|
Bivalvia | Unionoida | Unionoidea | Unionidae subfamily GONIDEINAE Ortmann, 1916: cladomics
8 tribes, 6 genera
Cladomics Summary
number of species = 196
number of subtaxa (rank: tribe) = 14
analyzed in 231 cladograms in 140 publications
maximum number of terminals in an analysis = 74, mean = 10.52
maximum number of subtaxa (tribe) in an analysis = 8, mean = 2.67
Basic Monophyly
number of cladograms that test basic monophyly (i.e., more than 1 included species) = 187
maximum number of included terminals = 74, mean = 12.76
maximum number of subtaxa (tribe) = 8, mean = 3.06
proportion of cladograms that support basic monophyly = 0.77
Rigorous Monophyly
number of cladograms that test rigorous monophyly = 136
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (tribe) taxon and at least one rigorous outgroup (subfamily).
maximum number of included terminals = 74, mean = 11.8
maximum number of subtaxa (tribe) = 8, mean = 3.31
proportion of cladograms that support rigorous monophyly = 0.71
Pairwise Cladogram Clustering
The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Zieritz et al., 2021 [Publ 4218], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 28 |
28 |
26 |
8 |
4 |
2 |
1 |
0.0 |
*** Top Monophyly Cladogram ***
+-- Amblema_plicata
+--------------------------------+
| +-- Lampsilis_ovata
|
| +-------------------------------- Chamberlainia_somsakpanhai **
--+ |
| | +----------------------------- Potomida_littoralis **
| | |
| | | +-- Gonidea_angulata **
+--+ | +--------------------+
| | | +-- Microcondylaea_bonellii **
| | +--+
| | | | +-------------------- Schepmania **
| | | | |
| | | +--+ +-- Indopseudodon_bogani **
+--+ | | +--------------+
| | | | +-- Indopseudodon_crebristriatus **
| | +--+
| | | +-------------- Pilsbryoconcha_exilis **
| | | |
| | +--+ +-- Pseudodon_mekongi **
| | | +--------+
| | | | +-- Pseudodon_vondembuschianus **
| | +--+
+--+ | +-------- Sundadontina **
| +--+
| | +----- Thaiconcha_callifera **
| +--+
| | +-- Bineurus_loeiensis **
| +--+
| +-- Bineurus_anodontinum **
|
| +----- Ctenodesma_borneensis **
| +--------------+
| | | +-- Khairuloconcha_lunbawangorum **
| | +--+
| | +-- Khairuloconcha_sahanae **
| |
+-----+ +-------- Ensidens_sagittarius **
| +--------+
| | | +----- Rectidens_sumatrensis **
| | +--+
| | | +-- Hyriopsis_bialata **
+--+ +--+
| +-- Hyriopsis_desowitzi **
|
| +-------------- Physunio_superbus **
| |
+--+ +----------- Trapezoideus_foliaceus **
| |
+--+ +-- Lens_contradens **
| +-----+
| | +-- Lens_eximius **
+--+
| +----- Yaukthwa_elongatula **
+--+
| +-- Yaukthwa_inlenensis **
+--+
+-- Yaukthwa_paiensis **
Huang et al., 2019 [Publ 3843], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 59 |
84 |
73 |
6 |
145 |
106 |
3 |
82.5 |
*** Top Sister-Group Cladogram ***
+----- Trapezidens_exolescens
+-----------------+
| | +-- Trapezidens_dolichorhynchus
| +--+
+--+ +-- Trapezidens_angustior
| |
| | +-------------------- Lamellidens_corrianus
| | |
| +--+ +----------------- Lamellidens_savadiensis
| | |
| +--+ +-- Lamellidens_lamellatus
| | +-----------+
| | | +-- Lamellidens_jenkinsianus
| +--+
| | +----------- Lamellidens_generosus
| +--+
| | +-------- Lamellidens_mainwaringi[2]
| +--+
+--------------+ | +----- Lamellidens_mainwaringi[1]
| | +--+
| | | +-- Lamellidens_ferrugineus
| | +--+
| | +-- Lamellidens_marginalis
| |
| | +----- Parreysia_keralaensis
| | +-----------------+
| | | | +-- Parreysia_nagpoorensis
| | | +--+
| | | +-- Parreysia_plagiosoma
| | |
| | | +-- Leoparreysia_tavoyensis
| | | |
| +--+ +-----------------+-- Leoparreysia_canefrii
| | | |
| | | +-- Leoparreysia_olivacea
| | |
| | | +-------- Coelatura_aegyptiaca[2]
| | | |
| | | +--------+ +----- Coelatura_aegyptiaca[1]
| +--+ | | |
| | | | +----- Nitia_teretiuscula
| | | +--+
| | | | +-- Coelatura_gabonensis
| | | | |
| | | +--+-- Coelatura_kunenensis
| | | |
| +--+ +-- Prisodontopsis_mweruensis
| |
| | +-------- Indonaia_shurtleffiana
| | |
| | +-----+ +----- Indonaia_bonneaudii
| | | | |
| | | +--+----- Indonaia_rugosa
| | | |
| | | | +-- Indonaia_andersoniana
| +--+ +--+
| | +-- Indonaia_caerulea
| |
| | +----------- Indochinella_pugio
| | |
| | | +----- Scabies_humilis
| +--+ +--+
| | | | +-- Scabies_phaselus
| | | +--+
| +--+ +-- Scabies_mandarinus
| |
| | +----- Radiatula_myitkyinae
| +--+
| | +-- Radiatula_chaudhurii
| +--+
| +-- Radiatula_mouhoti
|
| +-------------------------------- Acuticosta_chinensis
| |
| | +----------------------------- Lepidodesma_languilati
| | |
| | | +-------- Aculamprotula_tortuosa
| | | |
| | | +-----------+ +----- Aculamprotula_coreana
| | | | | |
| | | | +--+----- Aculamprotula_scripta
| | | | |
--+ | | | | +-- Aculamprotula_fibrosa
| | | | +--+
| | | | +-- Aculamprotula_tientsinensis
| +-----+ +--------+
| | | | | +-------- Unio_tumidus
| | | | | +--------+
| | | | | | | +----- Unio_crassus
| | | | | | +--+
| | | | | | | +-- Unio_delphinus
| | | | | | +--+
| | | | +--+ +-- Unio_pictorum
| | | | |
| | | | | +-- Schistodesmus_lampreyanus
| | | | | +-----------+
| | | | | | +-- Schistodesmus_spinosus
| | | | | |
| | | | +--+ +----- Nodularia_douglasiae[1]
| | +--+ | +-----+
| | | | | | +-- Nodularia_douglasiae[2]
| | | | | +--+
| | | +--+ +-- Nodularia_nuxpersicae
| | | |
| | | | +-------- Tchangsinaia_piscicula
| | | +--+
| | | | +----- Cuneopsis_rufescens
| | | +--+
| | | | +-- Cuneopsis_heudei
| | | +--+
| | | +-- Cuneopsis_celtiformis
| | |
| | | +-------- Lanceolaria_gladiola
| | | |
| | | +-----------------+ +----- Lanceolaria_grayii[1]
| | | | | |
| | | | +--+----- Lanceolaria_grayii[2]
| | | | |
| | | | | +-- Lanceolaria_lanceolata
| | | | +--+
| | | | +-- Lanceolaria_triformis
| | | |
| | | | +-------------------- Anemina_arcaeformis
| | | | |
| | | | +--+ +----------------- Cristaria_plicata
| | +--+ | | |
| | | | +--+ +-------------- Pletholophus_tenuis
| | | | | |
| | | | +--+ +-- Sinanodonta_angula
| | | | | +--------+
| | | | | | +-- Sinanodonta_lucida
| | | | +--+
| | | | | +-------- Sinanodonta_woodiana[2]
| | | | +--+
| | | | | +----- Sinanodonta_jourdyi
| | | | +--+
| | +--+ | +-- Sinanodonta_elliptica
| | | +--+
| | | +-- Sinanodonta_woodiana[1]
| | |
| | | +-------- Alasmidonta_marginata
| | | |
| | | +-------- Lasmigona_costata
| | | |
| | | +-------- Pseudanodonta_complanata
| | | |
| | | +-------- Simpsonaias_ambigua
+--+ | |
| +--------------+-------- Strophitus_undulatus
| |
| | +----- Anodonta_anatina
| +--+
| | | +-- Anodonta_cygnea
| | +--+
| | +-- Anodonta_nuttalliana
| |
| | +----- Pyganodon_grandis
| +--+
| | +-- Utterbackia_imbecillis
| +--+
| +-- Utterbackia_peninsularis
|
| +----------- Hyriopsis_desowitzi **
| |
| +----------- Rectidens_sumatrensis **
| |
| | +-- Hyriopsis_bialata **
| | |
| | +-- Hyriopsis_myersiana **
| +-----+--------+
| | | +-- Hyriopsis_panhai **
| | | |
| | | +-- Hyriopsis_khoratensis **
| | |
| | | +-- Ensidens_ingallsianus **
| | | +-----+
| | | | +-- Ensidens_telus **
| | +--+
| | | +----- Ensidens_sagittarius **
| +-----------------+ +--+
| | | | +-- Ensidens_spiculus **
| | | +--+
| | | +-- Ensidens_jaculus **
| | |
| | | +----- Physunio **
| | | +--------+
| | | | | +-- Physunio_modelli **
| | | | +--+
| | | | +-- Physunio_superbus **
| | | |
| | +--+ +-------- Yaukthwa_inlenensis **
| | | +--+
| | | | | +----- Yaukthwa_panhai **
| | | | +--+
| | | | | +-- Yaukthwa_nesemanni **
| | | | +--+
| | +--+ +-- Yaukthwa_rectangularis **
| | |
| | | +-- Lens_contradens **
| | | +-----+
| | | | +-- Lens_micropterus **
| | +--+
| | | +----- Lens_eximius **
| | +--+
| | | +-- Lens_comptus **
| | +--+
| | +-- Lens_novoselovi **
| |
| | +-- Megalonaias_nervosa
| | +--+
+--+ | +-- Uniomerus_carolinianus
| +-----------------------+
| | | +-- Quadrula_quadrula
| | +--+
| | +-- Quadrula_verrucosa
| |
| | +-------- Elliptoideus_sloatianus
| +--+ |
| | | +-----------------+ +-- Elliptio_complanata
| | | | | +--+
| | | | | | +-- Eurynia_dilatata
| | | | +--+
| | | | | +-- Fusconaia_flava
| | +--+ +--+
| | | +-- Pleurobema_sintoxia
| | |
| | | +----------------------- Amblema_plicata
| | | |
| | | | +----- Toxolasma_parvum
| | +--+ +--------------+
| | | | | +-- Obliquaria_reflexa
| | | | +--+
| | | | +-- Plectomerus_dombeyanus
| | +--+
| | | +----------------- Potamilus_leptodon
| | | |
| | | | +-------------- Potamilus_alatus
| | +--+ |
| | | | +----------- Venustaconcha_ellipsiformis
| | | | |
| | +--+ +----------- Leaunio_lienosus
| | | |
+--+ | | +-- Ellipsaria_lineolata
| | +--------+
| +--+ +-- Truncilla_truncata
| |
| | +-- Medionidus_conradicus
| +--------+
| | +-- Ptychobranchus_subtentus
| |
| | +-------- Cambarunio_iris
| +--+
| | +----- Ortmanniana_ligamentina
| +--+
| | +-- Lampsilis_cardium
| +--+
| +-- Lampsilis_ornata
|
| +----- Chamberlainia_somsakpanhai **
| +-----------------------+
| | | +-- Sinohyriopsis_cumingii **
| | +--+
| | +-- Sinohyriopsis_schlegelii **
| |
| | +-- Potomida_littoralis **
| | +-----+
| | | +-- Pronodularia_japanensis **
| | +-----------------+
+--+ | | +----- Lamprotula_leaii **
| | +--+
| | | +-- Lamprotula_caveata **
| | +--+
| | +-- Lamprotula_cornuumlunae **
| |
| | +-------- Sinosolenaia_carinata **
| | +--+
| | | | +----- Sinosolenaia_recognita **
| | | +--+
+--+ | | +-- Sinosolenaia_oleivora **
| | +--+
| +-----------+ +-- Sinosolenaia_iridinea **
| | |
| | | +----- Gonidea_angulata **
| | | +--+
| | | | | +-- Leguminaia_wheatleyi **
| | | | +--+
| | +--+ +-- Microcondylaea_bonellii **
| | |
| | | +-- Ptychorhynchus_pfisteri **
| | | |
+--+ +-----+-- Parvasolenaia_rivularis **
| |
| +-- Koreosolenaia_sitgyensis **
|
| +----- Indopseudodon_salwenianus **
| +--------------+
| | | +-- Indopseudodon_indawgyiensis **
| | +--+
| | +-- Indopseudodon_bogani **
| |
| | +-------- Pilsbryoconcha_exilis[3] **
+--+ +--------+
| | | +----- Pilsbryoconcha_exilis[2] **
| | +--+
| | | +-- Pilsbryoconcha_exilis[1] **
| | +--+
| | +-- Pilsbryoconcha_carinifera **
| |
+--+ +----------- Nyeinchanconcha_nyeinchani **
| +--+
| | | +-------- Thaiconcha_callifera **
| | +--+
| | | +----- Bineurus_loeiensis **
| | +--+
| | | +-- Bineurus_mouhotii **
+--+ +--+
| +-- Bineurus_exilis **
|
| +----- Pseudodon_vagulus **
| +--+
| | | +-- Pseudodon_mekongi **
| | +--+
+-----+ +-- Pseudodon_vondembuschianus **
|
| +-- Sundadontina_cumingii **
| |
+-----+-- Sundadontina_tumida[1] **
|
+-- Sundadontina_tumida[2] **
Bolotov et al., 2020 [Publ 4100], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 74 |
74 |
57 |
6 |
46 |
41 |
1 |
74.5 |
+-------- Trapezidens_exolescens
+--+
| | +----- Trapezidens_scutum
| +--+
| | +-- Trapezidens_dolichorhynchus
+--------------------------+ +--+
| | +-- Trapezidens_angustior
| |
| | +-------- Lamellidens_savadiensis
| +--+
| | +----- Lamellidens_generosus
| +--+
| | +-- Lamellidens_ferrugineus
| +--+
| +-- Lamellidens_mainwaringi
|
| +----------- Parreysia_keralaensis
| +--+
+--+ | | +-------- Parreysia_plagiosoma
| | | +--+
| | | | +----- Parreysia_nagpoorensis[2]
| | | +--+
| | | | +-- Parreysia_nagpoorensis[1]
| | +--------------------+ +--+
| | | | +-- Parreysia_rakhinensis
| | | |
| | | | +-- Leoparreysia_burmana
| | | | +-----+
| | | | | +-- Leoparreysia_whitteni
| | | +-----+
| | | | +----- Leoparreysia_tavoyensis
| | | +--+
| +--+ | +-- Leoparreysia_olivacea
| | +--+
| | +-- Leoparreysia_canefrii
| |
| | +-- Coelatura_aegyptiaca
| | +-----------------------------+
| | | +-- Nitia_teretiuscula
| | |
| | | +----------- Indonaia_shurtleffiana
| | | +-----------------+
| +--+ | | +-------- Indonaia_rugosa
| | | +--+
| | | | +----- Indonaia_andersoniana
| | | +--+
| | | | +-- Indonaia_subclathrata
| +--+ +--+
| | +-- Indonaia_caerulea
| |
| | +-------------------------- Indochinella_pugio
| | |
| | | +-------- Radiatula_crispisulcata
| | | +--------------+
| +--+ | | +----- Radiatula_mouhoti
| | | +--+
| | | | +-- Radiatula_chaudhurii
| | | +--+
| +--+ +-- Radiatula_myitkyinae
| |
| | +-------------------- Scabiellus_songkramensis
| | |
| | | +-- Unionetta_fabagina
| +--+ +--------------+
| | | +-- Harmandia_somboriensis
| | |
| +--+ +-------------- Scabies_nucleus
| | |
| | | +-- Scabies_crispata
| +--+ +--------+
| | | +-- Scabies_mandarinus
| | |
| +--+ +-- Scabies_humilis
| | +-----+
| | | +-- Scabies_pilata
| +--+
| | +----- Scabies_anceps
| +--+
| | +-- Scabies_phaselus
| +--+
--+ +-- Scabies_scobinatus
|
| +----------- Hyriopsis_desowitzi **
| |
| +--+ +-------- Rectidens_sumatrensis **
| | | |
| | +--+ +-- Hyriopsis_khoratensis **
| | | +--+
| | | | +-- Hyriopsis_bialata **
| | +--+
| | | +-- Hyriopsis_myersiana **
| +--------+ +--+
| | | +-- Hyriopsis_panhai **
| | |
| | | +-- Ensidens_telus **
| | | +-----+
| | | | +-- Ensidens_ingallsianus **
| | | |
| | +-----+ +-- Ensidens_jaculus **
| | | +--+
| | | | +-- Ensidens_spiculus **
| | +--+
| | | +-- Ensidens_dugasti **
| | +--+
| | +-- Ensidens_sagittarius **
| +-----+
| | | +-- Trapezoideus_lenya **
| | | +-----+
| | | | +-- Trapezoideus_foliaceus **
| | | +-----------+
| | | | | +----- Physunio_superbus **
| | | | +--+
| | | | | +-- Physunio_modelli **
| | | | +--+
| | | | +-- Physunio **
| | | |
| | | | +-- Yaukthwa_elongatula **
| | | | +-----------+
| | | | | +-- Yaukthwa_baniensis **
| | +--+ |
| | | +--+ +----- Yaukthwa_inlenensis **
| | | | | +-----+
| | | | | | | +-- Yaukthwa_avaensis **
| | | | | | +--+
| | | | +--+ +-- Yaukthwa_paiensis **
| | | | |
| | | | | +-------- Yaukthwa_panhai **
| | | | +--+
| | | | | +----- Yaukthwa_rectangularis **
| | +--+ +--+
| | | | +-- Yaukthwa_zayleymanensis **
| | | +--+
| | | +-- Yaukthwa_nesemanni **
| | |
| | | +----- Lens_misellus **
| | | +-----+
| | | | | +-- Lens_contradens **
| | | | +--+
| | | | +-- Lens_micropterus **
| | +-----+
| | | +----- Lens_pallegoixi **
| | | +--+
+-----------+ | | | +-- Lens_comptus **
| | | +--+
| +--+ +-- Lens_novoselovi **
| |
| | +----- Lens_maenamensis **
| +--+
| | +-- Lens_eximius **
| +--+
| +-- Lens_rolfbrandti **
|
| +-- Sinosolenaia_iridinea **
| |
| +-- Sinosolenaia_recognita[1] **
| |
| +-- Sinosolenaia_carinata **
| |
| +-- Leguminaia_wheatleyi **
| |
| +-- Microcondylaea_bonellii **
| |
| +-- Gonidea_angulata **
| |
| +-- Ptychorhynchus_pfisteri **
| |
| +-- Parvasolenaia_rivularis **
| |
| +-----------------------+-- Lamprotula_leaii **
| | |
| | +-- Lamprotula **
| | |
| | +-- Lamprotula_caveata **
| | |
| | +-- Potomida_littoralis **
| | |
| | +-- Pronodularia_japanensis **
| | |
| | +-- Sinohyriopsis_cumingii **
| | |
+--+ +-- Chamberlainia_somsakpanhai **
| |
| +-- Sinosolenaia_recognita[2] **
| |
| +-- Sinosolenaia_oleivora **
|
| +-------- Indopseudodon_kayinensis **
| +--------------+
| | | +----- Indopseudodon_indawgyiensis **
| | +--+
| | | +-- Indopseudodon_salwenianus **
| | +--+
| | +-- Indopseudodon_bogani **
+--+
| +----- Bineurus_loeiensis **
| +--------------+
| | | +-- Bineurus_exilis **
| | +--+
| | +-- Bineurus_mouhotii **
| |
+--+ +----------------- Nyeinchanconcha_nyeinchani **
| |
| | +-- Thaiconcha_callifera **
| | +-----+
| | | +-- Thaiconcha_munelliptica **
+--+ +-----+
| | | +----- Pilsbryoconcha_exilis[2] **
| | +--+
| | | +-- Pilsbryoconcha_exilis[1] **
| | +--+
| | +-- Pilsbryoconcha **
+--+
| +-------- Pseudodon_lenyanensis **
| +--+
| | | +----- Pseudodon_vagulus **
| | +--+
| | | +-- Pseudodon_vondembuschianus **
| | +--+
+--+ +-- Pseudodon_mekongi **
|
| +-- Sundadontina_taskaevi **
| +-----+
| | +-- Sundadontina_brandti **
+--+
| +----- Sundadontina_tumida **
+--+
| +-- Sundadontina_cumingii **
+--+
+-- Sundadontina_tanintharyiensis **
Bolotov et al., 2020 [Publ 4100], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 74 |
74 |
61 |
6 |
40 |
40 |
1 |
72.5 |
+-------- Trapezidens_exolescens
+--+
| | +----- Trapezidens_scutum
| +--+
| | +-- Trapezidens_dolichorhynchus
+--------------------------------+ +--+
| | +-- Trapezidens_angustior
| |
| | +-------- Lamellidens_savadiensis
| +--+
| | +----- Lamellidens_generosus
| +--+
| | +-- Lamellidens_ferrugineus
| +--+
| +-- Lamellidens_mainwaringi
|
| +----------- Parreysia_keralaensis
| +--+
+--+ | | +-------- Parreysia_nagpoorensis[2]
| | | +--+
| | | | +----- Parreysia_nagpoorensis[1]
| | | +--+
| | | | +-- Parreysia_rakhinensis
| | +--------------------------+ +--+
| | | | +-- Parreysia_plagiosoma
| | | |
| | | | +-- Leoparreysia_burmana
| | | | +-----+
| | | | | +-- Leoparreysia_whitteni
| | | +-----+
| | | | +----- Leoparreysia_olivacea
| | | +--+
| +--+ | +-- Leoparreysia_tavoyensis
| | +--+
| | +-- Leoparreysia_canefrii
| |
| | +-- Coelatura_aegyptiaca
| | +-----------------------------------+
| | | +-- Nitia_teretiuscula
| | |
| | | +-------- Indonaia_shurtleffiana
| +--+ +--------------------------+
| | | | +----- Indonaia_andersoniana
| | | +--+
| | | | +-- Indonaia_caerulea
| | | +--+
| +--+ +-- Indonaia_subclathrata
| |
| | +-------------------------------- Indochinella_pugio
| | |
| | | +----- Radiatula_mouhoti
| +--+ +-----------------------+
| | | | +-- Radiatula_myitkyinae
| | | +--+
| +--+ +-- Radiatula_chaudhurii
| |
| | +-------------------------- Radiatula_crispisulcata
| | |
| +--+ +----------------------- Scabiellus_songkramensis
| | |
| +--+ +-------------------- Harmandia_somboriensis
| | |
| +--+ +----------------- Unionetta_fabagina
| | |
| +--+ +-------------- Scabies_mandarinus
| | |
| +--+ +----------- Scabies_crispata
| | |
| +--+ +-------- Scabies_nucleus
| | |
| +--+ +-- Scabies_humilis
| | +--+
| | | +-- Scabies_pilata
| +--+
| | +-- Scabies_phaselus
| | |
| +--+-- Scabies_scobinatus
| |
--+ +-- Scabies_anceps
|
| +-------------- Hyriopsis_desowitzi **
| +--+
| | | +----------- Rectidens_sumatrensis **
| | +--+
| | | +-------- Hyriopsis_panhai **
| | +--+
| | | +----- Hyriopsis_myersiana **
| | +--+
| +--------+ | +-- Hyriopsis_khoratensis **
| | | +--+
| | | +-- Hyriopsis_bialata **
| | |
| | | +-- Ensidens_telus **
| | | +-----+
| | | | +-- Ensidens_ingallsianus **
| | | |
| | +--------+ +-- Ensidens_dugasti **
| | | +--+
| | | | +-- Ensidens_sagittarius **
| | +--+
| +--+ | +-- Ensidens_spiculus **
| | | +--+
| | | +-- Ensidens_jaculus **
| | |
| | | +----- Physunio **
| | | +-----------------+
| | | | | +-- Physunio_modelli **
| | | | +--+
| | | | +-- Physunio_superbus **
| | | |
| | | | +-- Trapezoideus_lenya **
| | | | +-----------------+
| | +--+ | +-- Trapezoideus_foliaceus **
| | | |
| | | | +----- Lens_misellus **
| | | | +-----+
| | | | | | +-- Lens_contradens **
| | | | | +--+
| | | | | +-- Lens_micropterus **
| | +--+ +-----+
| | | | | +----- Lens_maenamensis **
| | | | | +--+
| | | | | | | +-- Lens_eximius **
| | | | | | +--+
| | | | +--+ +-- Lens_rolfbrandti **
| | | | |
| | | | | +----- Lens_pallegoixi **
| | +--+ +--+
| | | | +-- Lens_novoselovi **
| | | +--+
| | | +-- Lens_comptus **
| | |
| | | +-- Yaukthwa_elongatula **
| | | +-----------+
| | | | +-- Yaukthwa_baniensis **
| | | |
| | +--+ +----- Yaukthwa_inlenensis **
| | | +-----+
+-----------------+ | | | +-- Yaukthwa_avaensis **
| | | +--+
| +--+ +-- Yaukthwa_paiensis **
| |
| | +-------- Yaukthwa_panhai **
| +--+
| | +----- Yaukthwa_rectangularis **
| +--+
| | +-- Yaukthwa_zayleymanensis **
| +--+
| +-- Yaukthwa_nesemanni **
|
| +-- Sinohyriopsis_cumingii **
| +--+
| | +-- Chamberlainia_somsakpanhai **
| |
| +--+ +-- Lamprotula_leaii **
| | | |
| | | +-- Lamprotula **
| | | |
| | +--+-- Lamprotula_caveata **
| | |
| | +-- Potomida_littoralis **
| | |
| | +-- Pronodularia_japanensis **
| +-----------------+
| | | +-- Sinosolenaia_iridinea **
| | | |
| | | +-- Sinosolenaia_recognita[1] **
| | | |
| | | +-- Sinosolenaia_carinata **
| | | |
| | | +-- Leguminaia_wheatleyi **
| | | |
| | | +-- Microcondylaea_bonellii **
| | +-----+
| | +-- Gonidea_angulata **
| | |
| | +-- Ptychorhynchus_pfisteri **
+--+ |
| +-- Parvasolenaia_rivularis **
| |
| +-- Sinosolenaia_recognita[2] **
| |
| +-- Sinosolenaia_oleivora **
|
| +-------- Indopseudodon_kayinensis **
| +--------------+
| | | +----- Indopseudodon_indawgyiensis **
| | +--+
| | | +-- Indopseudodon_salwenianus **
| | +--+
| | +-- Indopseudodon_bogani **
+--+
| +----- Bineurus_loeiensis **
| +--------------+
| | | +-- Bineurus_exilis **
| | +--+
| | +-- Bineurus_mouhotii **
| |
+--+ +----------------- Nyeinchanconcha_nyeinchani **
| |
| | +-- Thaiconcha_callifera **
| | +-----+
| | | +-- Thaiconcha_munelliptica **
+--+ +-----+
| | | +----- Pilsbryoconcha_exilis[2] **
| | +--+
| | | +-- Pilsbryoconcha_exilis[1] **
| | +--+
| | +-- Pilsbryoconcha **
+--+
| +-------- Pseudodon_lenyanensis **
| +--+
| | | +----- Pseudodon_vagulus **
| | +--+
| | | +-- Pseudodon_vondembuschianus **
| | +--+
+--+ +-- Pseudodon_mekongi **
|
| +-- Sundadontina_tanintharyiensis **
| +-----+
| | +-- Sundadontina_cumingii **
+--+
| +----- Sundadontina_tumida **
+--+
| +-- Sundadontina_taskaevi **
+--+
+-- Sundadontina_brandti **
Konopleva et al., 2023 [Publ 4262], fig. 8
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 70 |
70 |
68 |
5 |
14 |
14 |
1 |
69.5 |
+----------------------- Indonaia_andersoniana
|
+--------------+ +-------------------- Indochinella_pugio
| | |
| +--+ +----------------- Radiatula_mouhoti
| | |
| +--+ +-------------- Scabiellus_songkramensis
| | |
| | | +-- Unionetta_fabagina
| +--+ +--------+
| | | +-- Harmandia_somboriensis
| | |
| | | +-- Scabies_humilis
| +--+ +-----+
| | | +-- Scabies_pilata
| | |
| | | +----- Scabies_crispata
| | +--+
| +--+ | +-- Scabies_nucleus
| | +--+
| | +-- Scabies_mandarinus
| |
| | +-- Scabies_phaselus
| | |
| +-----+-- Scabies_anceps
| |
| +-- Scabies_scobinatus
|
| +-------- Indopseudodon_kayinensis **
| +-----------------------+
| | | +----- Indopseudodon_indawgyiensis **
| | +--+
| | | +-- Indopseudodon_salwenianus **
| | +--+
| | +-- Indopseudodon_bogani **
| |
| | +-- Pilsbryoconcha_schomburgki **
--+ +--+ +--------+
| | | | +-- Pilsbryoconcha_exilis[1] **
| | | +-----------------+
| | | | | +-------- Pilsbryoconcha **
| | | | +--+
| | | | | +----- Pilsbryoconcha_linguaeformis **
| | | | +--+
| | | | | +-- Pilsbryoconcha_exilis[2] **
| | +--+ +--+
| | | +-- Pilsbryoconcha_carinifera **
| | |
| | | +-------------------------- Songkhlanaia_tamodienica **
| | | |
| | | | +-- Namkongnaia_inkhavilayi **
| | | | +--------------------+
| | +--+ | +-- Namkongnaia_lemeslei **
| | | |
| | | | +-------- Pseudodon_lenyanensis **
| | | | +-----------+
| | | | | | +----- Pseudodon_vagulus **
| | +--+ | +--+
| | | | | +-- Pseudodon_vondembuschianus **
| | | | +--+
| | | | +-- Pseudodon_mekongi **
| | | |
| | | | +-- Sundadontina_taskaevi **
| | +--+ +--------+
| | | | +-- Sundadontina_brandti **
| | | +-----+
| | | | | +-------- Sundadontina_tumida **
| | | | +--+
| | | | | +----- Sundadontina_tanintharyiensis **
| | | | +--+
| | | | | +-- Sundadontina_plugpomenica **
| | +--+ +--+
| | | +-- Sundadontina_cumingii **
+--+ |
| | +-------------- Nyeinchanconcha_nyeinchani **
| | |
| | | +----- Thaiconcha_thaiensis **
| +--+ +-----+
| | | | +-- Thaiconcha_munelliptica **
| | | +--+
| +--+ +-- Thaiconcha_callifera **
| |
| | +-------- Bineurus_loeiensis **
| +--+
| | +----- Bineurus_mouhotii **
| +--+
| | +-- Bineurus_anodontinum **
| +--+
| +-- Bineurus_exilis **
|
| +-- Gonidea_angulata **
| +--+
| | +-- Leguminaia_wheatleyi **
| +-----------------------+
| | | +-- Lamprotula_leaii **
| | +--+
| | +-- Potomida_littoralis **
| |
| | +-- Ensidens_ingallsianus **
| | +-----+
| | | +-- Ensidens_telus **
| | |
| | +--------------+ +-- Ensidens_jaculus **
| | | | +--+
| | | | | +-- Ensidens_spiculus[1] **
| | | +--+
| | | | +-- Ensidens_spiculus[2] **
| | +--+ +--+
| | | | +-- Ensidens_dugasti **
+-----+ | |
| | | +-- Hyriopsis_desowitzi **
| | | +-----------------+
| | | | +-- Hyriopsis_delaportei **
| | +--+
| | | +----------------- Rectidens_sumatrensis **
| | | |
| | +--+ +-------------- Hyriopsis_panhai **
| | | |
| | +--+ +----------- Hyriopsis_myersiana **
| | | |
| | +--+ +-- Hyriopsis_khoratensis **
| | | +-----+
| | | | +-- Hyriopsis_bialata **
| | +--+
| | | +----- Hyriopsis_sakhonensis **
+--+ +--+
| | +-- Hyriopsis_phuphaniensis **
| +--+
| +-- Hyriopsis_kratiensis **
|
| +----- Physunio_modelli **
| +-----+
| | | +-- Physunio_superbus **
| | +--+
| | +-- Physunio_massini **
| +-----+
| | | +-- Trapezoideus_thachiadensis **
| | | +-----+
| | | | +-- Trapezoideus_subviridis **
| | +--+
| | | +----- Trapezoideus_mitanensis **
| | +--+
| | | +-- Trapezoideus_foliaceus **
+--------+ +--+
| +-- Trapezoideus_lenya **
|
| +-- Yaukthwa_nesemanni **
| +-----------+
| | +-- Yaukthwa_elongatula **
| |
| | +----- Lens_misellus **
+--+ +-----+
| | | +-- Lens_contradens **
| | +--+
| | +-- Lens_micropterus **
+--+
| +----- Lens_pallegoixi **
| +--+
| | | +-- Lens_novoselovi **
| | +--+
+--+ +-- Lens_comptus **
|
| +----- Lens_maenamensis **
+--+
| +-- Lens_eximius **
+--+
+-- Lens_rolfbrandti **
Huang et al., 2019 [Publ 3843], fig. S3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 58 |
126 |
123 |
6 |
143 |
104 |
3 |
124.5 |
+----- Trapezidens_exolescens
+-----+
| | +-- Trapezidens_angustior
| +--+
| +-- Trapezidens_dolichorhynchus
|
+--------------+ +-- Lamellidens_corrianus
| | +-----+
| | | +-- Lamellidens_savadiensis
| | |
| | | +----- Lamellidens_lamellatus
| +--+ |
| | +----- Lamellidens_jenkinsianus
| | |
| | | +-- Lamellidens_marginalis
| +--+--+
| | +-- Lamellidens_ferrugineus
| |
+-----------------------------+ | +-- Lamellidens_mainwaringi[1]
| | | |
| | +--+-- Lamellidens_mainwaringi[2]
| | |
| | +-- Lamellidens_generosus
| |
| | +----- Parreysia_keralaensis
| | +-----------------+
| | | | +-- Parreysia_nagpoorensis
| | | +--+
| | | +-- Parreysia_plagiosoma
| | |
| | | +----- Leoparreysia_canefrii
| +--+ +--------------+
| | | | +-- Leoparreysia_tavoyensis
| | | +--+
| | | +-- Leoparreysia_olivacea
| | |
| | | +----------- Coelatura_aegyptiaca[2]
| | | |
| +--+ +-----+ +-- Coelatura_aegyptiaca[1]
| | | | +-----+
| | | | | +-- Nitia_teretiuscula
| | | +--+
| | | | +----- Coelatura_kunenensis
| | | +--+
| | | | +-- Coelatura_gabonensis
| | | +--+
| +--+ +-- Prisodontopsis_mweruensis
| |
| | +-------- Indonaia_shurtleffiana
| | |
| | +-----+ +----- Indonaia_bonneaudii
| | | | |
| | | +--+----- Indonaia_rugosa
| | | |
| | | | +-- Indonaia_andersoniana
| +--+ +--+
| | +-- Indonaia_caerulea
| |
| | +----------- Indochinella_pugio
| | |
| | | +----- Scabies_phaselus
| +--+ +--+
| | | | +-- Scabies_humilis
| | | +--+
| +--+ +-- Scabies_mandarinus
| |
| | +----- Radiatula_mouhoti
| +--+
| | +-- Radiatula_myitkyinae
| +--+
--+ +-- Radiatula_chaudhurii
|
| +----- Indopseudodon_indawgyiensis **
| +--------------+
| | | +-- Indopseudodon_bogani **
| | +--+
| | +-- Indopseudodon_salwenianus **
| |
| | +-------- Pilsbryoconcha_exilis[3] **
| +--------------------------------+ +--------+
| | | | | +----- Pilsbryoconcha_exilis[2] **
| | | | +--+
| | | | | +-- Pilsbryoconcha_carinifera **
| | | | +--+
| | | | +-- Pilsbryoconcha_exilis[1] **
| | | |
| | +--+ +----------- Sundadontina_tumida **
| | | +--+
| | | | | +-------- Pseudodon_vagulus **
| | | | +--+
| | | | | +----- Pseudodon_vondembuschianus[1] **
| | | | +--+
| | | | | +-- Pseudodon_mekongi **
| | +--+ +--+
| | | +-- Pseudodon_vondembuschianus[2] **
| | |
| | | +-- Nyeinchanconcha_nyeinchani **
| | | +--------+
| | | | +-- Sundadontina_cumingii **
| | +--+
| | | +-------- Thaiconcha_callifera **
| | +--+
| | | +----- Bineurus_loeiensis **
| | +--+
| | | +-- Bineurus_mouhotii **
| | +--+
| | +-- Bineurus_exilis **
| |
| | +-- Ensidens_telus **
| | +-----+
| | | +-- Ensidens_ingallsianus **
| | +-----+
| | | | +----- Ensidens_sagittarius **
| | | +--+
| | | | +-- Ensidens_spiculus **
| | +--+ +--+
| | | | +-- Ensidens_jaculus **
| | | |
| | | | +----------- Hyriopsis_desowitzi **
| | | | |
| | | +--+ +-------- Rectidens_sumatrensis **
| | | | |
| | | +--+ +-- Hyriopsis_panhai **
+--+ | | +--+
| | | | +-- Hyriopsis_khoratensis **
| +--------------------------------+ +--+
| | | | +-- Hyriopsis_bialata **
| | | +--+
| | | +-- Hyriopsis_myersiana **
| | |
| | | +----- Physunio **
| | | +--------+
| | | | | +-- Physunio_superbus **
| | | | +--+
| | | | +-- Physunio_modelli **
| | | |
| | +--+ +-------- Yaukthwa_inlenensis **
| | | +--+
| | | | | +----- Yaukthwa_panhai **
| | | | +--+
| | | | | +-- Yaukthwa_nesemanni **
| | | | +--+
| | +--+ +-- Yaukthwa_rectangularis **
| | |
| | | +-- Lens_contradens **
| | | +-----+
| | | | +-- Lens_micropterus **
| | +--+
| | | +----- Lens_eximius **
| | +--+
| | | +-- Lens_novoselovi **
| | +--+
| | +-- Lens_comptus **
| |
| | +----- Chamberlainia_somsakpanhai **
| | +--------------------------------------+
| | | | +-- Sinohyriopsis_cumingii **
| | | +--+
| | | +-- Sinohyriopsis_schlegelii **
| | |
| | | +-- Potomida_littoralis **
| | | +-----+
| | | | +-- Pronodularia_japanensis **
| | | +-----+
| | | | | +----- Lamprotula_leaii **
| | | | +--+
| | | | | +-- Lamprotula_caveata **
+--+ | | +--+
| | | +-- Lamprotula_cornuumlunae **
| +--+ +--------------------------+
| | | | | +----- Sinosolenaia_recognita **
| | | | | +-----+
| | | | | | | +-- Sinosolenaia_oleivora **
| | | | | | +--+
| | | | | | +-- Sinosolenaia_iridinea **
| | | | +--+
| | | | | +----- Gonidea_angulata **
| | | | | +--+
| | | | | | | +-- Leguminaia_wheatleyi **
| | | | | | +--+
| | | | +--+ +-- Microcondylaea_bonellii **
| | | | |
| | | | | +----- Ptychorhynchus_pfisteri **
| | +--+ +--+
| | | | +-- Parvasolenaia_rivularis **
| | | +--+
| | | +-- Koreosolenaia_sitgyensis **
| | |
| | | +-- Megalonaias_nervosa
| | | +--+
| | | | +-- Uniomerus_carolinianus
| | | +--------------------------------+
| | | | | +-- Quadrula_quadrula
| | | | +--+
| | | | +-- Quadrula_verrucosa
| | | |
| | | | +-------- Elliptoideus_sloatianus
| | +--+ |
| | | +--------------------------+ +-- Elliptio_complanata
| | | | | +--+
| | | | | | +-- Eurynia_dilatata
| | | | +--+
| | | | | +-- Fusconaia_flava
| | +--+ +--+
| | | +-- Pleurobema_sintoxia
| | |
| | | +-------------------------------- Amblema_plicata
| | | |
| | +--+ +----------------------------- Toxolasma_parvum
| | | |
+--+ +--+ +-- Obliquaria_reflexa
| | +-----------------------+
| | | +-- Plectomerus_dombeyanus
| +--+
| | +----------------------- Potamilus_leptodon
| | |
| +--+ +-------------------- Potamilus_alatus
| | |
| | | +-- Medionidus_conradicus
| +--+ +--------------+
| | | +-- Ptychobranchus_subtentus
| | |
| +--+ +-- Ellipsaria_lineolata
| | +-----------+
| | | +-- Truncilla_truncata
| +--+
| | +----------- Leaunio_lienosus
| | |
| +--+ +-- Venustaconcha_ellipsiformis
| | +-----+
| | | +-- Cambarunio_iris
| +--+
| | +----- Ortmanniana_ligamentina
| +--+
| | +-- Lampsilis_cardium
| +--+
| +-- Lampsilis_ornata
|
| +----------------------------------- Acuticosta_chinensis
| |
| | +-------- Aculamprotula_tortuosa
| | |
