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Page last updated
Wed 06 Nov 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Unioninae | Unionini

genus Unio Philipsson in Retzius, 1788: cladomics
     45 valid species

Cladomics Summary

number of species = 45

analyzed in 211 cladograms in 123 publications
maximum number of terminals in an analysis = 23, mean = 3.24

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 144
maximum number of included terminals = 23, mean = 4.28
proportion of cladograms that support basic monophyly = 0.93


Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ tree


Lyubas et al., 2022 [Publ 4255], fig. 7

iT tT bi riS oT roT roS iD
23 23 20 23 8 0 0 0.0

*** Top Monophyly Cladogram ***

  +----------------------------- Unio_tumidus **      
  |                                                
  |                          +-- Unio_durieui **      
  |  +-----------------------+                     
  |  |                       +-- Unio_gibbus **       
--+  |                                             
  |  |           +-------------- Unio_damascensis **  
  |  |           |                                 
  |  |           +-------------- Unio_sesirmensis **  
  |  |  +--------+                                 
  +--+  |        |  +----------- Unio_ionicus **      
     |  |        |  |                              
     |  |        +--+        +-- Unio_bruguierianus **
     |  |           |  +-----+                     
     |  |           |  |     +-- Unio_tumidiformis ** 
     |  |           +--+                           
     |  |              |  +----- Unio_vicarius **     
     +--+              +--+                        
        |                 |  +-- Unio_carneus **      
        |                 +--+                     
        |                    +-- Unio_crassus **      
        |                                          
        |                 +----- Unio_caffer **       
        |  +--------------+                        
        |  |              |  +-- Unio_terminalis **   
        |  |              +--+                     
        +--+                 +-- Unio_delicatus **    
           |                                       
           |  +----------------- Unio_pictorum **     
           |  |                                    
           +--+  +-------------- Unio_eucirrus **     
              |  |                                 
              +--+           +-- Unio_elongatulus **  
                 |  +--------+                     
                 |  |        +-- Unio_mancus **       
                 +--+                              
                    |  +-------- Unio_ravoisieri **   
                    |  |                           
                    +--+     +-- Unio_delphinus **    
                       |  +--+                     
                       |  |  +-- Unio_foucauldianus **
                       +--+                        
                          |  +-- Unio_hueti **        
                          +--+                     
                             +-- Unio_tigridis **     

Chen et al., 2023 [Publ 4302], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 38 15 9 --

*** Top Sister-Group Cladogram ***

  +-------------------- Aculamprotula   
  |                                     
  |                 +-- Middendorffinaia
--+  +--------------+                   
  |  |              +-- Nodularia       
  |  |                                  
  +--+  +-------------- Cuneopsis       
     |  |                               
     |  |           +-- Diaurora        
     +--+-----------+                   
        |           +-- Schistodesmus   
        |                               
        |  +----------- Pseudocuneopsis 
        +--+                            
           |  +-------- Inversiunio     
           +--+                         
              |  +----- Tchangsinaia    
              +--+                      
                 |  +-- Unio_pictorum **   
                 +--+                   
                    +-- Unio_nanus **      

Lyubas et al., 2022 [Publ 4255], fig. S1

iT tT bi riS oT roT roS iD
23 23 14 23 8 0 0 10.0
  +----------------- Unio_tumidus **      
  |                                    
  |              +-- Unio_durieui **      
  +--------------+                     
  |              +-- Unio_gibbus **       
  |                                    
  |     +----------- Unio_sesirmensis **  
  |     |                              
  |  +--+  +-------- Unio_damascensis **  
--+  |  |  |                           
  |  |  |  |  +----- Unio_ionicus **      
  |  |  +--+  |                        
  |  |     |  |  +-- Unio_bruguierianus **
  |  |     |  +--+                     
  |  |     +--+  +-- Unio_tumidiformis ** 
  |  |        |                        
  |  |        |  +-- Unio_carneus **      
  |  |        |  |                     
  |  |        +--+-- Unio_crassus **      
  +--+           |                     
     |           +-- Unio_vicarius **     
     |                                 
     |     +-------- Unio_elongatulus **  
     |     |                           
     |     +-------- Unio_mancus **       
     |     |                           
     |     |     +-- Unio_pictorum **     
     |     +-----+                     
     |     |     +-- Unio_eucirrus **     
     |     |                           
     +-----+  +----- Unio_caffer **       
           +--+                        
           |  |  +-- Unio_terminalis **   
           |  +--+                     
           |     +-- Unio_delicatus **    
           |                           
           |  +----- Unio_ravoisieri **   
           |  |                        
           |  |  +-- Unio_delphinus **    
           +--+--+                     
              |  +-- Unio_foucauldianus **
              |                        
              |  +-- Unio_hueti **        
              +--+                     
                 +-- Unio_tigridis **     

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_AHE

iT tT bi riS oT roT roS iD
17 17 15 17 5 2 2 27.0
                             +-- Tchangsinaia      
  +--------------------------+                     
  |                          +-- Nodularia         
--+                                                
  |  +-------------------------- Unio_gibbus **       
  |  |                                             
  +--+  +----------------------- Unio_tumidus **      
     |  |                                          
     |  |                 +----- Unio_terminalis **   
     +--+  +--------------+                        
        |  |              |  +-- Unio_delphinus **    
        |  |              +--+                     
        |  |                 +-- Unio_pictorum **     
        +--+                                       
           |  +----------------- Unio_tumidiformis ** 
           |  |                                    
           |  |              +-- Unio_ionicus **      
           +--+  +-----------+                     
              |  |           +-- Unio_carneus **      
              |  |                                 
              |  |           +-- Unio_damascensis **  
              +--+  +--------+                     
                 |  |        +-- Unio_sesirmensis **  
                 |  |                              
                 |  |     +----- Unio_bruguierianus **
                 +--+  +--+                        
                    |  |  |  +-- Unio_gontierii **    
                    |  |  +--+                     
                    |  |     +-- Unio_mardinensis **  
                    +--+                           
                       |     +-- Unio_crassus **      
                       |  +--+                     
                       |  |  +-- Unio_nanus **        
                       +--+                        
                          |  +-- Unio_vicarius **     
                          +--+                     
                             +-- Unio_desectus **     

Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_MtG

iT tT bi riS oT roT roS iD
17 17 15 17 5 2 2 27.0
                                   +-- Tchangsinaia      
  +--------------------------------+                     
  |                                +-- Nodularia         
  |                                                      
--+                                +-- Unio_gibbus **       
  |  +-----------------------------+                     
  |  |                             +-- Unio_tumidus **      
  |  |                                                   
  +--+                          +----- Unio_terminalis **   
     |  +-----------------------+                        
     |  |                       |  +-- Unio_delphinus **    
     |  |                       +--+                     
     +--+                          +-- Unio_pictorum **     
        |                                                
        |  +-------------------------- Unio_tumidiformis ** 
        |  |                                             
        +--+  +----------------------- Unio_sesirmensis **  
           |  |                                          
           +--+  +-------------------- Unio_ionicus **      
              |  |                                       
              +--+  +----------------- Unio_damascensis **  
                 |  |                                    
                 +--+  +-------------- Unio_desectus **     
                    |  |                                 
                    |  |           +-- Unio_crassus **      
                    +--+  +--------+                     
                       |  |        +-- Unio_gontierii **    
                       |  |                              
                       +--+        +-- Unio_bruguierianus **
                          |  +-----+                     
                          |  |     +-- Unio_mardinensis **  
                          +--+                           
                             |  +----- Unio_vicarius **     
                             +--+                        
                                |  +-- Unio_carneus **      
                                +--+                     
                                   +-- Unio_nanus **        

Lopes-Lima et al., 2021 [Publ 4217], fig. 1

iT tT bi riS oT roT roS iD
16 16 14 16 9 0 0 18.0
  +----------------------- Unio_tumidus **      
  |                                          
  |              +-------- Unio_damascensis **  
  |  +-----------+                           
--+  |           |  +----- Unio_sesirmensis **  
  |  |           +--+                        
  |  |              |  +-- Unio_bruguierianus **
  |  |              +--+                     
  +--+                 +-- Unio_nanus **        
     |                                       
     |                 +-- Unio_delicatus **    
     |  +--------------+                     
     |  |              +-- Unio_terminalis **   
     |  |                                    
     +--+              +-- Unio_tigridis **     
        |  +-----------+                     
        |  |           +-- Unio_hueti **        
        |  |                                 
        +--+  +----------- Unio_ravoisieri **   
           |  |                              
           |  |        +-- Unio_pictorum **     
           +--+  +-----+                     
              |  |     +-- Unio_eucirrus **     
              |  |                           
              +--+     +-- Unio_elongatulus **  
                 |  +--+                     
                 |  |  +-- Unio_mancus **       
                 +--+                        
                    |  +-- Unio_foucauldianus **
                    +--+                     
                       +-- Unio_delphinus **    

Lopes-Lima et al., 2024 [Publ 4376], fig. 2

iT tT bi riS oT roT roS iD
14 14 12 18 0 0 0 29.0
                             +-- Unio_pictorum **        
  +--------------------------+                        
  |                          +-- Unio_delphinus **       
--+                                                   
  |  +-------------------------- Unio_tumidiformis **    
  |  |                                                
  +--+  +----------------------- Unio_sesirmensis **     
     |  |                                             
     +--+  +-------------------- Unio_desectus **        
        |  |                                          
        +--+  +----------------- Unio_damascensis **     
           |  |                                       
           |  |  +-------------- Unio_ionicus **         
           +--+  |                                    
              |  |           +-- Unio_nanus **           
              |  |  +--------+                        
              +--+  |        +-- Unio_carneus **         
                 |  |                                 
                 |  |        +-- Unio_gontierii **       
                 |  |        |                        
                 |  |        +-- Unio_crassus__1 **      
                 +--+--------+                        
                    |        +-- Unio_crassus__2 **      
                    |        |                        
                    |        +-- Unio_crassus__3 **      
                    |                                 
                    |  +-------- Unio_vicarius **        
                    |  |                              
                    +--+  +----- Unio_bruguierianus__1 **
                       |  |                           
                       +--+----- Unio_bruguierianus__2 **
                          |                           
                          |  +-- Unio_bruguierianus__3 **
                          +--+                        
                             +-- Unio_mardinensis **     

