
                 |  |                    |  +----- Plethobasus_cyphyus **     
                 |  |                    +--+                              
                 |  |                       |  +-- Elliptio_arca **           
                 |  |                       +--+                           
                 +--+                          +-- Elliptio_crassidens **     
                    |                                                      
                    |                       +----- Pleuronaia_gibber **       
                    |  +--------------------+                              
                    |  |                    |  +-- Pleuronaia_barnesiana **   
                    |  |                    +--+                           
                    +--+                       +-- Pleuronaia_dolabelloides **
                       |                                                   
                       |  +----------------------- Parvaspina_collina **      
                       |  |                                                
                       +--+  +-------------------- Fusconaia_subrotunda **    
                          |  |                                             
                          +--+                 +-- Fusconaia_cor **           
                             |  +--------------+                           
                             |  |              +-- Fusconaia_cuneolus **      
                             +--+                                          
                                |  +-------------- Fusconaia_ozarkensis **    
                                |  |                                       
                                +--+           +-- Fusconaia_burkei **        
                                   |  +--------+                           
                                   |  |        +-- Fusconaia_escambia **      
                                   +--+                                    
                                      |  +-------- Fusconaia_masoni **        
                                      +--+                                 
                                         |  +----- Fusconaia_cerina[2] **     
                                         +--+                              
                                            |  +-- Fusconaia_flava **         
                                            +--+                           
                                               +-- Fusconaia_cerina[1] **     

Campbell & Lydeard, 2012 [Publ 3308], fig. 2

iT tT bi riS oT roT roS iD
26 26 20 9 17 17 4 39.0
                       +----------- Quadrula_quadrula       
                       |                                    
  +--------------------+        +-- Pustulosa_infucata      
  |                    |  +-----+                           
  |                    |  |     +-- Pustulosa_kleiniana     
  |                    +--+                                 
  |                       |  +----- Cyclonaias_tuberculata  
  |                       +--+                              
  |                          |  +-- Pustulosa_succissa      
  |                          +--+                           
  |                             +-- Pustulosa_pustulosa     
  |                                                         
  |                             +-- Reginaia_ebenus         
  |                          +--+                           
  |                          |  +-- Reginaia_rotulata       
--+              +-----------+                              
  |              |           |  +-- Amblema_elliottii       
  |              |           +--+                           
  |  +-----------+              +-- Amblema_plicata         
  |  |           |                                          
  |  |           |  +-------------- Popenaias_popeii        
  |  |           |  |                                       
  |  |           +--+  +----------- Cyrtonaias_tampicoensis 
  |  |              |  |                                    
  |  |              +--+----------- Plectomerus_dombeyanus  
  |  |                 |                                    
  |  |                 |  +-------- Lemiox_rimosus          
  |  |                 +--+                                 
  |  |                    |  +----- Lampsilis_ovata         
  +--+                    +--+                              
     |                       |  +-- Obovaria_unicolor       
     |                       +--+                           
     |                          +-- Obovaria_olivaria       
     |                                                      
     |  +-------------------------- Elliptoideus_sloatianus ** 
     |  |                                                   
     |  |  +----------------------- Hemistena_lata **          
     |  |  |                                                
     |  |  +----------------------- Plethobasus_cyphyus **     
     |  |  |                                                
     +--+  |                    +-- Elliptio_arca **           
        |  +--------------------+                           
        |  |                    +-- Elliptio_crassidens **     
        |  |                                                
        |  |                    +-- Eurynia_dilatata **        
        |  +--------------------+                           
        |  |                    +-- Parvaspina_collina **      
        +--+                                                
           |                 +----- Pleuronaia_gibber **       
           |  +--------------+                              
           |  |              |  +-- Pleuronaia_barnesiana **   
           |  |              +--+                           
           |  |                 +-- Pleuronaia_dolabelloides **
           |  |                                             
           |  |                 +-- Pleurobema_cordatum **     
           |  |        +--------+                           
           |  |        |        +-- Pleurobema_sintoxia **     
           |  +--------+                                    
           |  |        |  +-------- Pleurobema_stabile **      
           +--+        +--+                                 
              |           |  +----- Pleurobema_pyriforme **    
              |           +--+                              
              |              |  +-- Pleurobema_clava **        
              |              +--+                           
              |                 +-- Pleurobema_decisum **      
              |                                             
              |              +----- Fusconaia_subrotunda **    
              |  +-----------+                              
              |  |           |  +-- Fusconaia_cor **           
              |  |           +--+                           
              +--+              +-- Fusconaia_cuneolus **      
                 |                                          
                 |  +-------------- Fusconaia_ozarkensis **    
                 |  |                                       
                 +--+           +-- Fusconaia_burkei **        
                    |  +--------+                           
                    |  |        +-- Fusconaia_escambia **      
                    +--+                                    
                       |  +-------- Fusconaia_masoni **        
                       +--+                                 
                          |  +----- Fusconaia_cerina[2] **     
                          +--+                              
                             |  +-- Fusconaia_flava **         
                             +--+                           
                                +-- Fusconaia_cerina[1] **     

Campbell & Lydeard, 2012 [Publ 3308], fig. 4

iT tT bi riS oT roT roS iD
26 26 24 9 17 17 4 43.0
                                         +----------- Quadrula_quadrula       
                                         |                                    
  +--------------------------------------+        +-- Pustulosa_infucata      
  |                                      |  +-----+                           
  |                                      |  |     +-- Pustulosa_kleiniana     
  |                                      +--+                                 
  |                                         |  +----- Cyclonaias_tuberculata  
  |                                         +--+                              
  |                                            |  +-- Pustulosa_succissa      
  |                                            +--+                           
  |                                               +-- Pustulosa_pustulosa     
  |                                                                           
  |                                            +----- Popenaias_popeii        
  |                                   +--------+                              
--+                                   |        |  +-- Amblema_elliottii       
  |                                   |        +--+                           
  |                                   |           +-- Amblema_plicata         
  |  +--------------------------------+                                       
  |  |                                |  +----------- Plectomerus_dombeyanus  
  |  |                                |  |                                    
  |  |                                |  |     +----- Cyrtonaias_tampicoensis 
  |  |                                |  +-----+                              
  |  |                                +--+     |  +-- Reginaia_ebenus         
  |  |                                   |     +--+                           
  |  |                                   |        +-- Reginaia_rotulata       
  |  |                                   |                                    
  +--+                                   |  +-------- Lemiox_rimosus          
     |                                   +--+                                 
     |                                      |  +----- Lampsilis_ovata         
     |                                      +--+                              
     |                                         |  +-- Obovaria_unicolor       
     |                                         +--+                           
     |                                            +-- Obovaria_olivaria       
     |                                                                        
     |  +-------------------------------------------- Elliptoideus_sloatianus ** 
     |  |                                                                     
     +--+  +----------------------------------------- Pleurobema_stabile **      
        |  |                                                                  
        +--+  +-------------------------------------- Hemistena_lata **          
           |  |                                                               
           +--+  +----------------------------------- Eurynia_dilatata **        
              |  |                                                            
              |  |                                +-- Pleurobema_cordatum **     
              +--+  +-----------------------------+                           
                 |  |                             +-- Pleurobema_sintoxia **     
                 |  |                                                         
                 |  |                          +----- Pleurobema_clava **        
                 +--+  +-----------------------+                              
                    |  |                       |  +-- Pleurobema_decisum **      
                    |  |                       +--+                           
                    |  |                          +-- Pleurobema_pyriforme **    
                    |  |                                                      
                    +--+                    +-------- Parvaspina_collina **      
                       |  +-----------------+                                 
                       |  |                 |  +----- Pleuronaia_gibber **       
                       |  |                 +--+                              
                       |  |                    |  +-- Pleuronaia_barnesiana **   
                       |  |                    +--+                           
                       +--+                       +-- Pleuronaia_dolabelloides **
                          |                                                   
                          |                    +----- Plethobasus_cyphyus **     
                          |  +-----------------+                              
                          |  |                 |  +-- Elliptio_arca **           
                          |  |                 +--+                           
                          +--+                    +-- Elliptio_crassidens **     
                             |                                                
                             |  +-------------------- Fusconaia_subrotunda **    
                             |  |                                             
                             +--+                 +-- Fusconaia_cor **           
                                |  +--------------+                           
                                |  |              +-- Fusconaia_cuneolus **      
                                +--+                                          
                                   |  +-------------- Fusconaia_ozarkensis **    
                                   |  |                                       
                                   +--+           +-- Fusconaia_burkei **        
                                      |  +--------+                           
                                      |  |        +-- Fusconaia_escambia **      
                                      +--+                                    
                                         |  +-------- Fusconaia_masoni **        
                                         +--+                                 
                                            |  +----- Fusconaia_cerina[2] **     
                                            +--+                              
                                               |  +-- Fusconaia_flava **         
                                               +--+                           
                                                  +-- Fusconaia_cerina[1] **     

Perkins et al., 2017 [Publ 3777], fig. 3

iT tT bi riS oT roT roS iD
26 26 10 7 2 2 2 46.0
  +----------------------- Plectomerus_dombeyanus  
  |                                                
  +----------------------- Uniomerus_declivis      
  |                                                
  |                    +-- Parvaspina_steinstansana **
  |                    |                           
  |                    +-- Parvaspina_collina[1] **   
--+                    |                           
  |  +-----------------+-- Parvaspina_collina[2] **   
  |  |                 |                           
  |  |                 +-- Parvaspina_collina[3] **   
  |  |                 |                           
  |  |                 +-- Parvaspina_collina[4] **   
  |  |                                             
  +--+  +----------------- Elliptio_arca **           
     |  |                                          
     |  +----------------- Hemistena_lata **          
     |  |                                          
     |  |              +-- Elliptio_arctata **        
     |  +--------------+                           
     |  |              +-- Elliptio_crassidens **     
     +--+                                          
        |  +-------------- Eurynia_dilatata **        
        |  |                                       
        |  |  +----------- Pleurobema_oviforme **     
        |  |  |                                    
        |  |  +----------- Pleurobema_clava **        
        |  |  |                                    
        +--+  +----------- Pleurobema_pyriforme **    
           |  |                                    
           |  +----------- Pleurobema_georgianum **   
           |  |                                    
           |  +----------- Pleurobema_beadleianum **  
           |  |                                    
           |  +----------- Pleurobema_hanleyianum **  
           +--+                                    
              +----------- Pleurobema_decisum **      
              |                                    
              |     +----- Pleurobema_rubellum **     
              +-----+                              
              |     |  +-- Pleurobema_sintoxia **     
              |     +--+                           
              |        +-- Pleurobema_rubrum **       
              |                                    
              |  +-------- Elliptio_spinosa **        
              |  |                                 
              +--+  +----- Fusconaia_escambia **      
                 |  |                              
                 |  +----- Fusconaia_flava **         
                 +--+                              
                    +----- Fusconaia_cerina **        
                    |                              
                    |  +-- Pleuronaia_gibber **       
                    +--+                           
                       +-- Pleuronaia_dolabelloides **

Pfeiffer et al., 2016 [Publ 3660], fig. 2

iT tT bi riS oT roT roS iD
22 28 23 9 17 16 3 45.0
                                   +-- Amblema_plicata         
                                +--+                           
                                |  +-- Amblema_neislerii       
  +-----------------------------+                              
  |                             |  +-- Reginaia_rotulata       
  |                             +--+                           
  |                                +-- Reginaia_ebenus         
  |                                                            
  |                                +-- Cyrtonaias_tampicoensis 
  |                          +-----+                           
  |                          |     +-- Glebula_rotundata       
--+                          |                                 
  |  +-----------------------+     +-- Lampsilis_teres         
  |  |                       |  +--+                           
  |  |                       |  |  +-- Leaunio_lienosus        
  |  |                       +--+                              
  |  |                          |  +-- Truncilla_macrodon      
  |  |                          +--+                           
  |  |                             +-- Potamilus_fragilis      
  +--+                                                         
     |  +----------------------------- Elliptoideus_sloatianus ** 
     |  |                                                      
     |  |              +-------------- Uniomerus_declivis      
     |  |  +-----------+                                       
     |  |  |           |  +----------- Quadrula_quadrula       
     |  |  |           +--+                                    
     +--+  |              |  +-------- Quadrula_verrucosa      
        |  |              +--+                                 
        |  |                 |  +----- Pustulosa_pustulosa     
        |  |                 +--+                              
        |  |                    |  +-- Pustulosa_petrina       
        |  |                    +--+                           
        +--+                       +-- Cyclonaias_tuberculata  
           |                                                   
           |                       +-- Elliptio_fumata **         
           |  +--------------------+                           
           |  |                    +-- Elliptio_crassidens **     
           |  |                                                
           |  |  +-------------------- Parvaspina_collina **      
           |  |  |                                             
           |  |  +-------------------- Plethobasus_cyphyus **     
           +--+  |                                             
              |  |                 +-- Hemistena_lata **          
              |  +-----------------+                           
              |  |                 +-- Eurynia_dilatata **        
              |  |                                             
              |  |              +----- Pleurobema_riddellii **    
              +--+           +--+                              
                 |           |  |  +-- Pleurobema_sintoxia **     
                 |           |  +--+                           
                 |  +--------+     +-- Pleurobema_cordatum **     
                 |  |        |                                 
                 |  |        |  +----- Pleuronaia_dolabelloides **
                 |  |        |  |                              
                 |  |        +--+  +-- Pleurobema_pyriforme **    
                 |  |           |  |                           
                 +--+           +--+-- Pleurobema_strodeanum **   
                    |              |                           
                    |              +-- Pleurobema_clava **        
                    |                                          
                    |  +-------------- Fusconaia_askewi **        
                    |  |                                       
                    |  |           +-- Fusconaia_mitchelli **     
                    +--+  +--------+                           
                       |  |        +-- Fusconaia_iheringi **      
                       |  |                                    
                       +--+        +-- Fusconaia_escambia **      
                          |  +-----+                           
                          |  |     +-- Fusconaia_burkei **        
                          +--+                                 
                             |  +----- Fusconaia_flava[2] **      
                             +--+                              
                                |  +-- Fusconaia_cerina **        
                                +--+                           
                                   +-- Fusconaia_flava[1] **      

Smith et al., 2020 [Publ 4136], fig. 1

iT tT bi riS oT roT roS iD
19 19 12 9 1 1 1 41.0
  +-------------------------- Quadrula_quadrula       
  |                                                   
--+  +----------------------- Elliptoideus_sloatianus ** 
  |  |                                                
  |  |  +-------------------- Elliptio_crassidens **     
  +--+  |                                             
     |  +-------------------- Plethobasus_cyphyus **     
     |  |                                             
     +--+  +----------------- Parvaspina_steinstansana **
        |  |                                          
        |  |              +-- Hemistena_lata **          
        |  +--------------+                           
        |  |              +-- Eurynia_dilatata **        
        +--+                                          
           |              +-- Pleuronaia_barnesiana **   
           |  +-----------+                           
           |  |           +-- Pleurobema_clava **        
           |  |                                       
           |  |  +----------- Fusconaia_subrotunda **    
           +--+  |                                    
              |  |        +-- Fusconaia_cor **           
              |  +--------+                           
              |  |        +-- Fusconaia_cuneolus **      
              +--+                                    
                 |  +-------- Fusconaia_flava **         
                 |  |                                 
                 |  +-------- Fusconaia_masoni **        
                 |  |                                 
                 |  |     +-- Fusconaia_burkei **        
                 +--+-----+                           
                    |     +-- Fusconaia_escambia **      
                    |                                 
                    |     +-- Fusconaia_iheringi **      
                    |  +--+                           
                    |  |  +-- Fusconaia_mitchelli **     
                    +--+                              
                       |  +-- Fusconaia_askewi **        
                       +--+                           
                          +-- Fusconaia_chunii **        

Kandl et al., 2001 [Publ 3384], fig. 4

iT tT bi riS oT roT roS iD
12 13 4 2 1 1 1 42.5
  +-------------- Elliptio **               
  |                                      
  +-------------- Uniomerus_carolinianus 
--+                                      
  |  +----------- Pleurobema_sintoxia **    
  |  |                                   
  +--+  +-------- Pleurobema_strodeanum **  
     |  |                                
     |  |  +----- Pleurobema_pyriforme[7] **
     +--+  |                             
        |  |  +-- Pleurobema_pyriforme[1] **
        |  |  |                          
        +--+  +-- Pleurobema_pyriforme[2] **
           |  |                          
           |  +-- Pleurobema_pyriforme[3] **
           |  |                          
           |  +-- Pleurobema_pyriforme[4] **
           +--+                          
              +-- Pleurobema_pyriforme[5] **
              |                          
              +-- Pleurobema[1] **          
              |                          
              +-- Pleurobema_pyriforme[6] **
              |                          
              +-- Pleurobema[2] **          

