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Page last updated
Wed 04 Dec 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Unionoida | Unionoidea | Unionidae | Gonideinae | Pseudodontini | Pseudodontina

genus Pseudodon Gould, 1844: cladomics
     10 valid species

Cladomics Summary

number of species = 10

analyzed in 82 cladograms in 46 publications
maximum number of terminals in an analysis = 7, mean = 1.95

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 32
maximum number of included terminals = 7, mean = 3.44
proportion of cladograms that support basic monophyly = 0.91

Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ tree

Jeratthitikul et al., 2024 [Publ 4387], fig. 1

iT tT bi riS oT roT roS iD
6 6 5 7 52 29 9 0.0

*** Top Monophyly Cladogram *** *** Top Sister-Group Cladogram ***

              +-- Pseudodon_inoscularis **        
     +--------+                                
     |        +-- Pseudodon_lenyanensis **        
     |                                         
  +--+        +-- Pseudodon_cambodjensis **       
  |  |  +-----+                                
  |  |  |     +-- Pseudodon_vagulus **            
  |  +--+                                      
  |     |  +----- Pseudodon_vondembuschianus__2 **
  |     +--+                                   
  |        |  +-- Pseudodon_vondembuschianus__1 **
  |        +--+                                
--+           +-- Pseudodon_mekongi **            
  |                                            
  |     +-------- Sundadontina                 
  |  +--+                                      
  |  |  |  +----- Nyeinchanconcha              
  |  |  +--+                                   
  |  |     |  +-- Thaiconcha                   
  |  |     +--+                                
  +--+        +-- Bineurus                     
     |                                         
     |        +-- Isannaia                     
     |  +-----+                                
     |  |     +-- Pilsbryoconcha               
     +--+                                      
        |  +----- Namkongnaia                  
        +--+                                   
           |  +-- Lannanaia                    
           +--+                                
              +-- Songkhlanaia

Zieritz et al., 2024 [Publ 4381], fig. 8A

iT tT bi riS oT roT roS iD
7 7 5 7 1 1 1 4.0 
  +-------------- Namkongnaia               
  |                                         
  |           +-- Pseudodon_vondembuschianus **
--+  +--------+                             
  |  |        +-- Pseudodon_mekongi **         
  |  |                                      
  +--+        +-- Pseudodon_lenyanensis **     
     |  +-----+                             
     |  |     +-- Pseudodon_inoscularis **     
     +--+                                   
        |  +----- Pseudodon_vagulus **         
        +--+                                
           |  +-- Pseudodon_walpolei **        
           +--+                             
              +-- Pseudodon_cambodjensis **

Bolotov et al., 2023 [Publ 4312], fig. 2

iT tT bi riS oT roT roS iD
6 6 5 7 33 28 7 3.0 
  +----------------------- Pilsbryoconcha               
  |                                                     
  |  +-------------------- Songkhlanaia                 
  |  |                                                  
--+  |        +----------- Namkongnaia                  
  |  |        |                                         
  |  |        |     +----- Pseudodon_vondembuschianus__2 **
  |  |  +-----+  +--+                                   
  |  |  |     |  |  |  +-- Pseudodon_vondembuschianus__1 **
  +--+  |     |  |  +--+                                
     |  |     |  |     +-- Pseudodon_mekongi **            
     |  |     +--+                                      
     |  |        |     +-- Pseudodon_vagulus **            
     |  |        |  +--+                                
     |  |        |  |  +-- Pseudodon_cambodjensis **       
     |  |        +--+                                   
     +--+           |  +-- Pseudodon_lenyanensis **        
        |           +--+                                
        |              +-- Pseudodon_inoscularis **        
        |                                               
        |           +----- Nyeinchanconcha              
        |  +--------+                                   
        |  |        |  +-- Thaiconcha                   
        |  |        +--+                                
        |  |           +-- Bineurus                     
        |  |                                            
        +--+     +-------- Sundadontina_sulcata         
           |  +--+                                      
           |  |  |  +----- Sundadontina_harmandi        
           |  |  +--+                                   
           |  |     |  +-- Sundadontina_brandti         
           +--+     +--+                                
              |        +-- Sundadontina_taskaevi        
              |                                         
              |  +-------- Sundadontina_tumida          
              +--+                                      
                 |  +----- Sundadontina_tanintharyiensis
                 +--+                                   
                    |  +-- Sundadontina_cumingii        
                    +--+                                
                       +-- Sundadontina_plugpomenica

