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Page last updated
Wed 22 Nov 2023

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Unionoida | Unionoidea | Unionidae | Gonideinae | Pseudodontini | Pseudodontina

genus Pilsbryoconcha Simpson, 1900: cladomics
     9 valid species

Cladomics Summary

number of species = 9

analyzed in 50 cladograms in 32 publications
maximum number of terminals in an analysis = 10, mean = 2.48

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 23
maximum number of included terminals = 10, mean = 4.22
proportion of cladograms that support basic monophyly = 0.87

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ tree

Bolotov et al., 2023 [Publ 4312], fig. S2

iT tT bi riS oT roT roS iD
10 10 8 10 32 27 7 0.0

*** Top Monophyly Cladogram *** *** Top Sister-Group Cladogram ***

                          +-- Namkongnaia_inkhavilayi        
           +--------------+                                  
           |              +-- Namkongnaia_lemeslei           
           |                                                 
  +--------+              +-- Thaiconcha_callifera           
  |        |              |                                  
  |        |  +-----------+-- Thaiconcha_thaiensis           
  |        |  |           |                                  
  |        +--+           +-- Thaiconcha_munelliptica        
  |           |                                              
  |           |  +----------- Bineurus_loeiensis             
  |           +--+                                           
  |              |  +-------- Bineurus_panhai                
  |              +--+                                        
  |                 |  +----- Bineurus_anodontinum           
  |                 +--+                                     
--+                    |  +-- Bineurus_exilis                
  |                    +--+                                  
  |                       +-- Bineurus_mouhotii              
  |                                                          
  |              +----------- Pseudodon_vagulus              
  |              |                                           
  |  +-----------+        +-- Pseudodon_lenyanensis          
  |  |           |  +-----+                                  
  |  |           |  |     +-- Pseudodon_inoscularis          
  |  |           +--+                                        
  |  |              |  +----- Pseudodon_cambodjensis         
  |  |              +--+                                     
  +--+                 |  +-- Pseudodon_vondembuschianus     
     |                 +--+                                  
     |                    +-- Pseudodon_mekongi              
     |                                                       
     |  +-------------------- Pilsbryoconcha_carinifera **      
     |  |                                                    
     +--+                 +-- Pilsbryoconcha_linguaeformis[2] **
        |  +--------------+                                  
        |  |              +-- Pilsbryoconcha **                 
        +--+                                                 
           |  +-------------- Pilsbryoconcha_linguaeformis[1] **
           |  |                                              
           +--+           +-- Pilsbryoconcha_schomburgki **     
              |  +--------+                                  
              |  |        +-- Pilsbryoconcha_exilis **          
              +--+                                           
                 |  +-------- Pilsbryoconcha_hoikaab **         
                 +--+                                        
                    |  +----- Pilsbryoconcha_mekongiana **      
                    +--+                                     
                       |  +-- Pilsbryoconcha_acuta **           
                       +--+                                  
                          +-- Pilsbryoconcha_kittitati **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1, 1

iT tT bi riS oT roT roS iD
8 8 6 8 68 7 6 6.0 
                    +-- Nyeinchanconcha_nyeinchani  
              +-----+                               
              |     +-- Sundadontina_cumingii       
  +-----------+                                     
  |           |  +----- Thaiconcha_callifera        
  |           +--+                                  
  |              |  +-- Bineurus_panhai             
  |              +--+                               
--+                 +-- Bineurus_mouhotii           
  |                                                 
  |                 +-- Namkongnaia_inkhavilayi     
  |  +--------------+                               
  |  |              +-- Pseudodon_vondembuschianus  
  |  |                                              
  +--+              +-- Pilsbryoconcha_exilis **       
     |  +-----------+                               
     |  |           +-- Pilsbryoconcha_schomburgki **  
     |  |                                           
     +--+           +-- Pilsbryoconcha_carinifera **   
        |  +--------+                               
        |  |        +-- Pilsbryoconcha_linguaeformis **
        +--+                                        
           |  +-------- Pilsbryoconcha_hoikaab **      
           +--+                                     
              |  +----- Pilsbryoconcha_mekongiana **   
              +--+                                  
                 |  +-- Pilsbryoconcha_acuta **        
                 +--+                               
                    +-- Pilsbryoconcha_kittitati **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2, 2