| | +-----------+ +-- Aculamprotula_coreana
| | | | +--+
| | | | | +-- Aculamprotula_scripta
| | | +--+
| | | | +-- Aculamprotula_fibrosa
| | | +--+
| | | +-- Aculamprotula_tientsinensis
| | +-----------+
| | | | +-------- Unio_tumidus
+-----------+ | | +--------+
| | | | | +----- Unio_crassus
| | | | +--+
| | | | | +-- Unio_delphinus
| | | | +--+
| | +--+ +-- Unio_pictorum
| | |
| | | +----- Nodularia_douglasiae[1]
| | | +--------+
| | | | | +-- Nodularia_douglasiae[2]
| | | | +--+
| | +--+ +-- Nodularia_nuxpersicae
| | |
+--+ | +----------- Schistodesmus_lampreyanus
| +--+
| | +-------- Tchangsinaia_piscicula
| +--+
| | +----- Cuneopsis_rufescens
| +--+
| | +-- Cuneopsis_heudei
| +--+
| +-- Cuneopsis_celtiformis
|
| +----------------------------- Lepidodesma_languilati
| |
| | +-------- Lanceolaria_gladiola
| | +-----------------+
| | | | +----- Lanceolaria_grayii
| | | +--+
+--+ | | +-- Lanceolaria_lanceolata
| | +--+
| | +-- Lanceolaria_triformis
| |
| | +-------------------- Anemina_arcaeformis
| | |
| | +--+ +----------------- Cristaria_plicata
+--+ | | |
| | +--+ +-------------- Pletholophus_tenuis
| | | |
| | +--+ +-- Sinanodonta_angula
| | | +--------+
| | | | +-- Sinanodonta_lucida
| | +--+
| | | +-------- Sinanodonta_woodiana[2]
| | +--+
| | | +----- Sinanodonta_jourdyi
+--+ +--+
| | +-- Sinanodonta_elliptica
| +--+
| +-- Sinanodonta_woodiana[1]
|
| +-- Alasmidonta_marginata
| +--------+
| | +-- Lasmigona_costata
| |
| | +----- Anodonta_anatina
| +-----+
| | | +-- Anodonta_cygnea
+-----------+ +--+
| +-- Anodonta_nuttalliana
|
| +----- Pseudanodonta_complanata
| +--+
| | | +-- Simpsonaias_ambigua
| | +--+
+--+ +-- Strophitus_undulatus
|
| +----- Pyganodon_grandis
+--+
| +-- Utterbackia_imbecillis
+--+
+-- Utterbackia_peninsularis
Huang et al., 2019 [Publ 3843], fig. S4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 58 |
127 |
70 |
6 |
144 |
105 |
3 |
127.5 |
+-- Parreysia_keralaensis
|
+--------------------+-- Parreysia_nagpoorensis
| |
| +-- Parreysia_plagiosoma
|
| +----- Trapezidens_exolescens
| +--------+
| | | +-- Trapezidens_angustior
| | +--+
+--------+ +-- Trapezidens_dolichorhynchus
| |
| | +----------- Lamellidens_corrianus
| | |
| +--+ +-------- Lamellidens_savadiensis
| | |
| | | +----- Lamellidens_marginalis
| +--+ |
| | +----- Lamellidens_ferrugineus
| | |
+--+ +--+ +-- Lamellidens_mainwaringi[1]
| | | |
| | | +-- Lamellidens_mainwaringi[2]
| | | |
| | +--+-- Lamellidens_lamellatus
| | |
| | +-- Lamellidens_generosus
| | |
| | +-- Lamellidens_jenkinsianus
| |
| | +----- Leoparreysia_tavoyensis
| | +--------------+
| | | | +-- Leoparreysia_canefrii
| | | +--+
| | | +-- Leoparreysia_olivacea
| | |
| | | +-------- Coelatura_aegyptiaca[2]
| | | |
| | | +-------- Nitia_teretiuscula
| +--+ +--------+
| | | | +----- Coelatura_aegyptiaca[1]
| | | | |
| | | +--+ +-- Coelatura_gabonensis
| | | | |
| | | +--+-- Coelatura_kunenensis
| | | |
| | | +-- Prisodontopsis_mweruensis
| +--+
| | +----- Indonaia_shurtleffiana
| | |
| | | +-- Indonaia_bonneaudii
| | +--------+--+
| | | | +-- Indonaia_rugosa
| | | |
| | | | +-- Indonaia_andersoniana
| | | +--+
| +--+ +-- Indonaia_caerulea
| |
| | +----------- Indochinella_pugio
| | |
| | | +----- Scabies_humilis
| +--+ +--+
| | | | +-- Scabies_phaselus
| | | +--+
| +--+ +-- Scabies_mandarinus
| |
| | +----- Radiatula_mouhoti
| +--+
| | +-- Radiatula_myitkyinae
| +--+
| +-- Radiatula_chaudhurii
|
| +-------------------- Chamberlainia_somsakpanhai **
| |
| +-------------------- Megalonaias_nervosa
| |
| +-------------------- Uniomerus_carolinianus
| |
| | +-- Quadrula_quadrula
| +-----------------+
| | +-- Quadrula_verrucosa
| |
| | +-- Sinohyriopsis_cumingii **
| +-----------------+
| | +-- Sinohyriopsis_schlegelii **
| |
| | +-- Potomida_littoralis **
| | +-----+
| | | +-- Pronodularia_japanensis **
| +-----------+
| | | +----- Lamprotula_leaii **
| | +--+
| | | +-- Lamprotula_caveata **
| | +--+
| | +-- Lamprotula_cornuumlunae **
| |
| | +-------- Sinosolenaia_carinata **
--+ | |
| | | +----- Gonidea_angulata **
| | +--+
| | | | +-- Leguminaia_wheatleyi **
| | | +--+
| | | +-- Microcondylaea_bonellii **
| +-----------+
| | | +----- Ptychorhynchus_pfisteri **
| | +--+
| | | | +-- Parvasolenaia_rivularis **
| | | +--+
| | | +-- Koreosolenaia_sitgyensis **
| | |
| | | +----- Sinosolenaia_recognita **
| | +--+
| | | +-- Sinosolenaia_iridinea **
| | +--+
| | +-- Sinosolenaia_oleivora **
| |
| | +----- Indopseudodon_salwenianus **
| | +-----+
| | | | +-- Indopseudodon_indawgyiensis **
| | | +--+
| | | +-- Indopseudodon_bogani **
| | |
| | | +-------- Nyeinchanconcha_nyeinchani **
| | | |
| | | +-------- Thaiconcha_callifera **
| +--------+ |
| | | +-------- Sundadontina_cumingii **
| | | |
| | | +-------- Sundadontina_tumida[1] **
| | | |
| | | +-------- Sundadontina_tumida[2] **
| | | |
| | | | +----- Pilsbryoconcha_exilis[2] **
| | +--+--+
| | | | +-- Pilsbryoconcha_carinifera **
| | | +--+
| | | +-- Pilsbryoconcha_exilis[1] **
| | |
| | | +----- Pseudodon_vagulus **
| | +--+
| | | | +-- Pseudodon_mekongi **
| | | +--+
| | | +-- Pseudodon_vondembuschianus **
| | |
| | | +----- Bineurus_loeiensis **
| | +--+
| | | +-- Bineurus_mouhotii **
| | +--+
| | +-- Bineurus_exilis **
| |
| | +-------------- Amblema_plicata
| | |
| | +-------------- Toxolasma_parvum
| | |
| | | +-- Obliquaria_reflexa
| | +-----------+
| | | +-- Plectomerus_dombeyanus
| | |
| | | +----------- Elliptoideus_sloatianus
| | +--+
| | | | +-------- Elliptio_complanata
| | | +--+
| | | | +----- Eurynia_dilatata
| +-----+ +--+
| | | | +-- Fusconaia_flava
| | | +--+
| | | +-- Pleurobema_sintoxia
| | |
| | | +-------- Ellipsaria_lineolata
| | | |
| | | +-------- Potamilus_leptodon
| | | |
| | | +-------- Medionidus_conradicus
| | | |
+-----+ | +-------- Potamilus_alatus
| | |
| | +-------- Ptychobranchus_subtentus
| +-----+
| +-------- Truncilla_truncata
| |
| +-------- Venustaconcha_ellipsiformis
| |
| +-------- Cambarunio_iris
| |
| +-------- Leaunio_lienosus
| |
| | +----- Ortmanniana_ligamentina
| +--+
| | +-- Lampsilis_cardium
| +--+
| +-- Lampsilis_ornata
|
| +-------------- Hyriopsis_desowitzi **
| |
| +-------------- Hyriopsis_myersiana **
| |
| +-------------- Hyriopsis_panhai **
| |
| +--+ +-- Hyriopsis_bialata **
| | +-----------+
| | | +-- Hyriopsis_khoratensis **
| | |
| | | +----------- Rectidens_sumatrensis **
| | | |
| | +--+ +-- Ensidens_telus **
| | | +-----+
| | | | +-- Ensidens_ingallsianus **
| | +--+
| | | +----- Ensidens_sagittarius **
| | +--+
+--+ | +-- Ensidens_spiculus **
| | +--+
| | +-- Ensidens_jaculus **
| |
| | +----------- Physunio_modelli **
| | |
| | +----------- Lens **
| | |
| | +----------- Physunio_superbus **
| | |
| | | +-------- Yaukthwa_inlenensis **
| | +--+
| +-----+ | +----- Yaukthwa_panhai **
| | +--+
| | | +-- Yaukthwa_nesemanni **
| | +--+
| | +-- Yaukthwa_rectangularis **
| |
| | +-- Lens_comptus **
| | +-----+
| | | +-- Lens_novoselovi **
| +--+
| | +----- Lens_eximius **
| +--+
| | +-- Lens_contradens **
| +--+
| +-- Lens_micropterus **
|
| +----------------- Acuticosta_chinensis
| |
| +----------------- Alasmidonta_marginata
| |
| +----------------- Anodonta_anatina
| |
| +----------------- Anemina_arcaeformis
| |
| +----------------- Anodonta_cygnea
| |
| +----------------- Cristaria_plicata
| |
| +----------------- Anodonta_nuttalliana
| |
| +----------------- Lasmigona_costata
| |
| +----------------- Lepidodesma_languilati
| |
| +----------------- Pseudanodonta_complanata
| |
| +----------------- Pyganodon_grandis
| |
| +----------------- Simpsonaias_ambigua
| |
| +----------------- Strophitus_undulatus
| |
| | +-- Schistodesmus_lampreyanus
| +--------------+
| | +-- Schistodesmus_spinosus
| |
| | +-- Utterbackia_imbecillis
| +--------------+
| | +-- Utterbackia_peninsularis
| |
| | +-------- Lanceolaria_gladiola
| +--------+
| | | +----- Lanceolaria_grayii
| | +--+
+--+ | +-- Lanceolaria_lanceolata
| +--+
| +-- Lanceolaria_triformis
|
| +-------- Unio_tumidus
+--------+
| | +----- Unio_crassus
| +--+
| | +-- Unio_delphinus
| +--+
| +-- Unio_pictorum
|
| +-------- Aculamprotula_tortuosa
| |
+--------+ +-- Aculamprotula_coreana
| | +--+
| | | +-- Aculamprotula_scripta
| +--+
| | +-- Aculamprotula_fibrosa
| +--+
| +-- Aculamprotula_tientsinensis
|
| +-------- Cuneopsis_heudei
| |
| +-------- Cuneopsis_celtiformis
| |
+--------+-------- Tchangsinaia_piscicula
| |
| +-------- Cuneopsis_rufescens
| |
| | +----- Nodularia_douglasiae[2]
| +--+
| | +-- Nodularia_douglasiae[1]
| +--+
| +-- Nodularia_nuxpersicae
|
| +-------------- Pletholophus_tenuis
| |
+--+ +-- Sinanodonta_angula
| +--------+
| | +-- Sinanodonta_lucida
+--+
| +-------- Sinanodonta_woodiana[1]
+--+
| +----- Sinanodonta_jourdyi
+--+
| +-- Sinanodonta_elliptica
+--+
+-- Sinanodonta_woodiana[2]
Bolotov et al., 2017 [Publ 3764], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 53 |
90 |
42 |
6 |
78 |
78 |
3 |
91.5 |
+----- Trapezidens_exolescens
+--------+
| | +-- Trapezidens_dolichorhynchus
| +--+
+-----------+ +-- Trapezidens_angustior
| |
| | +----------- Lamellidens_lamellatus
| | |
| +--+ +-------- Lamellidens_jenkinsianus
| | |
| | | +----- Lamellidens_generosus
| +--+ |
| | +----- Lamellidens_mainwaringi
| | |
| +--+ +-- Lamellidens_ferrugineus
| +--+
| | +-- Lamellidens_marginalis
| |
+--+ | +-- Lamellidens_savadiensis
| | +--+
| | +-- Lamellidens_corrianus[1]
| |
| | +-------- Parreysia_keralaensis
| | +--------------+
| | | | +----- Parreysia_plagiosoma
| | | +--+
| | | | +-- Parreysia_corrugata
| | | +--+
| | | +-- Parreysia_nagpoorensis
| | |
| | | +-------- Leoparreysia_olivacea
| | | +-----------+
| | | | | +----- Leoparreysia_tavoyensis[2]
| +--+ | +--+
| | | | +-- Leoparreysia_canefrii
| | | +--+
| | | +-- Leoparreysia_tavoyensis[1]
| | |
| | | +-- Nitia_teretiuscula
| | | |
| | | +-- Coelatura_gabonensis
| | | |
| | | +-- Prisodontopsis_mweruensis
| +--+ |
| | +-- Coelatura_kunenensis
| | |
| | +--------------+-- Coelatura_aegyptiaca[1]
| | | |
| | | +-- Coelatura_aegyptiaca[2]
| | | |
| | | +-- Prisodontopsis_aviculaeformis
| | | |
| | | +-- Coelatura_luapulaensis
| | | |
| | | +-- Coelatura_choziensis
| +--+
| | +-------- Indonaia_shurtleffiana
| | |
| | +-----+ +----- Indonaia_rugosa
| | | | |
| | | | | +-- Indonaia_andersoniana
| | | +--+--+
| | | | +-- Indonaia_caerulea
| | | |
| +--+ | +-- Indonaia_bonneaudii
| | +--+
| | +-- Lamellidens_corrianus[2]
| |
| | +----------- Indochinella_pugio
| | |
| | | +-- Scabies_humilis
| +--+ |
| | +-----+-- Scabies_phaselus
--+ | | |
| +--+ +-- Scabies_mandarinus
| |
| | +----- Radiatula_mouhoti
| +--+
| | +-- Radiatula_myitkyinae
| +--+
| +-- Radiatula_chaudhurii
|
| +----- Indopseudodon_indawgyiensis **
| +-----------+
| | | +-- Indopseudodon_salwenianus **
| | +--+
| | +-- Indopseudodon_bogani **
| |
| | +-- Nyeinchanconcha_nyeinchani **
| | +-----------+
| +-----+ | +-- Sundadontina_cumingii **
| | | |
| | | | +----- Bineurus_exilis **
| | | +--------+
| | | | | +-- Bineurus_mouhotii **
| | | | +--+
| | | | +-- Bineurus_loeiensis **
| | | |
| | +--+ +-------- Sundadontina_tumida **
| | +-----+
| | | | +----- Pseudodon_vagulus **
| | | +--+
| | | | +-- Pseudodon_mekongi **
| | | +--+
| | | +-- Pseudodon_vondembuschianus **
| | |
| | | +----------- Thaiconcha_callifera **
| | +--+
| | | +-------- Pilsbryoconcha_exilis[3] **
| | +--+
| | | +----- Pilsbryoconcha_exilis[2] **
| | +--+
| | | +-- Pilsbryoconcha_exilis[1] **
| | +--+
| | +-- Pilsbryoconcha_carinifera **
| |
| | +-- Ensidens_telus **
| | +-----+
| | | +-- Ensidens_ingallsianus **
| | +--+
| | | | +----- Ensidens_jaculus **
| | | +--+
| | | | +-- Ensidens_spiculus **
| | | +--+
| | +-----+ +-- Ensidens_sagittarius **
| | | |
| | | | +-------- Hyriopsis_desowitzi **
| | | | |
| | | | +-------- Rectidens_sumatrensis **
+-----+ | +--+
| | | +----- Hyriopsis_panhai **
| | | |
| | +--+ +-- Hyriopsis_myersiana **
| | | |
| +--+ +--+-- Hyriopsis_bialata **
| | | |
| | | +-- Hyriopsis_khoratensis **
| | |
| | | +----- Physunio **
| | | +--------+
| | | | | +-- Physunio_superbus **
| | | | +--+
| | | | +-- Physunio_modelli **
| | | |
| | +--+ +-------- Yaukthwa_inlenensis **
| | | +--+
| | | | | +----- Yaukthwa_panhai **
| | | | +--+
| | | | | +-- Yaukthwa_nesemanni **
| | | | +--+
| | +--+ +-- Yaukthwa_rectangularis **
| | |
| | | +-- Lens_contradens **
| | | +-----+
| | | | +-- Lens_micropterus **
| | +--+
| | | +----- Lens_eximius **
| | +--+
| | | +-- Lens_comptus **
| | +--+
| | +-- Lens_novoselovi **
| |
| | +-- Amblema_plicata
| | |
| | +-- Ortmanniana_ligamentina
| | |
| | +-- Lampsilis_cardium
| | |
| | +-- Cambarunio_iris
+--+ |
| +--+-- Elliptio_complanata
| | |
| | +-- Eurynia_dilatata
| | |
| | +-- Pleurobema_sintoxia
| | |
| | +-- Quadrula_quadrula
| | |
| | +-- Quadrula_verrucosa
| |
| | +-- Chamberlainia_somsakpanhai **
| +--+ |
| | | +-- Sinohyriopsis_cumingii **
| | | |
| | | +-- Lamprotula_caveata **
| | | |
| | | +-- Lamprotula_leaii **
| | | |
| | | +-- Potomida_littoralis **
| | | |
| | | +-- Pronodularia_japanensis **
| | | |
| | +--+-- Gonidea_angulata **
| | |
| | +-- Leguminaia_wheatleyi **
| | |
| | +-- Leguminaia **
| | |
| | +-- Microcondylaea_bonellii **
| | |
| | +-- Sinosolenaia_carinata **
| | |
| | +-- Sinosolenaia_oleivora **
| | |
| | +-- Sinosolenaia_recognita **
+-----------+
| +-- Unio_crassus
| |
| +-- Unio_pictorum
| |
| +-- Unio_tumidus
| |
| +-- Aculamprotula_tortuosa
| |
| +-- Cuneopsis_heudei
| +--+
| | +-- Tchangsinaia_piscicula
| | |
| | +-- Cuneopsis_rufescens
| | |
| | +-- Nodularia_douglasiae
| | |
| | +-- Nodularia_nuxpersicae
| | |
| | +-- Schistodesmus_lampreyanus
| |
| | +-- Alasmidonta_marginata
| | |
| | +-- Anodonta_anatina
+--+ |
| +-- Anodonta_cygnea
| |
| +-- Anodonta_nuttalliana
| |
| +-- Lasmigona_costata
| |
| +-- Pseudanodonta_complanata
| |
| +-- Pyganodon_grandis
| |
| +-- Simpsonaias_ambigua
| |
| +-- Strophitus_undulatus
+--+
+-- Sinanodonta_lauta
|
+-- Cristaria_plicata
|
+-- Cristaria_radiata
|
+-- Pletholophus_tenuis
|
+-- Sinanodonta_lucida
|
+-- Sinanodonta_jourdyi
|
+-- Lanceolaria_gladiola
|
+-- Lanceolaria_grayii[1]
|
+-- Lanceolaria_grayii[2]
Bolotov et al., 2017 [Publ 3761], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 51 |
87 |
85 |
6 |
90 |
65 |
3 |
86.5 |
+-- Trapezidens_dolichorhynchus
+-----------+
| +-- Trapezidens_exolescens
+-----------+
| | +----------- Lamellidens_corrianus
| | |
| +--+ +-------- Lamellidens_savadiensis
| | |
| +--+ +-- Lamellidens_jenkinsianus
| | +--+
| | | +-- Lamellidens_mainwaringi
| +--+
+--------+ | +-- Lamellidens_ferrugineus
| | +--+
| | +-- Lamellidens_marginalis
| |
| | +-- Parreysia_nagpoorensis
| | +--------------------+
| | | +-- Parreysia_plagiosoma
| | |
| | | +-- Leoparreysia_tavoyensis
| | | +-----------------+
| +--+ | +-- Leoparreysia_olivacea
| | |
| | | +-------------- Nitia_teretiuscula
| | | +--+
| | | | | +----------- Coelatura_aegyptiaca[2]
| +--+ | +--+
| | | | +-------- Coelatura_aegyptiaca[1]
| | | +--+
| | | | +----- Prisodontopsis_mweruensis
| | | +--+
| | | | +-- Coelatura_gabonensis
| +--+ +--+
| | +-- Coelatura_kunenensis
| |
| | +-------- Indonaia_shurtleffiana
| | +-----+
| | | | +----- Indonaia_andersoniana
| | | +--+
| | | | +-- Indonaia_rugosa
| | | +--+
| +--+ +-- Indonaia_bonneaudii
| |
| | +-- Indochinella_pugio
| | +--------+
| | | +-- Scabies_crispata
| | |
| +--+ +-- Radiatula_myitkyinae
| | +-----+
| | | +-- Radiatula_chaudhurii
| +--+
| | +----- Scabies_mandarinus
| +--+
| | +-- Scabies_humilis
| +--+
| +-- Scabies_phaselus
|
| +-- Ensidens_telus **
| +-----+
--+ | +-- Ensidens_ingallsianus **
| +-----+
| | | +----- Ensidens_sagittarius **
| | +--+
| | | +-- Ensidens_spiculus **
| | +--+
| +--+ +-- Ensidens_jaculus **
| | |
| | | +-- Hyriopsis_bialata **
| | | +--------+
| | | | +-- Hyriopsis_khoratensis **
| | +--+
| | | +-------- Hyriopsis_panhai **
| | +--+
| | | +----- Hyriopsis_myersiana **
| | +--+
| +--------------+ | +-- Rectidens_sumatrensis **
| | | +--+
| | | +-- Hyriopsis_desowitzi **
| | |
| | | +----- Physunio **
| | | +--------+
| | | | | +-- Physunio_modelli **
| | | | +--+
| | | | +-- Physunio_superbus **
| | | |
| | +--+ +-------- Yaukthwa_inlenensis **
| | | +--+
| | | | | +----- Yaukthwa_panhai **
| | | | +--+
| | | | | +-- Yaukthwa_rectangularis **
| | | | +--+
| | +--+ +-- Yaukthwa_nesemanni **
| | |
| | | +-- Lens_contradens **
| | | +-----+
| | | | +-- Lens_micropterus **
| | +--+
| | | +----- Lens_eximius **
| | +--+
| | | +-- Lens_comptus **
| | +--+
| | +-- Lens_novoselovi **
| |
| | +-------------------- Indopseudodon_indawgyiensis **
| | |
| | | +-------- Pilsbryoconcha_exilis[1] **
| | | +--------+
+--+ +--------+ | | +----- Pilsbryoconcha_exilis[3] **
| | | | +--+
| | | | | +-- Pilsbryoconcha_exilis[2] **
| | | | +--+
| | | | +-- Pilsbryoconcha_carinifera **
| | +--+
| | | +-------- Sundadontina_tumida **
| | | +-----+
| | | | | +----- Pseudodon_vagulus **
| | | | +--+
| | | | | +-- Pseudodon_mekongi **
| | | | +--+
| | +--+ +-- Pseudodon_vondembuschianus **
| | |
| | | +-- Nyeinchanconcha_nyeinchani **
| | | +--------+
| | | | +-- Sundadontina_cumingii **
| | +--+
| | | +-------- Thaiconcha_callifera **
| | +--+
| | | +----- Bineurus_exilis **
| | +--+
| | | +-- Bineurus_mouhotii **
| | +--+
| | +-- Bineurus_loeiensis **
| |
| | +-------- Amblema_plicata
| | +--+
| | | | +----- Ortmanniana_ligamentina
| | | +--+
| | | | +-- Lampsilis_cardium
| | | +--+
+--+ +-----+ +-- Cambarunio_iris
| | |
| | | +-- Quadrula_quadrula
| | | +-----+
| | | | +-- Quadrula_verrucosa
| | +--+
| | | +----- Pleurobema_sintoxia
| | +--+
| | | +-- Eurynia_dilatata
| | +--+
| | +-- Elliptio_complanata
| +--------+
| | | +-- Sinohyriopsis_cumingii **
| | | +-----+
| | | | +-- Chamberlainia_somsakpanhai **
| | | |
| | | +-----+ +-- Potomida_littoralis **
| | | | | +--+
| | | | | | +-- Pronodularia_japanensis **
| | | | +--+
| | | | | +-- Lamprotula_caveata **
| | | | +--+
| | +--+ +-- Lamprotula_leaii **
| | |
| | | +----- Sinosolenaia_carinata **
| | | +-----+
| | | | | +-- Sinosolenaia_oleivora **
| | | | +--+
| | +--+ +-- Sinosolenaia_recognita **
| | |
| | | +-------- Gonidea_angulata **
| | +--+
+--+ | +----- Microcondylaea_bonellii **
| +--+
| | +-- Leguminaia_wheatleyi **
| +--+
| +-- Leguminaia **
|
| +----- Unio_tumidus
| +--------+
| | | +-- Unio_pictorum
| | +--+
| | +-- Unio_crassus
| +--------+
| | | +----------- Aculamprotula_tortuosa
| | | |
| | | | +-- Nodularia_douglasiae
| | +--+ +-----+
| | | | +-- Nodularia_nuxpersicae
| | | |
| | +--+ +-- Cuneopsis_rufescens
| | | +--+
| | | | +-- Cuneopsis_heudei
| | +--+
+--+ | +-- Tchangsinaia_piscicula
| +--+
| +-- Schistodesmus_lampreyanus
|
| +-- Lanceolaria_grayii
| +-----------------+
| | +-- Lanceolaria_gladiola
| |
| | +-- Cristaria_plicata
| | +--------+
| | | +-- Cristaria_radiata
+--+ +-----+
| | | +-------- Pletholophus_tenuis
| | +--+
| | | +----- Sinanodonta_lucida
| | +--+
| | | +-- Sinanodonta_jourdyi
+--+ +--+
| +-- Sinanodonta_lauta
|
| +-------------- Anodonta_anatina
| |
| | +----- Pseudanodonta_complanata
+--+ +-----+
| | | +-- Anodonta_cygnea
| | +--+
| | +-- Anodonta_nuttalliana
+--+
| +-- Lasmigona_costata
| +-----+
| | +-- Alasmidonta_marginata
+--+
| +----- Simpsonaias_ambigua
+--+
| +-- Pyganodon_grandis
+--+
+-- Strophitus_undulatus
Bolotov et al., 2017 [Publ 3761], fig. S2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 51 |
139 |
123 |
6 |
90 |
65 |
3 |
144.5 |
+-------------------- Chamberlainia_somsakpanhai **
|
+-----------+ +----------------- Cumberlandia_monodonta
| | |
| | | +-------------- Gibbosula_laosensis
| +--+ |
| | | +-- Pseudunio_auricularius
| | +-----------+
| +--+ +-- Pseudunio_marocanus
| |
| | +-- Margaritifera_dahurica
| | +--------+
| | | +-- Margaritifera_margaritifera
| +--+
| | +-------- Margaritifera_falcata
| +--+
| | +----- Margaritifera_middendorffi
| +--+
| | +-- Margaritifera_laevis
| +--+
| +-- Margaritifera_marrianae
|
| +-- Tamsiella_schroeteriana
| +--+
| | +-- Etheria_elliptica
| +--------+
| | | +-- Aspatharia_pfeifferiana
| | +--+
| | +-- Chambardia_wahlbergi
| |
+-----------------+ +----- Diplodon_suavidicus
| | +-----+
| | | | +-- Castalia_ambigua
| | | +--+
| | | +-- Triplodon_corrugatus
| +--+
| | +----- Velesunio_ambiguus
| | +--+
| | | | +-- Alathyria_jacksoni
| | | +--+
| +--+ +-- Alathyria_profuga
| |
| | +----- Microdontia_anodontaeformis
| +--+
| | +-- Alathyria_pertexta
| +--+
| +-- Lortiella_froggatti
|
| +-- Parreysia_nagpoorensis
| +-----------+
| | +-- Parreysia_plagiosoma
| |
| +--------+ +-- Trapezidens_dolichorhynchus
| | | +--------+
| | | | +-- Trapezidens_exolescens
| | | |
| | +--+ +-------- Lamellidens_savadiensis
--+ | | |
| | | +-------- Lamellidens_corrianus
| | +--+
| | | +-- Lamellidens_marginalis
| | | +--+
| | | | +-- Lamellidens_ferrugineus
| +-----+ +--+
| | | | +-- Lamellidens_jenkinsianus
| | | +--+
| | | +-- Lamellidens_mainwaringi
| | |
| | | +-- Leoparreysia_olivacea
| | | +-----------------+
| | | | +-- Leoparreysia_tavoyensis
| | | |
| | | | +-------------- Nitia_teretiuscula
| | | | +--+
| | | | | | +----------- Coelatura_aegyptiaca[1]
| | +--+ | +--+
| | | | | +-------- Coelatura_aegyptiaca[2]
| | | | +--+
| | | | | +----- Coelatura_gabonensis
| | | | +--+
| | | | | +-- Coelatura_kunenensis
| | +--+ +--+
| | | +-- Prisodontopsis_mweruensis
| | |
| | | +-------- Indonaia_shurtleffiana
| | | +-----+
| | | | | +----- Indonaia_andersoniana
| | | | +--+
| | | | | +-- Indonaia_rugosa
| | | | +--+
| | +--+ +-- Indonaia_bonneaudii
| | |
| | | +-- Indochinella_pugio
| | | +--------+
| | | | +-- Scabies_crispata
| | | |
| | +--+ +-- Radiatula_myitkyinae
| | | +-----+
| | | | +-- Radiatula_chaudhurii
| | +--+
| | | +----- Scabies_mandarinus
| | +--+
| | | +-- Scabies_humilis
| | +--+
| | +-- Scabies_phaselus
| |
| | +----- Unio_tumidus
| | +-----+
| | | | +-- Unio_crassus
| | | +--+
| | | +-- Unio_pictorum
| | +-----+
+--+ | | +-------- Aculamprotula_tortuosa
| | | |
| | | +-------- Schistodesmus_lampreyanus
| | | |
| | +--+ +-- Nodularia_douglasiae
| | +-----+
| | | +-- Nodularia_nuxpersicae
| | |
| | | +----- Cuneopsis_heudei
| | +--+
| | | +-- Tchangsinaia_piscicula
| | +--+
| +--------+ +-- Cuneopsis_rufescens
| | |
| | | +-- Lanceolaria_grayii
| | | +-----------+
| | | | +-- Lanceolaria_gladiola
| | | |
| | | | +-- Cristaria_plicata
| | | | +--------+
| | | | | +-- Cristaria_radiata
| | | +--+
| | | | | +-------- Pletholophus_tenuis
| | | | +--+
| | | | | +----- Sinanodonta_lucida
| | +--+ +--+
| | | | +-- Sinanodonta_lauta
| | | +--+
| | | +-- Sinanodonta_jourdyi
| | |
| | | +-------- Anodonta_anatina
| | | |
| | | +-------- Pseudanodonta_complanata
| | | |
| | | | +-- Anodonta_cygnea
| | +-----+-----+
| | | +-- Anodonta_nuttalliana
| | |
| | | +----- Pyganodon_grandis
| | | |
| | | +----- Simpsonaias_ambigua
| | +--+
| | +----- Strophitus_undulatus
| | |
| | | +-- Alasmidonta_marginata
| | +--+
| | +-- Lasmigona_costata
| |
| | +-------- Lens_contradens **
| | +-----+
+--+ | | +----- Lens_eximius **
| | +--+
| | | +-- Lens_comptus **
| | +--+
| +--+ +-- Lens_novoselovi **
| | |
| | | +----- Physunio **
| | | +-----+
| | | | | +-- Physunio_modelli **
| | | | +--+
| | +--+ +-- Physunio_superbus **
| | |
| | | +-------- Yaukthwa_inlenensis **
| | +--+
| | | +----- Yaukthwa_panhai **
| +-----+ +--+
| | | | +-- Yaukthwa_rectangularis **
| | | +--+
| | | +-- Yaukthwa_nesemanni **
| | |
| | | +----- Hyriopsis_panhai **
| | | |
| | | +--------+----- Hyriopsis_khoratensis **
| | | | |
| | | | | +-- Hyriopsis_myersiana **
| | | | +--+
| | +--+ +-- Hyriopsis_bialata **
| | |
| | | +----------- Hyriopsis_desowitzi **
| | | |
| | | +----------- Rectidens_sumatrensis **
| | +--+
| | | +-- Ensidens_telus **
| | | +-----+
| | | | +-- Ensidens_ingallsianus **
| | +--+
| | | +----- Ensidens_sagittarius **
| | +--+
| | | +-- Ensidens_spiculus **
+--+ +--+
| +-- Ensidens_jaculus **
|
| +-------- Amblema_plicata
| +--+
| | | +----- Cambarunio_iris
| | +--+
| | | +-- Ortmanniana_ligamentina
| | +--+
| +--------+ +-- Lampsilis_cardium
| | |
| | | +-- Quadrula_quadrula
| | | +-----+
| | | | +-- Quadrula_verrucosa
| | +--+
| | | +----- Pleurobema_sintoxia
| | +--+
| | | +-- Elliptio_complanata
| | +--+
| | +-- Eurynia_dilatata
| |
| | +-------- Sinohyriopsis_cumingii **
| | |
| | +-----+ +-- Lamprotula_caveata **
+--+ | | +--+
| | | | +-- Lamprotula_leaii **
| | +--+
| | | +-- Potomida_littoralis **
| +--+ +--+
| | | +-- Pronodularia_japanensis **
| | |
| | | +----------- Sinosolenaia_carinata **
| | | |
| | | | +-- Sinosolenaia_oleivora **
| | +--+--------+
| | | +-- Sinosolenaia_recognita **
| | |
| | | +-------- Gonidea_angulata **
| | +--+
| | | +----- Microcondylaea_bonellii **
+--+ +--+
| | +-- Leguminaia_wheatleyi **
| +--+
| +-- Leguminaia **
|
| +-------- Pilsbryoconcha_exilis[3] **
| +-----+
| | | +----- Pilsbryoconcha_exilis[2] **
| | +--+
| | | +-- Pilsbryoconcha_exilis[1] **
| | +--+
| | +-- Pilsbryoconcha_carinifera **
| |
+--+ +----------- Indopseudodon_indawgyiensis **
| |
| | +-------- Thaiconcha_callifera **
| +--+
| | | +----- Bineurus_exilis **
| | +--+
| | | +-- Bineurus_mouhotii **
+--+ +--+
| +-- Bineurus_loeiensis **
|
| +----- Pseudodon_vagulus **
| +--+
| | | +-- Pseudodon_mekongi **
| | +--+
| | +-- Pseudodon_vondembuschianus **
+--+
| +-- Pseudodon_inoscularis **
| +--+
| | +-- Sundadontina_tumida **
+--+
| +-- Nyeinchanconcha_nyeinchani **
+--+
+-- Sundadontina_cumingii **
Dai et al., 2023 [Publ 4285], fig. 3-BI
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 43 |
43 |
39 |
6 |
2 |
0 |
0 |
55.5 |
+----- Indopseudodon_indawgyiensis **
+--------+
| | +-- Indopseudodon_bogani **
| +--+
| +-- Indopseudodon_salwenianus **
|
| +-------- Nyeinchanconcha_nyeinchani **
| +--+
| | | +----- Bineurus_loeiensis **
+--------+ | +--+
| | | | +-- Bineurus_mouhotii **
| | | +--+
| | | +-- Bineurus_exilis **
| | |
| | | +-- Thaiconcha_callifera **
| | | +-----+
| | | | +-- Thaiconcha_munelliptica **
| +--+--+
| | | +----- Sundadontina_tanintharyiensis **
| | +--+
| | | +-- Sundadontina_brandti **
| | +--+
| | +-- Sundadontina_taskaevi **
| |
| | +-- Pilsbryoconcha_carinifera **
| | +-----+
| | | +-- Pilsbryoconcha_exilis **
| +--+
--+ | +----- Pseudodon_lenyanensis **
| +--+
| | +-- Pseudodon_mekongi **
| +--+
| +-- Pseudodon_vagulus **
|
| +-- Hyriopsis_bialata **
| +-----+
| | +-- Ensidens_ingallsianus **
| +-----------+
| | | +----- Physunio_superbus **
| | +--+
| | | +-- Lens_contradens **
| | +--+
| | +-- Lens_comptus **
| |
| | +-- Inversidens_brandtii **
| | +--------------+
| | | +-- Inversidens_rentianensis **
+--+ |
| | +----- Chamberlainia_somsakpanhai **
| | +-----+
| | | | +-- Sinohyriopsis_cumingii **
| | | +--+
| +-----+ +-- Sinohyriopsis_schlegelii **
| | |
| | | +-------- Potomida_littoralis **
| | +--+
| | | +----- Pronodularia_japanensis **
+--+ +--+
| | +-- Lamprotula_leaii **
| +--+
| +-- Lamprotula_caveata **
|
| +-------- Sinosolenaia_carinata **
| +-----+
| | | +----- Sinosolenaia_recognita **
| | +--+
| | | +-- Sinosolenaia_iridinea **
| | +--+
| | +-- Sinosolenaia_oleivora **
+--+
| +----- Gonidea_angulata **
| +-----+
| | | +-- Leguminaia_wheatleyi **
| | +--+
| | +-- Microcondylaea_bonellii **
+--+
| +-- Koreosolenaia_sitgyensis **
| +-----+
| | +-- Parvasolenaia_rivularis **
+--+
| +----- Postolata_guangxiensis **
+--+
| +-- Ptychorhynchus_pfisteri **
+--+
+-- Obovalis_omiensis **
Dai et al., 2023 [Publ 4285], fig. 3-ML
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 43 |
43 |
41 |
6 |
2 |
0 |
0 |
56.5 |
+----- Indopseudodon_indawgyiensis **
+-----------+
| | +-- Indopseudodon_bogani **
| +--+
| +-- Indopseudodon_salwenianus **
|
| +-------- Nyeinchanconcha_nyeinchani **
+-----+ +-----+
| | | | +----- Bineurus_loeiensis **
| | | +--+
| | | | +-- Bineurus_mouhotii **
| | | +--+
| | | +-- Bineurus_exilis **
| | |
| +--+ +-- Thaiconcha_callifera **
| | +-----+
| | | +-- Thaiconcha_munelliptica **
| | +--+
| | | | +----- Sundadontina_tanintharyiensis **
| | | +--+
| | | | +-- Sundadontina_brandti **
| | | +--+
| +--+ +-- Sundadontina_taskaevi **
| |
| | +-- Pilsbryoconcha_carinifera **
| | +-----+
| | | +-- Pilsbryoconcha_exilis **
--+ +--+
| | +----- Pseudodon_lenyanensis **
| +--+
| | +-- Pseudodon_mekongi **
| +--+
| +-- Pseudodon_vagulus **
|
| +-- Hyriopsis_bialata **
| +-----+
| | +-- Ensidens_ingallsianus **
| +-----------+
| | | +----- Physunio_superbus **
| | +--+
| | | +-- Lens_contradens **
| | +--+
| | +-- Lens_comptus **
| |
| | +-- Inversidens_brandtii **
| | +-----------+
| | | +-- Inversidens_rentianensis **
+--+ |
| +--+ +----- Chamberlainia_somsakpanhai **
| | | +-----+
| | | | | +-- Sinohyriopsis_cumingii **
| | | | +--+
| | +--+ +-- Sinohyriopsis_schlegelii **
| | |
| | | +-------- Potomida_littoralis **
| | +--+
| | | +----- Pronodularia_japanensis **
| | +--+
+--+ | +-- Lamprotula_leaii **
| +--+
| +-- Lamprotula_caveata **
|
| +-------- Sinosolenaia_carinata **
| +-----+
| | | +----- Sinosolenaia_recognita **
| | +--+
| | | +-- Sinosolenaia_iridinea **
| | +--+
| | +-- Sinosolenaia_oleivora **
+--+
| +----- Gonidea_angulata **
| +-----+
| | | +-- Leguminaia_wheatleyi **
| | +--+
| | +-- Microcondylaea_bonellii **
+--+
| +-- Koreosolenaia_sitgyensis **
| +-----+
| | +-- Parvasolenaia_rivularis **
+--+
| +----- Postolata_guangxiensis **
+--+
| +-- Ptychorhynchus_pfisteri **
+--+
+-- Obovalis_omiensis **
Bolotov et al., 2023 [Publ 4312], fig. S2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 42 |
42 |
39 |
1 |
0 |
0 |
0 |
52.5 |
+----------- Indopseudodon_kayinensis **
+-----------------------+
| | +-------- Indopseudodon_indawgyiensis **
| +--+
| | +----- Indopseudodon_bogani **
| +--+
| | +-- Indopseudodon_salwenianus **
| +--+
| +-- Indopseudodon_crebristriatus **
|
--+ +-------------------------------- Songkhlanaia_tamodienica **
| |
| | +----------------- Nyeinchanconcha_nyeinchani **
| | |
| | +-----------+ +-------------- Sundadontina_tumida **
| | | | |
| | | | | +-------- Sundadontina **
| | | +--+ +--+
| | | | | | +----- Sundadontina_tanintharyiensis **
| | | | | +--+
+--+ | | | | +-- Sundadontina_plugpomenica **
| | +--+ +--+
| | | +-- Sundadontina_cumingii **
| | |
| | | +-------- Sundadontina_sulcata **
| | +--+
| | | +----- Sundadontina_harmandi **
| | +--+
| | | +-- Sundadontina_taskaevi **
| | +--+
| | +-- Sundadontina_brandti **
+--+
| +-- Namkongnaia_inkhavilayi **
| +--------------+
| | +-- Namkongnaia_lemeslei **
| |
| +--------+ +-- Thaiconcha_callifera **
| | | |
| | | +-----------+-- Thaiconcha_thaiensis **
| | | | |
| | +--+ +-- Thaiconcha_munelliptica **
| | |
| | | +----------- Bineurus_loeiensis **
| | +--+
| | | +-------- Bineurus_panhai **
