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Page last updated
Thu 24 Jul 2025

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

The MUSSEL Project Database is moving to a new server, with a new URL! The new address for this page will be: https://musselpdb.org/fmuotwaolcb/db.php?ty=validgen_cld&id=272

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genus Alasmidonta Say, 1818: cladomics
11 valid species

Cladomics Summary

number of species = 11

analyzed in 57 cladograms in 37 publications
maximum number of terminals in an analysis = 7, mean = 1.37

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 7
maximum number of included terminals = 7, mean = 4.0
proportion of cladograms that support basic monophyly = 0.86

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.

PCoA Ordination

NJ tree

Whelan et al., 2023 [Publ 4287], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
7	7	5	7	34	26	12	0.0

*** Top Monophyly Cladogram *** *** Top Sister-Group Cladogram ***

                          +-- Utterbackia             
  +-----------------------+                           
  |                       +-- Pyganodon               
  |                                                   
  |                 +-------- Utterbackiana           
  |                 |                                 
--+  +--------------+     +-- Lasmigona_costata       
  |  |              |  +--+                           
  |  |              |  |  +-- Pseudodontoideus        
  |  |              +--+                              
  |  |                 |  +-- Lasmigona_complanata    
  |  |                 +--+                           
  +--+                    +-- Arcidens                
     |                                                
     |                 +----- Pressodonta             
     |  +--------------+                              
     |  |              |  +-- Anodontoides            
     |  |              +--+                           
     |  |                 +-- Strophitus              
     +--+                                             
        |                 +-- Prolasmidonta           
        |  +--------------+                           
        |  |              +-- Platynaias              
        |  |                                          
        +--+  +-------------- Alasminota              
           |  |                                       
           |  |           +-- Alasmidonta_raveneliana ** 
           +--+  +--------+                           
              |  |        +-- Alasmidonta_marginata **   
              |  |                                    
              +--+        +-- Alasmidonta_varicosa **    
                 |  +-----+                           
                 |  |     +-- Alasmidonta_uwharriensis **
                 +--+                                 
                    |  +----- Alasmidonta_undulata **    
                    +--+                              
                       |  +-- Alasmidonta_triangulata ** 
                       +--+                           
                          +-- Alasmidonta_arcula **

Whelan et al., 2023 [Publ 4287], fig. 3

iT	tT	bi	riS	oT	roT	roS	iD
7	7	5	7	21	20	11	0.0

  +----------------------- Utterbackiana           
  |                                                
  |                    +-- Utterbackia             
  |  +-----------------+                           
--+  |                 +-- Pyganodon               
  |  |                                             
  |  |        +----------- Alasminota              
  |  |        |                                    
  |  |  +-----+        +-- Anodontoides            
  +--+  |     |  +-----+                           
     |  |     |  |     +-- Strophitus              
     |  |     +--+                                 
     |  |        |  +----- Pressodonta             
     |  |        +--+                              
     |  |           |  +-- Platynaias              
     +--+           +--+                           
        |              +-- Prolasmidonta           
        |                                          
        |              +-- Arcidens                
        |  +-----------+                           
        |  |           +-- Pseudodontoideus        
        |  |                                       
        +--+           +-- Alasmidonta_marginata **   
           |  +--------+                           
           |  |        +-- Alasmidonta_raveneliana ** 
           |  |                                    
           +--+        +-- Alasmidonta_varicosa **    
              |  +-----+                           
              |  |     +-- Alasmidonta_uwharriensis **
              +--+                                 
                 |  +----- Alasmidonta_undulata **    
                 +--+                              
                    |  +-- Alasmidonta_triangulata ** 
                    +--+                           
                       +-- Alasmidonta_arcula **

Whelan et al., 2023 [Publ 4287], fig. 4

iT	tT	bi	riS	oT	roT	roS	iD
5	5	2	5	22	20	10	3.0

  +-------------------- Pressodonta             
  |                                             
  |           +-------- Utterbackiana           
  |  +--------+                                 
  |  |        |  +----- Alasminota              
--+  |        +--+                              
  |  |           |  +-- Arcidens                
  |  |           +--+                           
  |  |              +-- Pseudodontoideus        
  |  |                                          
  +--+           +----- Alasmidonta_raveneliana ** 
     |           |                              
     |           |  +-- Alasmidonta_uwharriensis **
     |  +--------+--+                           
     |  |        |  +-- Alasmidonta_varicosa **    
     |  |        |                              
     |  |        |  +-- Alasmidonta_arcula **      
     +--+        +--+                           
        |           +-- Alasmidonta_triangulata ** 
        |                                       
        |  +----------- Platynaias              
        +--+                                    
           |  +-------- Strophitus              
           +--+                                 
              |  +----- Pyganodon               
              +--+                              
                 |  +-- Utterbackia             
                 +--+                           
                    +-- Anodontoides