Bolotov et al., 2023 [Publ 4312], fig. S1

iT tT bi riS oT roT roS iD
6 6 4 6 33 28 7 1.0 
  +-------------------------- Pilsbryoconcha               
  |                                                        
  |                    +----- Nyeinchanconcha              
  |  +-----------------+                                   
--+  |                 |  +-- Thaiconcha                   
  |  |                 +--+                                
  |  |                    +-- Bineurus                     
  |  |                                                     
  +--+  +-------------------- Songkhlanaia                 
     |  |                                                  
     |  |        +----------- Namkongnaia                  
     |  |        |                                         
     |  |        |        +-- Pseudodon_inoscularis **        
     |  |  +-----+  +-----+                                
     +--+  |     |  |     +-- Pseudodon_lenyanensis **        
        |  |     |  |                                      
        |  |     +--+     +-- Pseudodon_vagulus **            
        |  |        |  +--+                                
        |  |        |  |  +-- Pseudodon_cambodjensis **       
        |  |        +--+                                   
        +--+           |  +-- Pseudodon_vondembuschianus **   
           |           +--+                                
           |              +-- Pseudodon_mekongi **            
           |                                               
           |  +-------------- Sundadontina_tumida          
           |  |                                            
           |  |        +----- Sundadontina_tanintharyiensis
           +--+  +-----+                                   
              |  |     |  +-- Sundadontina_cumingii        
              |  |     +--+                                
              +--+        +-- Sundadontina_plugpomenica    
                 |                                         
                 |  +-------- Sundadontina_sulcata         
                 +--+                                      
                    |  +----- Sundadontina_harmandi        
                    +--+                                   
                       |  +-- Sundadontina_brandti         
                       +--+                                
                          +-- Sundadontina_taskaevi

Bolotov et al., 2023 [Publ 4312], fig. S2

iT tT bi riS oT roT roS iD
6 6 4 6 35 30 7 5.0 
  +-------------------------------- Songkhlanaia                   
  |                                                                
  |              +----------------- Nyeinchanconcha                
  |              |                                                 
  |  +-----------+  +-------------- Sundadontina_tumida            
  |  |           |  |                                              
  |  |           |  |        +----- Sundadontina_tanintharyiensis  
--+  |           +--+  +-----+                                     
  |  |              |  |     |  +-- Sundadontina_plugpomenica      
  |  |              |  |     +--+                                  
  |  |              +--+        +-- Sundadontina_cumingii          
  |  |                 |                                           
  |  |                 |  +-------- Sundadontina_sulcata           
  |  |                 +--+                                        
  |  |                    |  +----- Sundadontina_harmandi          
  +--+                    +--+                                     
     |                       |  +-- Sundadontina_taskaevi          
     |                       +--+                                  
     |                          +-- Sundadontina_brandti           
     |                                                             
     |                       +----- Namkongnaia                    
     |  +--------------------+                                     
     |  |                    |  +-- Thaiconcha                     
     |  |                    +--+                                  
     |  |                       +-- Bineurus                       
     |  |                                                          
     |  |              +----------- Pseudodon_vagulus **              
     +--+              |                                           
        |  +-----------+        +-- Pseudodon_lenyanensis **          
        |  |           |  +-----+                                  
        |  |           |  |     +-- Pseudodon_inoscularis **          
        |  |           +--+                                        
        |  |              |  +----- Pseudodon_cambodjensis **         
        |  |              +--+                                     
        +--+                 |  +-- Pseudodon_vondembuschianus **     
           |                 +--+                                  
           |                    +-- Pseudodon_mekongi **              
           |                                                       
           |  +-------------------- Pilsbryoconcha_carinifera      
           |  |                                                    
           +--+  +----------------- Pilsbryoconcha_linguaeformis__2
              |  |                                                 
              +--+  +-------------- Pilsbryoconcha_linguaeformis__1
                 |  |                                              
                 +--+           +-- Pilsbryoconcha_schomburgki     
                    |  +--------+                                  
                    |  |        +-- Pilsbryoconcha_exilis          
                    +--+                                           
                       |  +-------- Pilsbryoconcha_hoikaab         
                       +--+                                        
                          |  +----- Pilsbryoconcha_mekongiana      
                          +--+                                     
                             |  +-- Pilsbryoconcha_acuta           
                             +--+                                  
                                +-- Pilsbryoconcha_kittitati

Bolotov et al., 2020 [Publ 4100], fig. S1

iT tT bi riS oT roT roS iD
4 4 2 4 108 13 5 3.0 
  +-------------------- Bineurus                  
  |                                               
--+  +----------------- Nyeinchanconcha           
  |  |                                            
  |  |           +----- Thaiconcha                
  +--+  +--------+                                
     |  |        |  +-- Pilsbryoconcha_exilis__2  
     |  |        +--+                             
     +--+           +-- Pilsbryoconcha_exilis__1  
        |                                         
        |  +----------- Sundadontina              
        +--+                                      
           |  +-------- Pseudodon_lenyanensis **     
           +--+                                   
              |  +----- Pseudodon_vagulus **         
              +--+                                
                 |  +-- Pseudodon_vondembuschianus **
                 +--+                             
                    +-- Pseudodon_mekongi **

