MUSSELp
 
MUSSELpdb
 
 
Taxonomy
 
 
Geography
 
 
 

Page last updated
Wed 31 May 2023

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Gonideinae | Contradentini

genus Lens Simpson, 1900: cladomics
     13 valid species

Cladomics Summary

number of species = 13

analyzed in 41 cladograms in 27 publications
maximum number of terminal species in an analysis = 10, mean = 3.34

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 22
maximum number of included species = 10, mean = 5.36
proportion of cladograms that support basic monophyly = 0.91


Pfeiffer et al., 2021, fig. 12

iT tT bi riS oT roT roS iD
10 10 7 10 1 1 1 0.0

*** Top Monophyly Cladogram ***

  +-------------- Solenaia_emarginata
  |                                  
  |           +-- Lens_contradens **    
  |  +--------+                      
--+  |        +-- Lens_micropterus **   
  |  |                               
  |  |        +-- Lens_inornatus **     
  |  +--------+                      
  +--+        +-- Lens_misellus **      
     |                               
     |     +----- Lens_maenamensis **   
     |  +--+                         
     |  |  |  +-- Lens_rolfbrandti **   
     |  |  +--+                      
     +--+     +-- Lens_eximius **       
        |                            
        |  +----- Lens_pallegoixi **    
        +--+                         
           |  +-- Lens_comptus **       
           +--+                      
              +-- Lens_novoselovi **    

Pfeiffer et al., 2021, fig. 9

iT tT bi riS oT roT roS iD
8 8 6 8 21 7 4 5.0

*** Top Sister-Group Cladogram ***

  +----------------- Yaukthwa_paiensis  
  |                                     
  |  +-------------- Solenaia_emarginata
--+  |                                  
  |  |     +-------- Lens_misellus **      
  |  |  +--+                            
  +--+  |  |  +----- Lens_inornatus **     
     |  |  +--+                         
     |  |     |  +-- Lens_contradens **    
     |  |     +--+                      
     +--+        +-- Lens_micropterus **   
        |                               
        |        +-- Lens_maenamensis **   
        |     +--+                      
        |     |  +-- Lens_eximius **       
        +-----+                         
              |  +-- Lens_pallegoixi **    
              +--+                      
                 +-- Lens_comptus **       

Konopleva et al., 2019, fig. 3

iT tT bi riS oT roT roS iD
9 9 6 9 26 10 3 1.0
        +----------- Yaukthwa_elongatula   
        |                                  
  +-----+        +-- Yaukthwa_inlenensis   
  |     |  +-----+                         
  |     |  |     +-- Yaukthwa_paiensis     
  |     +--+                               
  |        |  +----- Yaukthwa_panhai       
  |        +--+                            
  |           |  +-- Yaukthwa_nesemanni    
  |           +--+                         
--+              +-- Yaukthwa_dalliana     
  |                                        
  |        +-------- Trapezoideus_foliaceus
  |  +-----+                               
  |  |     |  +----- Physunio_superbus     
  |  |     +--+                            
  |  |        |  +-- Physunio_modelli      
  |  |        +--+                         
  |  |           +-- Physunio              
  +--+                                     
     |        +----- Lens_misellus **         
     |  +-----+                            
     |  |     |  +-- Lens_contradens **       
     |  |     +--+                         
     |  |        +-- Lens_micropterus **      
     +--+                                  
        |     +----- Lens_pallegoixi **       
        |  +--+                            
        |  |  |  +-- Lens_novoselovi **       
        |  |  +--+                         
        +--+     +-- Lens_comptus **          
           |                               
           |  +----- Lens_maenamensis **      
           +--+                            
              |  +-- Lens_eximius **          
              +--+                         
                 +-- Lens_rolfbrandti **      

