Bivalvia | Unionoida | Unionoidea | Unionidae | Unioninae | Unionini genus Unio Philipsson in Retzius, 1788: cladomics 41 valid species
Cladomics Summary
number of species = 41
analyzed in 196 cladograms in 117 publications
maximum number of terminals in an analysis = 23, mean = 3.1
Basic Monophyly
number of cladograms that test basic monophyly (i.e., more than 1 included species) = 133
maximum number of included terminals = 23, mean = 4.09
proportion of cladograms that support basic monophyly = 0.92
Pairwise Cladogram Clustering
The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Lyubas et al., 2022 [Publ 4255], fig. 7
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
23 |
23 |
20 |
23 |
8 |
0 |
0 |
0.0 |
*** Top Monophyly Cladogram ***
+----------------------------- Unio_tumidus **
|
| +-- Unio_durieui **
| +-----------------------+
| | +-- Unio_gibbus **
--+ |
| | +-------------- Unio_damascensis **
| | |
| | +-------------- Unio_sesirmensis **
| | +--------+
+--+ | | +----------- Unio_ionicus **
| | | |
| | +--+ +-- Unio_bruguierianus **
| | | +-----+
| | | | +-- Unio_tumidiformis **
| | +--+
| | | +----- Unio_crassus[2] **
+--+ +--+
| | +-- Unio_carneus **
| +--+
| +-- Unio_crassus[1] **
|
| +----- Unio_caffer **
| +--------------+
| | | +-- Unio_terminalis **
| | +--+
+--+ +-- Unio_delicatus **
|
| +----------------- Unio_pictorum **
| |
+--+ +-------------- Unio_eucirrus **
| |
+--+ +-- Unio_elongatulus **
| +--------+
| | +-- Unio_mancus **
+--+
| +-------- Unio_ravoisieri **
| |
+--+ +-- Unio_delphinus **
| +--+
| | +-- Unio_foucauldianus **
+--+
| +-- Unio_hueti **
+--+
+-- Unio_tigridis **
Chen et al., 2023 [Publ 4302], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
38 |
15 |
9 |
-- |
*** Top Sister-Group Cladogram ***
+-- Inversiunio_yanagawensis
+-----+
| +-- Inversiunio_reinianus
--+
| +----- Tchangsinaia_piscicula
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_crassus **
Lyubas et al., 2022 [Publ 4255], fig. S1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
23 |
23 |
14 |
23 |
8 |
0 |
0 |
10.0 |
+----------------- Unio_tumidus **
|
| +-- Unio_durieui **
+--------------+
| +-- Unio_gibbus **
|
| +----------- Unio_sesirmensis **
| |
| +--+ +-------- Unio_damascensis **
--+ | | |
| | | | +----- Unio_ionicus **
| | +--+ |
| | | | +-- Unio_bruguierianus **
| | | +--+
| | +--+ +-- Unio_tumidiformis **
| | |
| | | +-- Unio_carneus **
| | | |
| | +--+-- Unio_crassus[1] **
+--+ |
| +-- Unio_crassus[2] **
|
| +-------- Unio_elongatulus **
| |
| +-------- Unio_mancus **
| |
| | +-- Unio_pictorum **
| +-----+
| | +-- Unio_eucirrus **
| |
+-----+ +----- Unio_caffer **
+--+
| | +-- Unio_terminalis **
| +--+
| +-- Unio_delicatus **
|
| +----- Unio_ravoisieri **
| |
| | +-- Unio_delphinus **
+--+--+
| +-- Unio_foucauldianus **
|
| +-- Unio_hueti **
+--+
+-- Unio_tigridis **
Lopes-Lima et al., 2021 [Publ 4217], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
16 |
16 |
14 |
16 |
9 |
0 |
0 |
16.0 |
+----------------------- Unio_tumidus **
|
| +-------- Unio_damascensis **
| +-----------+
--+ | | +----- Unio_sesirmensis **
| | +--+
| | | +-- Unio_bruguierianus **
| | +--+
+--+ +-- Unio_crassus **
|
| +-- Unio_delicatus **
| +--------------+
| | +-- Unio_terminalis **
| |
+--+ +-- Unio_tigridis **
| +-----------+
| | +-- Unio_hueti **
| |
+--+ +----------- Unio_ravoisieri **
| |
| | +-- Unio_pictorum **
+--+ +-----+
| | +-- Unio_eucirrus **
| |
+--+ +-- Unio_elongatulus **
| +--+
| | +-- Unio_mancus **
+--+
| +-- Unio_foucauldianus **
+--+
+-- Unio_delphinus **
Araujo et al., 2018 [Publ 3779], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
14 |
14 |
9 |
14 |
5 |
1 |
1 |
20.0 |
+-------------------- Nodularia_douglasiae
|
| +-- Unio_durieui **
--+ +--------------+
| | +-- Unio_gibbus **
| |
+--+ +-------------- Unio_tumidus **
| |
| | +-------- Unio_tumidiformis **
| | +--+
+--+ | | +----- Unio_bruguierianus **
| | +--+
| | | +-- Unio_crassus **
| | +--+
+--+ +-- Unio_ionicus **
|
| +-- Unio_delphinus **
| +--+
| | +-- Unio_foucauldianus **
| |
+-----+ +-- Unio_pictorum **
| |
| +-- Unio_elongatulus **
| |
+--+-- Unio_mancus **
|
+-- Unio_ravoisieri **
|
+-- Unio_tigridis **
Froufe et al., 2016 [Publ 3738], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
12 |
12 |
10 |
12 |
2 |
2 |
1 |
23.0 |
+-- Nodularia_douglasiae
+--------------------------+
| +-- Nodularia_breviconcha
|
--+ +-- Unio_gibbus **
| +-----------------------+
| | +-- Unio_durieui **
+--+
| +----------------------- Unio_tumidus **
| |
+--+ +-- Unio_crassus **
| +-----------------+
| | +-- Unio_tumidiformis **
+--+
| +----------------- Unio_delphinus **
+--+
| +-------------- Unio_foucauldianus **
+--+
| +----------- Unio_ravoisieri **
+--+
| +-------- Unio_mancus **
+--+
| +----- Unio_elongatulus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_caffer **
Prié & Puillandre, 2014 [Publ 3529], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
11 |
11 |
9 |
11 |
6 |
0 |
0 |
22.0 |
+----- Unio_tumidus **
+-----------+
| | +-- Unio_gibbus **
| +--+
| +-- Unio_durieui **
--+
| +----- Unio_tumidiformis **
| +--------+
| | | +-- Unio_crassus **
| | +--+
+--+ +-- Unio_courtillieri **
|
| +----------- Unio_delphinus **
+--+
| +-------- Unio_elongatulus **
+--+
| +----- Unio_mancus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_ravoisieri **
Lopes-Lima et al., 2021 [Publ 4217], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
10 |
10 |
8 |
10 |
0 |
0 |
0 |
17.0 |
+----------------- Unio_tumidus **
|
--+ +-------------- Unio_bruguierianus **
| |
| | +-- Unio_delicatus **
+--+ +--------+
| | +-- Unio_terminalis **
| |
+--+ +-- Unio_tigridis **
| +-----+
| | +-- Unio_hueti **
| |
+--+ +-- Unio_pictorum **
| +--+
| | +-- Unio_eucirrus **
+--+
| +-- Unio_elongatulus **
+--+
+-- Unio_mancus **
Khalloufi et al., 2011 [Publ 2748], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
9 |
9 |
6 |
9 |
2 |
0 |
0 |
18.0 |
+----- Unio_tumidus **
+-----+
| | +-- Unio_gibbus **
| +--+
| +-- Unio_durieui **
--+
| +-- Unio_tumidiformis **
| +-----+
| | +-- Unio_crassus **
+--+
| +----- Unio **
| |
+--+----- Unio_pictorum **
|
| +-- Unio_delphinus **
+--+
+-- Unio_mancus **
Lopes-Lima et al., 2017 [Publ 3734], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
9 |
9 |
5 |
9 |
8 |
0 |
0 |
20.0 |
+-- Unio_tumidus **
+--------+
| +-- Unio_gibbus **
|
--+ +-- Unio_crassus **
| +-----+
| | +-- Unio_tumidiformis **
+--+
| +----- Unio_ravoisieri **
| |
+--+ +-- Unio_mancus **
| |
| +-- Unio_pictorum **
+--+
+-- Unio_delphinus **
|
+-- Unio_elongatulus **
Araujo et al., 2009 [Publ 2589], fig. 7B
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
8 |
8 |
6 |
8 |
4 |
0 |
0 |
19.0 |
+-------------- Unio_gibbus **
|
--+ +----------- Unio_tumidus **
| |
| | +-- Unio_tumidiformis **
+--+ +-----+
| | +-- Unio_crassus **
| |
+--+ +-- Unio_pictorum[1] **
| +--+
| | +-- Unio_mancus **
+--+
| +-- Unio_pictorum[2] **
+--+
+-- Unio_caffer **
Reis et al., 2013 [Publ 3580], fig. 5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
8 |
8 |
6 |
8 |
8 |
0 |
0 |
19.0 |
+-------------- Unio_tumidus **
|
--+ +-- Unio_tumidiformis **
| +--------+
| | +-- Unio_crassus **
+--+
| +-------- Unio **
| |
+--+ +-- Unio_delphinus **
| +--+
| | +-- Unio_mancus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_caffer **
Marrone et al., 2019 [Publ 3907], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
8 |
8 |
4 |
8 |
0 |
0 |
0 |
23.0 |
+-------------- Unio_tumidus **
|
--+ +----------- Unio_delphinus **
| |
+--+ +-------- Unio_foucauldianus **
| |
+--+ +----- Unio_mancus **
| |
| +----- Unio_elongatulus **
+--+
+----- Unio_ravoisieri **
|