Kandl et al., 2001 [Publ 3384], fig. 5

iT tT bi riS oT roT roS iD
12 12 6 3 2 2 2 40.0
  +-------------------- Amblema_plicata        
  |                                            
  +-------------------- Quadrula_quadrula      
--+                                            
  |  +----------------- Elliptio_crassidens **    
  |  |                                         
  +--+  +-------------- Fusconaia_flava **        
     |  |                                      
     +--+  +----------- Pleurobema_sintoxia **    
        |  |                                   
        |  |        +-- Pleurobema_clava **       
        +--+  +-----+                          
           |  |     +-- Pleurobema_strodeanum **  
           |  |                                
           +--+  +----- Pleurobema_pyriforme[6] **
              |  |                             
              |  |  +-- Pleurobema_pyriforme[1] **
              +--+  |                          
                 |  +-- Pleurobema **             
                 |  |                          
                 |  +-- Pleurobema_pyriforme[2] **
                 +--+                          
                    +-- Pleurobema_pyriforme[3] **
                    |                          
                    +-- Pleurobema_pyriforme[4] **
                    |                          
                    +-- Pleurobema_pyriforme[5] **

Smith et al., 2020 [Publ 4136], fig. 3

iT tT bi riS oT roT roS iD
12 12 10 2 0 0 0 42.0
  +-------------- Pleurobema_clava **    
  |                                   
  |        +----- Fusconaia_subrotunda **
  |  +-----+                          
--+  |     |  +-- Fusconaia_cor **       
  |  |     +--+                       
  |  |        +-- Fusconaia_cuneolus **  
  |  |                                
  |  |        +-- Fusconaia_iheringi **  
  +--+     +--+                       
     |     |  +-- Fusconaia_mitchelli ** 
     |  +--+                          
     |  |  |  +-- Fusconaia_askewi **    
     |  |  +--+                       
     |  |     +-- Fusconaia_chunii **    
     +--+                             
        |     +-- Fusconaia_flava **     
        |  +--+                       
        |  |  +-- Fusconaia_masoni **    
        +--+                          
           |  +-- Fusconaia_burkei **    
           +--+                       
              +-- Fusconaia_escambia **  

Inoue et al., 2020 [Publ 3941], fig. 3

iT tT bi riS oT roT roS iD
8 8 6 6 36 35 3 41.0
                                            +-- Pustulosa_petrina       
  +-----------------------------------------+                           
  |                                         +-- Quadrula_quadrula       
  |                                                                     
  |                             +-------------- Parvaspina_collina **      
  |                             |                                       
  |                             |           +-- Eurynia_dilatata **        
  |  +--------------------------+  +--------+                           
  |  |                          |  |        +-- Hemistena_lata **          
--+  |                          |  |                                    
  |  |                          +--+        +-- Pleurobema_clava **        
  |  |                             |  +-----+                           
  |  |                             |  |     +-- Pleurobema_sintoxia **     
  |  |                             +--+                                 
  |  |                                |  +----- Pleuronaia_dolabelloides **
  |  |                                +--+                              
  |  |                                   |  +-- Fusconaia_flava **         
  +--+                                   +--+                           
     |                                      +-- Fusconaia_subrotunda **    
     |                                                                  
     |                                      +-- Amblema_plicata         
     |                                   +--+                           
     |                                   |  +-- Amblema_elliottii       
     |  +--------------------------------+                              
     |  |                                |  +-- Reginaia_ebenus         
     |  |                                +--+                           
     |  |                                   +-- Reginaia_rotulata       
     |  |                                                               
     +--+  +----------------------------------- Toxolasma_lividum       
        |  |                                                            
        |  |                          +-------- Potamilus_fragilis      
        |  |                          |                                 
        |  |  +-----------------------+     +-- Potamilus_ohiensis      
        +--+  |                       |  +--+                           
           |  |                       |  |  +-- Potamilus_amphichaenus  
           |  |                       +--+                              
           |  |                          |  +-- Potamilus_metnecktayi   
           |  |                          +--+                           
           |  |                             +-- Potamilus_purpuratus    
           +--+                                                         
              |                       +-------- Ellipsaria_lineolata    
              |  +--------------------+                                 
              |  |                    |  +----- Truncilla_macrodon      
              |  |                    +--+                              
              |  |                       |  +-- Truncilla_donaciformis  
              |  |                       +--+                           
              |  |                          +-- Truncilla_cognata       
              +--+                                                      
                 |  +-------------------------- Leaunio_lienosus        
                 |  |                                                   
                 |  |     +-------------------- Lampsilis_bracteata     
                 |  |     |                                             
                 |  |  +--+  +----------------- Lampsilis_fasciola      
                 |  |  |  |  |                                          
                 +--+  |  +--+  +-------------- Lampsilis_ornata        
                    |  |     |  |                                       
                    |  |     +--+           +-- Lampsilis_ovata         
                    |  |        |  +--------+                           
                    |  |        |  |        +-- Lampsilis_cariosa       
                    |  |        +--+                                    
                    |  |           |  +-------- Lampsilis_cardium[3]    
                    +--+           +--+                                 
                       |              |  +----- Lampsilis_cardium[2]    
                       |              +--+                              
                       |                 |  +-- Lampsilis_satura        
                       |                 +--+                           
                       |                    +-- Lampsilis_cardium[1]    
                       |                                                
                       |                    +-- Lampsilis_higginsii     
                       |     +--------------+                           
                       |     |              +-- Ortmanniana_ligamentina 
                       +-----+                                          
                             |  +-------------- Lampsilis_teres         
                             |  |                                       
                             +--+           +-- Lampsilis_radiata       
                                |  +--------+                           
                                |  |        +-- Lampsilis_siliquoidea   
                                +--+                                    
                                   |  +-------- Lampsilis_bergmanni     
                                   +--+                                 
                                      |  +----- Lampsilis_hydiana       
                                      +--+                              
                                         |  +-- Lampsilis_virescens     
                                         +--+                           
                                            +-- Lampsilis_straminea     

Inoue et al., 2020 [Publ 3941], fig. 4

iT tT bi riS oT roT roS iD
8 8 6 6 35 34 3 41.0
                                            +-- Reginaia_ebenus         
  +-----------------------------------------+                           
  |                                         +-- Reginaia_rotulata       
  |                                                                     
  |                             +-------------- Parvaspina_collina **      
  |                             |                                       
  |                             |           +-- Eurynia_dilatata **        
  |  +--------------------------+  +--------+                           
--+  |                          |  |        +-- Hemistena_lata **          
  |  |                          |  |                                    
  |  |                          +--+        +-- Pleurobema_clava **        
  |  |                             |  +-----+                           
  |  |                             |  |     +-- Pleurobema_sintoxia **     
  |  |                             +--+                                 
  |  |                                |  +----- Pleuronaia_dolabelloides **
  +--+                                +--+                              
     |                                   |  +-- Fusconaia_flava **         
     |                                   +--+                           
     |                                      +-- Fusconaia_subrotunda **    
     |                                                                  
     |                                      +-- Amblema_plicata         
     |  +-----------------------------------+                           
     |  |                                   +-- Amblema_elliottii       
     |  |                                                               
     +--+  +----------------------------------- Toxolasma_lividum       
        |  |                                                            
        |  |  +-------------------------------- Ellipsaria_lineolata    
        +--+  |                                                         
           |  |                          +----- Truncilla_cognata       
           |  |  +-----------------------+                              
           |  |  |                       |  +-- Truncilla_macrodon      
           +--+  |                       +--+                           
              |  |                          +-- Truncilla_donaciformis  
              |  |                                                      
              |  |                    +-------- Potamilus_fragilis      
              |  |                    |                                 
              +--+  +-----------------+     +-- Potamilus_ohiensis      
                 |  |                 |  +--+                           
                 |  |                 |  |  +-- Potamilus_amphichaenus  
                 |  |                 +--+                              
                 |  |                    |  +-- Potamilus_metnecktayi   
                 |  |                    +--+                           
                 |  |                       +-- Potamilus_purpuratus    
                 +--+                                                   
                    |  +----------------------- Leaunio_lienosus        
                    |  |                                                
                    |  |     +----------------- Lampsilis_bracteata     
                    |  |     |                                          
                    |  |  +--+              +-- Lampsilis_ornata        
                    |  |  |  |  +-----------+                           
                    +--+  |  |  |           +-- Lampsilis_fasciola      
                       |  |  +--+                                       
                       |  |     |  +----------- Lampsilis_ovata         
                       |  |     +--+                                    
                       |  |        |  +-------- Lampsilis_cardium[3]    
                       |  |        +--+                                 
                       +--+           |  +----- Lampsilis_cardium[2]    
                          |           +--+                              
                          |              |  +-- Lampsilis_satura        
                          |              +--+                           
                          |                 +-- Lampsilis_cardium[1]    
                          |                                             
                          |     +-------------- Lampsilis_teres         
                          |     |                                       
                          |     |           +-- Lampsilis_higginsii     
                          +-----+  +--------+                           
                                |  |        +-- Ortmanniana_ligamentina 
                                |  |                                    
                                +--+        +-- Lampsilis_radiata       
                                   |  +-----+                           
                                   |  |     +-- Lampsilis_siliquoidea   
                                   |  |                                 
                                   +--+     +-- Lampsilis_hydiana       
                                      |  +--+                           
                                      |  |  +-- Lampsilis_bergmanni     
                                      +--+                              
                                         |  +-- Lampsilis_virescens     
                                         +--+                           
                                            +-- Lampsilis_straminea     

Lydeard et al., 2000 [Publ 2654], fig. 2a

iT tT bi riS oT roT roS iD
6 6 4 2 11 9 3 39.0
              +----- Quadrula_nobilis   
  +-----------+                         
  |           |  +-- Pustulosa_infucata 
  |           +--+                      
--+              +-- Pustulosa_succissa 
  |                                     
  |  +-------------- Pleurobema_decisum ** 
  +--+                                  
     |  +----------- Fusconaia_flava **    
     +--+                               
        |  +-------- Fusconaia_burkei[1] **
        +--+                            
           |  +----- Fusconaia_burkei[2] **
           +--+                         
              |  +-- Fusconaia_burkei[3] **
              +--+                      
                 +-- Fusconaia_escambia ** 

Lydeard et al., 2000 [Publ 2654], fig. 2b

iT tT bi riS oT roT roS iD
6 6 4 2 11 9 3 39.0
                    +-- Reginaia_rotulata[1]
                    |                       
  +-----------------+-- Reginaia_rotulata[2]
  |                 |                       
  |                 +-- Reginaia_ebenus     
  |                                         
--+                 +-- Quadrula_nobilis    
  |                 |                       
  |  +--------------+-- Pustulosa_infucata  
  |  |              |                       
  +--+              +-- Pustulosa_succissa  
     |                                      
     |  +-------------- Pleurobema_decisum **  
     +--+                                   
        |  +----------- Fusconaia_flava **     
        +--+                                
           |  +-------- Fusconaia_burkei[1] ** 
           +--+                             
              |  +----- Fusconaia_burkei[2] ** 
              +--+                          
                 |  +-- Fusconaia_burkei[3] ** 
                 +--+                       
                    +-- Fusconaia_escambia **  

Lydeard et al., 2000 [Publ 2654], fig. 2c

iT tT bi riS oT roT roS iD
6 6 2 2 10 8 3 39.0
           +-- Reginaia_ebenus    
  +--------+                      
  |        +-- Reginaia_rotulata  
--+                               
  |  +-------- Pleurobema_decisum ** 
  |  |                            
  +--+  +----- Fusconaia_flava **    
     |  |                         
     +--+  +-- Fusconaia_burkei[1] **
        |  |                      
        |  +-- Fusconaia_burkei[2] **
        +--+                      
           +-- Fusconaia_burkei[3] **
           |                      
           +-- Fusconaia_escambia ** 

Lydeard et al., 2000 [Publ 2654], fig. 2d

iT tT bi riS oT roT roS iD
6 6 4 2 10 8 3 39.0
                 +----- Lampsilis_teres    
  +--------------+                         
  |              |  +-- Obovaria_unicolor  
  |              +--+                      
  |                 +-- Obovaria_olivaria  
--+                                        
  |                 +-- Quadrula_nobilis   
  |                 |                      
  |  +--------------+-- Pustulosa_infucata 
  |  |              |                      
  +--+              +-- Pustulosa_succissa 
     |                                     
     |  +-------------- Pleurobema_decisum ** 
     +--+                                  
        |  +----------- Fusconaia_flava **    
        +--+                               
           |  +-------- Fusconaia_burkei[1] **
           +--+                            
              |  +----- Fusconaia_burkei[2] **
              +--+                         
                 |  +-- Fusconaia_burkei[3] **
                 +--+                      
                    +-- Fusconaia_escambia ** 

Wu et al., 2018 [Publ 3836], fig. 2

iT tT bi riS oT roT roS iD
6 6 4 4 47 3 3 45.0
  +----------- Pleurobema_sintoxia **
  |                               
--+        +-- Elliptio_complanata **
  |  +-----+                      
  |  |     +-- Eurynia_dilatata **   
  +--+                            
     |  +----- Fusconaia_askewi **   
     +--+                         
        |  +-- Fusconaia_cerina **   
        +--+                      
           +-- Fusconaia_burkei **   

Pfeiffer et al., 2019 [Publ 3866], fig. 1

iT tT bi riS oT roT roS iD
6 6 4 5 33 32 4 42.0
                          +----------- Megalonaias_nervosa        
  +-----------------------+                                       
  |                       |  +-------- Uniomerus_declivis         
  |                       +--+                                    
  |                          |  +----- Quadrula_verrucosa         
  |                          +--+                                 
  |                             |  +-- Cyclonaias_tuberculata     
  |                             +--+                              
  |                                +-- Quadrula_quadrula          
  |                                                               
  |                    +-------------- Elliptoideus_sloatianus **    
--+  +-----------------+                                          
  |  |                 |  +----------- Elliptio_fumata **            
  |  |                 +--+                                       
  |  |                    |  +-------- Pleuronaia_barnesiana **      
  |  |                    +--+                                    
  |  |                       |  +----- Pleurobema_riddellii **       
  |  |                       +--+                                 
  |  |                          |  +-- Fusconaia_flava **            
  |  |                          +--+                              
  |  |                             +-- Pleurobema_strodeanum **      
  |  |                                                            
  +--+                             +-- Amblema_plicata            
     |                          +--+                              
     |                          |  +-- Amblema_neislerii          
     |  +-----------------------+                                 
     |  |                       |  +-- Reginaia_ebenus            
     |  |                       +--+                              
     |  |                          +-- Reginaia_rotulata          
     |  |                                                         
     |  |           +----------------- Popenaias_popeii           
     |  |           |                                             
     |  |  +--------+  +-------------- Psoronaias_guatemalensis[2]
     |  |  |        |  |                                          
     +--+  |        +--+  +----------- Psoronaias_guatemalensis[1]
        |  |           |  |                                       
        |  |           +--+  +-------- Psoronaias_semigranosa[3]  
        |  |              |  |                                    
        |  |              |  |  +----- Psoronaias_semigranosa[2]  
        |  |              +--+  |                                 
        |  |                 |  +----- Psoronaias_usumasintae[2]  
        |  |                 |  |                                 
        |  |                 +--+----- Psoronaias_percompressa    
        +--+                    |                                 
           |                    |  +-- Psoronaias_semigranosa[1]  
           |                    |  |                              
           |                    +--+-- Psoronaias_profunda        
           |                       |                              
           |                       +-- Psoronaias_usumasintae[1]  
           |                                                      
           |                       +-- Cyrtonaias_tampicoensis    
           |  +--------------------+                              
           |  |                    +-- Glebula_rotundata          
           |  |                                                   
           +--+  +-------------------- Potamilus_alatus           
              |  |                                                
              |  |                 +-- Truncilla_truncata         
              +--+  +--------------+                              
                 |  |              +-- Ptychobranchus_fasciolaris 
                 |  |                                             
                 +--+  +-------------- Pachynaias_spheniopsis     
                    |  |                                          
                    |  |           +-- Cyrtonaias_explicata       
                    +--+  +--------+                              
                       |  |        +-- Sphenonaias_microdon       
                       |  |                                       
                       +--+        +-- Lampsilis_cardium          
                          |  +-----+                              
                          |  |     +-- Ortmanniana_ligamentina    
                          +--+                                    
                             |  +----- Cambarunio_iris            
                             +--+                                 
                                |  +-- Epioblasma_triquetra       
                                +--+                              
                                   +-- Ligumia_recta              