Bolotov et al., 2020, fig. 1

iT tT bi riS oT roT roS iD
9 9 6 9 112 14 3 1.0
                    +-- Trapezoideus_lenya     
              +-----+                          
              |     +-- Trapezoideus_foliaceus 
  +-----------+                                
  |           |  +----- Physunio_superbus      
  |           +--+                             
  |              |  +-- Physunio_modelli       
  |              +--+                          
  |                 +-- Physunio               
  |                                            
  |                 +-- Yaukthwa_elongatula    
  |     +-----------+                          
  |     |           +-- Yaukthwa_baniensis     
--+     |                                      
  |  +--+        +----- Yaukthwa_inlenensis    
  |  |  |  +-----+                             
  |  |  |  |     |  +-- Yaukthwa_avaensis      
  |  |  |  |     +--+                          
  |  |  +--+        +-- Yaukthwa_paiensis      
  |  |     |                                   
  |  |     |  +-------- Yaukthwa_panhai        
  |  |     +--+                                
  |  |        |  +----- Yaukthwa_dalliana      
  +--+        +--+                             
     |           |  +-- Yaukthwa_zayleymanensis
     |           +--+                          
     |              +-- Yaukthwa_nesemanni     
     |                                         
     |           +----- Lens_misellus **          
     |     +-----+                             
     |     |     |  +-- Lens_contradens **        
     |     |     +--+                          
     |     |        +-- Lens_micropterus **       
     +-----+                                   
           |     +----- Lens_pallegoixi **        
           |  +--+                             
           |  |  |  +-- Lens_comptus **           
           |  |  +--+                          
           +--+     +-- Lens_novoselovi **        
              |                                
              |  +----- Lens_maenamensis **       
              +--+                             
                 |  +-- Lens_eximius **           
                 +--+                          
                    +-- Lens_rolfbrandti **       

Bolotov et al., 2020, fig. S1

iT tT bi riS oT roT roS iD
9 9 6 9 106 14 3 1.0
                    +-- Trapezoideus_lenya     
  +-----------------+                          
  |                 +-- Trapezoideus_foliaceus 
  |                                            
  |              +----- Lens_misellus **          
  |        +-----+                             
  |        |     |  +-- Lens_contradens **        
  |        |     +--+                          
  |        |        +-- Lens_micropterus **       
--+  +-----+                                   
  |  |     |     +----- Lens_maenamensis **       
  |  |     |  +--+                             
  |  |     |  |  |  +-- Lens_eximius **           
  |  |     |  |  +--+                          
  |  |     +--+     +-- Lens_rolfbrandti **       
  |  |        |                                
  |  |        |  +----- Lens_pallegoixi **        
  +--+        +--+                             
     |           |  +-- Lens_novoselovi **        
     |           +--+                          
     |              +-- Lens_comptus **           
     |                                         
     |              +-- Yaukthwa_elongatula    
     |  +-----------+                          
     |  |           +-- Yaukthwa_baniensis     
     |  |                                      
     +--+        +----- Yaukthwa_inlenensis    
        |  +-----+                             
        |  |     |  +-- Yaukthwa_avaensis      
        |  |     +--+                          
        +--+        +-- Yaukthwa_paiensis      
           |                                   
           |  +-------- Yaukthwa_panhai        
           +--+                                
              |  +----- Yaukthwa_dalliana      
              +--+                             
                 |  +-- Yaukthwa_zayleymanensis
                 +--+                          
                    +-- Yaukthwa_nesemanni     

Konopleva et al., 2023, fig. 8

iT tT bi riS oT roT roS iD
8 8 5 8 75 10 3 2.0
              +----- Physunio_modelli          
        +-----+                                
        |     |  +-- Physunio_superbus         
        |     +--+                             
        |        +-- Physunio_massini          
  +-----+                                      
  |     |        +-- Trapezoideus_thachiadensis
  |     |  +-----+                             
  |     |  |     +-- Trapezoideus_subviridis   
  |     +--+                                   
  |        |  +----- Trapezoideus_mitanensis   
  |        +--+                                
--+           |  +-- Trapezoideus_foliaceus    
  |           +--+                             
  |              +-- Trapezoideus_lenya        
  |                                            
  |              +-- Yaukthwa_nesemanni        
  |  +-----------+                             
  |  |           +-- Yaukthwa_elongatula       
  |  |                                         
  +--+           +-- Lens_contradens **           
     |  +--------+                             
     |  |        +-- Lens_misellus **             
     |  |                                      
     +--+     +----- Lens_pallegoixi **           
        |  +--+                                
        |  |  |  +-- Lens_novoselovi **           
        |  |  +--+                             
        +--+     +-- Lens_comptus **              
           |                                   
           |  +----- Lens_maenamensis **          
           +--+                                
              |  +-- Lens_eximius **              
              +--+                             
                 +-- Lens_rolfbrandti **          

Konopleva et al., 2019, fig. 1

iT tT bi riS oT roT roS iD
7 7 4 7 45 9 3 3.0
                 +----- Physunio_superbus     
  +--------------+                            
  |              |  +-- Physunio_modelli      
  |              +--+                         
--+                 +-- Physunio              
  |                                           
  |  +----------------- Trapezoideus_foliaceus
  |  |                                        
  +--+              +-- Lens_contradens **       
     |  +-----------+                         
     |  |           +-- Lens_micropterus **      
     +--+                                     
        |  +----------- Lens_misellus **         
        +--+                                  
           |  +-------- Lens_eximius **          
           +--+                               
              |  +----- Lens_pallegoixi **       
              +--+                            
                 |  +-- Lens_comptus **          
                 +--+                         
                    +-- Lens_novoselovi **       