iT tT bi riS oT roT roS iD
8 8 6 8 68 7 6 6.0 
                    +-- Nyeinchanconcha_nyeinchani  
              +-----+                               
              |     +-- Sundadontina_cumingii       
  +-----------+                                     
  |           |  +----- Thaiconcha_callifera        
  |           +--+                                  
  |              |  +-- Bineurus_panhai             
  |              +--+                               
--+                 +-- Bineurus_mouhotii           
  |                                                 
  |                 +-- Namkongnaia_inkhavilayi     
  |  +--------------+                               
  |  |              +-- Pseudodon_vondembuschianus  
  |  |                                              
  +--+              +-- Pilsbryoconcha_exilis **       
     |  +-----------+                               
     |  |           +-- Pilsbryoconcha_schomburgki **  
     |  |                                           
     +--+           +-- Pilsbryoconcha_carinifera **   
        |  +--------+                               
        |  |        +-- Pilsbryoconcha_linguaeformis **
        +--+                                        
           |  +-------- Pilsbryoconcha_hoikaab **      
           +--+                                     
              |  +----- Pilsbryoconcha_mekongiana **   
              +--+                                  
                 |  +-- Pilsbryoconcha_acuta **        
                 +--+                               
                    +-- Pilsbryoconcha_kittitati **

Bolotov et al., 2023 [Publ 4312], fig. 2

iT tT bi riS oT roT roS iD
8 8 6 8 32 27 7 10.0 
                          +-- Pilsbryoconcha_exilis **        
              +-----------+                                
              |           +-- Pilsbryoconcha_schomburgki **   
              |                                            
  +-----------+           +-- Pilsbryoconcha_carinifera **    
  |           |  +--------+                                
  |           |  |        +-- Pilsbryoconcha_linguaeformis ** 
  |           +--+                                         
  |              |  +-------- Pilsbryoconcha_acuta **         
  |              +--+                                      
  |                 |  +----- Pilsbryoconcha_kittitati **     
  |                 +--+                                   
  |                    |  +-- Pilsbryoconcha_mekongiana **    
  |                    +--+                                
--+                       +-- Pilsbryoconcha_hoikaab **       
  |                                                        
  |  +----------------------- Songkhlanaia_tamodienica     
  |  |                                                     
  |  |                    +-- Namkongnaia_inkhavilayi      
  |  |           +--------+                                
  |  |           |        +-- Namkongnaia_lemeslei         
  |  |           |                                         
  |  |           |     +----- Pseudodon_vondembuschianus[2]
  |  |  +--------+  +--+                                   
  |  |  |        |  |  |  +-- Pseudodon_vondembuschianus[1]
  +--+  |        |  |  +--+                                
     |  |        |  |     +-- Pseudodon_mekongi            
     |  |        +--+                                      
     |  |           |     +-- Pseudodon_vagulus            
     |  |           |  +--+                                
     |  |           |  |  +-- Pseudodon_cambodjensis       
     |  |           +--+                                   
     |  |              |  +-- Pseudodon_lenyanensis        
     |  |              +--+                                
     |  |                 +-- Pseudodon_inoscularis        
     +--+                                                  
        |     +-------------- Nyeinchanconcha_nyeinchani   
        |     |                                            
        |     |        +----- Thaiconcha_munelliptica      
        |  +--+  +-----+                                   
        |  |  |  |     |  +-- Thaiconcha_callifera         
        |  |  |  |     +--+                                
        |  |  +--+        +-- Thaiconcha_thaiensis         
        |  |     |                                         
        |  |     |  +-------- Bineurus_loeiensis           
        |  |     +--+                                      
        |  |        |  +----- Bineurus_mouhotii            
        |  |        +--+                                   
        +--+           |  +-- Bineurus_anodontinum         
           |           +--+                                
           |              +-- Bineurus_exilis              
           |                                               
           |        +-------- Sundadontina_sulcata         
           |  +-----+                                      
           |  |     |  +----- Sundadontina_harmandi        
           |  |     +--+                                   
           |  |        |  +-- Sundadontina_brandti         
           +--+        +--+                                
              |           +-- Sundadontina_taskaevi        
              |                                            
              |  +----------- Sundadontina_tumida          
              +--+                                         
                 |  +-------- Sundadontina                 
                 +--+                                      
                    |  +----- Sundadontina_tanintharyiensis
                    +--+                                   
                       |  +-- Sundadontina_cumingii        
                       +--+                                
                          +-- Sundadontina_plugpomenica