Serb et al., 2003 [Publ 2659], fig. 2

iT tT bi riS oT roT roS iD
5 27 18 4 24 24 3 57.0
                       +-- Lampsilis_siliquoidea  
  +--------------------+                          
  |                    +-- Lampsilis_ornata       
  |                                               
  |  +-------------------- Obliquaria_reflexa     
  |  |                                            
  |  +-------------------- Amblema_plicata        
  |  |                                            
  |  +-------------------- Reginaia_rotulata      
  |  |                                            
--+  +-------------------- Potamilus_alatus       
  |  |                                            
  |  |                 +-- Elliptoideus_sloatianus **
  |  +-----------------+                          
  |  |                 +-- Plectomerus_dombeyanus 
  |  |                                            
  |  |              +----- Medionidus_conradicus  
  |  +--------------+                             
  |  |              |  +-- Dromus_dromas          
  +--+              +--+                          
     |                 +-- Cyprogenia_aberti      
     |                                            
     |           +-------- Hemistena_lata **         
     +-----------+                                
     |           |  +----- Elliptio_complanata **    
     |           +--+                             
     |              |  +-- Fusconaia_burkei **       
     |              +--+                          
     |                 +-- Fusconaia_flava **        
     |                                            
     |  +----------------- Megalonaias_nervosa    
     |  |                                         
     |  |        +-------- Theliderma_cylindrica  
     |  |  +-----+                                
     +--+  |     |  +----- Theliderma_intermedia  
        |  |     +--+                             
        |  |        |  +-- Theliderma_sparsa      
        |  |        +--+                          
        |  |           +-- Theliderma_metanevra   
        +--+                                      
           |        +----- Quadrula_verrucosa     
           |  +-----+                             
           |  |     |  +-- Quadrula_quadrula      
           |  |     +--+                          
           |  |        +-- Quadrula_nobilis       
           +--+                                   
              |  +-------- Pustulosa_kieneriana   
              |  |                                
              |  +-------- Pustulosa_infucata     
              +--+                                
                 |     +-- Pustulosa_pustulosa    
                 |  +--+                          
                 |  |  +-- Pustulosa_succissa     
                 +--+                             
                    |  +-- Pustulosa_nodulata     
                    +--+                          
                       +-- Pustulosa_petrina      

Serb et al., 2003 [Publ 2659], fig. 3

iT tT bi riS oT roT roS iD
5 21 19 4 24 24 3 51.0
  +----------------------------------- Obliquaria_reflexa     
  |                                                           
  |  +-------------------------------- Amblema_plicata        
--+  |                                                        
  |  |                          +----- Reginaia_rotulata      
  |  |  +-----------------------+                             
  +--+  |                       |  +-- Elliptoideus_sloatianus **
     |  |                       +--+                          
     |  |                          +-- Plectomerus_dombeyanus 
     |  |                                                     
     +--+                    +-------- Hemistena_lata **         
        |  +-----------------+                                
        |  |                 |  +----- Elliptio_complanata **    
        |  |                 +--+                             
        |  |                    |  +-- Fusconaia_burkei **       
        |  |                    +--+                          
        +--+                       +-- Fusconaia_flava **        
           |                                                  
           |  +----------------------- Megalonaias_nervosa    
           |  |                                               
           |  |                    +-- Quadrula_verrucosa     
           +--+  +-----------------+                          
              |  |                 +-- Quadrula_nobilis       
              |  |                                            
              +--+  +----------------- Quadrula_quadrula      
                 |  |                                         
                 |  |        +-------- Theliderma_cylindrica  
                 +--+  +-----+                                
                    |  |     |  +----- Theliderma_sparsa      
                    |  |     +--+                             
                    |  |        |  +-- Theliderma_intermedia  
                    +--+        +--+                          
                       |           +-- Theliderma_metanevra   
                       |                                      
                       |  +----------- Pustulosa_kieneriana   
                       |  |                                   
                       +--+  +-------- Pustulosa_infucata     
                          |  |                                
                          +--+     +-- Pustulosa_succissa     
                             |  +--+                          
                             |  |  +-- Pustulosa_pustulosa    
                             +--+                             
                                |  +-- Pustulosa_nodulata     
                                +--+                          
                                   +-- Pustulosa_petrina      

Zanatta & Murphy, 2006 [Publ 2672], fig. 2

iT tT bi riS oT roT roS iD
5 5 2 5 52 50 3 44.0
                                            +-------- Pleuronaia_dolabelloides **   
                                            |                                    
  +-----------------------------------------+-------- Pleurobema_clava **           
  |                                         |                                    
  |                                         |  +----- Fusconaia_flava **            
  |                                         +--+                                 
  |                                            |  +-- Hemistena_lata **             
  |                                            +--+                              
  |                                               +-- Eurynia_dilatata **           
--+                                                                              
  |  +----------------------------------------------- Amblema_plicata            
  |  |                                                                           
  |  +----------------------------------------------- Popenaias_popeii           
  |  |                                                                           
  |  |                                         +----- Paetulunio_fabalis         
  |  |                                         |                                 
  |  |  +--------------------------------------+  +-- Toxolasma_parvum           
  +--+  |                                      |  |                              
     |  |                                      +--+-- Toxolasma_lividum          
     |  |                                         |                              
     |  |                                         +-- Toxolasma_texasiense       
     |  |                                                                        
     |  |                                      +----- Obliquaria_reflexa         
     |  |  +-----------------------------------+                                 
     +--+  |                                   |  +-- Glebula_rotundata          
        |  |                                   +--+                              
        |  |                                      +-- Cyrtonaias_tampicoensis    
        |  |                                                                     
        |  |                                   +----- Ptychobranchus_fasciolaris 
        |  |                                +--+                                 
        |  |                                |  |  +-- Dromus_dromas              
        |  |                                |  +--+                              
        +--+  +-----------------------------+     +-- Cyprogenia_stegaria        
           |  |                             |                                    
           |  |                             |  +----- Medionidus_conradicus      
           |  |                             +--+                                 
           |  |                                |  +-- Medionidus_acutissimus     
           |  |                                +--+                              
           |  |                                   +-- Lemiox_rimosus             
           |  |                                                                  
           |  |  +----------------------------------- Sagittunio_nasutus         
           |  |  |                                                               
           +--+  |                    +-------------- Potamilus_leptodon         
              |  |                    |                                          
              |  +--------------------+  +----------- Potamilus_fragilis         
              |  |                    |  |                                       
              |  |                    +--+        +-- Potamilus_alatus           
              |  |                       |  +-----+                              
              |  |                       |  |     +-- Potamilus_purpuratus       
              |  |                       +--+                                    
              |  |                          |  +----- Potamilus_inflatus         
              |  |                          +--+                                 
              +--+                             |  +-- Potamilus_ohiensis         
                 |                             +--+                              
                 |                                +-- Potamilus_amphichaenus     
                 |                                                               
                 |                                +-- Truncilla_truncata         
                 |  +-----------------------------+                              
                 |  |                             +-- Ellipsaria_lineolata       
                 |  |                                                            
                 |  |                             +-- Ortmanniana_pectorosa      
                 |  |                       +-----+                              
                 |  |                       |     +-- Lampsilis_fasciola         
                 |  |  +--------------------+                                    
                 +--+  |                    |  +----- Lampsilis_ornata           
                    |  |                    +--+                                 
                    |  |                       |  +-- Lampsilis_ovata            
                    |  |                       +--+                              
                    |  |                          +-- Lampsilis_cardium          
                    |  |                                                         
                    |  |                          +-- Ligumia_recta              
                    +--+                 +--------+                              
                       |                 |        +-- Cambarunio_iris            
                       |  +--------------+                                       
                       |  |              |  +-------- Lampsilis_teres            
                       |  |              |  |                                    
                       |  |              +--+     +-- Hamiota_australis          
                       |  |                 |  +--+                              
                       |  |                 |  |  +-- Hamiota_subangulata        
                       +--+                 +--+                                 
                          |                    |  +-- Hamiota_altilis            
                          |                    +--+                              
                          |                       +-- Hamiota_perovalis          
                          |                                                      
                          |  +----------------------- Leaunio_vanuxemensis       
                          |  |                                                   
                          +--+  +-------------------- Ortmanniana_ligamentina    
                             |  |                                                
                             +--+  +----------------- Lampsilis_siliquoidea      
                                |  |                                             
                                |  |  +-------------- Obovaria_olivaria          
                                +--+  |                                          
                                   |  +-------------- Venustaconcha_ellipsiformis
                                   |  |                                          
                                   |  |        +----- Obovaria_unicolor          
                                   +--+--------+                                 
                                      |        |  +-- Obovaria_subrotunda        
                                      |        +--+                              
                                      |           +-- Obovaria_arkansasensis     
                                      |                                          
                                      |  +----------- Epioblasma_brevidens       
                                      +--+                                       
                                         |  +-------- Epioblasma_triquetra       
                                         +--+                                    
                                            |  +----- Epioblasma_rangiana        
                                            +--+                                 
                                               |  +-- Epioblasma_walkeri         
                                               +--+                              
                                                  +-- Epioblasma_capsaeformis    

Huang et al., 2019 [Publ 3843], fig. 2

iT tT bi riS oT roT roS iD
5 5 3 5 199 20 3 42.0
                             +-- Megalonaias_nervosa        
                          +--+                              
                          |  +-- Uniomerus_carolinianus     
  +-----------------------+                                 
  |                       |  +-- Quadrula_quadrula          
  |                       +--+                              
  |                          +-- Quadrula_verrucosa         
  |                                                         
  |                    +-------- Elliptoideus_sloatianus **    
--+                    |                                    
  |  +-----------------+     +-- Elliptio_complanata **        
  |  |                 |  +--+                              
  |  |                 |  |  +-- Eurynia_dilatata **           
  |  |                 +--+                                 
  |  |                    |  +-- Fusconaia_flava **            
  +--+                    +--+                              
     |                       +-- Pleurobema_sintoxia **        
     |                                                      
     |  +----------------------- Amblema_plicata            
     |  |                                                   
     |  |                 +----- Toxolasma_parvum           
     +--+  +--------------+                                 
        |  |              |  +-- Obliquaria_reflexa         
        |  |              +--+                              
        |  |                 +-- Plectomerus_dombeyanus     
        +--+                                                
           |  +----------------- Potamilus_leptodon         
           |  |                                             
           |  |  +-------------- Potamilus_alatus           
           +--+  |                                          
              |  |  +----------- Venustaconcha_ellipsiformis
              |  |  |                                       
              +--+  +----------- Leaunio_lienosus           
                 |  |                                       
                 |  |        +-- Ellipsaria_lineolata       
                 |  +--------+                              
                 +--+        +-- Truncilla_truncata         
                    |                                       
                    |        +-- Medionidus_conradicus      
                    +--------+                              
                    |        +-- Ptychobranchus_subtentus   
                    |                                       
                    |  +-------- Cambarunio_iris            
                    +--+                                    
                       |  +----- Ortmanniana_ligamentina    
                       +--+                                 
                          |  +-- Lampsilis_cardium          
                          +--+                              
                             +-- Lampsilis_ornata           

Huang et al., 2019 [Publ 3843], fig. S3

iT tT bi riS oT roT roS iD
5 5 3 5 196 20 3 42.0
                                      +-- Megalonaias_nervosa        
                                   +--+                              
                                   |  +-- Uniomerus_carolinianus     
  +--------------------------------+                                 
  |                                |  +-- Quadrula_quadrula          
  |                                +--+                              
  |                                   +-- Quadrula_verrucosa         
  |                                                                  
  |                             +-------- Elliptoideus_sloatianus **    
--+                             |                                    
  |  +--------------------------+     +-- Elliptio_complanata **        
  |  |                          |  +--+                              
  |  |                          |  |  +-- Eurynia_dilatata **           
  |  |                          +--+                                 
  |  |                             |  +-- Fusconaia_flava **            
  +--+                             +--+                              
     |                                +-- Pleurobema_sintoxia **        
     |                                                               
     |  +-------------------------------- Amblema_plicata            
     |  |                                                            
     +--+  +----------------------------- Toxolasma_parvum           
        |  |                                                         
        +--+                          +-- Obliquaria_reflexa         
           |  +-----------------------+                              
           |  |                       +-- Plectomerus_dombeyanus     
           +--+                                                      
              |  +----------------------- Potamilus_leptodon         
              |  |                                                   
              +--+  +-------------------- Potamilus_alatus           
                 |  |                                                
                 |  |                 +-- Medionidus_conradicus      
                 +--+  +--------------+                              
                    |  |              +-- Ptychobranchus_subtentus   
                    |  |                                             
                    +--+              +-- Ellipsaria_lineolata       
                       |  +-----------+                              
                       |  |           +-- Truncilla_truncata         
                       +--+                                          
                          |  +----------- Leaunio_lienosus           
                          |  |                                       
                          +--+        +-- Venustaconcha_ellipsiformis
                             |  +-----+                              
                             |  |     +-- Cambarunio_iris            
                             +--+                                    
                                |  +----- Ortmanniana_ligamentina    
                                +--+                                 
                                   |  +-- Lampsilis_cardium          
                                   +--+                              
                                      +-- Lampsilis_ornata           

Huang et al., 2019 [Publ 3843], fig. S4

iT tT bi riS oT roT roS iD
5 5 3 5 197 20 3 40.0
  +-------------- Amblema_plicata            
  |                                          
  +-------------- Toxolasma_parvum           
  |                                          
  |           +-- Obliquaria_reflexa         
  +-----------+                              
  |           +-- Plectomerus_dombeyanus     
  |                                          
  |  +----------- Elliptoideus_sloatianus **    
  +--+                                       
  |  |  +-------- Elliptio_complanata **        
  |  +--+                                    
  |     |  +----- Eurynia_dilatata **           
--+     +--+                                 
  |        |  +-- Fusconaia_flava **            
  |        +--+                              
  |           +-- Pleurobema_sintoxia **        
  |                                          
  |     +-------- Ellipsaria_lineolata       
  |     |                                    
  |     +-------- Potamilus_leptodon         
  |     |                                    
  |     +-------- Medionidus_conradicus      
  |     |                                    
  |     +-------- Potamilus_alatus           
  |     |                                    
  |     +-------- Ptychobranchus_subtentus   
  +-----+                                    
        +-------- Truncilla_truncata         
        |                                    
        +-------- Venustaconcha_ellipsiformis
        |                                    
        +-------- Cambarunio_iris            
        |                                    
        +-------- Leaunio_lienosus           
        |                                    
        |  +----- Ortmanniana_ligamentina    
        +--+                                 
           |  +-- Lampsilis_cardium          
           +--+                              
              +-- Lampsilis_ornata           

