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Page last updated
Wed 04 Dec 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Gonideinae

tribe CONTRADENTINI Modell, 1942: cladomics
     6 genera

Cladomics Summary

number of species = 35
number of subtaxa (rank: genus) = 6

analyzed in 67 cladograms in 41 publications
maximum number of terminals in an analysis = 23, mean = 6.6
maximum number of subtaxa (genus) in an analysis = 5, mean = 2.7

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 58
maximum number of included terminals = 23, mean = 7.47
maximum number of subtaxa (genus) = 5, mean = 2.97
proportion of cladograms that support basic monophyly = 0.97

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 52
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (tribe).
maximum number of included terminals = 23, mean = 7.17
maximum number of subtaxa (genus) = 5, mean = 3.06
proportion of cladograms that support rigorous monophyly = 0.96

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ tree

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iT tT bi riS oT roT roS iD
5 5 3 5 52 12 5 0.0

*** Top Monophyly Cladogram *** 

  +----------------- Chamberlainiini
  |                                 
  |           +----- Lamprotulini   
--+  +--------+                     
  |  |        |  +-- Gonideini      
  |  |        +--+                  
  +--+           +-- Pseudodontini  
     |                              
     |  +----------- Rectidentini   
     |  |                           
     +--+        +-- Trapezoideus **   
        |  +-----+                  
        |  |     +-- Physunio **       
        +--+                        
           |  +----- Yaukthwa **       
           +--+                     
              |  +-- Lens **           
              +--+                  
                 +-- Solenaia **

Jeratthitikul et al., 2024 [Publ 4387], fig. 1

iT tT bi riS oT roT roS iD
3 3 1 3 55 55 7 4.0

*** Top Sister-Group Cladogram ***

           +-------- Lamprotulini   
  +--------+                        
  |        |  +----- Gonideini      
  |        +--+                     
  |           |  +-- Schepmaniini   
--+           +--+                  
  |              +-- Pseudodontini  
  |                                 
  |  +-------------- Chamberlainiini
  +--+                              
     |  +----------- Ctenodesmini   
     +--+                           
        |  +-------- Rectidentini   
        +--+                        
           |  +----- Physunio **       
           +--+                     
              |  +-- Lens **           
              +--+                  
                 +-- Trapezoideus **

Bolotov et al., 2020 [Publ 4100], fig. S1

iT tT bi riS oT roT roS iD
23 23 21 4 89 49 5 22.0 
        +-------- Pseudodontini    
  +-----+                          
  |     |  +----- Gonideini        
  |     +--+                       
  |        |  +-- Chamberlainiini  
--+        +--+                    
  |           +-- Lamprotulini     
  |                                
  |  +----------- Rectidentini     
  |  |                             
  +--+        +-- Physunio_modelli ** 
     |  +-----+                    
     |  |     +-- Physunio_superbus **
     +--+                          
        |  +----- Trapezoideus **     
        +--+                       
           |  +-- Lens **             
           +--+                    
              +-- Yaukthwa **

Bolotov et al., 2020 [Publ 4100], fig. 1

iT tT bi riS oT roT roS iD
23 23 21 4 95 49 5 20.0 
     +----------- Rectidentini             
     |                                     
  +--+        +-- Yaukthwa **                 
  |  |  +-----+                            
  |  |  |     +-- Lens **                     
  |  +--+                                  
  |     |  +----- Trapezoideus **             
  |     +--+                               
  |        |  +-- Physunio_superbus **        
  |        +--+                            
  |           +-- Physunio_modelli **         
--+                                        
  |        +----- Pseudodontini            
  |        |                               
  |        |  +-- Sinosolenaia_iridinea    
  |        |  |                            
  |        |  +-- Sinosolenaia_recognita__1
  |        |  |                            
  |        |  +-- Sinosolenaia_carinata    
  |        |  |                            
  |        |  +-- Leguminaia               
  +--------+  |                            
           |  +-- Microcondylaea           
           |  |                            
           |  +-- Gonidea                  
           |  |                            
           |  +-- Ptychorhynchus           
           |  |                            
           |  +-- Parvasolenaia            
           |  |                            
           +--+-- Lamprotula_leaii         
              |                            
              +-- Lamprotula               
              |                            
              +-- Lamprotula_caveata       
              |                            
              +-- Potomida                 
              |                            
              +-- Pronodularia             
              |                            
              +-- Sinohyriopsis            
              |                            
              +-- Chamberlainia            
              |                            
              +-- Sinosolenaia_recognita__2
              |                            
              +-- Sinosolenaia_oleivora