Inoue et al., 2014 [Publ 3635], fig. 3A

iT	tT	bi	riS	oT	roT	roS	iD
3	3	1	3	17	11	7	4.0

           +-------- Pseudodontoideus     
  +--------+                              
  |        |  +----- Lasmigona_costata    
  |        +--+                           
  |           |  +-- Lasmigona_complanata 
--+           +--+                        
  |              +-- Arcidens             
  |                                       
  |  +-------------- Strophitus           
  |  |                                    
  +--+           +-- Pyganodon            
     |  +--------+                        
     |  |        +-- Utterbackia          
     +--+                                 
        |  +-------- Platynaias           
        +--+                              
           |  +----- Alasmidonta_marginata **
           +--+                           
              |  +-- Alasmidonta_undulata ** 
              +--+                        
                 +-- Alasmidonta_varicosa **

Inoue et al., 2018 [Publ 3858], fig. 1_3a-c

iT	tT	bi	riS	oT	roT	roS	iD
2	20	0	2	102	14	8	24.0

  +-- Alasmidonta_marginata **      
  |                              
  +-- Prolasmidonta              
  |                              
  +-- Alasmidonta_undulata **       
  |                              
  +-- Beringiana                 
  |                              
  +-- Anodonta_kennerlyi         
  |                              
  +-- Anodonta_nuttalliana       
  |                              
  +-- Anodonta_oregonensis       
  |                              
  +-- Arcidens_confragosus       
  |                              
  +-- Arcidens_wheeleri          
  |                              
  +-- Lasmigona_complanata       
  |                              
--+-- Platynaias_compressa       
  |                              
  +-- Lasmigona_costata          
  |                              
  +-- Platynaias_decorata        
  |                              
  +-- Platynaias_subviridis      
  |                              
  +-- Pyganodon_grandis          
  |                              
  +-- Pyganodon_lacustris        
  |                              
  +-- Pseudodontoideus           
  |                              
  +-- Strophitus                 
  |                              
  +-- Utterbackia_imbecillis     
  |                              
  +-- Utterbackia_peninsularis__1
  |                              
  +-- Utterbackia_peninsularis__2

Chase et al., 2018 [Publ 3847], fig. 4

iT	tT	bi	riS	oT	roT	roS	iD
2	2	0	2	36	9	5	--

           +-- Pyganodon            
  +--------+                        
  |        +-- Utterbackia          
--+                                 
  |  +-------- Lasmigona            
  |  |                              
  +--+     +-- Alasmidonta_undulata ** 
     |  +--+                        
     |  |  +-- Alasmidonta_marginata **
     +--+                           
        |  +-- Strophitus           
        +--+                        
           +-- Platynaias

Breton et al., 2011 [Publ 3376], fig. 3

iT	tT	bi	riS	oT	roT	roS	iD
2	2	0	2	35	9	5	--

  +-------------- Lasmigona            
  |                                    
--+  +----------- Strophitus           
  |  |                                 
  +--+        +-- Pyganodon            
     |  +-----+                        
     |  |     +-- Utterbackia          
     +--+                              
        |  +----- Platynaias           
        +--+                           
           |  +-- Alasmidonta_marginata **
           +--+                        
              +-- Alasmidonta_undulata **

Wu et al., 2021 [Publ 4366], fig. 3-BI

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	72	6	4	--

        +-- Pyganodon           
  +-----+                       
  |     +-- Utterbackia         
--+                             
  |  +----- Prolasmidonta       
  +--+                          
     |  +-- Alasmidonta_varicosa **
     +--+                       
        +-- Platynaias