Zanatta & Murphy, 2006 [Publ 2672], fig. 1

iT tT bi riS oT roT roS iD
4 4 0 4 52 50 3 42.0
                                               +-- Amblema_plicata            
                                            +--+                              
                                            |  +-- Popenaias_popeii           
                                            |                                 
  +-----------------------------------------+  +-- Pleurobema_clava **           
  |                                         |  |                              
  |                                         |  +-- Eurynia_dilatata **           
  |                                         +--+                              
  |                                            +-- Pleuronaia_dolabelloides **   
  |                                            |                              
--+                                            +-- Fusconaia_flava **            
  |                                                                           
  |                                            +-- Cyrtonaias_tampicoensis    
  |  +-----------------------------------------+                              
  |  |                                         +-- Glebula_rotundata          
  |  |                                                                        
  |  |  +----------------------------------------- Paetulunio_fabalis         
  +--+  |                                                                     
     |  |                                +-------- Obliquaria_reflexa         
     |  |  +-----------------------------+                                    
     |  |  |                             |  +----- Toxolasma_texasiense       
     |  |  |                             +--+                                 
     +--+  |                                |  +-- Toxolasma_parvum           
        |  |                                +--+                              
        |  |                                   +-- Toxolasma_lividum          
        |  |                                                                  
        |  |  +----------------------------------- Sagittunio_nasutus         
        |  |  |                                                               
        +--+  |                                +-- Medionidus_conradicus      
           |  |                          +-----+                              
           |  |                          |     +-- Ptychobranchus_fasciolaris 
           |  |                 +--------+                                    
           |  |                 |        |  +----- Medionidus_acutissimus     
           |  |                 |        |  |                                 
           |  |                 |        +--+----- Lemiox_rimosus             
           |  |                 |           |                                 
           |  |  +--------------+           |  +-- Dromus_dromas              
           +--+  |              |           +--+                              
              |  |              |              +-- Cyprogenia_stegaria        
              |  |              |                                             
              |  |              |  +-------------- Potamilus_leptodon         
              |  |              |  |                                          
              |  |              +--+  +----------- Potamilus_fragilis         
              |  |                 |  |                                       
              |  |                 +--+        +-- Potamilus_alatus           
              |  |                    |  +-----+                              
              |  |                    |  |     +-- Potamilus_purpuratus       
              |  |                    +--+                                    
              +--+                       |  +----- Potamilus_inflatus         
                 |                       +--+                                 
                 |                          |  +-- Potamilus_ohiensis         
                 |                          +--+                              
                 |                             +-- Potamilus_amphichaenus     
                 |                                                            
                 |                          +----- Lampsilis_teres            
                 |  +-----------------------+                                 
                 |  |                       |  +-- Truncilla_truncata         
                 |  |                       +--+                              
                 |  |                          +-- Ellipsaria_lineolata       
                 |  |                                                         
                 |  |  +-------------------------- Leaunio_vanuxemensis       
                 +--+  |                                                      
                    |  |                       +-- Hamiota_australis          
                    |  |                    +--+                              
                    |  |                    |  +-- Hamiota_subangulata        
                    |  |              +-----+                                 
                    |  |              |     |  +-- Hamiota_altilis            
                    |  |              |     +--+                              
                    +--+              |        +-- Hamiota_perovalis          
                       |  +-----------+                                       
                       |  |           |        +-- Lampsilis_fasciola         
                       |  |           |  +-----+                              
                       |  |           |  |     +-- Ortmanniana_pectorosa      
                       |  |           +--+                                    
                       |  |              |  +----- Lampsilis_ornata           
                       |  |              +--+                                 
                       |  |                 |  +-- Lampsilis_ovata            
                       +--+                 +--+                              
                          |                    +-- Lampsilis_cardium          
                          |                                                   
                          |              +-------- Lampsilis_siliquoidea      
                          |  +-----------+                                    
                          |  |           |  +----- Ortmanniana_ligamentina    
                          |  |           +--+                                 
                          |  |              |  +-- Cambarunio_iris            
                          |  |              +--+                              
                          +--+                 +-- Ligumia_recta              
                             |                                                
                             |  +----------------- Venustaconcha_ellipsiformis
                             |  |                                             
                             |  |        +-------- Obovaria_olivaria          
                             +--+  +-----+                                    
                                |  |     |  +----- Obovaria_unicolor          
                                |  |     +--+                                 
                                |  |        |  +-- Obovaria_subrotunda        
                                +--+        +--+                              
                                   |           +-- Obovaria_arkansasensis     
                                   |                                          
                                   |  +----------- Epioblasma_brevidens       
                                   +--+                                       
                                      |  +-------- Epioblasma_triquetra       
                                      +--+                                    
                                         |  +----- Epioblasma_rangiana        
                                         +--+                                 
                                            |  +-- Epioblasma_walkeri         
                                            +--+                              
                                               +-- Epioblasma_capsaeformis    

Krebs et al., 2003 [Publ 2691], fig. 3

iT tT bi riS oT roT roS iD
4 5 3 4 30 18 3 43.0
                    +-------- Megalonaias_nervosa     
  +-----------------+                                 
  |                 |  +----- Quadrula_quadrula[2]    
  |                 +--+                              
  |                    |  +-- Quadrula_quadrula[1]    
  |                    +--+                           
  |                       +-- Theliderma_metanevra    
  |                                                   
--+              +----------- Elliptoideus_sloatianus ** 
  |  +-----------+                                    
  |  |           |  +-------- Pleurobema_pyriforme **    
  |  |           +--+                                 
  |  |              |  +----- Uniomerus_carolinianus  
  |  |              +--+                              
  |  |                 |  +-- Fusconaia_flava **         
  +--+                 +--+                           
     |                    +-- Eurynia_dilatata **        
     |                                                
     |  +-------------------- Amblema_plicata         
     |  |                                             
     |  |              +----- Toxolasma_parvum        
     |  |  +-----------+                              
     +--+  |           |  +-- Plectomerus_dombeyanus  
        |  |           +--+                           
        |  |              +-- Obliquaria_reflexa      
        |  |                                          
        +--+           +----- Ptychobranchus_subtentus
           |  +--------+                              
           |  |        |  +-- Potamilus_fragilis      
           |  |        +--+                           
           |  |           +-- Medionidus_conradicus   
           +--+                                       
              |           +-- Villosa_delumbis        
              |  +--------+                           
              |  |        +-- Ellipsaria_lineolata    
              +--+                                    
                 |  +-------- Lampsilis_teres         
                 +--+                                 
                    |  +----- Lampsilis_siliquoidea   
                    +--+                              
                       |  +-- Potamilus_purpuratus    
                       +--+                           
                          +-- Potamilus_alatus        

Boyer et al., 2011 [Publ 2718], fig. 3

iT tT bi riS oT roT roS iD
4 4 2 4 31 25 3 42.0
                       +-- Fusconaia_flava **        
                    +--+                          
                    |  +-- Pleurobema_sintoxia **    
     +--------------+                             
     |              |  +-- Eurynia_dilatata **       
     |              +--+                          
  +--+                 +-- Elliptio_crassidens **    
  |  |                                            
  |  |  +----------------- Megalonaias_nervosa    
  |  +--+                                         
  |     |  +-------------- Quadrula_quadrula      
  |     +--+                                      
  |        |  +----------- Quadrula_verrucosa     
  |        +--+                                   
  |           |  +-------- Theliderma_metanevra   
--+           +--+                                
  |              |  +----- Pustulosa_pustulosa    
  |              +--+                             
  |                 |  +-- Cyclonaias_tuberculata 
  |                 +--+                          
  |                    +-- Pustulosa_nodulata     
  |                                               
  |     +----------------- Amblema_plicata        
  |     |                                         
  |     |  +-------------- Toxolasma_parvum       
  +-----+  |                                      
        |  |  +----------- Obliquaria_reflexa     
        |  |  |                                   
        +--+  |  +-------- Obovaria_olivaria      
           |  |  |                                
           |  |  +-------- Lampsilis_cardium      
           |  |  |                                
           +--+  |     +-- Epioblasma_triquetra   
              |  +-----+                          
              |  |     +-- Lampsilis_siliquoidea  
              |  |                                
              |  |     +-- Ortmanniana_ligamentina
              |  +-----+                          
              |  |     +-- Lampsilis_higginsii    
              +--+                                
                 |     +-- Ligumia_recta          
                 +-----+                          
                 |     +-- Lampsilis_teres        
                 |                                
                 |  +----- Potamilus_fragilis     
                 +--+                             
                 |  |  +-- Potamilus_ohiensis     
                 |  +--+                          
                 |     +-- Potamilus_alatus       
                 |                                
                 |  +----- Ellipsaria_lineolata   
                 +--+                             
                    |  +-- Truncilla_truncata     
                    +--+                          
                       +-- Truncilla_donaciformis 

Boyer et al., 2011 [Publ 2718], fig. 4

iT tT bi riS oT roT roS iD
4 4 2 4 34 25 3 44.0
  +----------------------- Amblema_plicata        
  |                                               
  |                    +-- Reginaia_ebenus        
  |  +-----------------+                          
  |  |                 +-- Obliquaria_reflexa     
--+  |                                            
  |  |           +-------- Pleurobema_sintoxia **    
  |  +-----------+                                
  |  |           |  +----- Elliptio_crassidens **    
  |  |           +--+                             
  |  |              |  +-- Fusconaia_flava **        
  +--+              +--+                          
     |                 +-- Eurynia_dilatata **       
     |                                            
     |              +----- Potamilus_fragilis     
     |  +-----------+                             
     |  |           |  +-- Potamilus_ohiensis     
     |  |           +--+                          
     |  |              +-- Potamilus_alatus       
     |  |                                         
     +--+           +----- Ellipsaria_lineolata   
        |  +--------+                             
        |  |        |  +-- Truncilla_donaciformis 
        |  |        +--+                          
        |  |           +-- Truncilla_truncata     
        +--+                                      
           |  +----------- Lampsilis_cardium      
           |  |                                   
           |  |     +----- Ligumia_recta          
           |  +-----+                             
           +--+     |  +-- Obovaria_olivaria      
              |     +--+                          
              |        +-- Epioblasma_triquetra   
              |                                   
              |  +-------- Lampsilis_teres        
              +--+                                
                 |  +----- Lampsilis_siliquoidea  
                 +--+                             
                    |  +-- Ortmanniana_ligamentina
                    +--+                          
                       +-- Lampsilis_higginsii    

Johnson et al., 2023 [Publ 4323], fig. 3

iT tT bi riS oT roT roS iD
4 4 2 2 0 0 0 44.0
  +-------- Fusconaia_cerina **      
--+                               
  |  +----- Pleurobema_riddellii **  
  +--+                            
     |  +-- Pleurobema_beadleianum **
     +--+                         
        +-- Pleurobema_clava **      

Graf & Cummings, 2006 [Publ 1968], fig. 2

iT tT bi riS oT roT roS iD
3 3 1 3 44 11 3 41.0
  +-------------------- Amblema_plicata           
  |                                               
  |              +----- Eurynia_dilatata **          
--+  +-----------+                                
  |  |           |  +-- Fusconaia_flava **           
  |  |           +--+                             
  |  |              +-- Pleurobema_sintoxia **       
  +--+                                            
     |  +-------------- Obliquaria_reflexa        
     |  |                                         
     |  +-------------- Truncilla_truncata        
     +--+                                         
        +-------------- Ptychobranchus_fasciolaris
        |                                         
        |  +----------- Ligumia_recta             
        +--+                                      
           |  +-------- Cambarunio_iris           
           +--+                                   
              |  +----- Epioblasma_triquetra      
              +--+                                
                 |  +-- Lampsilis_cardium         
                 +--+                             
                    +-- Ortmanniana_ligamentina   

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

iT tT bi riS oT roT roS iD
3 3 1 3 27 14 3 41.0
           +-------- Quadrula_quadrula         
  +--------+                                   
  |        |  +----- Eurynia_dilatata **          
  |        +--+                                
  |           |  +-- Pleurobema_sintoxia **       
  |           +--+                             
--+              +-- Fusconaia_flava **           
  |                                            
  |  +-------------- Amblema_plicata           
  |  |                                         
  |  |  +----------- Ligumia_recta             
  |  |  |                                      
  +--+  +----------- Truncilla_truncata        
     |  |                                      
     |  |        +-- Ptychobranchus_fasciolaris
     |  +--------+                             
     +--+        +-- Sagittunio_nasutus        
        |                                      
        |     +----- Lampsilis_siliquoidea     
        |  +--+                                
        |  |  |  +-- Epioblasma_triquetra      
        |  |  +--+                             
        |  |     +-- Epioblasma_brevidens      
        +--+                                   
           |  +----- Ortmanniana_ligamentina   
           |  |                                
           |  +----- Cambarunio_iris           
           +--+                                
              +----- Leaunio_vanuxemensis      
              |                                
              |  +-- Lampsilis_cardium         
              +--+                             
                 +-- Lampsilis_fasciola        

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

iT tT bi riS oT roT roS iD
3 3 1 3 27 14 3 41.0
                    +-------- Quadrula_quadrula         
  +-----------------+                                   
  |                 |  +----- Eurynia_dilatata **          
  |                 +--+                                
  |                    |  +-- Pleurobema_sintoxia **       
--+                    +--+                             
  |                       +-- Fusconaia_flava **           
  |                                                     
  |  +----------------------- Amblema_plicata           
  |  |                                                  
  +--+  +-------------------- Truncilla_truncata        
     |  |                                               
     +--+  +----------------- Ligumia_recta             
        |  |                                            
        |  |              +-- Ptychobranchus_fasciolaris
        +--+  +-----------+                             
           |  |           +-- Sagittunio_nasutus        
           |  |                                         
           +--+  +----------- Lampsilis_siliquoidea     
              |  |                                      
              |  |     +----- Ortmanniana_ligamentina   
              +--+  +--+                                
                 |  |  |  +-- Lampsilis_cardium         
                 |  |  +--+                             
                 |  |     +-- Lampsilis_fasciola        
                 +--+                                   
                    |     +-- Epioblasma_brevidens      
                    |  +--+                             
                    |  |  +-- Epioblasma_triquetra      
                    +--+                                
                       |  +-- Cambarunio_iris           
                       +--+                             
                          +-- Leaunio_vanuxemensis      

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

iT tT bi riS oT roT roS iD
3 3 0 3 31 8 3 42.0
  +-------- Toxolasma_lividum      
  |                                
  +-------- Amblema_plicata        
  |                                
  |     +-- Glebula_rotundata      
  +-----+                          
  |     +-- Cyrtonaias_tampicoensis
  |                                
--+  +----- Potamilus_alatus       
  +--+                             
  |  |  +-- Ortmanniana_ligamentina
  |  +--+                          
  |     +-- Ligumia_recta          
  |                                
  |  +----- Quadrula_quadrula      
  |  |                             
  +--+  +-- Eurynia_dilatata **       
     |  |                          
     +--+-- Pleurobema_clava **       
        |                          
        +-- Fusconaia_flava **        

Hoeh et al., 2001 [Publ 2652], fig. 14.2

iT tT bi riS oT roT roS iD
3 28 16 3 28 8 3 59.0
  +-------------------- Amblema_plicata        
  |                                            
  +-------------------- Coelatura_aegyptiaca   
  |                                            
  +-------------------- Eurynia_dilatata **       
  |                                            
  +-------------------- Fusconaia_flava **        
  |                                            
  +-------------------- Gonidea_angulata       
  |                                            
  +-------------------- Pleurobema_clava **       
  |                                            
  +-------------------- Quadrula_quadrula      
  |                                            
--+        +----------- Ortmanniana_ligamentina
  +--------+                                   
  |        |  +-------- Glebula_rotundata      
  |        +--+                                
  |           |  +----- Potamilus_alatus       
  |           +--+                             
  |              |  +-- Ligumia_recta          
  |              +--+                          
  |                 +-- Toxolasma_lividum      
  |                                            
  |  +----------------- Cyrtonaias_tampicoensis
  |  |                                         
  |  |              +-- Unio_pictorum          
  |  |  +-----------+                          
  +--+  |           +-- Unio_tumidus           
     |  |                                      
     |  |        +----- Pyganodon_grandis      
     |  |  +-----+                             
     +--+  |     |  +-- Anodonta_cygnea        
        |  |     +--+                          
        |  |        +-- Strophitus_undulatus   
        |  |                                   
        |  |     +----- Diplodon_gratus        
        +--+     |                             
           |     +----- Echyridella_menziesii  
           |  +--+                             
           |  |  +----- Lortiella_rugata       
           |  |  |                             
           |  |  |  +-- Castalia_stevensi      
           |  |  +--+                          
           +--+     +-- Velesunio_angasi       
              |                                
              |     +-- Mutela_dubia           
              |  +--+                          
              |  |  +-- Mutela_rostrata        
              +--+                             
                 |  +-- Anodontites_guanarensis
                 |  |                          
                 +--+-- Anodontites_trigona    
                    |                          
                    +-- Monocondylaea_minuana  

Hoeh et al., 2001 [Publ 2652], fig. 14.4

iT tT bi riS oT roT roS iD
3 3 1 3 28 8 3 41.0
        +-------- Quadrula_quadrula      
  +-----+                                
  |     |  +----- Eurynia_dilatata **       
  |     +--+                             
  |        |  +-- Fusconaia_flava **        
  |        +--+                          
--+           +-- Pleurobema_clava **       
  |                                      
  |           +-- Cyrtonaias_tampicoensis
  |  +--------+                          
  |  |        +-- Glebula_rotundata      
  +--+                                   
     |  +-------- Toxolasma_lividum      
     +--+                                
        |  +----- Potamilus_alatus       
        +--+                             
           |  +-- Ligumia_recta          
           +--+                          
              +-- Ortmanniana_ligamentina

Hoeh et al., 2002 [Publ 2653], fig. 2

iT tT bi riS oT roT roS iD
3 3 1 3 28 8 3 41.0
  +----------- Amblema_plicata  
--+                             
  |  +-------- Quadrula_quadrula
  +--+                          
     |  +----- Eurynia_dilatata ** 
     +--+                       
        |  +-- Pleurobema_clava ** 
        +--+                    
           +-- Fusconaia_flava **  

Roe & Hoeh, 2003 [Publ 2655], fig. 4.2

iT tT bi riS oT roT roS iD
3 16 12 3 28 8 3 52.0
  +----------------------- Fusconaia_flava **        
  |                                               
  +----------------------- Amblema_plicata        
--+                                               
  |  +-------------------- Quadrula_quadrula      
  |  |                                            
  |  |  +----------------- Pleurobema_clava **       
  +--+  |                                         
     |  |  +-------------- Eurynia_dilatata **       
     |  |  |                                      
     +--+  |  +----------- Ortmanniana_ligamentina
        |  +--+                                   
        |  |  |  +-------- Glebula_rotundata      
        |  |  +--+                                
        |  |     |  +----- Potamilus_alatus       
        +--+     +--+                             
           |        |  +-- Ligumia_recta          
           |        +--+                          
           |           +-- Toxolasma_lividum      
           |                                      
           |  +----------- Cyrtonaias_tampicoensis
           |  |                                   
           +--+        +-- Unio_pictorum          
              |  +-----+                          
              |  |     +-- Unio_tumidus           
              +--+                                
                 |  +----- Pyganodon_grandis      
                 +--+                             
                    |  +-- Anodonta_cygnea        
                    +--+                          
                       +-- Strophitus_undulatus   