Bolotov et al., 2023 [Publ 4312], fig. S1

iT tT bi riS oT roT roS iD
8 8 6 8 31 26 7 10.0 
                          +-- Pilsbryoconcha_exilis **        
              +-----------+                                
              |           +-- Pilsbryoconcha_schomburgki **   
              |                                            
  +-----------+           +-- Pilsbryoconcha_carinifera **    
  |           |  +--------+                                
  |           |  |        +-- Pilsbryoconcha_linguaeformis ** 
  |           +--+                                         
  |              |  +-------- Pilsbryoconcha_kittitati **     
  |              +--+                                      
  |                 |  +----- Pilsbryoconcha_acuta **         
  |                 +--+                                   
  |                    |  +-- Pilsbryoconcha_hoikaab **       
  |                    +--+                                
  |                       +-- Pilsbryoconcha_mekongiana **    
  |                                                        
--+           +-------------- Nyeinchanconcha_nyeinchani   
  |           |                                            
  |           |        +----- Thaiconcha_thaiensis         
  |  +--------+  +-----+                                   
  |  |        |  |     |  +-- Thaiconcha_callifera         
  |  |        |  |     +--+                                
  |  |        +--+        +-- Thaiconcha_munelliptica      
  |  |           |                                         
  |  |           |  +-------- Bineurus_loeiensis           
  |  |           +--+                                      
  |  |              |  +----- Bineurus_mouhotii            
  |  |              +--+                                   
  +--+                 |  +-- Bineurus_anodontinum         
     |                 +--+                                
     |                    +-- Bineurus_exilis              
     |                                                     
     |  +-------------------- Songkhlanaia_tamodienica     
     |  |                                                  
     |  |                 +-- Namkongnaia_lemeslei         
     |  |        +--------+                                
     |  |        |        +-- Namkongnaia_inkhavilayi      
     |  |        |                                         
     +--+  +-----+        +-- Pseudodon_inoscularis        
        |  |     |  +-----+                                
        |  |     |  |     +-- Pseudodon_lenyanensis        
        |  |     |  |                                      
        |  |     +--+     +-- Pseudodon_vagulus            
        |  |        |  +--+                                
        |  |        |  |  +-- Pseudodon_cambodjensis       
        +--+        +--+                                   
           |           |  +-- Pseudodon_vondembuschianus   
           |           +--+                                
           |              +-- Pseudodon_mekongi            
           |                                               
           |  +-------------- Sundadontina_tumida          
           |  |                                            
           |  |     +-------- Sundadontina_sulcata         
           +--+  +--+                                      
              |  |  |  +----- Sundadontina_harmandi        
              |  |  +--+                                   
              |  |     |  +-- Sundadontina_brandti         
              +--+     +--+                                
                 |        +-- Sundadontina_taskaevi        
                 |                                         
                 |  +-------- Sundadontina_tanintharyiensis
                 +--+                                      
                    |  +----- Sundadontina                 
                    +--+                                   
                       |  +-- Sundadontina_cumingii        
                       +--+                                
                          +-- Sundadontina_plugpomenica

Konopleva et al., 2023 [Publ 4262], fig. 8

iT tT bi riS oT roT roS iD
6 6 4 6 78 21 7 12.0 
                             +-- Pilsbryoconcha_schomburgki **   
                    +--------+                                
                    |        +-- Pilsbryoconcha_exilis[1] **     
  +-----------------+                                         
  |                 |  +-------- Pilsbryoconcha **               
  |                 +--+                                      
  |                    |  +----- Pilsbryoconcha_linguaeformis ** 
  |                    +--+                                   
  |                       |  +-- Pilsbryoconcha_exilis[2] **     
--+                       +--+                                
  |                          +-- Pilsbryoconcha_carinifera **    
  |                                                           
  |  +-------------------------- Songkhlanaia_tamodienica     
  |  |                                                        
  |  |                       +-- Namkongnaia_inkhavilayi      
  |  |  +--------------------+                                
  +--+  |                    +-- Namkongnaia_lemeslei         
     |  |                                                     
     |  |              +-------- Pseudodon_lenyanensis        
     |  |  +-----------+                                      
     |  |  |           |  +----- Pseudodon_vagulus            
     +--+  |           +--+                                   
        |  |              |  +-- Pseudodon_vondembuschianus   
        |  |              +--+                                
        |  |                 +-- Pseudodon_mekongi            
        |  |                                                  
        |  |                 +-- Sundadontina_taskaevi        
        +--+        +--------+                                
           |        |        +-- Sundadontina_brandti         
           |  +-----+                                         
           |  |     |  +-------- Sundadontina_tumida          
           |  |     +--+                                      
           |  |        |  +----- Sundadontina_tanintharyiensis
           |  |        +--+                                   
           |  |           |  +-- Sundadontina_plugpomenica    
           +--+           +--+                                
              |              +-- Sundadontina_cumingii        
              |                                               
              |  +-------------- Nyeinchanconcha_nyeinchani   
              |  |                                            
              |  |        +----- Thaiconcha_thaiensis         
              +--+  +-----+                                   
                 |  |     |  +-- Thaiconcha_munelliptica      
                 |  |     +--+                                
                 +--+        +-- Thaiconcha_callifera         
                    |                                         
                    |  +-------- Bineurus_loeiensis           
                    +--+                                      
                       |  +----- Bineurus_mouhotii            
                       +--+                                   
                          |  +-- Bineurus_anodontinum         
                          +--+                                
                             +-- Bineurus_exilis