| +-- Unio_pictorum **
+--+
+-- Unio_tigridis **
Araujo et al., 2009 [Publ 2589], fig. 7A
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
7 |
8 |
3 |
7 |
5 |
1 |
1 |
22.0 |
+----------- Nodularia_douglasiae
|
| +-- Unio_gibbus **
--+ |
| +-----+-- Unio_tumidus **
| | |
| | +-- Anodonta_cygnea
+--+
| +----- Unio_pictorum[1] **
| |
| +----- Unio_pictorum[2] **
+--+
+----- Unio_mancus **
|
| +-- Unio_tumidiformis **
+--+
+-- Unio_crassus **
Araujo et al., 2009 [Publ 2589], fig. 7C
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
7 |
7 |
4 |
7 |
4 |
0 |
0 |
20.0 |
+-- Unio_gibbus **
+-----+
| +-- Unio_tumidus **
|
--+ +-- Unio_tumidiformis **
| +--+
| | +-- Unio_crassus **
+--+
| +-- Unio_pictorum[1] **
| |
+--+-- Unio_pictorum[2] **
|
+-- Unio_mancus **
Lopes-Lima et al., 2021 [Publ 4217], fig. 6
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
7 |
7 |
5 |
7 |
0 |
0 |
0 |
19.0 |
+----------------- Unio_tumidus **
--+
| +-------------- Unio_pictorum **
+--+
| +----------- Unio_sesirmensis **
+--+
| +-------- Unio_damascensis **
+--+
| +----- Unio_courtillieri **
+--+
| +-- Unio_bruguierianus **
+--+
+-- Unio_crassus **
Reis et al., 2013 [Publ 3580], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
6 |
6 |
4 |
6 |
8 |
0 |
0 |
19.0 |
+----------- Unio_tumidus **
|
--+ +-- Unio_tumidiformis **
| +-----+
| | +-- Unio_crassus **
+--+
| +----- Unio_delphinus **
+--+
| +-- Unio_mancus **
+--+
+-- Unio_pictorum **
Bashê, 2022 [Publ 4265], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
6 |
6 |
4 |
6 |
1 |
0 |
0 |
25.0 |
+-------------- Unio_pictorum[2] **
--+
| +----------- Unio_foucauldianus **
+--+
| +-------- Unio_delphinus **
+--+
| +----- Unio_tigridis **
+--+
| +-- Unio_elongatulus **
+--+
+-- Unio_pictorum[1] **
Wu et al., 2023 [Publ 4339], fig. 4, S5-aa
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
6 |
6 |
4 |
6 |
90 |
16 |
6 |
21.0 |
+-------- Aculamprotula_tortuosa
|
+--------------+ +-- Aculamprotula_coreana
| | +--+
| | | +-- Aculamprotula_tientsinensis
| +--+
| | +-- Aculamprotula_polysticta
| +--+
| +-- Aculamprotula_scripta
|
--+ +-------------- Unio_tumidus **
| +-----+
| | | +----------- Unio_crassus **
| | +--+
| | | +-------- Unio_elongatulus **
| | +--+
| | | +----- Unio_pictorum **
| | +--+
| | | +-- Unio_delphinus **
+--+ +--+
| +-- Unio_mancus **
|
| +-- Nodularia_breviconcha
| +--+
| | +-- Nodularia_nipponensis
| +-----------+
| | | +-- Pseudocuneopsis_capitata
| | +--+
+--+ +-- Pseudocuneopsis_sichuanensis
|
| +-------------- Tchangsinaia_piscicula
| |
+--+ +-- Schistodesmus_lampreyanus
| +--------+
| | +-- Schistodesmus_spinosus
+--+
| +-------- Cuneopsis_rufescens
+--+
| +----- Cuneopsis_celtiformis
+--+
| +-- Cuneopsis_demangei
+--+
+-- Cuneopsis_heudei
Wu et al., 2023 [Publ 4339], fig. 4, S5-nt
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
6 |
6 |
3 |
6 |
91 |
17 |
6 |
20.0 |
+-------- Aculamprotula_tortuosa
|
+-----------------+ +-- Aculamprotula_coreana
| | +--+
| | | +-- Aculamprotula_tientsinensis
| +--+
| | +-- Aculamprotula_polysticta
| +--+
| +-- Aculamprotula_scripta
|
--+ +----------- Unio_tumidus **
| |
| +-----------+ +-------- Unio_crassus **
| | | |
| | +--+ +----- Unio_elongatulus **
| | | |
| | +--+ +-- Unio_delphinus **
| | | |
| | +--+-- Unio_mancus **
+--+ |
| +-- Unio_pictorum **
|
| +-- Pseudocuneopsis_capitata
| +-----------------+
| | +-- Pseudocuneopsis_sichuanensis
| |
+--+ +-- Nodularia_breviconcha
| +--------------+
| | +-- Nodularia_nipponensis
| |
+--+ +----- Tchangsinaia_piscicula
| +--------+
| | | +-- Schistodesmus_lampreyanus[2]
| | +--+
+--+ +-- Schistodesmus_spinosus
|
| +----------- Schistodesmus_lampreyanus[1]
+--+
| +-------- Cuneopsis_rufescens
+--+
| +----- Cuneopsis_celtiformis
+--+
| +-- Cuneopsis_demangei
+--+
+-- Cuneopsis_heudei
Wu et al., 2024 [Publ 4350], fig. 8a, S1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
6 |
6 |
4 |
6 |
86 |
16 |
6 |
21.0 |
+-------- Aculamprotula_tortuosa
|
+--------------+ +-- Aculamprotula_coreana
| | +--+
| | | +-- Aculamprotula_tientsinensis
| +--+
| | +-- Aculamprotula_polysticta
| +--+
| +-- Aculamprotula_scripta
|
--+ +-------------- Unio_tumidus **
| +-----+
| | | +----------- Unio_crassus **
| | +--+
| | | +-------- Unio_elongatulus **
| | +--+
| | | +----- Unio_pictorum **
| | +--+
| | | +-- Unio_delphinus **
+--+ +--+
| +-- Unio_mancus **
|
| +-- Nodularia_breviconcha
| +--------------+
| | +-- Nodularia_douglasiae
| |
| | +-- Pseudocuneopsis_capitata
| +--------------+
+--+ +-- Pseudocuneopsis_sichuanensis
|
| +-- Schistodesmus_lampreyanus
| +-----------+
| | +-- Schistodesmus_spinosus
+--+
| +----------- Tchangsinaia_piscicula
+--+
| +-------- Cuneopsis_rufescens
+--+
| +----- Cuneopsis_celtiformis
+--+
| +-- Cuneopsis_demangei
+--+
+-- Cuneopsis_heudei
Wu et al., 2024 [Publ 4350], fig. 8b, S2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
6 |
6 |
4 |
6 |
86 |
16 |
6 |
21.0 |
+-------- Aculamprotula_tortuosa
|
+--------------+ +-- Aculamprotula_coreana
| | +--+
| | | +-- Aculamprotula_tientsinensis
| +--+
| | +-- Aculamprotula_polysticta
| +--+
| +-- Aculamprotula_scripta
|
--+ +-------------- Unio_tumidus **
| +-----+
| | | +----------- Unio_crassus **
| | +--+
| | | +-------- Unio_elongatulus **
| | +--+
| | | +----- Unio_pictorum **
| | +--+
| | | +-- Unio_delphinus **
+--+ +--+
| +-- Unio_mancus **
|
| +-- Nodularia_breviconcha
| +--------------+
| | +-- Nodularia_douglasiae
| |
| | +-- Pseudocuneopsis_capitata
| +--------------+
+--+ +-- Pseudocuneopsis_sichuanensis
|
| +-- Schistodesmus_lampreyanus
| +-----------+
| | +-- Schistodesmus_spinosus
+--+
| +----------- Tchangsinaia_piscicula
+--+
| +-------- Cuneopsis_rufescens
+--+
| +----- Cuneopsis_celtiformis
+--+
| +-- Cuneopsis_demangei
+--+
+-- Cuneopsis_heudei
Prie et al., 2012 [Publ 3487], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
3 |
5 |
1 |
0 |
0 |
18.0 |
+----------- Unio_tumidus **
--+
| +-------- Unio_crassus **
+--+
| +----- Unio_pictorum **
+--+
| +-- Unio_mancus **
+--+
+-- Unio_elongatulus **
Pfarrer et al., 2022 [Publ 4305], fig. 2-5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
3 |
5 |
7 |
0 |
0 |
20.0 |
+----------- Unio_tumidus **
--+
| +-------- Unio_crassus **
+--+
| +----- Unio_elongatulus **
+--+
| +-- Unio_mancus **
+--+
+-- Unio_pictorum **
Nagel et al., 1998 [Publ 1946], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
4 |
5 |
0 |
0 |
21.0 |
+-------- Unio_crassus **
--+
| +----- Unio_tumidus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_mancus **
Nagel et al., 1998 [Publ 1946], fig. 8
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
1 |
4 |
0 |
0 |
0 |
20.0 |
+----- Unio_tumidus **
|
--+ +-- Unio_pictorum **
| |
+--+-- Unio_crassus **
|
+-- Unio_mancus **
Soroka, 2005 [Publ 2726], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
4 |
1 |
0 |
0 |
19.0 |
+-------- Unio_tumidus **
--+
| +----- Unio_crassus **
+--+
| +-- Unio_caffer **
+--+
+-- Unio_pictorum **
Soroka, 2010 [Publ 3404], fig. 3a
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
4 |
5 |
0 |
0 |
19.0 |
+-------- Unio_tumidus **
--+
| +----- Unio_crassus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_mancus **
Soroka, 2010 [Publ 3404], fig. 3b
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
4 |
5 |
0 |
0 |
19.0 |
+-------- Unio_tumidus **
--+
| +----- Unio_crassus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_mancus **
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
1 |
4 |
77 |
7 |
5 |
20.0 |
+----- Unio_gibbus **
|
+-----------+----- Unio_tumidus **
| |
| | +-- Unio_crassus **
| +--+
--+ +-- Unio_pictorum **
|
| +-------------- Aculamprotula_tortuosa
| |