Jeratthitikul et al., 2021 [Publ 4219], fig. 1

iT tT bi riS oT roT roS iD
4 4 2 4 32 14 6 3.0 
                    +-- Pilsbryoconcha_exilis       
  +-----------------+                               
  |                 +-- Pilsbryoconcha_linguaeformis
  |                                                 
--+              +----- Nyeinchanconcha             
  |  +-----------+                                  
  |  |           |  +-- Thaiconcha                  
  |  |           +--+                               
  +--+              +-- Bineurus                    
     |                                              
     |  +-------------- Sundadontina                
     +--+                                           
        |  +----------- Namkongnaia                 
        +--+                                        
           |  +-------- Pseudodon_lenyanensis **       
           +--+                                     
              |  +----- Pseudodon_vagulus **           
              +--+                                  
                 |  +-- Pseudodon_vondembuschianus **  
                 +--+                               
                    +-- Pseudodon_mekongi **

Konopleva et al., 2023 [Publ 4262], fig. 8

iT tT bi riS oT roT roS iD
4 4 2 4 78 22 7 3.0 
                    +-- Pilsbryoconcha_schomburgki  
              +-----+                               
              |     +-- Pilsbryoconcha_exilis__1    
  +-----------+                                     
  |           |  +----- Pilsbryoconcha_linguaeformis
  |           +--+                                  
  |              |  +-- Pilsbryoconcha_exilis__2    
--+              +--+                               
  |                 +-- Pilsbryoconcha_carinifera   
  |                                                 
  |  +----------------- Songkhlanaia                
  |  |                                              
  +--+  +-------------- Namkongnaia                 
     |  |                                           
     |  |     +-------- Pseudodon_lenyanensis **       
     +--+  +--+                                     
        |  |  |  +----- Pseudodon_vagulus **           
        |  |  +--+                                  
        |  |     |  +-- Pseudodon_vondembuschianus **  
        +--+     +--+                               
           |        +-- Pseudodon_mekongi **           
           |                                        
           |  +-------- Sundadontina                
           +--+                                     
              |  +----- Nyeinchanconcha             
              +--+                                  
                 |  +-- Thaiconcha                  
                 +--+                               
                    +-- Bineurus

Bolotov et al., 2020 [Publ 4100], fig. 1

iT tT bi riS oT roT roS iD
4 4 2 4 114 13 5 3.0 
  +-------------------- Bineurus                  
  |                                               
--+  +----------------- Nyeinchanconcha           
  |  |                                            
  |  |           +----- Thaiconcha                
  +--+  +--------+                                
     |  |        |  +-- Pilsbryoconcha_exilis__2  
     |  |        +--+                             
     +--+           +-- Pilsbryoconcha_exilis__1  
        |                                         
        |  +----------- Sundadontina              
        +--+                                      
           |  +-------- Pseudodon_lenyanensis **     
           +--+                                   
              |  +----- Pseudodon_vagulus **         
              +--+                                
                 |  +-- Pseudodon_vondembuschianus **
                 +--+                             
                    +-- Pseudodon_mekongi **

Jeratthitikul et al., 2021 [Publ 4219], fig. 2

iT tT bi riS oT roT roS iD
4 4 2 4 32 14 6 3.0 
                    +-- Pilsbryoconcha_exilis       
  +-----------------+                               
  |                 +-- Pilsbryoconcha_linguaeformis
  |                                                 
  |                 +-- Nyeinchanconcha             
  |              +--+                               
--+              |  +-- Sundadontina_cumingii       
  |  +-----------+                                  
  |  |           |  +-- Thaiconcha                  
  |  |           +--+                               
  |  |              +-- Bineurus                    
  +--+                                              
     |           +----- Sundadontina_tumida         
     |  +--------+                                  
     |  |        |  +-- Sundadontina_brandti        
     |  |        +--+                               
     +--+           +-- Sundadontina_taskaevi       
        |                                           
        |  +----------- Namkongnaia                 
        +--+                                        
           |  +-------- Pseudodon_lenyanensis **       
           +--+                                     
              |  +----- Pseudodon_vagulus **           
              +--+                                  
                 |  +-- Pseudodon_vondembuschianus **  
                 +--+                               
                    +-- Pseudodon_mekongi **