Bolotov et al., 2017 [Publ 3761], fig. S1

iT tT bi riS oT roT roS iD
4 4 2 4 137 10 5 9.0 
           +-------- Pilsbryoconcha_exilis[1] **  
  +--------+                                   
  |        |  +----- Pilsbryoconcha_exilis[3] **  
  |        +--+                                
  |           |  +-- Pilsbryoconcha_exilis[2] **  
  |           +--+                             
  |              +-- Pilsbryoconcha_carinifera ** 
--+                                            
  |        +-------- Sundadontina_tumida       
  |  +-----+                                   
  |  |     |  +----- Pseudodon_vagulus         
  |  |     +--+                                
  |  |        |  +-- Pseudodon_mekongi         
  |  |        +--+                             
  +--+           +-- Pseudodon_vondembuschianus
     |                                         
     |           +-- Nyeinchanconcha_nyeinchani
     |  +--------+                             
     |  |        +-- Sundadontina_cumingii     
     +--+                                      
        |  +-------- Thaiconcha_callifera      
        +--+                                   
           |  +----- Bineurus_exilis           
           +--+                                
              |  +-- Bineurus_mouhotii         
              +--+                             
                 +-- Bineurus_loeiensis

Bolotov et al., 2017 [Publ 3761], fig. S2

iT tT bi riS oT roT roS iD
4 4 2 4 137 11 5 9.0 
  +-------- Pilsbryoconcha_exilis[3] ** 
--+                                  
  |  +----- Pilsbryoconcha_exilis[2] ** 
  +--+                               
     |  +-- Pilsbryoconcha_exilis[1] ** 
     +--+                            
        +-- Pilsbryoconcha_carinifera **

Bolotov et al., 2017 [Publ 3764], fig. S1

iT tT bi riS oT roT roS iD
4 4 2 4 127 10 5 9.0 
              +-- Nyeinchanconcha_nyeinchani
  +-----------+                             
  |           +-- Sundadontina_cumingii     
  |                                         
  |        +----- Bineurus_exilis           
  +--------+                                
  |        |  +-- Bineurus_mouhotii         
  |        +--+                             
  |           +-- Bineurus_loeiensis        
  |                                         
--+     +-------- Sundadontina_tumida       
  +-----+                                   
  |     |  +----- Pseudodon_vagulus         
  |     +--+                                
  |        |  +-- Pseudodon_mekongi         
  |        +--+                             
  |           +-- Pseudodon_vondembuschianus
  |                                         
  |  +----------- Thaiconcha_callifera      
  +--+                                      
     |  +-------- Pilsbryoconcha_exilis[3] **  
     +--+                                   
        |  +----- Pilsbryoconcha_exilis[2] **  
        +--+                                
           |  +-- Pilsbryoconcha_exilis[1] **  
           +--+                             
              +-- Pilsbryoconcha_carinifera **

Huang et al., 2019 [Publ 3843], fig. 2

iT tT bi riS oT roT roS iD
4 4 2 4 200 11 5 9.0 
           +-------- Pilsbryoconcha_exilis[3] **  
  +--------+                                   
  |        |  +----- Pilsbryoconcha_exilis[2] **  
  |        +--+                                
  |           |  +-- Pilsbryoconcha_exilis[1] **  
  |           +--+                             
  |              +-- Pilsbryoconcha_carinifera ** 
  |                                            
--+     +----------- Nyeinchanconcha_nyeinchani
  |  +--+                                      
  |  |  |  +-------- Thaiconcha_callifera      
  |  |  +--+                                   
  |  |     |  +----- Bineurus_loeiensis        
  |  |     +--+                                
  |  |        |  +-- Bineurus_mouhotii         
  +--+        +--+                             
     |           +-- Bineurus_exilis           
     |                                         
     |        +----- Pseudodon_vagulus         
     |     +--+                                
     |     |  |  +-- Pseudodon_mekongi         
     |     |  +--+                             
     +-----+     +-- Pseudodon_vondembuschianus
           |                                   
           |     +-- Sundadontina_cumingii     
           |     |                             
           +-----+-- Sundadontina_tumida[1]    
                 |                             
                 +-- Sundadontina_tumida[2]

Huang et al., 2019 [Publ 3843], fig. S3

iT tT bi riS oT roT roS iD
4 4 2 4 197 11 5 9.0 
           +-------- Pilsbryoconcha_exilis[3] **     
  +--------+                                      
  |        |  +----- Pilsbryoconcha_exilis[2] **     
  |        +--+                                   
  |           |  +-- Pilsbryoconcha_carinifera **    
  |           +--+                                
  |              +-- Pilsbryoconcha_exilis[1] **     
  |                                               
--+     +----------- Sundadontina_tumida          
  |  +--+                                         
  |  |  |  +-------- Pseudodon_vagulus            
  |  |  +--+                                      
  |  |     |  +----- Pseudodon_vondembuschianus[1]
  |  |     +--+                                   
  |  |        |  +-- Pseudodon_mekongi            
  +--+        +--+                                
     |           +-- Pseudodon_vondembuschianus[2]
     |                                            
     |           +-- Nyeinchanconcha_nyeinchani   
     |  +--------+                                
     |  |        +-- Sundadontina_cumingii        
     +--+                                         
        |  +-------- Thaiconcha_callifera         
        +--+                                      
           |  +----- Bineurus_loeiensis           
           +--+                                   
              |  +-- Bineurus_mouhotii            
              +--+                                
                 +-- Bineurus_exilis