Konopleva et al., 2019 [Publ 4041], fig. 3

iT tT bi riS oT roT roS iD
19 19 17 4 15 15 2 18.0 
  +----------------- Pseudodontini    
--+                                   
  |  +-------------- Rectidentini     
  +--+                                
     |  +----------- Yaukthwa **         
     +--+                             
        |  +-------- Lens **             
        +--+                          
           |  +----- Trapezoideus **     
           +--+                       
              |  +-- Physunio_superbus **
              +--+                    
                 +-- Physunio_modelli **

Konopleva et al., 2023 [Publ 4262], fig. 8

iT tT bi riS oT roT roS iD
19 19 17 4 63 49 4 17.0 
  +-------------- Pseudodontini
  |                            
--+           +-- Gonideini    
  |  +--------+                
  |  |        +-- Lamprotulini 
  +--+                         
     |  +-------- Rectidentini 
     |  |                      
     +--+     +-- Physunio **     
        |  +--+                
        |  |  +-- Trapezoideus ** 
        +--+                   
           |  +-- Yaukthwa **     
           +--+                
              +-- Lens **

Konopleva et al., 2019 [Publ 3900], fig. 1

iT tT bi riS oT roT roS iD
16 16 14 4 36 22 2 17.0 
  +----------- Rectidentini     
  |                             
--+  +-------- Yaukthwa **         
  |  |                          
  +--+     +-- Physunio_superbus **
     |  +--+                    
     |  |  +-- Physunio_modelli ** 
     +--+                       
        |  +-- Trapezoideus **     
        +--+                    
           +-- Lens **

Pfeiffer et al., 2021 [Publ 4126], fig. 9

iT tT bi riS oT roT roS iD
15 15 13 5 14 14 2 10.0 
  +-------------- Lamprotulini
  |                           
--+  +----------- Rectidentini
  |  |                        
  +--+        +-- Trapezoideus **
     |  +-----+               
     |  |     +-- Physunio **    
     +--+                     
        |  +----- Yaukthwa **    
        +--+                  
           |  +-- Solenaia **    
           +--+               
              +-- Lens **

Bolotov et al., 2022 [Publ 4250], fig. S5

iT tT bi riS oT roT roS iD
15 15 12 3 0 0 0 14.0 
  +----- Yaukthwa **    
--+                  
  |  +-- Physunio **    
  +--+               
     +-- Trapezoideus **

Zieritz et al., 2020 [Publ 4102], fig. 4

iT tT bi riS oT roT roS iD
14 14 12 4 17 17 6 15.0 
  +----------------------- Chamberlainiini
  |                                       
--+  +-------------------- Lamprotulini   
  |  |                                    
  +--+  +----------------- Gonideini      
     |  |                                 
     +--+  +-------------- Pseudodontini  
        |  |                              
        +--+           +-- Ctenodesmini   
           |  +--------+                  
           |  |        +-- Rectidentini   
           +--+                           
              |  +-------- Physunio **       
              +--+                        
                 |  +----- Yaukthwa **       
                 +--+                     
                    |  +-- Trapezoideus **   
                    +--+                  
                       +-- Lens **

Bolotov et al., 2017 [Publ 3761], fig. S1

iT tT bi riS oT roT roS iD
12 12 10 3 127 37 5 12.0 
  +-------- Rectidentini     
  |                          
--+     +-- Physunio_modelli ** 
  |  +--+                    
  |  |  +-- Physunio_superbus **
  +--+                       
     |  +-- Yaukthwa **         
     +--+                    
        +-- Lens **

Bolotov et al., 2022 [Publ 4250], fig. 8

iT tT bi riS oT roT roS iD
12 12 10 2 37 11 3 13.0 
  +----- Pseudodontini
--+                   
  |  +-- Trapezoideus ** 
  +--+                
     +-- Yaukthwa **

