Bivalvia | Unionoida | Unionoidea | Unionidae | Ambleminae | Lampsilini genus Leaunio Watters, 2018: cladomics 5 valid species
Cladomics Summary
number of species = 5
analyzed in 32 cladograms in 19 publications
maximum number of terminals in an analysis = 1, mean = 1.0
Basic Monophyly
number of cladograms that test basic monophyly (i.e., more than 1 included species) = 0
maximum number of included terminals = 0, mean = 0
proportion of cladograms that support basic monophyly = 0
Zanatta & Murphy, 2006 [Publ 2672], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
55 |
47 |
22 |
-- |
*** Top Monophyly Cladogram *** *** Top Sister-Group Cladogram ***
+-- Cyrtonaias
+--------------------------------+
| +-- Glebula
|
--+ +-------------------------------- Paetulunio
| |
| | +-- Obliquaria
+--+ +--------------------------+
| | +-- Toxolasma
| |
| | +-------------------------- Sagittunio
+--+ |
| | +----------- Potamilus
| | |
| | +-----------+ +-- Medionidus_conradicus
| | | | +-----+
+--+ | | | +-- Ptychobranchus
| | +--+
| | | +----- Medionidus_acutissimus
| | | |
| | +--+----- Lemiox
| | |
| | | +-- Dromus
+--+ +--+
| +-- Cyprogenia
|
| +----- Lampsilis_teres
| +--------------+
| | | +-- Truncilla
| | +--+
| | +-- Ellipsaria
| |
+--+ +----------------- Leaunio_vanuxemensis **
| |
| | +----------- Hamiota
| | |
| | +--+ +-- Lampsilis_fasciola
+--+ | | +-----+
| | | | +-- Ortmanniana_pectorosa
| | +--+
| | | +----- Lampsilis_ornata
| | +--+
| | | +-- Lampsilis_ovata
+--+ +--+
| +-- Lampsilis_cardium
|
| +----- Venustaconcha
| +-----+
| | | +-- Obovaria
| | +--+
+--+ +-- Epioblasma
|
| +-------- Lampsilis_siliquoidea
+--+
| +----- Ortmanniana_ligamentina
+--+
| +-- Cambarunio
+--+
+-- Ligumia
Campbell et al., 2005 [Publ 2660], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
101 |
42 |
22 |
-- |
+----- Obliquaria
+--------+
| | +-- Cyrtonaias
| +--+
| +-- Glebula
|
| +----------- Sagittunio
| |
| +----------- Leaunio_vanuxemensis **
| |
| +----------- Potamilus
| |
| +----------- Hamiota
| |
| | +-- Ellipsaria
--+ +--------+
| | +-- Truncilla
| |
| | +----- Lampsilis_teres
| +-----+
| | | +-- Ortmanniana_ligamentina
| | +--+
| | +-- Lampsilis_siliquoidea
| |
| | +----- Villosa
| +-----+
| | | +-- Ligumia
| | +--+
| | +-- Cambarunio
| |
+--+ +-------- Ortmanniana_pectorosa
+--+
| | +----- Lampsilis_ornata
| +--+
| | +-- Lampsilis_cardium
| +--+
| +-- Lampsilis_ovata
|
| +-------- Obovaria_olivaria
| |
| +-------- Venustaconcha_ellipsiformis__2
+--+
| +-------- Epioblasma
| |
| | +----- Obovaria_unicolor
| +--+
| | +-- Obovaria_arkansasensis
| +--+
| +-- Obovaria_subrotunda
|
| +-- Ptychobranchus
| +-----+
| | +-- Venustaconcha_ellipsiformis__1
| |
+--+ +----- Medionidus_conradicus
| |
| | +-- Lemiox
+--+--+
| +-- Medionidus_acutissimus
|
| +-- Dromus
+--+
+-- Cyprogenia
Wu et al., 2022 [Publ 4238], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
73 |
8 |
4 |
-- |
+----------------- Toxolasma
|
--+ +-------------- Potamilus
| |
+--+ +----------- Leaunio_lienosus **
| |
+--+ +-- Lampsilis_cardium
| +-----+
| | +-- Lampsilis_ornata
+--+
| +----- Venustaconcha
+--+
| +-- Lampsilis_siliquoidea
+--+
+-- Lampsilis_powellii
Dai et al., 2023 [Publ 4285], fig. 5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
52 |
8 |
4 |
-- |
+----------------- Toxolasma
|
--+ +-------------- Potamilus
| |
+--+ +----------- Leaunio_lienosus **