Araujo et al., 2018 [Publ 3779], fig. 1

iT tT bi riS oT roT roS iD
13 13 9 14 5 1 1 20.0
  +-------------------- Nodularia         
  |                                       
  |                 +-- Unio_durieui **      
--+  +--------------+                     
  |  |              +-- Unio_gibbus **       
  |  |                                    
  +--+  +-------------- Unio_tumidus **      
     |  |                                 
     |  |     +-------- Unio_tumidiformis ** 
     |  |  +--+                           
     +--+  |  |  +----- Unio_desectus **     
        |  |  +--+                        
        |  |     |  +-- Unio__1 **           
        |  |     +--+                     
        +--+        +-- Unio_ionicus **      
           |                              
           |        +-- Unio_delphinus **    
           |     +--+                     
           |     |  +-- Unio_foucauldianus **
           |     |                        
           +-----+  +-- Unio_pictorum **     
                 |  |                     
                 |  +-- Unio_elongatulus **  
                 |  |                     
                 +--+-- Unio_mancus **       
                    |                     
                    +-- Unio_ravoisieri **   
                    |                     
                    +-- Unio__2 **           

Froufe et al., 2016 [Publ 3738], fig. 3

iT tT bi riS oT roT roS iD
12 12 10 12 2 2 1 25.0
  +----------------------------- Nodularia         
  |                                                
--+                          +-- Unio_gibbus **       
  |  +-----------------------+                     
  |  |                       +-- Unio_durieui **      
  +--+                                             
     |  +----------------------- Unio_tumidus **      
     |  |                                          
     +--+                    +-- Unio **              
        |  +-----------------+                     
        |  |                 +-- Unio_tumidiformis ** 
        +--+                                       
           |  +----------------- Unio_delphinus **    
           +--+                                    
              |  +-------------- Unio_foucauldianus **
              +--+                                 
                 |  +----------- Unio_ravoisieri **   
                 +--+                              
                    |  +-------- Unio_mancus **       
                    +--+                           
                       |  +----- Unio_elongatulus **  
                       +--+                        
                          |  +-- Unio_pictorum **     
                          +--+                     
                             +-- Unio_caffer **       

Prié & Puillandre, 2014 [Publ 3529], fig. 2

iT tT bi riS oT roT roS iD
11 11 9 11 6 0 0 22.0
              +----- Unio_tumidus **     
  +-----------+                       
  |           |  +-- Unio_gibbus **      
  |           +--+                    
  |              +-- Unio_durieui **     
--+                                   
  |           +----- Unio_tumidiformis **
  |  +--------+                       
  |  |        |  +-- Unio **             
  |  |        +--+                    
  +--+           +-- Unio_crassus **     
     |                                
     |  +----------- Unio_delphinus **   
     +--+                             
        |  +-------- Unio_elongatulus ** 
        +--+                          
           |  +----- Unio_mancus **      
           +--+                       
              |  +-- Unio_pictorum **    
              +--+                    
                 +-- Unio_ravoisieri **  

Lopes-Lima et al., 2021 [Publ 4217], fig. 2

iT tT bi riS oT roT roS iD
10 10 8 10 0 0 0 17.0
  +----------------- Unio_tumidus **      
  |                                    
--+  +-------------- Unio_bruguierianus **
  |  |                                 
  |  |           +-- Unio_delicatus **    
  +--+  +--------+                     
     |  |        +-- Unio_terminalis **   
     |  |                              
     +--+        +-- Unio_tigridis **     
        |  +-----+                     
        |  |     +-- Unio_hueti **        
        |  |                           
        +--+     +-- Unio_pictorum **     
           |  +--+                     
           |  |  +-- Unio_eucirrus **     
           +--+                        
              |  +-- Unio_elongatulus **  
              +--+                     
                 +-- Unio_mancus **       

Lopes-Lima et al., 2017 [Publ 3734], fig. 1

iT tT bi riS oT roT roS iD
9 9 5 9 8 0 0 22.0
           +-- Unio_tumidus **     
  +--------+                    
  |        +-- Unio_gibbus **      
  |                             
--+        +-- Unio **             
  |  +-----+                    
  |  |     +-- Unio_tumidiformis **
  +--+                          
     |  +----- Unio_ravoisieri **  
     |  |                       
     +--+  +-- Unio_mancus **      
        |  |                    
        |  +-- Unio_pictorum **    
        +--+                    
           +-- Unio_delphinus **   
           |                    
           +-- Unio_elongatulus ** 

Khalloufi et al., 2011 [Publ 2748], fig. 2

iT tT bi riS oT roT roS iD
9 9 6 9 2 0 0 18.0
        +----- Unio_tumidus **     
  +-----+                       
  |     |  +-- Unio_gibbus **      
  |     +--+                    
  |        +-- Unio_durieui **     
--+                             
  |        +-- Unio_tumidiformis **
  |  +-----+                    
  |  |     +-- Unio_crassus **     
  +--+                          
     |  +----- Unio **             
     |  |                       
     +--+----- Unio_pictorum **    
        |                       
        |  +-- Unio_delphinus **   
        +--+                    
           +-- Unio_mancus **      

Marrone et al., 2019 [Publ 3907], fig. 1

iT tT bi riS oT roT roS iD
8 8 4 8 0 0 0 23.0
  +-------------- Unio_tumidus **      
  |                                 
--+  +----------- Unio_delphinus **    
  |  |                              
  +--+  +-------- Unio_foucauldianus **
     |  |                           
     +--+  +----- Unio_mancus **       
        |  |                        
        |  +----- Unio_elongatulus **  
        +--+                        
           +----- Unio_ravoisieri **   
           |                        
           |  +-- Unio_pictorum **     
           +--+                     
              +-- Unio_terminalis **   

Reis et al., 2013 [Publ 3580], fig. 5

iT tT bi riS oT roT roS iD
7 7 6 8 8 0 0 20.0
  +-------------- Unio_tumidus **     
  |                                
--+           +-- Unio_tumidiformis **
  |  +--------+                    
  |  |        +-- Unio__2 **          
  +--+                             
     |  +-------- Unio__1 **          
     |  |                          
     +--+     +-- Unio_delphinus **   
        |  +--+                    
        |  |  +-- Unio_mancus **      
        +--+                       
           |  +-- Unio_pictorum **    
           +--+                    
              +-- Unio_caffer **      

Lopes-Lima et al., 2021 [Publ 4217], fig. 6

iT tT bi riS oT roT roS iD
7 7 5 7 0 0 0 21.0
  +----------------- Unio_tumidus **    
--+                                  
  |  +-------------- Unio_pictorum **   
  +--+                               
     |  +----------- Unio_sesirmensis **
     +--+                            
        |  +-------- Unio_damascensis **
        +--+                         
           |  +----- Unio_crassus **    
           +--+                      
              |  +-- Unio **            
              +--+                   
                 +-- Unio_nanus **      

Araujo et al., 2009 [Publ 2589], fig. 7B

iT tT bi riS oT roT roS iD
7 7 6 8 4 0 0 19.0
  +-------------- Unio_gibbus **      
  |                                
--+  +----------- Unio_tumidus **     
  |  |                             
  |  |        +-- Unio_tumidiformis **
  +--+  +-----+                    
     |  |     +-- Unio_crassus **     
     |  |                          
     +--+     +-- Unio_pictorum__1 ** 
        |  +--+                    
        |  |  +-- Unio_mancus **      
        +--+                       
           |  +-- Unio_pictorum__2 ** 
           +--+                    
              +-- Unio_caffer **      

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iT tT bi riS oT roT roS iD
6 6 4 6 90 16 6 23.0
  +-------------------- Aculamprotula   
  |                                     
  |                 +-- Nodularia       
  |           +-----+                   
--+           |     +-- Pseudocuneopsis 
  |  +--------+                         
  |  |        |  +----- Tchangsinaia    
  |  |        +--+                      
  |  |           |  +-- Schistodesmus   
  +--+           +--+                   
     |              +-- Cuneopsis       
     |                                  
     |  +-------------- Unio_tumidus **    
     +--+                               
        |  +----------- Unio_nanus **      
        +--+                            
           |  +-------- Unio_elongatulus **
           +--+                         
              |  +----- Unio_pictorum **   
              +--+                      
                 |  +-- Unio_delphinus **  
                 +--+                   
                    +-- Unio_mancus **     

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iT tT bi riS oT roT roS iD
6 6 3 6 90 16 6 22.0
  +-------------------- Aculamprotula               
  |                                                 
  |        +----------- Unio_tumidus **                
  |        |                                        
  |  +-----+  +-------- Unio_nanus **                  
--+  |     |  |                                     
  |  |     +--+  +----- Unio_elongatulus **            
  |  |        |  |                                  
  |  |        +--+  +-- Unio_delphinus **              
  |  |           |  |                               
  +--+           +--+-- Unio_mancus **                 
     |              |                               
     |              +-- Unio_pictorum **               
     |                                              
     |  +-------------- Pseudocuneopsis             
     |  |                                           
     +--+  +----------- Nodularia                   
        |  |                                        
        +--+        +-- Schistodesmus_lampreyanus__1
           |  +-----+                               
           |  |     +-- Cuneopsis                   
           +--+                                     
              |  +----- Tchangsinaia                
              +--+                                  
                 |  +-- Schistodesmus_lampreyanus__2
                 +--+                               
                    +-- Schistodesmus_spinosus      

Dai et al., 2024 [Publ 4386], fig. 6

iT tT bi riS oT roT roS iD
6 6 4 6 34 14 6 21.0
  +-------------------- Aculamprotula   
  |                                     
  |        +----------- Pseudocuneopsis 
  |  +-----+                            
--+  |     |  +-------- Nodularia       
  |  |     +--+                         
  |  |        |  +----- Schistodesmus   
  |  |        +--+                      
  +--+           |  +-- Tchangsinaia    
     |           +--+                   
     |              +-- Cuneopsis       
     |                                  
     |  +-------------- Unio_tumidus **    
     +--+                               
        |  +----------- Unio_nanus **      
        +--+                            
           |  +-------- Unio_delphinus **  
           +--+                         
              |  +----- Unio_pictorum **   
              +--+                      
                 |  +-- Unio_elongatulus **
                 +--+                   
                    +-- Unio_mancus **     

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iT tT bi riS oT roT roS iD
6 6 4 6 86 16 6 23.0
  +-------------------- Aculamprotula   
  |                                     
  |           +-------- Nodularia       
  |           |                         
--+  +--------+-------- Pseudocuneopsis 
  |  |        |                         
  |  |        |  +----- Schistodesmus   
  |  |        +--+                      
  +--+           |  +-- Tchangsinaia    
     |           +--+                   
     |              +-- Cuneopsis       
     |                                  
     |  +-------------- Unio_tumidus **    
     +--+                               
        |  +----------- Unio_nanus **      
        +--+                            
           |  +-------- Unio_elongatulus **
           +--+                         
              |  +----- Unio_pictorum **   
              +--+                      
                 |  +-- Unio_delphinus **  
                 +--+                   
                    +-- Unio_mancus **     