Roe & Hoeh, 2003 [Publ 2655], fig. 4.3

iT tT bi riS oT roT roS iD
3 3 1 3 28 8 3 41.0
        +-------- Quadrula_quadrula      
  +-----+                                
  |     |  +----- Eurynia_dilatata **       
  |     +--+                             
  |        |  +-- Fusconaia_flava **        
  |        +--+                          
--+           +-- Pleurobema_clava **       
  |                                      
  |           +-- Cyrtonaias_tampicoensis
  |  +--------+                          
  |  |        +-- Glebula_rotundata      
  +--+                                   
     |  +-------- Toxolasma_lividum      
     +--+                                
        |  +----- Ortmanniana_ligamentina
        +--+                             
           |  +-- Ligumia_recta          
           +--+                          
              +-- Potamilus_alatus       

Lydeard et al., 1996 [Publ 2656], fig. 3

iT tT bi riS oT roT roS iD
3 4 1 3 24 15 3 42.0
                       +----- Megalonaias_nervosa     
  +--------------------+                              
  |                    |  +-- Quadrula_quadrula       
  |                    +--+                           
  |                       +-- Theliderma_metanevra    
  |                                                   
--+                    +----- Elliptoideus_sloatianus ** 
  |                    |                              
  |  +-----------------+  +-- Eurynia_dilatata **        
  |  |                 |  |                           
  |  |                 +--+-- Pleurobema_pyriforme **    
  |  |                    |                           
  +--+                    +-- Uniomerus_carolinianus  
     |                                                
     |  +-------------------- Amblema_plicata         
     |  |                                             
     +--+                 +-- Plectomerus_dombeyanus  
        |  +--------------+                           
        |  |              +-- Obliquaria_reflexa      
        +--+                                          
           |  +-------------- Potamilus_fragilis      
           |  |                                       
           +--+  +----------- Ptychobranchus_subtentus
              |  |                                    
              +--+  +-------- Medionidus_conradicus   
                 |  |                                 
                 +--+  +----- Potamilus_purpuratus    
                    |  |                              
                    |  |  +-- Ellipsaria_lineolata    
                    +--+--+                           
                       |  +-- Villosa_delumbis        
                       |                              
                       |  +-- Lampsilis_teres         
                       +--+                           
                          +-- Lampsilis_siliquoidea   

Lydeard et al., 1996 [Publ 2656], fig. 4

iT tT bi riS oT roT roS iD
3 4 2 3 25 15 3 42.0
                                +----- Megalonaias_nervosa     
  +-----------------------------+                              
  |                             |  +-- Quadrula_quadrula       
  |                             +--+                           
  |                                +-- Theliderma_metanevra    
  |                                                            
--+                          +-------- Elliptoideus_sloatianus ** 
  |  +-----------------------+                                 
  |  |                       |  +----- Uniomerus_carolinianus  
  |  |                       +--+                              
  |  |                          |  +-- Eurynia_dilatata **        
  +--+                          +--+                           
     |                             +-- Pleurobema_pyriforme **    
     |                                                         
     |  +----------------------------- Amblema_plicata         
     +--+                                                      
        |  +-------------------------- Plectomerus_dombeyanus  
        +--+                                                   
           |  +----------------------- Obliquaria_reflexa      
           +--+                                                
              |  +-------------------- Medionidus_conradicus   
              +--+                                             
                 |  +----------------- Ptychobranchus_subtentus
                 +--+                                          
                    |  +-------------- Potamilus_fragilis      
                    +--+                                       
                       |  +----------- Potamilus_purpuratus    
                       +--+                                    
                          |  +-------- Ellipsaria_lineolata    
                          +--+                                 
                             |  +----- Villosa_delumbis        
                             +--+                              
                                |  +-- Lampsilis_teres         
                                +--+                           
                                   +-- Lampsilis_siliquoidea   

Hoeh et al., 2009 [Publ 2716], fig. 3

iT tT bi riS oT roT roS iD
3 3 1 3 44 11 3 41.0
  +----------- Amblema_plicata    
  |                               
  |     +----- Obliquaria_reflexa 
--+  +--+                         
  |  |  |  +-- Quadrula_verrucosa 
  |  |  +--+                      
  +--+     +-- Quadrula_quadrula  
     |                            
     |  +----- Eurynia_dilatata **   
     +--+                         
        |  +-- Pleurobema_sintoxia **
        +--+                      
           +-- Fusconaia_flava **    

Hoeh et al., 2009 [Publ 2716], fig. 4

iT tT bi riS oT roT roS iD
3 3 0 3 43 10 2 42.0
                    +-- Eurynia_dilatata **          
                    |                             
              +-----+-- Pleurobema_sintoxia **       
              |     |                             
  +-----------+     +-- Fusconaia_flava **           
  |           |                                   
  |           |  +----- Obliquaria_reflexa        
  |           +--+                                
--+              |  +-- Quadrula_verrucosa        
  |              +--+                             
  |                 +-- Quadrula_quadrula         
  |                                               
  |  +----------------- Ptychobranchus_fasciolaris
  +--+                                            
     |  +-------------- Truncilla_truncata        
     +--+                                         
        |  +----------- Ortmanniana_ligamentina   
        +--+                                      
           |  +-------- Lampsilis_cardium         
           +--+                                   
              |  +----- Cambarunio_iris           
              +--+                                
                 |  +-- Ligumia_recta             
                 +--+                             
                    +-- Epioblasma_triquetra      

Hoeh et al., 2009 [Publ 2716], fig. 5

iT tT bi riS oT roT roS iD
3 3 1 3 43 10 3 41.0
        +----- Obliquaria_reflexa        
  +-----+                                
  |     |  +-- Quadrula_verrucosa        
  |     +--+                             
  |        +-- Quadrula_quadrula         
  |                                      
--+  +-------- Amblema_plicata           
  |  |                                   
  |  |  +----- Eurynia_dilatata **          
  |  +--+                                
  |  |  |  +-- Pleurobema_sintoxia **       
  +--+  +--+                             
     |     +-- Fusconaia_flava **           
     |                                   
     |  +----- Truncilla_truncata        
     |  |                                
     |  +----- Ptychobranchus_fasciolaris
     +--+                                
        |  +-- Ortmanniana_ligamentina   
        |  |                             
        |  +-- Cambarunio_iris           
        +--+                             
           +-- Ligumia_recta             
           |                             
           +-- Epioblasma_triquetra      

Rosenberg et al., 1994 [Publ 2719], fig. 3

iT tT bi riS oT roT roS iD
3 5 0 3 16 8 3 44.0
  +-- Elliptio_complanata **   
  |                         
  +-- Fusconaia_cerina **      
  |                         
--+-- Plectomerus_dombeyanus
  |                         
  +-- Pleurobema_cordatum **   
  |                         
  +-- Unio_pictorum         

Rosenberg et al., 1997 [Publ 2720], fig. 2A

iT tT bi riS oT roT roS iD
3 17 0 3 20 9 3 49.0
  +-- Gonidea_angulata      
  |                         
  +-- Pyganodon_grandis     
  |                         
  +-- Utterbackia_imbecillis
  |                         
  +-- Pyganodon_cataracta   
  |                         
  +-- Lampsilis_straminea   
  |                         
  +-- Lampsilis_teres       
  |                         
  +-- Megalonaias_nervosa   
  |                         
  +-- Obliquaria_reflexa    
  |                         
--+-- Theliderma_cylindrica 
  |                         
  +-- Quadrula_quadrula     
  |                         
  +-- Uniomerus_tetralasmus 
  |                         
  +-- Amblema_plicata       
  |                         
  +-- Elliptio_complanata **   
  |                         
  +-- Fusconaia_cerina **      
  |                         
  +-- Plectomerus_dombeyanus
  |                         
  +-- Pleurobema_cordatum **   
  |                         
  +-- Unio_pictorum         

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iT tT bi riS oT roT roS iD
3 3 1 3 78 6 3 42.0
     +-------- Amblema_plicata        
  +--+                                
  |  |  +----- Cambarunio_iris        
  |  +--+                             
  |     |  +-- Lampsilis_cardium      
  |     +--+                          
--+        +-- Ortmanniana_ligamentina
  |                                   
  |        +-- Quadrula_quadrula      
  |  +-----+                          
  |  |     +-- Quadrula_verrucosa     
  +--+                                
     |  +----- Elliptio_complanata **    
     +--+                             
        |  +-- Pleurobema_sintoxia **    
        +--+                          
           +-- Eurynia_dilatata **       

Bolotov et al., 2017 [Publ 3761], fig. S1

iT tT bi riS oT roT roS iD
3 3 1 3 138 6 3 44.0
     +-------- Amblema_plicata        
  +--+                                
  |  |  +----- Ortmanniana_ligamentina
  |  +--+                             
  |     |  +-- Lampsilis_cardium      
  |     +--+                          
--+        +-- Cambarunio_iris        
  |                                   
  |        +-- Quadrula_quadrula      
  |  +-----+                          
  |  |     +-- Quadrula_verrucosa     
  +--+                                
     |  +----- Pleurobema_sintoxia **    
     +--+                             
        |  +-- Eurynia_dilatata **       
        +--+                          
           +-- Elliptio_complanata **    

Bolotov et al., 2017 [Publ 3761], fig. S2

iT tT bi riS oT roT roS iD
3 3 1 3 138 6 3 44.0
     +-------- Amblema_plicata        
  +--+                                
  |  |  +----- Cambarunio_iris        
  |  +--+                             
  |     |  +-- Ortmanniana_ligamentina
  |     +--+                          
--+        +-- Lampsilis_cardium      
  |                                   
  |        +-- Quadrula_quadrula      
  |  +-----+                          
  |  |     +-- Quadrula_verrucosa     
  +--+                                
     |  +----- Pleurobema_sintoxia **    
     +--+                             
        |  +-- Elliptio_complanata **    
        +--+                          
           +-- Eurynia_dilatata **       

Bolotov et al., 2017 [Publ 3764], fig. S1

iT tT bi riS oT roT roS iD
3 9 0 3 128 6 3 48.0
  +-- Amblema_plicata        
  |                          
  +-- Ortmanniana_ligamentina
  |                          
  +-- Lampsilis_cardium      
  |                          
  +-- Cambarunio_iris        
  |                          
--+-- Elliptio_complanata **    
  |                          
  +-- Eurynia_dilatata **       
  |                          
  +-- Pleurobema_sintoxia **    
  |                          
  +-- Quadrula_quadrula      
  |                          
  +-- Quadrula_verrucosa     

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iT tT bi riS oT roT roS iD
3 3 1 3 54 12 4 43.0
                       +-- Quadrula_quadrula      
  +--------------------+                          
  |                    +-- Uniomerus_tetralasmus  
  |                                               
  |                 +----- Elliptoideus_sloatianus **
--+  +--------------+                             
  |  |              |  +-- Elliptio_complanata **    
  |  |              +--+                          
  |  |                 +-- Pleurobema_strodeanum **  
  |  |                                            
  +--+              +----- Reginaia_ebenus        
     |  +-----------+                             
     |  |           |  +-- Plectomerus_dombeyanus 
     |  |           +--+                          
     |  |              +-- Amblema_plicata        
     +--+                                         
        |              +-- Psoronaias_semigranosa 
        |  +-----------+                          
        |  |           +-- Popenaias_popeii       
        +--+                                      
           |  +----------- Obliquaria_reflexa     
           +--+                                   
              |  +-------- Cyrtonaias_tampicoensis
              +--+                                
                 |  +----- Leaunio_lienosus       
                 +--+                             
                    |  +-- Lampsilis_cardium      
                    +--+                          
                       +-- Pachynaias_spheniopsis 

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iT tT bi riS oT roT roS iD
3 3 1 3 54 12 4 43.0
              +----- Elliptoideus_sloatianus **
           +--+                             
           |  |  +-- Elliptio_complanata **    
           |  +--+                          
  +--------+     +-- Pleurobema_strodeanum **  
  |        |                                
  |        |  +----- Reginaia_ebenus        
  |        +--+                             
  |           |  +-- Plectomerus_dombeyanus 
--+           +--+                          
  |              +-- Amblema_plicata        
  |                                         
  |              +-- Psoronaias_semigranosa 
  |  +-----------+                          
  |  |           +-- Popenaias_popeii       
  +--+                                      
     |  +----------- Obliquaria_reflexa     
     +--+                                   
        |  +-------- Cyrtonaias_tampicoensis
        +--+                                
           |  +----- Leaunio_lienosus       
           +--+                             
              |  +-- Lampsilis_cardium      
              +--+                          
                 +-- Pachynaias_spheniopsis 

Jeratthitikul et al., 2022 [Publ 4260], fig. S1, 1

iT tT bi riS oT roT roS iD
3 3 1 3 73 6 3 42.0
     +-------- Amblema_plicata        
  +--+                                
  |  |  +----- Ortmanniana_ligamentina
  |  +--+                             
  |     |  +-- Cambarunio_iris        
  |     +--+                          
--+        +-- Lampsilis_cardium      
  |                                   
  |        +-- Quadrula_quadrula      
  |  +-----+                          
  |  |     +-- Quadrula_verrucosa     
  +--+                                
     |  +----- Elliptio_complanata **    
     +--+                             
        |  +-- Pleurobema_sintoxia **    
        +--+                          
           +-- Eurynia_dilatata **       

Jeratthitikul et al., 2022 [Publ 4260], fig. S2, 2

iT tT bi riS oT roT roS iD
3 3 1 3 73 6 3 44.0
     +-------- Amblema_plicata        
  +--+                                
  |  |  +----- Ortmanniana_ligamentina
  |  +--+                             
  |     |  +-- Lampsilis_cardium      
  |     +--+                          
--+        +-- Cambarunio_iris        
  |                                   
  |        +-- Quadrula_quadrula      
  |  +-----+                          
  |  |     +-- Quadrula_verrucosa     
  +--+                                
     |  +----- Pleurobema_sintoxia **    
     +--+                             
        |  +-- Eurynia_dilatata **       
        +--+                          
           +-- Elliptio_complanata **    

Huang et al., 2002 [Publ 2657], fig. 1

iT tT bi riS oT roT roS iD
2 3 0 2 32 12 3 43.0
  +----------------------------- Megalonaias_nervosa     
  |                                                      
  |                       +----- Elliptoideus_sloatianus ** 
--+  +--------------------+                              
  |  |                    |  +-- Uniomerus_carolinianus  
  |  |                    +--+                           
  +--+                       +-- Pleurobema_pyriforme **    
     |                                                   
     |  +----------------------- Amblema_plicata         
     |  |                                                
     +--+                    +-- Obliquaria_reflexa      
        |  +-----------------+                           
        |  |                 +-- Plectomerus_dombeyanus  
        +--+                                             
           |  +----------------- Medionidus_conradicus   
           +--+                                          
              |  +-------------- Potamilus_fragilis      
              +--+                                       
                 |  +----------- Ptychobranchus_subtentus
                 +--+                                    
                    |  +-------- Potamilus_purpuratus    
                    +--+                                 
                       |  +----- Lampsilis_teres         
                       +--+                              
                          |  +-- Villosa_delumbis        
                          +--+                           
                             +-- Ellipsaria_lineolata    

Huang et al., 2002 [Publ 2657], fig. 2

iT tT bi riS oT roT roS iD
2 3 0 2 32 12 3 43.0
  +----------- Megalonaias_nervosa     
  |                                    
  |     +----- Elliptoideus_sloatianus ** 
  |  +--+                              
--+  |  |  +-- Uniomerus_carolinianus  
  |  |  +--+                           
  |  |     +-- Pleurobema_pyriforme **    
  |  |                                 
  |  |  +----- Potamilus_fragilis      
  +--+  |                              
     |  +----- Medionidus_conradicus   
     |  |                              
     |  +----- Amblema_plicata         
     |  |                              
     |  |  +-- Obliquaria_reflexa      
     +--+--+                           
        |  +-- Plectomerus_dombeyanus  
        |                              
        |  +-- Villosa_delumbis        
        |  |                           
        |  +-- Ellipsaria_lineolata    
        |  |                           
        +--+-- Lampsilis_teres         
           |                           
           +-- Potamilus_purpuratus    
           |                           
           +-- Ptychobranchus_subtentus

