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Page last updated
Wed 06 Nov 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Ambleminae | Pleurobemini

genus Elliptio Rafinesque, 1819: cladomics
     45 valid species

Cladomics Summary

number of species = 45

analyzed in 73 cladograms in 44 publications
maximum number of terminals in an analysis = 19, mean = 1.88

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 20
maximum number of included terminals = 19, mean = 4.2
proportion of cladograms that support basic monophyly = 0.85


Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ tree


Inoue et al., 2018 [Publ 3858], fig. 1_3a-c

iT tT bi riS oT roT roS iD
19 57 49 20 85 39 8 0.0

*** Top Monophyly Cladogram *** *** Top Sister-Group Cladogram ***

  +----------------------------------------- Elliptoideus             
  |                                                                   
  |                                      +-- Elliptio_lanceolata **      
  |  +-----------------------------------+                            
  |  |                                   +-- Parvaspina               
  |  |                                                                
  |  |  +----------------------------------- Plethobasus              
  |  |  |                                                             
  |  |  +----------------------------------- Eurynia                  
--+  |  |                                                             
  |  |  +----------------------------------- Hemistena                
  |  |  |                                                             
  |  |  |                                +-- Elliptio_complanata **      
  |  |  |                                |                            
  |  |  |                                +-- Elliptio_hopetonensis **    
  |  |  |                                |                            
  |  |  |                                +-- Elliptio_producta__1 **     
  |  |  |                                |                            
  |  |  |                                +-- Elliptio_shepardiana **     
  +--+  |                                |                            
     |  |                                +-- Elliptio_angustata **       
     |  |                                |                            
     |  |                                +-- Elliptio_arca **            
     |  |                                |                            
     |  |                                +-- Elliptio_arctata **         
     |  |                                |                            
     |  |                                +-- Elliptio_jayensis **        
     |  |                                |                            
     |  |                                +-- Elliptio_crassidens **      
     |  |                                |                            
     |  +--------------------------------+-- Elliptio_dariensis **       
     |  |                                |                            
     |  |                                +-- Elliptio_fisheriana **      
     |  |                                |                            
     |  |                                +-- Elliptio_folliculata **     
     |  |                                |                            
     +--+                                +-- Elliptio_fumata **          
        |                                |                            
        |                                +-- Elliptio_icterina **        
        |                                |                            
        |                                +-- Elliptio_mcmichaeli **      
        |                                |                            
        |                                +-- Elliptio_producta__2 **     
        |                                |                            
        |                                +-- Elliptio_pullata **         
        |                                |                            
        |                                +-- Elliptio_purpurella **      
        |                                |                            
        |                                +-- Elliptio_congaraea **       
        |                                                             
        |  +-------------------------------- Pleuronaia               
        |  |                                                          
        |  |                             +-- Pleurobema_cordatum      
        |  |  +--------------------------+                            
        |  |  |                          +-- Fusconaia                
        |  |  |                                                       
        |  |  |  +-------------------------- Pleurobema_plenum        
        |  |  |  |                                                    
        |  |  |  |  +----------------------- Pleurobema_stabile       
        |  |  |  |  |                                                 
        |  |  |  |  |                 +----- Pleurobema_pyriforme     
        |  |  +--+  |              +--+                               
        |  |  |  |  |              |  |  +-- Pleurobema_georgianum__2 
        |  |  |  |  |              |  +--+                            
        |  |  |  |  |  +-----------+     +-- Pleurobema_athearni      
        |  |  |  |  |  |           |                                  
        +--+  |  +--+  |           |  +----- Pleurobema_decisum       
           |  |     |  |           |  |                               
           |  |     |  |           +--+  +-- Pleurobema_georgianum__1 
           |  |     |  |              |  |                            
           |  |     |  |              +--+-- Pleurobema_hanleyianum__1
           |  |     |  |                 |                            
           |  |     |  |                 +-- Pleurobema_hanleyianum__2
           |  |     |  |                                              
           |  |     +--+     +-------------- Pleurobema_beadleianum__2
           |  |        |     |                                        
           |  |        |  +--+  +----------- Pleurobema_strodeanum__2 
           |  |        |  |  |  |                                     
           |  |        |  |  +--+----------- Pleurobema_rubellum      
           |  |        |  |     |                                     
           |  |        |  |     |  +-------- Pleurobema_beadleianum__1
           |  |        |  |     +--+                                  
           +--+        |  |        |  +----- Pleurobema_taitianum     
              |        |  |        +--+                               
              |        |  |           |  +-- Pleurobema_perovatum     
              |        +--+           +--+                            
              |           |              +-- Pleurobema_strodeanum__1 
              |           |                                           
              |           |           +----- Pleurobema_oviforme__6   
              |           |           |                               
              |           |        +--+----- Pleurobema_clava__3      
              |           |        |  |                               
              |           |        |  |  +-- Pleurobema_oviforme__5   
              |           |        |  +--+                            
              |           +--------+     +-- Pleurobema_clava__2      
              |                    |                                  
              |                    |  +----- Pleurobema_oviforme__2   
              |                    |  |                               
              |                    |  +----- Pleurobema_oviforme__3   
              |                    +--+                               
              |                       +----- Pleurobema_oviforme__4   
              |                       |                               
              |                       |  +-- Pleurobema_clava__1      
              |                       +--+                            
              |                          +-- Pleurobema_oviforme__1   
              |                                                       
              |           +----------------- Pleurobema_riddellii     
              |           |                                           
              |           |  +-------------- Pleurobema_sintoxia__34  
              |           |  |                                        
              |           |  |  +----------- Pleurobema_sintoxia__31  
              |           |  |  |                                     
              +-----------+  |  +----------- Pleurobema_sintoxia__32  
                          |  |  |                                     
                          |  |  +----------- Pleurobema_sintoxia__33  
                          |  |  |                                     
                          |  |  |  +-------- Pleurobema__4            
                          |  |  |  |                                  
                          |  |  |  +-------- Pleurobema__5            
                          +--+  |  |                                  
                             |  |  +-------- Pleurobema_sintoxia__2   
                             |  |  |                                  
                             |  |  +-------- Pleurobema_sintoxia__3   
                             |  |  |                                  
                             |  |  +-------- Pleurobema__8            
                             |  |  |                                  
                             |  |  +-------- Pleurobema__9            
                             |  |  |                                  
                             |  |  +-------- Pleurobema__10           
                             |  |  |                                  
                             |  |  +-------- Pleurobema_sintoxia__5   
                             +--+  |                                  
                                |  +-------- Pleurobema_sintoxia__6   
                                |  |                                  
                                |  +-------- Pleurobema_sintoxia__7   
                                |  |                                  
                                |  +-------- Pleurobema_sintoxia__8   
                                |  |                                  
                                |  +-------- Pleurobema_sintoxia__9   
                                |  |                                  
                                |  +-------- Pleurobema_sintoxia__10  
                                |  |                                  
                                |  +-------- Pleurobema_sintoxia__11  
                                |  |                                  
                                |  +-------- Pleurobema_sintoxia__12  
                                |  |                                  
                                |  +-------- Pleurobema_sintoxia__13  
                                |  |                                  
                                |  +-------- Pleurobema_sintoxia__14  
                                |  |                                  
                                |  +-------- Pleurobema_sintoxia__15  
                                |  |                                  
                                |  +-------- Pleurobema_sintoxia__16  
                                +--+                                  
                                   +-------- Pleurobema_sintoxia__17  
                                   |                                  
                                   +-------- Pleurobema_sintoxia__18  
                                   |                                  
                                   +-------- Pleurobema_sintoxia__19  
                                   |                                  
                                   +-------- Pleurobema_sintoxia__20  
                                   |                                  
                                   +-------- Pleurobema_sintoxia__21  
                                   |                                  
                                   +-------- Pleurobema_sintoxia__22  
                                   |                                  
                                   +-------- Pleurobema_sintoxia__23  
                                   |                                  
                                   +-------- Pleurobema_sintoxia__24  
                                   |                                  
                                   +-------- Pleurobema_sintoxia__25  
                                   |                                  
                                   +-------- Pleurobema_sintoxia__26  
                                   |                                  
                                   +-------- Pleurobema_sintoxia__27  
                                   |                                  
                                   +-------- Pleurobema_sintoxia__28  
                                   |                                  
                                   +-------- Pleurobema_sintoxia__29  
                                   |                                  
                                   +-------- Pleurobema_sintoxia__30  
                                   |                                  
                                   |     +-- Pleurobema__6            
                                   +-----+                            
                                   |     +-- Pleurobema_sintoxia__1   
                                   |                                  
                                   |     +-- Pleurobema__7            
                                   +-----+                            
                                   |     +-- Pleurobema_sintoxia__4   
                                   |                                  
                                   |  +----- Pleurobema__3            
                                   +--+                               
                                      |  +-- Pleurobema__1            
                                      +--+                            
                                         +-- Pleurobema__2            