Konopleva et al., 2021 [Publ 4213], fig. 1

iT tT bi riS oT roT roS iD
4 4 2 4 23 15 6 3.0 
              +----- Pilsbryoconcha_exilis__2  
  +-----------+                                
  |           |  +-- Pilsbryoconcha_exilis__1  
  |           +--+                             
  |              +-- Namkongnaia               
--+                                            
  |           +----- Nyeinchanconcha           
  |  +--------+                                
  |  |        |  +-- Thaiconcha                
  |  |        +--+                             
  +--+           +-- Bineurus                  
     |                                         
     |  +----------- Sundadontina              
     +--+                                      
        |  +-------- Pseudodon_lenyanensis **     
        +--+                                   
           |  +----- Pseudodon_vagulus **         
           +--+                                
              |  +-- Pseudodon_vondembuschianus **
              +--+                             
                 +-- Pseudodon_mekongi **

Bolotov et al., 2017 [Publ 3761], fig. S2

iT tT bi riS oT roT roS iD
4 7 5 4 135 9 5 6.0 
     +----- Pseudodon_vagulus **         
  +--+                                
  |  |  +-- Pseudodon_mekongi **         
  |  +--+                             
  |     +-- Pseudodon_vondembuschianus **
--+                                   
  |     +-- Pseudodon_inoscularis **     
  |  +--+                             
  |  |  +-- Sundadontina_tumida       
  +--+                                
     |  +-- Nyeinchanconcha           
     +--+                             
        +-- Sundadontina_cumingii

Dai et al., 2023 [Publ 4285], fig. 3-BI

iT tT bi riS oT roT roS iD
3 3 1 3 42 11 5 4.0 
           +-- Nyeinchanconcha      
  +--------+                        
  |        +-- Bineurus             
  |                                 
  |        +-- Thaiconcha           
--+--------+                        
  |        +-- Sundadontina         
  |                                 
  |  +-------- Pilsbryoconcha       
  +--+                              
     |  +----- Pseudodon_lenyanensis **
     +--+                           
        |  +-- Pseudodon_mekongi **    
        +--+                        
           +-- Pseudodon_vagulus **

Huang et al., 2019 [Publ 3843], fig. S3

iT tT bi riS oT roT roS iD
3 3 2 4 193 9 5 4.0 
  +----------------- Pilsbryoconcha               
  |                                               
  |              +-- Nyeinchanconcha              
  |           +--+                                
--+           |  +-- Sundadontina_cumingii        
  |  +--------+                                   
  |  |        |  +-- Thaiconcha                   
  |  |        +--+                                
  +--+           +-- Bineurus                     
     |                                            
     |  +----------- Sundadontina_tumida          
     +--+                                         
        |  +-------- Pseudodon_vagulus **            
        +--+                                      
           |  +----- Pseudodon_vondembuschianus__1 **
           +--+                                   
              |  +-- Pseudodon_mekongi **            
              +--+                                
                 +-- Pseudodon_vondembuschianus__2 **

Dai et al., 2024 [Publ 4389], fig. 3

iT tT bi riS oT roT roS iD
3 3 1 3 44 8 5 4.0 
  +-------------- Pilsbryoconcha            
  |                                         
  |        +----- Pseudodon_cambodjensis **    
--+  +-----+                                
  |  |     |  +-- Pseudodon_mekongi **         
  |  |     +--+                             
  +--+        +-- Pseudodon_vondembuschianus **
     |                                      
     |  +-------- Sundadontina              
     +--+                                   
        |  +----- Nyeinchanconcha           
        +--+                                
           |  +-- Bineurus                  
           +--+                             
              +-- Thaiconcha

Konopleva et al., 2019 [Publ 3900], fig. 1

iT tT bi riS oT roT roS iD
3 8 0 3 49 5 2 10.0 
  +-- Bineurus_mouhotii         
  |                             
  +-- Pseudodon_vagulus **         
  |                             
  +-- Pseudodon_mekongi **         
  |                             
  +-- Pilsbryoconcha_exilis     
--+                             
  +-- Pilsbryoconcha_carinifera 
  |                             
  +-- Bineurus_loeiensis        
  |                             
  +-- Bineurus_exilis           
  |                             
  +-- Pseudodon_vondembuschianus **

Bolotov et al., 2017 [Publ 3761], fig. S1

iT tT bi riS oT roT roS iD
3 3 1 3 136 9 5 4.0 
  +-------------- Pilsbryoconcha            
  |                                         
  |     +-------- Sundadontina_tumida       
  |  +--+                                   
--+  |  |  +----- Pseudodon_vagulus **         
  |  |  +--+                                
  |  |     |  +-- Pseudodon_mekongi **         
  |  |     +--+                             
  +--+        +-- Pseudodon_vondembuschianus **
     |                                      
     |        +-- Nyeinchanconcha           
     |     +--+                             
     |     |  +-- Sundadontina_cumingii     
     +-----+                                
           |  +-- Thaiconcha                
           +--+                             
              +-- Bineurus