| | +--+
| | | +----- Bineurus_anodontinum **
| | +--+
+--+ | +-- Bineurus_exilis **
| +--+
| +-- Bineurus_mouhotii **
|
| +----------- Pseudodon_vagulus **
| |
| +-----------+ +-- Pseudodon_lenyanensis **
| | | +-----+
| | | | +-- Pseudodon_inoscularis **
| | +--+
| | | +----- Pseudodon_cambodjensis **
| | +--+
+--+ | +-- Pseudodon_vondembuschianus **
| +--+
| +-- Pseudodon_mekongi **
|
| +-------------------- Pilsbryoconcha_carinifera **
| |
+--+ +-- Pilsbryoconcha_linguaeformis[2] **
| +--------------+
| | +-- Pilsbryoconcha **
+--+
| +-------------- Pilsbryoconcha_linguaeformis[1] **
| |
+--+ +-- Pilsbryoconcha_schomburgki **
| +--------+
| | +-- Pilsbryoconcha_exilis **
+--+
| +-------- Pilsbryoconcha_hoikaab **
+--+
| +----- Pilsbryoconcha_mekongiana **
+--+
| +-- Pilsbryoconcha_acuta **
+--+
+-- Pilsbryoconcha_kittitati **
Bolotov et al., 2023 [Publ 4312], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 40 |
40 |
38 |
1 |
0 |
0 |
0 |
49.5 |
+----------- Indopseudodon_kayinensis **
+-----------------+
| | +-------- Indopseudodon_indawgyiensis **
| +--+
| | +----- Indopseudodon_bogani **
| +--+
| | +-- Indopseudodon_salwenianus **
| +--+
| +-- Indopseudodon_crebristriatus **
|
| +-- Pilsbryoconcha_exilis **
| +-----------+
--+ | +-- Pilsbryoconcha_schomburgki **
| |
| +-----------+ +-- Pilsbryoconcha_carinifera **
| | | +--------+
| | | | +-- Pilsbryoconcha_linguaeformis **
| | +--+
| | | +-------- Pilsbryoconcha_acuta **
| | +--+
| | | +----- Pilsbryoconcha_kittitati **
| | +--+
| | | +-- Pilsbryoconcha_mekongiana **
| | +--+
+--+ +-- Pilsbryoconcha_hoikaab **
|
| +----------------------- Songkhlanaia_tamodienica **
| |
| | +-- Namkongnaia_inkhavilayi **
| | +--------+
| | | +-- Namkongnaia_lemeslei **
| | |
| | | +----- Pseudodon_vondembuschianus[2] **
| | +--------+ +--+
| | | | | | +-- Pseudodon_vondembuschianus[1] **
+--+ | | | +--+
| | | | +-- Pseudodon_mekongi **
| | +--+
| | | +-- Pseudodon_vagulus **
| | | +--+
| | | | +-- Pseudodon_cambodjensis **
| | +--+
| | | +-- Pseudodon_lenyanensis **
| | +--+
| | +-- Pseudodon_inoscularis **
+--+
| +-------------- Nyeinchanconcha_nyeinchani **
| |
| | +----- Thaiconcha_munelliptica **
| +--+ +-----+
| | | | | +-- Thaiconcha_callifera **
| | | | +--+
| | +--+ +-- Thaiconcha_thaiensis **
| | |
| | | +-------- Bineurus_loeiensis **
| | +--+
| | | +----- Bineurus_mouhotii **
| | +--+
+--+ | +-- Bineurus_anodontinum **
| +--+
| +-- Bineurus_exilis **
|
| +-------- Sundadontina_sulcata **
| +-----+
| | | +----- Sundadontina_harmandi **
| | +--+
| | | +-- Sundadontina_brandti **
+--+ +--+
| +-- Sundadontina_taskaevi **
|
| +----------- Sundadontina_tumida **
+--+
| +-------- Sundadontina **
+--+
| +----- Sundadontina_tanintharyiensis **
+--+
| +-- Sundadontina_cumingii **
+--+
+-- Sundadontina_plugpomenica **
Bolotov et al., 2023 [Publ 4312], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 39 |
39 |
37 |
1 |
0 |
0 |
0 |
48.5 |
+----------- Indopseudodon_kayinensis **
+-----------------+
| | +-------- Indopseudodon_indawgyiensis **
| +--+
| | +----- Indopseudodon_bogani **
| +--+
| | +-- Indopseudodon_salwenianus **
| +--+
| +-- Indopseudodon_crebristriatus **
|
| +-- Pilsbryoconcha_exilis **
| +-----------+
| | +-- Pilsbryoconcha_schomburgki **
--+ |
| +-----------+ +-- Pilsbryoconcha_carinifera **
| | | +--------+
| | | | +-- Pilsbryoconcha_linguaeformis **
| | +--+
| | | +-------- Pilsbryoconcha_kittitati **
| | +--+
| | | +----- Pilsbryoconcha_acuta **
| | +--+
| | | +-- Pilsbryoconcha_hoikaab **
| | +--+
| | +-- Pilsbryoconcha_mekongiana **
| |
+--+ +-------------- Nyeinchanconcha_nyeinchani **
| |
| | +----- Thaiconcha_thaiensis **
| +--------+ +-----+
| | | | | +-- Thaiconcha_callifera **
| | | | +--+
| | +--+ +-- Thaiconcha_munelliptica **
| | |
| | | +-------- Bineurus_loeiensis **
| | +--+
| | | +----- Bineurus_mouhotii **
| | +--+
+--+ | +-- Bineurus_anodontinum **
| +--+
| +-- Bineurus_exilis **
|
| +-------------------- Songkhlanaia_tamodienica **
| |
| | +-- Namkongnaia_lemeslei **
| | +--------+
| | | +-- Namkongnaia_inkhavilayi **
| | |
+--+ +-----+ +-- Pseudodon_inoscularis **
| | | +-----+
| | | | +-- Pseudodon_lenyanensis **
| | | |
| | +--+ +-- Pseudodon_vagulus **
| | | +--+
| | | | +-- Pseudodon_cambodjensis **
+--+ +--+
| | +-- Pseudodon_vondembuschianus **
| +--+
| +-- Pseudodon_mekongi **
|
| +-------------- Sundadontina_tumida **
| |
| | +-------- Sundadontina_sulcata **
+--+ +--+
| | | +----- Sundadontina_harmandi **
| | +--+
| | | +-- Sundadontina_brandti **
+--+ +--+
| +-- Sundadontina_taskaevi **
|
| +-------- Sundadontina_tanintharyiensis **
+--+
| +----- Sundadontina **
+--+
| +-- Sundadontina_cumingii **
+--+
+-- Sundadontina_plugpomenica **
Konopleva et al., 2019 [Publ 3900], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 38 |
50 |
39 |
3 |
14 |
12 |
1 |
55.0 |
+-- Indopseudodon_indawgyiensis **
|
+--+-- Indopseudodon_bogani **
| |
| +-- Indopseudodon_salwenianus **
|
| +-- Bineurus_mouhotii **
+--------------------------+ |
| | +-- Pseudodon_vagulus **
| | |
| | +-- Pseudodon_mekongi **
| | |
| | +-- Pilsbryoconcha_exilis **
| +--+
| +-- Pilsbryoconcha_carinifera **
| |
| +-- Bineurus_loeiensis **
| |
| +-- Bineurus_exilis **
| |
| +-- Pseudodon_vondembuschianus **
|
| +----------- Indonaia_shurtleffiana
| +--+
| | | +-------- Indonaia_rugosa
| | +--+
--+ | | +----- Indonaia_andersoniana
| | +--+
| | | +-- Indonaia_caerulea
| +--------------+ +--+
| | | +-- Indonaia_subclathrata
| | |
| | | +----------- Indochinella_pugio
| | | |
| | | | +-- Scabies_humilis
| | +--+ |
| | | +-----+-- Scabies_mandarinus
| | | | |
| | +--+ +-- Scabies_phaselus
| | |
| | | +----- Radiatula_mouhoti
| | +--+
| | | +-- Radiatula_myitkyinae
| | +--+
| | +-- Radiatula_chaudhurii
| |
+--+ +-- Ensidens_ingallsianus **
| +-----+
| | +-- Ensidens_telus **
| +-----+
| | | +----- Ensidens_sagittarius **
| | +--+
| | | +-- Ensidens_jaculus **
| +-----------+ +--+
| | | +-- Ensidens_spiculus **
| | |
| | | +----------- Hyriopsis_desowitzi **
| | | |
| | +--+----------- Rectidens_sumatrensis **
| | |
| | | +-------- Hyriopsis_panhai **
| | +--+
| | | +----- Hyriopsis_myersiana **
| | +--+
+--+ | +-- Hyriopsis_khoratensis **
| +--+
| +-- Hyriopsis_bialata **
|
| +-- Yaukthwa_paiensis **
| +-----+
| | +-- Yaukthwa_inlenensis **
| +--------------+
| | | +----- Yaukthwa_panhai **
| | +--+
| | | +-- Yaukthwa_nesemanni **
| | +--+
+--+ +-- Yaukthwa_rectangularis **
|
| +----- Physunio_superbus **
| +--------------+
| | | +-- Physunio_modelli **
| | +--+
+--+ +-- Physunio **
|
| +----------------- Trapezoideus_foliaceus **
| |
+--+ +-- Lens_contradens **
| +-----------+
| | +-- Lens_micropterus **
+--+
| +----------- Lens_misellus **
+--+
| +-------- Lens_eximius **
+--+
| +----- Lens_pallegoixi **
+--+
| +-- Lens_comptus **
+--+
+-- Lens_novoselovi **
Bolotov et al., 2018 [Publ 3826], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 36 |
61 |
26 |
5 |
49 |
49 |
3 |
69.0 |
+-- Trapezidens_exolescens
+--------+
| +-- Trapezidens_dolichorhynchus
+--------------+
| | +-------- Lamellidens_savadiensis
| +--+
| | +----- Lamellidens_ferrugineus
| +--+
| | +-- Lamellidens_generosus
| +--+
+--------+ +-- Lamellidens_mainwaringi
| |
| | +-- Parreysia_keralaensis
| | +--------------------+
| | | +-- Parreysia_nagpoorensis
| | |
| | | +----- Leoparreysia_olivacea
| +--+ +--------------+
| | | | +-- Leoparreysia_canefrii
| | | +--+
| | | +-- Leoparreysia_tavoyensis
| | |
| +--+ +-- Coelatura_aegyptiaca
| | +--------------+
| | | +-- Nitia_teretiuscula
| | |
| | | +-------- Indonaia_shurtleffiana
| +--+ +-----+
| | | | +----- Indonaia_rugosa
| | | +--+
| | | | +-- Indonaia_caerulea
| | | +--+
| +--+ +-- Indonaia_andersoniana
| |
| | +----------- Indochinella_pugio
| | |
| | | +----- Radiatula_myitkyinae
| +--+ +--+
| | | | +-- Radiatula_mouhoti
| | | +--+
| +--+ +-- Radiatula_chaudhurii
| |
| | +----- Scabies_mandarinus
| +--+
| | +-- Scabies_humilis
| +--+
--+ +-- Scabies_phaselus
|
| +-- Indopseudodon_indawgyiensis **
| |
| +--+-- Indopseudodon_bogani **
| | |
| | +-- Indopseudodon_salwenianus **
| |
| | +-- Bineurus_mouhotii **
| +--------------------------+ |
| | | +-- Pseudodon_vagulus **
| | | |
| | | +-- Pseudodon_mekongi **
| | | |
| | | +-- Pilsbryoconcha_exilis **
| | +--+
| | +-- Pilsbryoconcha_carinifera **
| | |
| | +-- Bineurus_loeiensis **
| | |
| | +-- Bineurus_exilis **
| | |
| | +-- Pseudodon_vondembuschianus **
| |
| | +-- Ensidens_telus **
| | |
| | +-- Ensidens_ingallsianus **
| | |
| | +-- Ensidens_jaculus **
| | |
| | +-- Hyriopsis_desowitzi **
| | +--+
| | | +-- Hyriopsis_myersiana **
| | | |
| | | +-- Hyriopsis_khoratensis **
| | | |
+--+ | +-- Ensidens_spiculus **
| | |
| | +-- Ensidens_sagittarius **
| |
| +-----------------------+ +-- Lens_eximius **
| | | |
| | | +-- Physunio_modelli[1] **
| | | |
| | | +-- Yaukthwa_nesemanni **
| | | |
| | | +-- Yaukthwa_panhai **
| | | |
| | | +-- Lens_comptus **
| | | |
| | +--+-- Lens_novoselovi **
| | |
| | +-- Physunio_modelli[2] **
| | |
| | +-- Physunio **
| | |
| | +-- Yaukthwa_inlenensis **
| | |
| | +-- Lens_contradens **
| | |
| | +-- Lens_micropterus **
| |
| | +-- Lamprotula_leaii **
+--+ |
| +-- Potomida_littoralis **
| |
| +-- Pronodularia_japanensis **
| |
| +-- Gonidea_angulata **
| |
| +-- Leguminaia_wheatleyi **
| |
| +-- Microcondylaea_bonellii **
| |
| +-----------------------+-- Amblema_plicata
| | |
| | +-- Ortmanniana_ligamentina
| | |
| | +-- Lampsilis_cardium
| | |
| | +-- Cambarunio_iris
| | |
| | +-- Elliptio_complanata
| | |
| | +-- Pleurobema_sintoxia
| | |
+--+ +-- Quadrula_quadrula
|
| +----- Unio_tumidus
| +--+
| | | +-- Unio_crassus
| | +--+
| +--------------+ +-- Unio_pictorum
| | |
| | | +----- Nodularia_douglasiae[1]
| | +--+
| | | +-- Nodularia_nuxpersicae
| | +--+
+--+ +-- Nodularia_douglasiae[2]
|
| +-- Lanceolaria_gladiola
| +-----------------+
| | +-- Lanceolaria_grayii
| |
| | +-------- Cristaria_plicata
+--+ +--------+
| | | +----- Pletholophus_tenuis
| | +--+
| | | +-- Sinanodonta_jourdyi
+--+ +--+
| +-- Sinanodonta_lauta
|
| +-------------- Anodonta_anatina
+--+
| +----------- Pyganodon_grandis
+--+
| +-------- Simpsonaias_ambigua
+--+
| +----- Strophitus_undulatus
+--+
| +-- Alasmidonta_marginata
+--+
+-- Lasmigona_costata
Jeratthitikul et al., 2021 [Publ 4219], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 36 |
36 |
33 |
6 |
0 |
0 |
0 |
44.5 |
+----- Lens_contradens **
+--+
| | +-- Trapezoideus_foliaceus **
| +--+
+--------------------+ +-- Physunio_superbus **
| |
| | +----- Ensidens_ingallsianus **
| +--+
| | +-- Hyriopsis_bialata **
| +--+
| +-- Rectidens_sumatrensis **
|
| +----- Sinosolenaia_carinata **
| +-----+
| | | +-- Leguminaia_wheatleyi **
--+ | +--+
| | +-- Gonidea_angulata **
| +--------------+
| | | +-- Chamberlainia_somsakpanhai **
| | | +-----+
| | | | +-- Sinohyriopsis_cumingii **
| | +--+
| | | +----- Potomida_littoralis **
| | +--+
| | | +-- Pronodularia_japanensis **
| | +--+
+--+ +-- Lamprotula_leaii **
|
| +-------- Indopseudodon_kayinensis **
| +--------------+
| | | +----- Indopseudodon_indawgyiensis **
| | +--+
| | | +-- Indopseudodon_salwenianus **
| | +--+
| | +-- Indopseudodon_bogani **
| |
+--+ +----- Pilsbryoconcha_exilis **
| +--------------+
| | | +-- Pilsbryoconcha_linguaeformis **
| | +--+
| | +-- Pilsbryoconcha **
| |
| | +-------- Nyeinchanconcha_nyeinchani **
| | |
+--+ +--------+ +----- Thaiconcha_callifera **
| | | |
| | +--+ +-- Bineurus_panhai **
| | | |
| | +--+-- Bineurus_exilis **
| | |
| | +-- Bineurus_mouhotii **
+--+
| +-- Sundadontina_brandti **
| +--+
| | +-- Sundadontina_taskaevi **
| +--------+
| | | +-- Sundadontina_tumida **
| | +--+
| | +-- Sundadontina_cumingii **
+--+
| +-- Namkongnaia_lemeslei **
| +--------+
| | +-- Namkongnaia_inkhavilayi **
+--+
| +-------- Pseudodon_lenyanensis **
+--+
| +----- Pseudodon_vagulus **
+--+
| +-- Pseudodon_vondembuschianus **
+--+
+-- Pseudodon_mekongi **
Jeratthitikul et al., 2021 [Publ 4219], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 36 |
36 |
34 |
6 |
0 |
0 |
0 |
44.5 |
+----- Lens_contradens **
+--+
| | +-- Trapezoideus_foliaceus **
| +--+
+--------------------+ +-- Physunio_superbus **
| |
| | +----- Ensidens_ingallsianus **
| +--+
| | +-- Hyriopsis_bialata **
| +--+
| +-- Rectidens_sumatrensis **
|
| +----- Sinosolenaia_carinata **
| +-----+
| | | +-- Gonidea_angulata **
--+ | +--+
| | +-- Leguminaia_wheatleyi **
| +--------------+
| | | +-- Chamberlainia_somsakpanhai **
| | | +-----+
| | | | +-- Sinohyriopsis_cumingii **
| | +--+
| | | +----- Potomida_littoralis **
| | +--+
| | | +-- Lamprotula_leaii **
| | +--+
+--+ +-- Pronodularia_japanensis **
|
| +-------- Indopseudodon_kayinensis **
| +--------------+
| | | +----- Indopseudodon_indawgyiensis **
| | +--+
| | | +-- Indopseudodon_salwenianus **
| | +--+
| | +-- Indopseudodon_bogani **
| |
+--+ +----- Pilsbryoconcha_exilis **
| +--------------+
| | | +-- Pilsbryoconcha_linguaeformis **
| | +--+
| | +-- Pilsbryoconcha **
| |
| | +-- Nyeinchanconcha_nyeinchani **
| | +--------+
+--+ | +-- Sundadontina_cumingii **
| +-----+
| | | +-------- Thaiconcha_callifera **
| | +--+
| | | +----- Bineurus_mouhotii **
| | +--+
| | | +-- Bineurus_panhai **
| | +--+
+--+ +-- Bineurus_exilis **
|
| +----- Sundadontina_tumida **
| +--------+
| | | +-- Sundadontina_brandti **
| | +--+
| | +-- Sundadontina_taskaevi **
+--+
| +-- Namkongnaia_lemeslei **
| +--------+
| | +-- Namkongnaia_inkhavilayi **
+--+
| +-------- Pseudodon_lenyanensis **
+--+
| +----- Pseudodon_vagulus **
+--+
| +-- Pseudodon_vondembuschianus **
+--+
+-- Pseudodon_mekongi **
Wu et al., 2024 [Publ 4350], fig. 5a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 36 |
36 |
34 |
8 |
2 |
0 |
0 |
20.0 |
+-------------------------- Chamberlainia_somsakpanhai **
|
| +-- Khairuloconcha_lunbawangorum **
| +--------------+
| | +-- Khairuloconcha_sahanae **
| |
| | +-------- Ensidens_sagittarius **
| +-----+ +-----+
| | | | | +----- Rectidens_sumatrensis **
| | | | +--+
| | | | | +-- Hyriopsis_bialata **
| | | | +--+
--+ | +--+ +-- Hyriopsis_desowitzi **
| | |
| | | +-- Physunio_superbus **
| | | +--------+
| | | | +-- Trapezoideus_foliaceus **
| | | |
| | +--+ +-- Lens_contradens **
| | | +-----+
| | | | +-- Lens_eximius **
| | +--+
| | | +----- Yaukthwa_elongatula **
| | +--+
| | | +-- Yaukthwa_inlenensis **
+--+ +--+
| +-- Yaukthwa_paiensis **
|
| +-- Schepmania[1] **
| +--------------+
| | +-- Schepmania[2] **
| |
| +--+ +-- Indopseudodon_bogani **
| | | +-----------+
| | | | +-- Indopseudodon_kayinensis **
| | +--+
| | | +----------- Pilsbryoconcha_exilis **
| | | |
| | +--+ +-- Pseudodon_mekongi **
| | | +-----+
| | | | +-- Pseudodon_vondembuschianus **
| | +--+
| | | +----- Thaiconcha_callifera **
+--+ +--+
| | +-- Bineurus_anodontinum **
| +--+
| +-- Bineurus_loeiensis **
|
| +-- Inversidens_brandtii **
| +-----+
| | +-- Inversidens_rentianensis **
| +--------+
| | | +----- Potomida_littoralis **
| | +--+
| | | +-- Lamprotula_caveata **
+--+ +--+
| +-- Lamprotula_leaii **
|
| +-------------- Sinosolenaia_carinata **
| |
+--+ +-- Gonidea_angulata **
| +--------+
| | +-- Microcondylaea_bonellii **
+--+
| +-------- Parvasolenaia_rivularis **
+--+
| +----- Koreosolenaia_sitgyensis **
+--+
| +-- Postolata_guangxiensis **
+--+
+-- Ptychorhynchus_pfisteri **
Wu et al., 2024 [Publ 4350], fig. 5b
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 36 |
36 |
34 |
8 |
2 |
0 |
0 |
40.0 |
+-- Schepmania[1] **
+--------------------------------+
| +-- Schepmania[2] **
|
| +-- Indopseudodon_bogani **
| +-----------+
| | +-- Indopseudodon_kayinensis **
| +-----------------+
| | | +----------- Pilsbryoconcha_exilis **
| | | |
| | +--+ +-- Bineurus_anodontinum **
--+ | | +-----+
| | | | +-- Bineurus_loeiensis **
| | +--+
| | | +----- Thaiconcha_callifera **
| | +--+
| | | +-- Pseudodon_mekongi **
| | +--+
| | +-- Pseudodon_vondembuschianus **
| |
| | +-------------------------- Hyriopsis_bialata **
| | |
+--+ | +----- Rectidens_sumatrensis **
| +--+ +-----------------+
| | | | | +-- Khairuloconcha_lunbawangorum **
| | | | +--+
| | +--+ +-- Khairuloconcha_sahanae **
| | |
| | | +-------------------- Ensidens_sagittarius **
| | | |
| | +--+ +----------------- Hyriopsis_desowitzi **
| | | |
| | +--+ +-------------- Yaukthwa_elongatula **
| | | |
| | +--+ +----------- Trapezoideus_foliaceus **
| | | |
| | +--+ +-- Physunio_superbus **
+--+ | +-----+
| | | +-- Yaukthwa_inlenensis **
| +--+
| | +----- Yaukthwa_paiensis **
| +--+
| | +-- Lens_contradens **
| +--+
| +-- Lens_eximius **
|
| +-- Inversidens_brandtii **
| +-----------------+
| | +-- Inversidens_rentianensis **
| |
| | +-- Chamberlainia_somsakpanhai **
+--------+ +--+
| | +-- Potomida_littoralis **
| +-----------+
| | | +-- Lamprotula_caveata **
| | +--+
+--+ +-- Lamprotula_leaii **
|
| +-------------- Sinosolenaia_carinata **
| |
+--+ +-- Gonidea_angulata **
| +--------+
| | +-- Microcondylaea_bonellii **
+--+
| +-------- Koreosolenaia_sitgyensis **
+--+
| +----- Parvasolenaia_rivularis **
+--+
| +-- Postolata_guangxiensis **
+--+
+-- Ptychorhynchus_pfisteri **
Wu et al., 2024 [Publ 4350], fig. 5c
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 36 |
36 |
23 |
8 |
2 |
0 |
0 |
33.0 |
+----------------------------- Schepmania[1] **
|
+----------------------------- Schepmania[2] **
|
| +-- Indopseudodon_bogani **
| +-----------+
| | +-- Indopseudodon_kayinensis **
| +-----------+
| | | +----------- Pilsbryoconcha_exilis **
| | | |
| | +--+ +-------- Bineurus_anodontinum **
--+ | | |
| | +--+-------- Bineurus_loeiensis **
| | |
| | | +----- Thaiconcha_callifera **
| | +--+
| | | +-- Pseudodon_mekongi **
| | +--+
| | +-- Pseudodon_vondembuschianus **
| |
| | +-------- Ensidens_sagittarius **
| | |
| | +-------- Hyriopsis_desowitzi **
+--+ |
| +-------- Rectidens_sumatrensis **
| |
| +-------- Hyriopsis_bialata **
| |
| +--------------+ +-- Khairuloconcha_lunbawangorum **
| | +-----+
| | | +-- Khairuloconcha_sahanae **
| | |
| | | +----- Physunio_superbus **
| | | |
| | | +----- Yaukthwa_paiensis **
| | | |
| | | +----- Yaukthwa_inlenensis **
| | +--+
| | +----- Trapezoideus_foliaceus **
+--+ |
| +----- Yaukthwa_elongatula **
| |
| | +-- Lens_contradens **
| +--+
| +-- Lens_eximius **
|
| +-- Inversidens_rentianensis **
| +-----------------+
| | +-- Inversidens_brandtii **
| |
| | +-- Chamberlainia_somsakpanhai **
+--+ +--+
| | +-- Potomida_littoralis **
| +-----------+
| | | +-- Lamprotula_caveata **
| | +--+
+--+ +-- Lamprotula_leaii **
|
| +-------------- Sinosolenaia_carinata **
| |
+--+ +----------- Gonidea_angulata **
| |
+--+----------- Microcondylaea_bonellii **
|
| +-------- Koreosolenaia_sitgyensis **
+--+
| +----- Parvasolenaia_rivularis **
+--+
| +-- Postolata_guangxiensis **
+--+
+-- Ptychorhynchus_pfisteri **
Wu et al., 2024 [Publ 4350], fig. 6-7
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 36 |
36 |
34 |
8 |
2 |
0 |
0 |
20.0 |
+----------------------------- Chamberlainia_somsakpanhai **
|
| +-- Khairuloconcha_lunbawangorum **
| +--------------+
| | +-- Khairuloconcha_sahanae **
| |
| | +-------- Ensidens_sagittarius **
| +--------+ +-----+
| | | | | +----- Rectidens_sumatrensis **
| | | | +--+
| | | | | +-- Hyriopsis_bialata **
| | | | +--+
--+ | +--+ +-- Hyriopsis_desowitzi **
| | |
| | | +-- Physunio_superbus **
| | | +--------+
| | | | +-- Trapezoideus_foliaceus **
| | | |
| | +--+ +-- Lens_contradens **
| | | +-----+
| | | | +-- Lens_eximius **
| | +--+
| | | +----- Yaukthwa_elongatula **
| | +--+
+--+ | +-- Yaukthwa_inlenensis **
| +--+
| +-- Yaukthwa_paiensis **
|
| +-- Schepmania[1] **
| +--------------+
| | +-- Schepmania[2] **
| |
| +-----+ +-- Indopseudodon_bogani **
| | | +-----------+
| | | | +-- Indopseudodon_kayinensis **
| | +--+
| | | +----------- Pilsbryoconcha_exilis **
| | | |
| | +--+ +-- Pseudodon_mekongi **
| | | +-----+
| | | | +-- Pseudodon_vondembuschianus **
| | +--+
+--+ | +----- Thaiconcha_callifera **
| +--+
| | +-- Bineurus_anodontinum **
| +--+
| +-- Bineurus_loeiensis **
|
| +----- Potomida_littoralis **
| +--------------+
| | | +-- Lamprotula_caveata **
| | +--+
| | +-- Lamprotula_leaii **
+--+
| +-- Inversidens_brandtii **
| +--------------+
| | +-- Inversidens_rentianensis **
+--+
| +-------------- Sinosolenaia_carinata **
| |
+--+ +-- Gonidea_angulata **
| +--------+
| | +-- Microcondylaea_bonellii **
+--+
| +-------- Koreosolenaia_sitgyensis **
+--+
| +----- Parvasolenaia_rivularis **
+--+
| +-- Postolata_guangxiensis **
+--+
+-- Ptychorhynchus_pfisteri **
Konopleva et al., 2019 [Publ 4041], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 35 |
35 |
21 |
3 |
0 |
0 |
0 |
45.0 |
+-- Indopseudodon_indawgyiensis **
|
+--------------------+-- Indopseudodon_bogani **
| |
| +-- Indopseudodon_salwenianus **
|
| +-- Ensidens_ingallsianus **
| |
| +-- Ensidens_telus **
| |
| +-- Ensidens_dugasti **
| |
| +-- Hyriopsis_khoratensis **
| |
| +-- Hyriopsis_bialata **
| |
--+ +-- Hyriopsis_myersiana **
| |
| +-----------------+-- Hyriopsis_desowitzi **
| | |
| | +-- Rectidens_sumatrensis **
| | |
| | +-- Hyriopsis_panhai **
| | |
| | +-- Ensidens_jaculus[1] **
| | |
| | +-- Ensidens_jaculus[2] **
| | |
| | +-- Ensidens_spiculus **
| | |
+--+ +-- Ensidens_sagittarius **
|
| +----------- Yaukthwa_elongatula **
| |
| +-----+ +-- Yaukthwa_inlenensis **
| | | +-----+
| | | | +-- Yaukthwa_paiensis **
| | +--+
| | | +----- Yaukthwa_panhai **
| | +--+
| | | +-- Yaukthwa_nesemanni **
| | +--+
+--+ +-- Yaukthwa_rectangularis **
|
| +-------- Trapezoideus_foliaceus **
| +-----+
| | | +----- Physunio_superbus **
| | +--+
| | | +-- Physunio_modelli **
| | +--+
| | +-- Physunio **
+--+
| +----- Lens_misellus **
| +-----+
| | | +-- Lens_contradens **
| | +--+
| | +-- Lens_micropterus **
+--+
| +----- Lens_pallegoixi **
| +--+
| | | +-- Lens_novoselovi **
| | +--+
+--+ +-- Lens_comptus **
|
| +----- Lens_maenamensis **
+--+
| +-- Lens_eximius **
+--+
+-- Lens_rolfbrandti **
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 35 |
39 |
30 |
6 |
87 |
76 |
3 |
48.5 |
+-- Coelatura_aegyptiaca
|
+-----------------------------------+-- Leoparreysia_tavoyensis
| |
| +-- Radiatula_mouhoti
|
| +-- Lens_contradens **
| |
| +-- Ensidens_sagittarius **
| |
| +-----------------------+-- Physunio_modelli **
| | |
| | +-- Rectidens_sumatrensis **
| | |
| | +-- Yaukthwa_nesemanni **
| |
| | +-- Amblema_plicata
| | |
| | +-- Elliptio_complanata
| | +--------------------+
| +--------+ | +-- Lampsilis_cardium
| | | | |
| | | | +-- Quadrula_quadrula
| | | |
| | | | +----- Chamberlainia_somsakpanhai **
| | | | +-----------+
| | | | | | +-- Sinohyriopsis_cumingii **
| | | | | +--+
| | | | +--+ +-- Sinohyriopsis_schlegelii **
| | | | | |
| | | | | | +-------------- Pronodularia_japanensis **
| | +--+ | | |
| | | | +--+ +----------- Potomida_littoralis **
| | | | | |
| | | | +--+ +-------- Lamprotula_caveata **
| | | | | |
--+ | | | +--+ +----- Lamprotula_leaii[2] **
| | | | | |
| | | | +--+----- Lamprotula_leaii[3] **
| | | | |
| | | | | +-- Lamprotula_seomjinensis **
| | | | +--+
| | | | +-- Lamprotula_leaii[1] **
| | +--+
| | | +-- Indopseudodon_indawgyiensis **
| | | +--------+
| | | | +-- Indopseudodon_salwenianus **
| | | |
| | | +-----+ +-- Pilsbryoconcha_exilis **
| | | | | +-----+
| | | | | | +-- Pilsbryoconcha_carinifera **
| | | | +--+
| | | | | +----- Pseudodon_mekongi **
| | | | | |
| | | | +--+ +-- Nyeinchanconcha_nyeinchani **
| | | | | |
| | | | +--+-- Bineurus_mouhotii **
| | | | |
| | +--+ +-- Sundadontina_tumida **
| | |
| | | +-------- Inversidens_brandtii **
| | | |
| | | +-----+ +-- Obovalis_omiensis **
| | | | | +--+
| | | | | | +-- Ptychorhynchus_pfisteri **
| | | | +--+
| | | | | +-- Parvasolenaia_rivularis **
| | | | +--+
| | +--+ +-- Koreosolenaia_sitgyensis **
| | |
+--+ | +----- Gonidea_angulata **
| | +-----+
| | | | +-- Leguminaia_wheatleyi **
| | | +--+
| +--+ +-- Microcondylaea_bonellii **
| |
| | +-------- Sinosolenaia_carinata **
| +--+
| | +----- Sinosolenaia_recognita **
| +--+
| | +-- Sinosolenaia_iridinea **
| +--+
| +-- Sinosolenaia_oleivora **
|
| +-------------------------------- Acuticosta_chinensis
| |
| +-------------------------------- Lepidodesma_languilati
| |
| | +----------- Aculamprotula_tortuosa
| | +--------------+
| | | | +-------- Aculamprotula_scripta
| | | +--+
| | | | +----- Aculamprotula_coreana
| | | +--+
| | | | +-- Aculamprotula_fibrosa
| | | +--+
| +-----+ +-- Aculamprotula_tientsinensis
| | |
| | | +----------------------- Middendorffinaia_mongolica
| | | |
| | | | +----- Unio_gibbus
| | | | |
| | | | +--------------+----- Unio_tumidus
| | +--+ | |
| | | | | +-- Unio_crassus
| | | | +--+
| | | | +-- Unio_pictorum
| | | |
| | | | +-------------- Nodularia_breviconcha
| | +--+ +--+
| | | | | +----------- Nodularia[2]
| | | | +--+
| | | | | +-------- Nodularia_nuxpersicae
| | | | +--+
| | | | | +----- Nodularia_nipponensis
| | | | +--+
| | | | | +-- Nodularia_douglasiae
| | +--+ +--+
| | | +-- Nodularia[1]
| | |
+--+ | +-- Schistodesmus_lampreyanus
| | +-----------+
| | | +-- Schistodesmus_spinosus
| | |
| | | +----- Cuneopsis_rufescens
| | +--------+
| +--+ | +-- Cuneopsis_heudei
| | +--+
| | +-- Cuneopsis_celtiformis
| |
| | +----------- Tchangsinaia_piscicula
| +--+
| | +-------- Inversiunio_jokohamensis
| +--+
| | +----- Inversiunio_verrucosus
| +--+
| | +-- Inversiunio_reinianus
| +--+
| +-- Inversiunio_yanagawensis
|
| +----------------- Lanceolaria_oxyrhyncha
| |
| +-----------+ +-------------- Lanceolaria_gladiola
| | | |
| | +--+ +-- Lanceolaria_bilirata
| | | +--------+
| | | | +-- Lanceolaria_grayii
| | +--+
| | | +-------- Lanceolaria_acrorrhyncha
| | +--+
| | | +----- Lanceolaria_kihirai
| | +--+
| | | +-- Lanceolaria_lanceolata
| | +--+
| | +-- Lanceolaria_triformis
| |
| | +-------- Pseudanodonta_complanata
+--+ |
| | +----- Anodonta_cygnea
| +--+
| | | +-- Anodonta_anatina
| +-----------------+ +--+
| | | +-- Anodonta_nuttalliana
| | |
| | | +----- Pyganodon_grandis
| | | |
| | | +----- Simpsonaias_ambigua
| | +--+
| | +----- Strophitus_undulatus
| | |
| | | +-- Alasmidonta_marginata
| | +--+
| | +-- Lasmigona_costata
+--+
| +-- Cristaria_plicata
| +--------------------+
| | +-- Cristaria_clessini
| |
| | +-------------- Anemina_arcaeformis
| | |
| +--------+ +----------- Amuranodonta_kijaensis
| | | |
| | +--+ +-------- Buldowskia_suifunica
| | | |
| | +--+ +-- Buldowskia_flavotincta
+--+ | +--+
| | | +-- Buldowskia_shadini
| +--+
| | +-- Buldowskia_iwakawai
| +--+
| +-- Buldowskia_kamiyai
|
| +-------------------- Pletholophus_tenuis
| |
| | +-------- Beringiana_japonica
| | +--------+
+--+ | | +----- Beringiana_beringiana
| | +--+
| | | +-- Beringiana_fukuharai
| | +--+
+--+ +-- Beringiana_gosannensis
|
| +-- Sinanodonta_angula
| +-----------+
| | +-- Sinanodonta_lucida
| |
+--+ +----------- Sinanodonta_schrenkii
| |
| | +-- Sinanodonta_calipygos
| +--------+
+--+ +-- Sinanodonta_tumens
|
| +-------- Sinanodonta_woodiana[2]
| |
+--+ +----- Sinanodonta_elliptica
| |
+--+----- Sinanodonta_jourdyi
|
| +-- Sinanodonta_lauta
+--+
+-- Sinanodonta_woodiana[1]
Zieritz et al., 2020 [Publ 4102], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 31 |
31 |
27 |
7 |
0 |
0 |
0 |
37.0 |
+----------------------------------- Chamberlainia_somsakpanhai **
|
| +-------------------------------- Lamprotula_leaii **
--+ |
| | +----------------------------- Gonidea_angulata **
| | |
+--+ | +-- Pilsbryoconcha_carinifera **
| | +-----------------------+
| | | +-- Indopseudodon_indawgyiensis **
| | |
+--+ | +-------------- Khairuloconcha_lunbawangorum **
| | |
| | | +-- Ensidens_telus **
| | | +-----+
| | | | +-- Ensidens_ingallsianus **
| | +--------+ +--+
| | | | | | +----- Ensidens_sagittarius **
+--+ | | | +--+
| | | | | +-- Ensidens_spiculus **
| | | | +--+
| | +--+ +-- Ensidens_jaculus **
| | |
| | | +-------- Hyriopsis_desowitzi **
| | | |
| | | +-------- Rectidens_sumatrensis **
| | +--+
| | | +----- Hyriopsis_panhai **
+--+ | |
| +--+ +-- Hyriopsis_khoratensis **
| | |
| +--+-- Hyriopsis_bialata **
| |
| +-- Hyriopsis_myersiana **
|
| +-- Physunio_modelli **
| +-----------------+
| | +-- Physunio_superbus **
| |
| | +-- Yaukthwa_paiensis **
| | +-----+
+--+ | +-- Yaukthwa_inlenensis **
| +--------+
| | | +----- Yaukthwa_panhai **
| | +--+
| | | +-- Yaukthwa_rectangularis **
+--+ +--+
| +-- Yaukthwa_nesemanni **
|
| +-------------- Trapezoideus_foliaceus **
| |
+--+ +-- Lens_contradens **
| +--------+
| | +-- Lens_misellus **
+--+
| +-------- Lens_eximius **
+--+
| +----- Lens_pallegoixi **
+--+
| +-- Lens_comptus **
+--+
+-- Lens_novoselovi **
Jeratthitikul et al., 2022 [Publ 4260], fig. S1, 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 31 |
66 |
63 |
6 |
45 |
35 |
2 |
78.5 |
+-- Indopseudodon_indawgyiensis **
+--------------------+
| +-- Indopseudodon_kayinensis **
|
| +-- Nyeinchanconcha_nyeinchani **
| +-----+
+-----------+ | +-- Sundadontina_cumingii **
| | +-----------+
| | | | +----- Thaiconcha_callifera **
| | | +--+
| | | | +-- Bineurus_panhai **
| | | +--+
| +--+ +-- Bineurus_mouhotii **
| |
| | +-- Namkongnaia_inkhavilayi **
| | +--------------+
| | | +-- Pseudodon_vondembuschianus **
| | |
| +--+ +-- Pilsbryoconcha_exilis **
| | +-----------+
| | | +-- Pilsbryoconcha_schomburgki **
| | |
| +--+ +-- Pilsbryoconcha_carinifera **
| | +--------+
| | | +-- Pilsbryoconcha_linguaeformis **
| +--+
--+ | +-------- Pilsbryoconcha_hoikaab **
| +--+
| | +----- Pilsbryoconcha_mekongiana **
| +--+
| | +-- Pilsbryoconcha_acuta **
| +--+
| +-- Pilsbryoconcha_kittitati **
|
| +----- Lens_contradens **
| +--+
| | | +-- Trapezoideus_foliaceus **
| | +--+
| +-----------------------+ +-- Physunio_superbus **
| | |
| | | +-- Ensidens_ingallsianus **
| | | |
| | +-----+-- Hyriopsis_bialata **
| | |
| | +-- Rectidens_sumatrensis **
| |
| | +----- Sinosolenaia_carinata **
+--+ +-----------------------+
| | | +-- Leguminaia_wheatleyi **
| | +--+
| | +-- Gonidea_angulata **
| |
| | +-- Chamberlainia_somsakpanhai **
| | +-----+
| | | +-- Sinohyriopsis_cumingii **
| | +-----------------+
| | | | +----- Lamprotula_leaii **
+--+ | +--+
| | | +-- Potomida_littoralis **
| | +--+
| | +-- Pronodularia_japanensis **
| |
| | +-------- Amblema_plicata
| | +--+
| | | | +----- Ortmanniana_ligamentina
| | | +--+
| | | | +-- Cambarunio_iris
| | | +--+
+--+ +-----------+ +-- Lampsilis_cardium
| | |
| | | +-- Quadrula_quadrula
| | | +-----+
| | | | +-- Quadrula_verrucosa
| | +--+
| | | +----- Elliptio_complanata
| | +--+
| | | +-- Pleurobema_sintoxia
| | +--+
| | +-- Eurynia_dilatata
| |
| | +----- Unio_tumidus
+--+ +-----------+
| | | +-- Unio_pictorum
| | +--+
| +--+ +-- Unio_crassus
| | |