Perkins et al., 2017 [Publ 3777], fig. 2

iT tT bi riS oT roT roS iD
13 13 12 14 23 21 6 148.0
  +-------------------------------------- Pleurobema           
  |                                                            
--+  +----------------------------------- Pleuronaia           
  |  |                                                         
  +--+  +-------------------------------- Fusconaia            
     |  |                                                      
     +--+  +----------------------------- Parvaspina           
        |  |                                                   
        +--+                          +-- Eurynia              
           |  +-----------------------+                        
           |  |                       +-- Hemistena            
           +--+                                                
              |  +----------------------- Elliptio_producta__2 ** 
              |  |                                             
              +--+  +-------------------- Elliptio_producta__1 ** 
                 |  |                                          
                 +--+  +----------------- Elliptio_spinosa **     
                    |  |                                       
                    |  |  +-------------- Elliptio_icterina **    
                    +--+  |                                    
                       |  |     +-------- Elliptio_roanokensis ** 
                       |  |  +--+                              
                       +--+  |  |  +----- Elliptio_complanata **  
                          |  |  +--+                           
                          |  |     |  +-- Elliptio_crassidens **  
                          |  |     +--+                        
                          +--+        +-- Elliptio_pullata **     
                             |                                 
                             |        +-- Elliptio_congaraea **   
                             |  +-----+                        
                             |  |     +-- Elliptio_hopetonensis **
                             |  |                              
                             +--+     +-- Elliptio_folliculata ** 
                                |  +--+                        
                                |  |  +-- Elliptio_jayensis **    
                                +--+                           
                                   |  +-- Elliptio_arca **        
                                   +--+                        
                                      +-- Elliptio_arctata **     

Campbell & Lydeard, 2012 [Publ 3309], fig. 3

iT tT bi riS oT roT roS iD
8 8 5 8 65 33 8 145.0
  +----------------- Elliptoideus        
  |                                      
  |              +-- Eurynia             
  |  +-----------+                       
  |  |           +-- Parvaspina          
--+  |                                   
  |  |           +-- Hemistena           
  |  +-----------+                       
  |  |           +-- Plethobasus         
  |  |                                   
  |  |        +----- Pleuronaia          
  +--+--------+                          
     |        |  +-- Fusconaia           
     |        +--+                       
     |           +-- Pleurobema          
     |                                   
     |  +----------- Elliptio_shepardiana **
     |  |                                
     |  |     +----- Elliptio_crassidens ** 
     +--+  +--+                          
        |  |  |  +-- Elliptio_mcmichaeli ** 
        |  |  +--+                       
        +--+     +-- Elliptio_pullata **    
           |                             
           |  +----- Elliptio_complanata ** 
           |  |                          
           +--+  +-- Elliptio_purpurella ** 
              |  |                       
              +--+-- Elliptio_arca **       
                 |                       
                 +-- Elliptio_arctata **    