+--+ +-- Nodularia_nuxpersicae
| +--------+
| | +-- Nodularia_douglasiae
+--+
| +-------- Schistodesmus_lampreyanus
+--+
| +----- Tchangsinaia_piscicula
+--+
| +-- Cuneopsis_heudei
+--+
+-- Cuneopsis_rufescens
Huang et al., 2018 [Publ 3828], fig. 6
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
4 |
39 |
5 |
3 |
19.0 |
+----- Aculamprotula_tortuosa
+--------+
| | +-- Aculamprotula_coreana
| +--+
| +-- Aculamprotula_tientsinensis
--+
| +-- Tchangsinaia_piscicula
| +--------+
| | +-- Nodularia_douglasiae
+--+
| +-------- Unio_tumidus **
+--+
| +----- Unio_crassus **
+--+
| +-- Unio_delphinus **
+--+
+-- Unio_pictorum **
Soroka & Burzynski, 2017 [Publ 3837], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
4 |
20 |
6 |
4 |
19.0 |
+----- Aculamprotula_tortuosa
+--------+
| | +-- Aculamprotula_tientsinensis
| +--+
| +-- Aculamprotula_coreana
--+
| +----- Schistodesmus_lampreyanus
| +-----+
| | | +-- Tchangsinaia_piscicula
| | +--+
+--+ +-- Nodularia_douglasiae
|
| +-------- Unio_tumidus **
+--+
| +----- Unio_crassus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_delphinus **
Huang et al., 2019 [Publ 3843], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
4 |
200 |
14 |
5 |
19.0 |
+-------- Aculamprotula_tortuosa
|
+-----------+ +----- Aculamprotula_coreana
| | |
| +--+----- Aculamprotula_scripta
| |
| | +-- Aculamprotula_fibrosa
| +--+
| +-- Aculamprotula_tientsinensis
--+
| +-------- Unio_tumidus **
| +--------+
| | | +----- Unio_crassus **
| | +--+
| | | +-- Unio_delphinus **
| | +--+
+--+ +-- Unio_pictorum **
|
| +-- Schistodesmus_lampreyanus
| +-----------+
| | +-- Schistodesmus_spinosus
| |
+--+ +----- Nodularia_douglasiae[1]
| +-----+
| | | +-- Nodularia_douglasiae[2]
| | +--+
+--+ +-- Nodularia_nuxpersicae
|
| +-------- Tchangsinaia_piscicula
+--+
| +----- Cuneopsis_rufescens
+--+
| +-- Cuneopsis_heudei
+--+
+-- Cuneopsis_celtiformis
Huang et al., 2019 [Publ 3843], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
4 |
53 |
7 |
4 |
19.0 |
+-------- Aculamprotula_tortuosa
+-----+
| | +----- Aculamprotula_scripta
| +--+
| | +-- Aculamprotula_tientsinensis
| +--+
--+ +-- Aculamprotula_coreana
|
| +----- Nodularia_douglasiae
| +-----+
| | | +-- Tchangsinaia_piscicula
| | +--+
+--+ +-- Schistodesmus_lampreyanus
|
| +-------- Unio_tumidus **
+--+
| +----- Unio_crassus **
+--+
| +-- Unio_delphinus **
+--+
+-- Unio_pictorum **
Huang et al., 2019 [Publ 3843], fig. S3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
4 |
197 |
13 |
5 |
19.0 |
+-------- Aculamprotula_tortuosa
|
+-----------+ +-- Aculamprotula_coreana
| | +--+
| | | +-- Aculamprotula_scripta
| +--+
| | +-- Aculamprotula_fibrosa
| +--+
| +-- Aculamprotula_tientsinensis
--+
| +-------- Unio_tumidus **
| +--------+
| | | +----- Unio_crassus **
| | +--+
| | | +-- Unio_delphinus **
| | +--+
+--+ +-- Unio_pictorum **
|
| +----- Nodularia_douglasiae[1]
| +--------+
| | | +-- Nodularia_douglasiae[2]
| | +--+
+--+ +-- Nodularia_nuxpersicae
|
| +----------- Schistodesmus_lampreyanus
+--+
| +-------- Tchangsinaia_piscicula
+--+
| +----- Cuneopsis_rufescens
+--+
| +-- Cuneopsis_heudei
+--+
+-- Cuneopsis_celtiformis
Huang et al., 2019 [Publ 3843], fig. S4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
4 |
198 |
14 |
5 |
19.0 |
+-------- Unio_tumidus **
--+
| +----- Unio_crassus **
+--+
| +-- Unio_delphinus **
+--+
+-- Unio_pictorum **
Wang et al., 2018 [Publ 3846], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
4 |
28 |
4 |
4 |
19.0 |
+-------------- Aculamprotula_tientsinensis
|
| +----- Nodularia_douglasiae
--+ +-----+
| | | +-- Tchangsinaia_piscicula
| | +--+
+--+ +-- Cuneopsis_heudei
|
| +-------- Unio_tumidus **
+--+
| +----- Unio_crassus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_delphinus **
Burzynski & Soroka, 2018 [Publ 3862], fig. 5a
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
4 |
16 |
1 |
1 |
19.0 |
+----------- Aculamprotula_tortuosa
--+
| +-------- Unio_tumidus **
+--+
| +----- Unio_crassus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_delphinus **
Burzynski & Soroka, 2018 [Publ 3862], fig. 5b
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
4 |
6 |
1 |
1 |
19.0 |
+----------- Aculamprotula_tortuosa
--+
| +-------- Unio_tumidus **
+--+
| +----- Unio_crassus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_delphinus **
Aunins et al., 2018 [Publ 3863], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
4 |
37 |
7 |
5 |
19.0 |
+----- Aculamprotula_tortuosa
+--------+
| | +-- Aculamprotula_coreana
| +--+
| +-- Aculamprotula_tientsinensis
|
--+ +-------- Nodularia_douglasiae
| +--+
| | | +----- Tchangsinaia_piscicula
| | +--+
| | | +-- Cuneopsis_heudei
+--+ +--+
| +-- Schistodesmus_lampreyanus
|
| +-------- Unio_tumidus **
+--+
| +----- Unio_crassus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_delphinus **
Wu et al., 2019 [Publ 3864], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
4 |
43 |
9 |
6 |
19.0 |
+-------- Aculamprotula_tortuosa
+--------+
| | +----- Aculamprotula_scripta
| +--+
| | +-- Aculamprotula_tientsinensis
| +--+
| +-- Aculamprotula_coreana
--+
| +-------- Unio_tumidus **
| +-----+
| | | +----- Unio_crassus **
| | +--+
| | | +-- Unio_delphinus **
+--+ +--+
| +-- Unio_pictorum **
|
| +----------- Pseudocuneopsis_capitata
+--+
| +-------- Nodularia_douglasiae
+--+
| +----- Tchangsinaia_piscicula
+--+
| +-- Cuneopsis_heudei
+--+
+-- Schistodesmus_lampreyanus
Wang et al., 2018 [Publ 3865], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
4 |
28 |
3 |
3 |
19.0 |
+-------------- Aculamprotula_tientsinensis
|
--+ +-- Tchangsinaia_piscicula
| +--------+
| | +-- Nodularia_douglasiae
+--+
| +-------- Unio_tumidus **
+--+
| +----- Unio_crassus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_delphinus **
Klishko et al., 2019 [Publ 3886], fig. 9
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
1 |
4 |
7 |
4 |
2 |
20.0 |
+-------- Middendorffinaia_mongolica
+--+
| | +----- Nodularia_breviconcha
| +--+
| | +-- Nodularia_douglasiae
--+ +--+
| +-- Nodularia
|
| +----- Unio_tumidus **
| |
+-----+----- Unio_gibbus **
|
| +-- Unio_crassus **
+--+
+-- Unio_pictorum **
Cha et al., 2018 [Publ 3916], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
4 |
35 |
6 |
4 |
19.0 |
+----- Aculamprotula_tortuosa
+--------+
| | +-- Aculamprotula_coreana
| +--+
| +-- Aculamprotula_tientsinensis
--+
| +----- Schistodesmus_lampreyanus
| +-----+
| | | +-- Nodularia_douglasiae
| | +--+
+--+ +-- Tchangsinaia_piscicula
|
| +-------- Unio_tumidus **
+--+
| +----- Unio_crassus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_delphinus **
Froufe et al., 2019 [Publ 3936], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
4 |
41 |
6 |
4 |
19.0 |
+----- Aculamprotula_tortuosa
+--------+
| | +-- Aculamprotula_tientsinensis
| +--+
| +-- Aculamprotula_coreana
--+
| +----- Nodularia_douglasiae
| +-----+
| | | +-- Tchangsinaia_piscicula
| | +--+
+--+ +-- Schistodesmus_lampreyanus
|
| +-------- Unio_tumidus **
+--+
| +----- Unio_crassus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_delphinus **
Froufe et al., 2019 [Publ 3936], fig. 6
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
4 |
41 |
6 |
4 |
19.0 |
+----- Aculamprotula_tortuosa
+--------+
| | +-- Aculamprotula_tientsinensis
| +--+
| +-- Aculamprotula_coreana
--+
| +----- Nodularia_douglasiae
| +-----+
| | | +-- Tchangsinaia_piscicula
| | +--+
+--+ +-- Schistodesmus_lampreyanus
|
| +-------- Unio_tumidus **
+--+
| +----- Unio_crassus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_delphinus **
Liu et al., 2017 [Publ 3939], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
1 |
4 |
13 |
9 |
3 |
20.0 |