Chapman et al., 2008 [Publ 2667], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 19 18 4 --
                                         +-- Quadrula_quadrula          
  +--------------------------------------+                              
  |                                      +-- Pustulosa_pustulosa        
  |                                                                     
--+                                      +-- Pleurobema_sintoxia **        
  |  +-----------------------------------+                              
  |  |                                   +-- Fusconaia_flava **            
  +--+                                                                  
     |  +----------------------------------- Amblema_plicata            
     |  |                                                               
     +--+  +-------------------------------- Plectomerus_dombeyanus     
        |  |                                                            
        +--+  +----------------------------- Popenaias_popeii           
           |  |                                                         
           +--+  +-------------------------- Toxolasma_lividum          
              |  |                                                      
              +--+  +----------------------- Obliquaria_reflexa         
                 |  |                                                   
                 |  |                    +-- Cyrtonaias_tampicoensis    
                 +--+  +-----------------+                              
                    |  |                 +-- Glebula_rotundata          
                    |  |                                                
                    +--+                 +-- Ptychobranchus_fasciolaris 
                       |  +--------------+                              
                       |  |              +-- Lemiox_rimosus             
                       |  |                                             
                       +--+              +-- Venustaconcha_ellipsiformis
                          |  +-----------+                              
                          |  |           +-- Obovaria_olivaria          
                          +--+                                          
                             |  +----------- Hamiota_subangulata        
                             +--+                                       
                                |  +-------- Lampsilis_ovata            
                                +--+                                    
                                   |  +----- Ortmanniana_ligamentina    
                                   +--+                                 
                                      |  +-- Lampsilis_straminea        
                                      +--+                              
                                         +-- Lampsilis_hydiana          

Manendo et al., 2008 [Publ 2669], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 11 3 2 --
        +-- Lampsilis_siliquoidea   
  +-----+                           
  |     +-- Sagittunio_nasutus      
--+                                 
  |  +----- Amblema_plicata         
  +--+                              
     |  +-- Pleuronaia_dolabelloides **
     +--+                           
        +-- Fusconaia_flava **         

Rosenberg et al., 1997 [Publ 2720], fig. 2B

iT tT bi riS oT roT roS iD
2 17 0 2 21 9 3 49.0
  +-- Gonidea_angulata      
  |                         
  +-- Pyganodon_grandis     
  |                         
  +-- Utterbackia_imbecillis
  |                         
  +-- Pyganodon_cataracta   
  |                         
  +-- Lampsilis_straminea   
  |                         
  +-- Lampsilis_teres       
  |                         
  +-- Megalonaias_nervosa   
  |                         
  +-- Obliquaria_reflexa    
  |                         
--+-- Theliderma_cylindrica 
  |                         
  +-- Quadrula_quadrula     
  |                         
  +-- Uniomerus_tetralasmus 
  |                         
  +-- Amblema_plicata       
  |                         
  +-- Elliptio_complanata **   
  |                         
  +-- Fusconaia_cerina **      
  |                         
  +-- Plectomerus_dombeyanus
  |                         
  +-- Aculamprotula_tortuosa
  |                         
  +-- Unio_pictorum         

Chapman et al., 2008 [Publ 2744], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 19 18 4 --
                                         +-- Quadrula_quadrula          
  +--------------------------------------+                              
  |                                      +-- Pustulosa_pustulosa        
  |                                                                     
--+                                      +-- Pleurobema_sintoxia **        
  |  +-----------------------------------+                              
  |  |                                   +-- Fusconaia_flava **            
  +--+                                                                  
     |  +----------------------------------- Amblema_plicata            
     |  |                                                               
     +--+  +-------------------------------- Plectomerus_dombeyanus     
        |  |                                                            
        +--+  +----------------------------- Popenaias_popeii           
           |  |                                                         
           +--+  +-------------------------- Toxolasma_lividum          
              |  |                                                      
              +--+  +----------------------- Obliquaria_reflexa         
                 |  |                                                   
                 |  |                    +-- Cyrtonaias_tampicoensis    
                 +--+  +-----------------+                              
                    |  |                 +-- Glebula_rotundata          
                    |  |                                                
                    +--+                 +-- Ptychobranchus_fasciolaris 
                       |  +--------------+                              
                       |  |              +-- Lemiox_rimosus             
                       |  |                                             
                       +--+              +-- Venustaconcha_ellipsiformis
                          |  +-----------+                              
                          |  |           +-- Obovaria_olivaria          
                          +--+                                          
                             |  +----------- Hamiota_subangulata        
                             +--+                                       
                                |  +-------- Lampsilis_ovata            
                                +--+                                    
                                   |  +----- Ortmanniana_ligamentina    
                                   +--+                                 
                                      |  +-- Lampsilis_straminea        
                                      +--+                              
                                         +-- Lampsilis_hydiana          

Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11

iT tT bi riS oT roT roS iD
2 11 0 2 24 11 3 51.0
              +----- Eurynia_dilatata **          
  +-----------+                                
  |           |  +-- Cambarunio_iris           
  |           +--+                             
  |              +-- Ptychobranchus_fasciolaris
--+                                            
  |              +-- Amblema_plicata           
  |  +-----------+                             
  |  |           +-- Pleurobema_sintoxia **       
  +--+                                         
     |  +----------- Truncilla_truncata        
     |  |                                      
     +--+        +-- Lampsilis_cardium         
        |  +-----+                             
        |  |     +-- Epioblasma_triquetra      
        +--+                                   
           |  +----- Ligumia_recta             
           +--+                                
              |  +-- Obliquaria_reflexa        
              +--+                             
                 +-- Ortmanniana_ligamentina   

Hewitt et al., 2018 [Publ 3758], fig. 5

iT tT bi riS oT roT roS iD
2 2 0 2 7 0 0 --
  +-- Fusconaia_flava **    
--+                      
  +-- Pleurobema_rubellum **

Bolotov et al., 2018 [Publ 3826], fig. 2

iT tT bi riS oT roT roS iD
2 13 0 2 83 5 3 50.0
  +-- Lamprotula_leaii       
  |                          
  +-- Potomida_littoralis    
  |                          
  +-- Pronodularia_japanensis
  |                          
  +-- Gonidea_angulata       
  |                          
  +-- Leguminaia_wheatleyi   
  |                          
  +-- Microcondylaea_bonellii
  |                          
--+-- Amblema_plicata        
  |                          
  +-- Ortmanniana_ligamentina
  |                          
  +-- Lampsilis_cardium      
  |                          
  +-- Cambarunio_iris        
  |                          
  +-- Elliptio_complanata **    
  |                          
  +-- Pleurobema_sintoxia **    
  |                          
  +-- Quadrula_quadrula      

Burlakova et al., 2019 [Publ 3899], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 31 31 4 --
  +----------------------- Plectomerus_dombeyanus 
  |                                               
  |  +-------------------- Popenaias_popeii       
  |  |                                            
  |  |                 +-- Elliptio_crassidens **    
  |  +-----------------+                          
--+  |                 +-- Pleurobema_rubellum **    
  |  |                                            
  |  |                 +-- Cyrtonaias_tampicoensis
  |  |              +--+                          
  |  |              |  +-- Glebula_rotundata      
  |  +--------------+                             
  |  |              |  +-- Reginaia_ebenus        
  +--+              +--+                          
     |                 +-- Obliquaria_reflexa     
     |                                            
     |  +----------------- Toxolasma_lividum      
     |  |                                         
     |  |              +-- Potamilus_fragilis     
     |  |  +-----------+                          
     |  |  |           +-- Potamilus_metnecktayi  
     |  |  |                                      
     |  |  |        +----- Medionidus_acutissimus 
     |  |  +--------+                             
     +--+  |        |  +-- Lemiox_rimosus         
        |  |        +--+                          
        |  |           +-- Dromus_dromas          
        |  |                                      
        |  |        +----- Ellipsaria_lineolata   
        |  |        |                             
        |  +--------+  +-- Truncilla_cognata      
        |  |        |  |                          
        +--+        |  +-- Truncilla_donaciformis 
           |        +--+                          
           |           +-- Truncilla_macrodon     
           |           |                          
           |           +-- Truncilla_truncata     
           |                                      
           |  +----------- Cambarunio_simus       
           |  |                                   
           |  +----------- Leaunio_lienosus       
           |  |                                   
           |  +----------- Villosa_villosa        
           |  |                                   
           |  |        +-- Sagittunio_nasutus     
           |  +--------+                          
           +--+        +-- Lampsilis_ovata        
              |                                   
              |     +----- Venustaconcha_pleasii  
              +-----+                             
              |     |  +-- Epioblasma_rangiana    
              |     +--+                          
              |        +-- Obovaria_unicolor      
              |                                   
              |  +-------- Lampsilis_teres        
              +--+                                
                 |  +----- Lampsilis_siliquoidea  
                 +--+                             
                    |  +-- Lampsilis_hydiana      
                    +--+                          
                       +-- Lampsilis_straminea    

Lopes-Lima et al., 2019 [Publ 3901], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 26 22 3 --
                                +-- Lampsilis_ornata      
  +-----------------------------+                         
  |                             +-- Potamilus_leptodon    
  |                                                       
  |                             +-- Pleurobema_riddellii **  
--+  +--------------------------+                         
  |  |                          +-- Elliptio_crassidens **   
  |  |                                                    
  |  |                       +----- Megalonaias_nervosa   
  +--+                       |                            
     |  +--------------------+  +-- Uniomerus_carolinianus
     |  |                    |  |                         
     |  |                    +--+-- Uniomerus_declivis    
     |  |                       |                         
     +--+                       +-- Uniomerus_tetralasmus 
        |                                                 
        |  +----------------------- Quadrula_quadrula     
        |  |                                              
        |  |                    +-- Quadrula_verrucosa    
        +--+  +-----------------+                         
           |  |                 +-- Quadrula_nobilis      
           |  |                                           
           +--+                 +-- Theliderma_metanevra  
              |  +--------------+                         
              |  |              +-- Theliderma_johnsoni   
              |  |                                        
              +--+           +----- Pustulosa_kieneriana  
                 |  +--------+                            
                 |  |        |  +-- Pustulosa_infucata    
                 |  |        +--+                         
                 +--+           +-- Pustulosa_kleiniana   
                    |                                     
                    |  +----------- Cyclonaias_tuberculata
                    |  |                                  
                    +--+        +-- Pustulosa_pustulosa   
                       |  +-----+                         
                       |  |     +-- Pustulosa_succissa    
                       +--+                               
                          |  +----- Pustulosa_nodulata    
                          +--+                            
                             |  +-- Pustulosa_necki       
                             +--+                         
                                +-- Pustulosa_petrina     

Lopes-Lima et al., 2019 [Publ 3901], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 1 27 23 3 --
                             +-- Pleurobema_oviforme **   
                       +-----+                         
                       |     +-- Pleurobema_riddellii **  
                       |                               
  +--------------------+     +-- Lampsilis_ornata      
  |                    |  +--+                         
  |                    |  |  +-- Potamilus_leptodon    
  |                    +--+                            
  |                       |  +-- Amblema_elliottii     
  |                       +--+                         
--+                          +-- Amblema_plicata       
  |                                                    
  |  +-------------------------- Uniomerus_tetralasmus 
  |  |                                                 
  |  |              +----------- Theliderma_cylindrica 
  |  |  +-----------+                                  
  |  |  |           |  +-------- Theliderma_sparsa     
  +--+  |           +--+                               
     |  |              |  +----- Theliderma_intermedia 
     |  |              +--+                            
     |  |                 |  +-- Theliderma_metanevra  
     |  |                 +--+                         
     +--+                    +-- Theliderma_johnsoni   
        |                                              
        |                 +----- Quadrula_quadrula     
        |  +--------------+                            
        |  |              |  +-- Quadrula_nobilis      
        |  |              +--+                         
        |  |                 +-- Quadrula_verrucosa    
        +--+                                           
           |  +----------------- Pustulosa_kieneriana  
           |  |                                        
           |  |              +-- Pustulosa_infucata    
           +--+  +-----------+                         
              |  |           +-- Pustulosa_kleiniana   
              |  |                                     
              +--+           +-- Pustulosa_pustulosa   
                 |  +--------+                         
                 |  |        +-- Pustulosa_succissa    
                 +--+                                  
                    |  +-------- Cyclonaias_tuberculata
                    +--+                               
                       |  +----- Pustulosa_necki       
                       +--+                            
                          |  +-- Pustulosa_petrina     
                          +--+                         
                             +-- Pustulosa_nodulata    

Lopes-Lima et al., 2019 [Publ 3901], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 1 24 20 3 --
        +-- Amblema_elliottii   
  +-----+                       
  |     +-- Amblema_plicata     
  |                             
--+     +-- Pleurobema_riddellii **
  |  +--+                       
  |  |  +-- Pleurobema_oviforme ** 
  +--+                          
     |  +-- Lampsilis_ornata    
     +--+                       
        +-- Potamilus_leptodon  

Bolotov et al., 2020 [Publ 4084], fig. 1

iT tT bi riS oT roT roS iD
2 7 0 2 34 5 3 47.0
  +-- Amblema_plicata        
  |                          
  +-- Ortmanniana_ligamentina
  |                          
  +-- Lampsilis_cardium      
  |                          
--+-- Cambarunio_iris        
  |                          
  +-- Elliptio_complanata **    
  |                          
  +-- Pleurobema_sintoxia **    
  |                          
  +-- Quadrula_quadrula      

Smith et al., 2020 [Publ 4129], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 26 26 4 --
  +-------------------------------------- Quadrula_quadrula      
  |                                                              
  |                                   +-- Pleurobema_strodeanum **  
--+  +--------------------------------+                          
  |  |                                +-- Elliptoideus_sloatianus **
  |  |                                                           
  |  |                             +----- Reginaia_ebenus        
  +--+  +--------------------------+                             
     |  |                          |  +-- Amblema_plicata        
     |  |                          +--+                          
     |  |                             +-- Plectomerus_dombeyanus 
     +--+                                                        
        |                             +-- Popenaias_popeii       
        |  +--------------------------+                          
        |  |                          +-- Psoronaias_semigranosa 
        |  |                                                     
        +--+  +-------------------------- Obliquaria_reflexa     
           |  |                                                  
           |  |  +----------------------- Cyrtonaias_tampicoensis
           +--+  |                                               
              |  |                    +-- Pachynaias_spheniopsis 
              |  |  +-----------------+                          
              +--+  |                 +-- Lampsilis_cardium      
                 |  |                                            
                 |  |        +----------- Ellipsaria_lineolata   
                 |  |  +-----+                                   
                 +--+  |     |  +-------- Truncilla_truncata     
                    |  |     +--+                                
                    |  |        |  +----- Truncilla_donaciformis 
                    |  |        +--+                             
                    |  |           |  +-- Truncilla_cognata      
                    |  |           +--+                          
                    +--+              +-- Truncilla_macrodon     
                       |                                         
                       |  +-------------- Atlanticoncha_ochracea 
                       |  |                                      
                       |  |           +-- Potamilus_fragilis     
                       |  |     +-----+                          
                       |  |     |     +-- Potamilus_leptodon     
                       +--+  +--+                                
                          |  |  |  +----- Potamilus_metnecktayi  
                          |  |  +--+                             
                          |  |     |  +-- Potamilus_alatus       
                          |  |     +--+                          
                          +--+        +-- Potamilus_purpuratus   
                             |                                   
                             |        +-- Potamilus_capax        
                             |  +-----+                          
                             |  |     +-- Potamilus_inflatus     
                             +--+                                
                                |  +----- Potamilus_ohiensis     
                                +--+                             
                                   |  +-- Potamilus_streckersoni 
                                   +--+                          
                                      +-- Potamilus_amphichaenus 