| | | +-------------- Aculamprotula_tortuosa
| | | |
| | +--+ +-- Nodularia_douglasiae
| | | +--------+
| | | | +-- Nodularia_nuxpersicae
| | +--+
| | | +-------- Schistodesmus_lampreyanus
| | +--+
| | | +----- Cuneopsis_heudei
+--+ +--+
| | +-- Tchangsinaia_piscicula
| +--+
| +-- Cuneopsis_rufescens
|
| +-- Lanceolaria_grayii
| +--------------+
| | +-- Lanceolaria_gladiola
| |
| | +----------- Cristaria_plicata
| | +--+
+--+ | | +-------- Pletholophus_tenuis
| | +--+
| | | +----- Sinanodonta_lucida
| | +--+
| | | +-- Sinanodonta_lauta
| | +--+
+--+ +-- Sinanodonta_jourdyi
|
| +-------- Pseudanodonta_complanata
| +--+
| | | +----- Anodonta_nuttalliana
| | +--+
| | | +-- Anodonta_anatina
| | +--+
+--+ +-- Anodonta_cygnea
|
| +-- Pyganodon_grandis
| +-----+
| | +-- Simpsonaias_ambigua
+--+
| +----- Strophitus_undulatus
+--+
| +-- Alasmidonta_marginata
+--+
+-- Lasmigona_costata
Jeratthitikul et al., 2022 [Publ 4260], fig. S2, 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 31 |
66 |
64 |
6 |
45 |
35 |
2 |
77.5 |
+-- Indopseudodon_indawgyiensis **
+--------------------+
| +-- Indopseudodon_kayinensis **
|
| +-- Nyeinchanconcha_nyeinchani **
| +-----+
+-----------+ | +-- Sundadontina_cumingii **
| | +-----------+
| | | | +----- Thaiconcha_callifera **
| | | +--+
| | | | +-- Bineurus_panhai **
| | | +--+
| +--+ +-- Bineurus_mouhotii **
| |
| | +-- Namkongnaia_inkhavilayi **
| | +--------------+
| | | +-- Pseudodon_vondembuschianus **
| | |
| +--+ +-- Pilsbryoconcha_exilis **
| | +-----------+
| | | +-- Pilsbryoconcha_schomburgki **
| | |
| +--+ +-- Pilsbryoconcha_carinifera **
| | +--------+
| | | +-- Pilsbryoconcha_linguaeformis **
| +--+
--+ | +-------- Pilsbryoconcha_hoikaab **
| +--+
| | +----- Pilsbryoconcha_mekongiana **
| +--+
| | +-- Pilsbryoconcha_acuta **
| +--+
| +-- Pilsbryoconcha_kittitati **
|
| +----- Ensidens_ingallsianus **
| +--+
| | | +-- Rectidens_sumatrensis **
| | +--+
| +-----------------------+ +-- Hyriopsis_bialata **
| | |
| | | +----- Lens_contradens **
| | +--+
| | | +-- Trapezoideus_foliaceus **
| | +--+
| | +-- Physunio_superbus **
| |
| | +----- Sinosolenaia_carinata **
+--+ +-----------------------+
| | | +-- Gonidea_angulata **
| | +--+
| | +-- Leguminaia_wheatleyi **
| |
| | +-- Chamberlainia_somsakpanhai **
| | +-----+
| | | +-- Sinohyriopsis_cumingii **
| | +-----------------+
| | | | +----- Potomida_littoralis **
+--+ | +--+
| | | +-- Lamprotula_leaii **
| | +--+
| | +-- Pronodularia_japanensis **
| |
| | +-------- Amblema_plicata
| | +--+
| | | | +----- Ortmanniana_ligamentina
| | | +--+
| | | | +-- Lampsilis_cardium
| | | +--+
+--+ +-----------+ +-- Cambarunio_iris
| | |
| | | +-- Quadrula_quadrula
| | | +-----+
| | | | +-- Quadrula_verrucosa
| | +--+
| | | +----- Pleurobema_sintoxia
| | +--+
| | | +-- Eurynia_dilatata
| | +--+
| | +-- Elliptio_complanata
| |
| | +----- Unio_tumidus
+--+ +-----------+
| | | +-- Unio_pictorum
| | +--+
| +--+ +-- Unio_crassus
| | |
| | | +-------------- Aculamprotula_tortuosa
| | | |
| | +--+ +-- Nodularia_douglasiae
| | | +--------+
| | | | +-- Nodularia_nuxpersicae
| | +--+
| | | +-------- Schistodesmus_lampreyanus
| | +--+
| | | +----- Cuneopsis_heudei
+--+ +--+
| | +-- Tchangsinaia_piscicula
| +--+
| +-- Cuneopsis_rufescens
|
| +-- Lanceolaria_grayii
| +--------------+
| | +-- Lanceolaria_gladiola
| |
| | +----------- Cristaria_plicata
| | +--+
+--+ | | +-------- Pletholophus_tenuis
| | +--+
| | | +----- Sinanodonta_lucida
| | +--+
| | | +-- Sinanodonta_jourdyi
| | +--+
+--+ +-- Sinanodonta_lauta
|
| +-------- Pseudanodonta_complanata
| +--+
| | | +----- Anodonta_anatina
| | +--+
| | | +-- Anodonta_nuttalliana
| | +--+
+--+ +-- Anodonta_cygnea
|
| +-- Alasmidonta_marginata
| +-----+
| | +-- Lasmigona_costata
+--+
| +----- Pyganodon_grandis
+--+
| +-- Strophitus_undulatus
+--+
+-- Simpsonaias_ambigua
Pfeiffer et al., 2021 [Publ 4126], fig. 9
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 29 |
29 |
27 |
3 |
0 |
0 |
0 |
42.0 |
+-------------------------- Lamprotula_cornuumlunae **
|
| +-- Ensidens_ingallsianus **
| +--------+
| | +-- Ensidens_telus **
| +--------+
| | | +-------- Ensidens_spiculus **
| | +--+
| | | +----- Ensidens_dugasti **
| | +--+
| +--+ | +-- Ensidens_jaculus **
--+ | | +--+
| | | +-- Ensidens_sagittarius **
| | |
| | | +----------------- Rectidens_sumatrensis **
| | +--+
| | | +-------------- Hyriopsis_delaportei **
| | +--+
| | | +----------- Hyriopsis_myersiana **
| | +--+
| | | +-------- Hyriopsis_bialata **
| | +--+
| | | +----- Hyriopsis_khoratensis **
+--+ +--+
| | +-- Hyriopsis_kratiensis **
| +--+
| +-- Hyriopsis_sakhonensis **
|
| +-- Trapezoideus_subviridis **
| +-----+
| | +-- Trapezoideus_foliaceus **
| +-----------+
| | | +----- Physunio_superbus **
| | +--+
| | | +-- Physunio_modelli **
| | +--+
+--+ +-- Physunio_pellucidus **
|
| +----------------- Yaukthwa_paiensis **
| |
| | +-------------- Solenaia_emarginata **
+--+ |
| | +-------- Lens_misellus **
| | +--+
+--+ | | +----- Lens_inornatus **
| | +--+
| | | +-- Lens_contradens **
| | +--+
+--+ +-- Lens_micropterus **
|
| +-- Lens_maenamensis **
| +--+
| | +-- Lens_eximius **
+-----+
| +-- Lens_pallegoixi **
+--+
+-- Lens_comptus **
Konopleva et al., 2021 [Publ 4213], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 28 |
28 |
22 |
3 |
0 |
0 |
0 |
39.5 |
+-------------------------- Lamprotula_leaii **
|
+-------------------------- Leguminaia_wheatleyi **
|
+-------------------------- Potomida_littoralis **
|
+-------------------------- Gonidea_angulata **
|
--+ +-------- Indopseudodon_kayinensis **
| +--------------+
| | | +----- Indopseudodon_indawgyiensis **
| | +--+
| | | +-- Indopseudodon_salwenianus **
| | +--+
| | +-- Indopseudodon_bogani **
| |
| | +----- Pilsbryoconcha_exilis[2] **
+--+ +--------------+
| | | +-- Pilsbryoconcha_exilis[1] **
| | +--+
| | +-- Namkongnaia_lemeslei **
| |
| | +-------------- Nyeinchanconcha_nyeinchani **
| | |
| | | +-- Thaiconcha_munelliptica **
+--+ +--+ |
| | | +--------+-- Thaiconcha_thaiensis **
| | | | |
| | +--+ +-- Thaiconcha_callifera **
| | |
| | | +-------- Bineurus_loeiensis **
| | +--+
| | | +----- Bineurus_mouhotii **
| | +--+
+--+ | +-- Bineurus_anodontinum **
| +--+
| +-- Bineurus_exilis **
|
| +-------- Pseudodon_lenyanensis **
| +--+
| | | +----- Pseudodon_vagulus **
| | +--+
| | | +-- Pseudodon_vondembuschianus **
| | +--+
+-----+ +-- Pseudodon_mekongi **
|
| +-- Sundadontina_taskaevi **
| +-----+
| | +-- Sundadontina_brandti **
+--+
| +----- Sundadontina_tumida **
+--+
| +-- Sundadontina_cumingii **
+--+
+-- Sundadontina_tanintharyiensis **
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 27 |
37 |
32 |
6 |
54 |
42 |
3 |
50.0 |
+-------- Trapezidens_dolichorhynchus
+--------------------+
| | +----- Leoparreysia_tavoyensis
| +--+
| | +-- Coelatura_aegyptiaca
| +--+
| +-- Indochinella_pugio
|
| +----- Unio_gibbus
| |
| +-----------+----- Unio_tumidus
| | |
| | | +-- Unio_crassus
| | +--+
| +--+ +-- Unio_pictorum
| | |
| | | +-------------- Aculamprotula_tortuosa
| | | |
| | +--+ +-- Nodularia_nuxpersicae
| | | +--------+
| | | | +-- Nodularia_douglasiae
| | +--+
| | | +-------- Schistodesmus_lampreyanus
| | +--+
| | | +----- Tchangsinaia_piscicula
| | +--+
| +-----+ | +-- Cuneopsis_heudei
| | | +--+
| | | +-- Cuneopsis_rufescens
--+ | |
| | | +-------- Lanceolaria_gladiola
| | | +--------+
| | | | | +----- Lanceolaria_grayii[2]
| | | | +--+
| | | | | +-- Lanceolaria_lanceolata
| | | | +--+
| | | | +-- Lanceolaria_grayii[1]
| | | |
| | +--+ +----------- Cristaria_plicata
| | | +--+
| | | | | +-------- Anemina_arcaeformis
| | | | +--+
| | | | | +----- Pletholophus_tenuis
| | | | +--+
| | | | | +-- Sinanodonta_jourdyi
| | | | +--+
| | +--+ +-- Sinanodonta_lucida
| | |
| | | +----- Pseudanodonta_complanata
| | | |
| | | +-----+ +-- Anodonta_anatina
| | | | | |
| | | | +--+-- Anodonta_cygnea
| | | | |
| | +--+ +-- Anodonta_nuttalliana
| | |
| | | +-- Pyganodon_grandis
+--+ | +-----+
| | | +-- Simpsonaias_ambigua
| +--+
| | +----- Strophitus_undulatus
| +--+
| | +-- Alasmidonta_marginata
| +--+
| +-- Lasmigona_costata
|
| +-- Lens_contradens **
| +--+
| | +-- Lens_comptus **
| +--+
| | | +-- Physunio_modelli **
| | +--+
| | +-- unionoid_species
| |
| +--------------+ +----- Hyriopsis_desowitzi **
| | | |
| | | +----- Rectidens_sumatrensis **
| | | |
| | | | +-- Hyriopsis_khoratensis **
| | +--+--+
| | | +-- Hyriopsis_myersiana **
| | |
| | | +-- Ensidens_ingallsianus **
| | | |
| | +--+-- Ensidens_telus **
| | |
| | +-- Ensidens_jaculus **
| |
| | +-------- Amblema_plicata
+--+ +--+
| | | +----- Cambarunio_iris
| | +--+
| | | +-- Lampsilis_cardium
| | +--+
| +--------+ +-- Ortmanniana_ligamentina
| | |
| | | +-- Quadrula_quadrula
| | | +-----+
| | | | +-- Quadrula_verrucosa
| | +--+
| | | +----- Elliptio_complanata
| | +--+
| | | +-- Pleurobema_sintoxia
| | +--+
| | +-- Eurynia_dilatata
+--+
| +-- Chamberlainia_somsakpanhai **
| +-----+
| | +-- Sinohyriopsis_cumingii **
| |
| +--------+ +-- Lamprotula_caveata **
| | | +--+
| | | | +-- Lamprotula_leaii **
| | +--+
| | | +-- Potomida_littoralis **
| | +--+
| | +-- Pronodularia_japanensis **
+--+
| +-- Sinosolenaia_carinata **
| +-----+
| | +-- Sinosolenaia_oleivora **
| +-----+
| | | +----- Gonidea_angulata **
| | +--+
| | | +-- Microcondylaea_bonellii **
+--+ +--+
| +-- Leguminaia_wheatleyi **
|
| +----------- Parvasolenaia_rivularis **
| |
+--+ +-- Pilsbryoconcha_exilis **
| +-----+
| | +-- Pilsbryoconcha_acuta **
+--+
| +----- Pseudodon_vagulus **
+--+
| +-- Bineurus_exilis **
+--+
+-- Nyeinchanconcha_nyeinchani **
Bolotov et al., 2022 [Publ 4250], fig. 8
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 23 |
49 |
42 |
4 |
26 |
24 |
1 |
64.5 |
+-- Gibbosula_laosensis
|
+-- Margaritifera_dahurica
|
+-- Sinosolenaia_iridinea **
|
+-----------------------------+-- Leguminaia_wheatleyi **
| |
| +-- Gonidea_angulata **
| |
| +-- Lamprotula_leaii **
| |
| +-- Potomida_littoralis **
|
| +-- Pseudodon_lenyanensis **
| +--------+
| | +-- Sundadontina_tanintharyiensis **
| +--------+
| | | +-------- Indopseudodon_kayinensis **
| | +--+
| | | +----- Indopseudodon_indawgyiensis **
| | +--+
| | | +-- Indopseudodon_salwenianus **
| | +--+
| +--------+ +-- Indopseudodon_bogani **
--+ | |
| | | +----- Trapezoideus_mitanensis **
| | | +-----------+
| | | | | +-- Trapezoideus_foliaceus **
| | | | +--+
| | | | +-- Trapezoideus_lenya **
| | +--+
| | | +-- Yaukthwa_elongatula **
| | | +-----------+
| | | | +-- Yaukthwa_baniensis **
| | | |
| | +--+ +----- Yaukthwa_inlenensis **
| | | +-----+
| | | | | +-- Yaukthwa_paiensis **
| | | | +--+
| | +--+ +-- Yaukthwa_avaensis **
| | |
| | | +-------- Yaukthwa_panhai **
| | +--+
+--+ | +----- Yaukthwa_rectangularis **
| +--+
| | +-- Yaukthwa_nesemanni **
| +--+
| +-- Yaukthwa_zayleymanensis **
|
| +-------- Lamellidens_savadiensis
| +-----+
| | | +----- Lamellidens_ferrugineus
| | +--+
| | | +-- Lamellidens_generosus
| +-----------+ +--+
| | | +-- Lamellidens_mainwaringi
| | |
| | | +----------- Trapezidens_exolescens
| | +--+
| | | +-------- Trapezidens_scutum
| | +--+
| | | +----- Trapezidens_yeti
| | +--+
+--+ | +-- Trapezidens_angustior
| +--+
| +-- Trapezidens_dolichorhynchus
|
| +----------------------- Parreysia_rakhinensis
| |
| | +-- Leoparreysia_whitteni
| | +-----+
| | | +-- Leoparreysia_burmana
+--+ +-----------+
| | | +----- Leoparreysia_olivacea
| | +--+
| | | +-- Leoparreysia_canefrii
| | +--+
+--+ +-- Leoparreysia_tavoyensis
|
| +-- Indonaia_andersoniana
| +--------------+
| | +-- Indonaia_subclathrata
+--+
| +-------------- Indochinella_pugio
| |
+--+ +----------- Radiatula_crispisulcata
| |
+--+ +-- Radiatula_chaudhurii
| +-----+
| | +-- Radiatula_myitkyinae
+--+
| +----- Radiatula_mouhoti
+--+
| +-- Radiatula_chindwinensis
+--+
+-- Radiatula_myitthanensis
Jeratthitikul et al., 2019 [Publ 3911], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 21 |
22 |
14 |
3 |
1 |
1 |
1 |
31.0 |
+-- Pilsbryoconcha_exilis **
+--------------------------+
| +-- Scabies_crispata
|
--+ +-------------------------- Ensidens_ingallsianus **
| |
| | +-- Physunio_modelli **
+--+ +--------------------+
| | +-- Physunio_superbus **
| |
| | +-- Lens_maenamensis **
+--+ +-----------------+
| | +-- Lens_rolfbrandti **
| |
| | +----------------- Lens_contradens[12] **
+--+ |
| | +----- Lens_micropterus[3] **
| | |
| +-----------+ +-- Lens_contradens[10] **
| | | |
+--+ +--+-- Lens_micropterus[2] **
| |
| +-- Lens_contradens[11] **
|
| +-------------- Lens_contradens[9] **
| |
+--+ +----------- Lens_contradens[8] **
| |
+--+ +-------- Lens_contradens[7] **
| |
+--+ +----- Lens_contradens[6] **
| |
| | +-- Lens_contradens[1] **
+--+ |
| +-- Lens_contradens[2] **
| |
| +-- Lens_contradens[3] **
+--+
+-- Lens_contradens[4] **
|
+-- Lens_micropterus[1] **
|
+-- Lens_contradens[5] **
Jeratthitkul et al., 2021 [Publ 4212], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 21 |
21 |
19 |
2 |
0 |
0 |
0 |
39.0 |
+-------- Trapezoideus_foliaceus **
+-----+
| | +----- Physunio_superbus **
| +--+
| | +-- Physunio_pellucidus **
| +--+
+-----+ +-- Physunio_modelli **
| |
| | +-- Yaukthwa_nesemanni **
| | +--------+
| | | +-- Yaukthwa_panhai **
| +--+
| | +-------- Lens_contradens **
| +--+
| | +----- Lens_novoselovi **
--+ +--+
| | +-- Lens_rolfbrandti **
| +--+
| +-- Lens_maenamensis **
|
| +-- Ensidens_ingallsianus **
| +--------------+
| | +-- Ensidens_jaculus **
| |
+--+ +-- Hyriopsis_desowitzi **
| +-----------+
| | +-- Hyriopsis_delaportei **
| |
+--+ +----------- Rectidens_sumatrensis **
| |
| | +----- Hyriopsis_myersiana **
+--+ +--+
| | | +-- Hyriopsis_bialata **
| | +--+
+--+ +-- Hyriopsis_panhai **
|
| +----- Hyriopsis_khoratensis **
+--+
| +-- Hyriopsis_phuphaniensis **
+--+
+-- Hyriopsis_sakhonensis **
Wu et al., 2024 [Publ 4350], fig. 8a, S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 21 |
21 |
19 |
6 |
71 |
65 |
3 |
33.0 |
+-- Quadrula_quadrula
+--------------------------+
| +-- Uniomerus_tetralasmus
|
+--------+ +-- Elliptio_complanata
| | +-----------------------+
| | | +-- Pleurobema_oviforme
| +--+
| | +----------------------- Amblema_plicata
| | |
| +--+ +-------------------- Popenaias_popeii
| | |
| +--+ +----------------- Toxolasma_parvum
| | |
| | | +----- Potamilus_leptodon
| +--+ +--------+
| | | | +-- Potamilus_alatus
| | | +--+
| | | +-- Potamilus_streckersoni
| +--+
| | +----------- Leaunio_lienosus
| | |
| | | +----- Lampsilis_ornata
| +--+ +--+
| | | | +-- Lampsilis_cardium
| | | +--+
| +--+ +-- Lampsilis_cariosa
| |
| | +----- Venustaconcha_ellipsiformis
| +--+
| | +-- Lampsilis_powellii
| +--+
| +-- Lampsilis_siliquoidea
|
--+ +----- Chamberlainia_somsakpanhai **
| +--------------------+
| | | +-- Sinohyriopsis_cumingii **
| | +--+
| | +-- Sinohyriopsis_schlegelii **
| |
| | +-- Hyriopsis_bialata **
| | +--+
| +--------+ | +-- Rectidens_sumatrensis **
| | | +-----------------+
| | | | | +-- Lens_contradens **
| | | | +--+
| | | | +-- Physunio_superbus **
| | | |
| | | | +----------- Potomida_littoralis **
| | +--+ +--------+
| | | | | +-------- Pronodularia_japanensis **
| | | | +--+
| | | | | +----- Lamprotula_leaii **
| | | | +--+
| | | | | +-- Lamprotula_caveata **
| | +--+ +--+
| | | +-- Lamprotula_cornuumlunae **
| | |
| | | +-- Pseudodon_vondembuschianus **
| | | +--------------+
| | | | +-- Pilsbryoconcha_exilis **
| | +--+
| | | +-------------- Inversidens_rentianensis **
| | | |
+--+ +--+ +-- Parvasolenaia_rivularis **
| | +--------+
| | | +-- Ptychorhynchus_pfisteri **
| +--+
| | +-------- Microcondylaea_bonellii **
| +--+
| | +----- Sinosolenaia_carinata **
| +--+
| | +-- Sinosolenaia_iridinea **
| +--+
| +-- Sinosolenaia_oleivora **
|
| +-------------------------------- Lamellidens_marginalis
| |
| | +-- Lepidodesma_aligera
| | +--------------------------+
| | | +-- Lepidodesma_languilati
| | |
| | | +-------- Aculamprotula_tortuosa
| | | |
| | | +--------------+ +-- Aculamprotula_coreana
| | | | | +--+
+--+ | | | | +-- Aculamprotula_tientsinensis
| | | +--+
| | | | +-- Aculamprotula_polysticta
| | | +--+
| | | +-- Aculamprotula_scripta
| | |
| | +--+ +-------------- Unio_tumidus
| | | | +-----+
| | | | | | +----------- Unio_crassus
| | | | | +--+
+--+ | | | | +-------- Unio_elongatulus
| | | | +--+
| | | | | +----- Unio_pictorum
| | | | +--+
| | | | | +-- Unio_delphinus
| | +--+ +--+
| | | +-- Unio_mancus
| | |
| | | +-- Nodularia_breviconcha
| | | +--------------+
| | | | +-- Nodularia_douglasiae
| | | |
| | | | +-- Pseudocuneopsis_capitata
| | | +--------------+
| | +--+ +-- Pseudocuneopsis_sichuanensis
| | |
| | | +-- Schistodesmus_lampreyanus
| | | +-----------+
+--+ | | +-- Schistodesmus_spinosus
| +--+
| | +----------- Tchangsinaia_piscicula
| +--+
| | +-------- Cuneopsis_rufescens
| +--+
| | +----- Cuneopsis_celtiformis
| +--+
| | +-- Cuneopsis_demangei
| +--+
| +-- Cuneopsis_heudei
|
| +-------- Acuticosta_chinensis
| +--------------+
| | | +----- Lanceolaria_gladiola
| | +--+
| | | +-- Lanceolaria_grayii
| | +--+
| | +-- Lanceolaria_lanceolata
| |
| | +-------------- Cristaria_plicata
| | |
+--+ +-----+ +-- Anemina_arcaeformis
| | | +--------+
| | | | +-- Anemina_euscaphys
| | +--+
| | | +-------- Beringiana_fukuharai
| | +--+
| | | +----- Sinanodonta_lucida
| | +--+
| | | +-- Sinanodonta_tumens
+--+ +--+
| +-- Sinanodonta_woodiana
|
| +-------- Pseudanodonta_complanata
| +--------+
| | | +----- Anodonta_anatina
| | +--+
| | | +-- Anodonta_cygnea
| | +--+
+--+ +-- Anodonta_exulcerata
|
| +-------------- Anodonta_nuttalliana
| |
| | +----- Pyganodon_grandis
+--+ +-----+
| | | +-- Utterbackia_imbecillis
| | +--+
+--+ +-- Utterbackia_peninsularis
|
| +-------- Lasmigona_complanata
+--+
| +----- Prolasmidonta_heterodon
+--+
| +-- Alasmidonta_varicosa
+--+
+-- Platynaias_compressa
Wu et al., 2024 [Publ 4350], fig. 8b, S2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 21 |
21 |
19 |
6 |
71 |
65 |
3 |
33.0 |
+-- Quadrula_quadrula
+--------------------------+
| +-- Uniomerus_tetralasmus
|
+--------+ +-- Elliptio_complanata
| | +-----------------------+
| | | +-- Pleurobema_oviforme
| +--+
| | +----------------------- Amblema_plicata
| | |
| +--+ +-------------------- Popenaias_popeii
| | |
| +--+ +----------------- Toxolasma_parvum
| | |
| | | +----- Potamilus_leptodon
| +--+ +--------+
| | | | +-- Potamilus_alatus
| | | +--+
| +--+ +-- Potamilus_streckersoni
| |
| | +----------- Leaunio_lienosus
| | |
| +--+ +-------- Venustaconcha_ellipsiformis
| | |
| | | +-- Lampsilis_powellii
| +--+-----+
| | +-- Lampsilis_siliquoidea
| |
| | +----- Lampsilis_ornata
| +--+
| | +-- Lampsilis_cardium
| +--+
| +-- Lampsilis_cariosa
|
--+ +----- Chamberlainia_somsakpanhai **
| +--------------------+
| | | +-- Sinohyriopsis_cumingii **
| | +--+
| | +-- Sinohyriopsis_schlegelii **
| |
| | +-- Hyriopsis_bialata **
| | +--+
| +--------+ | +-- Rectidens_sumatrensis **
| | | +-----------------+
| | | | | +-- Lens_contradens **
| | | | +--+
| | | | +-- Physunio_superbus **
| | | |
| | | | +----------- Potomida_littoralis **
| | +--+ +--------+
| | | | | +-------- Pronodularia_japanensis **
| | | | +--+
| | | | | +----- Lamprotula_leaii **
| | | | +--+
| | | | | +-- Lamprotula_caveata **
| | +--+ +--+
| | | +-- Lamprotula_cornuumlunae **
| | |
| | | +-- Pseudodon_vondembuschianus **
| | | +--------------+
| | | | +-- Pilsbryoconcha_exilis **
| | +--+
| | | +-------------- Inversidens_rentianensis **
| | | |
+--+ +--+ +-- Parvasolenaia_rivularis **
| | +--------+
| | | +-- Ptychorhynchus_pfisteri **
| +--+
| | +-------- Microcondylaea_bonellii **
| +--+
| | +----- Sinosolenaia_carinata **
| +--+
| | +-- Sinosolenaia_iridinea **
| +--+
| +-- Sinosolenaia_oleivora **
|
| +-------------------------------- Lamellidens_marginalis
| |
| | +-- Lepidodesma_aligera
| | +--------------------------+
| | | +-- Lepidodesma_languilati
| | |
| | | +-------- Aculamprotula_tortuosa
| | | |
| | | +--------------+ +-- Aculamprotula_coreana
| | | | | +--+
+--+ | | | | +-- Aculamprotula_tientsinensis
| | | +--+
| | | | +-- Aculamprotula_polysticta
| | | +--+
| | | +-- Aculamprotula_scripta
| | |
| | +--+ +-------------- Unio_tumidus
| | | | +-----+
| | | | | | +----------- Unio_crassus
| | | | | +--+
+--+ | | | | +-------- Unio_elongatulus
| | | | +--+
| | | | | +----- Unio_pictorum
| | | | +--+
| | | | | +-- Unio_delphinus
| | +--+ +--+
| | | +-- Unio_mancus
| | |
| | | +-- Nodularia_breviconcha
| | | +--------------+
| | | | +-- Nodularia_douglasiae
| | | |
| | | | +-- Pseudocuneopsis_capitata
| | | +--------------+
| | +--+ +-- Pseudocuneopsis_sichuanensis
| | |
| | | +-- Schistodesmus_lampreyanus
+--+ | +-----------+
| | | +-- Schistodesmus_spinosus
| +--+
| | +----------- Tchangsinaia_piscicula
| +--+
| | +-------- Cuneopsis_rufescens
| +--+
| | +----- Cuneopsis_celtiformis
| +--+
| | +-- Cuneopsis_demangei
| +--+
| +-- Cuneopsis_heudei
|
| +-------- Acuticosta_chinensis
| +--------------+
| | | +----- Lanceolaria_gladiola
| | +--+
| | | +-- Lanceolaria_grayii
| | +--+
| | +-- Lanceolaria_lanceolata
| |
| | +-------------- Cristaria_plicata
+--+ |
| +-----+ +-- Anemina_arcaeformis
| | | +--------+
| | | | +-- Anemina_euscaphys
| | +--+
| | | +-------- Beringiana_fukuharai
| | +--+
| | | +----- Sinanodonta_lucida
| | +--+
+--+ | +-- Sinanodonta_tumens
| +--+
| +-- Sinanodonta_woodiana
|
| +----------------- Anodonta_nuttalliana
| |
| | +-------- Pseudanodonta_complanata
| | +-----+
+--+ | | +----- Anodonta_anatina
| | +--+
| | | +-- Anodonta_cygnea
| | +--+
+--+ +-- Anodonta_exulcerata
|
| +----- Pyganodon_grandis
| +-----+
| | | +-- Utterbackia_imbecillis
| | +--+
+--+ +-- Utterbackia_peninsularis
|
| +-------- Lasmigona_complanata
+--+
| +----- Prolasmidonta_heterodon
+--+
| +-- Alasmidonta_varicosa
+--+
+-- Platynaias_compressa
Wu et al., 2023 [Publ 4339], fig. 4, S5-aa
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 20 |
20 |
18 |
6 |
76 |
65 |
3 |
32.0 |
+-- Quadrula_quadrula
+--------------------------+
| +-- Uniomerus_tetralasmus
|
+--------+ +-- Elliptio_complanata
| | +-----------------------+
| | | +-- Pleurobema_oviforme
| +--+
| | +----------------------- Amblema_plicata
| | |
| +--+ +-------------------- Popenaias_popeii
| | |
| +--+ +----------------- Toxolasma_parvum
| | |
| | | +----- Potamilus_leptodon
| +--+ +--------+
| | | | +-- Potamilus_alatus
| | | +--+
| | | +-- Potamilus_streckersoni
| +--+
| | +----------- Leaunio_lienosus
| | |
| | | +----- Lampsilis_ornata
| +--+ +--+
| | | | +-- Lampsilis_cardium
| | | +--+
| +--+ +-- Lampsilis_cariosa
| |
| | +----- Venustaconcha_ellipsiformis
| +--+
| | +-- Lampsilis_powellii
| +--+
| +-- Lampsilis_siliquoidea
--+
| +----- Chamberlainia_somsakpanhai **
| +-----------------+
| | | +-- Sinohyriopsis_cumingii **
| | +--+
| | +-- Sinohyriopsis_schlegelii **
| |
| | +-- Hyriopsis_bialata **
| | +--+
| +-----------+ | +-- Rectidens_sumatrensis **
| | | +--------------+
| | | | | +-- Lens_contradens **
| | | | +--+
| | | | +-- Physunio_superbus **
| | | |
| | | | +----------- Potomida_littoralis **
| | +--+ +-----+
| | | | | +-------- Pronodularia_japanensis **
| | | | +--+
| | | | | +----- Lamprotula_leaii **
| | | | +--+
| | | | | +-- Lamprotula_caveata **
| | +--+ +--+
| | | +-- Lamprotula_cornuumlunae **
| | |
| | | +-- Pseudodon_vondembuschianus **
| | | +-----------+
| | | | +-- Pilsbryoconcha_exilis **
| | +--+
| | | +----------- Microcondylaea_bonellii **
+--+ | |
| +--+ +-- Parvasolenaia_rivularis **
| | +-----+
| | | +-- Ptychorhynchus_pfisteri **
| +--+
| | +----- Sinosolenaia_carinata **
| +--+
| | +-- Sinosolenaia_iridinea **
| +--+
| +-- Sinosolenaia_oleivora **
|
| +-------------------------------- Lamellidens_marginalis
| |
| | +-- Lepidodesma_aligera
| | +--------------------------+
| | | +-- Lepidodesma_languilati
| | |
| | | +-------- Aculamprotula_tortuosa
| | | |
| | | +--------------+ +-- Aculamprotula_coreana
| | | | | +--+
+--+ | | | | +-- Aculamprotula_tientsinensis
| | | +--+
| | | | +-- Aculamprotula_polysticta
| | | +--+
| | | +-- Aculamprotula_scripta
| | |
| | +--+ +-------------- Unio_tumidus
| | | | +-----+
| | | | | | +----------- Unio_crassus
| | | | | +--+
+--+ | | | | +-------- Unio_elongatulus
| | | | +--+
| | | | | +----- Unio_pictorum
| | | | +--+
| | | | | +-- Unio_delphinus
| | +--+ +--+
| | | +-- Unio_mancus
| | |
| | | +-- Nodularia_breviconcha
| | | +--+
| | | | +-- Nodularia_nipponensis
| | | +-----------+
| | | | | +-- Pseudocuneopsis_capitata
| | | | +--+
| | +--+ +-- Pseudocuneopsis_sichuanensis
| | |
| | | +-------------- Tchangsinaia_piscicula
| | | |
+--+ +--+ +-- Schistodesmus_lampreyanus
| | +--------+
| | | +-- Schistodesmus_spinosus
| +--+
| | +-------- Cuneopsis_rufescens
| +--+
| | +----- Cuneopsis_celtiformis
| +--+
| | +-- Cuneopsis_demangei
| +--+
| +-- Cuneopsis_heudei
|
| +-------- Acuticosta_chinensis
| +--------+
| | | +----- Lanceolaria_gladiola
| | +--+
| | | +-- Lanceolaria_grayii
| | +--+
| | +-- Lanceolaria_lanceolata
| |
| | +----- Cristaria_plicata
| | +-----+
| | | | +-- Anemina_arcaeformis
+--------+ | +--+
| +--+ +-- Anemina_euscaphys
| | |
| | | +-------- Beringiana_fukuharai
| | +--+
| | | +----- Sinanodonta_lucida
| | +--+
| | | +-- Sinanodonta_tumens
| | +--+
| | +-- Sinanodonta_woodiana
+--+
| +-- Anodonta_nuttalliana
| +-----+
| | +-- Pseudanodonta_complanata
| +--+
| | | +----- Anodonta_anatina
| | +--+
| | | +-- Anodonta_cygnea
| | +--+
| | +-- Anodonta_exulcerata
+--+
| +----- Pyganodon_grandis
| +--+
| | | +-- Utterbackia_peninsularis
| | +--+
| | +-- Utterbackia_imbecillis
+--+
| +-- Alasmidonta_varicosa
| +--+
| | +-- Platynaias_compressa
+--+
| +-- Prolasmidonta_heterodon
+--+
+-- Lasmigona_complanata
Wu et al., 2023 [Publ 4339], fig. 4, S5-nt
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 20 |
20 |
18 |
6 |
77 |
66 |
3 |
32.0 |
+-- Quadrula_quadrula
+--------------------------+
| +-- Uniomerus_tetralasmus
|
+-----------+ +-- Elliptio_complanata
| | +-----------------------+
| | | +-- Pleurobema_oviforme
| +--+
| | +----------------------- Amblema_plicata
| | |
| +--+ +-------------------- Popenaias_popeii
| | |
| +--+ +----------------- Toxolasma_parvum
| | |
| | | +----- Potamilus_leptodon
| +--+ +--------+
| | | | +-- Potamilus_alatus
| | | +--+
| | | +-- Potamilus_streckersoni
| +--+
| | +----------- Leaunio_lienosus
| | |
| | | +----- Lampsilis_ornata
| +--+ +--+
| | | | +-- Lampsilis_cardium
| | | +--+
| +--+ +-- Lampsilis_cariosa
| |
| | +----- Venustaconcha_ellipsiformis
| +--+
| | +-- Lampsilis_powellii
| +--+
| +-- Lampsilis_siliquoidea
--+
| +----- Chamberlainia_somsakpanhai **
| +-----------------+
| | | +-- Sinohyriopsis_cumingii **
| | +--+
| | +-- Sinohyriopsis_schlegelii **
| |
| | +-- Hyriopsis_bialata **
| | +--+
| +--------------+ | +-- Rectidens_sumatrensis **
| | | +--------------+
| | | | | +-- Lens_contradens **
| | | | +--+
| | | | +-- Physunio_superbus **
| | | |
| | | | +----------- Potomida_littoralis **
| | +--+ +-----+
| | | | | +-------- Pronodularia_japanensis **
| | | | +--+
| | | | | +----- Lamprotula_leaii **
| | | | +--+
| | | | | +-- Lamprotula_caveata **
| | +--+ +--+
| | | +-- Lamprotula_cornuumlunae **
| | |
| | | +-- Pseudodon_vondembuschianus **
| | | +-----------+
| | | | +-- Pilsbryoconcha_exilis **
| | | |
| | +--+ +-- Ptychorhynchus_pfisteri **
+--+ | +--------+
| | | +-- Microcondylaea_bonellii **
| +--+
| | +-------- Parvasolenaia_rivularis **
| +--+
| | +----- Sinosolenaia_carinata **
| +--+
| | +-- Sinosolenaia_iridinea **
| +--+
| +-- Sinosolenaia_oleivora **
|
| +----------------------------------- Lamellidens_marginalis
| |
| | +-- Lepidodesma_aligera
| | +-----------------------------+
| | | +-- Lepidodesma_languilati
| | |
| | | +-------- Acuticosta_chinensis
| | | +--------------+
| | | | | +----- Lanceolaria_gladiola
| | | | +--+
+--+ | | | +-- Lanceolaria_grayii
| | | +--+
| | | +-- Lanceolaria_lanceolata
| | |
| | | +-------------- Cristaria_plicata
| | +-----+ |
| | | | +-----+ +-- Anemina_arcaeformis
| | | | | | +--------+
| | | | | | | +-- Anemina_euscaphys
| | | | | +--+
+--+ | | | | +-------- Beringiana_fukuharai
| | | | +--+
| | | | | +----- Sinanodonta_lucida
| | | | +--+
| | +--+ | +-- Sinanodonta_tumens
| | | +--+
| | | +-- Sinanodonta_woodiana
| | |
| | | +----------------- Anodonta_nuttalliana
| | | |
| | | +----------------- Pseudanodonta_complanata
| | | |
| | +--+ +-------------- Anodonta_cygnea
| | | |
| | | | +-- Anodonta_exulcerata
| | | +-----------+
| | | | +-- Anodonta_anatina
| | +--+
+--+ | +----- Pyganodon_grandis
| | +-----+
| | | | +-- Utterbackia_imbecillis
| | | +--+
| +--+ +-- Utterbackia_peninsularis
| |
| | +-------- Lasmigona_complanata
| +--+
| | +----- Prolasmidonta_heterodon
| +--+
| | +-- Alasmidonta_varicosa
| +--+
| +-- Platynaias_compressa
|
| +-------- Aculamprotula_tortuosa
| |
| +-----------------+ +-- Aculamprotula_coreana
| | | +--+
| | | | +-- Aculamprotula_tientsinensis
| | +--+
| | | +-- Aculamprotula_polysticta
| | +--+
| | +-- Aculamprotula_scripta
| |
+--+ +----------- Unio_tumidus
| |
| +-----------+ +-------- Unio_crassus
| | | |
| | +--+ +----- Unio_elongatulus
| | | |
| | +--+ +-- Unio_delphinus
| | | |
| | +--+-- Unio_mancus
+--+ |
| +-- Unio_pictorum
|
| +-- Pseudocuneopsis_capitata
| +-----------------+
| | +-- Pseudocuneopsis_sichuanensis
| |
+--+ +-- Nodularia_breviconcha
| +--------------+
| | +-- Nodularia_nipponensis
| |
+--+ +----- Tchangsinaia_piscicula
| +--------+
| | | +-- Schistodesmus_lampreyanus[2]
| | +--+
+--+ +-- Schistodesmus_spinosus
|
| +----------- Schistodesmus_lampreyanus[1]
+--+
| +-------- Cuneopsis_rufescens
+--+
| +----- Cuneopsis_celtiformis
+--+
| +-- Cuneopsis_demangei
+--+
+-- Cuneopsis_heudei
Wu et al., 2021 [Publ 4215], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 19 |
19 |
17 |
4 |
5 |
5 |
2 |
43.5 |
+-------------------------- Indonaia_andersoniana
|
| +-------- Acuticosta_chinensis
| +--------------+
| | | +----- Nodularia_douglasiae
--+ | +--+
| | | +-- Inversiunio_jokohamensis
| | +--+
| | +-- Inversiunio_reinianus
| |
+--+ +----- Chamberlainia_somsakpanhai **
| +--------------+
| | | +-- Sundadontina_cumingii **
| | +--+
| | +-- Sinohyriopsis_schlegelii **
| |
| | +-- Indopseudodon_bogani **
| | +--+
+--+ | +-- Indopseudodon_salwenianus **
| +-----------+
| | | +-- Pseudodon_vagulus **
| | +--+
| | +-- Pilsbryoconcha_carinifera **
| |
| | +-------- Sinosolenaia_carinata **
+--+ |
| +-----+ +-- Lamprotula_caveata **
| | | +--+
| | | | +-- Lamprotula_leaii **
| | +--+
| | | +-- Potomida_littoralis **
| | +--+
+--+ +-- Pronodularia_japanensis **
|
| +-- Ptychorhynchus_pfisteri **
| +--------+
| | +-- Parvasolenaia_rivularis **
| |
+--+ +-- Inversidens_brandtii **
| +-----+