+----- Unio_tumidus **
|
+-----------+----- Unio_gibbus **
| |
| | +-- Unio_crassus **
| +--+
| +-- Unio_pictorum **
--+
| +-------------- Schistodesmus_lampreyanus
| |
| | +-- Cuneopsis_heudei
| +-----------+
+--+ +-- Cuneopsis_rufescens
|
| +----------- Nodularia_breviconcha
| |
+--+ +-------- Nodularia_douglasiae[3]
| |
+--+ +----- Nodularia_douglasiae[2]
| |
+--+ +-- Nodularia_douglasiae[1]
| |
+--+-- Nodularia_nuxpersicae
|
+-- Nodularia_nipponensis
Klishko et al., 2018 [Publ 3945], fig. 8
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
1 |
4 |
4 |
4 |
1 |
20.0 |
+-------- Nodularia_breviconcha
+--+
| | +----- Nodularia
| +--+
| | +-- Nodularia_douglasiae
--+ +--+
| +-- Nodularia_nipponensis
|
| +----- Unio_tumidus **
| |
+-----+----- Unio_gibbus **
|
| +-- Unio_crassus **
+--+
+-- Unio_pictorum **
Klishko et al., 2017 [Publ 3946], fig. 8
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
4 |
2 |
2 |
1 |
21.0 |
+-- Nodularia_douglasiae
+--------+
| +-- Nodularia_breviconcha
--+
| +-------- Unio_tumidus **
+--+
| +----- Unio_pictorum **
+--+
| +-- Unio_crassus **
+--+
+-- Unio_courtillieri **
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
1 |
4 |
118 |
22 |
7 |
20.0 |
+----------------------- Middendorffinaia_mongolica
|
| +----- Unio_gibbus **
| |
| +--------------+----- Unio_tumidus **
--+ | |
| | | +-- Unio_crassus **
| | +--+
| | +-- Unio_pictorum **
| |
| | +-------------- Nodularia_breviconcha
+--+ +--+
| | | +----------- Nodularia[2]
| | +--+
| | | +-------- Nodularia_nuxpersicae
| | +--+
| | | +----- Nodularia_nipponensis
| | +--+
| | | +-- Nodularia_douglasiae
+--+ +--+
| +-- Nodularia[1]
|
| +-- Schistodesmus_lampreyanus
| +-----------+
| | +-- Schistodesmus_spinosus
| |
| | +----- Cuneopsis_rufescens
| +--------+
+--+ | +-- Cuneopsis_heudei
| +--+
| +-- Cuneopsis_celtiformis
|
| +----------- Tchangsinaia_piscicula
+--+
| +-------- Inversiunio_jokohamensis
+--+
| +----- Inversiunio_verrucosus
+--+
| +-- Inversiunio_reinianus
+--+
+-- Inversiunio_yanagawensis
Fukata & Iigo, 2020 [Publ 4106], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
4 |
50 |
8 |
6 |
19.0 |
+----- Aculamprotula_tortuosa
+--------+
| | +-- Aculamprotula_tientsinensis
| +--+
| +-- Aculamprotula_coreana
|
--+ +-------- Unio_tumidus **
| +--+
| | | +----- Unio_crassus **
| | +--+
| | | +-- Unio_pictorum **
| | +--+
+--+ +-- Unio_delphinus **
|
| +-- Schistodesmus_lampreyanus
| +-----+
| | +-- Cuneopsis_heudei
+--+
| +----- Pseudocuneopsis_capitata
+--+
| +-- Nodularia_douglasiae
+--+
+-- Tchangsinaia_piscicula
Fukata & Iigo, 2019 [Publ 4108], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
4 |
10 |
6 |
6 |
19.0 |
+-------- Unio_tumidus **
+--+
| | +----- Unio_crassus **
| +--+
| | +-- Unio_delphinus **
--+ +--+
| +-- Unio_pictorum **
|
| +-------- Pseudocuneopsis_capitata
| |
+--+ +-- Nodularia_douglasiae
| +--+
| | +-- Tchangsinaia_piscicula
+--+
| +-- Schistodesmus_lampreyanus
+--+
+-- Cuneopsis_heudei
Wu et al., 2020 [Publ 4121], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
4 |
46 |
10 |
6 |
19.0 |
+-------- Aculamprotula_tortuosa
|
+--------+ +-- Aculamprotula_coreana
| | +--+
| | | +-- Aculamprotula_tientsinensis
| +--+
| | +-- Aculamprotula_polysticta
| +--+
--+ +-- Aculamprotula_scripta
|
| +-------- Unio_tumidus **
| +-----+
| | | +----- Unio_crassus **
| | +--+
| | | +-- Unio_delphinus **
+--+ +--+
| +-- Unio_pictorum **
|
| +----------- Pseudocuneopsis_capitata
+--+
| +-------- Nodularia_douglasiae
+--+
| +----- Tchangsinaia_piscicula
+--+
| +-- Cuneopsis_heudei
+--+
+-- Schistodesmus_lampreyanus
Kim et al., 2021 [Publ 4197], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
4 |
11 |
5 |
4 |
19.0 |
+-------- Unio_tumidus **
+--+
| | +----- Unio_crassus **
| +--+
| | +-- Unio_delphinus **
| +--+
--+ +-- Unio_pictorum **
|
| +-- Nodularia_douglasiae
| +--+
| | +-- Nodularia_breviconcha
+-----+
| +-- Schistodesmus_lampreyanus
+--+
+-- Cuneopsis_heudei
Zieritz et al., 2021 [Publ 4231], fig. 3-F, 5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
4 |
45 |
6 |
4 |
19.0 |
+----- Aculamprotula_tortuosa
+--------+
| | +-- Aculamprotula_tientsinensis
| +--+
| +-- Aculamprotula_coreana
--+
| +----- Nodularia_douglasiae
| +-----+
| | | +-- Schistodesmus_lampreyanus
| | +--+
+--+ +-- Tchangsinaia_piscicula
|
| +-------- Unio_tumidus **
+--+
| +----- Unio_crassus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_delphinus **
Wu et al., 2022 [Publ 4238], fig. 2d
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
4 |
35 |
19 |
7 |
19.0 |
+-------- Aculamprotula_tortuosa
|
+-----------------+ +-- Aculamprotula_tientsinensis
| | +--+
| | | +-- Aculamprotula_coreana
| +--+
| | +-- Aculamprotula_polysticta
| +--+
| +-- Aculamprotula_scripta
--+
| +-------- Unio_tumidus **
| +--------------+
| | | +----- Unio_crassus **
| | +--+
| | | +-- Unio_pictorum **
| | +--+
| | +-- Unio_delphinus **
+--+
| +-- Pseudocuneopsis_capitata
| +-----------------+
| | +-- Pseudocuneopsis_sichuanensis
| |
| | +-- Nodularia_breviconcha
+--+ +--------------+
| | +-- Nodularia_douglasiae
| |
| | +-- Schistodesmus_lampreyanus
+--+ +-----------+
| | +-- Schistodesmus_spinosus
| |
| | +-------- Cuneopsis_rufescens
+--+ +--+
| | | +----- Cuneopsis_celtiformis
| | +--+
| | | +-- Cuneopsis_heudei
+--+ +--+
| +-- Cuneopsis_demangei
|
| +-------- Tchangsinaia_piscicula
+--+
| +----- Inversiunio_jokohamensis
+--+
| +-- Inversiunio_reinianus
+--+
+-- Inversiunio_yanagawensis
Wu et al., 2022 [Publ 4238], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
4 |
70 |
16 |
6 |
19.0 |
+-------- Aculamprotula_tortuosa
|
+-----------------+ +-- Aculamprotula_tientsinensis
| | +--+
| | | +-- Aculamprotula_coreana
| +--+
| | +-- Aculamprotula_polysticta
| +--+
| +-- Aculamprotula_scripta
--+
| +-------- Unio_tumidus **
| +--------------+
| | | +----- Unio_crassus **
| | +--+
| | | +-- Unio_pictorum **
| | +--+
+--+ +-- Unio_delphinus **
|
| +-- Pseudocuneopsis_capitata
| +-----------------+
| | +-- Pseudocuneopsis_sichuanensis
| |
+--+ +-- Nodularia_douglasiae
| +--------------+
| | +-- Nodularia_breviconcha
| |
+--+ +-- Schistodesmus_lampreyanus
| +-----------+
| | +-- Schistodesmus_spinosus
+--+
| +----------- Tchangsinaia_piscicula
+--+
| +-------- Cuneopsis_rufescens
+--+
| +----- Cuneopsis_celtiformis
+--+
| +-- Cuneopsis_heudei
+--+
+-- Cuneopsis_demangei
Manendo et al., 2008 [Publ 2669], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
10 |
0 |
0 |
22.0 |
+----- Unio_crassus **
--+
| +-- Unio_pictorum **
+--+
+-- Unio_tumidus **
Källersjö et al., 2005 [Publ 2684], fig. 1A
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
0 |
3 |
4 |
0 |
0 |
21.0 |
+-- Unio_crassus **
|
--+-- Unio_pictorum **
|
+-- Unio_tumidus **
Källersjö et al., 2005 [Publ 2684], fig. 1B-F
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
4 |
0 |
0 |
20.0 |
+----- Unio_tumidus **
--+
| +-- Unio_crassus **
+--+
+-- Unio_pictorum **
Soroka, 2010 [Publ 3404], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
0 |
3 |
4 |
0 |
0 |
21.0 |
+-- Unio_tumidus **
|
--+-- Unio_pictorum **
|
+-- Unio_crassus **
Soroka, 2010 [Publ 3404], fig. 2-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
2 |
0 |
0 |
22.0 |
+----- Unio_crassus **
--+
| +-- Unio_tumidus **
+--+
+-- Unio_pictorum **
Soroka, 2010 [Publ 3404], fig. 3a-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
3 |
0 |
0 |
22.0 |
+----- Unio_crassus **
--+
| +-- Unio_tumidus **
+--+
+-- Unio_pictorum **
Soroka, 2010 [Publ 3404], fig. 3b-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
3 |
0 |
0 |
20.0 |
+----- Unio_tumidus **
--+
| +-- Unio_crassus **
+--+
+-- Unio_pictorum **