Huang et al., 2019 [Publ 3843], fig. S4

iT tT bi riS oT roT roS iD
12 12 7 3 186 44 5 13.0 
  +-------------- Rectidentini     
  |                                
  |  +----------- Physunio_modelli ** 
--+  |                             
  |  +----------- Lens **             
  |  |                             
  |  +----------- Physunio_superbus **
  +--+                             
     +----------- Yaukthwa **         
     |                             
     |        +-- Lens_comptus **     
     |  +-----+                    
     |  |     +-- Lens_novoselovi **  
     +--+                          
        |  +----- Lens_eximius **     
        +--+                       
           |  +-- Lens_contradens **  
           +--+                    
              +-- Lens_micropterus **

Huang et al., 2019 [Publ 3843], fig. S3

iT tT bi riS oT roT roS iD
12 12 10 3 184 43 5 12.0 
  +-------- Rectidentini     
  |                          
--+     +-- Physunio_superbus **
  |  +--+                    
  |  |  +-- Physunio_modelli ** 
  +--+                       
     |  +-- Yaukthwa **         
     +--+                    
        +-- Lens **

Bolotov et al., 2017 [Publ 3764], fig. S1

iT tT bi riS oT roT roS iD
12 12 10 3 113 39 5 12.0 
  +-------- Rectidentini     
  |                          
--+     +-- Physunio_superbus **
  |  +--+                    
  |  |  +-- Physunio_modelli ** 
  +--+                       
     |  +-- Yaukthwa **         
     +--+                    
        +-- Lens **

Zieritz et al., 2024 [Publ 4381], fig. 7A

iT tT bi riS oT roT roS iD
12 12 10 2 0 0 0 13.0 
  +-- Solenaia **
--+           
  +-- Lens **

Huang et al., 2019 [Publ 3843], fig. 2

iT tT bi riS oT roT roS iD
12 12 10 3 186 44 5 12.0 
  +-------- Rectidentini     
  |                          
--+     +-- Physunio_modelli ** 
  |  +--+                    
  |  |  +-- Physunio_superbus **
  +--+                       
     |  +-- Yaukthwa **         
     +--+                    
        +-- Lens **

Pfeiffer et al., 2021 [Publ 4126], fig. 12

iT tT bi riS oT roT roS iD
11 11 8 2 0 0 0 12.0 
  +-- Solenaia **
--+           
  +-- Lens **

Bolotov et al., 2017 [Publ 3761], fig. S2

iT tT bi riS oT roT roS iD
11 11 9 3 128 38 5 13.0 
  +----------- Rectidentini     
--+                             
  |  +-------- Lens **             
  +--+                          
     |  +----- Yaukthwa **         
     +--+                       
        |  +-- Physunio_modelli ** 
        +--+                    
           +-- Physunio_superbus **

Jeratthitikul et al., 2021 [Publ 4212], fig. 4

iT tT bi riS oT roT roS iD
10 10 8 4 11 11 1 8.0 
  +-------- Rectidentini
  |                     
--+     +-- Trapezoideus **
  |  +--+               
  |  |  +-- Physunio **    
  +--+                  
     |  +-- Yaukthwa **    
     +--+               
        +-- Lens **

Bolotov et al., 2018 [Publ 3826], fig. 2

iT tT bi riS oT roT roS iD
10 10 0 3 73 25 4 11.0 
  +----- Rectidentini       
  |                         
  |  +-- Lens_eximius **       
  |  |                      
  |  +-- Physunio_modelli__1 **
  |  |                      
--+  +-- Yaukthwa_nesemanni ** 
  |  |                      
  |  +-- Yaukthwa_panhai **    
  |  |                      
  |  +-- Lens_comptus **       
  |  |                      
  +--+-- Lens_novoselovi **    
     |                      
     +-- Physunio_modelli__2 **
     |                      
     +-- Physunio **           
     |                      
     +-- Yaukthwa_inlenensis **
     |                      
     +-- Lens_contradens **    
     |                      
     +-- Lens_micropterus **

Pfeiffer et al., 2021 [Publ 4126], fig. 25

iT tT bi riS oT roT roS iD
9 9 7 2 0 0 0 10.0 
  +-- Solenaia **
--+           
  +-- Yaukthwa **

Pfeiffer et al., 2021 [Publ 4126], fig. 18

iT tT bi riS oT roT roS iD
8 8 7 3 0 0 0 8.0 
  +----- Yaukthwa **    
--+                  
  |  +-- Trapezoideus **
  +--+               
     +-- Physunio **