| |
+--+ +-- Lampsilis_cardium
| +-----+
| | +-- Lampsilis_ornata
+--+
| +----- Venustaconcha
+--+
| +-- Lampsilis_powellii
+--+
+-- Lampsilis_siliquoidea
Inoue et al., 2020 [Publ 3941], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
41 |
26 |
6 |
-- |
+----------------------------------- Toxolasma
|
--+ +-------------------------------- Potamilus
| |
| | +-- Ellipsaria
+--+ +--------------------------+
| | +-- Truncilla
| |
| | +-------------------------- Leaunio_lienosus **
+--+ |
| | +-------------------- Lampsilis_bracteata
| | |
| | +--+ +----------------- Lampsilis_fasciola
| | | | |
+--+ | +--+ +-------------- Lampsilis_ornata
| | | |
| | +--+ +-- Lampsilis_ovata
| | | +--------+
| | | | +-- Lampsilis_cariosa
| | +--+
| | | +-------- Lampsilis_cardium__3
+--+ +--+
| | +----- Lampsilis_cardium__2
| +--+
| | +-- Lampsilis_satura
| +--+
| +-- Lampsilis_cardium__1
|
| +-- Lampsilis_higginsii
| +--------------+
| | +-- Ortmanniana
+-----+
| +-------------- Lampsilis_teres
| |
+--+ +-- Lampsilis_radiata
| +--------+
| | +-- Lampsilis_siliquoidea
+--+
| +-------- Lampsilis_bergmanni
+--+
| +----- Lampsilis_hydiana
+--+
| +-- Lampsilis_virescens
+--+
+-- Lampsilis_straminea
Pfeiffer et al., 2019 [Publ 3937], fig. 6
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
56 |
4 |
4 |
-- |
+----------- Obliquaria
--+
| +-------- Cyrtonaias
+--+
| +----- Leaunio_lienosus **
+--+
| +-- Lampsilis
+--+
+-- Pachynaias
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
29 |
11 |
8 |
-- |
+----------- Ligumia
|
+----------- Truncilla
|
| +-- Ptychobranchus
+--------+
--+ +-- Sagittunio
|
| +-- Lampsilis_siliquoidea
| +-----+
| | +-- Epioblasma
| |
+--+ +----- Ortmanniana
| |
| +----- Cambarunio
+--+
+----- Leaunio_vanuxemensis **
|
| +-- Lampsilis_cardium
+--+
+-- Lampsilis_fasciola
Wu et al., 2023 [Publ 4339], fig. 4_S5-aa
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
95 |
10 |
4 |
-- |
+----------------- Toxolasma
|
--+ +-------------- Potamilus
| |
+--+ +----------- Leaunio_lienosus **
| |
| | +----- Lampsilis_ornata
+--+ +--+
| | | +-- Lampsilis_cardium
| | +--+
+--+ +-- Lampsilis_cariosa
|
| +----- Venustaconcha
+--+
| +-- Lampsilis_powellii
+--+
+-- Lampsilis_siliquoidea
Huang et al., 2019 [Publ 3843], fig. S4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
197 |
13 |
11 |
-- |
+----- Ellipsaria
|
+----- Potamilus_leptodon
|
+----- Medionidus
|
+----- Potamilus_alatus
|
+----- Ptychobranchus
--+
+----- Truncilla
|
+----- Venustaconcha
|
+----- Cambarunio
|
+----- Leaunio_lienosus **
|
| +-- Ortmanniana
+--+
+-- Lampsilis
Porto-Hannes et al., 2019 [Publ 3969], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
17 |
16 |
5 |
-- |
+----------------- Lampsilis_teres
|
+----------------- Obovaria
|
| +-- Leaunio_lienosus **
+--------------+
| +-- Cambarunio
|
| +-------- Ortmanniana
| |
--+--------+ +-- Lampsilis_hydiana
| | +--+
| | | +-- Lampsilis_virescens
| +--+
| | +-- Lampsilis_siliquoidea
| +--+
| +-- Lampsilis_radiata
|
| +-------------- Lampsilis_bracteata
| |
+--+ +----------- Lampsilis_ornata
| |
+--+ +-------- Lampsilis_ovata
| |
+--+ +----- Lampsilis_cardium__3
| |
+--+ +-- Lampsilis_satura
| |
+--+-- Lampsilis_cardium__1
|
+-- Lampsilis_cardium__2
Porto-Hannes et al., 2019 [Publ 3969], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
17 |
16 |
5 |
-- |
+----------------------- Obliquaria