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iT tT bi riS oT roT roS iD
6 6 4 6 86 16 6 23.0
  +-------------------- Aculamprotula   
  |                                     
  |           +-------- Nodularia       
  |           |                         
--+  +--------+-------- Pseudocuneopsis 
  |  |        |                         
  |  |        |  +----- Schistodesmus   
  |  |        +--+                      
  +--+           |  +-- Tchangsinaia    
     |           +--+                   
     |              +-- Cuneopsis       
     |                                  
     |  +-------------- Unio_tumidus **    
     +--+                               
        |  +----------- Unio_nanus **      
        +--+                            
           |  +-------- Unio_elongatulus **
           +--+                         
              |  +----- Unio_pictorum **   
              +--+                      
                 |  +-- Unio_delphinus **  
                 +--+                   
                    +-- Unio_mancus **     

Reis et al., 2013 [Publ 3580], fig. 4

iT tT bi riS oT roT roS iD
6 6 4 6 8 0 0 19.0
  +----------- Unio_tumidus **     
  |                             
--+        +-- Unio_tumidiformis **
  |  +-----+                    
  |  |     +-- Unio_crassus **     
  +--+                          
     |  +----- Unio_delphinus **   
     +--+                       
        |  +-- Unio_mancus **      
        +--+                    
           +-- Unio_pictorum **    

Araujo et al., 2009 [Publ 2589], fig. 7A

iT tT bi riS oT roT roS iD
6 7 3 7 5 1 1 22.0
  +----------- Nodularia        
  |                             
  |        +-- Unio_gibbus **      
--+        |                    
  |  +-----+-- Unio_tumidus **     
  |  |     |                    
  |  |     +-- Anodonta         
  +--+                          
     |  +----- Unio_pictorum__1 ** 
     |  |                       
     |  +----- Unio_pictorum__2 ** 
     +--+                       
        +----- Unio_mancus **      
        |                       
        |  +-- Unio_tumidiformis **
        +--+                    
           +-- Unio_crassus **     

Araujo et al., 2009 [Publ 2589], fig. 7C

iT tT bi riS oT roT roS iD
6 6 4 7 4 0 0 20.0
        +-- Unio_gibbus **      
  +-----+                    
  |     +-- Unio_tumidus **     
  |                          
--+     +-- Unio_tumidiformis **
  |  +--+                    
  |  |  +-- Unio_crassus **     
  +--+                       
     |  +-- Unio_pictorum__1 ** 
     |  |                    
     +--+-- Unio_pictorum__2 ** 
        |                    
        +-- Unio_mancus **      

Wu et al., 2024 [Publ 4379], fig. 5

iT tT bi riS oT roT roS iD
6 6 4 6 48 23 6 21.0
  +-------------------- Aculamprotula   
  |                                     
  |        +----------- Pseudocuneopsis 
  |  +-----+                            
--+  |     |  +-------- Nodularia       
  |  |     +--+                         
  |  |        |  +----- Schistodesmus   
  |  |        +--+                      
  +--+           |  +-- Tchangsinaia    
     |           +--+                   
     |              +-- Cuneopsis       
     |                                  
     |  +-------------- Unio_tumidus **    
     +--+                               
        |  +----------- Unio_nanus **      
        +--+                            
           |  +-------- Unio_delphinus **  
           +--+                         
              |  +----- Unio_pictorum **   
              +--+                      
                 |  +-- Unio_elongatulus **
                 +--+                   
                    +-- Unio_mancus **     

Bashê, 2022 [Publ 4265], fig. 1

iT tT bi riS oT roT roS iD
5 5 4 6 1 0 0 25.0
  +-------------- Unio_pictorum__2 **  
--+                                 
  |  +----------- Unio_foucauldianus **
  +--+                              
     |  +-------- Unio_delphinus **    
     +--+                           
        |  +----- Unio_tigridis **     
        +--+                        
           |  +-- Unio_elongatulus **  
           +--+                     
              +-- Unio_pictorum__1 **  

Pfarrer et al., 2022 [Publ 4305], fig. 2-5

iT tT bi riS oT roT roS iD
5 5 3 5 7 0 0 22.0
  +----------- Unio_tumidus **    
--+                            
  |  +-------- Unio_nanus **      
  +--+                         
     |  +----- Unio_elongatulus **
     +--+                      
        |  +-- Unio_mancus **     
        +--+                   
           +-- Unio_pictorum **   

Prie et al., 2012 [Publ 3487], fig. 2

iT tT bi riS oT roT roS iD
5 5 3 5 1 0 0 18.0
  +----------- Unio_tumidus **    
--+                            
  |  +-------- Unio_crassus **    
  +--+                         
     |  +----- Unio_pictorum **   
     +--+                      
        |  +-- Unio_mancus **     
        +--+                   
           +-- Unio_elongatulus **

Wu et al., 2021 [Publ 4366], fig. 4

iT tT bi riS oT roT roS iD
4 4 2 4 65 11 6 21.0
  +----------------- Aculamprotula  
  |                                 
  |        +-------- Unio_tumidus **   
--+  +-----+                        
  |  |     |  +----- Unio_nanus **     
  |  |     +--+                     
  |  |        |  +-- Unio_pictorum **  
  +--+        +--+                  
     |           +-- Unio_delphinus ** 
     |                              
     |  +----------- Pseudocuneopsis
     +--+                           
        |  +-------- Nodularia      
        +--+                        
           |  +----- Schistodesmus  
           +--+                     
              |  +-- Tchangsinaia   
              +--+                  
                 +-- Cuneopsis      

Nagel et al., 1998 [Publ 1946], fig. 8

iT tT bi riS oT roT roS iD
4 4 1 4 0 0 0 20.0
  +----- Unio_tumidus ** 
  |                   
--+  +-- Unio_pictorum **
  |  |                
  +--+-- Unio_crassus ** 
     |                
     +-- Unio_mancus **  

Huang et al., 2019 [Publ 3843], fig. 2

iT tT bi riS oT roT roS iD
4 4 2 4 194 14 5 19.0
  +-------------- Aculamprotula 
  |                             
  |     +-------- Unio_tumidus **  
--+  +--+                       
  |  |  |  +----- Unio_crassus **  
  |  |  +--+                    
  |  |     |  +-- Unio_delphinus **
  +--+     +--+                 
     |        +-- Unio_pictorum ** 
     |                          
     |  +-------- Schistodesmus 
     +--+                       
        |  +----- Nodularia     
        +--+                    
           |  +-- Tchangsinaia  
           +--+                 
              +-- Cuneopsis     

Huang et al., 2019 [Publ 3843], fig. S3

iT tT bi riS oT roT roS iD
4 4 2 4 192 13 5 19.0
  +-------------- Aculamprotula 
  |                             
  |     +-------- Unio_tumidus **  
--+  +--+                       
  |  |  |  +----- Unio_crassus **  
  |  |  +--+                    
  |  |     |  +-- Unio_delphinus **
  +--+     +--+                 
     |        +-- Unio_pictorum ** 
     |                          
     |  +-------- Nodularia     
     +--+                       
        |  +----- Schistodesmus 
        +--+                    
           |  +-- Tchangsinaia  
           +--+                 
              +-- Cuneopsis     

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iT tT bi riS oT roT roS iD
4 4 1 4 76 7 5 20.0
           +----- Unio_gibbus **  
           |                   
  +--------+----- Unio_tumidus ** 
  |        |                   
  |        |  +-- Unio_crassus ** 
--+        +--+                
  |           +-- Unio_pictorum **
  |                            
  |  +----------- Aculamprotula
  +--+                         
     |  +-------- Nodularia    
     +--+                      
        |  +----- Schistodesmus
        +--+                   
           |  +-- Tchangsinaia 
           +--+                
              +-- Cuneopsis    

Klishko et al., 2017 [Publ 3946], fig. 8

iT tT bi riS oT roT roS iD
4 4 2 4 2 2 1 21.0
  +----------- Nodularia    
--+                         
  |  +-------- Unio_tumidus ** 
  +--+                      
     |  +----- Unio_pictorum **
     +--+                   
        |  +-- Unio_nanus **   
        +--+                
           +-- Unio_crassus ** 

Huang et al., 2018 [Publ 3828], fig. 6

iT tT bi riS oT roT roS iD
4 4 2 4 39 5 3 19.0
  +-------------- Aculamprotula 
  |                             
--+           +-- Tchangsinaia  
  |  +--------+                 
  |  |        +-- Nodularia     
  +--+                          
     |  +-------- Unio_tumidus **  
     +--+                       
        |  +----- Unio_crassus **  
        +--+                    
           |  +-- Unio_delphinus **
           +--+                 
              +-- Unio_pictorum ** 

Soroka, 2010 [Publ 3404], fig. 3b

iT tT bi riS oT roT roS iD
4 4 2 4 5 0 0 21.0
  +-------- Unio_tumidus ** 
--+                      
  |  +----- Unio **         
  +--+                   
     |  +-- Unio_pictorum **
     +--+                
        +-- Unio_mancus **  

Liu et al., 2017 [Publ 3939], fig. 2

iT tT bi riS oT roT roS iD
4 4 1 4 11 7 3 20.0
        +-- Schistodesmus
        |                
  +-----+-- Cuneopsis    
  |     |                
  |     +-- Nodularia    
--+                      
  |  +----- Unio_tumidus ** 
  |  |                   
  +--+----- Unio_gibbus **  
     |                   
     |  +-- Unio_crassus ** 
     +--+                
        +-- Unio_pictorum **

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iT tT bi riS oT roT roS iD
4 4 2 4 45 6 4 21.0
  +-------------- Aculamprotula 
  |                             
  |        +----- Nodularia     
--+  +-----+                    
  |  |     |  +-- Schistodesmus 
  |  |     +--+                 
  +--+        +-- Tchangsinaia  
     |                          
     |  +-------- Unio_tumidus **  
     +--+                       
        |  +----- Unio_nanus **    
        +--+                    
           |  +-- Unio_pictorum ** 
           +--+                 
              +-- Unio_delphinus **

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

iT tT bi riS oT roT roS iD
4 4 2 4 20 6 4 21.0
  +-------------- Aculamprotula 
  |                             
  |        +----- Schistodesmus 
--+  +-----+                    
  |  |     |  +-- Tchangsinaia  
  |  |     +--+                 
  +--+        +-- Nodularia     
     |                          
     |  +-------- Unio_tumidus **  
     +--+                       
        |  +----- Unio_nanus **    
        +--+                    
           |  +-- Unio_pictorum ** 
           +--+                 
              +-- Unio_delphinus **