Soroka, 2010 [Publ 3404], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
4 |
0 |
0 |
20.0 |
+----- Unio_tumidus **
--+
| +-- Unio_pictorum **
+--+
+-- Unio_crassus **
Soroka, 2010 [Publ 3404], fig. 5a
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
5 |
0 |
0 |
22.0 |
+----- Unio_crassus **
--+
| +-- Unio_pictorum **
+--+
+-- Unio_tumidus **
Soroka, 2010 [Publ 3404], fig. 5b
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
5 |
0 |
0 |
20.0 |
+----- Unio_tumidus **
--+
| +-- Unio_pictorum **
+--+
+-- Unio_crassus **
Soroka, 2010 [Publ 3404], fig. 8a
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
3 |
0 |
0 |
20.0 |
+----- Unio_tumidus **
--+
| +-- Unio_pictorum **
+--+
+-- Unio_crassus **
Soroka, 2010 [Publ 3404], fig. 8b
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
3 |
0 |
0 |
20.0 |
+----- Unio_tumidus **
--+
| +-- Unio_pictorum **
+--+
+-- Unio_crassus **
Zieritz et al., 2012 [Publ 3673], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
5 |
0 |
0 |
22.0 |
+----- Unio_crassus **
--+
| +-- Unio_tumidus **
+--+
+-- Unio_pictorum **
Plazzi et al., 2011 [Publ 3676], fig. 5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
15 |
0 |
0 |
20.0 |
+----- Unio_tumidus **
--+
| +-- Unio_crassus **
+--+
+-- Unio_pictorum **
Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
8 |
5 |
3 |
23 |
1 |
1 |
27.0 |
+-------------- Unio_pictorum[2] **
|
--+ +-- Unio_pictorum[1] **
| +--------+
| | +-- Unio_caffer **
+--+
| +-------- Anodonta
| |
+--+ +----- Lasmigona_costata
| |
+--+ +-- Strophitus_undulatus
| |
+--+-- Alasmidonta_marginata
|
+-- Pyganodon_grandis
Bolotov et al., 2017 [Publ 3761], fig. S1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
138 |
7 |
5 |
20.0 |
+----- Unio_tumidus **
+--------+
| | +-- Unio_pictorum **
| +--+
| +-- Unio_crassus **
--+
| +----------- Aculamprotula_tortuosa
| |
| | +-- Nodularia_douglasiae
+--+ +-----+
| | +-- Nodularia_nuxpersicae
| |
+--+ +-- Cuneopsis_rufescens
| +--+
| | +-- Cuneopsis_heudei
+--+
| +-- Tchangsinaia_piscicula
+--+
+-- Schistodesmus_lampreyanus
Bolotov et al., 2017 [Publ 3761], fig. S2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
138 |
7 |
5 |
20.0 |
+----- Unio_tumidus **
+-----+
| | +-- Unio_crassus **
| +--+
| +-- Unio_pictorum **
--+
| +-------- Aculamprotula_tortuosa
| |
| +-------- Schistodesmus_lampreyanus
| |
+--+ +-- Nodularia_douglasiae
+-----+
| +-- Nodularia_nuxpersicae
|
| +----- Cuneopsis_heudei
+--+
| +-- Tchangsinaia_piscicula
+--+
+-- Cuneopsis_rufescens
Bolotov et al., 2017 [Publ 3764], fig. S1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
10 |
0 |
3 |
128 |
7 |
5 |
28.0 |
+-- Unio_crassus **
|
+-- Unio_pictorum **
|
+-- Unio_tumidus **
|
+-- Aculamprotula_tortuosa
|
+-- Cuneopsis_heudei
--+
+-- Tchangsinaia_piscicula
|
+-- Cuneopsis_rufescens
|
+-- Nodularia_douglasiae
|
+-- Nodularia_nuxpersicae
|
+-- Schistodesmus_lampreyanus
Froufe et al., 2017 [Publ 3802], fig. 7
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
7 |
0 |
0 |
22.0 |
+----- Unio_mancus **
--+
| +-- Unio_elongatulus **
+--+
+-- Unio_pictorum **
Dyduch-Falniowska & Koziol, 1989 [Publ 3814], fig. 5, 7
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
2 |
0 |
0 |
22.0 |
+----- Unio_pictorum **
--+
| +-- Unio_tumidus **
+--+
+-- Unio_crassus **
Bolotov et al., 2018 [Publ 3826], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
82 |
3 |
1 |
20.0 |
+----- Unio_tumidus **
+--+
| | +-- Unio_crassus **
| +--+
--+ +-- Unio_pictorum **
|
| +----- Nodularia_douglasiae[1]
+--+
| +-- Nodularia_nuxpersicae
+--+
+-- Nodularia_douglasiae[2]
Huang et al., 2018 [Publ 3828], fig. 6-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
14 |
1 |
1 |
20.0 |
+-------- Aculamprotula_tortuosa
--+
| +----- Unio_tumidus **
+--+
| +-- Unio_crassus **
+--+
+-- Unio_delphinus **
Huang et al., 2019 [Publ 3843], fig. 3-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
20 |
2 |
2 |
20.0 |
+----------- Schistodesmus_lampreyanus
--+
| +-------- Aculamprotula_tortuosa
+--+
| +----- Unio_tumidus **
+--+
| +-- Unio_crassus **
+--+
+-- Unio_delphinus **
Bogatov et al., 2018 [Publ 3859], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
5 |
3 |
3 |
3 |
0 |
0 |
22.0 |
+-------- Unio_tumidus **
|
--+ +-- Anodonta_anatina
| +--+
| | +-- Pseudanodonta_complanata
+--+
| +-- Unio_crassus **
+--+
+-- Unio_pictorum **
Bogatov et al., 2018 [Publ 3859], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
5 |
3 |
3 |
3 |
0 |
0 |
24.0 |
+----------- Unio_crassus **
--+
| +-------- Unio_pictorum **
+--+
| +----- Unio_tumidus **
+--+
| +-- Anodonta_anatina
+--+
+-- Pseudanodonta_complanata
Froufe et al., 2019 [Publ 3936], fig. 3-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
16 |
0 |
0 |
20.0 |
+----- Unio_tumidus **
--+
| +-- Unio_crassus **
+--+
+-- Unio_delphinus **
Froufe et al., 2019 [Publ 3936], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
16 |
0 |
0 |
20.0 |
+----- Unio_tumidus **
--+
| +-- Unio_crassus **
+--+
+-- Unio_delphinus **
Bolotov et al., 2020 [Publ 4084], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
33 |
2 |
2 |
20.0 |
+----------- Middendorffinaia_mongolica
--+
| +-------- Nodularia_douglasiae
+--+
| +----- Unio_tumidus **
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_crassus **
Sano et al., 2020 [Publ 4138], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
70 |
12 |
5 |
20.0 |
+----- Aculamprotula_tortuosa
+-----------+
| | +-- Aculamprotula_tientsinensis
| +--+
| +-- Aculamprotula_coreana
|
--+ +----- Unio_crassus **
| +--------+
| | | +-- Unio_delphinus **
| | +--+
| | +-- Unio_pictorum **
| |
+--+ +-------- Inversiunio_jokohamensis
| +--+
| | | +----- Inversiunio_yanagawensis[2]
| | +--+
| | | +-- Inversiunio_reinianus
| | +--+
+--+ +-- Inversiunio_yanagawensis[1]
|
| +-- Tchangsinaia_piscicula
| +-----+
| | +-- Schistodesmus_lampreyanus
+--+
| +----- Nodularia_douglasiae[2]
+--+
| +-- Nodularia_nipponensis
+--+
+-- Nodularia_douglasiae[1]
Sano et al., 2020 [Publ 4138], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
69 |
11 |
5 |
20.0 |
+----- Aculamprotula_tortuosa
+-----------+
| | +-- Aculamprotula_tientsinensis
| +--+
| +-- Aculamprotula_coreana
|
--+ +----- Unio_crassus **
| +--------+
| | | +-- Unio_delphinus **
| | +--+
| | +-- Unio_pictorum **
| |
+--+ +-------- Inversiunio_jokohamensis
| +--+
| | | +----- Inversiunio_yanagawensis[2]
| | +--+
| | | +-- Inversiunio_reinianus
| | +--+
+--+ +-- Inversiunio_yanagawensis[1]
|
| +-- Nodularia_nipponensis
| +--+
| | +-- Nodularia_douglasiae
+-----+
| +-- Schistodesmus_lampreyanus
+--+
+-- Tchangsinaia_piscicula
Sano et al., 2020 [Publ 4138], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
69 |
11 |
5 |
20.0 |
+----- Aculamprotula_tortuosa
+--------------+
| | +-- Aculamprotula_tientsinensis
| +--+
| +-- Aculamprotula_coreana
|
--+ +----- Unio_crassus **
| +-----------+
| | | +-- Unio_delphinus **
| | +--+
| | +-- Unio_pictorum **
+--+
| +-- Nodularia_douglasiae
| +-----------+
| | +-- Nodularia_nipponensis
| |
+--+ +-- Schistodesmus_lampreyanus
| +--------+
| | +-- Tchangsinaia_piscicula
+--+
| +-------- Inversiunio_jokohamensis
+--+
| +----- Inversiunio_yanagawensis[2]
+--+
| +-- Inversiunio_reinianus
+--+
+-- Inversiunio_yanagawensis[1]
Guerra et al., 2019 [Publ 4168], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
38 |
2 |
2 |
20.0 |
+----------- Aculamprotula_tortuosa
--+
| +-------- Schistodesmus_lampreyanus
+--+
| +----- Unio_tumidus **
+--+
| +-- Unio_crassus **
+--+
+-- Unio_delphinus **
Guerra et al., 2019 [Publ 4168], fig. 1-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
19 |
2 |
2 |
20.0 |
+----------- Schistodesmus_lampreyanus
--+