Huang et al., 2019 [Publ 3843], fig. S4

iT tT bi riS oT roT roS iD
3 3 1 3 199 11 5 9.0 
  +-------- Nyeinchanconcha_nyeinchani
  |                                   
  +-------- Thaiconcha_callifera      
  |                                   
  +-------- Sundadontina_cumingii     
  |                                   
  +-------- Sundadontina_tumida[1]    
  |                                   
  +-------- Sundadontina_tumida[2]    
  |                                   
  |  +----- Pilsbryoconcha_exilis[2] **  
--+--+                                
  |  |  +-- Pilsbryoconcha_carinifera ** 
  |  +--+                             
  |     +-- Pilsbryoconcha_exilis[1] **  
  |                                   
  |  +----- Pseudodon_vagulus         
  +--+                                
  |  |  +-- Pseudodon_mekongi         
  |  +--+                             
  |     +-- Pseudodon_vondembuschianus
  |                                   
  |  +----- Bineurus_loeiensis        
  +--+                                
     |  +-- Bineurus_mouhotii         
     +--+                             
        +-- Bineurus_exilis

Bolotov et al., 2020 [Publ 4100], fig. 1

iT tT bi riS oT roT roS iD
3 3 1 3 117 15 5 11.0 
              +-- Thaiconcha_callifera         
        +-----+                                
        |     +-- Thaiconcha_munelliptica      
  +-----+                                      
  |     |  +----- Pilsbryoconcha_exilis[2] **     
  |     +--+                                   
  |        |  +-- Pilsbryoconcha_exilis[1] **     
  |        +--+                                
  |           +-- Pilsbryoconcha **               
--+                                            
  |     +-------- Pseudodon_lenyanensis        
  |  +--+                                      
  |  |  |  +----- Pseudodon_vagulus            
  |  |  +--+                                   
  |  |     |  +-- Pseudodon_vondembuschianus   
  |  |     +--+                                
  +--+        +-- Pseudodon_mekongi            
     |                                         
     |        +-- Sundadontina_taskaevi        
     |  +-----+                                
     |  |     +-- Sundadontina_brandti         
     +--+                                      
        |  +----- Sundadontina_tumida          
        +--+                                   
           |  +-- Sundadontina_cumingii        
           +--+                                
              +-- Sundadontina_tanintharyiensis

Bolotov et al., 2020 [Publ 4100], fig. S1

iT tT bi riS oT roT roS iD
3 3 1 3 111 15 5 11.0 
              +-- Thaiconcha_callifera         
        +-----+                                
        |     +-- Thaiconcha_munelliptica      
  +-----+                                      
  |     |  +----- Pilsbryoconcha_exilis[2] **     
  |     +--+                                   
  |        |  +-- Pilsbryoconcha_exilis[1] **     
  |        +--+                                
  |           +-- Pilsbryoconcha **               
--+                                            
  |     +-------- Pseudodon_lenyanensis        
  |  +--+                                      
  |  |  |  +----- Pseudodon_vagulus            
  |  |  +--+                                   
  |  |     |  +-- Pseudodon_vondembuschianus   
  |  |     +--+                                
  +--+        +-- Pseudodon_mekongi            
     |                                         
     |        +-- Sundadontina_tanintharyiensis
     |  +-----+                                
     |  |     +-- Sundadontina_cumingii        
     +--+                                      
        |  +----- Sundadontina_tumida          
        +--+                                   
           |  +-- Sundadontina_taskaevi        
           +--+                                
              +-- Sundadontina_brandti

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

iT tT bi riS oT roT roS iD
3 3 1 3 33 15 6 9.0 
                 +----- Pilsbryoconcha_exilis **       
  +--------------+                                  
  |              |  +-- Pilsbryoconcha_linguaeformis **
  |              +--+                               
  |                 +-- Pilsbryoconcha **              
  |                                                 
  |           +-------- Nyeinchanconcha_nyeinchani  
  |           |                                     
--+  +--------+  +----- Thaiconcha_callifera        
  |  |        |  |                                  
  |  |        +--+  +-- Bineurus_panhai             
  |  |           |  |                               
  |  |           +--+-- Bineurus_exilis             
  |  |              |                               
  |  |              +-- Bineurus_mouhotii           
  +--+                                              
     |              +-- Sundadontina_brandti        
     |           +--+                               
     |           |  +-- Sundadontina_taskaevi       
     |  +--------+                                  
     |  |        |  +-- Sundadontina_tumida         
     |  |        +--+                               
     |  |           +-- Sundadontina_cumingii       
     +--+                                           
        |           +-- Namkongnaia_lemeslei        
        |  +--------+                               
        |  |        +-- Namkongnaia_inkhavilayi     
        +--+                                        
           |  +-------- Pseudodon_lenyanensis       
           +--+                                     
              |  +----- Pseudodon_vagulus           
              +--+                                  
                 |  +-- Pseudodon_vondembuschianus  
                 +--+                               
                    +-- Pseudodon_mekongi