Zieritz et al., 2024 [Publ 4381], fig. 2

iT tT bi riS oT roT roS iD
8 8 6 4 38 27 7 5.0 
              +-- Schepmaniini   
        +-----+                  
        |     +-- Pseudodontini  
  +-----+                        
  |     |  +----- Chamberlainiini
  |     +--+                     
  |        |  +-- Lamprotulini   
--+        +--+                  
  |           +-- Gonideini      
  |                              
  |  +----------- Ctenodesmini   
  |  |                           
  +--+  +-------- Rectidentini   
     |  |                        
     +--+     +-- Physunio **       
        |  +--+                  
        |  |  +-- Trapezoideus **   
        +--+                     
           |  +-- Yaukthwa **       
           +--+                  
              +-- Lens **

Zieritz et al., 2021 [Publ 4218], fig. 5

iT tT bi riS oT roT roS iD
7 7 5 4 25 21 7 6.0 
  +----------------------- Chamberlainiini
  |                                       
--+  +-------------------- Lamprotulini   
  |  |                                    
  |  |              +----- Gonideini      
  +--+  +-----------+                     
     |  |           |  +-- Schepmaniini   
     |  |           +--+                  
     +--+              +-- Pseudodontini  
        |                                 
        |  +-------------- Ctenodesmini   
        +--+                              
           |  +----------- Rectidentini   
           +--+                           
              |  +-------- Physunio **       
              +--+                        
                 |  +----- Trapezoideus **   
                 +--+                     
                    |  +-- Lens **           
                    +--+                  
                       +-- Yaukthwa **

Wu et al., 2024 [Publ 4350], fig. 6-7

iT tT bi riS oT roT roS iD
7 7 5 4 30 28 7 4.0 
  +----------------- Chamberlainiini
  |                                 
  |              +-- Schepmaniini   
  |           +--+                  
--+           |  +-- Pseudodontini  
  |  +--------+                     
  |  |        |  +-- Lamprotulini   
  |  |        +--+                  
  +--+           +-- Gonideini      
     |                              
     |  +----------- Ctenodesmini   
     |  |                           
     +--+  +-------- Rectidentini   
        |  |                        
        +--+     +-- Physunio **       
           |  +--+                  
           |  |  +-- Trapezoideus **   
           +--+                     
              |  +-- Lens **           
              +--+                  
                 +-- Yaukthwa **

Wu et al., 2024 [Publ 4350], fig. 5b

iT tT bi riS oT roT roS iD
7 7 5 4 30 28 7 6.0 
  +-------------------------------- Schepmaniini       
  |                                                    
  |  +----------------------------- Pseudodontini      
  |  |                                                 
--+  |     +----------------------- Hyriopsis_bialata  
  |  |     |                                           
  |  |  +--+                    +-- Rectidens          
  |  |  |  |  +-----------------+                      
  |  |  |  |  |                 +-- Ctenodesmini       
  +--+  |  +--+                                        
     |  |     |  +----------------- Ensidens           
     |  |     |  |                                     
     |  |     +--+  +-------------- Hyriopsis_desowitzi
     |  |        |  |                                  
     |  |        +--+  +----------- Yaukthwa_elongatula **
     |  |           |  |                               
     |  |           +--+  +-------- Trapezoideus **       
     +--+              |  |                            
        |              +--+     +-- Physunio **           
        |                 |  +--+                      
        |                 |  |  +-- Yaukthwa_inlenensis **
        |                 +--+                         
        |                    |  +-- Yaukthwa_paiensis **  
        |                    +--+                      
        |                       +-- Lens **               
        |                                              
        |     +-------------------- Inversidens        
        |     |                                        
        |     |              +----- Lamprotula         
        +-----+  +-----------+                         
              |  |           |  +-- Chamberlainiini    
              |  |           +--+                      
              +--+              +-- Potomida           
                 |                                     
                 |  +-------------- Sinosolenaia       
                 |  |                                  
                 +--+           +-- Gonidea            
                    |  +--------+                      
                    |  |        +-- Microcondylaea     
                    +--+                               
                       |  +-------- Koreosolenaia      
                       +--+                            
                          |  +----- Parvasolenaia      
                          +--+                         
                             |  +-- Postolata          
                             +--+                      
                                +-- Ptychorhynchus