| | +-- Inversidens_rentianensis **
+--+
| +----- Gonidea_angulata **
+--+
| +-- Microcondylaea_bonellii **
+--+
+-- Leguminaia_wheatleyi **
Dai et al., 2023 [Publ 4285], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 19 |
19 |
17 |
6 |
34 |
32 |
2 |
31.0 |
+-- Quadrula_quadrula
+--------------------------+
| +-- Uniomerus_tetralasmus
|
+--+ +-- Elliptio_complanata
| | +-----------------------+
| | | +-- Pleurobema_oviforme
| +--+
| | +----------------------- Amblema_plicata
| | |
| +--+ +-------------------- Popenaias_popeii
| | |
| +--+ +----------------- Toxolasma_parvum
| | |
| +--+ +-- Potamilus_leptodon
| | +-----------+
| | | +-- Potamilus_alatus
| +--+
| | +----------- Leaunio_lienosus
| | |
| +--+ +-- Lampsilis_cardium
| | +-----+
| | | +-- Lampsilis_ornata
| +--+
| | +----- Venustaconcha_ellipsiformis
| +--+
| | +-- Lampsilis_powellii
| +--+
--+ +-- Lampsilis_siliquoidea
|
| +-------------------- Lepidodesma_languilati
| |
| | +-------------- Aculamprotula_tientsinensis
| | |
| | +--+ +----------- Unio_crassus
| | | | |
| +--+ | +--+ +-- Pseudocuneopsis_capitata
| | | | | +-----+
| | | | | | +-- Nodularia_douglasiae
| | | | +--+
| | | | | +----- Schistodesmus_lampreyanus
| | | | +--+
| | +--+ | +-- Cuneopsis_celtiformis
| | | +--+
| | | +-- Tchangsinaia_piscicula
| | |
| | | +-- Acuticosta_chinensis
| | | +-----------+
| | | | +-- Lanceolaria_grayii
| | | |
| | +--+ +----- Cristaria_plicata
| | | +-----+
| | | | | +-- Anemina_arcaeformis
+--------+ | | +--+
| +--+ +-- Sinanodonta_woodiana
| |
| | +-------- Anodonta_cygnea
| +--+
| | +----- Platynaias_compressa
| +--+
| | +-- Pyganodon_grandis
| +--+
| +-- Utterbackia_imbecillis
|
| +----- Chamberlainia_somsakpanhai **
| +--------------+
| | | +-- Sinohyriopsis_cumingii **
| | +--+
| | +-- Sinohyriopsis_schlegelii **
| |
| | +-- Hyriopsis_bialata **
+--+ +--+
| | +-- Rectidens_sumatrensis **
| +-----------+
| | | +-- Lens_contradens **
| | +--+
| | +-- Physunio_superbus **
| |
+--+ +-------- Potomida_littoralis **
| +-----+
| | | +----- Pronodularia_japanensis **
| | +--+
| | | +-- Lamprotula_caveata **
| | +--+
+--+ +-- Lamprotula_leaii **
|
| +-- Pseudodon_vondembuschianus **
| +--------+
| | +-- Pilsbryoconcha_exilis **
| |
+--+ +----- Parvasolenaia_rivularis **
| +--+
| | | +-- Postolata_guangxiensis **
| | +--+
+--+ +-- Ptychorhynchus_pfisteri **
|
| +----- Microcondylaea_bonellii **
+--+
| +-- Sinosolenaia_oleivora **
+--+
+-- Sinosolenaia_carinata **
Huang et al., 2019 [Publ 3843], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 18 |
18 |
13 |
3 |
57 |
55 |
1 |
40.0 |
+----- Sinohyriopsis_cumingii[2] **
+--------------+
| | +-- Sinohyriopsis_cumingii[1] **
| +--+
| +-- Sinohyriopsis_schlegelii **
|
| +----- Parvasolenaia_rivularis **
+--------------+ +-----------+
| | | | +-- Parvasolenaia_triangularis **
| | | +--+
| | | +-- Ptychorhynchus_pfisteri **
| | |
| | | +-------- Sinosolenaia_recognita **
| +--+ |
| | +-----+ +----- Sinosolenaia_oleivora[2] **
| | | | |
| | | +--+ +-- Sinosolenaia_oleivora[1] **
| | | | |
| | | +--+-- Sinosolenaia_iridinea **
| +--+ |
| | +-- Sinosolenaia_oleivora[3] **
| |
| | +----------- Sinosolenaia_carinata **
| | |
| +--+ +-------- Lamprotula_leaii **
| | |
| +--+ +----- Lamprotula_cornuumlunae[1] **
| | |
| | +----- Lamprotula_cornuumlunae[3] **
| +--+
| +----- Lamprotula_cornuumlunae[4] **
--+ |
| | +-- Lamprotula_cornuumlunae[2] **
| +--+
| +-- Lamprotula_caveata **
|
| +-------------- Anemina_arcaeformis[2]
| |
| +-----------------+ +----------- Anemina_arcaeformis[5]
| | | |
| | | | +-------- Anemina_arcaeformis[3]
| | +--+ |
| | | +-------- Anemina_arcaeformis[7]
| | | |
| | +--+ +----- Anemina_arcaeformis[4]
| | | |
| | | +----- Anemina_arcaeformis[6]
| | | |
| | +--+----- Anemina_arcaeformis[8]
| | |
| | +----- Anemina_arcaeformis[9]
| | |
| | | +-- Anemina_arcaeformis[1]
| | +--+
| | +-- Anemina_euscaphys
+--+
| +-- Sinanodonta_angula
| +-----------+
| | +-- Sinanodonta_lucida
| +--------------+
| | | +----------- Sinanodonta_woodiana[2]
| | +--+
| | | +-------- Sinanodonta_jourdyi
| | +--+
| | | +----- Sinanodonta_elliptica
| | +--+
| | | +-- Sinanodonta_woodiana[1]
| | +--+
| | +-- Sinanodonta_lauta
| |
| | +-------------- Acuticosta_ovata[1]
| | |
| | | +----------- Acuticosta_chinensis[3]
+--+ +-----------+ |
| | | +----------- Acuticosta_chinensis[4]
| | | |
| | +--+----------- Acuticosta_chinensis[6]
| | |
| | | +-- Acuticosta_chinensis[1]
| | | +-----+
| | | | +-- Acuticosta_ovata[2]
| | +--+
| | | +----- Acuticosta_chinensis[5]
| | | |
| | +--+----- Acuticosta_chinensis[7]
| | |
| | | +-- Acuticosta_chinensis[2]
+--+ +--+
| +-- Acuticosta_ovata[3]
|
| +-- Lepidodesma_languilati
| +--------------------+
| | +-- Cristaria_plicata
| |
| | +----------- Lanceolaria_gladiola
| | |
| | | +-------- Lanceolaria_grayii[5]
| | +--------+ |
| | | | | +-- Lanceolaria_lanceolata
| | | | +-----+
| | | | | +-- Lanceolaria_triformis
+--+ | +--+
| | | +----- Lanceolaria_grayii[4]
| | | |
| | | | +-- Lanceolaria_eucylindrica[1]
| | | | |
| | +--+ +-- Lanceolaria_eucylindrica[2]
| | | |
| | | +-- Lanceolaria_eucylindrica[3]
| | | |
| | +--+-- Lanceolaria_eucylindrica[4]
+--+ |
| +-- Lanceolaria_grayii[1]
| |
| +-- Lanceolaria_grayii[2]
| |
| +-- Lanceolaria_grayii[3]
|
| +----------- Aculamprotula_tortuosa
| +-----+
| | | +-------- Aculamprotula_tientsinensis
| | +--+
| | | +----- Aculamprotula_fibrosa
| | +--+
| | | +-- Aculamprotula_coreana
+--+ +--+
| +-- Aculamprotula_scripta
|
| +----- Nodularia_douglasiae[1]
| +--------+
| | | +-- Nodularia_nuxpersicae
| | +--+
+--+ +-- Nodularia_douglasiae[2]
|
| +----------- Tchangsinaia_piscicula
| |
+--+ +-- Schistodesmus_spinosus
| +-----+
| | +-- Schistodesmus_lampreyanus
+--+
| +----- Cuneopsis_rufescens
+--+
| +-- Cuneopsis_heudei
+--+
+-- Cuneopsis_celtiformis
Pfeiffer et al., 2019 [Publ 3937], fig. 4-5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 17 |
17 |
15 |
6 |
40 |
33 |
3 |
26.0 |
+-- Quadrula_quadrula
+--------------------+
| +-- Uniomerus_tetralasmus
|
| +----- Elliptoideus_sloatianus
+-----+ +--------------+
| | | | +-- Elliptio_complanata
| | | +--+
| | | +-- Pleurobema_strodeanum
| | |
| +--+ +----- Reginaia_ebenus
| | +-----------+
| | | | +-- Plectomerus_dombeyanus
| | | +--+
| | | +-- Amblema_plicata
| +--+
| | +-- Psoronaias_semigranosa
| | +-----------+
| | | +-- Popenaias_popeii
| +--+
| | +----------- Obliquaria_reflexa
| +--+
| | +-------- Cyrtonaias_tampicoensis
| +--+
| | +----- Leaunio_lienosus
| +--+
| | +-- Lampsilis_cardium
| +--+
| +-- Pachynaias_spheniopsis
|
--+ +-- Chamberlainia_hainesiana **
| +--------------------+
| | +-- Sinohyriopsis_cumingii **
| |
| | +----------------- Lamprotula_cornuumlunae **
| | +--+
| | | | +-------------- Gonidea_angulata **
| +--+ | +--+
| | | | | +----------- Indopseudodon_crebristriatus **
| | | | +--+
| | | | | +-------- Bineurus_anodontinum **
| | | | +--+
| | | | | +----- Pilsbryoconcha_exilis **
| | | | +--+
| | +--+ | +-- Pseudodon_vondembuschianus **
| | | +--+
| | | +-- Pseudodon_inoscularis **
| | |
| | | +----- Ensidens_ingallsianus **
| | | +-----+
| | | | | +-- Hyriopsis_bialata **
| | | | +--+
| | | | +-- Rectidens_sumatrensis **
| | +--------+
| | | +-- Trapezoideus_foliaceus **
| | | +-----+
+--+ | | +-- Physunio_superbus **
| +--+
| | +----- Yaukthwa_paiensis **
| +--+
| | +-- Lens_contradens **
| +--+
| +-- Solenaia_emarginata **
|
| +----------- Lanceolaria_lanceolata
| |
| | +----- Unio_gibbus
| +--------+ +--+
| | | | | +-- Nodularia_douglasiae
| | | | +--+
| | | | +-- Tchangsinaia_piscicula
| | +--+
| | | +-- Cristaria_plicata
| | | +--+
| | | | +-- Sinanodonta_woodiana
+-----+ +--+
| | +-- Anodonta_cygnea
| +--+
| +-- Pyganodon_grandis
|
| +-- Trapezidens_dolichorhynchus
| +--------------+
| | +-- Lamellidens_savadiensis
| |
+--+ +-- Leoparreysia_tavoyensis
| +-----------+
| | +-- Leoparreysia_olivacea
| |
+--+ +-- Coelatura_choziensis
| +--------+
| | +-- Prisodontopsis_aviculaeformis
+--+
| +-------- Indonaia_caerulea
+--+
| +----- Indochinella_pugio
+--+
| +-- Scabies_phaselus
+--+
+-- Harmandia_somboriensis
Pfeiffer et al., 2019 [Publ 3937], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 17 |
17 |
15 |
6 |
40 |
33 |
3 |
26.0 |
+-- Quadrula_quadrula
+-----------------+
| +-- Uniomerus_tetralasmus
|
| +----- Elliptoideus_sloatianus
| +--+
| | | +-- Elliptio_complanata
+--+ | +--+
| | +--------+ +-- Pleurobema_strodeanum
| | | |
| | | | +----- Reginaia_ebenus
| | | +--+
| | | | +-- Plectomerus_dombeyanus
| +--+ +--+
| | +-- Amblema_plicata
| |
| | +-- Psoronaias_semigranosa
| | +-----------+
| | | +-- Popenaias_popeii
| +--+
| | +----------- Obliquaria_reflexa
| +--+
| | +-------- Cyrtonaias_tampicoensis
| +--+
| | +----- Leaunio_lienosus
| +--+
| | +-- Lampsilis_cardium
| +--+
| +-- Pachynaias_spheniopsis
|
| +-- Chamberlainia_hainesiana **
| +-----------------+
| | +-- Sinohyriopsis_cumingii **
| |
| | +-- Gonidea_angulata **
| | +-----------+
| | | +-- Lamprotula_cornuumlunae **
| | +--+
+--+ | | +----------- Indopseudodon_crebristriatus **
| | | +--+
| | | | +-------- Pilsbryoconcha_exilis **
| | | +--+
--+ | | | +----- Bineurus_anodontinum **
| | | +--+
| | | | +-- Pseudodon_vondembuschianus **
| +--+ +--+
| | +-- Pseudodon_inoscularis **
| |
| | +----- Ensidens_ingallsianus **
| | +-----+
| | | | +-- Hyriopsis_bialata **
| | | +--+
| | | +-- Rectidens_sumatrensis **
| +-----+
| | +-- Trapezoideus_foliaceus **
| | +-----+
| | | +-- Physunio_superbus **
| +--+
| | +----- Yaukthwa_paiensis **
| +--+
| | +-- Lens_contradens **
| +--+
| +-- Solenaia_emarginata **
|
| +-------- Lanceolaria_lanceolata
| +--+
| | | +----- Unio_gibbus
| | +--+
| | | +-- Tchangsinaia_piscicula
| | +--+
| +--------+ +-- Nodularia_douglasiae
| | |
| | | +-- Cristaria_plicata
| | | +--+
| | | | +-- Sinanodonta_woodiana
| | +-----+
| | | +-- Anodonta_cygnea
+--+ +--+
| +-- Pyganodon_grandis
|
| +-- Trapezidens_dolichorhynchus
| +--------------+
| | +-- Lamellidens_savadiensis
| |
+--+ +-- Leoparreysia_tavoyensis
| +-----------+
| | +-- Leoparreysia_olivacea
| |
+--+ +-- Coelatura_choziensis
| +--------+
| | +-- Prisodontopsis_aviculaeformis
+--+
| +-------- Indonaia_caerulea
+--+
| +----- Indochinella_pugio
+--+
| +-- Scabies_phaselus
+--+
+-- Harmandia_somboriensis
Zieritz et al., 2021 [Publ 4231], fig. 3-F, 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 17 |
17 |
15 |
6 |
32 |
29 |
2 |
29.0 |
+----------- Quadrula_quadrula
|
+-----------------+ +-------- Toxolasma_parvum
| | |
| +--+ +-- Potamilus_alatus
| | +--+
| | | +-- Potamilus_leptodon
| +--+
| | +-- Venustaconcha_ellipsiformis
| +--+
| +-- Lampsilis_ornata
|
| +----- Chamberlainia_somsakpanhai **
| +--------------+
| | | +-- Sinohyriopsis_cumingii **
| | +--+
| | +-- Sinohyriopsis_schlegelii **
| |
| | +-- Lens_contradens **
--+ +-----+ +--+
| | | | +-- Physunio_superbus **
| | | +-----------+
| | | | | +-- Hyriopsis_bialata **
| | | | +--+
| | | | +-- Rectidens_sumatrensis **
| | | |
| | +--+ +-------- Potomida_littoralis **
| | | +-----+
| | | | | +----- Pronodularia_japanensis **
| | | | +--+
| | | | | +-- Lamprotula_caveata **
| | | | +--+
| | +--+ +-- Lamprotula_leaii **
| | |
| | | +-- Pilsbryoconcha_exilis **
| | | +--------+
+--+ | | +-- Pseudodon_vondembuschianus **
| +--+
| | +-------- Ptychorhynchus_pfisteri **
| +--+
| | +----- Microcondylaea_bonellii **
| +--+
| | +-- Sinosolenaia_recognita **
| +--+
| +-- Sinosolenaia_carinata **
|
| +----------------------- Lepidodesma_languilati
| |
| | +----- Aculamprotula_tortuosa
| | +--------+
| | | | +-- Aculamprotula_tientsinensis
| | | +--+
| | | +-- Aculamprotula_coreana
| | +-----+
+--+ | | +----- Nodularia_douglasiae
| | | +-----+
| | | | | +-- Schistodesmus_lampreyanus
| | | | +--+
| | +--+ +-- Tchangsinaia_piscicula
| | |
| | | +-------- Unio_tumidus
| | +--+
| | | +----- Unio_crassus
+--+ +--+
| | +-- Unio_pictorum
| +--+
| +-- Unio_delphinus
|
| +-- Lanceolaria_grayii
| +--------------+
| | +-- Lanceolaria_lanceolata
| |
| | +----------- Anodonta_anatina
+--+ +--+
| | | +-------- Platynaias_compressa
| | +--+
| | | +----- Pyganodon_grandis
| | +--+
+--+ | +-- Utterbackia_imbecillis
| +--+
| +-- Utterbackia_peninsularis
|
| +-------- Cristaria_plicata
| |
+-----+ +-- Sinanodonta_lucida
| +--+
| | +-- Sinanodonta_woodiana
+--+
| +-- Anemina_arcaeformis
+--+
+-- Anemina_euscaphys
Bolotov et al., 2022 [Publ 4250], fig. S5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 15 |
15 |
12 |
1 |
0 |
0 |
0 |
35.0 |
+-- Physunio_modelli **
+-----+
| +-- Physunio_superbus **
+--------+
| | +----- Trapezoideus_foliaceus **
| +--+
| | +-- Trapezoideus_lenya **
| +--+
| +-- Trapezoideus_mitanensis **
--+
| +-- Yaukthwa_elongatula **
| +-----------+
| | +-- Yaukthwa_baniensis **
| |
| | +-------- Yaukthwa_panhai **
+--+ +--+
| | | +----- Yaukthwa_rectangularis **
| | +--+
| | | +-- Yaukthwa_nesemanni **
+--+ +--+
| +-- Yaukthwa_zayleymanensis **
|
| +----- Yaukthwa_dalliana **
| |
+-----+ +-- Yaukthwa_paiensis **
| |
+--+-- Yaukthwa_inlenensis **
|
+-- Yaukthwa_avaensis **
Kongim et al., 2023 [Publ 4290], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 15 |
15 |
13 |
4 |
0 |
0 |
0 |
34.0 |
+----- Chamberlainia_somsakpanhai **
+--+
| | +-- Chamberlainia_hainesiana **
| +--+
+-----+ +-- Chamberlainia_duclerci **
| |
| | +----- Sinohyriopsis_cumingii **
| +--+
| | +-- Sinohyriopsis_schlegelii **
| +--+
--+ +-- Sinohyriopsis_goliath **
|
| +----- Lamprotula_leaii **
| +-----+
| | | +-- Pronodularia_japanensis **
| | +--+
| | +-- Potomida_littoralis **
+--+
| +----- Sinosolenaia_carinata **
| +--+
| | | +-- Leguminaia_wheatleyi **
| | +--+
+--+ +-- Gonidea_angulata **
|
| +----- Indopseudodon_indawgyiensis **
+--+
| +-- Namkongnaia_inkhavilayi **
+--+
+-- Pilsbryoconcha_exilis **
Pfeiffer & Graf, 2015 [Publ 3620], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
14 |
12 |
5 |
26 |
16 |
3 |
33.0 |
+----- Unio_pictorum
+-----------------+
| | +-- Pyganodon_grandis
| +--+
| +-- Lasmigona_costata
|
| +----------- Quadrula_quadrula
| +--------+
--+ | | +-------- Amblema_plicata
| | +--+
| | | +----- Potamilus_alatus
| | +--+
| | | +-- Lampsilis_cardium
| | +--+
| | +-- Truncilla_truncata
+--+
| +-------- Ensidens_telus **
| +--------+
| | | +----- Lens_micropterus **
| | +--+
| | | +-- Physunio **
| | +--+
| | +-- Physunio_modelli **
+--+
| +-------- Gonidea_angulata **
| +-----+
| | | +----- Potomida_littoralis **
| | +--+
| | | +-- Pronodularia_japanensis **
+--+ +--+
| +-- Lamprotula_caveata **
|
| +----------- Pilsbryoconcha_carinifera **
| |
+--+ +-- Pseudodon_vondembuschianus **
| +-----+
| | +-- Pseudodon_vagulus **
+--+
| +----- Sundadontina_tumida **
+--+
| +-- Nyeinchanconcha_nyeinchani **
+--+
+-- Bineurus_exilis **
Huang et al., 2019 [Publ 3843], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
42 |
39 |
3 |
30 |
28 |
1 |
66.0 |
+----- Lamprotula_leaii **
|
+-----------+ +-- Lamprotula_caveata[1] **
| | |
| +--+-- Lamprotula_cornuumlunae **
| |
+--------------------+ +-- Lamprotula_caveata[2] **
| |
| | +-- Parvasolenaia_rivularis **
| | +-----------+
| | | +-- Parvasolenaia_triangularis **
| +--+
| | +----------- Ptychorhynchus_pfisteri **
| +--+
| | +-------- Sinosolenaia_carinata **
| +--+
--+ | +----- Sinosolenaia_recognita **
| +--+
| | +-- Sinosolenaia_oleivora **
| +--+
| +-- Sinosolenaia_iridinea **
|
| +----- Sinohyriopsis_cumingii[1] **
| +-----------------------------+
| | | +-- Sinohyriopsis_cumingii[2] **
| | +--+
| | +-- Sinohyriopsis_schlegelii **
+--+
| +-------------------------------- Anemina_arcaeformis
| |
| | +----------------------------- Acuticosta_chinensis
| | |
+--+ | +-- Sinanodonta_angula
| | +--------+
| | | +-- Sinanodonta_lucida
| | +--------------+
| | | | +-------- Sinanodonta_jourdyi
+--+ | +--+
| | | +----- Sinanodonta_woodiana[2]
| | +--+
| | | +-- Sinanodonta_elliptica
| | +--+
| | +-- Sinanodonta_woodiana[1]
| |
+--+ +----------- Aculamprotula_tortuosa
| +-----------+
| | | +-------- Aculamprotula_scripta
| | +--+
| | | +----- Aculamprotula_coreana
| | +--+
| | | +-- Aculamprotula_fibrosa
| | +--+
| | +-- Aculamprotula_tientsinensis
+--+
| +----------- Cristaria_plicata
| +--------+
| | | +-------- Lanceolaria_gladiola
| | +--+
| | | +----- Lanceolaria_grayii
| | +--+
| | | +-- Lanceolaria_lanceolata
+--+ +--+
| +-- Lanceolaria_triformis
|
| +----------------- Lepidodesma_languilati
| |
| | +-- Schistodesmus_lampreyanus
+--+ +-----------+
| | +-- Schistodesmus_spinosus
| |
+--+ +----- Nodularia_douglasiae[1]
| +-----+
| | | +-- Nodularia_douglasiae[2]
| | +--+
+--+ +-- Nodularia_nuxpersicae
|
| +-------- Cuneopsis_rufescens
+--+
| +----- Tchangsinaia_piscicula
+--+
| +-- Cuneopsis_heudei
+--+
+-- Cuneopsis_celtiformis
Wu et al., 2022 [Publ 4238], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 14 |
14 |
12 |
4 |
60 |
51 |
2 |
32.0 |
+-- Uniomerus_tetralasmus
+--------------------------+
| +-- Quadrula_quadrula
|
+--------+ +-- Pleurobema_oviforme
| | +-----------------------+
| | | +-- Elliptio_complanata
| +--+
| | +----------------------- Amblema_plicata
| | |
| +--+ +-------------------- Popenaias_popeii
| | |
| +--+ +----------------- Toxolasma_parvum
| | |
| +--+ +-- Potamilus_alatus
| | +-----------+
| | | +-- Potamilus_leptodon
| +--+
| | +----------- Leaunio_lienosus
| | |
| +--+ +-- Lampsilis_cardium
| | +-----+
| | | +-- Lampsilis_ornata
| +--+
| | +----- Venustaconcha_ellipsiformis
| +--+
| | +-- Lampsilis_siliquoidea
| +--+
--+ +-- Lampsilis_powellii
|
| +----- Chamberlainia_somsakpanhai **
| +-----------+
| | | +-- Sinohyriopsis_schlegelii **
| | +--+
| | +-- Sinohyriopsis_cumingii **
| |
| +-----------------+ +-------- Potomida_littoralis **
| | | +-----+
| | | | | +----- Pronodularia_japanensis **
| | | | +--+
| | | | | +-- Lamprotula_caveata **
| | | | +--+
| | +--+ +-- Lamprotula_leaii **
| | |
| | | +-- Pilsbryoconcha_exilis **
| | | +--------+
| | | | +-- Pseudodon_vondembuschianus **
| | | |
| | +--+ +-- Ptychorhynchus_pfisteri **
| | | +-----+
| | | | +-- Parvasolenaia_rivularis **
| | +--+
| | | +----- Microcondylaea_bonellii **
+--+ +--+
| | +-- Sinosolenaia_carinata **
| +--+
| +-- Sinosolenaia_recognita **
|
| +-------------------------------- Lepidodesma_languilati
| |
| | +-------- Acuticosta_chinensis
| | +--------+
| | | | +----- Lanceolaria_gladiola
| | | +--+
| | | | +-- Lanceolaria_grayii
| | | +--+
| | | +-- Lanceolaria_lanceolata
| | |
| | +-----------+ +-------- Cristaria_plicata
| | | | |
+--+ | | +-----+ +-- Sinanodonta_woodiana
| | | | | +--+
| | | | | | +-- Sinanodonta_lucida
| | | | +--+
| | | | | +-- Anemina_arcaeformis
| | +--+ +--+
| | | +-- Anemina_euscaphys
| | |
| | | +-- Anodonta_cygnea
| | | +--------+
| | | | +-- Anodonta_anatina
| | +--+
| | | +-------- Platynaias_compressa
+--+ +--+
| | +----- Pyganodon_grandis
| +--+
| | +-- Utterbackia_imbecillis
| +--+
| +-- Utterbackia_peninsularis
|
| +-------- Aculamprotula_tortuosa
| |
| +-----------------+ +-- Aculamprotula_tientsinensis
| | | +--+
| | | | +-- Aculamprotula_coreana
| | +--+
| | | +-- Aculamprotula_polysticta
| | +--+
| | +-- Aculamprotula_scripta
+--+
| +-------- Unio_tumidus
| +--------------+
| | | +----- Unio_crassus
| | +--+
| | | +-- Unio_pictorum
| | +--+
+--+ +-- Unio_delphinus
|
| +-- Pseudocuneopsis_capitata
| +-----------------+
| | +-- Pseudocuneopsis_sichuanensis
| |
+--+ +-- Nodularia_douglasiae
| +--------------+
| | +-- Nodularia_breviconcha
| |
+--+ +-- Schistodesmus_lampreyanus
| +-----------+
| | +-- Schistodesmus_spinosus
+--+
| +----------- Tchangsinaia_piscicula
+--+
| +-------- Cuneopsis_rufescens
+--+
| +----- Cuneopsis_celtiformis
+--+
| +-- Cuneopsis_heudei
+--+
+-- Cuneopsis_demangei
Froufe et al., 2019 [Publ 3936], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 13 |
13 |
11 |
4 |
32 |
29 |
2 |
31.0 |
+----------- Quadrula_quadrula
|
+-----------------+ +-------- Toxolasma_parvum
| | |
| +--+ +-- Lampsilis_ornata
| | +--+
| | | +-- Venustaconcha_ellipsiformis
| +--+
| | +-- Potamilus_leptodon
| +--+
| +-- Potamilus_alatus
|
| +----- Chamberlainia_somsakpanhai **
| +-----------+
| | | +-- Sinohyriopsis_cumingii **
| | +--+
| | +-- Sinohyriopsis_schlegelii **
--+ |
| +--------+ +-------- Potomida_littoralis **
| | | +-----+
| | | | | +----- Pronodularia_japanensis **
| | | | +--+
| | | | | +-- Lamprotula_leaii **
| | | | +--+
| | +--+ +-- Lamprotula_caveata **
| | |
| | | +-- Pilsbryoconcha_exilis **
| | | +--------+
| | | | +-- Pseudodon_vondembuschianus **
| | +--+
| | | +-------- Ptychorhynchus_pfisteri **
| | +--+
+--+ | +----- Microcondylaea_bonellii **
| +--+
| | +-- Sinosolenaia_carinata **
| +--+
| +-- Sinosolenaia_oleivora **
|
| +----------------------- Lepidodesma_languilati
| |
| | +----- Aculamprotula_tortuosa
| | +--------+
| | | | +-- Aculamprotula_tientsinensis
| | | +--+
| | | +-- Aculamprotula_coreana
| | +-----+
+--+ | | +----- Nodularia_douglasiae
| | | +-----+
| | | | | +-- Tchangsinaia_piscicula
| | | | +--+
| | +--+ +-- Schistodesmus_lampreyanus
| | |
| | | +-------- Unio_tumidus
| | +--+
| | | +----- Unio_crassus
+--+ +--+
| | +-- Unio_pictorum
| +--+
| +-- Unio_delphinus
|
| +-- Lanceolaria_lanceolata
| +--------------+
| | +-- Lanceolaria_grayii
| |
| | +-------- Cristaria_plicata
+--+ |
| +-----+ +-- Anemina_arcaeformis
| | | +--+
| | | | +-- Anemina_euscaphys
| | +--+
+--+ | +-- Sinanodonta_lucida
| +--+
| +-- Sinanodonta_woodiana
|
| +----------- Anodonta_anatina
+--+
| +-------- Lasmigona_complanata
+--+
| +----- Pyganodon_grandis
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Utterbackia_imbecillis
Froufe et al., 2019 [Publ 3936], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 13 |
13 |
11 |
4 |
32 |
29 |
2 |
31.0 |
+----------- Quadrula_quadrula
|
+-----------------+ +-------- Toxolasma_parvum
| | |
| +--+ +-- Potamilus_leptodon
| | +--+
| | | +-- Potamilus_alatus
| +--+
| | +-- Lampsilis_ornata
| +--+
| +-- Venustaconcha_ellipsiformis
|
| +----- Chamberlainia_somsakpanhai **
| +-----------+
| | | +-- Sinohyriopsis_cumingii **
| | +--+
| | +-- Sinohyriopsis_schlegelii **
--+ |
| +--------+ +-------- Potomida_littoralis **
| | | +-----+
| | | | | +----- Pronodularia_japanensis **
| | | | +--+
| | | | | +-- Lamprotula_leaii **
| | | | +--+
| | +--+ +-- Lamprotula_caveata **
| | |
| | | +-- Pilsbryoconcha_exilis **
| | | +--------+
| | | | +-- Pseudodon_vondembuschianus **
| | +--+
| | | +-------- Ptychorhynchus_pfisteri **
| | +--+
+--+ | +----- Microcondylaea_bonellii **
| +--+
| | +-- Sinosolenaia_carinata **
| +--+
| +-- Sinosolenaia_oleivora **
|
| +----------------------- Lepidodesma_languilati
| |
| | +----- Aculamprotula_tortuosa
| | +--------+
| | | | +-- Aculamprotula_tientsinensis
| | | +--+
| | | +-- Aculamprotula_coreana
| | +-----+
+--+ | | +----- Nodularia_douglasiae
| | | +-----+
| | | | | +-- Tchangsinaia_piscicula
| | | | +--+
| | +--+ +-- Schistodesmus_lampreyanus
| | |
| | | +-------- Unio_tumidus
| | +--+
| | | +----- Unio_crassus
+--+ +--+
| | +-- Unio_pictorum
| +--+
| +-- Unio_delphinus
|
| +-- Lanceolaria_lanceolata
| +--------------+
| | +-- Lanceolaria_grayii
| |
| | +----------- Anodonta_anatina
+--+ +--+
| | | +-------- Lasmigona_complanata
| | +--+
| | | +----- Pyganodon_grandis
| | +--+
+--+ | +-- Utterbackia_peninsularis
| +--+
| +-- Utterbackia_imbecillis
|
| +-------- Cristaria_plicata
| |
+-----+ +-- Sinanodonta_lucida
| +--+
| | +-- Sinanodonta_woodiana
+--+
| +-- Anemina_arcaeformis
+--+
+-- Anemina_euscaphys
Lopes-Lima et al., 2020 [Publ 4080], fig. 17
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 13 |
13 |
3 |
1 |
2 |
0 |
0 |
30.0 |
+----------- Chamberlainia_somsakpanhai **
|
| +-------- Sinohyriopsis_cumingii[10] **
--+ |
| | +----- Sinohyriopsis_cumingii[1] **
| | |
| | +----- Sinohyriopsis_cumingii[2] **
+--+ |
| +----- Sinohyriopsis_cumingii[3] **
| |
| +----- Sinohyriopsis_cumingii[4] **
| |
| +----- Sinohyriopsis_cumingii[5] **
+--+
+----- Sinohyriopsis_cumingii[6] **
|
+----- Sinohyriopsis_cumingii[7] **
|
+----- Sinohyriopsis_cumingii[8] **
|
+----- Sinohyriopsis_cumingii[9] **
|
| +-- Sinohyriopsis_schlegelii **
+--+
+-- Sinohyriopsis_goliath **
Konopleva et al., 2023 [Publ 4269], fig. S1, 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 13 |
13 |
11 |
3 |
43 |
42 |
3 |
35.0 |
+-- Lampsilis_ovata
+--------------+
| +-- Amblema_plicata
|
| +----------- Sinosolenaia_recognita **
| |
| +--+ +-- Ptychorhynchus_pfisteri **
+-----------+ | | +-----+
| | | | | +-- Parvasolenaia_rivularis **
| | | +--+
| | | | +----- Gonidea_angulata **
| | | +--+
| | | | +-- Microcondylaea_bonellii **
| +--+ +--+
| | +-- Leguminaia_wheatleyi **
| |
| | +----- Chamberlainia_somsakpanhai **
| | +-----+
| | | | +-- Sinohyriopsis_schlegelii **
| | | +--+
| +--+ +-- Sinohyriopsis_cumingii **
| |
| | +-------- Potomida_littoralis **
| +--+
| | +----- Pronodularia_japanensis **
| +--+
| | +-- Lamprotula_caveata **
--+ +--+
| +-- Lamprotula_leaii **
|
| +----------------- Middendorffinaia_mongolica
| |
| | +-------- Aculamprotula_tortuosa
| | +-----+
| +--------+ | | +----- Aculamprotula_scripta
| | | | +--+
| | | | | +-- Aculamprotula_tientsinensis
| | | | +--+
| | +--+ +-- Aculamprotula_coreana
| | |
| | | +----- Unio_tumidus
| | | +-----+
| | | | | +-- Unio_pictorum
| | | | +--+
| | +--+ +-- Unio_crassus
| | |
| | | +-------- Nodularia_douglasiae
+--+ +--+
| | +----- Tchangsinaia_piscicula
| +--+
| | +-- Cuneopsis_rufescens
| +--+
| +-- Cuneopsis_heudei
|
| +-------- Lepidodesma_languilati
| +--------------+
| | | +----- Lanceolaria_gladiola
| | +--+
| | | +-- Lanceolaria_grayii
| | +--+
+--+ +-- Lanceolaria_triformis
|
| +-------------------- Acuticosta_chinensis
| |
| | +----- Anodonta_anatina
| | +-----------+
+--+ | | +-- Pseudanodonta_complanata
| | +--+
| | +-- Anodonta_cygnea
| |
| | +-------- Anemina_arcaeformis
+--+ +-----+
| | | +----- Amuranodonta_kijaensis
| | +--+
| | | +-- Buldowskia_suifunica
| | +--+
| | +-- Buldowskia_shadini
+--+
| +-------- Beringiana_beringiana
| +--+
| | | +----- Sinanodonta_schrenkii
| | +--+
| | | +-- Sinanodonta_woodiana
+--+ +--+
| +-- Sinanodonta_lauta
|
| +-------- Cristaria_radiata
| |
+--+ +-- Cristaria_truncata
| +--+
| | +-- Cristaria_plicata
+--+
| +-- Cristaria_bellua
+--+
+-- Cristaria_clessini
Jeratthitikul & Sutcharit, 2023 [Publ 4289], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 13 |
13 |
11 |
1 |
0 |
0 |
0 |
34.5 |
+-- Indopseudodon_indawgyiensis **
+--------------+
| +-- Indopseudodon_kayinensis **
|
| +----------- Bineurus_mouhotii **
| +--+
--+ | | +-------- Bineurus_loeiensis **
| | +--+
| | | +----- Bineurus_panhai **
| | +--+
| | | +-- Bineurus_anodontinum **
+--+ +--+
| +-- Bineurus_exilis **
|
| +----- Namkongnaia_inkhavilayi **
| +--+
| | | +-- Sundadontina_cumingii **
| | +--+
+-----+ +-- Nyeinchanconcha_nyeinchani **
|
| +----- Pseudodon_vondembuschianus **
+--+
| +-- Pilsbryoconcha_exilis **
+--+
+-- Thaiconcha_callifera **
Pfeiffer & Graf, 2013 [Publ 3531], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
13 |
10 |
5 |
23 |
20 |
3 |
33.0 |
+-- Cristaria_plicata
+-----+
| +-- Sinanodonta_woodiana
+--------+
| | +----- Strophitus_undulatus
| +--+
| | +-- Platynaias_compressa
+-----------+ +--+
| | +-- Pyganodon_grandis
| |
| | +-------------- Acuticosta_chinensis
| | |
| +--+ +-- Unio_pictorum
| | +--------+
| | | +-- Nodularia_douglasiae
| +--+
| | +-------- Aculamprotula_tortuosa
| | |
| +--+ +----- Aculamprotula_scripta
| | |
--+ +--+ +-- Aculamprotula_tientsinensis
| | |
| +--+-- Aculamprotula_fibrosa
| |
| +-- Aculamprotula_zonata
|
| +----------- Quadrula_quadrula
| +--------------+
| | | +-------- Eurynia_dilatata
| | +--+
| | | +----- Amblema_plicata
| | +--+
| | | +-- Lampsilis_ornata
+--+ +--+
| +-- Truncilla_truncata
|
| +-- Sinohyriopsis_cumingii **
| +--------------------+
| | +-- Sinohyriopsis_schlegelii **
| |
+--+ +----- Gonidea_angulata **
| +--------------+
| | | +-- Pseudodon_vagulus **
| | +--+
+--+ +-- Sundadontina_tumida **
|
| +----------------- Physunio **
| |
+--+ +-------------- Pronodularia_japanensis **
| |
+--+ +----------- Potomida_littoralis **
| |
+--+ +-------- Lamprotula_leaii **
| |
+--+ +----- Lamprotula_caveata[2] **
| |
+--+ +-- Lamprotula_cornuumlunae **
| |
+--+-- Gibbosula_rochechouartii
|
+-- Lamprotula_caveata[1] **
Fukata & Iigo, 2020 [Publ 4106], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
10 |
4 |
42 |
39 |
2 |
34.0 |
+-- Quadrula_quadrula
+-----------------+
| +-- Uniomerus_tetralasmus
+--+
| | +----------------- Elliptio_complanata
| | |
| +--+ +-- Potamilus_leptodon
| | +-----------+
| | | +-- Potamilus_alatus
| +--+
| | +----------- Leaunio_lienosus
| | |
| +--+ +-- Lampsilis_cardium
| | +-----+
| | | +-- Lampsilis_ornata
+--+ +--+
| | | +----- Venustaconcha_ellipsiformis
| | +--+
| | | +-- Lampsilis_powellii
| | +--+
| | +-- Lampsilis_siliquoidea
| |
| | +-- Sinohyriopsis_cumingii **
| | +-----------+
| | | +-- Sinohyriopsis_schlegelii **
| | |
| | | +-- Pilsbryoconcha_exilis **
| +--------+ +--------+
| | | +-- Pseudodon_vondembuschianus **
| | |
| | | +-- Lamprotula_caveata **
| | | +--+
| +--+ | +-- Lamprotula_leaii **
| | +--+
--+ | | | +-- Pronodularia_japanensis **
| | | +--+
| | | +-- Potomida_littoralis **
| +--+
| | +-- Sinosolenaia_carinata **
| | +--+
| | | +-- Sinosolenaia_oleivora **
| +--+
| | +-- Parvasolenaia_rivularis **
| +--+
| +-- Ptychorhynchus_pfisteri **
|
| +----------------------- Lepidodesma_languilati
| |
| | +----- Lanceolaria_gladiola
| | +--------------+
| | | | +-- Lanceolaria_grayii
| | | +--+
| | | +-- Lanceolaria_lanceolata
+--+ |
| | +-------------- Acuticosta_chinensis
| | |
| | | +-------- Cristaria_plicata
| | | |
| | | +--+ +-- Sinanodonta_woodiana
| | +--+ | | +--+
+--+ | | | | | +-- Sinanodonta_lucida
| | | | +--+
| | | | | +-- Anemina_arcaeformis
| | | | +--+
| | +--+ +-- Anemina_euscaphys
| | |
| | | +-- Anodonta_anatina
| | | +-----+
| | | | +-- Anodonta_cygnea
| | | |
| | +--+ +-- Pyganodon_grandis
+--+ | +--+
| | | +-- Utterbackia_peninsularis
| +--+
| | +-- Alasmidonta_varicosa
| +--+
| +-- Prolasmidonta_heterodon
|
| +----- Aculamprotula_tortuosa
| +--------+
| | | +-- Aculamprotula_tientsinensis
| | +--+
| | +-- Aculamprotula_coreana
| |
+--+ +-------- Unio_tumidus
| +--+
| | | +----- Unio_crassus
| | +--+
| | | +-- Unio_pictorum
| | +--+
+--+ +-- Unio_delphinus
|
| +-- Schistodesmus_lampreyanus
| +-----+
| | +-- Cuneopsis_heudei
+--+
| +----- Pseudocuneopsis_capitata
+--+
| +-- Nodularia_douglasiae
+--+
+-- Tchangsinaia_piscicula
Zieritz et al., 2021 [Publ 4231], fig. 3-M, 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
10 |
6 |
11 |
10 |
2 |
26.0 |
+----- Quadrula_quadrula
+--------------+