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

iT tT bi riS oT roT roS iD
3 3 1 3 33 15 6 9.0 
                 +----- Pilsbryoconcha_exilis **       
  +--------------+                                  
  |              |  +-- Pilsbryoconcha_linguaeformis **
  |              +--+                               
  |                 +-- Pilsbryoconcha **              
  |                                                 
  |                 +-- Nyeinchanconcha_nyeinchani  
  |        +--------+                               
--+        |        +-- Sundadontina_cumingii       
  |  +-----+                                        
  |  |     |  +-------- Thaiconcha_callifera        
  |  |     +--+                                     
  |  |        |  +----- Bineurus_mouhotii           
  |  |        +--+                                  
  |  |           |  +-- Bineurus_panhai             
  |  |           +--+                               
  +--+              +-- Bineurus_exilis             
     |                                              
     |           +----- Sundadontina_tumida         
     |  +--------+                                  
     |  |        |  +-- Sundadontina_brandti        
     |  |        +--+                               
     |  |           +-- Sundadontina_taskaevi       
     +--+                                           
        |           +-- Namkongnaia_lemeslei        
        |  +--------+                               
        |  |        +-- Namkongnaia_inkhavilayi     
        +--+                                        
           |  +-------- Pseudodon_lenyanensis       
           +--+                                     
              |  +----- Pseudodon_vagulus           
              +--+                                  
                 |  +-- Pseudodon_vondembuschianus  
                 +--+                               
                    +-- Pseudodon_mekongi

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iT tT bi riS oT roT roS iD
2 2 0 2 79 3 3 -- 
        +-- Pilsbryoconcha_exilis **     
  +-----+                             
  |     +-- Pilsbryoconcha_acuta **      
--+                                   
  |  +----- Pseudodon_vagulus         
  +--+                                
     |  +-- Bineurus_exilis           
     +--+                             
        +-- Nyeinchanconcha_nyeinchani

Bolotov et al., 2018 [Publ 3826], fig. 2

iT tT bi riS oT roT roS iD
2 8 0 2 83 6 2 14.0 
  +-- Bineurus_mouhotii         
  |                             
  +-- Pseudodon_vagulus         
  |                             
  +-- Pseudodon_mekongi         
  |                             
  +-- Pilsbryoconcha_exilis **     
--+                             
  +-- Pilsbryoconcha_carinifera ** 
  |                             
  +-- Bineurus_loeiensis        
  |                             
  +-- Bineurus_exilis           
  |                             
  +-- Pseudodon_vondembuschianus

Konopleva et al., 2019 [Publ 3900], fig. 1

iT tT bi riS oT roT roS iD
2 8 0 2 50 6 2 14.0 
  +-- Bineurus_mouhotii         
  |                             
  +-- Pseudodon_vagulus         
  |                             
  +-- Pseudodon_mekongi         
  |                             
  +-- Pilsbryoconcha_exilis **     
--+                             
  +-- Pilsbryoconcha_carinifera ** 
  |                             
  +-- Bineurus_loeiensis        
  |                             
  +-- Bineurus_exilis           
  |                             
  +-- Pseudodon_vondembuschianus

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iT tT bi riS oT roT roS iD
2 2 0 2 120 4 4 -- 
        +-- Pilsbryoconcha_exilis **     
  +-----+                             
  |     +-- Pilsbryoconcha_carinifera ** 
--+                                   
  |  +----- Pseudodon_mekongi         
  |  |                                
  +--+  +-- Nyeinchanconcha_nyeinchani
     |  |                             
     +--+-- Bineurus_mouhotii         
        |                             
        +-- Sundadontina_tumida