| +-------- Aculamprotula_tortuosa
+--+
| +----- Unio_tumidus **
+--+
| +-- Unio_crassus **
+--+
+-- Unio_delphinus **
Guerra et al., 2019 [Publ 4168], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
38 |
2 |
2 |
20.0 |
+-------- Schistodesmus_lampreyanus
--+
| +----- Unio_tumidus **
+--+
| +-- Unio_crassus **
+--+
+-- Unio_delphinus **
Guerra et al., 2019 [Publ 4168], fig. 2-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
19 |
2 |
2 |
20.0 |
+----------- Aculamprotula_tortuosa
--+
| +-------- Schistodesmus_lampreyanus
+--+
| +----- Unio_tumidus **
+--+
| +-- Unio_crassus **
+--+
+-- Unio_delphinus **
Zieritz et al., 2021 [Publ 4231], fig. 3-M, 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
20 |
0 |
0 |
20.0 |
+----- Unio_tumidus **
--+
| +-- Unio_delphinus **
+--+
+-- Unio_crassus **
Wu et al., 2022 [Publ 4238], fig. 2e
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
14 |
9 |
6 |
20.0 |
+-------------------- Aculamprotula_tortuosa
|
| +----- Unio_tumidus **
--+ +-----------+
| | | +-- Unio_crassus **
| | +--+
| | +-- Unio_delphinus **
+--+
| +-- Pseudocuneopsis_sichuanensis
| +-----------+
| | +-- Pseudocuneopsis_capitata
+--+
| +----------- Nodularia_douglasiae
| |
+--+ +-------- Tchangsinaia_piscicula
| |
+--+ +-- Cuneopsis_celtiformis
| +--+
| | +-- Cuneopsis_heudei
+--+
| +-- Schistodesmus_lampreyanus
+--+
+-- Schistodesmus_spinosus
Jeratthitikul et al., 2022 [Publ 4260], fig. S1, 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
73 |
7 |
5 |
20.0 |
+----- Unio_tumidus **
+-----------+
| | +-- Unio_pictorum **
| +--+
--+ +-- Unio_crassus **
|
| +-------------- Aculamprotula_tortuosa
| |
+--+ +-- Nodularia_douglasiae
| +--------+
| | +-- Nodularia_nuxpersicae
+--+
| +-------- Schistodesmus_lampreyanus
+--+
| +----- Cuneopsis_heudei
+--+
| +-- Tchangsinaia_piscicula
+--+
+-- Cuneopsis_rufescens
Jeratthitikul et al., 2022 [Publ 4260], fig. S2, 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
73 |
7 |
5 |
20.0 |
+----- Unio_tumidus **
+-----------+
| | +-- Unio_pictorum **
| +--+
--+ +-- Unio_crassus **
|
| +-------------- Aculamprotula_tortuosa
| |
+--+ +-- Nodularia_douglasiae
| +--------+
| | +-- Nodularia_nuxpersicae
+--+
| +-------- Schistodesmus_lampreyanus
+--+
| +----- Cuneopsis_heudei
+--+
| +-- Tchangsinaia_piscicula
+--+
+-- Cuneopsis_rufescens
Konopleva et al., 2023 [Publ 4269], fig. S1, 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
3 |
53 |
9 |
5 |
20.0 |
+-------- Aculamprotula_tortuosa
+-----+
| | +----- Aculamprotula_scripta
| +--+
| | +-- Aculamprotula_tientsinensis
| +--+
--+ +-- Aculamprotula_coreana
|
| +----- Unio_tumidus **
| +-----+
| | | +-- Unio_pictorum **
| | +--+
+--+ +-- Unio_crassus **
|
| +-------- Nodularia_douglasiae
+--+
| +----- Tchangsinaia_piscicula
+--+
| +-- Cuneopsis_rufescens
+--+
+-- Cuneopsis_heudei
Graf & Cummings, 2006 [Publ 1968], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
45 |
0 |
0 |
-- |
+-- Unio_pictorum **
--+
+-- Unio_caffer **
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
28 |
0 |
0 |
-- |
+-- Unio_pictorum **
--+
+-- Unio_caffer **
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
28 |
0 |
0 |
-- |
+-- Unio_pictorum **
--+
+-- Unio_caffer **
Bogan & Hoeh, 2000 [Publ 2650], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
32 |
0 |
0 |
-- |
+-- Unio_pictorum **
--+
+-- Unio_tumidus **
Graf, 2002 [Publ 2651], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
16 |
0 |
0 |
-- |
+-- Unio_pictorum **
--+
+-- Unio_caffer **
Hoeh et al., 2001 [Publ 2652], fig. 14.2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
29 |
0 |
0 |
-- |
+-- Unio_pictorum **
--+
+-- Unio_tumidus **
Hoeh et al., 2001 [Publ 2652], fig. 14.4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
5 |
0 |
2 |
29 |
0 |
0 |
24.0 |
+-------- Unio_pictorum **
|
--+-------- Unio_tumidus **
|
| +----- Anodonta_cygnea
+--+
| +-- Strophitus_undulatus
+--+
+-- Pyganodon_grandis
Hoeh et al., 2002 [Publ 2653], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
5 |
0 |
2 |
29 |
0 |
0 |
24.0 |
+----------- Unio_pictorum **
--+
| +-------- Unio_tumidus **
+--+
| +----- Anodonta_cygnea
+--+
| +-- Pyganodon_grandis
+--+
+-- Strophitus_undulatus
Roe & Hoeh, 2003 [Publ 2655], fig. 4.2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
29 |
0 |
0 |
-- |
+-- Unio_pictorum **
--+
+-- Unio_tumidus **
Roe & Hoeh, 2003 [Publ 2655], fig. 4.3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
29 |
0 |
0 |
-- |
+-- Unio_pictorum **
--+
+-- Unio_tumidus **
Hoeh et al., 2009 [Publ 2716], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
45 |
0 |
0 |
-- |
+-- Unio_pictorum **
--+
+-- Unio_caffer **
Hoeh et al., 2009 [Publ 2716], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
44 |
0 |
0 |
-- |
+-- Unio_pictorum **
--+
+-- Unio_caffer **
Hoeh et al., 2009 [Publ 2716], fig. 5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
44 |
0 |
0 |
-- |
+-- Unio_pictorum **
--+
+-- Unio_caffer **
Whelan et al., 2011 [Publ 3380], fig. 1 ML
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
35 |
0 |
0 |
-- |
+-- Unio_pictorum **
--+
+-- Unio_caffer **
Whelan et al., 2011 [Publ 3380], fig. 1 MP
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
35 |
0 |
0 |
-- |
+-- Unio_pictorum **
--+
+-- Unio_caffer **
Soroka, 2010 [Publ 3404], fig. 8a-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
1 |
0 |
0 |
-- |
+-- Unio_pictorum **
--+
+-- Unio_tumidus **
Soroka, 2010 [Publ 3404], fig. 8b-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
1 |
0 |
0 |
-- |
+-- Unio_pictorum **
--+
+-- Unio_tumidus **
Bolotov et al., 2016 [Publ 3743], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
28 |
0 |
0 |
-- |
+-- Unio_pictorum **
--+
+-- Unio_ravoisieri **
Bolotov et al., 2016 [Publ 3743], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
28 |
0 |
0 |
-- |
+-- Unio_pictorum **
--+
+-- Unio_ravoisieri **
Burzynski et al., 2017 [Publ 3774], fig. 1 bottom
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
11 |
1 |
1 |
-- |
+----- Aculamprotula_tortuosa
--+
| +-- Unio_crassus **
+--+
+-- Unio_tumidus **
Burzynski et al., 2017 [Publ 3774], fig. 1 top
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
11 |
1 |
1 |
-- |
+----- Aculamprotula_tortuosa
--+
| +-- Unio_crassus **
+--+
+-- Unio_tumidus **
Fonseca et al., 2016 [Publ 3775], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
30 |
5 |
4 |
-- |
+-- Aculamprotula_tortuosa
+--------+
| +-- Aculamprotula_coreana
|
--+ +-- Unio_pictorum **
| +-----+
| | +-- Unio_delphinus **
+--+
| +----- Schistodesmus_lampreyanus
+--+
| +-- Nodularia_douglasiae
+--+
+-- Tchangsinaia_piscicula
Lopes-Lima et al., 2017 [Publ 3792], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
31 |
5 |
4 |
-- |
+-- Aculamprotula_tortuosa
+--------+
| +-- Aculamprotula_coreana
|
--+ +-- Unio_delphinus **
| +-----+
| | +-- Unio_pictorum **
+--+
| +----- Schistodesmus_lampreyanus
+--+
| +-- Tchangsinaia_piscicula
+--+
+-- Nodularia_douglasiae
Wu et al., 2018 [Publ 3836], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
51 |
12 |
5 |
-- |
+----------- Aculamprotula_tortuosa
+-----+
| | +-------- Aculamprotula_scripta
| +--+
| | +----- Aculamprotula_coreana
| +--+
| | +-- Aculamprotula_fibrosa
--+ +--+
| +-- Aculamprotula_tientsinensis
|
| +-- Unio_crassus **
| +-----------+
| | +-- Unio_pictorum **
| |
+--+ +----- Nodularia_douglasiae
| +-----+
| | | +-- Schistodesmus_lampreyanus
| | +--+
+--+ +-- Schistodesmus_spinosus
|
| +-------- Tchangsinaia_piscicula
+--+
| +----- Cuneopsis_rufescens
+--+
| +-- Cuneopsis_celtiformis
+--+
+-- Cuneopsis_heudei
Chase et al., 2018 [Publ 3847], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