Liu et al., 2024 [Publ 4380], fig. 5

iT tT bi riS oT roT roS iD
7 7 5 4 30 28 7 4.0 
                    +-- Schepmaniini   
  +-----------------+                  
  |                 +-- Pseudodontini  
--+                                    
  |  +----------------- Gonideini      
  |  |                                 
  +--+              +-- Chamberlainiini
     |  +-----------+                  
     |  |           +-- Lamprotulini   
     +--+                              
        |  +----------- Ctenodesmini   
        |  |                           
        +--+  +-------- Rectidentini   
           |  |                        
           +--+     +-- Physunio **       
              |  +--+                  
              |  |  +-- Trapezoideus **   
              +--+                     
                 |  +-- Lens **           
                 +--+                  
                    +-- Yaukthwa **

Wu et al., 2024 [Publ 4388], fig. 5

iT tT bi riS oT roT roS iD
7 7 5 4 31 29 7 4.0 
  +----------------- Chamberlainiini
  |                                 
  |        +-------- Lamprotulini   
--+  +-----+                        
  |  |     |  +----- Gonideini      
  |  |     +--+                     
  |  |        |  +-- Schepmaniini   
  +--+        +--+                  
     |           +-- Pseudodontini  
     |                              
     |  +----------- Ctenodesmini   
     |  |                           
     +--+  +-------- Rectidentini   
        |  |                        
        +--+     +-- Physunio **       
           |  +--+                  
           |  |  +-- Trapezoideus **   
           +--+                     
              |  +-- Lens **           
              +--+                  
                 +-- Yaukthwa **

Wu et al., 2024 [Publ 4350], fig. 5a

iT tT bi riS oT roT roS iD
7 7 5 4 30 28 7 4.0 
  +-------------------------- Chamberlainiini
  |                                          
  |              +----------- Ctenodesmini   
  |              |                           
  |  +-----------+  +-------- Rectidentini   
--+  |           |  |                        
  |  |           +--+     +-- Physunio **       
  |  |              |  +--+                  
  |  |              |  |  +-- Trapezoideus **   
  |  |              +--+                     
  |  |                 |  +-- Lens **           
  +--+                 +--+                  
     |                    +-- Yaukthwa **       
     |                                       
     |                    +-- Schepmaniini   
     |  +-----------------+                  
     |  |                 +-- Pseudodontini  
     |  |                                    
     +--+                 +-- Inversidens    
        |  +--------------+                  
        |  |              +-- Lamprotulini   
        +--+                                 
           |  +-------------- Sinosolenaia   
           |  |                              
           +--+           +-- Gonidea        
              |  +--------+                  
              |  |        +-- Microcondylaea 
              +--+                           
                 |  +-------- Parvasolenaia  
                 +--+                        
                    |  +----- Koreosolenaia  
                    +--+                     
                       |  +-- Postolata      
                       +--+                  
                          +-- Ptychorhynchus

Wu et al., 2024 [Publ 4350], fig. 5c

iT tT bi riS oT roT roS iD
7 7 1 4 30 28 7 6.0 
  +----------------------------- Schepmania_nieuwenhuisi__1
  |                                                        
  +----------------------------- Schepmania_nieuwenhuisi__2
  |                                                        
  |  +-------------------------- Pseudodontini             
  |  |                                                     
--+  |                    +----- Ensidens                  
  |  |                    |                                
  |  |                    +----- Hyriopsis_desowitzi       
  |  |                    |                                
  |  |                    +----- Rectidens                 
  |  |                    |                                
  |  |                    +----- Hyriopsis_bialata         
  +--+  +-----------------+                                
     |  |                 +----- Ctenodesmini              
     |  |                 |                                
     |  |                 |  +-- Physunio **                  
     |  |                 |  |                             
     |  |                 |  +-- Yaukthwa_paiensis **         
     |  |                 |  |                             
     |  |                 |  +-- Yaukthwa_inlenensis **       
     |  |                 +--+                             
     +--+                    +-- Trapezoideus **              
        |                    |                             
        |                    +-- Yaukthwa_elongatula **       
        |                    |                             
        |                    +-- Lens **                      
        |                                                  
        |  +-------------------- Inversidens               
        |  |                                               
        |  |              +----- Lamprotula                
        +--+  +-----------+                                
           |  |           |  +-- Chamberlainiini           
           |  |           +--+                             
           +--+              +-- Potomida                  
              |                                            
              |  +-------------- Sinosolenaia              
              |  |                                         
              +--+  +----------- Gonidea                   
                 |  |                                      
                 +--+----------- Microcondylaea            
                    |                                      
                    |  +-------- Koreosolenaia             
                    +--+                                   
                       |  +----- Parvasolenaia             
                       +--+                                
                          |  +-- Postolata                 
                          +--+                             
                             +-- Ptychorhynchus