Konopleva et al., 2021 [Publ 4213], fig. 1

iT tT bi riS oT roT roS iD
2 3 0 2 26 18 6 11.0 
                 +----- Pilsbryoconcha_exilis[2] **     
  +--------------+                                   
  |              |  +-- Pilsbryoconcha_exilis[1] **     
  |              +--+                                
  |                 +-- Namkongnaia_lemeslei         
  |                                                  
  |     +-------------- Nyeinchanconcha_nyeinchani   
  |     |                                            
  |     |           +-- Thaiconcha_munelliptica      
--+  +--+           |                                
  |  |  |  +--------+-- Thaiconcha_thaiensis         
  |  |  |  |        |                                
  |  |  +--+        +-- Thaiconcha_callifera         
  |  |     |                                         
  |  |     |  +-------- Bineurus_loeiensis           
  |  |     +--+                                      
  |  |        |  +----- Bineurus_mouhotii            
  |  |        +--+                                   
  +--+           |  +-- Bineurus_anodontinum         
     |           +--+                                
     |              +-- Bineurus_exilis              
     |                                               
     |        +-------- Pseudodon_lenyanensis        
     |     +--+                                      
     |     |  |  +----- Pseudodon_vagulus            
     |     |  +--+                                   
     |     |     |  +-- Pseudodon_vondembuschianus   
     |     |     +--+                                
     +-----+        +-- Pseudodon_mekongi            
           |                                         
           |        +-- Sundadontina_taskaevi        
           |  +-----+                                
           |  |     +-- Sundadontina_brandti         
           +--+                                      
              |  +----- Sundadontina_tumida          
              +--+                                   
                 |  +-- Sundadontina_cumingii        
                 +--+                                
                    +-- Sundadontina_tanintharyiensis

Dai et al., 2023 [Publ 4285], fig. 3-BI

iT tT bi riS oT roT roS iD
2 2 0 2 43 12 5 -- 
     +-------- Nyeinchanconcha_nyeinchani   
  +--+                                      
  |  |  +----- Bineurus_loeiensis           
  |  +--+                                   
  |     |  +-- Bineurus_mouhotii            
  |     +--+                                
  |        +-- Bineurus_exilis              
  |                                         
  |        +-- Thaiconcha_callifera         
  |  +-----+                                
  |  |     +-- Thaiconcha_munelliptica      
--+--+                                      
  |  |  +----- Sundadontina_tanintharyiensis
  |  +--+                                   
  |     |  +-- Sundadontina_brandti         
  |     +--+                                
  |        +-- Sundadontina_taskaevi        
  |                                         
  |        +-- Pilsbryoconcha_carinifera **    
  |  +-----+                                
  |  |     +-- Pilsbryoconcha_exilis **        
  +--+                                      
     |  +----- Pseudodon_lenyanensis        
     +--+                                   
        |  +-- Pseudodon_mekongi            
        +--+                                
           +-- Pseudodon_vagulus

Dai et al., 2023 [Publ 4285], fig. 3-ML

iT tT bi riS oT roT roS iD
2 2 0 2 43 12 5 -- 
           +-- Thaiconcha_callifera         
     +-----+                                
     |     +-- Thaiconcha_munelliptica      
  +--+                                      
  |  |  +----- Sundadontina_tanintharyiensis
  |  +--+                                   
  |     |  +-- Sundadontina_brandti         
  |     +--+                                
--+        +-- Sundadontina_taskaevi        
  |                                         
  |        +-- Pilsbryoconcha_carinifera **    
  |  +-----+                                
  |  |     +-- Pilsbryoconcha_exilis **        
  +--+                                      
     |  +----- Pseudodon_lenyanensis        
     +--+                                   
        |  +-- Pseudodon_mekongi            
        +--+                                
           +-- Pseudodon_vagulus

Graf & Cummings, 2006 [Publ 1968], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 46 1 1 -- 
  +-- Pilsbryoconcha_exilis **     
--+                             
  +-- Pseudodon_vondembuschianus

Graf, 2002 [Publ 2651], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 17 1 1 -- 
  +-- Pilsbryoconcha_exilis **     
--+                             
  +-- Pseudodon_vondembuschianus

Hoeh et al., 2009 [Publ 2716], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 46 1 1 -- 
  +-- Pilsbryoconcha_exilis **     
--+                             
  +-- Pseudodon_vondembuschianus

Hoeh et al., 2009 [Publ 2716], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 45 1 1 -- 
  +-- Pilsbryoconcha_exilis **     
--+                             
  +-- Pseudodon_vondembuschianus

Hoeh et al., 2009 [Publ 2716], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 45 1 1 -- 
  +-- Pilsbryoconcha_exilis **     
--+                             
  +-- Pseudodon_vondembuschianus

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 39 5 4 -- 
  +----------- Pilsbryoconcha_carinifera ** 
  |                                      
--+        +-- Pseudodon_vondembuschianus
  |  +-----+                             
  |  |     +-- Pseudodon_vagulus         
  +--+                                   
     |  +----- Sundadontina_tumida       
     +--+                                
        |  +-- Nyeinchanconcha_nyeinchani
        +--+                             
           +-- Bineurus_exilis