37 |
0 |
0 |
-- |
+-- Unio_delphinus **
--+
+-- Unio_tumidus **
Chase et al., 2018 [Publ 3847], fig. 5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
8 |
0 |
0 |
-- |
+-- Unio_delphinus **
--+
+-- Unio_tumidus **
Chase et al., 2018 [Publ 3861], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
9 |
0 |
0 |
-- |
+-- Unio_delphinus **
--+
+-- Unio_tumidus **
Chase et al., 2018 [Publ 3861], fig. 1-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
9 |
0 |
0 |
-- |
+-- Unio_delphinus **
--+
+-- Unio_tumidus **
Riccardi et al., 2019 [Publ 4089], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
18 |
0 |
0 |
-- |
+-- Unio_mancus **
--+
+-- Unio_elongatulus **
Tomilova et al., 2020 [Publ 4097], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
6 |
0 |
0 |
-- |
+-- Unio_tumidus **
--+
+-- Unio_pictorum **
Sayenko et al., 2021 [Publ 4189], fig. 6
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
11 |
11 |
3 |
-- |
+-- Unio_pictorum **
+-----------------+
| +-- Unio_crassus **
|
| +-------- Inversiunio_jokohamensis
--+ |
| +--------+ +----- Inversiunio_yanagawensis[2]
| | | |
| | +--+----- Inversiunio_verrucosus
| | |
+--+ | +-- Inversiunio_reinianus
| +--+
| +-- Inversiunio_yanagawensis[1]
|
| +-------------- Middendorffinaia_mongolica
+--+
| +----------- Nodularia_breviconcha
+--+
| +-------- Nodularia_douglasiae[2]
+--+
| +----- Nodularia_nuxpersicae
+--+
| +-- Nodularia_douglasiae[1]
+--+
+-- Nodularia_nipponensis
Sonowal et al., 2021 [Publ 4221], fig. 3a
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
4 |
0 |
2 |
13 |
1 |
1 |
23.0 |
+-- Anodonta_cygnea
|
+-- Unio_ravoisieri **
--+
+-- Unio_pictorum **
|
+-- Aculamprotula_tientsinensis
Sonowal et al., 2021 [Publ 4221], fig. 3b
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
9 |
0 |
2 |
13 |
1 |
1 |
28.0 |
+-- Aculamprotula_tientsinensis
|
+-- Lamprotula_leaii
|
+-- Lampsilis_cardium
|
+-- Potomida_littoralis
|
--+-- Unio_pictorum **
|
+-- Anodonta_cygnea
|
+-- Unio_ravoisieri **
|
+-- Lamellidens_mainwaringi
|
+-- Elliptio_complanata
Sonowal et al., 2021 [Publ 4221], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
4 |
0 |
2 |
14 |
1 |
1 |
23.0 |
+----------- Aculamprotula_tientsinensis
--+
| +-------- Unio_pictorum **
+--+
| +----- Anodonta_cygnea
+--+
| +-- Unio_ravoisieri **
+--+
+-- Lamellidens_corrianus
Chen et al., 2023 [Publ 4302], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
38 |
15 |
9 |
-- |
+-- Inversiunio_yanagawensis
+-----+
| +-- Inversiunio_reinianus
--+
| +----- Tchangsinaia_piscicula
+--+
| +-- Unio_pictorum **
+--+
+-- Unio_crassus **
Dai et al., 2024 [Publ 4358], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
50 |
15 |
6 |
-- |
+-- Aculamprotula_scripta
+-----------------------+
| +-- Aculamprotula_tientsinensis
|
| +-- Unio_crassus **
--+ +--------------------+
| | +-- Unio_delphinus **
| |
| | +-------- Pseudocuneopsis_capitata
+--+ +-----------+
| | | +----- Pseudocuneopsis_yangshuoensis
| | +--+
| | | +-- Pseudocuneopsis_yemaoi
| | +--+
+--+ +-- Pseudocuneopsis_sichuanensis
|
| +-- Nodularia_breviconcha
| +--------------+
| | +-- Nodularia_douglasiae
| |
+--+ +-- Schistodesmus_lampreyanus
| +-----------+
| | +-- Schistodesmus_spinosus
+--+
| +----------- Tchangsinaia_piscicula
+--+
| +-------- Cuneopsis_rufescens
+--+
| +----- Cuneopsis_celtiformis
+--+
| +-- Cuneopsis_heudei
+--+
+-- Cuneopsis_demangei
Nagel et al., 1998 [Publ 1946], fig. 9
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
5 |
0 |
0 |
-- |
-- Unio
Araujo et al., 2009 [Publ 2395], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
7 |
0 |
0 |
-- |
-- Unio
Graf, 2000 [Publ 2647], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
17 |
0 |
0 |
-- |
-- Unio
Graf & Ó Foighil, 2000 [Publ 2649], fig. 1, 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
11 |
0 |
0 |
-- |
-- Unio_pictorum
Chong et al., 2008 [Publ 2668], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
20 |
0 |
0 |
-- |
-- Unio_tumidus
Taylor et al., 2007 [Publ 2696], fig. 1-3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
3 |
0 |
0 |
-- |
-- Unio_pictorum
Huff et al., 2004 [Publ 2697], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
13 |
0 |
0 |
-- |
-- Unio_ravoisieri
Huff et al., 2004 [Publ 2697], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
10 |
0 |
0 |
-- |
-- Unio_ravoisieri
Huff et al., 2004 [Publ 2697], fig. 5-6
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
11 |
0 |
0 |
-- |
-- Unio_ravoisieri
Rosenberg et al., 1994 [Publ 2719], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
18 |
0 |
0 |
-- |
-- Unio_pictorum
Rosenberg et al., 1997 [Publ 2720], fig. 2A
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
22 |
0 |
0 |
-- |
-- Unio_pictorum
Rosenberg et al., 1997 [Publ 2720], fig. 2B
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
22 |
1 |
1 |
-- |
-- Unio_pictorum
Pfeiffer & Graf, 2013 [Publ 3531], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
34 |
6 |
2 |
-- |
+-- Unio_pictorum **
+--------+
| +-- Nodularia_douglasiae
--+
| +-------- Aculamprotula_tortuosa
| |
+--+ +----- Aculamprotula_scripta
| |
+--+ +-- Aculamprotula_tientsinensis
| |
+--+-- Aculamprotula_fibrosa
|
+-- Aculamprotula_zonata
Graf et al., 2014 [Publ 3591], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
26 |
0 |
0 |
-- |
-- Unio_pictorum
Marshall et al., 2014 [Publ 3593], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
13 |
0 |
0 |
-- |
-- Unio_pictorum
Graf et al., 2015 [Publ 3619], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
19 |
0 |
0 |
-- |
-- Unio_pictorum
Pfeiffer & Graf, 2015 [Publ 3620], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
39 |
0 |
0 |
-- |
-- Unio_pictorum
Bieler et al., 2014 [Publ 3622], fig. 26, 27, 29, 30
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
4 |
0 |
0 |
-- |
-- Unio_pictorum
Bieler et al., 2014 [Publ 3622], fig. 28
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
4 |
0 |
0 |
-- |
-- Unio_pictorum
Huang et al., 2013 [Publ 3629], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
15 |
0 |
0 |
-- |
-- Unio_pictorum
González & Giribet, 2015 [Publ 3632], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
5 |
0 |
0 |
-- |
-- Unio_pictorum
González & Giribet, 2015 [Publ 3632], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
5 |
0 |
0 |
-- |
-- Unio_pictorum
González & Giribet, 2015 [Publ 3632], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
5 |
0 |
0 |
-- |
-- Unio_pictorum
González & Giribet, 2015 [Publ 3632], fig. 6
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
5 |
0 |
0 |
-- |
-- Unio_pictorum
Bolotov et al., 2015 [Publ 3646], fig. 2A
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
9 |
0 |
0 |
-- |
-- Unio_pictorum
Bolotov et al., 2015 [Publ 3646], fig. 2B
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
8 |
0 |
0 |
-- |
-- Unio_pictorum
Xue et al., 2016 [Publ 3659], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
15 |
3 |
3 |
-- |
+----- Schistodesmus_lampreyanus
--+
| +-- Nodularia_douglasiae
+--+
+-- Unio_pictorum **
Giribet & Distel, 2003 [Publ 3670], fig. 3.5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
5 |
0 |
0 |
-- |
-- Unio_ravoisieri
Zhou et al., 2016 [Publ 3710], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
19 |
1 |
1 |
-- |
+-- Schistodesmus_lampreyanus
--+
+-- Unio_pictorum **
Wu et al., 2016 [Publ 3711], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
20 |
4 |
2 |
-- |
+-- Unio_pictorum **
+-----+
| +-- Nodularia_douglasiae
--+
| +----- Aculamprotula_tortuosa
+--+
| +-- Aculamprotula_coreana
+--+
+-- Aculamprotula_tientsinensis
Wang et al., 2016 [Publ 3714], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