Huang et al., 2019 [Publ 3843], fig. S4

iT tT bi riS oT roT roS iD
3 3 1 3 195 9 5 4.0 
  +-------- Nyeinchanconcha           
  |                                   
  +-------- Thaiconcha                
  |                                   
  +-------- Sundadontina_cumingii     
  |                                   
  +-------- Sundadontina_tumida__1    
--+                                   
  +-------- Sundadontina_tumida__2    
  |                                   
  +-------- Pilsbryoconcha            
  |                                   
  +-------- Bineurus                  
  |                                   
  |  +----- Pseudodon_vagulus **         
  +--+                                
     |  +-- Pseudodon_mekongi **         
     +--+                             
        +-- Pseudodon_vondembuschianus **

Bolotov et al., 2018 [Publ 3826], fig. 2

iT tT bi riS oT roT roS iD
3 8 0 3 80 5 2 10.0 
  +-- Bineurus_mouhotii         
  |                             
  +-- Pseudodon_vagulus **         
  |                             
  +-- Pseudodon_mekongi **         
  |                             
  +-- Pilsbryoconcha_exilis     
--+                             
  +-- Pilsbryoconcha_carinifera 
  |                             
  +-- Bineurus_loeiensis        
  |                             
  +-- Bineurus_exilis           
  |                             
  +-- Pseudodon_vondembuschianus **

Zieritz et al., 2024 [Publ 4381], fig. 2

iT tT bi riS oT roT roS iD
3 3 1 3 43 5 4 6.0 
        +----- Sundadontina              
  +-----+                                
  |     |  +-- Thaiconcha                
  |     +--+                             
--+        +-- Bineurus                  
  |                                      
  |  +-------- Pilsbryoconcha            
  +--+                                   
     |  +----- Pseudodon_walpolei **        
     +--+                                
        |  +-- Pseudodon_mekongi **         
        +--+                             
           +-- Pseudodon_vondembuschianus **

Dai et al., 2023 [Publ 4285], fig. 3-ML

iT tT bi riS oT roT roS iD
3 3 1 3 42 11 5 4.0 
              +-- Nyeinchanconcha      
  +-----------+                        
  |           +-- Bineurus             
  |                                    
--+           +-- Thaiconcha           
  |  +--------+                        
  |  |        +-- Sundadontina         
  +--+                                 
     |  +-------- Pilsbryoconcha       
     +--+                              
        |  +----- Pseudodon_lenyanensis **
        +--+                           
           |  +-- Pseudodon_mekongi **    
           +--+                        
              +-- Pseudodon_vagulus **

Huang et al., 2019 [Publ 3843], fig. 2

iT tT bi riS oT roT roS iD
3 3 1 3 195 9 5 4.0 
  +-------------- Pilsbryoconcha            
  |                                         
  |        +----- Nyeinchanconcha           
--+  +-----+                                
  |  |     |  +-- Thaiconcha                
  |  |     +--+                             
  +--+        +-- Bineurus                  
     |                                      
     |  +-------- Sundadontina              
     +--+                                   
        |  +----- Pseudodon_vagulus **         
        +--+                                
           |  +-- Pseudodon_mekongi **         
           +--+                             
              +-- Pseudodon_vondembuschianus **

Bolotov et al., 2017 [Publ 3764], fig. S1

iT tT bi riS oT roT roS iD
3 3 1 3 122 9 5 4.0 
  +----------- Bineurus                  
  |                                      
  |        +-- Nyeinchanconcha           
  +--------+                             
  |        +-- Sundadontina_cumingii     
--+                                      
  |        +-- Thaiconcha                
  +--------+                             
  |        +-- Pilsbryoconcha            
  |                                      
  |  +-------- Sundadontina_tumida       
  +--+                                   
     |  +----- Pseudodon_vagulus **         
     +--+                                
        |  +-- Pseudodon_mekongi **         
        +--+                             
           +-- Pseudodon_vondembuschianus **

Wu et al., 2024 [Publ 4350], fig. 6-7

iT tT bi riS oT roT roS iD
2 2 0 2 35 4 3 -- 
  +-------- Pilsbryoconcha            
  |                                   
--+     +-- Pseudodon_mekongi **         
  |  +--+                             
  |  |  +-- Pseudodon_vondembuschianus **
  +--+                                
     |  +-- Thaiconcha                
     +--+                             
        +-- Bineurus