Cha et al., 2018 [Publ 3916], fig. 1

iT tT bi riS oT roT roS iD
4 4 2 4 35 6 4 21.0
  +-------------- Aculamprotula 
  |                             
  |        +----- Schistodesmus 
--+  +-----+                    
  |  |     |  +-- Nodularia     
  |  |     +--+                 
  +--+        +-- Tchangsinaia  
     |                          
     |  +-------- Unio_tumidus **  
     +--+                       
        |  +----- Unio_nanus **    
        +--+                    
           |  +-- Unio_pictorum ** 
           +--+                 
              +-- Unio_delphinus **

Huang et al., 2019 [Publ 3843], fig. S4

iT tT bi riS oT roT roS iD
4 4 2 4 194 14 5 19.0
  +-------- Unio_tumidus **  
--+                       
  |  +----- Unio_crassus **  
  +--+                    
     |  +-- Unio_delphinus **
     +--+                 
        +-- Unio_pictorum ** 

Burzynski & Soroka, 2018 [Publ 3862], fig. 5b

iT tT bi riS oT roT roS iD
4 4 2 4 6 1 1 21.0
  +----------- Aculamprotula 
--+                          
  |  +-------- Unio_tumidus **  
  +--+                       
     |  +----- Unio_nanus **    
     +--+                    
        |  +-- Unio_pictorum ** 
        +--+                 
           +-- Unio_delphinus **

Nagel et al., 1998 [Publ 1946], fig. 1

iT tT bi riS oT roT roS iD
4 4 2 4 5 0 0 21.0
  +-------- Unio_crassus ** 
--+                      
  |  +----- Unio_tumidus ** 
  +--+                   
     |  +-- Unio_pictorum **
     +--+                
        +-- Unio_mancus **  

Wang et al., 2018 [Publ 3865], fig. 2

iT tT bi riS oT roT roS iD
4 4 2 4 28 3 3 21.0
  +-------------- Aculamprotula 
  |                             
--+           +-- Tchangsinaia  
  |  +--------+                 
  |  |        +-- Nodularia     
  +--+                          
     |  +-------- Unio_tumidus **  
     +--+                       
        |  +----- Unio_nanus **    
        +--+                    
           |  +-- Unio_pictorum ** 
           +--+                 
              +-- Unio_delphinus **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iT tT bi riS oT roT roS iD
4 4 1 4 116 22 7 20.0
  +----------------- Aculamprotula   
  |                                  
  |  +-------------- Middendorffinaia
--+  |                               
  |  |        +----- Unio_gibbus **     
  |  |        |                      
  +--+  +-----+----- Unio_tumidus **    
     |  |     |                      
     |  |     |  +-- Unio_crassus **    
     |  |     +--+                   
     +--+        +-- Unio_pictorum **   
        |                            
        |  +-------- Nodularia       
        |  |                         
        +--+  +----- Schistodesmus   
           |  |                      
           +--+----- Cuneopsis       
              |                      
              |  +-- Tchangsinaia    
              +--+                   
                 +-- Inversiunio     

Wu et al., 2021 [Publ 4366], fig. 3-BI

iT tT bi riS oT roT roS iD
4 4 2 4 70 11 6 21.0
  +----------------- Aculamprotula  
  |                                 
  |        +-------- Unio_tumidus **   
--+  +-----+                        
  |  |     |  +----- Unio_nanus **     
  |  |     +--+                     
  |  |        |  +-- Unio_pictorum **  
  +--+        +--+                  
     |           +-- Unio_delphinus ** 
     |                              
     |  +----------- Pseudocuneopsis
     +--+                           
        |  +-------- Nodularia      
        +--+                        
           |  +----- Schistodesmus  
           +--+                     
              |  +-- Cuneopsis      
              +--+                  
                 +-- Tchangsinaia   

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iT tT bi riS oT roT roS iD
4 4 2 4 50 8 6 21.0
  +-------------- Aculamprotula  
  |                              
  |     +-------- Unio_tumidus **   
  |  +--+                        
--+  |  |  +----- Unio_nanus **     
  |  |  +--+                     
  |  |     |  +-- Unio_pictorum **  
  |  |     +--+                  
  +--+        +-- Unio_delphinus ** 
     |                           
     |        +-- Schistodesmus  
     |  +-----+                  
     |  |     +-- Cuneopsis      
     +--+                        
        |  +----- Pseudocuneopsis
        +--+                     
           |  +-- Nodularia      
           +--+                  
              +-- Tchangsinaia   

Wu et al., 2022 [Publ 4238], fig. 3

iT tT bi riS oT roT roS iD
4 4 2 4 70 16 6 21.0
  +----------------- Aculamprotula  
  |                                 
  |        +-------- Unio_tumidus **   
--+  +-----+                        
  |  |     |  +----- Unio_nanus **     
  |  |     +--+                     
  |  |        |  +-- Unio_pictorum **  
  +--+        +--+                  
     |           +-- Unio_delphinus ** 
     |                              
     |  +----------- Pseudocuneopsis
     +--+                           
        |  +-------- Nodularia      
        +--+                        
           |  +----- Schistodesmus  
           +--+                     
              |  +-- Tchangsinaia   
              +--+                  
                 +-- Cuneopsis      

Wang et al., 2018 [Publ 3846], fig. 2

iT tT bi riS oT roT roS iD
4 4 2 4 28 4 4 21.0
  +-------------- Aculamprotula 
  |                             
  |        +----- Nodularia     
--+  +-----+                    
  |  |     |  +-- Tchangsinaia  
  |  |     +--+                 
  +--+        +-- Cuneopsis     
     |                          
     |  +-------- Unio_tumidus **  
     +--+                       
        |  +----- Unio_nanus **    
        +--+                    
           |  +-- Unio_pictorum ** 
           +--+                 
              +-- Unio_delphinus **

Wu et al., 2020 [Publ 4121], fig. 3

iT tT bi riS oT roT roS iD
4 4 2 4 46 10 6 21.0
  +----------------- Aculamprotula  
  |                                 
  |        +-------- Unio_tumidus **   
--+  +-----+                        
  |  |     |  +----- Unio_nanus **     
  |  |     +--+                     
  |  |        |  +-- Unio_delphinus ** 
  +--+        +--+                  
     |           +-- Unio_pictorum **  
     |                              
     |  +----------- Pseudocuneopsis
     +--+                           
        |  +-------- Nodularia      
        +--+                        
           |  +----- Tchangsinaia   
           +--+                     
              |  +-- Cuneopsis      
              +--+                  
                 +-- Schistodesmus  

Froufe et al., 2019 [Publ 3936], fig. 3

iT tT bi riS oT roT roS iD
4 4 2 4 41 6 4 21.0
  +-------------- Aculamprotula 
  |                             
  |        +----- Nodularia     
--+  +-----+                    
  |  |     |  +-- Tchangsinaia  
  |  |     +--+                 
  +--+        +-- Schistodesmus 
     |                          
     |  +-------- Unio_tumidus **  
     +--+                       
        |  +----- Unio_nanus **    
        +--+                    
           |  +-- Unio_pictorum ** 
           +--+                 
              +-- Unio_delphinus **

Aunins et al., 2018 [Publ 3863], fig. 1

iT tT bi riS oT roT roS iD
4 4 2 4 37 7 5 21.0
  +-------------- Aculamprotula 
  |                             
  |     +-------- Nodularia     
--+  +--+                       
  |  |  |  +----- Tchangsinaia  
  |  |  +--+                    
  |  |     |  +-- Cuneopsis     
  +--+     +--+                 
     |        +-- Schistodesmus 
     |                          
     |  +-------- Unio_tumidus **  
     +--+                       
        |  +----- Unio_nanus **    
        +--+                    
           |  +-- Unio_pictorum ** 
           +--+                 
              +-- Unio_delphinus **

Soroka, 2010 [Publ 3404], fig. 3a

iT tT bi riS oT roT roS iD
4 4 2 4 5 0 0 21.0
  +-------- Unio_tumidus ** 
--+                      
  |  +----- Unio **         
  +--+                   
     |  +-- Unio_pictorum **
     +--+                
        +-- Unio_mancus **  

Kim et al., 2021 [Publ 4197], fig. 1

iT tT bi riS oT roT roS iD
4 4 2 4 13 7 4 21.0
        +----- Nodularia     
  +-----+                    
  |     |  +-- Schistodesmus 
  |     +--+                 
--+        +-- Cuneopsis     
  |                          
  |  +-------- Unio_tumidus **  
  +--+                       
     |  +----- Unio_nanus **    
     +--+                    
        |  +-- Unio_delphinus **
        +--+                 
           +-- Unio_pictorum ** 

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

iT tT bi riS oT roT roS iD
4 4 2 4 16 1 1 21.0
  +----------- Aculamprotula 
--+                          
  |  +-------- Unio_tumidus **  
  +--+                       
     |  +----- Unio_nanus **    
     +--+                    
        |  +-- Unio_pictorum ** 
        +--+                 
           +-- Unio_delphinus **

Wu et al., 2019 [Publ 3864], fig. 3

iT tT bi riS oT roT roS iD
4 4 2 4 43 9 6 21.0
  +----------------- Aculamprotula  
  |                                 
  |        +-------- Unio_tumidus **   
--+  +-----+                        
  |  |     |  +----- Unio_nanus **     
  |  |     +--+                     
  |  |        |  +-- Unio_delphinus ** 
  +--+        +--+                  
     |           +-- Unio_pictorum **  
     |                              
     |  +----------- Pseudocuneopsis
     +--+                           
        |  +-------- Nodularia      
        +--+                        
           |  +----- Tchangsinaia   
           +--+                     
              |  +-- Cuneopsis      
              +--+                  
                 +-- Schistodesmus  

Wu et al., 2021 [Publ 4366], fig. 3-ML

iT tT bi riS oT roT roS iD
4 4 2 4 70 11 6 21.0
  +----------------- Aculamprotula  
  |                                 
  |        +-------- Unio_tumidus **   
--+  +-----+                        
  |  |     |  +----- Unio_nanus **     
  |  |     +--+                     
  |  |        |  +-- Unio_pictorum **  
  +--+        +--+                  
     |           +-- Unio_delphinus ** 
     |                              
     |  +----------- Pseudocuneopsis
     +--+                           
        |  +-------- Nodularia      
        +--+                        
           |  +----- Tchangsinaia   
           +--+                     
              |  +-- Schistodesmus  
              +--+                  
                 +-- Cuneopsis      