Jeratthitikul et al., 2019 [Publ 3911], fig. 4

iT tT bi riS oT roT roS iD
6 6 11 2 3 2 2 18.0 
  +----- Rectidentini
--+                  
  |  +-- Physunio **    
  +--+               
     +-- Lens **

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iT tT bi riS oT roT roS iD
5 5 3 5 52 12 5 0.0 
  +----------------- Chamberlainiini
  |                                 
  |           +----- Pseudodontini  
--+  +--------+                     
  |  |        |  +-- Gonideini      
  |  |        +--+                  
  +--+           +-- Lamprotulini   
     |                              
     |  +----------- Rectidentini   
     |  |                           
     +--+        +-- Trapezoideus **   
        |  +-----+                  
        |  |     +-- Physunio **       
        +--+                        
           |  +----- Yaukthwa **       
           +--+                     
              |  +-- Lens **           
              +--+                  
                 +-- Solenaia **

Bolotov et al., 2019 [Publ 3974], fig. 1

iT tT bi riS oT roT roS iD
5 5 3 1 13 0 0 9.0 
-- (Yaukthwa

Wu et al., 2021 [Publ 4366], fig. 2_I

iT tT bi riS oT roT roS iD
4 4 2 3 87 25 5 6.0 
  +-------- Rectidentini
--+                     
  |  +----- Physunio **    
  +--+                  
     |  +-- Lens **        
     +--+               
        +-- Yaukthwa **

Pfeiffer et al., 2018 [Publ 3827], fig. 1

iT tT bi riS oT roT roS iD
4 4 1 3 3 3 1 5.0 
  +----- Rectidentini
  |                  
--+  +-- Physunio **    
  |  |               
  +--+-- Trapezoideus **
     |               
     +-- Lens **

Wu et al., 2021 [Publ 4366], fig. 6

iT tT bi riS oT roT roS iD
4 4 2 3 87 25 5 6.0 
  +-------- Rectidentini
--+                     
  |  +----- Physunio **    
  +--+                  
     |  +-- Lens **        
     +--+               
        +-- Yaukthwa **

Dai et al., 2024 [Publ 4389], fig. 3

iT tT bi riS oT roT roS iD
3 3 1 2 44 39 7 4.0 
        +-------- Ctenodesmini   
  +-----+                        
  |     |  +----- Rectidentini   
  |     +--+                     
  |        |  +-- Physunio **       
--+        +--+                  
  |           +-- Lens **           
  |                              
  |  +----------- Chamberlainiini
  +--+                           
     |  +-------- Lamprotulini   
     +--+                        
        |  +----- Gonideini      
        +--+                     
           |  +-- Schepmaniini   
           +--+                  
              +-- Pseudodontini

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

iT tT bi riS oT roT roS iD
3 3 1 3 33 33 5 2.0 
     +-------- Rectidentini   
  +--+                        
  |  |  +----- Lens **           
  |  +--+                     
  |     |  +-- Trapezoideus **   
--+     +--+                  
  |        +-- Physunio **       
  |                           
  |  +-------- Pseudodontini  
  +--+                        
     |  +----- Gonideini      
     +--+                     
        |  +-- Chamberlainiini
        +--+                  
           +-- Lamprotulini

Dai et al., 2023 [Publ 4285], fig. 3-BI

iT tT bi riS oT roT roS iD
3 3 1 2 42 40 5 4.0 
  +----------------------- Pseudodontini  
  |                                       
  |                 +----- Rectidentini   
--+  +--------------+                     
  |  |              |  +-- Physunio **       
  |  |              +--+                  
  |  |                 +-- Lens **           
  +--+                                    
     |  +----------------- Inversidens    
     |  |                                 
     |  |              +-- Chamberlainiini
     |  +--------------+                  
     +--+              +-- Lamprotulini   
        |                                 
        |  +-------------- Sinosolenaia   
        |  |                              
        |  |        +----- Gonidea        
        +--+  +-----+                     
           |  |     |  +-- Leguminaia     
           |  |     +--+                  
           |  |        +-- Microcondylaea 
           +--+                           
              |        +-- Koreosolenaia  
              |  +-----+                  
              |  |     +-- Parvasolenaia  
              +--+                        
                 |  +----- Postolata      
                 +--+                     
                    |  +-- Ptychorhynchus 
                    +--+                  
                       +-- Obovalis