| | +-- Venustaconcha_ellipsiformis
| +--+
| +-- Potamilus_alatus
|
| +----- Unio_tumidus
| +-----+
| | | +-- Unio_delphinus
--+ | +--+
| +-----+ +-- Unio_crassus
| | |
| | | +-------- Sinanodonta_woodiana
| | +--+
| | | +----- Anodonta_anatina
| | +--+
| | | +-- Utterbackia_peninsularis
| | +--+
+--+ +-- Pyganodon_grandis
|
| +-- Chamberlainia_somsakpanhai **
| +-----------+
| | +-- Sinohyriopsis_cumingii **
| |
| | +-- Lens_contradens **
| | +--+
+--+ | +-- Physunio_superbus **
| +-----+
| | | +-- Hyriopsis_bialata **
| | +--+
| | +-- Rectidens_sumatrensis **
+--+
| +-- Pilsbryoconcha_exilis **
| +-----+
| | +-- Pseudodon_vondembuschianus **
| |
+--+ +-- Pronodularia_japanensis **
| +--+
| | +-- Potomida_littoralis **
+--+
| +-- Microcondylaea_bonellii **
+--+
+-- Sinosolenaia_carinata **
Pavan-Kumar et al., 2022 [Publ 4263], fig. S5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 12 |
12 |
10 |
4 |
27 |
25 |
3 |
32.0 |
+-------------------- Lamellidens_marginalis
|
| +-- Anodonta_anatina
| +--------+
| | +-- Pyganodon_grandis
| +-----+
+--+ | | +-------- Cristaria_plicata
| | | | |
| | | +--+ +-- Anemina_arcaeformis
| | | | +--+
| | | | | +-- Anemina_euscaphys
| | | +--+
| | | | +-- Sinanodonta_lucida
| +--+ +--+
| | +-- Sinanodonta_woodiana
| |
| | +----- Acuticosta_chinensis
| | +--------+
| | | | +-- Lanceolaria_lanceolata
| | | +--+
| | | +-- Lanceolaria_gladiola
| +--+
| | +----- Lepidodesma_languilati
| | +-----+
| | | | +-- Aculamprotula_tientsinensis
| | | +--+
--+ +--+ +-- Aculamprotula_tortuosa
| |
| | +-------- Unio_pictorum
| +--+
| | +----- Schistodesmus_lampreyanus
| +--+
| | +-- Nodularia_douglasiae
| +--+
| +-- Tchangsinaia_piscicula
|
| +-------------- Quadrula_quadrula
| |
| +--+ +----------- Toxolasma_parvum
| | | |
| | +--+ +-------- Potamilus_leptodon
| | | |
| | +--+ +-- Lampsilis_cardium
| | | +--+
| | | | +-- Lampsilis_ornata
| | +--+
| | | +-- Lampsilis_siliquoidea
+-----+ +--+
| +-- Venustaconcha_ellipsiformis
|
| +----- Chamberlainia_somsakpanhai **
| +--------+
| | | +-- Sinohyriopsis_schlegelii **
| | +--+
| | +-- Sinohyriopsis_cumingii **
| |
| | +-- Sinosolenaia_carinata **
+--+ +--+
| | +-- Sinosolenaia_oleivora **
| +-----+
| | | +-- Parvasolenaia_rivularis **
| | +--+
| | +-- Microcondylaea_bonellii **
+--+
| +-- Pseudodon_vondembuschianus **
| +-----+
| | +-- Pilsbryoconcha_exilis **
+--+
| +----- Potomida_littoralis **
+--+
| +-- Lamprotula_leaii **
+--+
+-- Pronodularia_japanensis **
Pfeiffer et al., 2021 [Publ 4126], fig. 12
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
11 |
8 |
1 |
0 |
0 |
0 |
35.0 |
+-------------- Solenaia_emarginata **
|
| +-- Lens_contradens **
| +--------+
--+ | +-- Lens_micropterus **
| |
| | +-- Lens_inornatus **
| +--------+
+--+ +-- Lens_misellus **
|
| +----- Lens_maenamensis **
| +--+
| | | +-- Lens_rolfbrandti **
| | +--+
+--+ +-- Lens_eximius **
|
| +----- Lens_pallegoixi **
+--+
| +-- Lens_comptus **
+--+
+-- Lens_novoselovi **
Pfeiffer et al., 2021 [Publ 4126], fig. 27
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
11 |
7 |
2 |
0 |
0 |
0 |
31.0 |
+----------------- Khairuloconcha_lunbawangorum **
|
--+ +-------------- Rectidens_sumatrensis **
| |
| | +-- Hyriopsis_delaportei **
+--+ +--------+
| | +-- Hyriopsis_desowitzi **
| |
+--+ +-------- Hyriopsis_khoratensis **
| |
| | +-- Hyriopsis_kratiensis **
| +-----+
+--+ +-- Hyriopsis_phuphaniensis **
|
| +----- Hyriopsis_sakhonensis **
| |
+--+ +-- Hyriopsis_myersiana **
| |
+--+-- Hyriopsis_panhai **
|
+-- Hyriopsis_bialata **
Chen et al., 2020 [Publ 4170], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
34 |
32 |
4 |
23 |
22 |
2 |
58.0 |
+----- Prolasmidonta_heterodon
+--+
| | +-- Alasmidonta_varicosa
| +--+
| +-- Anemina_euscaphys
+-----------------+
| | +-- Lamprotula_caveata **
| | +--+
| | | +-- Chamberlainia_somsakpanhai **
| +--+
| | +-- Pseudodon_vondembuschianus **
| +--+
| +-- Pilsbryoconcha_exilis **
|
| +-------- Schistodesmus_lampreyanus
| +-----+
| | | +----- Lepidodesma_languilati
| | +--+
--+ | | +-- Aculamprotula_coreana
| | +--+
| +--------+ +-- Aculamprotula_tortuosa
| | |
| | | +----- Platynaias_compressa
| | | +-----+
| | | | | +-- Utterbackia_imbecillis
| | | | +--+
| | +--+ +-- Pyganodon_grandis
| | |
| | | +-------- Sinanodonta_lucida
| | +--+
| | | +----- Cristaria_plicata
| | +--+
+--+ | +-- Anodonta_cygnea
| +--+
| +-- Anodonta_anatina
|
| +----------------- Gibbosula_crassa
| +--+
| | | +-------------- Quadrula_quadrula
| | +--+
| | | +----------- Toxolasma_parvum
| | +--+
| | | +-------- Lampsilis_ornata
| | +--+
| | | +----- Potamilus_leptodon
+--+ +--+
| | +-- Lampsilis_siliquoidea
| +--+
| +-- Lampsilis_powellii
|
| +-- Utterbackia_peninsularis
| +--------------+
| | +-- Potamilus_alatus
+--+
| +-------------- Lamprotula_leaii **
| |
+--+ +-- Sinohyriopsis_cumingii **
| +--------+
| | +-- Sinohyriopsis_schlegelii **
+--+
| +-------- Parvasolenaia_rivularis **
+--+
| +----- Sinosolenaia_carinata **
+--+
| +-- Sinosolenaia_iridinea **
+--+
+-- Sinosolenaia_oleivora **
Pavan-Kumar et al., 2022 [Publ 4263], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
11 |
9 |
4 |
25 |
23 |
3 |
29.0 |
+-------------- Quadrula_quadrula
|
+--------------+ +----------- Toxolasma_parvum
| | |
| +--+ +-------- Potamilus_leptodon
| | |
| +--+ +-- Lampsilis_cardium
| | +--+
| | | +-- Lampsilis_ornata
| +--+
| | +-- Lampsilis_siliquoidea
| +--+
| +-- Venustaconcha_ellipsiformis
|
| +----- Chamberlainia_somsakpanhai **
--+ +-----------+
| | | +-- Sinohyriopsis_cumingii **
| | +--+
| | +-- Sinohyriopsis_schlegelii **
| +--------+
| | | +-- Lamprotula_leaii **
| | | +-----------+
| | | | +-- Potomida_littoralis **
| | | |
| | +--+ +-- Pseudodon_vondembuschianus **
| | | +--------+
| | | | +-- Pilsbryoconcha_exilis **
| | +--+
| | | +-------- Parvasolenaia_rivularis **
+--+ +--+
| | +----- Microcondylaea_bonellii **
| +--+
| | +-- Sinosolenaia_carinata **
| +--+
| +-- Sinosolenaia_oleivora **
|
| +----------------------- Lamellidens_marginalis
| |
| | +-------------------- Lepidodesma_languilati
+--+ |
| | +-- Aculamprotula_tientsinensis
| | +--------+
| | | +-- Aculamprotula_tortuosa
+--+ +-----+
| | | +-------- Unio_pictorum
| | +--+
| | | +----- Schistodesmus_lampreyanus
| | +--+
| | | +-- Nodularia_douglasiae
+--+ +--+
| +-- Tchangsinaia_piscicula
|
| +-- Lanceolaria_lanceolata
| +-----------+
| | +-- Lanceolaria_gladiola
+--+
| +----------- Acuticosta_chinensis
| |
+--+ +-- Anodonta_anatina
| +-----+
| | +-- Pyganodon_grandis
+--+
| +----- Cristaria_plicata
+--+
| +-- Sinanodonta_lucida
+--+
+-- Anemina_euscaphys
Dai et al., 2024 [Publ 4358], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 11 |
11 |
9 |
4 |
41 |
39 |
2 |
29.0 |
+-- Quadrula_quadrula
+--------------------+
| +-- Uniomerus_tetralasmus
|
+--------------+ +-- Elliptio_complanata
| | +-----------------+
| | | +-- Pleurobema_oviforme
| +--+
| | +----------------- Amblema_plicata
| | |
| +--+ +-------------- Popenaias_popeii
| | |
| +--+ +----------- Toxolasma_parvum
| | |
| +--+ +-- Potamilus_leptodon
| | +-----+
| | | +-- Potamilus_alatus
| +--+
| | +----- Leaunio_lienosus
| +--+
| | +-- Lampsilis_cardium
| +--+
--+ +-- Venustaconcha_ellipsiformis
|
| +-- Chamberlainia_somsakpanhai **
| +--------------+
| | +-- Sinohyriopsis_cumingii **
| |
| | +----- Potomida_littoralis **
| +-----------------+ +--------+
| | | | | +-- Lamprotula_leaii **
| | | | +--+
| | | | +-- Pronodularia_japanensis **
| | +--+
| | | +-- Pseudodon_vondembuschianus **
| | | +--------+
| | | | +-- Pilsbryoconcha_exilis **
| | +--+
| | | +-------- Microcondylaea_bonellii **
| | +--+
| | | +----- Sinosolenaia_oleivora **
+--+ +--+
| | +-- Ptychorhynchus_pfisteri **
| +--+
| +-- Parvasolenaia_rivularis **
|
| +-------------------------------- Lepidodesma_languilati
| |
| | +----------------- Lanceolaria_gladiola
| | |
| | +-----------+ +-------------- Acuticosta_chinensis
| | | | |
| | | | | +----- Cristaria_plicata
+--+ | +--+ +-----+
| | | | | +-- Anemina_arcaeformis
| | | | +--+
| | +--+ +-- Sinanodonta_woodiana
| | |
| | | +-------- Anodonta_anatina
| | +--+
+--+ | +----- Platynaias_compressa
| +--+
| | +-- Pyganodon_grandis
| +--+
| +-- Utterbackia_imbecillis
|
| +-- Aculamprotula_scripta
| +-----------------------+
| | +-- Aculamprotula_tientsinensis
| |
| | +-- Unio_crassus
+--+ +--------------------+
| | +-- Unio_delphinus
| |
| | +-------- Pseudocuneopsis_capitata
+--+ +-----------+
| | | +----- Pseudocuneopsis_yangshuoensis
| | +--+
| | | +-- Pseudocuneopsis_yemaoi
| | +--+
+--+ +-- Pseudocuneopsis_sichuanensis
|
| +-- Nodularia_breviconcha
| +--------------+
| | +-- Nodularia_douglasiae
| |
+--+ +-- Schistodesmus_lampreyanus
| +-----------+
| | +-- Schistodesmus_spinosus
+--+
| +----------- Tchangsinaia_piscicula
+--+
| +-------- Cuneopsis_rufescens
+--+
| +----- Cuneopsis_celtiformis
+--+
| +-- Cuneopsis_heudei
+--+
+-- Cuneopsis_demangei
Konopleva et al., 2017 [Publ 3807], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
3 |
21 |
11 |
1 |
32.0 |
+-------- Ensidens_sagittarius **
+-----+
| | +----- Lens_micropterus **
| +--+
| | +-- Physunio **
+--+ +--+
| | +-- Physunio_modelli **
| |
| | +----------- Sundadontina_tumida **
| | |
| +--+ +-- Nyeinchanconcha_nyeinchani **
| | +-----+
| | | +-- Bineurus_exilis **
| +--+
| | +----- Pilsbryoconcha_carinifera **
| +--+
--+ | +-- Pseudodon_vagulus **
| +--+
| +-- Pseudodon_vondembuschianus **
|
| +-- Trapezidens_exolescens
| +-----+
| | +-- Trapezidens_dolichorhynchus
| +-----+
| | | +----- Lamellidens_savadiensis
| | +--+
| | | +-- Lamellidens_jenkinsianus
| | +--+
+--+ +-- Lamellidens_mainwaringi
|
| +-- Leoparreysia_tavoyensis
| +--------+
| | +-- Leoparreysia_olivacea
+--+
| +-------- Coelatura_aegyptiaca
+--+
| +----- Indochinella_pugio
+--+
| +-- Radiatula_chaudhurii
+--+
+-- Scabies_mandarinus
Huang et al., 2019 [Publ 3843], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
3 |
47 |
37 |
2 |
34.0 |
+-- Sinohyriopsis_cumingii **
+-----------+
| +-- Sinohyriopsis_schlegelii **
|
| +-------- Potomida_littoralis **
+--------+ +--+
| | | | +----- Pronodularia_japanensis **
| | | +--+
| | | | +-- Lamprotula_caveata **
| | | +--+
| +--+ +-- Lamprotula_leaii **
| |
| | +-- Ptychorhynchus_pfisteri **
| | +--+
+--------+ | | +-- Parvasolenaia_rivularis **
| | +-----+
| | | +-- Sinosolenaia_oleivora **
| | +--+
| | +-- Sinosolenaia_carinata **
| |
| | +-- Quadrula_quadrula
| | +-----------------+
| | | +-- Uniomerus_tetralasmus
| +--+
| | +----------------- Elliptio_complanata
| | |
| +--+ +-------------- Toxolasma_parvum
| | |
| +--+ +-- Potamilus_leptodon
| | +--------+
| | | +-- Potamilus_alatus
--+ +--+
| | +-------- Leaunio_lienosus
| +--+
| | +----- Venustaconcha_ellipsiformis
| +--+
| | +-- Lampsilis_cardium
| +--+
| +-- Lampsilis_ornata
|
| +----------------------------- Lepidodesma_languilati
| |
| | +-------- Aculamprotula_tortuosa
| | +-----+
| | | | +----- Aculamprotula_scripta
| | | +--+
| | | | +-- Aculamprotula_tientsinensis
| | | +--+
| | +-----------+ +-- Aculamprotula_coreana
+--+ | |
| | | +----- Nodularia_douglasiae
| | | +-----+
| | | | | +-- Tchangsinaia_piscicula
| | | | +--+
| | +--+ +-- Schistodesmus_lampreyanus
| | |
| | | +-------- Unio_tumidus
| | +--+
+--+ | +----- Unio_crassus
| +--+
| | +-- Unio_delphinus
| +--+
| +-- Unio_pictorum
|
| +----- Lanceolaria_gladiola
| +-----------------+
| | | +-- Lanceolaria_lanceolata
| | +--+
| | +-- Lanceolaria_grayii
+--+
| +-------------------- Acuticosta_chinensis
| |
| | +-------- Cristaria_plicata
| | |
+--+ +--------+ +-- Sinanodonta_lucida
| | | +--+
| | | | +-- Sinanodonta_woodiana
| | +--+
+--+ | +-- Anemina_arcaeformis
| +--+
| +-- Anemina_euscaphys
|
| +-------------- Anodonta_cygnea
+--+
| +----------- Anodonta_anatina
+--+
| +-------- Lasmigona_costata
+--+
| +----- Pyganodon_grandis
+--+
| +-- Utterbackia_imbecillis
+--+
+-- Utterbackia_peninsularis
Muanta et al., 2019 [Publ 3921], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
8 |
1 |
0 |
0 |
0 |
34.0 |
+-------- Hyriopsis_desowitzi **
+-----+
| | +----- Hyriopsis_myersiana **
| +--+
| | +-- Hyriopsis_khoratensis **
| +--+
--+ +-- Hyriopsis_panhai **
|
| +-- Ensidens_ingallsianus **
| +--------+
| | +-- Ensidens_telus **
+--+
| +-------- Ensidens_sagittarius[2] **
+--+
| +----- Ensidens_jaculus **
+--+
| +-- Ensidens_sagittarius[1] **
+--+
+-- Ensidens_dugasti **
Pfeiffer et al., 2021 [Publ 4126], fig. 18
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
7 |
1 |
0 |
0 |
0 |
32.0 |
+-------------- Yaukthwa_paiensis **
|
| +----- Trapezoideus_foliaceus **
--+ +-----+
| | | +-- Trapezoideus_lenya **
| | +--+
| | +-- Trapezoideus_subviridis **
+--+
| +-- Physunio_massini **
| +-----+
| | +-- Physunio_superbus **
+--+
| +----- Physunio_modelli[3] **
| |
+--+ +-- Physunio_modelli[1] **
| |
+--+-- Physunio_modelli[2] **
|
+-- Physunio_pellucidus **
Sano et al., 2020 [Publ 4138], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 10 |
10 |
6 |
3 |
38 |
34 |
1 |
37.0 |
+-------- Sinosolenaia_iridinea **
|
+--+ +----- Inversidens_brandtii **
| | |
| +--+ +-- Obovalis_omiensis[1] **
| | |
| +--+-- Obovalis_omiensis[2] **
+--+ |
| | +-- Ptychorhynchus_pfisteri **
| |
| | +-- Lamprotula_caveata **
| | +--+
| | | +-- Pronodularia_japanensis **
| +-----+
| | +-- Sinohyriopsis_schlegelii[1] **
| | |
| +--+-- Sinohyriopsis_schlegelii[2] **
| |
| +-- Sinohyriopsis_cumingii **
|
| +-------- Lanceolaria_grayii[1]
| |
| +-------- Lanceolaria_grayii[2]
| |
| +-------- Cristaria_plicata[1]
| |
| +-------- Cristaria_plicata[2]
--+ |
| +-------- Sinanodonta_lauta
| |
| +-------- Beringiana_japonica
| |
| +-------- Beringiana_fukuharai[1]
| |
| +-------- Beringiana_beringiana
| |
| +-------- Beringiana_fukuharai[2]
| |
| +-------- Beringiana_fukuharai[3]
| |
| +-------- Buldowskia_iwakawai[1]
| |
| +-------- Buldowskia_iwakawai[2]
| |
| +-------- Sinanodonta_tumens[1]
| |
| +-------- Sinanodonta_tumens[2]
| |
+-----+-------- Pletholophus_tenuis
|
+-------- Anemina_arcaeformis
|
+-------- Lanceolaria_lanceolata
|
+-------- Cristaria_plicata[3]
|
+-------- Lanceolaria_gladiola
|
+-------- Sinanodonta_angula
|
+-------- Lepidodesma_languilati
|
| +-- Aculamprotula_tortuosa
+-----+
| +-- Acuticosta_chinensis
|
| +----- Cuneopsis_celtiformis
| |
+--+ +-- Nodularia_douglasiae[1]
| | |
| +--+-- Nodularia_douglasiae[2]
| |
| +-- Nodularia_nipponensis
|
| +----- Schistodesmus_lampreyanus
| |
| | +-- Inversiunio_reinianus[1]
+--+ |
| +-- Inversiunio_reinianus[2]
| |
| +-- Inversiunio_jokohamensis[1]
+--+
+-- Inversiunio_jokohamensis[2]
|
+-- Inversiunio_jokohamensis[3]
|
+-- Inversiunio_jokohamensis[4]
Wu et al., 2018 [Publ 3836], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
42 |
36 |
3 |
44 |
42 |
2 |
65.0 |
+-------------------------------- Quadrula_quadrula
|
| +-- Amblema_plicata
+-----------------------------+
| +-- Obovaria_arkansasensis
|
| +----------- Pleurobema_sintoxia
| |
+--------------------+ +-- Elliptio_complanata
| | +-----+
| | | +-- Eurynia_dilatata
| +--+
| | +----- Fusconaia_askewi
| +--+
| | +-- Fusconaia_cerina
| +--+
--+ +-- Fusconaia_burkei
|
| +----- Potomida_littoralis **
| +-----+
| | | +-- Lamprotula_caveata **
| | +--+
| +-----------------+ +-- Lamprotula_leaii **
| | |
| | | +-------- Sinosolenaia_carinata **
| | | |
| | +--+ +----- Gonidea_angulata **
| | | |
| | +--+----- Sinosolenaia_oleivora **
| | |
| | | +-- Parvasolenaia_rivularis **
| | +--+
| | +-- Ptychorhynchus_pfisteri **
+--+
| +-------------------------- Sinohyriopsis_cumingii **
| |
| | +----------------------- Lepidodesma_languilati
| | |
| | | +----------- Aculamprotula_tortuosa
| | | +-----+
| | | | | +-------- Aculamprotula_scripta
| | | | +--+
| | | | | +----- Aculamprotula_coreana
| | | | +--+
+--+ | | | +-- Aculamprotula_fibrosa
| +-----+ +--+
| | | +-- Aculamprotula_tientsinensis
| | |
| | | +-- Unio_crassus
| | | +-----------+
| | | | +-- Unio_pictorum
| | | |
| | +--+ +----- Nodularia_douglasiae
| | | +-----+
| | | | | +-- Schistodesmus_lampreyanus
+--+ | | +--+
| +--+ +-- Schistodesmus_spinosus
| |
| | +-------- Tchangsinaia_piscicula
| +--+
| | +----- Cuneopsis_rufescens
| +--+
| | +-- Cuneopsis_celtiformis
| +--+
| +-- Cuneopsis_heudei
|
| +----------- Acuticosta_chinensis
| +--------+
| | | +-------- Lanceolaria_gladiola
| | +--+
| | | +----- Lanceolaria_grayii
| | +--+
| | | +-- Lanceolaria_lanceolata
| | +--+
| | +-- Lanceolaria_triformis
+--+
| +-------- Anemina_arcaeformis
| |
| +-------- Cristaria_plicata
| +--------+
| | | +-- Sinanodonta_woodiana
| | | +--+
| | | | +-- Sinanodonta_elliptica
| | +--+
+--+ | +-- Sinanodonta_lucida
| +--+
| +-- Sinanodonta_angula
|
| +-------------- Anodonta_cygnea
| |
+--+ +----------- Anodonta_anatina
| |
+--+ +-------- Strophitus_undulatus
| |
+--+ +----- Platynaias_compressa
| |
+--+----- Alasmidonta_marginata
|
| +-- Pyganodon_cataracta
+--+
+-- Pyganodon_grandis
Cha et al., 2018 [Publ 3916], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
7 |
3 |
30 |
27 |
2 |
33.0 |
+----------- Quadrula_quadrula
+-----------------+
| | +-------- Toxolasma_parvum
| +--+
| | +----- Lampsilis_ornata
| +--+
| | +-- Potamilus_alatus
| +--+
| +-- Potamilus_leptodon
|
| +-- Sinohyriopsis_cumingii **
| +-----------+
| | +-- Sinohyriopsis_schlegelii **
--+ |
| +-----------+ +----- Ptychorhynchus_pfisteri **
| | | +-----+
| | | | | +-- Sinosolenaia_carinata **
| | | | +--+
| | +--+ +-- Sinosolenaia_oleivora **
| | |
| | | +-------- Potomida_littoralis **
| | +--+
| | | +----- Pronodularia_japanensis **
| | +--+
| | | +-- Lamprotula_caveata **
+--+ +--+
| +-- Lamprotula_leaii **
|
| +----------------------- Lepidodesma_languilati
| |
| | +----- Aculamprotula_tortuosa
| | +--------+
| | | | +-- Aculamprotula_coreana
| | | +--+
| | | +-- Aculamprotula_tientsinensis
| | +-----+
+--+ | | +----- Schistodesmus_lampreyanus
| | | +-----+
| | | | | +-- Nodularia_douglasiae
| | | | +--+
| | +--+ +-- Tchangsinaia_piscicula
| | |
| | | +-------- Unio_tumidus
| | +--+
+--+ | +----- Unio_crassus
| +--+
| | +-- Unio_pictorum
| +--+
| +-- Unio_delphinus
|
| +----------------- Lanceolaria_grayii
| |
| | +----------- Anodonta_anatina
| | +--+
+--+ | | +-------- Platynaias_compressa
| | +--+
| | | +----- Pyganodon_grandis
| | +--+
+--+ | +-- Utterbackia_peninsularis
| +--+
| +-- Utterbackia_imbecillis
|
| +-------- Cristaria_plicata
| |
+-----+ +-- Anemina_arcaeformis
| +--+
| | +-- Anemina_euscaphys
+--+
| +-- Sinanodonta_lucida
+--+
+-- Sinanodonta_woodiana
Pfeiffer et al., 2021 [Publ 4126], fig. 25
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
7 |
1 |
0 |
0 |
0 |
31.0 |
+-------------- Solenaia_emarginata **
|
| +-- Yaukthwa_elongatula **
--+ +--------+
| | +-- Yaukthwa_baniensis **
| |
+--+ +----- Yaukthwa_panhai **
| +--+
| | | +-- Yaukthwa_rectangularis **
| | +--+
+--+ +-- Yaukthwa_nesemanni **
|
| +----- Yaukthwa_avaensis **
+--+
| +-- Yaukthwa_inlenensis **
+--+
+-- Yaukthwa_paiensis **
Sano et al., 2020 [Publ 4138], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
7 |
3 |
64 |
41 |
3 |
32.0 |
+----------------- Lampsilis_cardium
|
| +----- Sinohyriopsis_cumingii[2] **
+--------------------+ +--------+
| | | | +-- Sinohyriopsis_schlegelii **
| | | +--+
| +--+ +-- Sinohyriopsis_cumingii[1] **
| |
| | +----------- Inversidens_brandtii **
| | |
| +--+ +-- Ptychorhynchus_pfisteri **
| | +-----+
| | | +-- Obovalis_omiensis **
| +--+
| | +----- Lamprotula_leaii **
| +--+
--+ | +-- Pronodularia_japanensis **
| +--+
| +-- Potomida_littoralis **
|
| +----- Buldowskia_iwakawai
| +-----------------------------+
| | | +-- Anemina_arcaeformis
| | +--+
| | +-- Anemina_euscaphys
| |
| | +-------------- Pletholophus_tenuis
| | |
| | | +----- Beringiana_fukuharai
| | +-----------------+ +-----+
+--+ | | | | +-- Beringiana_japonica
| | | | +--+
| | +--+ +-- Beringiana_beringiana
| | |
| | | +-------- Sinanodonta_lucida
| | +--+
| | | +----- Sinanodonta_tumens
| | +--+
| | | +-- Sinanodonta_lauta
+--+ +--+
| +-- Sinanodonta_woodiana
|
| +----------------------------- Cristaria_plicata
| |
| | +-- Anodonta_cygnea
| | +--------+
| | | +-- Anodonta_anatina
| | |
| | +--------------+ +----- Alasmidonta_varicosa
+--+ | | +--+
| | | | | +-- Prolasmidonta_heterodon
| | | | +--+
| | +--+ +-- Platynaias_compressa
| | |
| | | +----- Pyganodon_grandis
| | +--+
+--+ | +-- Utterbackia_imbecillis
| +--+
| +-- Utterbackia_peninsularis
|
| +----- Lanceolaria_grayii[1]
| +-----------------+
| | | +-- Lanceolaria_lanceolata
| | +--+
| | +-- Lanceolaria_grayii[2]
+--+
| +-------------------- Lepidodesma_languilati
| |
| | +----- Aculamprotula_tortuosa
| | +-----------+
+--+ | | +-- Aculamprotula_tientsinensis
| | +--+
| | +-- Aculamprotula_coreana
| |
+--+ +----- Unio_crassus
| +--------+
| | | +-- Unio_delphinus
| | +--+
| | +-- Unio_pictorum
| |
+--+ +-------- Inversiunio_jokohamensis
| +--+
| | | +----- Inversiunio_yanagawensis[2]
| | +--+
| | | +-- Inversiunio_reinianus
| | +--+
+--+ +-- Inversiunio_yanagawensis[1]
|
| +-- Tchangsinaia_piscicula
| +-----+
| | +-- Schistodesmus_lampreyanus
+--+
| +----- Nodularia_douglasiae[2]
+--+
| +-- Nodularia_nipponensis
+--+
+-- Nodularia_douglasiae[1]
Sano et al., 2020 [Publ 4138], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
10 |
8 |
3 |
63 |
40 |
3 |
33.0 |
+----------------------------------------- Scabies_crispata
|
| +----- Sinohyriopsis_cumingii[2] **
| +--------+
| | | +-- Sinohyriopsis_schlegelii **
| | +--+
| | +-- Sinohyriopsis_cumingii[1] **
| +-----------------------+
| | | +----------- Lampsilis_cardium
--+ | | |
| | | | +----- Lamprotula_leaii **
| | +--+ +--+
| | | | | +-- Potomida_littoralis **
| | | | +--+
| | +--+ +-- Pronodularia_japanensis **
| | |
| | | +----- Inversidens_brandtii **
| | +--+
| | | +-- Obovalis_omiensis **
+--+ +--+
| +-- Ptychorhynchus_pfisteri **
|
| +----- Buldowskia_iwakawai
| +-----------------------------+
| | | +-- Anemina_arcaeformis
| | +--+
| | +-- Anemina_euscaphys
| |
| | +-------- Pletholophus_tenuis
| | +--+
| | | | +----- Beringiana_fukuharai
| | | +--+
+--+ | | +-- Beringiana_japonica
| +--------------------+ +--+
| | | +-- Beringiana_beringiana
| | |
| | | +-------- Sinanodonta_lucida
| | +--+
| | | +----- Sinanodonta_tumens
| | +--+
| | | +-- Sinanodonta_woodiana
| | +--+
+--+ +-- Sinanodonta_lauta
|
| +----------------------------- Cristaria_plicata
| |
| | +-- Anodonta_anatina
| | +--------+
| | | +-- Anodonta_cygnea
| | |
| | +--------------+ +----- Pyganodon_grandis
+--+ | | +--+
| | | | | +-- Utterbackia_imbecillis
| | | | +--+
| | +--+ +-- Utterbackia_peninsularis
| | |
| | | +----- Alasmidonta_varicosa
| | +--+
+--+ | +-- Platynaias_compressa
| +--+
| +-- Prolasmidonta_heterodon
|
| +----- Lanceolaria_grayii[1]
| +-----------------+
| | | +-- Lanceolaria_lanceolata
| | +--+
| | +-- Lanceolaria_grayii[2]
+--+
| +-------------------- Lepidodesma_languilati
| |
| | +----- Aculamprotula_tortuosa
| | +-----------+
+--+ | | +-- Aculamprotula_tientsinensis
| | +--+
| | +-- Aculamprotula_coreana
| |
+--+ +----- Unio_crassus
| +--------+
| | | +-- Unio_delphinus
| | +--+
| | +-- Unio_pictorum
| |
+--+ +-------- Inversiunio_jokohamensis
| +--+
| | | +----- Inversiunio_yanagawensis[2]
| | +--+
| | | +-- Inversiunio_reinianus
| | +--+
+--+ +-- Inversiunio_yanagawensis[1]
|
| +-- Nodularia_nipponensis
| +--+
| | +-- Nodularia_douglasiae
+-----+
| +-- Schistodesmus_lampreyanus
+--+
+-- Tchangsinaia_piscicula
Sano et al., 2020 [Publ 4138], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
7 |
3 |
63 |
40 |
3 |
32.0 |
+----------------- Lampsilis_cardium
|
| +----- Sinohyriopsis_cumingii[2] **
+-----------------------+ +--------+
| | | | +-- Sinohyriopsis_schlegelii **
| | | +--+
| +--+ +-- Sinohyriopsis_cumingii[1] **
| |
| | +----------- Inversidens_brandtii **
| | |
| +--+ +-- Obovalis_omiensis **
| | +-----+
| | | +-- Ptychorhynchus_pfisteri **
| +--+
| | +----- Lamprotula_leaii **
| +--+
--+ | +-- Pronodularia_japanensis **
| +--+
| +-- Potomida_littoralis **
|
| +----- Buldowskia_iwakawai
| +--------------------------------+
| | | +-- Anemina_arcaeformis
| | +--+
| | +-- Anemina_euscaphys
| |
| | +-------- Sinanodonta_lucida
| | +--+
| | | | +----- Sinanodonta_tumens
| | | +--+
+--+ | | +-- Sinanodonta_woodiana
| +-----------------------+ +--+
| | | +-- Sinanodonta_lauta
| | |
| | | +-------- Pletholophus_tenuis
| | +--+
| | | +----- Beringiana_fukuharai
| | +--+
| | | +-- Beringiana_japonica
| | +--+
+--+ +-- Beringiana_beringiana
|
| +-------------------------------- Cristaria_plicata
| |
| | +-- Anodonta_cygnea
| | +--------+
| | | +-- Anodonta_anatina
| | |
| | +-----------------+ +----- Alasmidonta_varicosa
+--+ | | +--+
| | | | | +-- Prolasmidonta_heterodon
| | | | +--+
| | +--+ +-- Platynaias_compressa
| | |
| | | +----- Pyganodon_grandis
| | +--+
+--+ | +-- Utterbackia_imbecillis
| +--+
| +-- Utterbackia_peninsularis
|
| +-------------------------- Lepidodesma_languilati
| |
| | +----- Lanceolaria_grayii[1]
| | +-----------------+
+--+ | | +-- Lanceolaria_lanceolata
| | +--+
| | +-- Lanceolaria_grayii[2]
| |
+--+ +----- Aculamprotula_tortuosa
| +--------------+
| | | +-- Aculamprotula_tientsinensis
| | +--+
| | +-- Aculamprotula_coreana
| |
+--+ +----- Unio_crassus
| +-----------+
| | | +-- Unio_delphinus
| | +--+
| | +-- Unio_pictorum
+--+
| +-- Nodularia_douglasiae
| +-----------+
| | +-- Nodularia_nipponensis
| |
+--+ +-- Schistodesmus_lampreyanus
| +--------+
| | +-- Tchangsinaia_piscicula
+--+
| +-------- Inversiunio_jokohamensis
+--+
| +----- Inversiunio_yanagawensis[2]
+--+
| +-- Inversiunio_reinianus
+--+
+-- Inversiunio_yanagawensis[1]
Wu et al., 2020 [Publ 4147], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
7 |
3 |
21 |
20 |
2 |
33.0 |
+----- Quadrula_quadrula
+--------------------+
| | +-- Toxolasma_parvum
| +--+
| +-- Venustaconcha_ellipsiformis
|
| +-- Sinohyriopsis_schlegelii **
| +--------------+
| | +-- Sinohyriopsis_cumingii **
| |
| +-----+ +-- Sinosolenaia_oleivora **
--+ | | +-----------+
| | | | +-- Sinosolenaia_carinata **
| | +--+
| | | +----------- Pronodularia_japanensis **
| | +--+
| | | +-------- Lamprotula_leaii **
| | +--+
| | | +----- Lamprotula_caveata[1] **
| | +--+
| | | +-- Lamprotula_cornuumlunae **
| | +--+
+--+ +-- Lamprotula_caveata[2] **
|
| +----- Cristaria_plicata
| +-----+
| | | +-- Sinanodonta_lucida
| | +--+
| +--------+ +-- Anemina_arcaeformis
| | |
| | | +-------- Anodonta_anatina
| | +--+
| | | +----- Pyganodon_grandis
| | +--+
| | | +-- Utterbackia_peninsularis
+--+ +--+
| +-- Utterbackia_imbecillis
|
| +-------- Aculamprotula_tortuosa
| +--------+
| | | +----- Aculamprotula_scripta
| | +--+
| | | +-- Aculamprotula_tientsinensis
+--+ +--+
| +-- Aculamprotula_coreana
|
| +-------------- Unio_pictorum
+--+
| +----------- Pseudocuneopsis_capitata
+--+
| +-------- Nodularia_douglasiae
+--+
| +----- Schistodesmus_lampreyanus
+--+
| +-- Cuneopsis_heudei
+--+
+-- Tchangsinaia_piscicula
Lopes-Lima et al., 2021 [Publ 4217], fig. 14
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 9 |
9 |
4 |
1 |
0 |
0 |
0 |
29.0 |
+-- Gonidea_angulata **
+--------+
| +-- Microcondylaea_bonellii **
|
--+ +-------- Leguminaia_anatolica **
| |
| | +----- Leguminaia_wheatleyi[2] **
+--+ |
| +----- Leguminaia_wheatleyi[3] **
| |
+--+----- Leguminaia_wheatleyi[4] **
|
+----- Leguminaia_wheatleyi[5] **
|
| +-- Leguminaia_saulcyi **
+--+
+-- Leguminaia_wheatleyi[1] **
Sano et al., 2017 [Publ 3799], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
6 |
3 |
28 |
24 |
1 |
35.0 |
+-- Obovalis_omiensis **
+--------+
| +-- Ptychorhynchus_pfisteri **
|
+-----------+ +-- Inversidens_brandtii **
| | +-----+
| | | +-- Sinosolenaia_oleivora **
| | |
| +--+ +-- Pronodularia_japanensis **
| | +--+
| | | +-- Lamprotula_leaii **
| +--+
| | +-- Sinohyriopsis_schlegelii **
| +--+
| +-- Sinohyriopsis_cumingii **
|
| +-- Cristaria_plicata
--+ +-----------+
| | +-- Buldowskia_iwakawai
| |
| +-----+ +----- Beringiana_fukuharai
| | | +-----+
| | | | | +-- Beringiana_japonica
| | | | +--+
| | +--+ +-- Beringiana_beringiana
| | |
| | | +-------- Pletholophus_tenuis
| | +--+
| | | +----- Sinanodonta_woodiana
| | +--+
+--+ | +-- Sinanodonta_lauta
| +--+
| +-- Sinanodonta_tumens
|
| +-------- Tchangsinaia_piscicula
| +--------+
| | | +----- Nodularia_nipponensis
| | +--+
| | | +-- Nodularia_douglasiae
| | +--+
| | +-- Nodularia_breviconcha
+--+
| +-- Schistodesmus_lampreyanus
| +-----------+
| | +-- Acuticosta_ovata
| |
| | +-------- Lepidodesma_languilati
+--+ +--+
| | | +----- Lanceolaria_grayii[1]
| | +--+
| | | +-- Lanceolaria_lanceolata
+--+ +--+
| +-- Lanceolaria_grayii[2]
|
| +-------- Inversiunio_jokohamensis
| |
+--+ +----- Inversiunio_yanagawensis[3]
| |
+--+ +-- Inversiunio_yanagawensis[1]
| |
+--+-- Inversiunio_yanagawensis[2]
|
+-- Inversiunio_reinianus
Pfeiffer et al., 2018 [Publ 3827], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
3 |
2 |
0 |
0 |
0 |
28.5 |
+----- Physunio **
|
+--+----- Trapezoideus **
| |
| | +-- Lens_comptus **
| +--+
--+ +-- Lens_contradens **
|
| +-- Ensidens **
| |
| +-- Hyriopsis[1] **
+-----+
+-- Hyriopsis[2] **
|
+-- Rectidens **
Huang et al., 2018 [Publ 3828], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
6 |
3 |
35 |
25 |
2 |
32.0 |
+----------- Quadrula_quadrula
|
+-----+ +-------- Toxolasma_parvum
| | |
| +--+ +-- Lampsilis_ornata
| | +--+
| | | +-- Venustaconcha_ellipsiformis
| +--+
| | +-- Potamilus_leptodon
+--+ +--+
| | +-- Potamilus_alatus
| |
| | +-- Sinohyriopsis_cumingii **
| | +-----------+
| | | +-- Sinohyriopsis_schlegelii **
| | |
| +--+ +-- Sinosolenaia_oleivora **
| | +--------+
| | | +-- Sinosolenaia_carinata **
| +--+
| | +-------- Potomida_littoralis **
| +--+
| | +----- Pronodularia_japanensis **
| +--+
| | +-- Lamprotula_caveata **
--+ +--+
| +-- Lamprotula_leaii **
|
| +----- Aculamprotula_tortuosa
| +--------+
| | | +-- Aculamprotula_coreana
| | +--+
| | +-- Aculamprotula_tientsinensis
| +--+
| | | +-- Tchangsinaia_piscicula
| | | +--------+
| | | | +-- Nodularia_douglasiae
| | +--+
| | | +-------- Unio_tumidus
| | +--+
| | | +----- Unio_crassus
| | +--+
+--+ | +-- Unio_delphinus
| +--+
| +-- Unio_pictorum
|
| +----------- Anodonta_anatina
| +--+
| | | +-------- Platynaias_compressa
| | +--+
| | | +----- Pyganodon_grandis
| | +--+
+--+ | +-- Utterbackia_imbecillis
| +--+
| +-- Utterbackia_peninsularis
|
| +-------- Cristaria_plicata
| |
+-----+ +-- Anemina_arcaeformis
| +--+
| | +-- Anemina_euscaphys
+--+
| +-- Sinanodonta_woodiana
+--+
+-- Sinanodonta_lucida
Wang et al., 2018 [Publ 3846], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
6 |
3 |
24 |
24 |
2 |
32.0 |
+----------- Quadrula_quadrula
|
+--+ +-------- Toxolasma_parvum