Campbell & Lydeard, 2012 [Publ 3309], fig. 1-2

iT tT bi riS oT roT roS iD
8 8 4 8 65 33 8 144.0
  +----------------------------------- Elliptoideus          
  |                                                          
  |  +-------------------------------- Pleurobema_stabile    
  |  |                                                       
  |  |                             +-- Pleurobema_clava      
--+  |                 +-----------+                         
  |  |                 |           +-- Pleurobema_oviforme   
  |  |                 |                                     
  |  |  +--------------+  +----------- Pleurobema_pyriforme  
  |  |  |              |  |                                  
  |  |  |              |  |        +-- Pleurobema_athearni   
  +--+  |              |  |     +--+                         
     |  |              +--+     |  +-- Pleurobema_georgianum 
     |  |                 |  +--+                            
     |  |                 |  |  |  +-- Pleurobema_decisum    
     |  |                 |  |  +--+                         
     |  |                 |  |     +-- Pleurobema_hanleyianum
     |  |                 +--+                               
     |  |                    |  +----- Pleurobema_beadleianum
     |  |                    |  |                            
     +--+                    |  +----- Pleurobema_strodeanum 
        |                    +--+                            
        |                       +----- Pleurobema_rubellum   
        |                       |                            
        |                       |  +-- Pleurobema_perovatum  
        |                       +--+                         
        |                          +-- Pleurobema_taitianum  
        |                                                    
        |                       +----- Pleurobema_riddellii  
        |  +--------------------+                            
        |  |                    |  +-- Pleurobema_sintoxia   
        |  |                    +--+                         
        +--+                       +-- Pleurobema_cordatum   
           |                                                 
           |  +----------------------- Fusconaia             
           |  |                                              
           +--+  +-------------------- Pleuronaia            
              |  |                                           
              |  |                 +-- Eurynia               
              +--+  +--------------+                         
                 |  |              +-- Parvaspina            
                 |  |                                        
                 +--+              +-- Hemistena             
                    |  +-----------+                         
                    |  |           +-- Plethobasus           
                    |  |                                     
                    +--+  +----------- Elliptio_shepardiana **  
                       |  |                                  
                       |  |  +-------- Elliptio_complanata **   
                       +--+  |                               
                          |  |  +----- Elliptio_purpurella **   
                          |  +--+                            
                          |  |  |  +-- Elliptio_arca **         
                          +--+  +--+                         
                             |     +-- Elliptio_arctata **      
                             |                               
                             |     +-- Elliptio_mcmichaeli **   
                             |     |                         
                             +-----+-- Elliptio_pullata **      
                                   |                         
                                   +-- Elliptio_crassidens **   

Perkins et al., 2017 [Publ 3777], fig. 3

iT tT bi riS oT roT roS iD
4 20 7 4 20 18 6 145.0
  +----------------- Parvaspina            
  |                                        
  |  +-------------- Elliptio_arca **         
  |  |                                     
--+  +-------------- Hemistena             
  |  |                                     
  |  |           +-- Elliptio_arctata **      
  |  +-----------+                         
  |  |           +-- Elliptio_crassidens **   
  +--+                                     
     |  +----------- Eurynia               
     |  |                                  
     |  |  +-------- Pleurobema_oviforme   
     |  |  |                               
     |  |  +-------- Pleurobema_clava      
     |  |  |                               
     +--+  +-------- Pleurobema_pyriforme  
        |  |                               
        |  +-------- Pleurobema_georgianum 
        |  |                               
        |  +-------- Pleurobema_beadleianum
        |  |                               
        +--+-------- Pleurobema_hanleyianum
           |                               
           +-------- Pleurobema_decisum    
           |                               
           |     +-- Pleurobema_sintoxia   
           +-----+                         
           |     +-- Pleurobema_rubellum   
           |                               
           |  +----- Elliptio_spinosa **      
           |  |                            
           +--+  +-- Fusconaia_escambia    
              |  |                         
              |  +-- Fusconaia_flava       
              +--+                         
                 +-- Fusconaia_cerina      
                 |                         
                 +-- Pleuronaia            

Campbell et al., 2008 [Publ 2675], fig. 5

iT tT bi riS oT roT roS iD
3 23 5 3 22 19 7 147.0
        +-- Elliptio_arctata **         
  +-----+                            
  |     +-- Elliptio_crassidens **      
  |                                  
  |  +----- Pleurobema_decisum       
  |  |                               
  |  +----- Pleurobema_rubellum      
  |  |                               
  |  +----- Pleurobema_georgianum__2 
  |  |                               
  |  +----- Pleurobema_stabile       
  |  |                               
  |  +----- Pleurobema_pyriforme     
  |  |                               
--+  +----- Pleurobema_oviforme__2   
  |  |                               
  |  +----- Pleurobema_hanleyianum__2
  |  |                               
  |  +----- Pleuronaia               
  |  |                               
  |  +----- Pleurobema_georgianum__3 
  |  |                               
  |  +----- Pleurobema_cordatum      
  |  |                               
  |  +----- Pleurobema_beadleianum   
  |  |                               
  |  +----- Hemistena                
  +--+                               
     +----- Fusconaia_flava          
     |                               
     +----- Fusconaia_cerina         
     |                               
     +----- Eurynia                  
     |                               
     +----- Elliptio_arca **            
     |                               
     +----- Cyrtonaias               
     |                               
     |  +-- Plethobasus              
     +--+                            
     |  +-- Parvaspina               
     |                               
     |  +-- Pleurobema_sintoxia      
     |  |                            
     +--+-- Pleurobema_oviforme__1   
     |  |                            
     |  +-- Pleurobema_clava         
     |                               
     |  +-- Pleurobema_athearni      
     |  |                            
     +--+-- Pleurobema_georgianum__1 
        |                            
        +-- Pleurobema_hanleyianum__1