|
| +-- Cambarunio
| +-----------------+
| | +-- Leaunio_lienosus **
--+ |
| | +----------- Lampsilis_teres
| | |
| +--------+ +-------- Ortmanniana
| | | |
| | +--+ +-- Lampsilis_hydiana
+--+ | +--+
| | | +-- Lampsilis_virescens
| +--+
| | +-- Lampsilis_siliquoidea
| +--+
| +-- Lampsilis_radiata
|
| +----------------- Obovaria
| |
+--+ +-------------- Lampsilis_bracteata
| |
+--+ +----------- Lampsilis_ornata
| |
+--+ +-------- Lampsilis_ovata
| |
+--+ +----- Lampsilis_cardium__3
| |
+--+ +-- Lampsilis_satura
| |
+--+-- Lampsilis_cardium__1
|
+-- Lampsilis_cardium__2
Wu et al., 2024 [Publ 4350], fig. 8b_S2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
91 |
10 |
4 |
-- |
+----------------- Toxolasma
|
--+ +-------------- Potamilus
| |
+--+ +----------- Leaunio_lienosus **
| |
+--+ +-------- Venustaconcha
| |
| | +-- Lampsilis_powellii
+--+-----+
| +-- Lampsilis_siliquoidea
|
| +----- Lampsilis_ornata
+--+
| +-- Lampsilis_cardium
+--+
+-- Lampsilis_cariosa
Curole & Kocher, 2005 [Publ 2682], fig. 1-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
11 |
4 |
3 |
-- |
+-- Potamilus
+--+
| +-- Glebula
--+
| +-- Leaunio_lienosus **
+--+
+-- Lampsilis
Huang et al., 2019 [Publ 3843], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
197 |
13 |
11 |
-- |
+-------------- Potamilus_leptodon
|
| +----------- Potamilus_alatus
--+ |
| | +-------- Venustaconcha
| | |
+--+ +-------- Leaunio_lienosus **
| |
| | +-- Ellipsaria
| +-----+
+--+ +-- Truncilla
|
| +-- Medionidus
+-----+
| +-- Ptychobranchus
|
| +----- Cambarunio
+--+
| +-- Ortmanniana
+--+
+-- Lampsilis
Huang et al., 2019 [Publ 3843], fig. S3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
195 |
13 |
11 |
-- |
+-------------------- Potamilus_leptodon
|
--+ +----------------- Potamilus_alatus
| |
| | +-- Medionidus
+--+ +-----------+
| | +-- Ptychobranchus
| |
+--+ +-- Ellipsaria
| +--------+
| | +-- Truncilla
+--+
| +-------- Leaunio_lienosus **
| |
+--+ +-- Venustaconcha
| +--+
| | +-- Cambarunio
+--+
| +-- Ortmanniana
+--+
+-- Lampsilis
Huang et al., 2019 [Publ 3843], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
56 |
6 |
4 |
-- |
+----------- Toxolasma
--+
| +-------- Potamilus
+--+
| +----- Leaunio_lienosus **
+--+
| +-- Venustaconcha
+--+
+-- Lampsilis
Inoue et al., 2020 [Publ 3941], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
40 |
25 |
6 |
-- |
+----------------------------------- Toxolasma
|
--+ +-------------------------------- Ellipsaria
| |
+--+ +----------------------------- Truncilla
| |
| | +-------------------------- Potamilus
+--+ |
| | +----------------------- Leaunio_lienosus **
| | |
+--+ | +----------------- Lampsilis_bracteata
| | |
| | +--+ +-- Lampsilis_ornata
| | | | +-----------+
+--+ | | | +-- Lampsilis_fasciola
| | +--+
| | | +----------- Lampsilis_ovata
| | +--+
| | | +-------- Lampsilis_cardium__3
| | +--+
+--+ | +----- Lampsilis_cardium__2
| +--+
| | +-- Lampsilis_satura
| +--+
| +-- Lampsilis_cardium__1
|
| +-------------- Lampsilis_teres
| |
| | +-- Lampsilis_higginsii
+-----+ +--------+
| | +-- Ortmanniana
| |
+--+ +-- Lampsilis_radiata
| +-----+
| | +-- Lampsilis_siliquoidea
| |
+--+ +-- Lampsilis_hydiana
| +--+
| | +-- Lampsilis_bergmanni
+--+
| +-- Lampsilis_virescens
+--+
+-- Lampsilis_straminea
Pfeiffer et al., 2016 [Publ 3660], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
37 |
5 |
5 |