Smith et al., 2020 [Publ 4129], fig. 2-3

iT tT bi riS oT roT roS iD
2 2 0 2 26 26 4 --
  +----------------------------------------- Quadrula_quadrula      
  |                                                                 
  |                                      +-- Pleurobema_strodeanum **  
--+  +-----------------------------------+                          
  |  |                                   +-- Elliptoideus_sloatianus **
  |  |                                                              
  |  |                                +----- Reginaia_ebenus        
  +--+  +-----------------------------+                             
     |  |                             |  +-- Amblema_plicata        
     |  |                             +--+                          
     |  |                                +-- Plectomerus_dombeyanus 
     +--+                                                           
        |                                +-- Popenaias_popeii       
        |  +-----------------------------+                          
        |  |                             +-- Psoronaias_semigranosa 
        |  |                                                        
        +--+  +----------------------------- Obliquaria_reflexa     
           |  |                                                     
           |  |  +-------------------------- Cyrtonaias_tampicoensis
           +--+  |                                                  
              |  |                       +-- Pachynaias_spheniopsis 
              |  |  +--------------------+                          
              +--+  |                    +-- Lampsilis_cardium      
                 |  |                                               
                 |  |           +----------- Ellipsaria_lineolata   
                 |  |  +--------+                                   
                 +--+  |        |  +-------- Truncilla_truncata     
                    |  |        +--+                                
                    |  |           |  +----- Truncilla_donaciformis 
                    |  |           +--+                             
                    |  |              |  +-- Truncilla_cognata      
                    +--+              +--+                          
                       |                 +-- Truncilla_macrodon     
                       |                                            
                       |  +----------------- Atlanticoncha_ochracea 
                       |  |                                         
                       |  |              +-- Potamilus_capax        
                       +--+  +-----------+                          
                          |  |           +-- Potamilus_inflatus     
                          |  |                                      
                          |  |        +----- Potamilus_ohiensis     
                          +--+  +-----+                             
                             |  |     |  +-- Potamilus_streckersoni 
                             |  |     +--+                          
                             |  |        +-- Potamilus_amphichaenus 
                             +--+                                   
                                |        +-- Potamilus_fragilis     
                                |  +-----+                          
                                |  |     +-- Potamilus_leptodon     
                                +--+                                
                                   |  +----- Potamilus_metnecktayi  
                                   +--+                             
                                      |  +-- Potamilus_alatus       
                                      +--+                          
                                         +-- Potamilus_purpuratus   

Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 17 12 4 --
                          +-- Uniomerus_tetralasmus      
  +-----------------------+                              
  |                       +-- Quadrula_quadrula          
  |                                                      
--+                       +-- Elliptio_complanata **        
  |  +--------------------+                              
  |  |                    +-- Pleurobema_oviforme **        
  +--+                                                   
     |  +-------------------- Amblema_plicata            
     |  |                                                
     +--+  +----------------- Popenaias_popeii           
        |  |                                             
        +--+  +-------------- Toxolasma_parvum           
           |  |                                          
           |  |           +-- Potamilus_alatus           
           +--+  +--------+                              
              |  |        +-- Potamilus_leptodon         
              |  |                                       
              +--+        +-- Lampsilis_cardium          
                 |  +-----+                              
                 |  |     +-- Lampsilis_ornata           
                 +--+                                    
                    |  +----- Venustaconcha_ellipsiformis
                    +--+                                 
                       |  +-- Lampsilis_powellii         
                       +--+                              
                          +-- Lampsilis_siliquoidea      

Porto-Hannes et al., 2021 [Publ 4227], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 14 14 3 --
  +-------------------- Quadrula_quadrula      
  |                                            
  +-------------------- Quadrula_verrucosa     
  |                                            
--+                 +-- Eurynia_dilatata **       
  |  +--------------+                          
  |  |              +-- Pleurobema_sintoxia **    
  |  |                                         
  |  |              +-- Amblema_plicata        
  +--+  +-----------+                          
     |  |           +-- Amblema_elliottii      
     |  |                                      
     |  |           +-- Obovaria_olivaria      
     +--+  +--------+                          
        |  |        +-- Ortmanniana_ligamentina
        |  |                                   
        |  |        +-- Obliquaria_reflexa     
        |  +--------+                          
        +--+        +-- Lampsilis_ornata       
           |                                   
           |     +----- Lampsilis_radiata[2]   
           |  +--+                             
           |  |  |  +-- Lampsilis_siliquoidea  
           |  |  +--+                          
           +--+     +-- Lampsilis_radiata[1]   
              |                                
              |  +----- Lampsilis_virescens    
              +--+                             
                 |  +-- Lampsilis_hydiana      
                 +--+                          
                    +-- Lampsilis_straminea    

Wu et al., 2022 [Publ 4238], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 72 13 4 --
                             +-- Uniomerus_tetralasmus      
  +--------------------------+                              
  |                          +-- Quadrula_quadrula          
  |                                                         
--+                          +-- Pleurobema_oviforme **        
  |  +-----------------------+                              
  |  |                       +-- Elliptio_complanata **        
  +--+                                                      
     |  +----------------------- Amblema_plicata            
     |  |                                                   
     +--+  +-------------------- Popenaias_popeii           
        |  |                                                
        +--+  +----------------- Toxolasma_parvum           
           |  |                                             
           +--+              +-- Potamilus_alatus           
              |  +-----------+                              
              |  |           +-- Potamilus_leptodon         
              +--+                                          
                 |  +----------- Leaunio_lienosus           
                 |  |                                       
                 +--+        +-- Lampsilis_cardium          
                    |  +-----+                              
                    |  |     +-- Lampsilis_ornata           
                    +--+                                    
                       |  +----- Venustaconcha_ellipsiformis
                       +--+                                 
                          |  +-- Lampsilis_siliquoidea      
                          +--+                              
                             +-- Lampsilis_powellii         

Mejia-Trujillo & Smith, 2022 [Publ 4274], fig. 2-F

iT tT bi riS oT roT roS iD
2 2 0 2 13 13 4 --
  +-------- Amblema_plicata      
  |                              
--+     +-- Quadrula_quadrula    
  |  +--+                        
  |  |  +-- Uniomerus_tetralasmus
  +--+                           
     |  +-- Pleurobema_oviforme **  
     +--+                        
        +-- Elliptio_complanata **  

Dai et al., 2023 [Publ 4285], fig. 5

iT tT bi riS oT roT roS iD
2 2 0 2 51 13 4 --
                             +-- Quadrula_quadrula          
  +--------------------------+                              
  |                          +-- Uniomerus_tetralasmus      
  |                                                         
--+                          +-- Elliptio_complanata **        
  |  +-----------------------+                              
  |  |                       +-- Pleurobema_oviforme **        
  +--+                                                      
     |  +----------------------- Amblema_plicata            
     |  |                                                   
     +--+  +-------------------- Popenaias_popeii           
        |  |                                                
        +--+  +----------------- Toxolasma_parvum           
           |  |                                             
           +--+              +-- Potamilus_leptodon         
              |  +-----------+                              
              |  |           +-- Potamilus_alatus           
              +--+                                          
                 |  +----------- Leaunio_lienosus           
                 |  |                                       
                 +--+        +-- Lampsilis_cardium          
                    |  +-----+                              
                    |  |     +-- Lampsilis_ornata           
                    +--+                                    
                       |  +----- Venustaconcha_ellipsiformis
                       +--+                                 
                          |  +-- Lampsilis_powellii         
                          +--+                              
                             +-- Lampsilis_siliquoidea      

Smith, 2021 [Publ 4331], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 1 18 7 3 --
                 +-- Uniomerus_tetralasmus      
  +--------------+                              
  |              +-- Megalonaias_nervosa        
  |                                             
--+              +-- Elliptio_crassidens **        
  |  +-----------+                              
  |  |           +-- Elliptio_complanata **        
  +--+                                          
     |  +----------- Amblema_plicata            
     +--+                                       
        |  +-------- Potamilus_streckersoni     
        +--+                                    
           |  +----- Venustaconcha_ellipsiformis
           +--+                                 
              |  +-- Lampsilis_siliquoidea      
              +--+                              
                 +-- Lampsilis_cardium          

Wu et al., 2023 [Publ 4339], fig. 4, S5-aa

iT tT bi riS oT roT roS iD
2 2 0 2 94 15 4 --
                             +-- Quadrula_quadrula          
  +--------------------------+                              
  |                          +-- Uniomerus_tetralasmus      
  |                                                         
--+                          +-- Elliptio_complanata **        
  |  +-----------------------+                              
  |  |                       +-- Pleurobema_oviforme **        
  +--+                                                      
     |  +----------------------- Amblema_plicata            
     |  |                                                   
     +--+  +-------------------- Popenaias_popeii           
        |  |                                                
        +--+  +----------------- Toxolasma_parvum           
           |  |                                             
           |  |           +----- Potamilus_leptodon         
           +--+  +--------+                                 
              |  |        |  +-- Potamilus_alatus           
              |  |        +--+                              
              |  |           +-- Potamilus_streckersoni     
              +--+                                          
                 |  +----------- Leaunio_lienosus           
                 |  |                                       
                 |  |     +----- Lampsilis_ornata           
                 +--+  +--+                                 
                    |  |  |  +-- Lampsilis_cardium          
                    |  |  +--+                              
                    +--+     +-- Lampsilis_cariosa          
                       |                                    
                       |  +----- Venustaconcha_ellipsiformis
                       +--+                                 
                          |  +-- Lampsilis_powellii         
                          +--+                              
                             +-- Lampsilis_siliquoidea      

Wu et al., 2023 [Publ 4339], fig. 4, S5-nt

iT tT bi riS oT roT roS iD
2 2 0 2 95 15 4 --
                             +-- Quadrula_quadrula          
  +--------------------------+                              
  |                          +-- Uniomerus_tetralasmus      
  |                                                         
--+                          +-- Elliptio_complanata **        
  |  +-----------------------+                              
  |  |                       +-- Pleurobema_oviforme **        
  +--+                                                      
     |  +----------------------- Amblema_plicata            
     |  |                                                   
     +--+  +-------------------- Popenaias_popeii           
        |  |                                                
        +--+  +----------------- Toxolasma_parvum           
           |  |                                             
           |  |           +----- Potamilus_leptodon         
           +--+  +--------+                                 
              |  |        |  +-- Potamilus_alatus           
              |  |        +--+                              
              |  |           +-- Potamilus_streckersoni     
              +--+                                          
                 |  +----------- Leaunio_lienosus           
                 |  |                                       
                 |  |     +----- Lampsilis_ornata           
                 +--+  +--+                                 
                    |  |  |  +-- Lampsilis_cardium          
                    |  |  +--+                              
                    +--+     +-- Lampsilis_cariosa          
                       |                                    
                       |  +----- Venustaconcha_ellipsiformis
                       +--+                                 
                          |  +-- Lampsilis_powellii         
                          +--+                              
                             +-- Lampsilis_siliquoidea      

Wu et al., 2024 [Publ 4350], fig. 8a, S1

iT tT bi riS oT roT roS iD
2 2 0 2 90 15 4 --
                             +-- Quadrula_quadrula          
  +--------------------------+                              
  |                          +-- Uniomerus_tetralasmus      
  |                                                         
--+                          +-- Elliptio_complanata **        
  |  +-----------------------+                              
  |  |                       +-- Pleurobema_oviforme **        
  +--+                                                      
     |  +----------------------- Amblema_plicata            
     |  |                                                   
     +--+  +-------------------- Popenaias_popeii           
        |  |                                                
        +--+  +----------------- Toxolasma_parvum           
           |  |                                             
           |  |           +----- Potamilus_leptodon         
           +--+  +--------+                                 
              |  |        |  +-- Potamilus_alatus           
              |  |        +--+                              
              |  |           +-- Potamilus_streckersoni     
              +--+                                          
                 |  +----------- Leaunio_lienosus           
                 |  |                                       
                 |  |     +----- Lampsilis_ornata           
                 +--+  +--+                                 
                    |  |  |  +-- Lampsilis_cardium          
                    |  |  +--+                              
                    +--+     +-- Lampsilis_cariosa          
                       |                                    
                       |  +----- Venustaconcha_ellipsiformis
                       +--+                                 
                          |  +-- Lampsilis_powellii         
                          +--+                              
                             +-- Lampsilis_siliquoidea      

Wu et al., 2024 [Publ 4350], fig. 8b, S2

iT tT bi riS oT roT roS iD
2 2 0 2 90 15 4 --
                             +-- Quadrula_quadrula          
  +--------------------------+                              
  |                          +-- Uniomerus_tetralasmus      
  |                                                         
--+                          +-- Elliptio_complanata **        
  |  +-----------------------+                              
  |  |                       +-- Pleurobema_oviforme **        
  +--+                                                      
     |  +----------------------- Amblema_plicata            
     |  |                                                   
     +--+  +-------------------- Popenaias_popeii           
        |  |                                                
        +--+  +----------------- Toxolasma_parvum           
           |  |                                             
           |  |           +----- Potamilus_leptodon         
           +--+  +--------+                                 
              |  |        |  +-- Potamilus_alatus           
              |  |        +--+                              
              +--+           +-- Potamilus_streckersoni     
                 |                                          
                 |  +----------- Leaunio_lienosus           
                 |  |                                       
                 +--+  +-------- Venustaconcha_ellipsiformis
                    |  |                                    
                    |  |     +-- Lampsilis_powellii         
                    +--+-----+                              
                       |     +-- Lampsilis_siliquoidea      
                       |                                    
                       |  +----- Lampsilis_ornata           
                       +--+                                 
                          |  +-- Lampsilis_cardium          
                          +--+                              
                             +-- Lampsilis_cariosa          

Dai et al., 2024 [Publ 4358], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 2 50 10 4 --
                       +-- Quadrula_quadrula          
  +--------------------+                              
  |                    +-- Uniomerus_tetralasmus      
  |                                                   
--+                    +-- Elliptio_complanata **        
  |  +-----------------+                              
  |  |                 +-- Pleurobema_oviforme **        
  +--+                                                
     |  +----------------- Amblema_plicata            
     |  |                                             
     +--+  +-------------- Popenaias_popeii           
        |  |                                          
        +--+  +----------- Toxolasma_parvum           
           |  |                                       
           +--+        +-- Potamilus_leptodon         
              |  +-----+                              
              |  |     +-- Potamilus_alatus           
              +--+                                    
                 |  +----- Leaunio_lienosus           
                 +--+                                 
                    |  +-- Lampsilis_cardium          
                    +--+                              
                       +-- Venustaconcha_ellipsiformis

Graf, 2000 [Publ 2647], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 17 1 1 --
-- Elliptio

Graf, 2002 [Publ 2651], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 17 3 2 --
     +-- Lampsilis_cardium 
  +--+                     
  |  +-- Obliquaria_reflexa
--+                        
  |  +-- Eurynia_dilatata **  
  +--+                     
     +-- Amblema_plicata   

Walker et al., 2006 [Publ 2666], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 14 5 1 --
-- Fusconaia_flava

Walker et al., 2006 [Publ 2666], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 14 5 1 --
  +-------------- Fusconaia_flava **        
--+                                      
  |  +----------- Potamilus_purpuratus   
  +--+                                   
     |  +-------- Cyrtonaias_tampicoensis
     +--+                                
        |  +----- Ligumia_recta          
        +--+                             
           |  +-- Ortmanniana_ligamentina
           +--+                          
              +-- Lampsilis_teres        

Walker et al., 2006 [Publ 2666], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 14 5 1 --
  +-------------- Fusconaia_flava **        
--+                                      
  |  +----------- Cyrtonaias_tampicoensis
  +--+                                   
     |  +-------- Potamilus_purpuratus   
     +--+                                
        |  +----- Ligumia_recta          
        +--+                             
           |  +-- Ortmanniana_ligamentina
           +--+                          
              +-- Lampsilis_teres        

Walker et al., 2006 [Publ 2666], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 37 5 1 --
  +----------- Cyrtonaias_tampicoensis
  |                                   
--+----------- Fusconaia_flava **        
  |                                   
  |  +-------- Potamilus_purpuratus   
  +--+                                
     |  +----- Lampsilis_teres        
     +--+                             
        |  +-- Ligumia_recta          
        +--+                          
           +-- Ortmanniana_ligamentina

Walker et al., 2006 [Publ 2666], fig. 6-M

iT tT bi riS oT roT roS iD
1 1 0 1 14 5 1 --
  +-------------- Fusconaia_flava **        
--+                                      
  |  +----------- Potamilus_purpuratus   
  +--+                                   
     |  +-------- Cyrtonaias_tampicoensis
     +--+                                
        |  +----- Ligumia_recta          
        +--+                             
           |  +-- Lampsilis_teres        
           +--+                          
              +-- Ortmanniana_ligamentina

Hoeh et al., 2002 [Publ 2673], fig. 1-F

iT tT bi riS oT roT roS iD
1 1 0 1 8 5 1 --
  +-------------- Fusconaia_flava **        
--+                                      
  |  +----------- Cyrtonaias_tampicoensis
  +--+                                   
     |  +-------- Ortmanniana_ligamentina
     +--+                                
        |  +----- Lampsilis_teres        
        +--+                             
           |  +-- Potamilus_purpuratus   
           +--+                          
              +-- Ligumia_recta          

Hoeh et al., 2002 [Publ 2673], fig. 1-M

iT tT bi riS oT roT roS iD
1 1 0 1 8 5 1 --
  +-------------- Fusconaia_flava **        
--+                                      
  |  +----------- Potamilus_purpuratus   
  +--+                                   
     |  +-------- Cyrtonaias_tampicoensis
     +--+                                
        |  +----- Ligumia_recta          
        +--+                             
           |  +-- Lampsilis_teres        
           +--+                          
              +-- Ortmanniana_ligamentina