| | |
| +--+ +-- Venustaconcha_ellipsiformis
| | +--+
| | | +-- Lampsilis_ornata
| +--+
| | +-- Potamilus_leptodon
+--------+ +--+
| | +-- Potamilus_alatus
| |
| | +-- Sinohyriopsis_cumingii **
| | +--------+
| | | +-- Sinohyriopsis_schlegelii **
| | |
| +--+ +----- Potomida_littoralis **
| | +--+
| | | | +-- Lamprotula_caveata **
| | | +--+
| +--+ +-- Lamprotula_leaii **
| |
| | +----- Ptychorhynchus_pfisteri **
| +--+
--+ | +-- Sinosolenaia_carinata **
| +--+
| +-- Sinosolenaia_oleivora **
|
| +----- Anodonta_anatina
| +-----+
| | | +-- Pyganodon_grandis
| | +--+
| +--------+ +-- Platynaias_compressa
| | |
| | | +-------- Cristaria_plicata
| | | |
| | +--+ +-- Anemina_euscaphys
| | | +--+
| | | | +-- Anemina_arcaeformis
| | +--+
+--+ | +-- Sinanodonta_woodiana
| +--+
| +-- Sinanodonta_lucida
|
| +-- Lanceolaria_grayii
| +--------------+
| | +-- Lanceolaria_lanceolata
| |
+--+ +-------------- Aculamprotula_tientsinensis
| |
| | +----- Nodularia_douglasiae
+--+ +-----+
| | | +-- Tchangsinaia_piscicula
| | +--+
+--+ +-- Cuneopsis_heudei
|
| +-------- Unio_tumidus
+--+
| +----- Unio_crassus
+--+
| +-- Unio_pictorum
+--+
+-- Unio_delphinus
Wu et al., 2019 [Publ 3864], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
6 |
3 |
39 |
33 |
2 |
32.0 |
+----------- Quadrula_quadrula
|
+-----------------+ +-------- Toxolasma_parvum
| | |
| +--+ +-- Lampsilis_ornata
| | +--+
| | | +-- Venustaconcha_ellipsiformis
| +--+
| | +-- Potamilus_leptodon
| +--+
| +-- Potamilus_alatus
|
| +-- Sinohyriopsis_cumingii **
| +--------+
--+ | +-- Sinohyriopsis_schlegelii **
| |
| +--------------+ +----- Potomida_littoralis **
| | | +--+
| | | | | +-- Lamprotula_leaii **
| | | | +--+
| | +--+ +-- Pronodularia_japanensis **
| | |
| | | +----- Ptychorhynchus_pfisteri **
| | +--+
| | | +-- Sinosolenaia_oleivora **
| | +--+
| | +-- Sinosolenaia_carinata **
+--+
| +----------------------- Lepidodesma_languilati
| |
| | +-------- Aculamprotula_tortuosa
| | +--------+
| | | | +----- Aculamprotula_scripta
| | | +--+
| | | | +-- Aculamprotula_tientsinensis
| | | +--+
| | | +-- Aculamprotula_coreana
| | +--+
+--+ | | +-------- Unio_tumidus
| | | +-----+
| | | | | +----- Unio_crassus
| | | | +--+
| | | | | +-- Unio_delphinus
| | +--+ +--+
| | | +-- Unio_pictorum
| | |
| | | +----------- Pseudocuneopsis_capitata
| | +--+
| | | +-------- Nodularia_douglasiae
+--+ +--+
| | +----- Tchangsinaia_piscicula
| +--+
| | +-- Cuneopsis_heudei
| +--+
| +-- Schistodesmus_lampreyanus
|
| +----- Acuticosta_chinensis
| +-----------+
| | | +-- Lanceolaria_grayii
| | +--+
| | +-- Lanceolaria_lanceolata
| |
+--+ +----------- Anodonta_anatina
| +--+
| | | +-------- Platynaias_compressa
| | +--+
| | | +----- Pyganodon_grandis
| | +--+
+--+ | +-- Utterbackia_peninsularis
| +--+
| +-- Utterbackia_imbecillis
|
| +-------- Cristaria_plicata
| |
+-----+ +-- Anemina_arcaeformis
| +--+
| | +-- Anemina_euscaphys
+--+
| +-- Sinanodonta_lucida
+--+
+-- Sinanodonta_woodiana
Wang et al., 2018 [Publ 3865], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
6 |
3 |
24 |
24 |
2 |
32.0 |
+----------- Quadrula_quadrula
|
+--+ +-------- Toxolasma_parvum
| | |
| +--+ +-- Potamilus_leptodon
| | +--+
| | | +-- Potamilus_alatus
| +--+
| | +-- Venustaconcha_ellipsiformis
+-----+ +--+
| | +-- Lampsilis_ornata
| |
| | +-- Sinohyriopsis_schlegelii **
| | +--------+
| | | +-- Sinohyriopsis_cumingii **
| | |
| +--+ +----- Ptychorhynchus_pfisteri **
| | +--+
| | | | +-- Sinosolenaia_oleivora **
| | | +--+
| +--+ +-- Sinosolenaia_carinata **
| |
| | +----- Potomida_littoralis **
--+ +--+
| | +-- Lamprotula_caveata **
| +--+
| +-- Lamprotula_leaii **
|
| +-------------- Aculamprotula_tientsinensis
| |
| +--+ +-- Tchangsinaia_piscicula
| | | +--------+
| | | | +-- Nodularia_douglasiae
| | +--+
| | | +-------- Unio_tumidus
| | +--+
| | | +----- Unio_crassus
| | +--+
+--+ | +-- Unio_pictorum
| +--+
| +-- Unio_delphinus
|
| +----- Acuticosta_chinensis
| +--------+
| | | +-- Lanceolaria_grayii
| | +--+
| | +-- Lanceolaria_lanceolata
+--+
| +----- Anodonta_anatina
| +-----+
| | | +-- Pyganodon_grandis
| | +--+
+--+ +-- Platynaias_compressa
|
| +-------- Cristaria_plicata
| |
+--+ +-- Anemina_euscaphys
| +--+
| | +-- Anemina_arcaeformis
+--+
| +-- Sinanodonta_woodiana
+--+
+-- Sinanodonta_lucida
Froufe et al., 2019 [Publ 3936], fig. 3-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
6 |
4 |
11 |
10 |
2 |
26.0 |
+----- Quadrula_quadrula
+-----------+
| | +-- Potamilus_alatus
| +--+
| +-- Venustaconcha_ellipsiformis
|
| +----- Unio_tumidus
| +-----+
| | | +-- Unio_crassus
--+ | +--+
| +--+ +-- Unio_delphinus
| | |
| | | +-------- Sinanodonta_woodiana
| | +--+
| | | +----- Anodonta_anatina
| | +--+
| | | +-- Utterbackia_peninsularis
+--+ +--+
| +-- Pyganodon_grandis
|
| +-- Chamberlainia_somsakpanhai **
| +--------+
| | +-- Sinohyriopsis_cumingii **
| |
+--+ +-- Potomida_littoralis **
| +-----+
| | +-- Pronodularia_japanensis **
| |
+--+ +-- Microcondylaea_bonellii **
| +--+
| | +-- Sinosolenaia_carinata **
+--+
| +-- Pilsbryoconcha_exilis **
+--+
+-- Pseudodon_vondembuschianus **
Froufe et al., 2019 [Publ 3936], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
6 |
4 |
11 |
10 |
2 |
26.0 |
+----- Quadrula_quadrula
+-----------+
| | +-- Potamilus_alatus
| +--+
| +-- Venustaconcha_ellipsiformis
|
| +----- Unio_tumidus
| +-----+
| | | +-- Unio_crassus
--+ | +--+
| +--+ +-- Unio_delphinus
| | |
| | | +-------- Sinanodonta_woodiana
| | +--+
| | | +----- Anodonta_anatina
| | +--+
| | | +-- Utterbackia_peninsularis
+--+ +--+
| +-- Pyganodon_grandis
|
| +-- Chamberlainia_somsakpanhai **
| +--------+
| | +-- Sinohyriopsis_cumingii **
| |
+--+ +-- Potomida_littoralis **
| +-----+
| | +-- Pronodularia_japanensis **
| |
+--+ +-- Microcondylaea_bonellii **
| +--+
| | +-- Sinosolenaia_carinata **
+--+
| +-- Pilsbryoconcha_exilis **
+--+
+-- Pseudodon_vondembuschianus **
Wu et al., 2020 [Publ 4121], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
6 |
3 |
42 |
34 |
2 |
32.0 |
+----------- Quadrula_quadrula
|
+-----------------+ +-------- Toxolasma_parvum
| | |
| +--+ +-- Venustaconcha_ellipsiformis
| | +--+
| | | +-- Lampsilis_ornata
| +--+
| | +-- Potamilus_alatus
| +--+
| +-- Potamilus_leptodon
|
| +-- Sinohyriopsis_schlegelii **
| +--------+
--+ | +-- Sinohyriopsis_cumingii **
| |
| +--------------+ +----- Potomida_littoralis **
| | | +--+
| | | | | +-- Pronodularia_japanensis **
| | | | +--+
| | +--+ +-- Lamprotula_leaii **
| | |
| | | +----- Ptychorhynchus_pfisteri **
| | +--+
| | | +-- Sinosolenaia_oleivora **
| | +--+
| | +-- Sinosolenaia_carinata **
+--+
| +----------------------- Lepidodesma_languilati
| |
| | +----- Acuticosta_chinensis
| | +-----------+
| | | | +-- Lanceolaria_lanceolata
| | | +--+
| | | +-- Lanceolaria_grayii
| | |
| | +--+ +----------- Anodonta_anatina
| | | | +--+
| | | | | | +-------- Platynaias_compressa
+--+ | | | +--+
| | | | | +----- Pyganodon_grandis
| | | | +--+
| | +--+ | +-- Utterbackia_peninsularis
| | | +--+
| | | +-- Utterbackia_imbecillis
| | |
| | | +-------- Cristaria_plicata
| | | |
| | +-----+ +-- Sinanodonta_woodiana
| | | +--+
+--+ | | +-- Sinanodonta_lauta
| +--+
| | +-- Anemina_arcaeformis
| +--+
| +-- Anemina_euscaphys
|
| +-------- Aculamprotula_tortuosa
| |
| +--------+ +-- Aculamprotula_coreana
| | | +--+
| | | | +-- Aculamprotula_tientsinensis
| | +--+
| | | +-- Aculamprotula_polysticta
| | +--+
+--+ +-- Aculamprotula_scripta
|
| +-------- Unio_tumidus
| +-----+
| | | +----- Unio_crassus
| | +--+
| | | +-- Unio_delphinus
+--+ +--+
| +-- Unio_pictorum
|
| +----------- Pseudocuneopsis_capitata
+--+
| +-------- Nodularia_douglasiae
+--+
| +----- Tchangsinaia_piscicula
+--+
| +-- Cuneopsis_heudei
+--+
+-- Schistodesmus_lampreyanus
Goncalves et al., 2022 [Publ 4253], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
8 |
6 |
2 |
0 |
0 |
0 |
32.0 |
+-------------- Lamprotula_caveata **
|
| +----- Chamberlainia_somsakpanhai **
--+ +-----+
| | | +-- Chamberlainia_hainesiana **
| | +--+
+--+ +-- Chamberlainia_duclerci **
|
| +-------- Sinohyriopsis_cumingii[2] **
+--+
| +----- Sinohyriopsis_cumingii[1] **
+--+
| +-- Sinohyriopsis_schlegelii **
+--+
+-- Sinohyriopsis_goliath **
Froufe et al., 2015 [Publ 3664], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
12 |
10 |
3 |
5 |
5 |
2 |
36.0 |
+-- Sinosolenaia_oleivora **
+-----+
| +-- Sinosolenaia_carinata **
+-----+
| | +----- Potomida_littoralis **
| +--+
| | +-- Lamprotula_leaii **
| +--+
--+ +-- Pronodularia_japanensis **
|
| +-- Sinohyriopsis_cumingii **
| +--------+
| | +-- Sinohyriopsis_schlegelii **
| |
+--+ +-- Venustaconcha_ellipsiformis
| +-----+
| | +-- Quadrula_quadrula
+--+
| +----- Anodonta_anatina
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Pyganodon_grandis
Froufe et al., 2016 [Publ 3727], fig. 3, 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
3 |
2 |
0 |
0 |
0 |
31.0 |
+----- Potomida_littoralis **
+--+
| | +-- Potomida_acarnanica **
| +--+
| +-- Potomida_semirugata **
--+
| +-- Pronodularia_japanensis **
| |
| +-- Lamprotula_leaii **
+-----+
+-- Leguminaia_wheatleyi **
|
+-- Microcondylaea_bonellii **
Fonseca et al., 2016 [Publ 3775], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
3 |
25 |
23 |
2 |
31.0 |
+-------- Quadrula_quadrula
+--------------+
| | +----- Toxolasma_parvum
| +--+
| | +-- Lampsilis_ornata
| +--+
| +-- Venustaconcha_ellipsiformis
|
| +-- Sinohyriopsis_cumingii **
| +--------+
| | +-- Sinohyriopsis_schlegelii **
| |
--+ +--------+ +-- Sinosolenaia_oleivora **
| | | +-----+
| | | | +-- Sinosolenaia_carinata **
| | +--+
| | | +----- Potomida_littoralis **
| | +--+
| | | +-- Lamprotula_leaii **
| | +--+
| | +-- Pronodularia_japanensis **
| |
| | +-- Lanceolaria_lanceolata
| | +-----------+
+--+ | +-- Lanceolaria_grayii
| |
| +--+ +-- Aculamprotula_tortuosa
| | | +--------+
| | | | +-- Aculamprotula_coreana
| | | |
| | +--+ +-- Unio_pictorum
| | | +-----+
| | | | +-- Unio_delphinus
| | +--+
| | | +----- Schistodesmus_lampreyanus
| | +--+
+--+ | +-- Nodularia_douglasiae
| +--+
| +-- Tchangsinaia_piscicula
|
| +-------- Cristaria_plicata
| |
| +-----+ +-- Anemina_euscaphys
| | | +--+
| | | | +-- Anemina_arcaeformis
| | +--+
+--+ | +-- Sinanodonta_woodiana
| +--+
| +-- Sinanodonta_lucida
|
| +----------- Anodonta_anatina
+--+
| +-------- Platynaias_compressa
+--+
| +----- Pyganodon_grandis
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Utterbackia_imbecillis
Zhou et al., 2017 [Publ 3788], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
3 |
10 |
10 |
2 |
32.0 |
+-------- Lanceolaria_lanceolata
|
+--------+ +-- Nodularia_douglasiae
| | +--+
| | | +-- Tchangsinaia_piscicula
| +--+
| | +-- Aculamprotula_tientsinensis
| +--+
--+ +-- Aculamprotula_tortuosa
|
| +----- Quadrula_quadrula
| +--------+
| | | +-- Potamilus_leptodon
| | +--+
| | +-- Lampsilis_ornata
+--+
| +-- Sinohyriopsis_cumingii **
| +--------+
| | +-- Sinohyriopsis_schlegelii **
| |
+--+ +-- Lamprotula_caveata **
| +-----+
| | +-- Lamprotula_leaii **
+--+
| +----- Ptychorhynchus_pfisteri **
+--+
| +-- Sinosolenaia_carinata **
+--+
+-- Sinosolenaia_oleivora **
Lopes-Lima et al., 2017 [Publ 3792], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
3 |
26 |
23 |
2 |
31.0 |
+-------- Quadrula_quadrula
+--------------+
| | +----- Toxolasma_parvum
| +--+
| | +-- Lampsilis_ornata
| +--+
| +-- Venustaconcha_ellipsiformis
|
| +-- Sinohyriopsis_cumingii **
| +--------+
| | +-- Sinohyriopsis_schlegelii **
| |
--+ +--------+ +-- Sinosolenaia_oleivora **
| | | +-----+
| | | | +-- Sinosolenaia_carinata **
| | +--+
| | | +----- Potomida_littoralis **
| | +--+
| | | +-- Pronodularia_japanensis **
| | +--+
| | +-- Lamprotula_leaii **
| |
| | +-- Lanceolaria_lanceolata
| | +-----------+
+--+ | +-- Lanceolaria_grayii
| |
| +--+ +-- Aculamprotula_tortuosa
| | | +--------+
| | | | +-- Aculamprotula_coreana
| | | |
| | +--+ +-- Unio_delphinus
| | | +-----+
| | | | +-- Unio_pictorum
| | +--+
| | | +----- Schistodesmus_lampreyanus
| | +--+
+--+ | +-- Tchangsinaia_piscicula
| +--+
| +-- Nodularia_douglasiae
|
| +----------- Anodonta_anatina
| +--+
| | | +-------- Platynaias_compressa
| | +--+
| | | +----- Pyganodon_grandis
| | +--+
+--+ | +-- Utterbackia_peninsularis
| +--+
| +-- Utterbackia_imbecillis
|
| +-------- Cristaria_plicata
| |
+-----+ +-- Sinanodonta_lucida
| +--+
| | +-- Sinanodonta_woodiana
+--+
| +-- Anemina_arcaeformis
+--+
+-- Anemina_euscaphys
Huang et al., 2019 [Publ 3843], fig. 3-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
3 |
16 |
13 |
2 |
31.0 |
+----- Quadrula_quadrula
+-----------------+
| | +-- Venustaconcha_ellipsiformis
| +--+
| +-- Potamilus_alatus
|
| +----------- Sinohyriopsis_cumingii **
| |
| | +----- Potomida_littoralis **
--+ +--------+ +--+
| | | | | +-- Lamprotula_leaii **
| | | | +--+
| | +--+ +-- Pronodularia_japanensis **
| | |
| | | +----- Parvasolenaia_rivularis **
| | +--+
| | | +-- Sinosolenaia_oleivora **
+--+ +--+
| +-- Sinosolenaia_carinata **
|
| +-------- Sinanodonta_woodiana
| +--------+
| | | +----- Anodonta_anatina
| | +--+
| | | +-- Pyganodon_grandis
+--+ +--+
| +-- Utterbackia_peninsularis
|
| +-------------- Lanceolaria_lanceolata
+--+
| +----------- Schistodesmus_lampreyanus
+--+
| +-------- Aculamprotula_tortuosa
+--+
| +----- Unio_tumidus
+--+
| +-- Unio_crassus
+--+
+-- Unio_delphinus
Aunins et al., 2018 [Publ 3863], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
3 |
34 |
31 |
2 |
31.0 |
+-------- Quadrula_quadrula
+--------------------+
| | +----- Toxolasma_parvum
| +--+
| | +-- Lampsilis_ornata
| +--+
| +-- Venustaconcha_ellipsiformis
|
| +-- Sinohyriopsis_cumingii **
| +--------+
| | +-- Sinohyriopsis_schlegelii **
--+ |
| +--------------+ +-- Lamprotula_caveata **
| | | +-----+
| | | | +-- Potomida_littoralis **
| | +--+
| | | +----- Sinosolenaia_oleivora **
| | +--+
| | | +-- Ptychorhynchus_pfisteri **
| | +--+
| | +-- Sinosolenaia_carinata **
+--+
| +----------------------- Lepidodesma_languilati
| |
| | +----- Aculamprotula_tortuosa
| | +--------+
| | | | +-- Aculamprotula_coreana
| | | +--+
| | | +-- Aculamprotula_tientsinensis
| | |
| | +-----+ +-------- Nodularia_douglasiae
+--+ | | +--+
| | | | | +----- Tchangsinaia_piscicula
| | | | +--+
| | | | | +-- Cuneopsis_heudei
| | +--+ +--+
| | | +-- Schistodesmus_lampreyanus
| | |
| | | +-------- Unio_tumidus
| | +--+
| | | +----- Unio_crassus
+--+ +--+
| | +-- Unio_pictorum
| +--+
| +-- Unio_delphinus
|
| +----- Acuticosta_chinensis
| +-----------+
| | | +-- Lanceolaria_grayii
| | +--+
| | +-- Lanceolaria_lanceolata
| |
| | +-------- Cristaria_plicata
+--+ |
| +-----+ +-- Anemina_arcaeformis
| | | +--+
| | | | +-- Anemina_euscaphys
| | +--+
| | | +-- Sinanodonta_woodiana
| | +--+
+--+ +-- Sinanodonta_lucida
|
| +-- Anodonta_anatina
| +--------+
| | +-- Anodonta_cygnea
| |
+--+ +-- Prolasmidonta_heterodon
| +-----+
| | +-- Alasmidonta_varicosa
+--+
| +----- Pyganodon_grandis
+--+
| +-- Utterbackia_imbecillis
+--+
+-- Utterbackia_peninsularis
Lopes-Lima et al., 2020 [Publ 4080], fig. 16
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
2 |
2 |
0 |
0 |
31.0 |
+----------------- Chamberlainia_somsakpanhai **
--+
| +-------------- Sinohyriopsis_schlegelii **
+--+
| +----------- Pronodularia_japanensis **
+--+
| +-------- Lamprotula_caveata **
+--+
| +----- Lamprotula_leaii[2] **
+--+
| +-- Lamprotula_seomjinensis **
+--+
+-- Lamprotula_leaii[1] **
Pfeiffer et al., 2021 [Publ 4126], fig. 30
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
1 |
0 |
0 |
0 |
31.0 |
+-------------- Rectidens_sumatrensis **
|
--+ +-- Ensidens_telus **
| +--------+
| | +-- Ensidens_ingallsianus **
+--+
| +-------- Ensidens_dugasti **
+--+
| +----- Ensidens_sagittarius **
+--+
| +-- Ensidens_spiculus **
+--+
+-- Ensidens_jaculus **
Whelan et al., 2011 [Publ 3380], fig. 1 ML
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
15 |
13 |
4 |
31 |
19 |
3 |
39.0 |
+-- Lamellidens_savadiensis
+-----------------+
| +-- Trapezidens_dolichorhynchus
+-----+
| | +----------------- Leoparreysia_tavoyensis
| | |
| +--+ +-- Radiatula_chaudhurii
| | +-----------+
| | | +-- Indochinella_pugio
| +--+
| | +----------- Nitia_teretiuscula
| +--+
| | +-------- Coelatura_aegyptiaca[2]
| +--+
--+ | +----- Coelatura_aegyptiaca[1]
| +--+
| | +-- Coelatura_gabonensis
| +--+
| +-- Prisodontopsis_aviculaeformis
|
| +-- Pseudodon_vondembuschianus **
| +--------------------+
| | +-- Lens_contradens **
| |
| | +-- Pyganodon_grandis
| | +--+
+--+ | +-- Lasmigona_costata
| +--------------+
| | | +-- Unio_pictorum
| | +--+
| | +-- Unio_caffer
+--+
| +----------------- Gonidea_angulata **
| |
| | +----- Pronodularia_japanensis **
+--+ +--------+
| | | +-- Potomida_littoralis **
| | +--+
+--+ +-- Potomida_semirugata **
|
| +----------- Pleurobema_sintoxia
+--+
| +-------- Quadrula_quadrula
+--+
| +----- Amblema_plicata
+--+
| +-- Truncilla_truncata
+--+
+-- Lampsilis_cardium
Whelan et al., 2011 [Publ 3380], fig. 1 MP
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
15 |
12 |
4 |
31 |
19 |
3 |
39.0 |
+-- Lamellidens_savadiensis
+--------------+
| +-- Trapezidens_dolichorhynchus
+--------+
| | +-------------- Leoparreysia_tavoyensis
| | |
| +--+ +-- Radiatula_chaudhurii
| | +--------+
| | | +-- Indochinella_pugio
| +--+
| | +-------- Nitia_teretiuscula
| | |
| +--+ +----- Coelatura_aegyptiaca[1]
--+ | |
| +--+----- Coelatura_aegyptiaca[2]
| |
| | +-- Coelatura_gabonensis
| +--+
| +-- Prisodontopsis_aviculaeformis
|
| +----------------------- Lens_contradens **
| |
| | +-------------------- Pseudodon_vondembuschianus **
+--+ |
| | +-- Pyganodon_grandis
| | +--+
+--+ | +-- Lasmigona_costata
| +-----------+
| | | +-- Unio_pictorum
| | +--+
| | +-- Unio_caffer
+--+
| +-------------- Gonidea_angulata **
| |
| | +----- Pronodularia_japanensis **
+--+ +-----+
| | | +-- Potomida_littoralis **
| | +--+
+--+ +-- Potomida_semirugata **
|
| +-------- Quadrula_quadrula
| |
+--+-------- Pleurobema_sintoxia
|
| +----- Amblema_plicata
+--+
| +-- Lampsilis_cardium
+--+
+-- Truncilla_truncata
Zhou et al., 2016 [Publ 3710], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
3 |
14 |
13 |
2 |
31.0 |
+-------- Quadrula_quadrula
+--+
| | +----- Toxolasma_parvum
| +--+
| | +-- Venustaconcha_ellipsiformis
| +--+
--+ +-- Lampsilis_ornata
|
| +-- Sinohyriopsis_cumingii **
| +-----+
| | +-- Sinohyriopsis_schlegelii **
| |
+--+ +-- Lamprotula_leaii **
| +--+
| | +-- Pronodularia_japanensis **
+--+
| +-- Sinosolenaia_carinata **
+--+
+-- Sinosolenaia_oleivora **
Wu et al., 2016 [Publ 3711], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
3 |
15 |
15 |
2 |
31.0 |
+----- Quadrula_quadrula
+-----+
| | +-- Venustaconcha_ellipsiformis
| +--+
| +-- Lampsilis_ornata
+-----------+
| | +-- Sinohyriopsis_schlegelii **
| | +-----+
| | | +-- Sinohyriopsis_cumingii **
| | |
| +--+ +-- Sinosolenaia_carinata **
| | +--+
| | | +-- Sinosolenaia_oleivora **
| +--+
| | +-- Lamprotula_leaii **
--+ +--+
| +-- Pronodularia_japanensis **
|
| +-- Unio_pictorum
| +-----+
| | +-- Nodularia_douglasiae
| +-----------+
| | | +----- Aculamprotula_tortuosa
| | +--+
| | | +-- Aculamprotula_coreana
+--+ +--+
| +-- Aculamprotula_tientsinensis
|
| +----------------- Sinanodonta_lucida
+--+
| +-------------- Anemina_arcaeformis
+--+
| +----------- Cristaria_plicata
+--+
| +-------- Anodonta_anatina
+--+
| +----- Pyganodon_grandis
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Utterbackia_imbecillis
Zieritz et al., 2016 [Publ 3804], fig. S2-1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
3 |
1 |
0 |
0 |
0 |
28.0 |
+----------- Rectidens_sumatrensis **
|
--+ +-------- Hyriopsis_desowitzi **
| |
+--+ +----- Hyriopsis_khoratensis **
| |
+--+ +-- Hyriopsis_myersiana **
| |
+--+-- Hyriopsis_panhai **
|
+-- Hyriopsis_bialata **
Bolotov et al., 2020 [Publ 4084], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
0 |
2 |
30 |
26 |
2 |
29.0 |
+-- Lamprotula_leaii **
|
+-- Potomida_littoralis **
|
+-- Pronodularia_japanensis **
+-----------------------+
| +-- Gonidea_angulata **
| |
| +-- Leguminaia_wheatleyi **
| |
| +-- Microcondylaea_bonellii **
|
--+ +----------- Middendorffinaia_mongolica
| +-----------+
| | | +-------- Nodularia_douglasiae
| | +--+
| | | +----- Unio_tumidus
| | +--+
| | | +-- Unio_pictorum
| | +--+
+--+ +-- Unio_crassus
|
| +-- Lanceolaria_grayii
| +-----------------+
| | +-- Lanceolaria
| |
| | +----- Anodonta_anatina
+--+ +-----------+
| | | +-- Pseudanodonta_complanata
| | +--+
| | +-- Anodonta_cygnea
| |
+--+ +-------- Amuranodonta_kijaensis
| +-----+
| | | +----- Anemina_arcaeformis
| | +--+
| | | +-- Buldowskia_suifunica
+--+ +--+
| +-- Buldowskia_shadini
|
| +----------- Cristaria_plicata
+--+
| +-------- Beringiana_beringiana
+--+
| +----- Sinanodonta_schrenkii
+--+
| +-- Sinanodonta_lauta
+--+
+-- Sinanodonta_woodiana
Wu et al., 2018 [Publ 4143], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
3 |
16 |
16 |
2 |
31.0 |
+----- Quadrula_quadrula
+-----+
| | +-- Venustaconcha_ellipsiformis
| +--+
| +-- Lampsilis_ornata
+-----------+
| | +-- Sinohyriopsis_schlegelii **
| | +-----+
| | | +-- Sinohyriopsis_cumingii **
| | |
| +--+ +-- Sinosolenaia_oleivora **
| | +--+
| | | +-- Sinosolenaia_carinata **
| +--+
| | +-- Pronodularia_japanensis **
--+ +--+
| +-- Lamprotula_leaii **
|
| +-- Unio_pictorum
| +--------+
| | +-- Nodularia_douglasiae
| +--------+
| | | +-------- Aculamprotula_tortuosa
| | +--+
| | | +----- Aculamprotula_scripta
| | +--+
+--+ | +-- Aculamprotula_coreana
| +--+
| +-- Aculamprotula_tientsinensis
|
| +----------------- Sinanodonta_lucida
+--+
| +-------------- Anemina_arcaeformis
+--+
| +----------- Cristaria_plicata
+--+
| +-------- Anodonta_anatina
+--+
| +----- Pyganodon_grandis
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Utterbackia_imbecillis
Lopes-Lima et al., 2021 [Publ 4217], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
2 |
19 |
19 |
1 |
31.0 |
+----- Potomida_littoralis **
+--+
| | +-- Potomida_semirugata **
| +--+
+--------------------+ +-- Potomida_acarnanica **
| |
| | +----- Leguminaia_anatolica **
| +--+
| | +-- Leguminaia_saulcyi **
| +--+
--+ +-- Leguminaia_wheatleyi **
|
| +----- Anodonta_anatina
| +--------------------+
| | | +-- Anodonta_cygnea
| | +--+
| | +-- Anodonta_seddoni
+--+
| +----------------------- Unio_tumidus
| |
| | +-------- Unio_damascensis
| | +-----------+
+--+ | | +----- Unio_sesirmensis
| | +--+
| | | +-- Unio_bruguierianus
| | +--+
+--+ +-- Unio_crassus
|
| +-- Unio_delicatus
| +--------------+
| | +-- Unio_terminalis
| |
+--+ +-- Unio_tigridis
| +-----------+
| | +-- Unio_hueti
| |
+--+ +----------- Unio_ravoisieri
| |
| | +-- Unio_pictorum
+--+ +-----+
| | +-- Unio_eucirrus
| |
+--+ +-- Unio_elongatulus
| +--+
| | +-- Unio_mancus
+--+
| +-- Unio_foucauldianus
+--+
+-- Unio_delphinus
Zhou et al., 2007 [Publ 2658], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
6 |
4 |
2 |
18 |
15 |
1 |
32.0 |
+-- Sinanodonta_woodiana
+--------------------+
| +-- Cristaria_plicata
|
| +-- Sinohyriopsis_cumingii **
| +--------+
| | +-- Sinohyriopsis_schlegelii **
| +--------+
--+ | | +-------- Lamprotula_leaii **
| | +--+
| | | +----- Lamprotula_caveata **
| | +--+
| | | +-- Gibbosula_rochechouartii
| | +--+
| | +-- Lamprotula_cornuumlunae **
| |
+--+ +-- Schistodesmus_lampreyanus
| +-----+
| | +-- Schistodesmus_spinosus
| |
| +--------+ +-- Lanceolaria_triformis
| | | +--+
| | | | +-- Lanceolaria_lanceolata
| | +--+
| | | +-- Acuticosta_ovata
+--+ +--+
| +-- Acuticosta_chinensis
|
| +-- Tchangsinaia_piscicula
| +-----------+
| | +-- Nodularia_douglasiae
+--+
| +----------- Aculamprotula_tortuosa
+--+
| +-------- Aculamprotula_scripta
+--+
| +----- Aculamprotula_tientsinensis
+--+
| +-- Aculamprotula_zonata
+--+
+-- Aculamprotula_fibrosa
Guerra et al., 2017 [Publ 3772], fig. 2a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
3 |
20 |
14 |
2 |
30.0 |
+-------- Quadrula_quadrula
+--+
| | +----- Toxolasma_parvum
| +--+
| | +-- Venustaconcha_ellipsiformis
| +--+
+-----------+ +-- Lampsilis_ornata
| |
| | +-- Sinohyriopsis_schlegelii **
| | +-----+
| | | +-- Sinohyriopsis_cumingii **
| +--+
| | +----- Pronodularia_japanensis **
--+ +--+
| | +-- Sinosolenaia_oleivora **
| +--+
| +-- Sinosolenaia_carinata **
|
| +-- Aculamprotula_tortuosa
| +-----------------+
| | +-- Unio_pictorum
+--+
| +----------------- Sinanodonta_woodiana
| |
+--+ +-------------- Cristaria_plicata
| |
+--+ +----------- Anodonta_anatina
| |
+--+ +-- Platynaias_subviridis
| +-----+
| | +-- Platynaias_compressa
+--+
| +----- Pyganodon_grandis
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Utterbackia_imbecillis
Guerra et al., 2017 [Publ 3772], fig. 2b
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
3 |
20 |
14 |
2 |
30.0 |
+-------- Quadrula_quadrula
+-----------------+
| | +----- Toxolasma_parvum
| +--+
| | +-- Venustaconcha_ellipsiformis
| +--+
| +-- Lampsilis_ornata
|
--+ +-- Sinohyriopsis_schlegelii **
| +-----+
| | +-- Sinohyriopsis_cumingii **
| +--------------+
| | | +----- Pronodularia_japanensis **
| | +--+
| | | +-- Sinosolenaia_oleivora **
+--+ +--+
| +-- Sinosolenaia_carinata **
|
| +-------------------- Unio_pictorum
| |
+--+ +----------------- Aculamprotula_tortuosa
| |
+--+ +-- Cristaria_plicata
| +-----------+
| | +-- Sinanodonta_woodiana
+--+
| +----------- Anodonta_anatina
| |
+--+ +-- Platynaias_subviridis
| +-----+
| | +-- Platynaias_compressa
+--+
| +----- Pyganodon_grandis
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Utterbackia_imbecillis
Burzynski et al., 2017 [Publ 3774], fig. 1 bottom
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
3 |
8 |
8 |
2 |
29.0 |
+----- Aculamprotula_tortuosa
+--+
| | +-- Unio_crassus
| +--+
+--------+ +-- Unio_tumidus
| |
| | +----- Anodonta_anatina
| +--+
| | +-- Utterbackia_peninsularis
--+ +--+
| +-- Pyganodon_grandis
|
| +-- Quadrula_quadrula
| +-----------+
| | +-- Venustaconcha_ellipsiformis
+--+
| +----------- Sinohyriopsis_cumingii **
+--+
| +-------- Sinosolenaia_carinata **
+--+
| +----- Potomida_littoralis **
+--+
| +-- Lamprotula_leaii **
+--+
+-- Pronodularia_japanensis **
Burzynski et al., 2017 [Publ 3774], fig. 1 top
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
3 |
8 |
8 |
2 |
29.0 |
+----- Aculamprotula_tortuosa
+--+
| | +-- Unio_crassus
| +--+
+--------+ +-- Unio_tumidus
| |
| | +----- Anodonta_anatina
| +--+
| | +-- Utterbackia_peninsularis
--+ +--+
| +-- Pyganodon_grandis
|
| +-- Quadrula_quadrula
| +-----------+
| | +-- Venustaconcha_ellipsiformis
+--+
| +----------- Sinohyriopsis_cumingii **
+--+
| +-------- Sinosolenaia_carinata **
+--+
| +----- Lamprotula_leaii **
+--+
| +-- Potomida_littoralis **
+--+
+-- Pronodularia_japanensis **
Araujo et al., 2017 [Publ 3801], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
2 |
9 |
8 |
3 |
27.0 |
+-- Amblema_plicata
+-----------+
| +-- Lampsilis_teres
|
| +-- Schistodesmus_lampreyanus
| +--+
--+ | +-- Aculamprotula_tortuosa
| +-----+
| | | +-- Anodonta_anatina
| | +--+
| | +-- Unio_delphinus
+--+
| +-- Gonidea_angulata **
| +-----+
| | +-- Microcondylaea_bonellii **
+--+
| +----- Potomida_littoralis **
+--+
| +-- Lamprotula_leaii **
+--+
+-- Pronodularia_japanensis **
Burzynski & Soroka, 2018 [Publ 3862], fig. 5a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
3 |
15 |
13 |
2 |
29.0 |
+----- Quadrula_quadrula
+-----------------+
| | +-- Potamilus_alatus
| +--+
| +-- Venustaconcha_ellipsiformis
|
| +----------- Sinohyriopsis_cumingii **
--+ +--------+
| | | +-------- Sinosolenaia_carinata **
| | +--+
| | | +----- Potomida_littoralis **
| | +--+
| | | +-- Lamprotula_leaii **
| | +--+
+--+ +-- Pronodularia_japanensis **
|
| +----- Sinanodonta_woodiana
| |
| +-----------+----- Anodonta_anatina
| | |
| | | +-- Utterbackia_peninsularis
+--+ +--+
| +-- Pyganodon_grandis
|
| +-------------- Lanceolaria_lanceolata
+--+
| +----------- Aculamprotula_tortuosa
+--+
| +-------- Unio_tumidus
+--+
| +----- Unio_crassus
+--+
| +-- Unio_pictorum
+--+
+-- Unio_delphinus
Bolotov et al., 2019 [Publ 3974], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
1 |
13 |
13 |
1 |
29.0 |
+----------- Yaukthwa_inlenensis **
+--------+
| | +-------- Yaukthwa_elongatula **
| +--+
| | +----- Yaukthwa_panhai **
| +--+
| | +-- Yaukthwa_nesemanni **
| +--+
| +-- Yaukthwa_rectangularis **
--+
| +-- Leoparreysia_whitteni
| +-----+
| | +-- Leoparreysia_burmana
| +--------+
| | | +----- Leoparreysia_olivacea
| | +--+
| | | +-- Leoparreysia_tavoyensis
| | +--+
+--+ +-- Leoparreysia_canefrii
|
| +-- Nitia_teretiuscula
| +-----------+
| | +-- Coelatura_aegyptiaca
+--+
| +----------- Indochinella_pugio
| |
+--+ +-- Radiatula_myitkyinae
| +-----+
| | +-- Radiatula_chaudhurii
+--+
| +----- Radiatula_chindwinensis
+--+
| +-- Radiatula_mouhoti
+--+
+-- Radiatula_myitthanensis
Wu et al., 2020 [Publ 4147], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
2 |
9 |
6 |
2 |
30.0 |
+----------------- Lampsilis_cardium
|
| +----------- Sinosolenaia_carinata **
| +--+
--+ | | +-------- Potomida_littoralis **
| | +--+
| | | +----- Lamprotula_leaii **
| | +--+
+--+ | +-- Lamprotula_caveata **
| +--+
| +-- Lamprotula_cornuumlunae **
|
| +----------- Unio_pictorum
+--+
| +-------- Aculamprotula_tortuosa
+--+
| +----- Aculamprotula_scripta
+--+
| +-- Aculamprotula_tientsinensis
+--+
+-- Aculamprotula_fibrosa
Graf & Cummings, 2006 [Publ 1968], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
25 |
19 |
3 |
43 |
21 |
3 |
47.0 |
+-- Pilsbryoconcha_exilis **
+-----------------------------+
| +-- Pseudodon_vondembuschianus **
|
| +----------------------------- Coelatura_aegyptiaca
| |
| +----------------------------- Lens_contradens **
--+ |
| | +-- Unio_pictorum
| | +--------+
| | | +-- Unio_caffer
| +-----------------+
| | | +-------- Lasmigona_costata
+--+ +--+
| | +----- Alasmidonta_marginata
| +--+
| | +-- Pyganodon_grandis
| +--+
| +-- Strophitus_undulatus
|
| +-------------------------- Gonidea_angulata **
| |
| | +-- Quadrula_verrucosa
+--+ +--------------------+
| | +-- Quadrula_quadrula
| |
+--+ +-------------------- Amblema_plicata
| |
| | +----- Eurynia_dilatata
+--+ +-----------+
| | | +-- Fusconaia_flava
| | +--+
| | +-- Pleurobema_sintoxia
+--+
| +-------------- Obliquaria_reflexa
| |
| +-------------- Truncilla_truncata
+--+
+-------------- Ptychobranchus_fasciolaris
|
| +----------- Ligumia_recta
+--+
| +-------- Cambarunio_iris
+--+
| +----- Epioblasma_triquetra
+--+
| +-- Lampsilis_cardium
+--+
+-- Ortmanniana_ligamentina
Graf, 2002 [Publ 2651], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
12 |
10 |
3 |
14 |
8 |
2 |
34.0 |
+-- Pilsbryoconcha_exilis **
+--------------+
| +-- Pseudodon_vondembuschianus **
|
--+ +-------------- Lens_contradens **
| |