-- |
+-- Cyrtonaias
+-----+
| +-- Glebula
|
--+ +-- Lampsilis
| +--+
| | +-- Leaunio_lienosus **
+--+
| +-- Truncilla
+--+
+-- Potamilus
Zanatta & Murphy, 2006 [Publ 2672], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
56 |
47 |
22 |
-- |
+-- Paetulunio
+-----------------------------------+
| +-- Toxolasma
|
| +----- Obliquaria
| +-----------------------------+
--+ | | +-- Glebula
| | +--+
| | +-- Cyrtonaias
| |
| | +----- Ptychobranchus
| | +--+
+--+ | | +-- Dromus
| | +--+
| +-----------------------+ +-- Cyprogenia
| | |
| | | +----- Medionidus_conradicus
| | +--+
| | | +-- Medionidus_acutissimus
| | +--+
+--+ +-- Lemiox
|
| +----------------------------- Sagittunio
| |
| +----------------------------- Potamilus
| |
| | +-- Truncilla
+--+ +-----------------------+
| | +-- Ellipsaria
| |
| | +-- Ortmanniana_pectorosa
| | +-----+
| | | +-- Lampsilis_fasciola
+--+ +--------------+
| | | +----- Lampsilis_ornata
| | +--+
| | | +-- Lampsilis_ovata
| | +--+
| | +-- Lampsilis_cardium
+--+
| +-- Ligumia
| +--+
| | +-- Cambarunio
| +--------------+
| | | +-- Lampsilis_teres
| | +--+
+--+ +-- Hamiota
|
| +----------------- Leaunio_vanuxemensis **
| |
+--+ +-------------- Ortmanniana_ligamentina
| |
+--+ +----------- Lampsilis_siliquoidea
| |
+--+ +-------- Obovaria_olivaria
| |
| +-------- Venustaconcha
+--+
+-------- Epioblasma
|
| +----- Obovaria_unicolor
+--+
| +-- Obovaria_subrotunda
+--+
+-- Obovaria_arkansasensis
Wu et al., 2024 [Publ 4350], fig. 8a_S1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
91 |
10 |
4 |
-- |
+----------------- Toxolasma
|
--+ +-------------- Potamilus
| |
+--+ +----------- Leaunio_lienosus **
| |
| | +----- Lampsilis_ornata
+--+ +--+
| | | +-- Lampsilis_cardium
| | +--+
+--+ +-- Lampsilis_cariosa
|
| +----- Venustaconcha
+--+
| +-- Lampsilis_powellii
+--+
+-- Lampsilis_siliquoidea
Inoue et al., 2013 [Publ 3528], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
16 |
9 |
5 |
-- |
+-------- Lampsilis
|
--+ +----- Leaunio_vanuxemensis **
| |
+--+----- Obovaria
|
| +-- Villosa
+--+
+-- Cambarunio
Wu et al., 2023 [Publ 4339], fig. 4_S5-nt
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
95 |
10 |
4 |
-- |
+----------------- Toxolasma
|
--+ +-------------- Potamilus
| |
+--+ +----------- Leaunio_lienosus **
| |
| | +----- Lampsilis_ornata
+--+ +--+
| | | +-- Lampsilis_cardium
| | +--+
+--+ +-- Lampsilis_cariosa
|
| +----- Venustaconcha
+--+
| +-- Lampsilis_powellii
+--+
+-- Lampsilis_siliquoidea
Keogh & Simons, 2019 [Publ 3935], fig. S1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
9 |
7 |
2 |
-- |
+----- Potamilus
--+
| +-- Leaunio_lienosus **
+--+
+-- Lampsilis
Pfeiffer et al., 2019 [Publ 3937], fig. 4-5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
56 |
4 |
4 |
-- |
+----------- Obliquaria
--+
| +-------- Cyrtonaias
+--+
| +----- Leaunio_lienosus **
+--+
| +-- Lampsilis
+--+
+-- Pachynaias
Wu et al., 2021 [Publ 4366], fig. 3-BI
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
73 |
8 |
4 |
-- |
+----------------- Toxolasma
|
--+ +-------------- Potamilus
| |
+--+ +----------- Leaunio_lienosus **
| |
+--+ +-- Lampsilis_ornata
| +-----+
| | +-- Lampsilis_cardium
+--+
| +----- Venustaconcha
+--+
| +-- Lampsilis_siliquoidea
+--+
+-- Lampsilis_powellii
Curole & Kocher, 2005 [Publ 2682], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
10 |
4 |
3 |
-- |