Campbell et al., 2008 [Publ 2675], fig. 4

iT tT bi riS oT roT roS iD
3 3 1 3 22 19 7 146.0
  +----- Elliptio_arca **      
--+                         
  |  +-- Elliptio_crassidens **
  +--+                      
     +-- Elliptio_arctata **   

Campbell et al., 2005 [Publ 2660], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 100 28 8 --
  +-------------------------------- Elliptoideus             
  |                                                          
  |  +----------------------------- Plethobasus              
  |  |                                                       
--+  +----------------------------- Parvaspina               
  |  |                                                       
  |  |                          +-- Elliptio_arca **            
  |  +--------------------------+                            
  +--+                          +-- Elliptio_crassidens **      
     |                                                       
     |  +-------------------------- Eurynia                  
     |  |                                                    
     |  +-------------------------- Hemistena                
     |  |                                                    
     |  |  +----------------------- Fusconaia                
     +--+  |                                                 
        |  |                 +----- Pleurobema_sintoxia__2   
        |  +-----------------+                               
        |  |                 |  +-- Pleurobema_sintoxia__1   
        |  |                 +--+                            
        +--+                    +-- Pleurobema_cordatum      
           |                                                 
           |  +-------------------- Pleuronaia               
           |  |                                              
           |  |           +-------- Pleurobema_rubellum      
           |  |  +--------+                                  
           +--+  |        |  +----- Pleurobema_strodeanum    
              |  |        +--+                               
              |  |           |  +-- Pleurobema_perovatum     
              |  |           +--+                            
              |  |              +-- Pleurobema_taitianum     
              +--+                                           
                 |              +-- Pleurobema_oviforme__1   
                 |              |                            
                 |  +-----------+-- Pleurobema_oviforme__2   
                 |  |           |                            
                 |  |           +-- Pleurobema_clava         
                 +--+                                        
                    |  +----------- Pleurobema_pyriforme     
                    |  |                                     
                    +--+        +-- Pleurobema_athearni      
                       |  +-----+                            
                       |  |     +-- Pleurobema_georgianum__2 
                       +--+                                  
                          |  +----- Pleurobema_decisum       
                          |  |                               
                          +--+  +-- Pleurobema_hanleyianum__1
                             |  |                            
                             +--+-- Pleurobema_hanleyianum__2
                                |                            
                                +-- Pleurobema_georgianum__1 

Campbell & Lydeard, 2012 [Publ 3308], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 2 40 23 8 --
  +----------------------------- Elliptoideus        
  |                                                  
--+  +-------------------------- Pleurobema_stabile  
  |  |                                               
  +--+  +----------------------- Hemistena           
     |  |                                            
     +--+  +-------------------- Eurynia             
        |  |                                         
        |  |                 +-- Pleurobema_cordatum 
        +--+  +--------------+                       
           |  |              +-- Pleurobema_sintoxia 
           |  |                                      
           |  |           +----- Pleurobema_clava    
           +--+  +--------+                          
              |  |        |  +-- Pleurobema_decisum  
              |  |        +--+                       
              |  |           +-- Pleurobema_pyriforme
              +--+                                   
                 |           +-- Parvaspina          
                 |  +--------+                       
                 |  |        +-- Pleuronaia          
                 +--+                                
                    |  +-------- Fusconaia           
                    +--+                             
                       |  +----- Plethobasus         
                       +--+                          
                          |  +-- Elliptio_arca **       
                          +--+                       
                             +-- Elliptio_crassidens ** 

Campbell et al., 2008 [Publ 2675], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 24 22 7 --
                    +-- Pleurobema_cordatum     
                    |                           
        +-----------+-- Pleurobema_sintoxia__1  
        |           |                           
        |           +-- Pleurobema_sintoxia__2  
  +-----+                                       
  |     |  +----------- Fusconaia               
  |     |  |                                    
  |     +--+  +-------- Pleuronaia              
  |        |  |                                 
  |        |  |  +----- Eurynia                 
  |        +--+  |                              
  |           |  +----- Hemistena               
  |           |  |                              
  |           +--+----- Plethobasus             
--+              |                              
  |              +----- Parvaspina              
  |              |                              
  |              |  +-- Elliptio_arca **           
  |              +--+                           
  |                 +-- Elliptio_crassidens **     
  |                                             
  |  +----------------- Pleurobema_stabile      
  |  |                                          
  |  |  +-------------- Pleurobema_pyriforme    
  |  |  |                                       
  +--+  |           +-- Pleurobema_georgianum__2
     |  +-----------+                           
     |  |           +-- Pleurobema_athearni     
     |  |                                       
     |  |        +----- Pleurobema_decisum      
     +--+--------+                              
        |        |  +-- Pleurobema_hanleyianum  
        |        +--+                           
        |           +-- Pleurobema_georgianum__1
        |                                       
        |           +-- Pleurobema_clava        
        |  +--------+                           
        |  |        +-- Pleurobema_oviforme     
        +--+                                    
           |  +-------- Pleurobema_rubellum     
           |  |                                 
           +--+-------- Pleurobema_strodeanum   
              |                                 
              |  +----- Pleurobema_beadleianum  
              +--+                              
                 |  +-- Pleurobema_taitianum    
                 +--+                           
                    +-- Pleurobema_perovatum    