2 |
0 |
0 |
-- |
-- Unio_tumidus
Soroka & Burzynski, 2018 [Publ 3762], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
6 |
1 |
1 |
-- |
+-- Unio_tumidus **
--+
+-- Aculamprotula_tortuosa
Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
6 |
1 |
1 |
-- |
+-- Unio_tumidus **
--+
+-- Aculamprotula_tortuosa
Guerra et al., 2017 [Publ 3772], fig. 2a
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
24 |
1 |
1 |
-- |
+-- Aculamprotula_tortuosa
--+
+-- Unio_pictorum **
Guerra et al., 2017 [Publ 3772], fig. 2b
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
24 |
1 |
1 |
-- |
-- Unio_pictorum
Fonseca et al., 2016 [Publ 3775], fig. 1-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
8 |
0 |
0 |
-- |
-- Unio_delphinus
Lopes-Lima et al., 2017 [Publ 3792], fig. 2-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
9 |
0 |
0 |
-- |
-- Unio_delphinus
Wen et al., 2017 [Publ 3795], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
15 |
2 |
1 |
-- |
-- Unio_pictorum
Araujo et al., 2017 [Publ 3801], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
13 |
2 |
2 |
-- |
-- Unio_delphinus
Combosch et al., 2017 [Publ 3806], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
14 |
0 |
0 |
-- |
-- Unio_pictorum
Combosch et al., 2017 [Publ 3806], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
14 |
0 |
0 |
-- |
-- Unio_pictorum
Vikhrev et al., 2018 [Publ 3812], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
12 |
0 |
0 |
-- |
-- Unio_ravoisieri
Huang et al., 2018 [Publ 3828], fig. 3, 5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
28 |
1 |
1 |
-- |
+-- Unio_pictorum **
--+
+-- Aculamprotula_tientsinensis
Lopes-Lima et al., 2018 [Publ 3829], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
27 |
0 |
0 |
-- |
-- Unio_pictorum
Lopes-Lima et al., 2018 [Publ 3829], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
27 |
0 |
0 |
-- |
-- Unio_pictorum
Pfeiffer et al., 2019 [Publ 3937], fig. 4-5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
56 |
2 |
2 |
-- |
+----- Unio_gibbus **
--+
| +-- Nodularia_douglasiae
+--+
+-- Tchangsinaia_piscicula
Pfeiffer et al., 2019 [Publ 3937], fig. 6
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
56 |
2 |
2 |
-- |
+----- Unio_gibbus **
--+
| +-- Tchangsinaia_piscicula
+--+
+-- Nodularia_douglasiae
Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
9 |
0 |
0 |
-- |
-- Unio_delphinus
Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
5 |
0 |
0 |
-- |
-- Unio_delphinus
Klunzinger et al., 2021 [Publ 4086], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
5 |
0 |
0 |
-- |
-- Unio_pictorum
Wu et al., 2020 [Publ 4121], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
33 |
4 |
1 |
-- |
+----------- Unio_pictorum **
--+
| +-------- Aculamprotula_tortuosa
+--+
| +----- Aculamprotula_polysticta
+--+
| +-- Aculamprotula_fibrosa
+--+
+-- Aculamprotula_tientsinensis
Wu et al., 2020 [Publ 4121], fig. S1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
33 |
4 |
1 |
-- |
+----------- Unio_pictorum **
--+
| +-------- Aculamprotula_tortuosa
+--+
| +----- Aculamprotula_polysticta
+--+
| +-- Aculamprotula_tientsinensis
+--+
+-- Aculamprotula_fibrosa
Wu et al., 2018 [Publ 4143], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
21 |
5 |
2 |
-- |
+-- Unio_pictorum **
+--------+
| +-- Nodularia_douglasiae
--+
| +-------- Aculamprotula_tortuosa
+--+
| +----- Aculamprotula_scripta
+--+
| +-- Aculamprotula_coreana
+--+
+-- Aculamprotula_tientsinensis
Wu et al., 2020 [Publ 4147], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
29 |
9 |
6 |
-- |
+-------- Aculamprotula_tortuosa
+--------+
| | +----- Aculamprotula_scripta
| +--+
| | +-- Aculamprotula_tientsinensis
--+ +--+
| +-- Aculamprotula_coreana
|
| +-------------- Unio_pictorum **
+--+
| +----------- Pseudocuneopsis_capitata
+--+
| +-------- Nodularia_douglasiae
+--+
| +----- Schistodesmus_lampreyanus
+--+
| +-- Cuneopsis_heudei
+--+
+-- Tchangsinaia_piscicula
Wu et al., 2020 [Publ 4147], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
13 |
4 |
1 |
-- |
+----------- Unio_pictorum **
--+
| +-------- Aculamprotula_tortuosa
+--+
| +----- Aculamprotula_scripta
+--+
| +-- Aculamprotula_tientsinensis
+--+
+-- Aculamprotula_fibrosa
Choi et al., 2020 [Publ 4169], fig. 2a
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
10 |
10 |
4 |
-- |
+-- Unio_pictorum **
+--+
| +-- Tchangsinaia_piscicula
--+
| +-- Cuneopsis_heudei
+--+
+-- Cuneopsis_rufescens
Choi et al., 2020 [Publ 4169], fig. 3a
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
4 |
4 |
2 |
-- |
+-- Cuneopsis_heudei
+-----+
| +-- Unio_pictorum **
--+
| +----- Nodularia_breviconcha
+--+
| +-- Nodularia_douglasiae
+--+
+-- Nodularia_nipponensis
Pavan-Kumar et al., 2022 [Publ 4263], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
35 |
5 |
4 |
-- |
+-- Aculamprotula_tientsinensis
+--------+
| +-- Aculamprotula_tortuosa
--+
| +-------- Unio_pictorum **
+--+
| +----- Schistodesmus_lampreyanus
+--+
| +-- Nodularia_douglasiae
+--+
+-- Tchangsinaia_piscicula
Pavan-Kumar et al., 2022 [Publ 4263], fig. S5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
38 |
5 |
4 |
-- |
+-------- Unio_pictorum **
--+
| +----- Schistodesmus_lampreyanus
+--+
| +-- Nodularia_douglasiae
+--+
+-- Tchangsinaia_piscicula
Dai et al., 2023 [Publ 4285], fig. 5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
52 |
6 |
6 |
-- |
+-------------- Aculamprotula_tientsinensis
|
--+ +----------- Unio_crassus **
| |
+--+ +-- Pseudocuneopsis_capitata
| +-----+
| | +-- Nodularia_douglasiae
+--+
| +----- Schistodesmus_lampreyanus
+--+
| +-- Cuneopsis_celtiformis
+--+
+-- Tchangsinaia_piscicula
Wu et al., 2023 [Publ 4296], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
22 |
12 |
7 |
-- |
+----------------------- Unio_tumidus **
|
| +-- Aculamprotula_scripta
--+ +-----------------+
| | +-- Aculamprotula_tientsinensis
| |
+--+ +-- Cuneopsis_heudei
| +--------------+
| | +-- Cuneopsis_rufescens
| |
+--+ +----- Nodularia_douglasiae
| +--------+
| | | +-- Schistodesmus_lampreyanus
| | +--+
+--+ +-- Schistodesmus_spinosus
|
| +----------- Tchangsinaia_piscicula
+--+
| +-------- Inversiunio_yanagawensis
+--+
| +----- Pseudocuneopsis_capitata
+--+
| +-- Pseudocuneopsis_yangshuoensis
+--+
+-- Pseudocuneopsis_sichuanensis
Liu et al., 2023 [Publ 4310], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
23 |
13 |
7 |
-- |
+-------------------------- Unio_tumidus **
|
| +-- Aculamprotula_scripta
--+ +--------------------+
| | +-- Aculamprotula_tientsinensis
| |
+--+ +-- Cuneopsis_heudei
| +-----------------+
| | +-- Cuneopsis_rufescens
| |
+--+ +----- Nodularia_douglasiae
| +-----------+
| | | +-- Schistodesmus_lampreyanus
| | +--+
+--+ +-- Schistodesmus_spinosus
|
| +-------------- Tchangsinaia_piscicula
+--+
| +----------- Inversiunio_yanagawensis
+--+
| +-------- Pseudocuneopsis_capitata
+--+
| +----- Pseudocuneopsis_sichuanensis
+--+
| +-- Pseudocuneopsis_wuana
+--+
+-- Pseudocuneopsis_yangshuoensis
Liu et al., 2023 [Publ 4310], fig. 5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
23 |
13 |
7 |
-- |
+----------------- Unio_tumidus **
|
| +-- Aculamprotula_tientsinensis
+--------------+
| +-- Aculamprotula_scripta
|
--+ +-- Cuneopsis_rufescens
| +-----------+
| | +-- Cuneopsis_heudei
| |
| | +----- Nodularia_douglasiae
| +--------+
+--+ | +-- Schistodesmus_lampreyanus
| +--+
| +-- Schistodesmus_spinosus
|
| +----------- Tchangsinaia_piscicula
| |
+--+----------- Inversiunio_yanagawensis
|
| +-------- Pseudocuneopsis_capitata
+--+
| +----- Pseudocuneopsis_sichuanensis
+--+
| +-- Pseudocuneopsis_wuana
+--+
+-- Pseudocuneopsis_yangshuoensis
|