Huang et al., 2019 [Publ 3843], fig. S4

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	198	6	5	--

-- Alasmidonta_marginata

Hou et al., 2024 [Publ 4400], fig. 4

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	75	6	5	--

           +-- Pyganodon           
  +--------+                       
  |        +-- Utterbackia         
--+                                
  |  +-------- Lasmigona           
  +--+                             
     |  +----- Prolasmidonta       
     +--+                          
        |  +-- Alasmidonta_varicosa **
        +--+                       
           +-- Platynaias

Sano et al., 2020 [Publ 4138], fig. 4

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	67	5	4	--

        +-- Pyganodon           
  +-----+                       
  |     +-- Utterbackia         
--+                             
  |  +----- Alasmidonta_varicosa **
  +--+                          
     |  +-- Prolasmidonta       
     +--+                       
        +-- Platynaias

Dai et al., 2025 [Publ 4440], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	44	1	1	--

  +-- Alasmidonta_marginata **
--+                        
  +-- Lasmigona

Bolotov et al., 2017 [Publ 3761], fig. S1

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	138	4	4	--

        +-- Lasmigona            
  +-----+                        
  |     +-- Alasmidonta_marginata **
--+                              
  |  +----- Simpsonaias          
  +--+                           
     |  +-- Pyganodon            
     +--+                        
        +-- Strophitus

Hoeh et al., 2009 [Publ 2716], fig. 4

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	45	3	3	--

  +-------- Lasmigona            
--+                              
  |  +----- Alasmidonta_marginata **
  +--+                           
     |  +-- Strophitus           
     +--+                        
        +-- Pyganodon

Huang et al., 2019 [Publ 3843], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	198	6	5	--

-- Alasmidonta_marginata

Huang et al., 2002 [Publ 2657], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	33	4	4	--

  +----- Utterbackia            
  |                             
--+----- Utterbackiana          
  |                             
  |  +-- Alasmidonta_triangulata **
  +--+                          
     +-- Lasmigona

Dai et al., 2025 [Publ 4440], fig. 1

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	44	1	1	--

  +-- Alasmidonta_marginata **
--+                        
  +-- Lasmigona

Chong et al., 2008 [Publ 2668], fig. 3

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	17	7	7	--

  +-- Pseudodontoideus     
  |                        
--+-- Alasmidonta_marginata **
  |                        
  +-- Lasmigona

Graf & Cummings, 2006 [Publ 1968], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	46	3	3	--

  +-------- Lasmigona            
--+                              
  |  +----- Alasmidonta_marginata **
  +--+                           
     |  +-- Pyganodon            
     +--+                        
        +-- Strophitus

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	79	4	4	--

        +-- Pyganodon            
  +-----+                        
  |     +-- Simpsonaias          
--+                              
  |  +----- Strophitus           
  +--+                           
     |  +-- Alasmidonta_marginata **
     +--+                        
        +-- Lasmigona

Huang et al., 2002 [Publ 2657], fig. 1

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	33	4	4	--

  +----------- Utterbackiana          
--+                                   
  |  +-------- Utterbackia            
  +--+                                
     |  +----- Strophitus             
     +--+                             
        |  +-- Alasmidonta_triangulata **
        +--+                          
           +-- Lasmigona

Wu et al., 2024 [Publ 4388], fig. 6

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	53	1	1	--

  +-- Alasmidonta_varicosa **
--+                       
  +-- Utterbackia

Sano et al., 2020 [Publ 4138], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	67	5	4	--

        +-- Pyganodon           
  +-----+                       
  |     +-- Utterbackia         
--+                             
  |  +----- Alasmidonta_varicosa **
  +--+                          
     |  +-- Prolasmidonta       
     +--+                       
        +-- Platynaias

Boyer et al., 2011 [Publ 2718], fig. 4

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	37	8	7	--

  +-------------- Strophitus           
  |                                    
  |        +----- Lasmigona_complanata 
--+  +-----+                           
  |  |     |  +-- Lasmigona_costata    
  |  |     +--+                        
  +--+        +-- Arcidens             
     |                                 
     |  +-------- Utterbackia          
     |  |                              
     +--+-------- Alasmidonta_marginata **
        |                              
        |  +----- Pyganodon            
        +--+                           
           |  +-- Anodontoides         
           +--+                        
              +-- Platynaias

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	94	6	5	--

        +-- Pyganodon           
  +-----+                       
  |     +-- Utterbackia         
  |                             
--+     +-- Alasmidonta_varicosa **
  |  +--+                       
  |  |  +-- Platynaias          
  +--+                          
     |  +-- Prolasmidonta       
     +--+                       
        +-- Lasmigona