Campbell et al., 2008 [Publ 2675], fig. 6

iT tT bi riS oT roT roS iD
2 2 0 2 24 22 7 --
  +----------------- Parvaspina         
  |                                     
--+  +-------------- Hemistena          
  |  |                                  
  |  |        +----- Plethobasus        
  +--+  +-----+                         
     |  |     |  +-- Elliptio_arca **      
     |  |     +--+                      
     +--+        +-- Elliptio_crassidens **
        |                               
        |  +-------- Fusconaia          
        +--+                            
           |  +----- Pleurobema         
           +--+                         
              |  +-- Eurynia            
              +--+                      
                 +-- Pleuronaia         

Pfeiffer et al., 2016 [Publ 3660], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 36 19 8 --
                 +-- Elliptio_fumata **      
  +--------------+                        
  |              +-- Elliptio_crassidens **  
  |                                       
  |  +-------------- Parvaspina           
--+  |                                    
  |  +-------------- Plethobasus          
  |  |                                    
  |  |           +-- Hemistena            
  +--+-----------+                        
     |           +-- Eurynia              
     |                                    
     |  +----------- Fusconaia            
     |  |                                 
     |  |     +----- Pleurobema_riddellii 
     +--+  +--+                           
        |  |  |  +-- Pleurobema_sintoxia  
        |  |  +--+                        
        +--+     +-- Pleurobema_cordatum  
           |                              
           |  +----- Pleuronaia           
           |  |                           
           +--+  +-- Pleurobema_pyriforme 
              |  |                        
              +--+-- Pleurobema_strodeanum
                 |                        
                 +-- Pleurobema_clava     

Campbell & Lydeard, 2012 [Publ 3308], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 40 23 8 --
  +----------------------- Elliptoideus        
  |                                            
  |  +-------------------- Pleurobema_stabile  
--+  |                                         
  |  |              +----- Pleurobema_pyriforme
  |  |  +-----------+                          
  +--+  |           |  +-- Pleurobema_clava    
     |  |           +--+                       
     |  |              +-- Pleurobema_decisum  
     |  |                                      
     +--+              +-- Pleurobema_cordatum 
        |  +-----------+                       
        |  |           +-- Pleurobema_sintoxia 
        |  |                                   
        |  |        +----- Pleuronaia          
        +--+  +-----+                          
           |  |     |  +-- Parvaspina          
           |  |     +--+                       
           |  |        +-- Fusconaia           
           +--+                                
              |        +-- Eurynia             
              |  +-----+                       
              |  |     +-- Hemistena           
              +--+                             
                 |  +----- Plethobasus         
                 +--+                          
                    |  +-- Elliptio_arca **       
                    +--+                       
                       +-- Elliptio_crassidens ** 

Burlakova et al., 2012 [Publ 3486], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 39 33 7 --
  +----------------------------------- Parvaspina            
  |                                                          
  |                             +----- Plethobasus           
--+  +--------------------------+                            
  |  |                          |  +-- Elliptio_arca **         
  |  |                          +--+                         
  |  |                             +-- Elliptio_crassidens **   
  +--+                                                       
     |  +----------------------------- Fusconaia             
     |  |                                                    
     |  |                          +-- Eurynia               
     +--+  +-----------------------+                         
        |  |                       +-- Hemistena             
        |  |                                                 
        |  |              +----------- Pleuronaia            
        +--+  +-----------+                                  
           |  |           |  +-------- Pleurobema_riddellii  
           |  |           +--+                               
           |  |              |  +----- Pleurobema_sintoxia__2
           |  |              +--+                            
           +--+                 |  +-- Pleurobema_sintoxia__1
              |                 +--+                         
              |                    +-- Pleurobema_cordatum   
              |                                              
              |  +-------------------- Pleurobema_stabile    
              |  |                                           
              +--+  +----------------- Pleurobema_pyriforme  
                 |  |                                        
                 |  |           +----- Pleurobema_georgianum 
                 +--+  +--------+                            
                    |  |        |  +-- Pleurobema_hanleyianum
                    |  |        +--+                         
                    |  |           +-- Pleurobema_decisum    
                    +--+                                     
                       |           +-- Pleurobema_clava      
                       |  +--------+                         
                       |  |        +-- Pleurobema_oviforme   
                       +--+                                  
                          |  +-------- Pleurobema_beadleianum
                          |  |                               
                          +--+     +-- Pleurobema_rubellum   
                             |  +--+                         
                             |  |  +-- Pleurobema_strodeanum 
                             +--+                            
                                |  +-- Pleurobema_perovatum  
                                +--+                         
                                   +-- Pleurobema_taitianum  

Campbell & Lydeard, 2012 [Publ 3308], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 40 23 8 --
  +-------- Elliptoideus       
  |                            
  |  +----- Hemistena          
  |  |                         
--+  +----- Plethobasus        
  |  |                         
  |  |  +-- Elliptio_arca **      
  |  +--+                      
  |  |  +-- Elliptio_crassidens **
  +--+                         
     |  +-- Eurynia            
     +--+                      
     |  +-- Parvaspina         
     |                         
     |  +-- Pleuronaia         
     |  |                      
     +--+-- Pleurobema         
        |                      
        +-- Fusconaia          