Krebs, 2004 [Publ 2692], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 4 2 2 --
  +-- Lampsilis_siliquoidea
--+                        
  +-- Fusconaia_flava **      

Krebs, 2004 [Publ 2692], fig. 2-M

iT tT bi riS oT roT roS iD
1 1 0 1 3 1 1 --
  +-- Lampsilis_siliquoidea
--+                        
  +-- Fusconaia_flava **      

Roe & Lydeard, 1998 [Publ 2693], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 10 10 1 --
  +-------------- Fusconaia_cerina **       
  |                                      
  +-------------- Obliquaria_reflexa     
--+                                      
  |  +----------- Potamilus_capax        
  |  |                                   
  |  +----------- Lampsilis_ornata       
  +--+                                   
     |  +-------- Potamilus_fragilis     
     |  |                                
     |  |  +----- Potamilus_inflatus     
     +--+  |                             
        |  |  +-- Potamilus_ohiensis     
        |  +--+                          
        +--+  +-- Potamilus_amphichaenus 
           |                             
           |  +-- Potamilus_purpuratus[1]
           |  |                          
           +--+-- Potamilus_purpuratus[2]
              |                          
              +-- Potamilus_alatus       

Hoeh et al., 1996 [Publ 2694], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 4 0 0 --
-- Fusconaia_flava

Turner et al., 2000 [Publ 2722], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 15 13 3 --
                       +-- Amblema_plicata         
  +--------------------+                           
  |                    +-- Eurynia_dilatata **        
--+                                                
  |  +-------------------- Medionidus_conradicus   
  |  |                                             
  +--+  +----------------- Lampsilis_teres         
     |  |                                          
     |  +----------------- Villosa_delumbis        
     +--+                                          
        |              +-- Lampsilis_satura        
        |  +-----------+                           
        |  |           +-- Lampsilis_cardium       
        +--+                                       
           |  +----------- Lampsilis_reeveiana     
           |  |                                    
           +--+  +-------- Lampsilis_hydiana[1]    
              |  |                                 
              +--+-------- Lampsilis_hydiana[2]    
                 |                                 
                 |  +----- Lampsilis_siliquoidea[2]
                 +--+                              
                    |  +-- Lampsilis_siliquoidea[1]
                    +--+                           
                       +-- Lampsilis_powellii      

Campbell, 2000 [Publ 2724], fig. 1-5

iT tT bi riS oT roT roS iD
1 1 0 1 2 0 0 --
-- Elliptio_complanata

Breton et al., 2011 [Publ 3376], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 38 18 3 --
  +-------------------------- Megalonaias_nervosa        
  |                                                      
--+  +----------------------- Fusconaia_flava **            
  |  |                                                   
  +--+  +-------------------- Reginaia_ebenus            
     |  |                                                
     |  |           +-------- Toxolasma_paulum[2]        
     +--+  +--------+                                    
        |  |        |  +----- Toxolasma_lividum          
        |  |        +--+                                 
        |  |           |  +-- Toxolasma_parvum           
        +--+           +--+                              
           |              +-- Toxolasma_paulum[1]        
           |                                             
           |  +-------------- Ellipsaria_lineolata       
           |  |                                          
           +--+  +----------- Truncilla_macrodon         
              |  |                                       
              |  |        +-- Lemiox_rimosus             
              +--+  +-----+                              
                 |  |     +-- Potamilus_metnecktayi      
                 |  |                                    
                 +--+     +-- Cambarunio_iris            
                    |  +--+                              
                    |  |  +-- Venustaconcha_ellipsiformis
                    +--+                                 
                       |  +-- Lampsilis_powellii         
                       |  |                              
                       +--+-- Lampsilis_ornata           
                          |                              
                          +-- Lampsilis_siliquoidea      

Whelan et al., 2011 [Publ 3380], fig. 1 ML

iT tT bi riS oT roT roS iD
1 1 0 1 36 4 3 --
  +----------- Pleurobema_sintoxia **
--+                               
  |  +-------- Quadrula_quadrula  
  +--+                            
     |  +----- Amblema_plicata    
     +--+                         
        |  +-- Truncilla_truncata 
        +--+                      
           +-- Lampsilis_cardium  

Whelan et al., 2011 [Publ 3380], fig. 1 MP

iT tT bi riS oT roT roS iD
1 1 0 1 36 4 3 --
  +-------- Quadrula_quadrula  
  |                            
--+-------- Pleurobema_sintoxia **
  |                            
  |  +----- Amblema_plicata    
  +--+                         
     |  +-- Lampsilis_cardium  
     +--+                      
        +-- Truncilla_truncata 

Inoue et al., 2013 [Publ 3528], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 16 14 3 --
                       +-- Eurynia_dilatata **      
  +--------------------+                         
  |                    +-- Quadrula_quadrula     
  |                                              
  |              +-------- Paetulunio_fabalis    
--+  +-----------+                               
  |  |           |  +----- Amblema_plicata       
  |  |           +--+                            
  |  |              |  +-- Reginaia_ebenus       
  +--+              +--+                         
     |                 +-- Reginaia_rotulata     
     |                                           
     |  +----------------- Lampsilis_ornata      
     |  |                                        
     +--+  +-------------- Leaunio_vanuxemensis  
        |  |                                     
        |  |           +-- Villosa_villosa       
        +--+-----------+                         
           |           +-- Cambarunio_iris       
           |                                     
           |  +----------- Obovaria_olivaria     
           +--+                                  
              |  +-------- Obovaria_arkansasensis
              +--+                               
                 |  +----- Obovaria_retusa       
                 +--+                            
                    |  +-- Obovaria_unicolor     
                    +--+                         
                       +-- Obovaria_subrotunda   

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 34 4 3 --
  +----------- Quadrula_quadrula 
--+                              
  |  +-------- Eurynia_dilatata **  
  +--+                           
     |  +----- Amblema_plicata   
     +--+                        
        |  +-- Lampsilis_ornata  
        +--+                     
           +-- Truncilla_truncata

Inoue et al., 2014 [Publ 3635], fig. 3A

iT tT bi riS oT roT roS iD
1 1 0 1 20 3 3 --
-- Eurynia_dilatata

Giribet & Distel, 2003 [Publ 3670], fig. 3.5

iT tT bi riS oT roT roS iD
1 1 0 1 5 1 1 --
-- Elliptio_complanata

Plazzi et al., 2011 [Publ 3676], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 17 4 2 --
  +-------- Quadrula_quadrula          
  |                                    
--+-------- Parvaspina_collina **         
  |                                    
  |  +----- Lampsilis_ornata           
  +--+                                 
     |  +-- Epioblasma_rangiana        
     +--+                              
        +-- Venustaconcha_ellipsiformis

Soroka, 2008 [Publ 3695], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 3 0 0 --
-- Fusconaia_flava

Soroka, 2008 [Publ 3695], fig. 1-M

iT tT bi riS oT roT roS iD
1 1 0 1 3 0 0 --
-- Fusconaia_flava

Amaro et al., 2017 [Publ 3791], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 17 6 2 --
-- Fusconaia_flava

Amaro et al., 2017 [Publ 3791], fig. 2-M

iT tT bi riS oT roT roS iD
1 1 0 1 9 3 2 --
     +-- Fusconaia_flava **            
  +--+                              
  |  +-- Quadrula_quadrula          
--+                                 
  |  +-- Lampsilis_siliquoidea      
  +--+                              
     +-- Venustaconcha_ellipsiformis

Huang et al., 2019 [Publ 3843], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 56 9 2 --
                    +-- Quadrula_quadrula          
  +-----------------+                              
  |                 +-- Uniomerus_tetralasmus      
--+                                                
  |  +----------------- Elliptio_complanata **        
  |  |                                             
  +--+  +-------------- Toxolasma_parvum           
     |  |                                          
     +--+           +-- Potamilus_leptodon         
        |  +--------+                              
        |  |        +-- Potamilus_alatus           
        +--+                                       
           |  +-------- Leaunio_lienosus           
           +--+                                    
              |  +----- Venustaconcha_ellipsiformis
              +--+                                 
                 |  +-- Lampsilis_cardium          
                 +--+                              
                    +-- Lampsilis_ornata           

Johnson et al., 2018 [Publ 3844], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 15 15 2 --
  +-------------------------------- Amblema_plicata       
  |                                                       
--+  +----------------------------- Elliptio_crassidens **   
  |  |                                                    
  +--+  +-------------------------- Megalonaias_nervosa   
     |  |                                                 
     +--+  +----------------------- Uniomerus_tetralasmus 
        |  |                                              
        +--+  +-------------------- Theliderma_metanevra  
           |  |                                           
           |  |                 +-- Quadrula_quadrula     
           +--+  +--------------+                         
              |  |              +-- Quadrula_verrucosa    
              |  |                                        
              |  |           +----- Pustulosa_kieneriana  
              +--+  +--------+                            
                 |  |        |  +-- Pustulosa_infucata    
                 |  |        +--+                         
                 |  |           +-- Pustulosa_kleiniana   
                 +--+                                     
                    |           +-- Pustulosa_pustulosa   
                    |  +--------+                         
                    |  |        +-- Pustulosa_succissa    
                    +--+                                  
                       |  +-------- Cyclonaias_tuberculata
                       +--+                               
                          |  +----- Pustulosa_necki       
                          +--+                            
                             |  +-- Pustulosa_nodulata    
                             +--+                         
                                +-- Pustulosa_petrina     

Chase et al., 2018 [Publ 3847], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 38 16 3 --
  +-------- Fusconaia_flava **       
--+                               
  |  +----- Megalonaias_nervosa   
  +--+                            
     |  +-- Cyclonaias_tuberculata
     +--+                         
        +-- Pustulosa_pustulosa   

Keogh & Simons, 2019 [Publ 3935], fig. S1

iT tT bi riS oT roT roS iD
1 1 0 1 9 9 2 --
                    +-- Amblema_plicata      
  +-----------------+                        
  |                 +-- Fusconaia_mitchelli **  
--+                                          
  |  +----------------- Potamilus_fragilis   
  |  |                                       
  +--+  +-------------- Leaunio_lienosus     
     |  |                                    
     +--+           +-- Lampsilis_floridensis
        |  +--------+                        
        |  |        +-- Lampsilis_teres      
        +--+                                 
           |  +-------- Lampsilis_sietmani   
           +--+                              
              |  +----- Lampsilis_siliquoidea
              +--+                           
                 |  +-- Lampsilis_hydiana    
                 +--+                        
                    +-- Lampsilis_straminea  

Metzger et al., 2018 [Publ 3952], fig. 1C

iT tT bi riS oT roT roS iD
1 1 0 1 2 1 1 --
  +-- Elliptio_complanata **  
--+                        
  +-- Lampsilis_siliquoidea

Hewitt et al., 2019 [Publ 3962], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 15 13 1 --
  +----------------------- Fusconaia_flava **      
  |                                             
--+  +-------------------- Lampsilis_siliquoidea
  |  |                                          
  |  |  +----------------- Lampsilis_fasciola   
  +--+  |                                       
     |  |              +-- Epioblasma_rangiana  
     |  +--------------+                        
     +--+              +-- Lampsilis_radiata    
        |                                       
        |  +-------------- Lampsilis_ornata     
        |  |                                    
        +--+           +-- Lampsilis_cardium[1] 
           |  +--------+                        
           |  |        +-- Lampsilis_ovata[1]   
           +--+                                 
              |  +-------- Lampsilis_cardium[6] 
              |  |                              
              +--+  +----- Lampsilis_cardium[5] 
                 |  |                           
                 +--+  +-- Lampsilis_cardium[2] 
                    |  |                        
                    |  +-- Lampsilis_ovata[2]   
                    +--+                        
                       +-- Lampsilis_cardium[3] 
                       |                        
                       +-- Lampsilis_cardium[4] 

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iT tT bi riS oT roT roS iD
1 1 0 1 121 3 3 --
  +-- Amblema_plicata    
  |                      
  +-- Elliptio_complanata **
--+                      
  +-- Lampsilis_cardium  
  |                      
  +-- Quadrula_quadrula  

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 53 10 2 --
                    +-- Quadrula_quadrula          
  +-----------------+                              
  |                 +-- Uniomerus_tetralasmus      
--+                                                
  |  +----------------- Elliptio_complanata **        
  |  |                                             
  +--+              +-- Potamilus_leptodon         
     |  +-----------+                              
     |  |           +-- Potamilus_alatus           
     +--+                                          
        |  +----------- Leaunio_lienosus           
        |  |                                       
        +--+        +-- Lampsilis_cardium          
           |  +-----+                              
           |  |     +-- Lampsilis_ornata           
           +--+                                    
              |  +----- Venustaconcha_ellipsiformis
              +--+                                 
                 |  +-- Lampsilis_powellii         
                 +--+                              
                    +-- Lampsilis_siliquoidea      

Sonowal et al., 2021 [Publ 4221], fig. 3a

iT tT bi riS oT roT roS iD
1 1 0 1 14 1 1 --
-- Elliptio_complanata

Sonowal et al., 2021 [Publ 4221], fig. 3b

iT tT bi riS oT roT roS iD
1 1 0 1 14 1 1 --
-- Elliptio_complanata

Sonowal et al., 2021 [Publ 4221], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 15 1 1 --
-- Elliptio_complanata

Porto-Hannes et al., 2021 [Publ 4227], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 11 11 3 --
  +-------------------- Quadrula_quadrula      
  |                                            
  +-------------------- Pustulosa_pustulosa    
--+                                            
  |  +----------------- Pleurobema_sintoxia **    
  |  |                                         
  +--+  +-------------- Amblema_plicata        
     |  |                                      
     +--+  +----------- Obovaria_olivaria      
        |  |                                   
        +--+  +-------- Ortmanniana_ligamentina
           |  |                                
           +--+  +----- Lampsilis_hydiana      
              |  |                             
              |  +----- Lampsilis_straminea    
              +--+                             
                 |  +-- Lampsilis_siliquoidea  
                 |  |                          
                 |  +-- Lampsilis_radiata      
                 +--+                          
                    +-- Lampsilis[1]           
                    |                          
                    +-- Lampsilis[2]           

Neemuchwala et al., 2023 [Publ 4292], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 26 25 4 --
                       +-------- Pleurobema_strodeanum ** 
  +--------------------+                               
  |                    |  +----- Amblema_plicata       
  |                    +--+                            
  |                       |  +-- Lampsilis_cardium     
  |                       +--+                         
--+                          +-- Popenaias_popeii      
  |                                                    
  |  +-------------------------- Megalonaias_nervosa   
  |  |                                                 
  |  |                 +-------- Uniomerus_carolinianus
  |  |  +--------------+                               
  +--+  |              |  +----- Uniomerus_columbensis 
     |  |              +--+                            
     |  |                 |  +-- Uniomerus_tetralasmus 
     |  |                 +--+                         
     |  |                    +-- Uniomerus_declivis    
     +--+                                              
        |              +-------- Theliderma_cylindrica 
        |  +-----------+                               
        |  |           |  +----- Theliderma_metanevra  
        |  |           +--+                            
        |  |              |  +-- Theliderma_sparsa     
        |  |              +--+                         
        +--+                 +-- Theliderma_johnsoni   
           |                                           
           |  +----------------- Cyclonaias_tuberculata
           |  |                                        
           |  |        +-------- Quadrula_nobilis      
           |  |  +-----+                               
           +--+  |     |  +----- Quadrula_verrucosa    
              |  |     +--+                            
              |  |        |  +-- Quadrula_fragosa      
              |  |        +--+                         
              |  |           +-- Quadrula_quadrula     
              +--+                                     
                 |        +----- Pustulosa_nodulata    
                 |  +-----+                            
                 |  |     |  +-- Pustulosa_necki       
                 |  |     +--+                         
                 |  |        +-- Pustulosa_petrina     
                 +--+                                  
                    |        +-- Pustulosa_kieneriana  
                    |  +-----+                         
                    |  |     +-- Pustulosa_pustulosa   
                    +--+                               
                       |  +----- Pustulosa_succissa    
                       +--+                            
                          |  +-- Pustulosa_infucata    
                          +--+                         
                             +-- Pustulosa_kleiniana   

Wu et al., 2023 [Publ 4339], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 18 5 2 --
           +-- Elliptio_complanata **
  +--------+                      
  |        +-- Quadrula_quadrula  
--+                               
  |  +-------- Obliquaria_reflexa 
  +--+                            
     |  +----- Truncilla_truncata 
     +--+                         
        |  +-- Lampsilis_cardium  
        +--+                      
           +-- Potamilus_alatus   

Page updated: 07:02:15 Thu 04 Apr 2024

 
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