Fukata & Iigo, 2019 [Publ 4108], fig. 1

iT tT bi riS oT roT roS iD
4 4 2 4 12 8 6 21.0
     +-------- Unio_tumidus **   
  +--+                        
  |  |  +----- Unio_nanus **     
  |  +--+                     
  |     |  +-- Unio_delphinus ** 
--+     +--+                  
  |        +-- Unio_pictorum **  
  |                           
  |  +-------- Pseudocuneopsis
  |  |                        
  +--+     +-- Schistodesmus  
     |  +--+                  
     |  |  +-- Cuneopsis      
     +--+                     
        |  +-- Tchangsinaia   
        +--+                  
           +-- Nodularia      

Klishko et al., 2018 [Publ 3945], fig. 8

iT tT bi riS oT roT roS iD
4 4 1 4 4 4 1 22.0
  +-------- Nodularia    
  |                      
--+  +----- Unio_tumidus ** 
  |  |                   
  +--+----- Unio_gibbus **  
     |                   
     |  +-- Unio_nanus **   
     +--+                
        +-- Unio_pictorum **

Soroka, 2005 [Publ 2726], fig. 3

iT tT bi riS oT roT roS iD
4 4 2 4 1 0 0 21.0
  +-------- Unio_tumidus ** 
--+                      
  |  +----- Unio_nanus **   
  +--+                   
     |  +-- Unio_caffer **  
     +--+                
        +-- Unio_pictorum **

Wu et al., 2022 [Publ 4238], fig. 2d

iT tT bi riS oT roT roS iD
4 4 2 4 35 19 7 21.0
  +-------------------- Aculamprotula  
  |                                    
  |           +-------- Unio_tumidus **   
--+  +--------+                        
  |  |        |  +----- Unio_nanus **     
  |  |        +--+                     
  |  |           |  +-- Unio_pictorum **  
  +--+           +--+                  
     |              +-- Unio_delphinus ** 
     |                                 
     |  +-------------- Pseudocuneopsis
     +--+                              
        |  +----------- Nodularia      
        +--+                           
           |  +-------- Schistodesmus  
           +--+                        
              |  +----- Cuneopsis      
              +--+                     
                 |  +-- Tchangsinaia   
                 +--+                  
                    +-- Inversiunio    

Huang et al., 2019 [Publ 3843], fig. 3

iT tT bi riS oT roT roS iD
4 4 2 4 53 7 4 21.0
  +-------------- Aculamprotula 
  |                             
  |        +----- Nodularia     
--+  +-----+                    
  |  |     |  +-- Tchangsinaia  
  |  |     +--+                 
  +--+        +-- Schistodesmus 
     |                          
     |  +-------- Unio_tumidus **  
     +--+                       
        |  +----- Unio_nanus **    
        +--+                    
           |  +-- Unio_delphinus **
           +--+                 
              +-- Unio_pictorum ** 

Froufe et al., 2019 [Publ 3936], fig. 6

iT tT bi riS oT roT roS iD
4 4 2 4 41 6 4 21.0
  +-------------- Aculamprotula 
  |                             
  |        +----- Nodularia     
--+  +-----+                    
  |  |     |  +-- Tchangsinaia  
  |  |     +--+                 
  +--+        +-- Schistodesmus 
     |                          
     |  +-------- Unio_tumidus **  
     +--+                       
        |  +----- Unio_nanus **    
        +--+                    
           |  +-- Unio_pictorum ** 
           +--+                 
              +-- Unio_delphinus **

Klishko et al., 2019 [Publ 3886], fig. 9

iT tT bi riS oT roT roS iD
4 4 1 4 7 4 2 22.0
        +-- Middendorffinaia
  +-----+                   
  |     +-- Nodularia       
--+                         
  |  +----- Unio_tumidus **    
  |  |                      
  +--+----- Unio_gibbus **     
     |                      
     |  +-- Unio_nanus **      
     +--+                   
        +-- Unio_pictorum **   

Soroka, 2010 [Publ 3404], fig. 2

iT tT bi riS oT roT roS iD
3 3 0 3 4 0 0 22.0
  +-- Unio_tumidus ** 
  |                
--+-- Unio_pictorum **
  |                
  +-- Unio_nanus **   

Bolotov et al., 2018 [Publ 3826], fig. 2

iT tT bi riS oT roT roS iD
3 3 1 3 80 2 1 20.0
  +-------- Nodularia    
--+                      
  |  +----- Unio_tumidus ** 
  +--+                   
     |  +-- Unio_crassus ** 
     +--+                
        +-- Unio_pictorum **

Huang et al., 2019 [Publ 3843], fig. 3-M

iT tT bi riS oT roT roS iD
3 3 1 3 20 2 2 22.0
  +----------- Schistodesmus 
--+                          
  |  +-------- Aculamprotula 
  +--+                       
     |  +----- Unio_tumidus **  
     +--+                    
        |  +-- Unio_nanus **    
        +--+                 
           +-- Unio_delphinus **

Källersjö et al., 2005 [Publ 2684], fig. 1A

iT tT bi riS oT roT roS iD
3 3 0 3 4 0 0 21.0
  +-- Unio_crassus ** 
  |                
--+-- Unio_pictorum **
  |                
  +-- Unio_tumidus ** 

Dyduch-Falniowska & Koziol, 1989 [Publ 3814], fig. 5_7

iT tT bi riS oT roT roS iD
3 3 1 3 2 0 0 22.0
  +----- Unio_pictorum **
--+                   
  |  +-- Unio_tumidus ** 
  +--+                
     +-- Unio_crassus ** 

Soroka, 2010 [Publ 3404], fig. 4

iT tT bi riS oT roT roS iD
3 3 1 3 4 0 0 22.0
  +----- Unio_tumidus ** 
--+                   
  |  +-- Unio_pictorum **
  +--+                
     +-- Unio_nanus **   

Plazzi et al., 2011 [Publ 3676], fig. 5

iT tT bi riS oT roT roS iD
3 3 1 3 15 0 0 20.0
  +----- Unio_tumidus ** 
--+                   
  |  +-- Unio_crassus ** 
  +--+                
     +-- Unio_pictorum **

Manendo et al., 2008 [Publ 2669], fig. 1

iT tT bi riS oT roT roS iD
3 3 1 3 10 0 0 22.0
  +----- Unio_crassus ** 
--+                   
  |  +-- Unio_pictorum **
  +--+                
     +-- Unio_tumidus ** 

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iT tT bi riS oT roT roS iD
3 3 1 3 53 9 5 20.0
  +-------------- Middendorffinaia
  |                               
--+  +----------- Aculamprotula   
  |  |                            
  |  |     +----- Unio_tumidus **    
  +--+  +--+                      
     |  |  |  +-- Unio_pictorum **   
     |  |  +--+                   
     +--+     +-- Unio_crassus **    
        |                         
        |  +----- Nodularia       
        +--+                      
           |  +-- Tchangsinaia    
           +--+                   
              +-- Cuneopsis       

Zieritz et al., 2012 [Publ 3673], fig. 4

iT tT bi riS oT roT roS iD
3 3 1 3 5 0 0 22.0
  +----- Unio_crassus ** 
--+                   
  |  +-- Unio_tumidus ** 
  +--+                
     +-- Unio_pictorum **

Kilikowska et al., 2020 [Publ 4335], fig. 4a

iT tT bi riS oT roT roS iD
3 3 1 3 0 0 0 22.0
  +----- Unio_tumidus **
--+                  
  |  +-- Unio_crassus **
  +--+               
     +-- Unio_nanus **  

Soroka, 2010 [Publ 3404], fig. 5b

iT tT bi riS oT roT roS iD
3 3 1 3 5 0 0 22.0
  +----- Unio_tumidus ** 
--+                   
  |  +-- Unio_pictorum **
  +--+                
     +-- Unio_nanus **   

Soroka, 2010 [Publ 3404], fig. 8b

iT tT bi riS oT roT roS iD
3 3 1 3 3 0 0 22.0
  +----- Unio_tumidus ** 
--+                   
  |  +-- Unio_pictorum **
  +--+                
     +-- Unio_nanus **   

Wu et al., 2022 [Publ 4238], fig. 2e

iT tT bi riS oT roT roS iD
3 3 1 3 14 9 6 22.0
  +----------------- Aculamprotula  
  |                                 
  |           +----- Unio_tumidus **   
--+  +--------+                     
  |  |        |  +-- Unio_nanus **     
  |  |        +--+                  
  +--+           +-- Unio_delphinus ** 
     |                              
     |  +----------- Pseudocuneopsis
     +--+                           
        |  +-------- Nodularia      
        +--+                        
           |  +----- Tchangsinaia   
           +--+                     
              |  +-- Cuneopsis      
              +--+                  
                 +-- Schistodesmus  

Bolotov et al., 2020 [Publ 4084], fig. 1

iT tT bi riS oT roT roS iD
3 3 1 3 33 2 2 20.0
  +----------- Middendorffinaia
--+                            
  |  +-------- Nodularia       
  +--+                         
     |  +----- Unio_tumidus **    
     +--+                      
        |  +-- Unio_pictorum **   
        +--+                   
           +-- Unio_crassus **    

Guerra et al., 2019 [Publ 4168], fig. 2-M

iT tT bi riS oT roT roS iD
3 3 1 3 19 2 2 22.0
  +----------- Aculamprotula 
--+                          
  |  +-------- Schistodesmus 
  +--+                       
     |  +----- Unio_tumidus **  
     +--+                    
        |  +-- Unio_nanus **    
        +--+                 
           +-- Unio_delphinus **

Soroka, 2010 [Publ 3404], fig. 2-M

iT tT bi riS oT roT roS iD
3 3 1 3 2 0 0 22.0
  +----- Unio_nanus **   
--+                   
  |  +-- Unio_tumidus ** 
  +--+                
     +-- Unio_pictorum **

Soroka, 2010 [Publ 3404], fig. 3a-M

iT tT bi riS oT roT roS iD
3 3 1 3 3 0 0 22.0
  +----- Unio_nanus **   
--+                   
  |  +-- Unio_tumidus ** 
  +--+                
     +-- Unio_pictorum **

Soroka, 2010 [Publ 3404], fig. 3b-M

iT tT bi riS oT roT roS iD
3 3 1 3 3 0 0 22.0
  +----- Unio_tumidus ** 
--+                   
  |  +-- Unio_nanus **   
  +--+                
     +-- Unio_pictorum **

Guerra et al., 2019 [Publ 4168], fig. 1-M

iT tT bi riS oT roT roS iD
3 3 1 3 19 2 2 22.0
  +----------- Schistodesmus 
--+                          
  |  +-------- Aculamprotula 
  +--+                       
     |  +----- Unio_tumidus **  
     +--+                    
        |  +-- Unio_nanus **    
        +--+                 
           +-- Unio_delphinus **