Burlakova et al., 2012 [Publ 3486], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 33 33 7 --
  +----------------------------------- Parvaspina            
  |                                                          
  |  +-------------------------------- Pleurobema_plenum     
--+  |                                                       
  |  |                             +-- Plethobasus           
  |  |                       +-----+                         
  |  |                       |     +-- Hemistena             
  +--+  +--------------------+                               
     |  |                    |  +----- Eurynia               
     |  |                    +--+                            
     |  |                       |  +-- Elliptio_arca **         
     |  |                       +--+                         
     +--+                          +-- Elliptio_crassidens **   
        |                                                    
        |  +-------------------------- Pleurobema_cordatum__2
        |  |                                                 
        |  |  +----------------------- Fusconaia             
        +--+  |                                              
           |  |              +-------- Pleurobema_stabile    
           |  |              |                               
           |  |  +-----------+  +----- Pleurobema_riddellii  
           +--+  |           |  |                            
              |  |           +--+  +-- Pleurobema_cordatum__1
              |  |              |  |                         
              |  |              |  +-- Pleurobema_sintoxia__1
              |  |              +--+                         
              +--+                 +-- Pleurobema_sintoxia__2
                 |                 |                         
                 |                 +-- Pleurobema_sintoxia__3
                 |                                           
                 |  +----------------- Pleuronaia            
                 |  |                                        
                 |  |              +-- Pleurobema_georgianum 
                 +--+  +-----------+                         
                    |  |           +-- Pleurobema_pyriforme  
                    |  |                                     
                    +--+           +-- Pleurobema_decisum    
                       |  +--------+                         
                       |  |        +-- Pleurobema_hanleyianum
                       |  |                                  
                       +--+        +-- Pleurobema_clava      
                          |  +-----+                         
                          |  |     +-- Pleurobema_oviforme   
                          |  |                               
                          +--+  +----- Pleurobema_rubellum   
                             |  |                            
                             |  +----- Pleurobema_beadleianum
                             +--+                            
                                +----- Pleurobema_strodeanum 
                                |                            
                                |  +-- Pleurobema_perovatum  
                                +--+                         
                                   +-- Pleurobema_taitianum  

Campbell & Lydeard, 2012 [Publ 3308], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 41 23 8 --
                       +-- Elliptio_arca **       
  +--------------------+                       
  |                    +-- Elliptio_crassidens ** 
  |                                            
--+  +-------------------- Eurynia             
  |  |                                         
  |  |  +----------------- Parvaspina          
  +--+  |                                      
     |  |              +-- Hemistena           
     |  +--------------+                       
     +--+              +-- Plethobasus         
        |                                      
        |  +-------------- Pleuronaia          
        |  |                                   
        |  |           +-- Pleurobema_cordatum 
        +--+  +--------+                       
           |  |        +-- Fusconaia           
           |  |                                
           +--+        +-- Pleurobema_stabile  
              |  +-----+                       
              |  |     +-- Pleurobema_sintoxia 
              +--+                             
                 |  +----- Pleurobema_decisum  
                 +--+                          
                    |  +-- Pleurobema_clava    
                    +--+                       
                       +-- Pleurobema_pyriforme

Burlakova et al., 2012 [Publ 3486], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 34 33 7 --
  +----------------------- Hemistena             
  |                                              
  +----------------------- Pleurobema_stabile    
  |                                              
  |  +-------------------- Eurynia               
--+  |                                           
  |  |           +-------- Pleurobema_riddellii  
  |  |           |                               
  |  |  +--------+  +----- Pleurobema_sintoxia__2
  |  |  |        |  |                            
  +--+  |        +--+----- Pleurobema_sintoxia__3
     |  |           |                            
     |  |           |  +-- Pleurobema_cordatum   
     |  |           +--+                         
     |  |              +-- Pleurobema_sintoxia__1
     |  |                                        
     +--+  +-------------- Parvaspina            
        |  |                                     
        |  +-------------- Pleuronaia            
        |  |                                     
        |  +-------------- Fusconaia             
        |  |                                     
        |  |        +----- Plethobasus           
        |  +--------+                            
        +--+        |  +-- Elliptio_arca **         
           |        +--+                         
           |           +-- Elliptio_crassidens **   
           |                                     
           |           +-- Pleurobema_clava      
           |  +--------+                         
           |  |        +-- Pleurobema_oviforme   
           |  |                                  
           |  |  +-------- Pleurobema_decisum    
           +--+  |                               
              |  +-------- Pleurobema_pyriforme  
              |  |                               
              |  +-------- Pleurobema_hanleyianum
              +--+                               
                 +-------- Pleurobema_georgianum 
                 |                               
                 |  +----- Pleurobema_rubellum   
                 |  |                            
                 |  +----- Pleurobema_beadleianum
                 +--+                            
                    +----- Pleurobema_strodeanum 
                    |                            
                    |  +-- Pleurobema_perovatum  
                    +--+                         
                       +-- Pleurobema_taitianum  

Smith, 2021 [Publ 4331], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 18 0 0 --
  +-- Elliptio_crassidens **
--+                      
  +-- Elliptio_complanata **

Campbell et al., 2005 [Publ 2660], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 100 28 8 --
  +-------------- Elliptoideus       
  |                                  
--+  +----------- Hemistena          
  |  |                               
  |  |     +----- Plethobasus        
  +--+  +--+                         
     |  |  |  +-- Elliptio_arca **      
     |  |  +--+                      
     |  |     +-- Elliptio_crassidens **
     +--+                            
        |  +----- Pleuronaia         
        |  |                         
        |  +----- Fusconaia          
        +--+                         
           +----- Pleurobema         
           |                         
           |  +-- Eurynia            
           +--+                      
              +-- Parvaspina         

Wu et al., 2022 [Publ 4238], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 73 1 1 --
  +-- Pleurobema         
--+                      
  +-- Elliptio_complanata **

Johnson et al., 2018 [Publ 3844], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 15 0 0 --
-- Elliptio_crassidens

Mejia-Trujillo & Smith, 2022 [Publ 4274], fig. 2-F

iT tT bi riS oT roT roS iD
1 1 0 1 14 1 1 --
  +-- Pleurobema         
--+                      
  +-- Elliptio_complanata **

Dai et al., 2023 [Publ 4285], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 52 1 1 --
  +-- Elliptio_complanata **
--+                      
  +-- Pleurobema         

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iT tT bi riS oT roT roS iD
1 1 0 1 75 2 2 --
  +----- Elliptio_complanata **
--+                         
  |  +-- Pleurobema         
  +--+                      
     +-- Eurynia            