Wu et al., 2024 [Publ 4350], fig. 5b

iT tT bi riS oT roT roS iD
2 2 0 2 35 4 3 -- 
  +----------- Pilsbryoconcha            
--+                                      
  |  +-------- Bineurus                  
  +--+                                   
     |  +----- Thaiconcha                
     +--+                                
        |  +-- Pseudodon_mekongi **         
        +--+                             
           +-- Pseudodon_vondembuschianus **

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 38 4 4 -- 
  +----------- Pilsbryoconcha            
  |                                      
--+        +-- Pseudodon_vondembuschianus **
  |  +-----+                             
  |  |     +-- Pseudodon_vagulus **         
  +--+                                   
     |  +----- Sundadontina              
     +--+                                
        |  +-- Nyeinchanconcha           
        +--+                             
           +-- Bineurus

Wu et al., 2024 [Publ 4388], fig. 5

iT tT bi riS oT roT roS iD
2 2 0 2 36 4 3 -- 
  +-------- Pilsbryoconcha            
  |                                   
--+     +-- Pseudodon_mekongi **         
  |  +--+                             
  |  |  +-- Pseudodon_vondembuschianus **
  +--+                                
     |  +-- Thaiconcha                
     +--+                             
        +-- Bineurus

Konopleva et al., 2017 [Publ 3807], fig. 6

iT tT bi riS oT roT roS iD
2 2 0 2 29 4 4 -- 
  +----------- Sundadontina              
  |                                      
--+        +-- Nyeinchanconcha           
  |  +-----+                             
  |  |     +-- Bineurus                  
  +--+                                   
     |  +----- Pilsbryoconcha            
     +--+                                
        |  +-- Pseudodon_vagulus **         
        +--+                             
           +-- Pseudodon_vondembuschianus **

Wu et al., 2024 [Publ 4350], fig. 5a

iT tT bi riS oT roT roS iD
2 2 0 2 35 4 3 -- 
  +-------- Pilsbryoconcha            
  |                                   
--+     +-- Pseudodon_mekongi **         
  |  +--+                             
  |  |  +-- Pseudodon_vondembuschianus **
  +--+                                
     |  +-- Thaiconcha                
     +--+                             
        +-- Bineurus

Wu et al., 2024 [Publ 4350], fig. 5c

iT tT bi riS oT roT roS iD
2 2 0 2 35 4 3 -- 
  +----------- Pilsbryoconcha            
  |                                      
--+  +-------- Bineurus_anodontinum      
  |  |                                   
  +--+-------- Bineurus_loeiensis        
     |                                   
     |  +----- Thaiconcha                
     +--+                                
        |  +-- Pseudodon_mekongi **         
        +--+                             
           +-- Pseudodon_vondembuschianus **

Zieritz et al., 2021 [Publ 4218], fig. 5

iT tT bi riS oT roT roS iD
2 2 0 2 30 5 4 -- 
  +----------- Pilsbryoconcha            
  |                                      
--+        +-- Pseudodon_mekongi **         
  |  +-----+                             
  |  |     +-- Pseudodon_vondembuschianus **
  +--+                                   
     |  +----- Sundadontina              
     +--+                                
        |  +-- Thaiconcha                
        +--+                             
           +-- Bineurus

Liu et al., 2024 [Publ 4380], fig. 5

iT tT bi riS oT roT roS iD
2 2 0 2 35 4 3 -- 
  +-------- Pilsbryoconcha            
  |                                   
--+     +-- Pseudodon_mekongi **         
  |  +--+                             
  |  |  +-- Pseudodon_vondembuschianus **
  +--+                                
     |  +-- Thaiconcha                
     +--+                             
        +-- Bineurus

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iT tT bi riS oT roT roS iD
1 1 0 1 91 1 1 -- 
  +-- Pseudodon_vondembuschianus **
--+                             
  +-- Pilsbryoconcha

Bolotov et al., 2022 [Publ 4239], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 57 2 2 -- 
-- Pseudodon_lenyanensis

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iT tT bi riS oT roT roS iD
1 1 0 1 56 3 3 -- 
  +-------- Bineurus                  
--+                                   
  |  +----- Pilsbryoconcha            
  +--+                                
     |  +-- Pseudodon_vondembuschianus **
     +--+                             
        +-- Sundadontina

Wu et al., 2021 [Publ 4366], fig. 3-BI

iT tT bi riS oT roT roS iD
1 1 0 1 73 1 1 -- 
  +-- Pilsbryoconcha            
--+                             
  +-- Pseudodon_vondembuschianus **

Dai et al., 2023 [Publ 4285], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 52 1 1 -- 
  +-- Pseudodon_vondembuschianus **
--+                             
  +-- Pilsbryoconcha

Walker et al., 2006 [Publ 2666], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 14 0 0 -- 
-- Pseudodon_vondembuschianus