Soroka, 2010 [Publ 3404], fig. 8a

iT tT bi riS oT roT roS iD
3 3 1 3 3 0 0 22.0
  +----- Unio_tumidus ** 
--+                   
  |  +-- Unio_pictorum **
  +--+                
     +-- Unio_nanus **   

Sano et al., 2020 [Publ 4138], fig. 3

iT tT bi riS oT roT roS iD
3 3 1 3 66 10 5 22.0
  +-------------- Aculamprotula 
  |                             
  |        +----- Unio_nanus **    
--+  +-----+                    
  |  |     |  +-- Unio_delphinus **
  |  |     +--+                 
  +--+        +-- Unio_pictorum ** 
     |                          
     |  +-------- Inversiunio   
     +--+                       
        |  +----- Nodularia     
        +--+                    
           |  +-- Schistodesmus 
           +--+                 
              +-- Tchangsinaia  

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

iT tT bi riS oT roT roS iD
3 3 1 3 20 0 0 22.0
  +----- Unio_tumidus **  
--+                    
  |  +-- Unio_delphinus **
  +--+                 
     +-- Unio_nanus **    

Soroka, 2010 [Publ 3404], fig. 5a

iT tT bi riS oT roT roS iD
3 3 1 3 5 0 0 22.0
  +----- Unio_nanus **   
--+                   
  |  +-- Unio_pictorum **
  +--+                
     +-- Unio_tumidus ** 

Bolotov et al., 2017 [Publ 3761], fig. S1

iT tT bi riS oT roT roS iD
3 3 1 3 136 7 5 20.0
           +----- Unio_tumidus ** 
  +--------+                   
  |        |  +-- Unio_pictorum **
  |        +--+                
--+           +-- Unio_crassus ** 
  |                            
  |  +----------- Aculamprotula
  +--+                         
     |  +-------- Nodularia    
     +--+                      
        |  +----- Cuneopsis    
        +--+                   
           |  +-- Tchangsinaia 
           +--+                
              +-- Schistodesmus

Froufe et al., 2019 [Publ 3936], fig. 3-M

iT tT bi riS oT roT roS iD
3 3 1 3 16 0 0 22.0
  +----- Unio_tumidus **  
--+                    
  |  +-- Unio_nanus **    
  +--+                 
     +-- Unio_delphinus **

Froufe et al., 2017 [Publ 3802], fig. 7

iT tT bi riS oT roT roS iD
3 3 1 3 7 0 0 22.0
  +----- Unio_mancus **     
--+                      
  |  +-- Unio_elongatulus **
  +--+                   
     +-- Unio_pictorum **   

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iT tT bi riS oT roT roS iD
3 3 1 3 73 7 5 20.0
              +----- Unio_tumidus **       
  +-----------+                         
  |           |  +-- Unio_pictorum **      
  |           +--+                      
--+              +-- Unio_crassus **       
  |                                     
  |  +-------------- Aculamprotula      
  +--+                                  
     |  +----------- Nodularia          
     +--+                               
        |  +-------- Schistodesmus      
        +--+                            
           |  +----- Cuneopsis_heudei   
           +--+                         
              |  +-- Tchangsinaia       
              +--+                      
                 +-- Cuneopsis_rufescens

Huang et al., 2018 [Publ 3828], fig. 6-M

iT tT bi riS oT roT roS iD
3 3 1 3 14 1 1 20.0
  +-------- Aculamprotula 
--+                       
  |  +----- Unio_tumidus **  
  +--+                    
     |  +-- Unio_crassus **  
     +--+                 
        +-- Unio_delphinus **

Bogatov et al., 2018 [Publ 3859], fig. 1

iT tT bi riS oT roT roS iD
3 5 3 3 3 0 0 22.0
  +-------- Unio_tumidus ** 
  |                      
--+     +-- Anodonta     
  |  +--+                
  |  |  +-- Pseudanodonta
  +--+                   
     |  +-- Unio_crassus ** 
     +--+                
        +-- Unio_pictorum **

Bogatov et al., 2018 [Publ 3859], fig. 2

iT tT bi riS oT roT roS iD
3 5 3 3 3 0 0 24.0
  +----------- Unio_crassus ** 
--+                         
  |  +-------- Unio_pictorum **
  +--+                      
     |  +----- Unio_tumidus ** 
     +--+                   
        |  +-- Anodonta     
        +--+                
           +-- Pseudanodonta

Källersjö et al., 2005 [Publ 2684], fig. 1B-F

iT tT bi riS oT roT roS iD
3 3 1 3 4 0 0 20.0
  +----- Unio_tumidus ** 
--+                   
  |  +-- Unio_crassus ** 
  +--+                
     +-- Unio_pictorum **

Bolotov et al., 2017 [Publ 3761], fig. S2

iT tT bi riS oT roT roS iD
3 3 1 3 136 7 5 20.0
        +----- Unio_tumidus **       
  +-----+                         
  |     |  +-- Unio_crassus **       
  |     +--+                      
  |        +-- Unio_pictorum **      
--+                               
  |  +-------- Aculamprotula      
  |  |                            
  |  +-------- Schistodesmus      
  +--+                            
     +-------- Nodularia          
     |                            
     |  +----- Cuneopsis_heudei   
     +--+                         
        |  +-- Tchangsinaia       
        +--+                      
           +-- Cuneopsis_rufescens

Bolotov et al., 2017 [Publ 3764], fig. S1

iT tT bi riS oT roT roS iD
3 10 0 3 122 7 5 28.0
  +-- Unio_crassus **         
  |                        
  +-- Unio_pictorum **        
  |                        
  +-- Unio_tumidus **         
  |                        
  +-- Aculamprotula        
  |                        
  +-- Cuneopsis_heudei     
--+                        
  +-- Tchangsinaia         
  |                        
  +-- Cuneopsis_rufescens  
  |                        
  +-- Nodularia_douglasiae 
  |                        
  +-- Nodularia_nuxpersicae
  |                        
  +-- Schistodesmus        

Sano et al., 2020 [Publ 4138], fig. 2

iT tT bi riS oT roT roS iD
3 3 1 3 66 10 5 22.0
  +-------------- Aculamprotula 
  |                             
  |        +----- Unio_nanus **    
--+  +-----+                    
  |  |     |  +-- Unio_delphinus **
  |  |     +--+                 
  +--+        +-- Unio_pictorum ** 
     |                          
     |  +-------- Inversiunio   
     +--+                       
        |  +----- Nodularia     
        +--+                    
           |  +-- Tchangsinaia  
           +--+                 
              +-- Schistodesmus 

Froufe et al., 2019 [Publ 3936], fig. 4

iT tT bi riS oT roT roS iD
3 3 1 3 16 0 0 22.0
  +----- Unio_tumidus **  
--+                    
  |  +-- Unio_nanus **    
  +--+                 
     +-- Unio_delphinus **

Guerra et al., 2019 [Publ 4168], fig. 1

iT tT bi riS oT roT roS iD
3 3 1 3 38 2 2 22.0
  +----------- Aculamprotula 
--+                          
  |  +-------- Schistodesmus 
  +--+                       
     |  +----- Unio_tumidus **  
     +--+                    
        |  +-- Unio_nanus **    
        +--+                 
           +-- Unio_delphinus **

Sano et al., 2020 [Publ 4138], fig. 4

iT tT bi riS oT roT roS iD
3 3 1 3 66 10 5 22.0
  +-------------- Aculamprotula 
  |                             
  |        +----- Unio_nanus **    
--+  +-----+                    
  |  |     |  +-- Unio_delphinus **
  |  |     +--+                 
  +--+        +-- Unio_pictorum ** 
     |                          
     |  +-------- Nodularia     
     +--+                       
        |  +----- Inversiunio   
        +--+                    
           |  +-- Schistodesmus 
           +--+                 
              +-- Tchangsinaia  

Guerra et al., 2019 [Publ 4168], fig. 2

iT tT bi riS oT roT roS iD
3 3 1 3 38 2 2 22.0
  +-------- Schistodesmus 
--+                       
  |  +----- Unio_tumidus **  
  +--+                    
     |  +-- Unio_nanus **    
     +--+                 
        +-- Unio_delphinus **

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iT tT bi riS oT roT roS iD
3 3 1 3 73 7 5 20.0
              +----- Unio_tumidus **       
  +-----------+                         
  |           |  +-- Unio_pictorum **      
  |           +--+                      
--+              +-- Unio_crassus **       
  |                                     
  |  +-------------- Aculamprotula      
  +--+                                  
     |  +----------- Nodularia          
     +--+                               
        |  +-------- Schistodesmus      
        +--+                            
           |  +----- Cuneopsis_heudei   
           +--+                         
              |  +-- Tchangsinaia       
              +--+                      
                 +-- Cuneopsis_rufescens

Wu et al., 2021 [Publ 4366], fig. 6

iT tT bi riS oT roT roS iD
2 2 0 2 89 9 5 --
  +-------------- Aculamprotula
  |                            
--+           +-- Unio_pictorum **
  |  +--------+                
  |  |        +-- Unio_tumidus ** 
  +--+                         
     |  +-------- Nodularia    
     +--+                      
        |  +----- Cuneopsis    
        +--+                   
           |  +-- Tchangsinaia 
           +--+                
              +-- Schistodesmus

Sonowal et al., 2021 [Publ 4221], fig. 4

iT tT bi riS oT roT roS iD
2 4 0 2 14 1 1 23.0
  +----------- Aculamprotula  
--+                           
  |  +-------- Unio_pictorum **  
  +--+                        
     |  +----- Anodonta       
     +--+                     
        |  +-- Unio_ravoisieri **
        +--+                  
           +-- Lamellidens    

Bolotov et al., 2016 [Publ 3743], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 2 28 0 0 --
  +-- Unio_pictorum **  
--+                  
  +-- Unio_ravoisieri **

Fonseca et al., 2016 [Publ 3775], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 30 5 4 --
  +----------- Aculamprotula 
  |                          
--+        +-- Unio_pictorum ** 
  |  +-----+                 
  |  |     +-- Unio_delphinus **
  +--+                       
     |  +----- Schistodesmus 
     +--+                    
        |  +-- Nodularia     
        +--+                 
           +-- Tchangsinaia  

Sonowal et al., 2021 [Publ 4221], fig. 3b

iT tT bi riS oT roT roS iD
2 9 0 2 13 1 1 28.0
  +-- Aculamprotula  
  |                  
  +-- Lamprotula     
  |                  
  +-- Lampsilis      
  |                  
  +-- Potomida       
  |                  
--+-- Unio_pictorum **  
  |                  
  +-- Anodonta       
  |                  
  +-- Unio_ravoisieri **
  |                  
  +-- Lamellidens    
  |                  
  +-- Elliptio       