Huang et al., 2019, fig. S4

iT tT bi riS oT roT roS iD
6 12 7 6 193 6 2 12.0
  +----------- Physunio_modelli      
  |                                  
  +----------- Lens **                  
  |                                  
  +----------- Physunio_superbus     
  |                                  
  |  +-------- Yaukthwa_inlenensis   
  +--+                               
--+  |  +----- Yaukthwa_panhai       
  |  +--+                            
  |     |  +-- Yaukthwa_nesemanni    
  |     +--+                         
  |        +-- Yaukthwa_rectangularis
  |                                  
  |        +-- Lens_comptus **          
  |  +-----+                         
  |  |     +-- Lens_novoselovi **       
  +--+                               
     |  +----- Lens_eximius **          
     +--+                            
        |  +-- Lens_contradens **       
        +--+                         
           +-- Lens_micropterus **      

Zieritz et al., 2020, fig. 4

iT tT bi riS oT roT roS iD
6 6 3 6 25 8 3 4.0
                 +-- Yaukthwa_paiensis     
           +-----+                         
           |     +-- Yaukthwa_inlenensis   
  +--------+                               
  |        |  +----- Yaukthwa_panhai       
  |        +--+                            
  |           |  +-- Yaukthwa_dalliana     
--+           +--+                         
  |              +-- Yaukthwa_nesemanni    
  |                                        
  |  +-------------- Trapezoideus_foliaceus
  |  |                                     
  +--+           +-- Lens_contradens **       
     |  +--------+                         
     |  |        +-- Lens_misellus **         
     +--+                                  
        |  +-------- Lens_eximius **          
        +--+                               
           |  +----- Lens_pallegoixi **       
           +--+                            
              |  +-- Lens_comptus **          
              +--+                         
                 +-- Lens_novoselovi **       

Bolotov et al., 2017, fig. S1

iT tT bi riS oT roT roS iD
5 5 2 5 133 7 2 5.0
           +----- Physunio              
  +--------+                            
  |        |  +-- Physunio_modelli      
  |        +--+                         
  |           +-- Physunio_superbus     
  |                                     
--+     +-------- Yaukthwa_inlenensis   
  |  +--+                               
  |  |  |  +----- Yaukthwa_panhai       
  |  |  +--+                            
  |  |     |  +-- Yaukthwa_rectangularis
  |  |     +--+                         
  +--+        +-- Yaukthwa_nesemanni    
     |                                  
     |        +-- Lens_contradens **       
     |  +-----+                         
     |  |     +-- Lens_micropterus **      
     +--+                               
        |  +----- Lens_eximius **          
        +--+                            
           |  +-- Lens_comptus **          
           +--+                         
              +-- Lens_novoselovi **       

Bolotov et al., 2017, fig. S1

iT tT bi riS oT roT roS iD
5 5 2 5 120 7 2 5.0
           +----- Physunio              
  +--------+                            
  |        |  +-- Physunio_superbus     
  |        +--+                         
  |           +-- Physunio_modelli      
  |                                     
--+     +-------- Yaukthwa_inlenensis   
  |  +--+                               
  |  |  |  +----- Yaukthwa_panhai       
  |  |  +--+                            
  |  |     |  +-- Yaukthwa_nesemanni    
  |  |     +--+                         
  +--+        +-- Yaukthwa_rectangularis
     |                                  
     |        +-- Lens_contradens **       
     |  +-----+                         
     |  |     +-- Lens_micropterus **      
     +--+                               
        |  +----- Lens_eximius **          
        +--+                            
           |  +-- Lens_comptus **          
           +--+                         
              +-- Lens_novoselovi **       

Bolotov et al., 2018, fig. 2

iT tT bi riS oT roT roS iD
5 11 0 5 80 6 2 12.0
  +-- Lens_eximius **       
  |                      
  +-- Physunio_modelli[1]
  |                      
  +-- Yaukthwa_nesemanni 
  |                      
  +-- Yaukthwa_panhai    
  |                      
  +-- Lens_comptus **       
  |                      
--+-- Lens_novoselovi **    
  |                      
  +-- Physunio_modelli[2]
  |                      
  +-- Physunio           
  |                      
  +-- Yaukthwa_inlenensis
  |                      
  +-- Lens_contradens **    
  |                      
  +-- Lens_micropterus **   