Sano et al., 2020 [Publ 4138], fig. 3

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	67	5	4	--

        +-- Pyganodon           
  +-----+                       
  |     +-- Utterbackia         
--+                             
  |  +----- Alasmidonta_varicosa **
  +--+                          
     |  +-- Platynaias          
     +--+                       
        +-- Prolasmidonta

Hoeh et al., 2009 [Publ 2716], fig. 5

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	45	3	3	--

  +-------- Lasmigona            
--+                              
  |  +----- Alasmidonta_marginata **
  +--+                           
     |  +-- Strophitus           
     +--+                        
        +-- Pyganodon

Wu et al., 2021 [Publ 4366], fig. 4

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	67	5	4	--

        +-- Pyganodon           
  +-----+                       
  |     +-- Utterbackia         
--+                             
  |  +----- Prolasmidonta       
  +--+                          
     |  +-- Platynaias          
     +--+                       
        +-- Alasmidonta_varicosa **

Aunins et al., 2018 [Publ 3863], fig. 1

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	39	4	3	--

     +-- Prolasmidonta       
  +--+                       
  |  +-- Alasmidonta_varicosa **
--+                          
  |  +-- Pyganodon           
  +--+                       
     +-- Utterbackia

Hoeh et al., 2009 [Publ 2716], fig. 3

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	46	3	3	--

  +-------- Lasmigona            
--+                              
  |  +----- Alasmidonta_marginata **
  +--+                           
     |  +-- Strophitus           
     +--+                        
        +-- Pyganodon

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	90	6	5	--

           +-- Pyganodon           
  +--------+                       
  |        +-- Utterbackia         
--+                                
  |  +-------- Lasmigona           
  +--+                             
     |  +----- Prolasmidonta       
     +--+                          
        |  +-- Alasmidonta_varicosa **
        +--+                       
           +-- Platynaias

Lydeard et al., 1996 [Publ 2656], fig. 4

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	27	4	4	--

  +----- Strophitus             
--+                             
  |  +-- Lasmigona              
  +--+                          
     +-- Alasmidonta_triangulata **

Araujo et al., 2018 [Publ 3779], fig. 1

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	17	1	1	--

  +-- Pyganodon            
--+                        
  +-- Alasmidonta_marginata **

Smith et al., 2018 [Publ 3790], fig. 1

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	17	16	7	--

           +-- Utterbackia            
  +--------+                          
  |        +-- Pyganodon              
  |                                   
--+        +-- Anodontoides           
  |  +-----+                          
  |  |     +-- Strophitus             
  |  |                                
  +--+     +-- Alasmidonta_triangulata **
     |  +--+                          
     |  |  +-- Alasminota             
     +--+                             
        |  +-- Pseudodontoideus       
        +--+                          
           +-- Utterbackiana

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	29	4	3	--

     +-- Alasmidonta_marginata **
  +--+                        
  |  +-- Lasmigona            
--+                           
  |  +-- Strophitus           
  +--+                        
     +-- Pyganodon

Wu et al., 2021 [Publ 4366], fig. 3-ML

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	72	6	4	--

        +-- Pyganodon           
  +-----+                       
  |     +-- Utterbackia         
--+                             
  |  +----- Prolasmidonta       
  +--+                          
     |  +-- Alasmidonta_varicosa **
     +--+                       
        +-- Platynaias

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	90	6	5	--

           +-- Pyganodon           
  +--------+                       
  |        +-- Utterbackia         
--+                                
  |  +-------- Lasmigona           
  +--+                             
     |  +----- Prolasmidonta       
     +--+                          
        |  +-- Alasmidonta_varicosa **
        +--+                       
           +-- Platynaias

Wu et al., 2018 [Publ 3836], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	52	4	3	--

  +----- Strophitus           
  |                           
--+  +-- Platynaias           
  |  |                        
  +--+-- Alasmidonta_marginata **
     |                        
     +-- Pyganodon

Boyer et al., 2011 [Publ 2718], fig. 3

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	34	5	4	--

  +----- Alasmidonta_marginata **
  |                           
  +----- Pyganodon            
--+                           
  +----- Lasmigona            
  |                           
  |  +-- Platynaias           
  +--+                        
     +-- Strophitus