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iT tT bi riS oT roT roS iD
3 3 1 2 37 11 4 5.5 
     +----- Rectidentini    
  +--+                      
  |  |  +-- Lens **            
  |  +--+                   
--+     +-- Physunio_modelli **
  |                         
  |  +----- Pseudodontini   
  +--+                      
     |  +-- Gonideini       
     +--+                   
        +-- Lamprotulini

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

iT tT bi riS oT roT roS iD
3 3 1 3 33 33 5 2.0 
     +-------- Rectidentini   
  +--+                        
  |  |  +----- Lens **           
  |  +--+                     
  |     |  +-- Trapezoideus **   
--+     +--+                  
  |        +-- Physunio **       
  |                           
  |  +-------- Pseudodontini  
  +--+                        
     |  +----- Gonideini      
     +--+                     
        |  +-- Chamberlainiini
        +--+                  
           +-- Lamprotulini

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iT tT bi riS oT roT roS iD
3 5 0 3 117 30 5 7.0 
  +-- Lens **     
  |            
  +-- Ensidens 
  |            
--+-- Physunio ** 
  |            
  +-- Rectidens
  |            
  +-- Yaukthwa **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iT tT bi riS oT roT roS iD
3 4 2 2 77 24 5 6.0 
  +-------- Rectidentini
--+                     
  |  +----- Lens **        
  +--+                  
     |  +-- Physunio **    
     +--+               
        +-- unionoid

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iT tT bi riS oT roT roS iD
3 3 1 3 73 28 5 2.0 
  +-------- Rectidentini
--+                     
  |  +----- Lens **        
  +--+                  
     |  +-- Trapezoideus **
     +--+               
        +-- Physunio **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iT tT bi riS oT roT roS iD
3 3 1 3 73 28 5 2.0 
  +-------- Rectidentini
--+                     
  |  +----- Lens **        
  +--+                  
     |  +-- Trapezoideus **
     +--+               
        +-- Physunio **

Konopleva et al., 2017 [Publ 3807], fig. 6

iT tT bi riS oT roT roS iD
3 3 1 2 28 7 2 5.5 
  +-------- Pseudodontini   
--+                         
  |  +----- Rectidentini    
  +--+                      
     |  +-- Lens **            
     +--+                   
        +-- Physunio_modelli **

Dai et al., 2023 [Publ 4285], fig. 3-ML

iT tT bi riS oT roT roS iD
3 3 1 2 42 40 5 4.0 
  +----------------------- Pseudodontini  
  |                                       
  |                 +----- Rectidentini   
  |  +--------------+                     
--+  |              |  +-- Physunio **       
  |  |              +--+                  
  |  |                 +-- Lens **           
  |  |                                    
  +--+              +----- Inversidens    
     |  +-----------+                     
     |  |           |  +-- Chamberlainiini
     |  |           +--+                  
     |  |              +-- Lamprotulini   
     +--+                                 
        |  +-------------- Sinosolenaia   
        |  |                              
        |  |        +----- Gonidea        
        +--+  +-----+                     
           |  |     |  +-- Leguminaia     
           |  |     +--+                  
           |  |        +-- Microcondylaea 
           +--+                           
              |        +-- Koreosolenaia  
              |  +-----+                  
              |  |     +-- Parvasolenaia  
              +--+                        
                 |  +----- Postolata      
                 +--+                     
                    |  +-- Ptychorhynchus 
                    +--+                  
                       +-- Obovalis

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iT tT bi riS oT roT roS iD
2 2 0 2 90 19 5 -- 
  +----------- Chamberlainiini
  |                           
  |     +----- Rectidentini   
--+  +--+                     
  |  |  |  +-- Lens **           
  |  |  +--+                  
  +--+     +-- Physunio **       
     |                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontini  
        +--+                  
           +-- Gonideini

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

iT tT bi riS oT roT roS iD
2 2 0 2 21 10 5 -- 
  +----------- Chamberlainiini
  |                           
  |     +----- Rectidentini   
--+  +--+                     
  |  |  |  +-- Lens **           
  |  |  +--+                  
  +--+     +-- Physunio **       
     |                        
     |  +----- Pseudodontini  
     +--+                     
        |  +-- Lamprotulini   
        +--+                  
           +-- Gonideini