Huang et al., 2019, fig. 2

iT tT bi riS oT roT roS iD
5 5 2 5 194 7 2 5.0
           +----- Physunio              
  +--------+                            
  |        |  +-- Physunio_modelli      
  |        +--+                         
  |           +-- Physunio_superbus     
  |                                     
--+     +-------- Yaukthwa_inlenensis   
  |  +--+                               
  |  |  |  +----- Yaukthwa_panhai       
  |  |  +--+                            
  |  |     |  +-- Yaukthwa_nesemanni    
  |  |     +--+                         
  +--+        +-- Yaukthwa_rectangularis
     |                                  
     |        +-- Lens_contradens **       
     |  +-----+                         
     |  |     +-- Lens_micropterus **      
     +--+                               
        |  +----- Lens_eximius **          
        +--+                            
           |  +-- Lens_comptus **          
           +--+                         
              +-- Lens_novoselovi **       

Huang et al., 2019, fig. S3

iT tT bi riS oT roT roS iD
5 5 2 5 192 7 2 5.0
           +----- Physunio              
  +--------+                            
  |        |  +-- Physunio_superbus     
  |        +--+                         
  |           +-- Physunio_modelli      
  |                                     
--+     +-------- Yaukthwa_inlenensis   
  |  +--+                               
  |  |  |  +----- Yaukthwa_panhai       
  |  |  +--+                            
  |  |     |  +-- Yaukthwa_nesemanni    
  |  |     +--+                         
  +--+        +-- Yaukthwa_rectangularis
     |                                  
     |        +-- Lens_contradens **       
     |  +-----+                         
     |  |     +-- Lens_micropterus **      
     +--+                               
        |  +----- Lens_eximius **          
        +--+                            
           |  +-- Lens_novoselovi **       
           +--+                         
              +-- Lens_comptus **          

Bolotov et al., 2017, fig. S2

iT tT bi riS oT roT roS iD
4 4 2 4 134 7 2 6.0
        +-------- Lens_contradens **       
  +-----+                               
  |     |  +----- Lens_eximius **          
  |     +--+                            
  |        |  +-- Lens_comptus **          
  |        +--+                         
--+           +-- Lens_novoselovi **       
  |                                     
  |        +----- Physunio              
  |  +-----+                            
  |  |     |  +-- Physunio_modelli      
  |  |     +--+                         
  +--+        +-- Physunio_superbus     
     |                                  
     |  +-------- Yaukthwa_inlenensis   
     +--+                               
        |  +----- Yaukthwa_panhai       
        +--+                            
           |  +-- Yaukthwa_rectangularis
           +--+                         
              +-- Yaukthwa_nesemanni    

Jeratthitkul et al., 2021, fig. 4

iT tT bi riS oT roT roS iD
4 4 2 4 17 6 3 6.0
        +-------- Trapezoideus_foliaceus
  +-----+                               
  |     |  +----- Physunio_superbus     
  |     +--+                            
  |        |  +-- Physunio_pellucidus   
  |        +--+                         
--+           +-- Physunio_modelli      
  |                                     
  |           +-- Yaukthwa_nesemanni    
  |  +--------+                         
  |  |        +-- Yaukthwa_panhai       
  +--+                                  
     |  +-------- Lens_contradens **       
     +--+                               
        |  +----- Lens_novoselovi **       
        +--+                            
           |  +-- Lens_rolfbrandti **      
           +--+                         
              +-- Lens_maenamensis **      

Jeratthitikul et al., 2019, fig. 4

iT tT bi riS oT roT roS iD
3 3 1 3 5 2 1 7.0
        +-- Physunio_modelli 
  +-----+                    
  |     +-- Physunio_superbus
--+                          
  |  +----- Lens_contradens **  
  +--+                       
     |  +-- Lens_maenamensis ** 
     +--+                    
        +-- Lens_rolfbrandti ** 

Lopes-Lima et al., 2017, fig. 1-2

iT tT bi riS oT roT roS iD
2 2 0 2 79 1 1 --
  +-- Lens_contradens **
--+                  
  +-- Lens_comptus **   

Pfeiffer et al., 2018, fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 6 2 2 --
  +----- Physunio       
  |                     
--+----- Trapezoideus   
  |                     
  |  +-- Lens_comptus **   
  +--+                  
     +-- Lens_contradens **