Wu et al., 2021 [Publ 4366], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 90 2 1 -- 
  +----- Pilsbryoconcha_exilis    
--+                               
  |  +-- Pseudodon_vagulus **        
  +--+                            
     +-- Pilsbryoconcha_carinifera

Walker et al., 2006 [Publ 2666], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 14 0 0 -- 
-- Pseudodon_vondembuschianus

Chen et al., 2020 [Publ 4170], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 33 1 1 -- 
  +-- Pseudodon_vondembuschianus **
--+                             
  +-- Pilsbryoconcha

Bolotov et al., 2022 [Publ 4252], fig. S1

iT tT bi riS oT roT roS iD
1 1 0 1 60 2 2 -- 
-- Pseudodon_lenyanensis

Froufe et al., 2019 [Publ 3936], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 44 1 1 -- 
  +-- Pilsbryoconcha            
--+                             
  +-- Pseudodon_vondembuschianus **

Graf & Cummings, 2006 [Publ 1968], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 46 1 1 -- 
  +-- Pilsbryoconcha            
--+                             
  +-- Pseudodon_vondembuschianus **

Wu et al., 2021 [Publ 4215], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 23 2 2 -- 
  +-- Pseudodon_vagulus **
--+                    
  +-- Pilsbryoconcha

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

iT tT bi riS oT roT roS iD
1 1 0 1 22 1 1 -- 
  +-- Pilsbryoconcha            
--+                             
  +-- Pseudodon_vondembuschianus **

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 35 1 1 -- 
  +-- Pseudodon_vondembuschianus **
--+                             
  +-- Pilsbryoconcha

Hoeh et al., 2009 [Publ 2716], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 45 1 1 -- 
  +-- Pilsbryoconcha            
--+                             
  +-- Pseudodon_vondembuschianus **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iT tT bi riS oT roT roS iD
1 1 0 1 79 4 3 -- 
  +-------- Pilsbryoconcha   
--+                          
  |  +----- Pseudodon_vagulus **
  +--+                       
     |  +-- Bineurus         
     +--+                    
        +-- Nyeinchanconcha

Bolotov et al., 2022 [Publ 4239], fig. S2

iT tT bi riS oT roT roS iD
1 1 0 1 57 2 2 -- 
-- Pseudodon_lenyanensis

Graf, 2002 [Publ 2651], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 17 1 1 -- 
  +-- Pilsbryoconcha            
--+                             
  +-- Pseudodon_vondembuschianus **

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iT tT bi riS oT roT roS iD
1 1 0 1 75 14 6 -- 
     +----- Pilsbryoconcha            
  +--+                                
  |  |  +-- Namkongnaia               
  |  +--+                             
  |     +-- Pseudodon_vondembuschianus **
--+                                   
  |     +-- Nyeinchanconcha           
  |  +--+                             
  |  |  +-- Sundadontina              
  +--+                                
     |  +-- Thaiconcha                
     +--+                             
        +-- Bineurus

Wu et al., 2022 [Publ 4238], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 73 1 1 -- 
  +-- Pilsbryoconcha            
--+                             
  +-- Pseudodon_vondembuschianus **

Matos et al., 2024 [Publ 4383], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 19 1 1 -- 
  +-- Pseudodon_vondembuschianus **
--+                             
  +-- Pilsbryoconcha

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iT tT bi riS oT roT roS iD
1 1 0 1 95 1 1 -- 
  +-- Pseudodon_vondembuschianus **
--+                             
  +-- Pilsbryoconcha

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 56 3 3 -- 
  +-------- Pilsbryoconcha            
--+                                   
  |  +----- Bineurus                  
  +--+                                
     |  +-- Pseudodon_vondembuschianus **
     +--+                             
        +-- Sundadontina

Froufe et al., 2019 [Publ 3936], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 18 1 1 -- 
  +-- Pilsbryoconcha            
--+                             
  +-- Pseudodon_vondembuschianus **

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iT tT bi riS oT roT roS iD
1 1 0 1 48 1 1 -- 
  +-- Pilsbryoconcha            
--+                             
  +-- Pseudodon_vondembuschianus **

Jeratthitikul & Sutcharit, 2023 [Publ 4289], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 12 10 6 -- 
  +----------- Bineurus                  
  |                                      
  |     +----- Namkongnaia               
--+  +--+                                
  |  |  |  +-- Sundadontina              
  |  |  +--+                             
  +--+     +-- Nyeinchanconcha           
     |                                   
     |  +----- Pseudodon_vondembuschianus **
     +--+                                
        |  +-- Pilsbryoconcha            
        +--+                             
           +-- Thaiconcha