Matos et al., 2024 [Publ 4384], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 18 16 5 -- 
  +----------- Chamberlainiini
  |                           
  |     +----- Rectidentini   
--+  +--+                     
  |  |  |  +-- Physunio **       
  |  |  +--+                  
  +--+     +-- Lens **           
     |                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontini  
        +--+                  
           +-- Gonideini

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iT tT bi riS oT roT roS iD
2 2 0 2 94 18 5 -- 
  +----------- Chamberlainiini
  |                           
  |     +----- Rectidentini   
--+  +--+                     
  |  |  |  +-- Lens **           
  |  |  +--+                  
  +--+     +-- Physunio **       
     |                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontini  
        +--+                  
           +-- Gonideini

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iT tT bi riS oT roT roS iD
2 2 0 2 90 19 5 -- 
  +----------- Chamberlainiini
  |                           
  |     +----- Rectidentini   
--+  +--+                     
  |  |  |  +-- Lens **           
  |  |  +--+                  
  +--+     +-- Physunio **       
     |                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontini  
        +--+                  
           +-- Gonideini

Dai et al., 2023 [Publ 4285], fig. 5

iT tT bi riS oT roT roS iD
2 2 0 2 51 17 5 -- 
  +----------- Chamberlainiini
  |                           
  |     +----- Rectidentini   
--+  +--+                     
  |  |  |  +-- Lens **           
  |  |  +--+                  
  +--+     +-- Physunio **       
     |                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontini  
        +--+                  
           +-- Gonideini

Matos et al., 2024 [Publ 4383], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 18 16 5 -- 
  +----------- Chamberlainiini
  |                           
  |     +----- Rectidentini   
--+  +--+                     
  |  |  |  +-- Physunio **       
  |  |  +--+                  
  +--+     +-- Lens **           
     |                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontini  
        +--+                  
           +-- Gonideini

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iT tT bi riS oT roT roS iD
2 2 0 2 94 18 5 -- 
  +----------- Chamberlainiini
  |                           
  |     +----- Rectidentini   
--+  +--+                     
  |  |  |  +-- Lens **           
  |  |  +--+                  
  +--+     +-- Physunio **       
     |                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontini  
        +--+                  
           +-- Gonideini

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iT tT bi riS oT roT roS iD
2 2 0 2 47 15 5 -- 
  +----------- Chamberlainiini
  |                           
  |     +----- Rectidentini   
--+  +--+                     
  |  |  |  +-- Lens **           
  |  |  +--+                  
  +--+     +-- Physunio **       
     |                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontini  
        +--+                  
           +-- Gonideini

Wu et al., 2024 [Publ 4388], fig. 6

iT tT bi riS oT roT roS iD
2 2 0 2 52 21 5 -- 
  +----------- Chamberlainiini
  |                           
  |     +----- Rectidentini   
--+  +--+                     
  |  |  |  +-- Lens **           
  |  |  +--+                  
  +--+     +-- Physunio **       
     |                        
     |  +----- Lamprotulini   
     +--+                     
        |  +-- Pseudodontini  
        +--+                  
           +-- Gonideini

Hoeh et al., 2009 [Publ 2716], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 45 3 2 -- 
-- Lens_contradens

Hoeh et al., 2009 [Publ 2716], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 46 3 2 -- 
-- Lens_contradens

Hoeh et al., 2009 [Publ 2716], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 45 3 2 -- 
-- Lens_contradens

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iT tT bi riS oT roT roS iD
1 1 0 1 35 5 3 -- 
  +-- Pseudodontini
--+                
  +-- Lens **

Graf, 2002 [Publ 2651], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 17 3 2 -- 
-- Lens_contradens

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iT tT bi riS oT roT roS iD
1 1 0 1 35 5 3 -- 
-- Lens_contradens

Smith, 2021 [Publ 4331], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 19 2 2 -- 
  +----- Chamberlainiini
--+                     
  |  +-- Lens **           
  +--+                  
     +-- Lamprotulini

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 33 10 4 -- 
-- Physunio

Graf & Cummings, 2006 [Publ 1968], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 46 3 2 -- 
-- Lens_contradens

Page updated: 14:54:21 Wed 04 Dec 2024
 
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