Konopleva et al., 2017 [Publ 3807], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 30 5 4 -- 
        +-- Nyeinchanconcha_nyeinchani
  +-----+                             
  |     +-- Bineurus_exilis           
--+                                   
  |  +----- Pilsbryoconcha_carinifera ** 
  +--+                                
     |  +-- Pseudodon_vagulus         
     +--+                             
        +-- Pseudodon_vondembuschianus

Jeratthitikul et al., 2019 [Publ 3911], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 21 0 0 -- 
-- Pilsbryoconcha_exilis

Froufe et al., 2019 [Publ 3936], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 44 1 1 -- 
  +-- Pilsbryoconcha_exilis **     
--+                             
  +-- Pseudodon_vondembuschianus

Froufe et al., 2019 [Publ 3936], fig. 3-M

iT tT bi riS oT roT roS iD
1 1 0 1 18 1 1 -- 
  +-- Pilsbryoconcha_exilis **     
--+                             
  +-- Pseudodon_vondembuschianus

Froufe et al., 2019 [Publ 3936], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 18 1 1 -- 
  +-- Pilsbryoconcha_exilis **     
--+                             
  +-- Pseudodon_vondembuschianus

Froufe et al., 2019 [Publ 3936], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 44 1 1 -- 
  +-- Pilsbryoconcha_exilis **     
--+                             
  +-- Pseudodon_vondembuschianus

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iT tT bi riS oT roT roS iD
1 1 0 1 56 3 2 -- 
  +-------- Bineurus_anodontinum      
--+                                   
  |  +----- Pilsbryoconcha_exilis **     
  +--+                                
     |  +-- Pseudodon_vondembuschianus
     +--+                             
        +-- Pseudodon_inoscularis

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 56 3 2 -- 
  +-------- Pilsbryoconcha_exilis **     
--+                                   
  |  +----- Bineurus_anodontinum      
  +--+                                
     |  +-- Pseudodon_vondembuschianus
     +--+                             
        +-- Pseudodon_inoscularis

Zieritz et al., 2020 [Publ 4102], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 30 0 0 -- 
-- Pilsbryoconcha_carinifera

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 53 1 1 -- 
  +-- Pilsbryoconcha_exilis **     
--+                             
  +-- Pseudodon_vondembuschianus

Chen et al., 2020 [Publ 4170], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 33 1 1 -- 
  +-- Pseudodon_vondembuschianus
--+                             
  +-- Pilsbryoconcha_exilis **

Wu et al., 2021 [Publ 4215], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 23 2 2 -- 
  +-- Pseudodon_vagulus        
--+                            
  +-- Pilsbryoconcha_carinifera **

Zieritz et al., 2021 [Publ 4218], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 31 6 4 -- 
  +-------------- Pilsbryoconcha_exilis **     
  |                                         
--+           +-- Pseudodon_mekongi         
  |  +--------+                             
  |  |        +-- Pseudodon_vondembuschianus
  +--+                                      
     |  +-------- Sundadontina              
     +--+                                   
        |  +----- Thaiconcha_callifera      
        +--+                                
           |  +-- Bineurus_loeiensis        
           +--+                             
              +-- Bineurus_anodontinum

Zieritz et al., 2021 [Publ 4231], fig. 3-F, 5

iT tT bi riS oT roT roS iD
1 1 0 1 48 1 1 -- 
  +-- Pilsbryoconcha_exilis **     
--+                             
  +-- Pseudodon_vondembuschianus

Zieritz et al., 2021 [Publ 4231], fig. 3-M, 4

iT tT bi riS oT roT roS iD
1 1 0 1 22 1 1 -- 
  +-- Pilsbryoconcha_exilis **     
--+                             
  +-- Pseudodon_vondembuschianus

Wu et al., 2022 [Publ 4238], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 73 1 1 -- 
  +-- Pilsbryoconcha_exilis **     
--+                             
  +-- Pseudodon_vondembuschianus

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 35 1 1 -- 
  +-- Pseudodon_vondembuschianus
--+                             
  +-- Pilsbryoconcha_exilis **

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iT tT bi riS oT roT roS iD
1 1 0 1 38 1 1 -- 
  +-- Pseudodon_vondembuschianus
--+                             
  +-- Pilsbryoconcha_exilis **

Dai et al., 2023 [Publ 4285], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 52 1 1 -- 
  +-- Pseudodon_vondembuschianus
--+                             
  +-- Pilsbryoconcha_exilis **

Jeratthitikul & Sutcharit, 2023 [Publ 4289], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 12 10 6 -- 
  +----- Pseudodon_vondembuschianus
--+                                
  |  +-- Pilsbryoconcha_exilis **     
  +--+                             
     +-- Thaiconcha_callifera

Kongim et al., 2023 [Publ 4290], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 14 1 1 -- 
  +-- Namkongnaia_inkhavilayi
--+                          
  +-- Pilsbryoconcha_exilis **

Page updated: 11:30:03 Wed 22 Nov 2023
 
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