Bolotov et al., 2022 [Publ 4239], fig. S1

iT tT bi riS oT roT roS iD
1 1 0 1 57 2 2 -- 
-- Pseudodon_lenyanensis

Wu et al., 2024 [Publ 4388], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 53 1 1 -- 
  +-- Pseudodon_vondembuschianus **
--+                             
  +-- Pilsbryoconcha

Hoeh et al., 2009 [Publ 2716], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 46 1 1 -- 
  +-- Pilsbryoconcha            
--+                             
  +-- Pseudodon_vondembuschianus **

Froufe et al., 2019 [Publ 3936], fig. 3-M

iT tT bi riS oT roT roS iD
1 1 0 1 18 1 1 -- 
  +-- Pilsbryoconcha            
--+                             
  +-- Pseudodon_vondembuschianus **

Wu et al., 2021 [Publ 4366], fig. 2_I

iT tT bi riS oT roT roS iD
1 1 0 1 90 2 1 -- 
  +----- Pilsbryoconcha_exilis    
--+                               
  |  +-- Pseudodon_vagulus **        
  +--+                            
     +-- Pilsbryoconcha_carinifera

Wu et al., 2021 [Publ 4366], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 68 1 1 -- 
  +-- Pilsbryoconcha            
--+                             
  +-- Pseudodon_vondembuschianus **

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iT tT bi riS oT roT roS iD
1 1 0 1 35 0 0 -- 
-- Pseudodon_vondembuschianus

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iT tT bi riS oT roT roS iD
1 1 0 1 95 1 1 -- 
  +-- Pseudodon_vondembuschianus **
--+                             
  +-- Pilsbryoconcha

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iT tT bi riS oT roT roS iD
1 1 0 1 38 1 1 -- 
  +-- Pseudodon_vondembuschianus **
--+                             
  +-- Pilsbryoconcha

Wu et al., 2021 [Publ 4366], fig. 3-ML

iT tT bi riS oT roT roS iD
1 1 0 1 73 1 1 -- 
  +-- Pilsbryoconcha            
--+                             
  +-- Pseudodon_vondembuschianus **

Dai et al., 2024 [Publ 4358], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 51 1 1 -- 
  +-- Pseudodon_vondembuschianus **
--+                             
  +-- Pilsbryoconcha

Bolotov et al., 2022 [Publ 4250], fig. 8

iT tT bi riS oT roT roS iD
1 1 0 1 48 1 1 -- 
  +-- Pseudodon_lenyanensis **
--+                        
  +-- Sundadontina

Bolotov et al., 2022 [Publ 4252], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 60 2 2 -- 
-- Pseudodon_lenyanensis

Hoeh et al., 2009 [Publ 2716], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 45 1 1 -- 
  +-- Pilsbryoconcha            
--+                             
  +-- Pseudodon_vondembuschianus **

Froufe et al., 2019 [Publ 3936], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 44 1 1 -- 
  +-- Pilsbryoconcha            
--+                             
  +-- Pseudodon_vondembuschianus **

Matos et al., 2024 [Publ 4384], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 19 1 1 -- 
  +-- Pseudodon_vondembuschianus **
--+                             
  +-- Pilsbryoconcha

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iT tT bi riS oT roT roS iD
1 1 0 1 91 1 1 -- 
  +-- Pseudodon_vondembuschianus **
--+                             
  +-- Pilsbryoconcha

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iT tT bi riS oT roT roS iD
1 1 0 1 119 5 4 -- 
  +-------- Pilsbryoconcha   
  |                          
--+  +----- Pseudodon_mekongi **
  |  |                       
  +--+  +-- Nyeinchanconcha  
     |  |                    
     +--+-- Bineurus         
        |                    
        +-- Sundadontina

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iT tT bi riS oT roT roS iD
1 1 0 1 75 14 6 -- 
     +----- Pilsbryoconcha            
  +--+                                
  |  |  +-- Namkongnaia               
  |  +--+                             
  |     +-- Pseudodon_vondembuschianus **
--+                                   
  |     +-- Nyeinchanconcha           
  |  +--+                             
  |  |  +-- Sundadontina              
  +--+                                
     |  +-- Thaiconcha                
     +--+                             
        +-- Bineurus

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iT tT bi riS oT roT roS iD
1 1 0 1 35 0 0 -- 
-- Pseudodon_vondembuschianus

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 33 1 1 -- 
  +-- Pseudodon_vagulus **
--+                    
  +-- Sundadontina

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 53 1 1 -- 
  +-- Pilsbryoconcha            
--+                             
  +-- Pseudodon_vondembuschianus **

Walker et al., 2006 [Publ 2666], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 14 0 0 -- 
-- Pseudodon_vondembuschianus

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