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 32 0 0 --
  +-- Unio_pictorum **
--+                
  +-- Unio_tumidus ** 

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 31 5 4 --
  +----------- Aculamprotula 
  |                          
--+        +-- Unio_delphinus **
  |  +-----+                 
  |  |     +-- Unio_pictorum ** 
  +--+                       
     |  +----- Schistodesmus 
     +--+                    
        |  +-- Tchangsinaia  
        +--+                 
           +-- Nodularia     

Chase et al., 2018 [Publ 3861], fig. 1-M

iT tT bi riS oT roT roS iD
2 2 0 2 9 0 0 --
  +-- Unio_delphinus **
--+                 
  +-- Unio_tumidus **  

Dai et al., 2024 [Publ 4358], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 2 50 15 6 --
  +----------------- Aculamprotula  
  |                                 
--+              +-- Unio_nanus **     
  |  +-----------+                  
  |  |           +-- Unio_delphinus ** 
  +--+                              
     |  +----------- Pseudocuneopsis
     +--+                           
        |  +-------- Nodularia      
        +--+                        
           |  +----- Schistodesmus  
           +--+                     
              |  +-- Tchangsinaia   
              +--+                  
                 +-- Cuneopsis      

Hoeh et al., 2009 [Publ 2716], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 2 44 0 0 --
  +-- Unio_pictorum **
--+                
  +-- Unio_caffer **  

Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11

iT tT bi riS oT roT roS iD
2 7 0 3 23 1 1 27.0
  +-------------- Unio_pictorum__2 **
  |                               
--+           +-- Unio_pictorum__1 **
  |  +--------+                   
  |  |        +-- Unio_caffer **     
  +--+                            
     |  +-------- Anodonta        
     |  |                         
     +--+  +----- Lasmigona       
        |  |                      
        +--+  +-- Strophitus      
           |  |                   
           +--+-- Alasmidonta     
              |                   
              +-- Pyganodon       

Tomilova et al., 2020 [Publ 4097], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 6 0 0 --
  +-- Unio_tumidus ** 
--+                
  +-- Unio_pictorum **

Graf & Cummings, 2006 [Publ 1968], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 45 0 0 --
  +-- Unio_pictorum **
--+                
  +-- Unio_caffer **  

Burzynski et al., 2017 [Publ 3774], fig. 1-bottom

iT tT bi riS oT roT roS iD
2 2 0 2 11 1 1 --
  +----- Aculamprotula
--+                   
  |  +-- Unio_nanus **   
  +--+                
     +-- Unio_tumidus ** 

Hoeh et al., 2002 [Publ 2653], fig. 2

iT tT bi riS oT roT roS iD
2 5 0 2 29 0 0 24.0
  +----------- Unio_pictorum **
--+                         
  |  +-------- Unio_tumidus ** 
  +--+                      
     |  +----- Anodonta     
     +--+                   
        |  +-- Pyganodon    
        +--+                
           +-- Strophitus   

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

iT tT bi riS oT roT roS iD
2 2 0 2 28 0 0 --
  +-- Unio_pictorum **
--+                
  +-- Unio_caffer **  

Hoeh et al., 2001 [Publ 2652], fig. 14f4

iT tT bi riS oT roT roS iD
2 5 0 2 29 0 0 24.0
  +-------- Unio_pictorum **
  |                      
--+-------- Unio_tumidus ** 
  |                      
  |  +----- Anodonta     
  +--+                   
     |  +-- Strophitus   
     +--+                
        +-- Pyganodon    

Hoeh et al., 2001 [Publ 2652], fig. 14f2

iT tT bi riS oT roT roS iD
2 2 0 2 29 0 0 --
  +-- Unio_pictorum **
--+                
  +-- Unio_tumidus ** 

Sonowal et al., 2021 [Publ 4221], fig. 3a

iT tT bi riS oT roT roS iD
2 4 0 2 13 1 1 23.0
  +-- Anodonta       
  |                  
  +-- Unio_ravoisieri **
--+                  
  +-- Unio_pictorum **  
  |                  
  +-- Aculamprotula  

Chase et al., 2018 [Publ 3861], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 9 0 0 --
  +-- Unio_delphinus **
--+                 
  +-- Unio_tumidus **  

Burzynski et al., 2017 [Publ 3774], fig. 1-top

iT tT bi riS oT roT roS iD
2 2 0 2 11 1 1 --
  +----- Aculamprotula
--+                   
  |  +-- Unio_nanus **   
  +--+                
     +-- Unio_tumidus ** 

Riccardi et al., 2019 [Publ 4089], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 18 0 0 --
  +-- Unio_mancus **     
--+                   
  +-- Unio_elongatulus **

Wu et al., 2021 [Publ 4366], fig. 2_I

iT tT bi riS oT roT roS iD
2 2 0 2 89 9 5 --
  +-------------- Aculamprotula
  |                            
--+           +-- Unio_pictorum **
  |  +--------+                
  |  |        +-- Unio_tumidus ** 
  +--+                         
     |  +-------- Nodularia    
     +--+                      
        |  +----- Cuneopsis    
        +--+                   
           |  +-- Tchangsinaia 
           +--+                
              +-- Schistodesmus

Wu et al., 2018 [Publ 3836], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 51 12 5 --
  +----------- Aculamprotula
  |                         
  |        +-- Unio_crassus ** 
--+  +-----+                
  |  |     +-- Unio_pictorum **
  |  |                      
  +--+     +-- Nodularia    
     |  +--+                
     |  |  +-- Schistodesmus
     +--+                   
        |  +-- Tchangsinaia 
        +--+                
           +-- Cuneopsis    

Chen et al., 2023 [Publ 4302], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 38 15 9 --
  +-------------------- Aculamprotula   
  |                                     
  |                 +-- Middendorffinaia
--+  +--------------+                   
  |  |              +-- Nodularia       
  |  |                                  
  +--+  +-------------- Cuneopsis       
     |  |                               
     |  |           +-- Diaurora        
     +--+-----------+                   
        |           +-- Schistodesmus   
        |                               
        |  +----------- Pseudocuneopsis 
        +--+                            
           |  +-------- Inversiunio     
           +--+                         
              |  +----- Tchangsinaia    
              +--+                      
                 |  +-- Unio_pictorum **   
                 +--+                   
                    +-- Unio_nanus **      

Chase et al., 2018 [Publ 3847], fig. 5

iT tT bi riS oT roT roS iD
2 2 0 2 8 0 0 --
  +-- Unio_delphinus **
--+                 
  +-- Unio_tumidus **  

Graf, 2002 [Publ 2651], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 16 0 0 --
  +-- Unio_pictorum **
--+                
  +-- Unio_caffer **  

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iT tT bi riS oT roT roS iD
2 2 0 2 34 0 0 --
  +-- Unio_pictorum **
--+                
  +-- Unio_caffer **  

Soroka, 2010 [Publ 3404], fig. 8b-M

iT tT bi riS oT roT roS iD
2 2 0 2 1 0 0 --
  +-- Unio_pictorum **
--+                
  +-- Unio_tumidus ** 

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iT tT bi riS oT roT roS iD
2 2 0 2 34 0 0 --
  +-- Unio_pictorum **
--+                
  +-- Unio_caffer **  

Chase et al., 2018 [Publ 3847], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 2 36 0 0 --
  +-- Unio_delphinus **
--+                 
  +-- Unio_tumidus **  

Hoeh et al., 2009 [Publ 2716], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 45 0 0 --
  +-- Unio_pictorum **
--+                
  +-- Unio_caffer **  

Sayenko et al., 2021 [Publ 4189], fig. 6

iT tT bi riS oT roT roS iD
2 2 0 2 11 11 3 --
        +-- Unio_pictorum **   
  +-----+                   
  |     +-- Unio_nanus **      
--+                         
  |  +----- Inversiunio     
  +--+                      
     |  +-- Middendorffinaia
     +--+                   
        +-- Nodularia       

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

iT tT bi riS oT roT roS iD
2 2 0 2 29 0 0 --
  +-- Unio_pictorum **
--+                
  +-- Unio_tumidus ** 

Bolotov et al., 2016 [Publ 3743], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 28 0 0 --
  +-- Unio_pictorum **  
--+                  
  +-- Unio_ravoisieri **

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

iT tT bi riS oT roT roS iD
2 2 0 2 28 0 0 --
  +-- Unio_pictorum **
--+                
  +-- Unio_caffer **  

Hoeh et al., 2009 [Publ 2716], fig. 5

iT tT bi riS oT roT roS iD
2 2 0 2 44 0 0 --
  +-- Unio_pictorum **
--+                
  +-- Unio_caffer **  

Soroka, 2010 [Publ 3404], fig. 8a-M

iT tT bi riS oT roT roS iD
2 2 0 2 1 0 0 --
  +-- Unio_pictorum **
--+                
  +-- Unio_tumidus ** 

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

iT tT bi riS oT roT roS iD
2 2 0 2 29 0 0 --
  +-- Unio_pictorum **
--+                
  +-- Unio_tumidus ** 

Zieritz et al., 2024 [Publ 4381], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 45 0 0 --
-- Unio_crassus

Rosenberg et al., 1997 [Publ 2720], fig. 2A

iT tT bi riS oT roT roS iD
1 1 0 1 21 0 0 --
-- Unio_pictorum

Nagel et al., 1998 [Publ 1946], fig. 9

iT tT bi riS oT roT roS iD
1 1 0 1 5 0 0 --
-- Unio

Dai et al., 2024 [Publ 4386], fig. 5-BI

iT tT bi riS oT roT roS iD
1 1 0 1 20 10 6 --
  +----------- Unio_crassus **    
  |                            
--+  +-------- Middendorffinaia
  |  |                         
  +--+  +----- Cuneopsis       
     |  |                      
     +--+----- Nodularia       
        |                      
        |  +-- Inversiunio     
        +--+                   
           +-- Schistodesmus   

González & Giribet, 2015 [Publ 3632], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 5 0 0 --
-- Unio_pictorum

Bolotov et al., 2015 [Publ 3646], fig. 2B

iT tT bi riS oT roT roS iD
1 1 0 1 8 0 0 --
-- Unio_pictorum

Araujo et al., 2009 [Publ 2395], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 7 0 0 --
-- Unio

Giribet & Distel, 2003 [Publ 3670], fig. 3f5

iT tT bi riS oT roT roS iD
1 1 0 1 5 0 0 --
-- Unio_ravoisieri

Araujo et al., 2017 [Publ 3801], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 13 2 2 --
-- Unio_delphinus