+-------- Leaunio_lienosus **
--+
| +----- Lampsilis_straminea
+--+
| +-- Lampsilis_teres
+--+
+-- Potamilus
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
29 |
11 |
8 |
-- |
+-------------------- Truncilla
|
--+ +----------------- Ligumia
| |
| | +-- Ptychobranchus
+--+ +-----------+
| | +-- Sagittunio
| |
+--+ +----------- Lampsilis_siliquoidea
| |
| | +----- Ortmanniana
+--+ +--+
| | | +-- Lampsilis_cardium
| | +--+
+--+ +-- Lampsilis_fasciola
|
| +----- Epioblasma
+--+
| +-- Cambarunio
+--+
+-- Leaunio_vanuxemensis **
Dai et al., 2024 [Publ 4358], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
51 |
5 |
4 |
-- |
+----------- Toxolasma
--+
| +-------- Potamilus
+--+
| +----- Leaunio_lienosus **
+--+
| +-- Lampsilis
+--+
+-- Venustaconcha
Wu et al., 2021 [Publ 4366], fig. 3-ML
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
73 |
8 |
4 |
-- |
+----------------- Toxolasma
|
--+ +-------------- Potamilus
| |
+--+ +----------- Leaunio_lienosus **
| |
+--+ +-- Lampsilis_ornata
| +-----+
| | +-- Lampsilis_cardium
+--+
| +----- Venustaconcha
+--+
| +-- Lampsilis_siliquoidea
+--+
+-- Lampsilis_powellii
Burlakova et al., 2019 [Publ 3899], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
32 |
25 |
17 |
-- |
+----------------- Toxolasma
|
| +-------------- Potamilus
| |
| | +-- Ellipsaria
| +-----------+
--+ | +-- Truncilla
| |
| | +----- Medionidus
| +--------+
| | | +-- Lemiox
| | +--+
+--+ +-- Dromus
|
| +----------- Cambarunio
| |
| +----------- Leaunio_lienosus **
| |
| +----------- Villosa
| |
| | +-- Sagittunio
| +--------+
+--+ +-- Lampsilis_ovata
|
| +----- Venustaconcha
+-----+
| | +-- Epioblasma
| +--+
| +-- Obovaria
|
| +-------- Lampsilis_teres
+--+
| +----- Lampsilis_siliquoidea
+--+
| +-- Lampsilis_hydiana
+--+
+-- Lampsilis_straminea
Fukata & Iigo, 2020 [Publ 4106], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
53 |
7 |
3 |
-- |
+-------------- Potamilus
|
--+ +----------- Leaunio_lienosus **
| |
+--+ +-- Lampsilis_cardium
| +-----+
| | +-- Lampsilis_ornata
+--+
| +----- Venustaconcha
+--+
| +-- Lampsilis_powellii
+--+
+-- Lampsilis_siliquoidea
Campbell et al., 2005 [Publ 2660], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
101 |
42 |
22 |
-- |
+-------- Toxolasma
+--------------------+
| | +----- Obliquaria
| +--+
| | +-- Cyrtonaias
| +--+
| +-- Glebula
|
| +-------------- Potamilus
| |
| | +-------- Medionidus_acutissimus
--+ | +--+
| +-----------+ | | +----- Lemiox
| | | | +--+
| | | | | +-- Dromus
| | | | +--+
| | +--+ +-- Cyprogenia
| | |
| | | +-- Sagittunio
| | | +--+
| | | | +-- Medionidus_conradicus
+--+ +-----+
| | +-- Ptychobranchus
| +--+
| +-- Venustaconcha_ellipsiformis__2
|
| +-- Ellipsaria
| +--------------------+
| | +-- Truncilla
| |
| | +-------------------- Villosa
+--+ |
| | +-- Obovaria_olivaria
| | +--------+
| | | +-- Venustaconcha_ellipsiformis__1
| | +-----+
+--+ | | +-------- Epioblasma
| | +--+
| | | +----- Obovaria_unicolor
| | +--+
| | | +-- Obovaria_arkansasensis
| | +--+
| | +-- Obovaria_subrotunda
+--+
| +-------- Hamiota
| |
| +-----+ +-- Lampsilis_cardium
| | | +--+
| | | | +-- Lampsilis_ovata
| | +--+
| | | +-- Ortmanniana_pectorosa
+--+ +--+
| +-- Lampsilis_ornata
|
| +-- Ligumia
| +--------+
| | +-- Cambarunio
+--+
| +-------- Lampsilis_teres
+--+
| +----- Ortmanniana_ligamentina
+--+
| +-- Lampsilis_siliquoidea
+--+
+-- Leaunio_vanuxemensis **
|