Zieritz et al., 2021, fig. 5

iT tT bi riS oT roT roS iD
2 2 0 2 29 5 3 --
  +----------- Trapezoideus_foliaceus
  |                                  
--+        +-- Lens_contradens **       
  |  +-----+                         
  |  |     +-- Lens_eximius **          
  +--+                               
     |  +----- Yaukthwa_elongatula   
     +--+                            
        |  +-- Yaukthwa_inlenensis   
        +--+                         
           +-- Yaukthwa_paiensis     

Dai et al., 2023, fig. 3-BI

iT tT bi riS oT roT roS iD
2 2 0 2 42 1 1 --
  +----- Physunio_superbus
--+                       
  |  +-- Lens_contradens **  
  +--+                    
     +-- Lens_comptus **     

Dai et al., 2023, fig. 3-ML

iT tT bi riS oT roT roS iD
2 2 0 2 42 1 1 --
  +----- Physunio_superbus
--+                       
  |  +-- Lens_contradens **  
  +--+                    
     +-- Lens_comptus **     

Graf & Cummings, 2006, fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 46 0 0 --
-- Lens_contradens

Graf, 2002, fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 17 0 0 --
-- Lens_contradens

Hoeh et al., 2009, fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 46 0 0 --
-- Lens_contradens

Hoeh et al., 2009, fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 45 0 0 --
-- Lens_contradens

Hoeh et al., 2009, fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 45 0 0 --
-- Lens_contradens

Whelan et al., 2011, fig. 1 ML

iT tT bi riS oT roT roS iD
1 1 0 1 35 0 0 --
-- Lens_contradens

Whelan et al., 2011, fig. 1 MP

iT tT bi riS oT roT roS iD
1 1 0 1 35 0 0 --
-- Lens_contradens

Pfeiffer & Graf, 2015, fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 39 2 1 --
  +----- Lens_micropterus **
--+                      
  |  +-- Physunio        
  +--+                   
     +-- Physunio_modelli

Konopleva et al., 2017, fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 30 2 1 --
  +----- Lens_micropterus **
--+                      
  |  +-- Physunio        
  +--+                   
     +-- Physunio_modelli

Pfeiffer et al., 2019, fig. 4-5

iT tT bi riS oT roT roS iD
1 1 0 1 56 4 4 --
  +----- Yaukthwa_paiensis  
--+                         
  |  +-- Lens_contradens **    
  +--+                      
     +-- Solenaia_emarginata

Pfeiffer et al., 2019, fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 56 4 4 --
  +----- Yaukthwa_paiensis  
--+                         
  |  +-- Lens_contradens **    
  +--+                      
     +-- Solenaia_emarginata

Lopes-Lima et al., 2020, fig. 2-4

iT tT bi riS oT roT roS iD
1 1 0 1 121 2 2 --
-- Lens_contradens

Jeratthitikul et al., 2021, fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 36 2 2 --
  +----- Lens_contradens **       
--+                            
  |  +-- Trapezoideus_foliaceus
  +--+                         
     +-- Physunio_superbus     

Jeratthitikul et al., 2021, fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 36 2 2 --
  +----- Lens_contradens **       
--+                            
  |  +-- Trapezoideus_foliaceus
  +--+                         
     +-- Physunio_superbus     

Zieritz et al., 2021, fig. 3-F, 5

iT tT bi riS oT roT roS iD
1 1 0 1 48 1 1 --
  +-- Lens_contradens **  
--+                    
  +-- Physunio_superbus

Zieritz et al., 2021, fig. 3-M, 4

iT tT bi riS oT roT roS iD
1 1 0 1 22 1 1 --
  +-- Lens_contradens **  
--+                    
  +-- Physunio_superbus

Jeratthitikul et al., 2022, fig. S1, 1

iT tT bi riS oT roT roS iD
1 1 0 1 75 2 2 --
  +----- Lens_contradens **       
--+                            
  |  +-- Trapezoideus_foliaceus
  +--+                         
     +-- Physunio_superbus     

Jeratthitikul et al., 2022, fig. S2, 2

iT tT bi riS oT roT roS iD
1 1 0 1 75 2 2 --
  +----- Lens_contradens **       
--+                            
  |  +-- Trapezoideus_foliaceus
  +--+                         
     +-- Physunio_superbus     

Dai et al., 2023, fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 52 1 1 --
  +-- Lens_contradens **  
--+                    
  +-- Physunio_superbus

Page updated: 14:10:29 Wed 31 May 2023

 
NSF icon MUSSEL icon
"Making the world a better place, one mollusk at a time."