Campbell, 2000 [Publ 2724], fig. 1-5

iT tT bi riS oT roT roS iD
1 1 0 1 2 0 0 --
-- Elliptio_complanata

Bolotov et al., 2018 [Publ 3826], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 82 1 1 --
-- Elliptio_complanata

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 56 2 2 --
  +----- Elliptoideus       
--+                         
  |  +-- Elliptio_complanata **
  +--+                      
     +-- Pleurobema         

Wu et al., 2021 [Publ 4366], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 90 0 0 --
-- Elliptio_complanata

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iT tT bi riS oT roT roS iD
1 1 0 1 95 1 1 --
  +-- Elliptio_complanata **
--+                      
  +-- Pleurobema         

Boyer et al., 2011 [Publ 2718], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 37 3 3 --
  +-------- Pleurobema         
--+                            
  |  +----- Elliptio_crassidens **
  +--+                         
     |  +-- Fusconaia          
     +--+                      
        +-- Eurynia            

Sonowal et al., 2021 [Publ 4221], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 15 0 0 --
-- Elliptio_complanata

Lopes-Lima et al., 2019 [Publ 3901], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 27 1 1 --
  +-- Pleurobema         
--+                      
  +-- Elliptio_crassidens **

Rosenberg et al., 1997 [Publ 2720], fig. 2A

iT tT bi riS oT roT roS iD
1 1 0 1 21 2 2 --
-- Elliptio_complanata

Sonowal et al., 2021 [Publ 4221], fig. 3a

iT tT bi riS oT roT roS iD
1 1 0 1 14 0 0 --
-- Elliptio_complanata

Smith et al., 2020 [Publ 4136], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 19 18 8 --
  +-------------- Elliptoideus       
  |                                  
--+  +----------- Elliptio_crassidens **
  |  |                               
  |  +----------- Plethobasus        
  +--+                               
     |  +-------- Parvaspina         
     |  |                            
     |  |     +-- Hemistena          
     +--+-----+                      
        |     +-- Eurynia            
        |                            
        |  +----- Fusconaia          
        +--+                         
           |  +-- Pleuronaia         
           +--+                      
              +-- Pleurobema         

Huang et al., 2019 [Publ 3843], fig. S4

iT tT bi riS oT roT roS iD
1 1 0 1 197 4 4 --
  +----------- Elliptoideus       
--+                               
  |  +-------- Elliptio_complanata **
  +--+                            
     |  +----- Eurynia            
     +--+                         
        |  +-- Fusconaia          
        +--+                      
           +-- Pleurobema         

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iT tT bi riS oT roT roS iD
1 1 0 1 91 1 1 --
  +-- Elliptio_complanata **
--+                      
  +-- Pleurobema         

Rosenberg et al., 1994 [Publ 2719], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 18 2 2 --
-- Elliptio_complanata

Kandl et al., 2001 [Publ 3384], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 8 6 2 --
  +----------------- Elliptio_crassidens **    
  |                                         
--+  +-------------- Fusconaia              
  |  |                                      
  +--+  +----------- Pleurobema_sintoxia    
     |  |                                   
     |  |        +-- Pleurobema_clava       
     +--+  +-----+                          
        |  |     +-- Pleurobema_strodeanum  
        |  |                                
        +--+  +----- Pleurobema_pyriforme__6
           |  |                             
           |  |  +-- Pleurobema_pyriforme__1
           +--+  |                          
              |  +-- Pleurobema_pyriforme__2
              |  |                          
              +--+-- Pleurobema_pyriforme__3
                 |                          
                 +-- Pleurobema_pyriforme__4
                 |                          
                 +-- Pleurobema_pyriforme__5

Huang et al., 2019 [Publ 3843], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 197 4 4 --
  +-------- Elliptoideus       
  |                            
--+     +-- Elliptio_complanata **
  |  +--+                      
  |  |  +-- Eurynia            
  +--+                         
     |  +-- Fusconaia          
     +--+                      
        +-- Pleurobema         

Metzger et al., 2018 [Publ 3952], fig. 1C

iT tT bi riS oT roT roS iD
1 1 0 1 2 0 0 --
-- Elliptio_complanata

Burdick & White, 2007 [Publ 2679], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 29 27 6 --
                    +----- Eurynia               
  +-----------------+                            
  |                 |  +-- Plethobasus           
  |                 +--+                         
  |                    +-- Elliptio_crassidens **   
  |                                              
  |  +-------------------- Pleurobema_plenum     
--+  |                                           
  |  +-------------------- Pleuronaia            
  |  |                                           
  |  |                 +-- Pleurobema_cordatum__1
  |  |           +-----+                         
  |  |           |     +-- Fusconaia             
  +--+  +--------+                               
     |  |        |  +----- Pleurobema_sintoxia__1
     |  |        |  |                            
     |  |        +--+  +-- Pleurobema_cordatum__2
     |  |           |  |                         
     |  |           +--+-- Pleurobema_sintoxia__2
     |  |              |                         
     +--+              +-- Pleurobema_sintoxia__3
        |                                        
        |  +-------------- Pleurobema_stabile    
        |  |                                     
        |  |           +-- Pleurobema_rubellum   
        |  |     +-----+                         
        |  |     |     +-- Pleurobema_strodeanum 
        +--+  +--+                               
           |  |  |  +----- Pleurobema_beadleianum
           |  |  +--+                            
           |  |     |  +-- Pleurobema_perovatum  
           |  |     +--+                         
           +--+        +-- Pleurobema_taitianum  
              |                                  
              |        +-- Pleurobema_clava      
              |  +-----+                         
              |  |     +-- Pleurobema_oviforme   
              |  |                               
              +--+     +-- Pleurobema_decisum    
                 |  +--+                         
                 |  |  +-- Pleurobema_hanleyianum
                 +--+                            
                    |  +-- Pleurobema_pyriforme  
                    +--+                         
                       +-- Pleurobema_georgianum 