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	29	4	3	--

     +-- Alasmidonta_marginata **
  +--+                        
  |  +-- Lasmigona            
--+                           
  |  +-- Strophitus           
  +--+                        
     +-- Pyganodon

Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	24	3	3	--

  +----- Lasmigona            
  |                           
--+  +-- Strophitus           
  |  |                        
  +--+-- Alasmidonta_marginata **
     |                        
     +-- Pyganodon

Chen et al., 2023 [Publ 4302], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	39	1	1	--

  +-- Alasmidonta_marginata **
--+                        
  +-- Lasmigona

Lydeard et al., 1996 [Publ 2656], fig. 3

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	26	5	4	--

        +-- Utterbackia            
  +-----+                          
  |     +-- Utterbackiana          
--+                                
  |  +----- Strophitus             
  +--+                             
     |  +-- Lasmigona              
     +--+                          
        +-- Alasmidonta_triangulata **

Bolotov et al., 2017 [Publ 3764], fig. S1

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	124	4	4	--

-- Alasmidonta_marginata

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	94	6	5	--

           +-- Pyganodon           
  +--------+                       
  |        +-- Utterbackia         
--+                                
  |  +-------- Lasmigona           
  +--+                             
     |  +----- Prolasmidonta       
     +--+                          
        |  +-- Alasmidonta_varicosa **
        +--+                       
           +-- Platynaias

Bolotov et al., 2017 [Publ 3761], fig. S2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	138	4	4	--

  +----- Pyganodon            
  |                           
  +----- Simpsonaias          
--+                           
  +----- Strophitus           
  |                           
  |  +-- Alasmidonta_marginata **
  +--+                        
     +-- Lasmigona

Krebs et al., 2003 [Publ 2691], fig. 3

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	32	8	6	--

-- Alasmidonta_triangulata

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	120	4	4	--

  +----- Pyganodon            
  |                           
  +----- Simpsonaias          
--+                           
  +----- Strophitus           
  |                           
  |  +-- Alasmidonta_marginata **
  +--+                        
     +-- Lasmigona

Chen et al., 2023 [Publ 4302], fig. 1

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	39	1	1	--

  +-- Alasmidonta_marginata **
--+                        
  +-- Lasmigona

Wu et al., 2024 [Publ 4379], fig. 5

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	52	6	5	--

           +-- Pyganodon           
  +--------+                       
  |        +-- Utterbackia         
--+                                
  |  +-------- Lasmigona           
  +--+                             
     |  +----- Prolasmidonta       
     +--+                          
        |  +-- Alasmidonta_varicosa **
        +--+                       
           +-- Platynaias

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	75	4	4	--

        +-- Pyganodon            
  +-----+                        
  |     +-- Simpsonaias          
--+                              
  |  +----- Strophitus           
  +--+                           
     |  +-- Alasmidonta_marginata **
     +--+                        
        +-- Lasmigona

Huang et al., 2019 [Publ 3843], fig. S3

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	196	6	5	--

  +-- Alasmidonta_marginata **
--+                        
  +-- Lasmigona

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	75	4	4	--

        +-- Alasmidonta_marginata **
  +-----+                        
  |     +-- Lasmigona            
--+                              
  |  +----- Pyganodon            
  +--+                           
     |  +-- Strophitus           
     +--+                        
        +-- Simpsonaias

Chen et al., 2020 [Publ 4170], fig. 1

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	33	5	4	--

-- Alasmidonta_varicosa

Graf, 2002 [Publ 2651], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	17	1	1	--

  +-- Alasmidonta_marginata **
--+                        
  +-- Pyganodon

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	52	3	3	--

     +-- Pyganodon           
  +--+                       
  |  +-- Utterbackia         
--+                          
  |  +-- Alasmidonta_varicosa **
  +--+                       
     +-- Prolasmidonta

Bolotov et al., 2018 [Publ 3826], fig. 2

iT	tT	bi	riS	oT	roT	roS	iD
1	1	0	1	82	4	4	--

  +----------- Pyganodon            
--+                                 
  |  +-------- Simpsonaias          
  +--+                              
     |  +----- Strophitus           
     +--+                           
        |  +-- Alasmidonta_marginata **
        +--+                        
           +-- Lasmigona

Page updated: 09:54:13 Thu 24 Jul 2025

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