González & Giribet, 2015 [Publ 3632], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 5 0 0 --
-- Unio_pictorum

Choi et al., 2020 [Publ 4169], fig. 2a

iT tT bi riS oT roT roS iD
1 1 0 1 11 11 4 --
  +----------- Nodularia    
--+                         
  |  +-------- Schistodesmus
  +--+                      
     |  +----- Cuneopsis    
     +--+                   
        |  +-- Unio_pictorum **
        +--+                
           +-- Tchangsinaia 

González & Giribet, 2015 [Publ 3632], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 5 0 0 --
-- Unio_pictorum

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iT tT bi riS oT roT roS iD
1 1 0 1 56 2 2 --
  +----- Unio_gibbus ** 
--+                  
  |  +-- Nodularia   
  +--+               
     +-- Tchangsinaia

Wu et al., 2020 [Publ 4147], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 13 4 1 --
  +-- Unio_pictorum **
--+                
  +-- Aculamprotula

Xue et al., 2016 [Publ 3659], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 15 3 3 --
  +-------- Aculamprotula
--+                      
  |  +----- Schistodesmus
  +--+                   
     |  +-- Nodularia    
     +--+                
        +-- Unio_pictorum **

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 27 0 0 --
-- Unio_pictorum

Klunzinger et al., 2021 [Publ 4086], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 5 0 0 --
-- Unio_pictorum

Wang et al., 2016 [Publ 3714], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 2 0 0 --
-- Unio_tumidus

Bieler et al., 2014 [Publ 3622], fig. 28

iT tT bi riS oT roT roS iD
1 1 0 1 4 0 0 --
-- Unio_pictorum

Wu et al., 2023 [Publ 4296], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 22 12 7 --
  +----------------- Unio_tumidus **   
  |                                 
--+  +-------------- Aculamprotula  
  |  |                              
  +--+  +----------- Cuneopsis      
     |  |                           
     +--+        +-- Nodularia      
        |  +-----+                  
        |  |     +-- Schistodesmus  
        +--+                        
           |  +----- Tchangsinaia   
           +--+                     
              |  +-- Inversiunio    
              +--+                  
                 +-- Pseudocuneopsis

Wu et al., 2018 [Publ 4143], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 21 5 2 --
  +----- Aculamprotula
--+                   
  |  +-- Unio_pictorum **
  +--+                
     +-- Nodularia    

Huff et al., 2004 [Publ 2697], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 11 0 0 --
-- Unio_ravoisieri

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 33 6 2 --
  +----- Aculamprotula
--+                   
  |  +-- Unio_pictorum **
  +--+                
     +-- Nodularia    

Wen et al., 2017 [Publ 3795], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 15 2 1 --
-- Unio_pictorum

Guerra et al., 2017 [Publ 3772], fig. 2b

iT tT bi riS oT roT roS iD
1 1 0 1 24 1 1 --
-- Unio_pictorum

Huff et al., 2004 [Publ 2697], fig. 5-6

iT tT bi riS oT roT roS iD
1 1 0 1 11 0 0 --
-- Unio_ravoisieri

Vikhrev et al., 2018 [Publ 3812], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 12 0 0 --
-- Unio_ravoisieri

Huff et al., 2004 [Publ 2697], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 10 0 0 --
-- Unio_ravoisieri

Liu et al., 2023 [Publ 4310], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 23 13 7 --
  +-------- Unio_tumidus **   
  |                        
  +-------- Aculamprotula  
  |                        
--+  +----- Cuneopsis      
  |  |                     
  |  |  +-- Nodularia      
  |  +--+                  
  +--+  +-- Schistodesmus  
     |                     
     |  +-- Tchangsinaia   
     |  |                  
     +--+-- Inversiunio    
        |                  
        +-- Pseudocuneopsis

Zhou et al., 2016 [Publ 3710], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 19 1 1 --
  +-- Schistodesmus
--+                
  +-- Unio_pictorum **

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 39 0 0 --
-- Unio_pictorum

Huang et al., 2013 [Publ 3629], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 15 0 0 --
-- Unio_pictorum

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 27 0 0 --
-- Unio_pictorum

Rosenberg et al., 1997 [Publ 2720], fig. 2B

iT tT bi riS oT roT roS iD
1 1 0 1 21 1 1 --
-- Unio_pictorum

Marshall et al., 2014 [Publ 3593], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 13 0 0 --
-- Unio_pictorum

Graf et al., 2014 [Publ 3591], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 10 0 0 --
-- Unio_pictorum

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 56 2 2 --
  +----- Unio_gibbus ** 
--+                  
  |  +-- Tchangsinaia
  +--+               
     +-- Nodularia   

Guerra et al., 2017 [Publ 3772], fig. 2a

iT tT bi riS oT roT roS iD
1 1 0 1 24 1 1 --
  +-- Aculamprotula
--+                
  +-- Unio_pictorum **

Graf et al., 2015 [Publ 3619], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 19 0 0 --
-- Unio_pictorum

Combosch et al., 2017 [Publ 3806], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 14 0 0 --
-- Unio_pictorum

Combosch et al., 2017 [Publ 3806], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 14 0 0 --
-- Unio_pictorum

Rosenberg et al., 1994 [Publ 2719], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 18 0 0 --
-- Unio_pictorum

Chong et al., 2008 [Publ 2668], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 17 0 0 --
-- Unio_tumidus

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iT tT bi riS oT roT roS iD
1 1 0 1 38 5 4 --
  +-------- Unio_pictorum **
--+                      
  |  +----- Schistodesmus
  +--+                   
     |  +-- Nodularia    
     +--+                
        +-- Tchangsinaia 

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1-M

iT tT bi riS oT roT roS iD
1 1 0 1 5 0 0 --
-- Unio_delphinus

Bolotov et al., 2015 [Publ 3646], fig. 2A

iT tT bi riS oT roT roS iD
1 1 0 1 8 0 0 --
-- Unio_pictorum

Lopes-Lima et al., 2017 [Publ 3792], fig. 2-M

iT tT bi riS oT roT roS iD
1 1 0 1 9 0 0 --
-- Unio_delphinus

Taylor et al., 2007 [Publ 2696], fig. 1-3

iT tT bi riS oT roT roS iD
1 1 0 1 3 0 0 --
-- Unio_pictorum

Fonseca et al., 2016 [Publ 3775], fig. 1-M

iT tT bi riS oT roT roS iD
1 1 0 1 8 0 0 --
-- Unio_delphinus

Graf, 2000 [Publ 2647], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 16 0 0 --
-- Unio

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 9 0 0 --
-- Unio_delphinus

Dai et al., 2023 [Publ 4285], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 52 6 6 --
  +-------------- Aculamprotula  
  |                              
--+  +----------- Unio_nanus **     
  |  |                           
  +--+        +-- Pseudocuneopsis
     |  +-----+                  
     |  |     +-- Nodularia      
     +--+                        
        |  +----- Schistodesmus  
        +--+                     
           |  +-- Cuneopsis      
           +--+                  
              +-- Tchangsinaia   

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 35 5 4 --
  +----------- Aculamprotula
--+                         
  |  +-------- Unio_pictorum **
  +--+                      
     |  +----- Schistodesmus
     +--+                   
        |  +-- Nodularia    
        +--+                
           +-- Tchangsinaia 

Wu et al., 2020 [Publ 4121], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 33 4 1 --
  +-- Unio_pictorum **
--+                
  +-- Aculamprotula

Soroka & Burzynski, 2018 [Publ 3762], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 6 1 1 --
  +-- Unio_tumidus ** 
--+                
  +-- Aculamprotula

Wu et al., 2020 [Publ 4121], fig. S1

iT tT bi riS oT roT roS iD
1 1 0 1 33 4 1 --
  +-- Unio_pictorum **
--+                
  +-- Aculamprotula

Huang et al., 2018 [Publ 3828], fig. 3_5

iT tT bi riS oT roT roS iD
1 1 0 1 28 1 1 --
  +-- Unio_pictorum **
--+                
  +-- Aculamprotula

González & Giribet, 2015 [Publ 3632], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 5 0 0 --
-- Unio_pictorum

Dai et al., 2024 [Publ 4386], fig. 5-ML

iT tT bi riS oT roT roS iD
1 1 0 1 20 10 6 --
  +----------------- Aculamprotula   
--+                                  
  |  +-------------- Unio_crassus **    
  +--+                               
     |  +----------- Middendorffinaia
     +--+                            
        |  +-------- Nodularia       
        +--+                         
           |  +----- Cuneopsis       
           +--+                      
              |  +-- Inversiunio     
              +--+                   
                 +-- Schistodesmus   

Wu et al., 2016 [Publ 3711], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 20 4 2 --
  +----- Aculamprotula
--+                   
  |  +-- Unio_pictorum **
  +--+                
     +-- Nodularia    

Wu et al., 2020 [Publ 4147], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 28 9 6 --
  +----------------- Aculamprotula  
--+                                 
  |  +-------------- Unio_pictorum **  
  +--+                              
     |  +----------- Pseudocuneopsis
     +--+                           
        |  +-------- Nodularia      
        +--+                        
           |  +----- Schistodesmus  
           +--+                     
              |  +-- Cuneopsis      
              +--+                  
                 +-- Tchangsinaia   

Bieler et al., 2014 [Publ 3622], fig. 26_27_29_30

iT tT bi riS oT roT roS iD
1 1 0 1 4 0 0 --
-- Unio_pictorum

Choi et al., 2020 [Publ 4169], fig. 3a

iT tT bi riS oT roT roS iD
1 1 0 1 5 5 2 --
  +----- Nodularia    
--+                   
  |  +-- Cuneopsis    
  +--+                
     +-- Unio_pictorum **

Dai et al., 2024 [Publ 4386], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 12 9 5 --
-- Unio_crassus

Liu et al., 2023 [Publ 4310], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 23 13 7 --
  +----------------- Unio_tumidus **   
  |                                 
--+  +-------------- Aculamprotula  
  |  |                              
  +--+  +----------- Cuneopsis      
     |  |                           
     +--+        +-- Nodularia      
        |  +-----+                  
        |  |     +-- Schistodesmus  
        +--+                        
           |  +----- Tchangsinaia   
           +--+                     
              |  +-- Inversiunio    
              +--+                  
                 +-- Pseudocuneopsis

Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2

iT tT bi riS oT roT roS iD
1 1 0 1 10 0 0 --
-- Unio_pictorum

Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M

iT tT bi riS oT roT roS iD
1 1 0 1 6 1 1 --
  +-- Unio_tumidus ** 
--+                
  +-- Aculamprotula

Page updated: 08:38:02 Wed 06 Nov 2024

 
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