Huang et al., 2019 [Publ 3843], fig. S3

iT tT bi riS oT roT roS iD
1 1 0 1 195 4 4 --
  +-------- Elliptoideus       
  |                            
--+     +-- Elliptio_complanata **
  |  +--+                      
  |  |  +-- Eurynia            
  +--+                         
     |  +-- Fusconaia          
     +--+                      
        +-- Pleurobema         

Bolotov et al., 2017 [Publ 3761], fig. S2

iT tT bi riS oT roT roS iD
1 1 0 1 138 2 2 --
  +----- Pleurobema         
--+                         
  |  +-- Elliptio_complanata **
  +--+                      
     +-- Eurynia            

Giribet & Distel, 2003 [Publ 3670], fig. 3f5

iT tT bi riS oT roT roS iD
1 1 0 1 5 0 0 --
-- Elliptio_complanata

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iT tT bi riS oT roT roS iD
1 1 0 1 79 2 2 --
  +----- Elliptio_complanata **
--+                         
  |  +-- Pleurobema         
  +--+                      
     +-- Eurynia            

Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 18 1 1 --
  +-- Elliptio_complanata **
--+                      
  +-- Pleurobema         

Huang et al., 2019 [Publ 3843], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 56 0 0 --
-- Elliptio_complanata

Bolotov et al., 2017 [Publ 3764], fig. S1

iT tT bi riS oT roT roS iD
1 1 0 1 124 2 2 --
-- Elliptio_complanata

Sonowal et al., 2021 [Publ 4221], fig. 3b

iT tT bi riS oT roT roS iD
1 1 0 1 14 0 0 --
-- Elliptio_complanata

Serb et al., 2003 [Publ 2659], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 28 4 3 --
  +----- Hemistena          
--+                         
  |  +-- Elliptio_complanata **
  +--+                      
     +-- Fusconaia          

Kandl et al., 2001 [Publ 3384], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 5 4 1 --
-- Elliptio

Wu et al., 2023 [Publ 4339], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 18 0 0 --
-- Elliptio_complanata

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iT tT bi riS oT roT roS iD
1 1 0 1 119 0 0 --
-- Elliptio_complanata

Serb et al., 2003 [Publ 2659], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 28 4 3 --
  +----- Hemistena          
--+                         
  |  +-- Elliptio_complanata **
  +--+                      
     +-- Fusconaia          

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iT tT bi riS oT roT roS iD
1 1 0 1 91 1 1 --
  +-- Elliptio_complanata **
--+                      
  +-- Pleurobema         

Wu et al., 2021 [Publ 4366], fig. 2_I

iT tT bi riS oT roT roS iD
1 1 0 1 90 0 0 --
-- Elliptio_complanata

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iT tT bi riS oT roT roS iD
1 1 0 1 95 1 1 --
  +-- Elliptio_complanata **
--+                      
  +-- Pleurobema         

Wu et al., 2018 [Publ 3836], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 52 5 3 --
  +-------- Pleurobema         
--+                            
  |  +----- Fusconaia          
  +--+                         
     |  +-- Elliptio_complanata **
     +--+                      
        +-- Eurynia            

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iT tT bi riS oT roT roS iD
1 1 0 1 56 2 2 --
  +----- Elliptoideus       
--+                         
  |  +-- Elliptio_complanata **
  +--+                      
     +-- Pleurobema         

Wu et al., 2021 [Publ 4366], fig. 3-BI

iT tT bi riS oT roT roS iD
1 1 0 1 73 1 1 --
  +-- Pleurobema         
--+                      
  +-- Elliptio_complanata **

Boyer et al., 2011 [Publ 2718], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 34 3 3 --
     +-- Fusconaia          
  +--+                      
  |  +-- Pleurobema         
--+                         
  |  +-- Eurynia            
  +--+                      
     +-- Elliptio_crassidens **

Pfeiffer et al., 2019 [Publ 3866], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 34 5 4 --
  +-------------- Elliptoideus         
--+                                    
  |  +----------- Elliptio_fumata **      
  +--+                                 
     |  +-------- Pleuronaia           
     +--+                              
        |  +----- Pleurobema_riddellii 
        +--+                           
           |  +-- Fusconaia            
           +--+                        
              +-- Pleurobema_strodeanum

Bolotov et al., 2017 [Publ 3761], fig. S1

iT tT bi riS oT roT roS iD
1 1 0 1 138 2 2 --
  +----- Pleurobema         
--+                         
  |  +-- Eurynia            
  +--+                      
     +-- Elliptio_complanata **

Dai et al., 2024 [Publ 4358], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 51 1 1 --
  +-- Elliptio_complanata **
--+                      
  +-- Pleurobema         

Graf, 2000 [Publ 2647], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 16 0 0 --
-- Elliptio

Wu et al., 2021 [Publ 4366], fig. 3-ML

iT tT bi riS oT roT roS iD
1 1 0 1 73 1 1 --
  +-- Pleurobema         
--+                      
  +-- Elliptio_complanata **

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 53 0 0 --
-- Elliptio_complanata

Bolotov et al., 2020 [Publ 4084], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 35 1 1 --
-- Elliptio_complanata

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iT tT bi riS oT roT roS iD
1 1 0 1 75 2 2 --
  +----- Pleurobema         
--+                         
  |  +-- Eurynia            
  +--+                      
     +-- Elliptio_complanata **

Burlakova et al., 2019 [Publ 3899], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 32 1 1 --
  +-- Elliptio_crassidens **
--+                      
  +-- Pleurobema         

Rosenberg et al., 1997 [Publ 2720], fig. 2B

iT tT bi riS oT roT roS iD
1 1 0 1 21 1 1 --
-- Elliptio_complanata

Page updated: 08:36:45 Wed 06 Nov 2024

 
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