| | +-- Unio_pictorum
| | +--+
+--+ | +-- Unio_caffer
| +-----+
| | | +-- Alasmidonta_marginata
| | +--+
+--+ +-- Pyganodon_grandis
|
| +-------- Gonidea_angulata **
| |
+--+ +-- Lampsilis_cardium
| +--+
| | +-- Obliquaria_reflexa
+--+
| +-- Eurynia_dilatata
+--+
+-- Amblema_plicata
Huang et al., 2002 [Publ 2657], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
18 |
16 |
3 |
30 |
28 |
2 |
45.0 |
+-- Sinanodonta_woodiana
+-----------+
| +-- Cristaria_plicata
+--+
| | +----------- Utterbackiana_couperiana
| +--+
| | +-------- Utterbackia_imbecillis
| +--+
| | +----- Strophitus_radiatus
| +--+
| | +-- Alasmidonta_triangulata
+--------------------------+ +--+
| | +-- Lasmigona_costata
| |
| | +----- Acuticosta_ovata
| | +--+
| | | | +-- Nodularia_douglasiae
| | | +--+
| | | +-- Tchangsinaia_piscicula
| +--------+
--+ | +-- Lanceolaria_grayii
| | +--+
| | | +-- Lanceolaria_lanceolata
| +--+
| | +-- Schistodesmus_lampreyanus
| +--+
| +-- Lepidodesma_languilati
|
| +----------------------------------------- Sinosolenaia_oleivora **
| |
+--+ +-------------------------------------- Lamprotula_leaii **
| |
+--+ +----------------------------------- Sinohyriopsis_cumingii **
| |
+--+ +-------------------------------- Ptychorhynchus_pfisteri **
| |
+--+ +----------------------------- Megalonaias_nervosa
| |
| | +----- Elliptoideus_sloatianus
+--+ +--------------------+
| | | +-- Uniomerus_carolinianus
| | +--+
+--+ +-- Pleurobema_pyriforme
|
| +----------------------- Amblema_plicata
| |
+--+ +-- Obliquaria_reflexa
| +-----------------+
| | +-- Plectomerus_dombeyanus
+--+
| +----------------- Medionidus_conradicus
+--+
| +-------------- Potamilus_fragilis
+--+
| +----------- Ptychobranchus_subtentus
+--+
| +-------- Potamilus_purpuratus
+--+
| +----- Lampsilis_teres
+--+
| +-- Villosa_delumbis
+--+
+-- Ellipsaria_lineolata
Huang et al., 2002 [Publ 2657], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
18 |
7 |
3 |
30 |
28 |
2 |
44.0 |
+-------- Strophitus_radiatus
|
+-------- Schistodesmus_lampreyanus
|
+-------- Lepidodesma_languilati
|
+-------- Acuticosta_ovata
|
| +-- Sinanodonta_woodiana
+-----+
| +-- Cristaria_plicata
+--------+
| | +-- Lanceolaria_grayii
| +-----+
| | +-- Lanceolaria_lanceolata
| |
| | +-- Nodularia_douglasiae
| +-----+
| | +-- Tchangsinaia_piscicula
| |
| | +----- Utterbackia_imbecillis
| | |
--+ +--+----- Utterbackiana_couperiana
| |
| | +-- Alasmidonta_triangulata
| +--+
| +-- Lasmigona_costata
|
| +-------------- Ptychorhynchus_pfisteri **
| |
| +-------------- Sinohyriopsis_cumingii **
| |
| +-------------- Lamprotula_leaii **
| |
+--+-------------- Sinosolenaia_oleivora **
|
| +----------- Megalonaias_nervosa
| |
| | +----- Elliptoideus_sloatianus
| | +--+
+--+ | | +-- Uniomerus_carolinianus
| | +--+
| | +-- Pleurobema_pyriforme
| |
| | +----- Potamilus_fragilis
+--+ |
| +----- Medionidus_conradicus
| |
| +----- Amblema_plicata
| |
| | +-- Obliquaria_reflexa
+--+--+
| +-- Plectomerus_dombeyanus
|
| +-- Villosa_delumbis
| |
| +-- Ellipsaria_lineolata
| |
+--+-- Lampsilis_teres
|
+-- Potamilus_purpuratus
|
+-- Ptychobranchus_subtentus
Campbell et al., 2005 [Publ 2660], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
115 |
102 |
3 |
113 |
111 |
2 |
133.0 |
+-------------------------------------------------- Sinohyriopsis_cumingii **
|
| +----------- Anodonta_oregonensis
| |
+--------------------------------------+ +-- Anodonta_cygnea
| | +-----+
| | | +-- Alasmidonta_holstonia
| +--+
| | +----- Pyganodon_grandis
| +--+
| | +-- Lasmigona_costata
| +--+
| +-- Pseudodontoideus_subvexus
|
| +----- Gonidea_angulata **
| +-----------------------------------------+
| | | +-- Potomida_littoralis **
--+ | +--+
| | +-- Pronodularia_japanensis **
| |
| | +-- Megalonaias_nervosa
| | +-----------+
| | | +-- Uniomerus_declivis
| | |
| | +-----------------------------+ +----------- Theliderma_metanevra
| | | | |
| | | | | +----- Quadrula_verrucosa
| | | +--+ +--+
| | | | | | +-- Quadrula_quadrula
| | | | | +--+
| | | | | +-- Quadrula_nobilis
| | | +--+
| | | | +----- Pustulosa_kieneriana
| | | | |
| | | | | +-- Cyclonaias_tuberculata
+--+ | +--+--+
| | | +-- Pustulosa_succissa
| | |
| | | +-- Pustulosa_infucata
| | +--+
| | +-- Pustulosa_kleiniana
| |
| | +----------------------------------- Elliptoideus_sloatianus
| | |
| | | +-------------------------------- Hemistena_lata
| | | |
| | +-----+ | +----- Plethobasus_cyphyus
| | | | | +-----------------------+
| | | | | | | +-- Elliptio_arca
| | | | | | +--+
| | | | | | +-- Elliptio_crassidens
| | | +--+ |
| | | | | +-- Eurynia_dilatata
| | | | | +-----------------------+
| | | | | | +-- Parvaspina_collina
+--+ | | | |
| | | | | +----- Pleuronaia_gibber
| | | | +--------------------+
| | | | | | +-- Pleuronaia_barnesiana
| | +--+ | +--+
| | | | +-- Pleuronaia_dolabelloides
| | | |
| | | | +----------- Fusconaia_subrotunda
| | | | |
| | | | | +-- Fusconaia_cor
| | | +--------------+ +-----+
| | | | | | +-- Fusconaia_cuneolus
| | | | | |
| | | | | | +-- Fusconaia_cerina[1]
| | | | +--+ |
| | | | | +--+-- Fusconaia_cerina[2]
| | +--+ | | |
| | | | | +-- Fusconaia_flava
| | | +--+
| | | | +-- Fusconaia_burkei[1]
| | | | |
| | | | +-- Fusconaia_burkei[2]
| | | +--+
| | | +-- Fusconaia_burkei[3]
| | | |
| | | +-- Fusconaia_escambia
| | |
| | | +-- Pleurobema_rubrum[2]
| | | +--+
| | | | +-- Pleurobema_cordatum
| | | +-----------------+
| | | | | +-- Pleurobema_rubrum[1]
+--+ | | +--+
| | | +-- Pleurobema_sintoxia
| +--+
| | +-------------------- Pleurobema_pyriforme
| | |
| | | +-- Pleurobema_georgianum[2]
| | | |
| +--+ +--------------+-- Pleurobema_georgianum[3]
| | | |
| | | +-- Pleurobema_athearni
| | |
| | | +----- Pleurobema_decisum
| +--+ |
| | +--------+ +-- Pleurobema_hanleyianum[1]
| | | | |
| | | +--+-- Pleurobema_hanleyianum[2]
| | | |
| | | +-- Pleurobema_georgianum[1]
| +--+
| | +-- Pleurobema_oviforme[1]
| | |
| | +--------+-- Pleurobema_oviforme[2]
| | | |
| +--+ +-- Pleurobema_clava
| |
| | +-------- Pleurobema_strodeanum
| +--+
| | +----- Pleurobema_rubellum
| +--+
| | +-- Pleurobema_perovatum
| +--+
| +-- Pleurobema_taitianum
|
| +----- Plectomerus_dombeyanus
| +--------------------------------+
| | | +-- Reginaia_ebenus
| | +--+
| | +-- Reginaia_rotulata
| |
| | +----- Popenaias_popeii
| | +-----------------------------+
| | | | +-- Amblema_elliottii
+--+ | +--+
| | +-- Amblema_plicata
| |
| | +-- Toxolasma_parvum
| | +-----+
| | | +-- Toxolasma_texasiense
| | +-----------------------+
| | | | +----- Obliquaria_reflexa
+--+ | +--+
| | | +-- Cyrtonaias_tampicoensis
| | +--+
| | +-- Glebula_rotundata
| |
| | +-- Potamilus_fragilis
| | +--+
| | | +-- Potamilus_leptodon
| | +-----------+
| | | | +-- Potamilus_alatus
| | | +--+
+--+ | +-- Potamilus_purpuratus
| |
| +-----------+ +-- Sagittunio_nasutus
| | | +--+
| | | | +-- Medionidus_conradicus
| | | +--------+
| | | | | +-- Ptychobranchus_fasciolaris
| | | | +--+
| | +--+ +-- Venustaconcha_ellipsiformis[2]
| | |
| | | +----------- Medionidus_acutissimus
| | +--+
| | | +-------- Lemiox_rimosus
| | +--+
+--+ | +----- Dromus_dromas
| +--+
| | +-- Cyprogenia_aberti
| +--+
| +-- Cyprogenia_stegaria
|
| +-- Ellipsaria_lineolata
| +-----------------------+
| | +-- Truncilla_truncata
| |
| | +----------------------- Villosa_villosa
| | |
| | | +-- Obovaria_olivaria
+--+ | +--------------+
| | | +-- Venustaconcha_ellipsiformis[1]
| | |
| | +--+ +----- Obovaria_unicolor
| | | | +--------+
| | | | | | +-- Obovaria_arkansasensis
+--+ | | | +--+
| | +--+ +-- Obovaria_subrotunda
| | |
| | | +----------- Epioblasma_brevidens
| | +--+
| | | +-------- Epioblasma_triquetra
| | +--+
| | | +----- Epioblasma_walkeri[2]
| | +--+
| | | +-- Epioblasma_capsaeformis
+--+ +--+
| +-- Epioblasma_walkeri[1]
|
| +-- Ligumia_recta
| +--------+
| | +-- Cambarunio_iris
| +--+
| | | +-------- Lampsilis_teres
| | +--+
| | | +----- Ortmanniana_ligamentina
| | +--+
| | | +-- Lampsilis_siliquoidea
| | +--+
| | +-- Leaunio_vanuxemensis
+-----+
| +-- Lampsilis_cardium
| +--+
| | +-- Lampsilis_ovata
| +-----+
| | | +-- Ortmanniana_pectorosa
| | +--+
| | +-- Lampsilis_ornata
| |
+--+ +----- Hamiota_australis[2]
| +--+
| | | +-- Hamiota_australis[1]
| | +--+
| | +-- Hamiota_subangulata
+--+
| +----- Hamiota_altilis[2]
| |
| | +-- Hamiota_altilis[1]
+--+--+
| +-- Hamiota_perovalis[1]
|
| +-- Hamiota_altilis[3]
+--+
+-- Hamiota_perovalis[2]
Campbell et al., 2005 [Publ 2660], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
113 |
91 |
3 |
111 |
109 |
2 |
132.0 |
+----------- Sinohyriopsis_cumingii **
|
+--------------------------------+ +-------- Pyganodon_grandis
| | |
| | | +-- Lasmigona_costata
| +--+-----+
| | +-- Pseudodontoideus_subvexus
| |
| | +----- Alasmidonta_holstonia
| +--+
| | +-- Anodonta_cygnea
| +--+
| +-- Anodonta_oregonensis
|
| +----- Gonidea_angulata **
| +-----------------------------------+
| | | +-- Potomida_littoralis **
| | +--+
--+ | +-- Pronodularia_japanensis **
| |
| | +-------------------- Megalonaias_nervosa
| | |
| | +-----------------+ +----------------- Uniomerus_declivis
| | | | |
| | | +--+ +-------------- Theliderma_metanevra
| | | | |
| | | +--+ +----------- Quadrula_quadrula
| | | | |
| | | | | +-- Quadrula_nobilis
| | | +--+ +-----+
| | | | | +-- Quadrula_verrucosa
| | | | |
| | | +--+ +----- Pustulosa_kieneriana
| | | | |
+--+ | | | +-- Cyclonaias_tuberculata
| | +--+--+
| | | +-- Pustulosa_succissa
| | |
| | | +-- Pustulosa_infucata
| | +--+
| | +-- Pustulosa_kleiniana
| |
| | +-------------------------------- Elliptoideus_sloatianus
| | |
| | | +----------------------------- Plethobasus_cyphyus
| | | |
| | | +----------------------------- Parvaspina_collina
| | +--+ |
| | | | | +-- Elliptio_arca
| | | | +--------------------------+
| | | | | +-- Elliptio_crassidens
| | | | |
| | | | | +-------------------------- Eurynia_dilatata
| | | +--+ |
+--+ | | +-------------------------- Hemistena_lata
| | | |
| | | | +-- Pleurobema_rubrum[1]
| | | | +--+
| | | | | +-- Pleurobema_sintoxia
| | | | +-----------------+
| | | | | | +-- Pleurobema_rubrum[2]
| | | | | +--+
| | +--+ | +-- Pleurobema_cordatum
| | | |
| | | | +-------------- Fusconaia_subrotunda
| | | | |
| | | | | +-- Fusconaia_cor
| | | +--------+ +--------+
| | | | | | +-- Fusconaia_cuneolus
| | | | | |
| | | | +--+ +-- Fusconaia_cerina
| | | | | +-----+
| | | | | | +-- Fusconaia_flava
| | +--+ +--+
| | | | +----- Fusconaia_burkei[2]
| | | +--+
| | | | +-- Fusconaia_escambia
| | | +--+
| | | +-- Fusconaia_burkei[1]
| | |
| | | +----- Pleuronaia_gibber
| | | +--------------+
| | | | | +-- Pleuronaia_barnesiana
| | | | +--+
| | | | +-- Pleuronaia_dolabelloides
| | | |
| | | | +-------- Pleurobema_rubellum
+--+ +--+ +--------+
| | | | +----- Pleurobema_strodeanum
| | | +--+
| | | | +-- Pleurobema_perovatum
| | | +--+
| | | +-- Pleurobema_taitianum
| +--+
| | +-- Pleurobema_oviforme[1]
| | |
| | +-----------+-- Pleurobema_oviforme[2]
| | | |
| | | +-- Pleurobema_clava
| +--+
| | +----------- Pleurobema_pyriforme
| | |
| +--+ +-- Pleurobema_athearni
| | +-----+
| | | +-- Pleurobema_georgianum[2]
| +--+
| | +----- Pleurobema_decisum
| | |
| +--+ +-- Pleurobema_hanleyianum[1]
| | |
| +--+-- Pleurobema_hanleyianum[2]
| |
| +-- Pleurobema_georgianum[1]
|
| +-------- Popenaias_popeii
| |
| +--------------+ +-- Amblema_elliottii
| | | +--+
| | | | +-- Amblema_plicata
| | +--+
| | | +-- Reginaia_ebenus
| | | |
| | +--+-- Reginaia_rotulata[1]
| | |
| | +-- Reginaia_rotulata[2]
| |
| | +----- Plectomerus_dombeyanus
| | +--------------+
+-----------+ | | +-- Toxolasma_parvum
| | +--+
| | +-- Toxolasma_texasiense
| |
| | +----- Obliquaria_reflexa
| | +-----------+
| | | | +-- Cyrtonaias_tampicoensis
| | | +--+
| | | +-- Glebula_rotundata
| | |
| | | +-------------- Sagittunio_nasutus
| | | |
+--+ | +-------------- Leaunio_vanuxemensis
| | |
| | | +-- Ellipsaria_lineolata
| | +-----------+
| | | +-- Truncilla_truncata
| | |
| | | +----- Lampsilis_teres
| | +--------+
| | | | +-- Ortmanniana_ligamentina
| | | +--+
| | | +-- Lampsilis_siliquoidea
| | |
| | | +----- Villosa_villosa
| | +--------+
+--+ | | +-- Ligumia_recta
| | +--+
| | +-- Cambarunio_iris
| |
| | +-------- Ortmanniana_pectorosa
| +-----+
| | | +----- Lampsilis_ornata
| | +--+
| | | +-- Lampsilis_cardium
| | +--+
| | +-- Lampsilis_ovata
| |
| | +-- Potamilus_fragilis
| | +--+
| | | +-- Potamilus_leptodon
| +--------+
| | | +-- Potamilus_alatus
| | +--+
| | +-- Potamilus_purpuratus
| |
| | +-- Ptychobranchus_fasciolaris
+--+ +--------+
| | +-- Venustaconcha_ellipsiformis[1]
| |
+--+ +-------- Medionidus_conradicus
| | |
| | | +-- Lemiox_rimosus
| +--+-----+
| | +-- Medionidus_acutissimus
| |
| | +----- Dromus_dromas
| +--+
| | +-- Cyprogenia_aberti
| +--+
| +-- Cyprogenia_stegaria
|
| +----- Hamiota_australis[2]
| +-----+
| | | +-- Hamiota_australis[1]
| | +--+
| | +-- Hamiota_subangulata
+--+
| | +-- Hamiota_altilis[3]
| | +-----+
| | | +-- Hamiota_perovalis[2]
| +--+
| | +----- Hamiota_altilis[2]
| +--+
| | +-- Hamiota_altilis[1]
| +--+
| +-- Hamiota_perovalis[1]
|
| +----------- Obovaria_olivaria
| |
| +----------- Venustaconcha_ellipsiformis[2]
| |
| | +----- Obovaria_unicolor
| +-----+
+--+ | +-- Obovaria_arkansasensis
| +--+
| +-- Obovaria_subrotunda
|
| +-- Epioblasma_brevidens
| +-----+
| | +-- Epioblasma_triquetra
+--+
| +----- Epioblasma_walkeri[1]
+--+
| +-- Epioblasma_walkeri[2]
+--+
+-- Epioblasma_capsaeformis
Hoeh et al., 2009 [Publ 2716], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
18 |
16 |
3 |
43 |
21 |
3 |
38.0 |
+-- Unio_pictorum
+--------+
| +-- Unio_caffer
+-----------------+
| | +-------- Lasmigona_costata
| +--+
| | +----- Alasmidonta_marginata
| +--+
| | +-- Strophitus_undulatus
| +--+
--+ +-- Pyganodon_grandis
|
| +-------------------------- Lens_contradens **
| |
| | +----- Gonidea_angulata **
| | +-----------------+
| | | | +-- Pilsbryoconcha_exilis **
+--+ | +--+
| | +-- Pseudodon_vondembuschianus **
| |
| | +----------- Amblema_plicata
| | |
+--+ | +----- Obliquaria_reflexa
| +--------+ +--+
| | | | | +-- Quadrula_verrucosa
| | | | +--+
| | +--+ +-- Quadrula_quadrula
| | |
| | | +----- Eurynia_dilatata
+--+ +--+
| | +-- Pleurobema_sintoxia
| +--+
| +-- Fusconaia_flava
|
| +----------------- Ptychobranchus_fasciolaris
+--+
| +-------------- Truncilla_truncata
+--+
| +----------- Ortmanniana_ligamentina
+--+
| +-------- Lampsilis_cardium
+--+
| +----- Cambarunio_iris
+--+
| +-- Ligumia_recta
+--+
+-- Epioblasma_triquetra
Hoeh et al., 2009 [Publ 2716], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
23 |
20 |
3 |
42 |
20 |
3 |
45.0 |
+-- Pilsbryoconcha_exilis **
+-----------------------------+
| +-- Pseudodon_vondembuschianus **
|
| +----------------------------- Lens_contradens **
--+ |
| | +-- Unio_pictorum
| | +--------+
| | | +-- Unio_caffer
| | +--------------+
+--+ | | +-------- Lasmigona_costata
| | +--+
| | | +----- Alasmidonta_marginata
| | +--+
| | | +-- Strophitus_undulatus
| | +--+
+--+ +-- Pyganodon_grandis
|
| +----------------------- Gonidea_angulata **
| |
| | +-- Eurynia_dilatata
| | |
| | +-----+-- Pleurobema_sintoxia
+--+ | |
| +-----------+ +-- Fusconaia_flava
| | |
| | | +----- Obliquaria_reflexa
| | +--+
+--+ | +-- Quadrula_verrucosa
| +--+
| +-- Quadrula_quadrula
|
| +----------------- Ptychobranchus_fasciolaris
+--+
| +-------------- Truncilla_truncata
+--+
| +----------- Ortmanniana_ligamentina
+--+
| +-------- Lampsilis_cardium
+--+
| +----- Cambarunio_iris
+--+
| +-- Ligumia_recta
+--+
+-- Epioblasma_triquetra
Hoeh et al., 2009 [Publ 2716], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
23 |
17 |
3 |
42 |
20 |
3 |
45.0 |
+-- Pilsbryoconcha_exilis **
+--------------------+
| +-- Pseudodon_vondembuschianus **
|
| +-------------------- Lens_contradens **
--+ |
| | +-- Unio_pictorum
| | +--------+
| | | +-- Unio_caffer
+--+ +-----+
| | | +-------- Lasmigona_costata
| | +--+
| | | +----- Alasmidonta_marginata
| | +--+
| | | +-- Strophitus_undulatus
+--+ +--+
| +-- Pyganodon_grandis
|
| +-------------- Gonidea_angulata **
| |
| | +----- Obliquaria_reflexa
| | +-----+
+--+ | | +-- Quadrula_verrucosa
| | +--+
| | +-- Quadrula_quadrula
| |
+--+ +-------- Amblema_plicata
| |
| | +----- Eurynia_dilatata
| +--+
| | | +-- Pleurobema_sintoxia
+--+ +--+
| +-- Fusconaia_flava
|
| +----- Truncilla_truncata
| |
| +----- Ptychobranchus_fasciolaris
+--+
| +-- Ortmanniana_ligamentina
| |
| +-- Cambarunio_iris
+--+
+-- Ligumia_recta
|
+-- Epioblasma_triquetra
Ouyang et al., 2011 [Publ 3408], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
2 |
9 |
9 |
1 |
30.0 |
+-------- Lamprotula_caveata **
+-----+
| | +----- Sinosolenaia_oleivora **
| +--+
| | +-- Sinosolenaia_carinata **
| +--+
| +-- Parvasolenaia_rivularis **
--+
| +----- Anemina_arcaeformis
| +-----+
| | | +-- Sinanodonta_elliptica
| | +--+
| | +-- Sinanodonta_woodiana
+--+
| +-- Nodularia_douglasiae
| +-----+
| | +-- Cuneopsis_heudei
+--+
| +----- Lanceolaria_grayii
| |
+--+----- Acuticosta_chinensis
|
| +-- Lanceolaria_gladiola
+--+
+-- Aculamprotula_tortuosa
Huang et al., 2013 [Publ 3629], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
3 |
12 |
11 |
2 |
29.0 |
+-------------- Unio_pictorum
|
+--+ +-- Sinanodonta_woodiana
| | +--------+
| | | +-- Cristaria_plicata
| +--+
| | +-------- Platynaias_compressa
| +--+
| | +----- Pyganodon_grandis
| +--+
| | +-- Utterbackia_imbecillis
--+ +--+
| +-- Utterbackia_peninsularis
|
| +-------- Quadrula_quadrula
| +--+
| | | +----- Toxolasma_parvum
| | +--+
| | | +-- Lampsilis_ornata
| | +--+
+-----+ +-- Venustaconcha_ellipsiformis
|
| +-- Sinosolenaia_carinata **
| +--+
| | +-- Pronodularia_japanensis **
+-----+
| +-- Sinohyriopsis_cumingii **
+--+
+-- Sinohyriopsis_schlegelii **
Sano et al., 2017 [Publ 3799], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
3 |
23 |
17 |
1 |
31.0 |
+-------- Obovalis_omiensis **
+-----------------+
| | +----- Inversidens_brandtii **
| +--+
| | +-- Pronodularia_japanensis **
| +--+
| +-- Sinohyriopsis_schlegelii **
|
| +-- Cristaria_plicata
--+ +-----------+
| | +-- Buldowskia_iwakawai
| +--------+
| | | +----------- Pletholophus_tenuis
| | | |
| | +--+ +-- Sinanodonta_tumens
| | | +-----+
| | | | +-- Sinanodonta_lauta
| | +--+
+--+ | +----- Beringiana_fukuharai
| +--+
| | +-- Beringiana_japonica
| +--+
| +-- Beringiana_beringiana
|
| +-------------------- Lanceolaria_grayii
| |
+--+ +-- Nodularia_nipponensis
| +--------------+
| | +-- Nodularia_douglasiae
+--+
| +-------------- Inversiunio_jokohamensis
+--+
| +----------- Inversiunio_yanagawensis[1]
+--+
| +-------- Inversiunio_yanagawensis[2]
+--+
| +----- Inversiunio_yanagawensis[3]
+--+
| +-- Inversiunio_yanagawensis[4]
+--+
+-- Inversiunio_reinianus
Huang et al., 2018 [Publ 3828], fig. 6-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
2 |
13 |
11 |
2 |
29.0 |
+----- Quadrula_quadrula
+-----------+
| | +-- Venustaconcha_ellipsiformis
| +--+
| +-- Potamilus_alatus
|
| +-------- Sinosolenaia_carinata **
--+ +-----+
| | | +----- Potomida_littoralis **
| | +--+
| | | +-- Lamprotula_leaii **
| | +--+
| | +-- Pronodularia_japanensis **
+--+
| +-------- Sinanodonta_woodiana
| +--+
| | | +----- Anodonta_anatina
| | +--+
| | | +-- Pyganodon_grandis
+--+ +--+
| +-- Utterbackia_peninsularis
|
| +-------- Aculamprotula_tortuosa
+--+
| +----- Unio_tumidus
+--+
| +-- Unio_crassus
+--+
+-- Unio_delphinus
Lopes-Lima et al., 2020 [Publ 4080], fig. 14
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
2 |
2 |
0 |
0 |
30.0 |
+-- Parvasolenaia_rivularis **
+--+
| +-- Koreosolenaia_sitgyensis **
--+
| +-- Lamprotula_leaii **
+--+
+-- Lamprotula_seomjinensis **
Mabuchi & Nishida, 2020 [Publ 4120], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
3 |
33 |
33 |
1 |
31.0 |
+-------- Obovalis_omiensis **
+-----------+
| | +----- Inversidens_brandtii **
| +--+
| | +-- Sinohyriopsis_schlegelii **
| +--+
| +-- Pronodularia_japanensis **
--+
| +----- Nodularia_douglasiae
| +-----------+
| | | +-- Lanceolaria_grayii
| | +--+
| | +-- Inversiunio_reinianus
| |
+--+ +-------------- Cristaria_plicata
| |
| | +----------- Sinanodonta_tumens
| | |
| | | +-- Sinanodonta_lauta[5]
| | | |
+--+ | +-- Sinanodonta_lauta[6]
| | |
| | +-- Beringiana_japonica[15]
| | |
| | +-- Beringiana_japonica[16]
| | +-----+
| | | +-- Beringiana_japonica[17]
+--+ | |
| | +-- Beringiana_japonica[18]
| | |
| | +-- Beringiana_japonica[19]
| | |
| | +-- Beringiana_japonica[20]
| |
| | +----- Beringiana_japonica[1]
| | |
| | +----- Beringiana_japonica[2]
| | |
+--+ +----- Beringiana_japonica[3]
| |
| +----- Beringiana_japonica[4]
| |
| +----- Beringiana_japonica[5]
| |
| +----- Sinanodonta_lauta[1]
| |
| +----- Beringiana_japonica[6]
| |
| +----- Sinanodonta_lauta[2]
| |
| +----- Sinanodonta_calipygos[1]
| |
+--+----- Beringiana_japonica[7]
|
+----- Sinanodonta_lauta[4]
|
+----- Beringiana_japonica[8]
|
+----- Beringiana_japonica[9]
|
+----- Beringiana_japonica[10]
|
+----- Beringiana_japonica[11]
|
+----- Beringiana_japonica[12]
|
+----- Beringiana_japonica[13]
|
+----- Beringiana_japonica[14]
|
| +-- Sinanodonta_lauta[3]
+--+
+-- Sinanodonta_calipygos[2]
Wu et al., 2020 [Publ 4121], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
2 |
30 |
7 |
3 |
29.0 |
+-- Lampsilis_cardium
+--------------+
| +-- Scabies_crispata
|
| +-------- Sinosolenaia_carinata **
--+ +-----+
| | | +----- Potomida_littoralis **
| | +--+
| | | +-- Lamprotula_caveata **
+--+ +--+
| +-- Lamprotula_leaii **
|
| +----------- Unio_pictorum
+--+
| +-------- Aculamprotula_tortuosa
+--+
| +----- Aculamprotula_polysticta
+--+
| +-- Aculamprotula_fibrosa
+--+
+-- Aculamprotula_tientsinensis
Wu et al., 2020 [Publ 4121], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
2 |
30 |
7 |
3 |
29.0 |
+-- Lampsilis_cardium
+--------------+
| +-- Scabies_crispata
|
| +-------- Sinosolenaia_carinata **
--+ +-----+
| | | +----- Potomida_littoralis **
| | +--+
| | | +-- Lamprotula_caveata **
+--+ +--+
| +-- Lamprotula_leaii **
|
| +----------- Unio_pictorum
+--+
| +-------- Aculamprotula_tortuosa
+--+
| +----- Aculamprotula_polysticta
+--+
| +-- Aculamprotula_tientsinensis
+--+
+-- Aculamprotula_fibrosa
Guerra et al., 2019 [Publ 4168], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
2 |
37 |
17 |
2 |
29.0 |
+-------- Quadrula_quadrula
+--------------+
| | +----- Toxolasma_parvum
| +--+
| | +-- Venustaconcha_ellipsiformis
| +--+
| +-- Potamilus_alatus
|
--+ +-------- Sinosolenaia_carinata **
| +-----------+
| | | +----- Pronodularia_japanensis **
| | +--+
| | | +-- Lamprotula_leaii **
| | +--+
| | +-- Potomida_littoralis **
| |
+--+ +-------------- Lanceolaria_lanceolata
| +--+
| | | +----------- Aculamprotula_tortuosa
| | +--+
| | | +-------- Schistodesmus_lampreyanus
| | +--+
| | | +----- Unio_tumidus
+--+ +--+
| | +-- Unio_crassus
| +--+
| +-- Unio_delphinus
|
| +-------------- Sinanodonta_woodiana
| |
+--+ +----------- Anodonta_anatina
| |
+--+ +-- Platynaias_compressa
| +-----+
| | +-- Platynaias_subviridis
+--+
| +----- Pyganodon_grandis
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Utterbackia_imbecillis
Guerra et al., 2019 [Publ 4168], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
2 |
18 |
13 |
2 |
29.0 |
+----- Quadrula_quadrula
+-----------------+
| | +-- Venustaconcha_ellipsiformis
| +--+
| +-- Potamilus_alatus
|
| +-------- Sinosolenaia_carinata **
--+ +-----------+
| | | +----- Potomida_littoralis **
| | +--+
| | | +-- Lamprotula_leaii **
| | +--+
| | +-- Pronodularia_japanensis **
+--+
| +-------- Sinanodonta_woodiana
| +--------+
| | | +----- Anodonta_anatina
| | +--+
| | | +-- Utterbackia_peninsularis
+--+ +--+
| +-- Pyganodon_grandis
|
| +-------------- Lanceolaria_lanceolata
+--+
| +----------- Schistodesmus_lampreyanus
+--+
| +-------- Aculamprotula_tortuosa
+--+
| +----- Unio_tumidus
+--+
| +-- Unio_crassus
+--+
+-- Unio_delphinus
Guerra et al., 2019 [Publ 4168], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
2 |
37 |
17 |
2 |
29.0 |
+-------- Quadrula_quadrula
+--------------------+
| | +----- Toxolasma_parvum
| +--+
| | +-- Venustaconcha_ellipsiformis
| +--+
| +-- Potamilus_alatus
--+
| +-------- Sinosolenaia_carinata **
| +-----------------+
| | | +----- Pronodularia_japanensis **
| | +--+
| | | +-- Lamprotula_leaii **
| | +--+
+--+ +-- Potomida_littoralis **
|
| +----------------------- Aculamprotula_tortuosa
| |
| | +-------- Schistodesmus_lampreyanus
+--+ +-----------+
| | | +----- Unio_tumidus
| | +--+
| | | +-- Unio_crassus
+--+ +--+
| +-- Unio_delphinus
|
| +----------------- Lanceolaria_lanceolata
| |
+--+ +-------------- Sinanodonta_woodiana
| |
+--+ +----------- Anodonta_anatina
| |
+--+ +-- Platynaias_compressa
| +-----+
| | +-- Platynaias_subviridis
+--+
| +----- Pyganodon_grandis
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Utterbackia_imbecillis
Guerra et al., 2019 [Publ 4168], fig. 2-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
2 |
18 |
13 |
2 |
29.0 |
+----- Quadrula_quadrula
+-----------------+
| | +-- Venustaconcha_ellipsiformis
| +--+
| +-- Potamilus_alatus
|
| +-------- Sinosolenaia_carinata **
--+ +-----------+
| | | +----- Potomida_littoralis **
| | +--+
| | | +-- Lamprotula_leaii **
| | +--+
| | +-- Pronodularia_japanensis **
+--+
| +-- Utterbackia_peninsularis
| +--+
| | +-- Pyganodon_grandis
| +-----------+
| | | +-- Anodonta_anatina
| | +--+
+--+ +-- Sinanodonta_woodiana
|
| +-------------- Lanceolaria_lanceolata
+--+
| +----------- Aculamprotula_tortuosa
+--+
| +-------- Schistodesmus_lampreyanus
+--+
| +----- Unio_tumidus
+--+
| +-- Unio_crassus
+--+
+-- Unio_delphinus
Bolotov et al., 2022 [Publ 4239], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
0 |
1 |
54 |
54 |
1 |
29.5 |
+-- Pseudodon_lenyanensis **
|
+-- Nyeinchanconcha_nyeinchani **
+-----------------------------------------------+
| +-- Indopseudodon_kayinensis **
| |
| +-- Sundadontina_tanintharyiensis **
|
| +-- Trapezidens_exolescens
| +-----+
| | +-- Trapezidens_scutum
| +-----------+
| | | +----- Trapezidens_angustior
| | +--+
| | | +-- Trapezidens_dolichorhynchus
| +--------------------------+ +--+
| | | +-- Trapezidens_yeti
| | |
--+ | | +----------------- Lamellidens_corrianus
| | | |
| | +--+ +-------------- Lamellidens_nongyangensis
| | | |
| | +--+ +----------- Lamellidens_savadiensis
| | | |
| | | | +-- Lamellidens_ferrugineus
| | +--+ +-----+
| | | | +-- Lamellidens_marginalis
| | | |
| | +--+ +-- Lamellidens_lamellatus
| | | +--+
| | | | +-- Lamellidens_jenkinsianus
| | +--+
| | | +-- Lamellidens_generosus
| | +--+
+--+ +-- Lamellidens_mainwaringi
|
| +----------- Parreysia_keralaensis
| +--+
| | | +-------- Parreysia_plagiosoma
| | +--+
| | | +----- Parreysia_nagpoorensis
| | +--+
| | | +-- Parreysia_corrugata
| +-----------------------------+ +--+
| | | +-- Parreysia_rakhinensis
| | |
| | | +-- Leoparreysia_burmana
| | | +-----+
| | | | +-- Leoparreysia_whitteni
| | +-----+
| | | +----- Leoparreysia_olivacea
| | +--+
| | | +-- Leoparreysia_canefrii
| | +--+
+--+ +-- Leoparreysia_tavoyensis
|
| +-------- Nitia_teretiuscula
| +--------------------------------+
| | | +----- Coelatura_aegyptiaca
| | +--+
| | | +-- Coelatura_kunenensis
| | +--+
| | +-- Nyassunio_nyassaensis
| |
| | +-- Indonaia_cylindrica
| | +--------+
+--+ | +-- Indonaia_gratiosa
| |
| +--------------------------+ +----- Indonaia_andersoniana
| | | +--+
| | | | | +-- Indonaia_caerulea
| | | | +--+
| | +--+ +-- Indonaia_subclathrata
| | |
| | | +----- Indonaia_shurtleffiana
| | +--+
+--+ | +-- Indonaia_bonneaudii
| +--+
| +-- Indonaia_rugosa
|
| +----------------------------------- Indochinella_pugio
| |
| | +----------- Radiatula_crispisulcata
| | |
+--+ +--------------------+ +-- Radiatula_chaudhurii
| | | +-----+
| | | | +-- Radiatula_myitkyinae
| | +--+
| | | +----- Radiatula_mouhoti
+--+ +--+
| | +-- Radiatula_chindwinensis
| +--+
| +-- Radiatula_myitthanensis
|
| +----------------------------- Scabiellus_songkramensis
+--+
| +-------------------------- Harmandia_somboriensis
+--+
| +----------------------- Unionetta_fabagina
+--+
| +-------------------- Scabies_crispata
+--+
| +----------------- Scabies_mandarinus
+--+
| +-------------- Scabies_nucleus
+--+
| +----------- Scabies_pilata
+--+
| +-------- Scabies_humilis
+--+
| +----- Scabies_anceps
+--+
| +-- Scabies_phaselus
+--+
+-- Scabies_scobinatus
Bolotov et al., 2022 [Publ 4239], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
0 |
1 |
54 |
54 |
1 |
29.5 |
+-- Pseudodon_lenyanensis **
|
+-- Nyeinchanconcha_nyeinchani **
+--------------------------------------------+
| +-- Indopseudodon_kayinensis **
| |
| +-- Sundadontina_tanintharyiensis **
|
| +-------- Trapezidens_exolescens
| |
| +--------+ +----- Trapezidens_scutum
| | | |
| | +--+ +-- Trapezidens_angustior
| | | |
| | +--+-- Trapezidens_dolichorhynchus
| | |
| +--------------------------+ +-- Trapezidens_yeti
--+ | |
| | | +----- Lamellidens_savadiensis
| | | +--------+
| | | | | +-- Lamellidens_corrianus
| | | | +--+
| | | | +-- Lamellidens_nongyangensis
| | +--+
| | | +-- Lamellidens_ferrugineus
| | | +--------+
| | | | +-- Lamellidens_marginalis
| | +--+
| | | +-------- Lamellidens_lamellatus
| | +--+
| | | +----- Lamellidens_jenkinsianus
+--+ +--+
| | +-- Lamellidens_generosus
| +--+
| +-- Lamellidens_mainwaringi
|
| +----------- Parreysia_keralaensis
| +-----------------------------+
| | | +-------- Parreysia_corrugata
| | +--+
| | | +----- Parreysia_rakhinensis
| | +--+
| | | +-- Parreysia_plagiosoma
| | +--+
| | +-- Parreysia_nagpoorensis
| |
| | +-- Leoparreysia_burmana
+--+ +-----+
| | +-- Leoparreysia_whitteni
| +-----------------------------+
| | | +----- Leoparreysia_olivacea
| | +--+
| | | +-- Leoparreysia_canefrii
| | +--+
| | +-- Leoparreysia_tavoyensis
| |
| | +-------- Nitia_teretiuscula
+--+ +--------------------------+
| | | +----- Coelatura_aegyptiaca
| | +--+
| | | +-- Coelatura_kunenensis
| | +--+
| | +-- Nyassunio_nyassaensis
| |
| | +----------------- Indonaia_cylindrica
| | |
+--+ +--------------+ +-------------- Indonaia_gratiosa
| | | |
| | +--+ +----------- Indonaia_andersoniana
| | | |
| | +--+ +-- Indonaia_caerulea
| | | +-----+
| | | | +-- Indonaia_subclathrata
| | +--+
| | | +----- Indonaia_rugosa
+--+ +--+
| | +-- Indonaia_bonneaudii
| +--+
| +-- Indonaia_shurtleffiana
|
| +----------------------------- Indochinella_pugio
| |
| | +-------- Radiatula_crispisulcata
| | |
| | +-----------------+ +-- Radiatula_chaudhurii
+--+ | | +--+
| | | | +-- Radiatula_myitkyinae
| | +--+
| | | +-- Radiatula_myitthanensis
| | | |
+--+ +--+-- Radiatula_mouhoti
| |
| +-- Radiatula_chindwinensis
|
| +----------------------- Scabiellus_songkramensis
| |
+--+ +-------------------- Harmandia_somboriensis
| |
+--+ +----------------- Unionetta_fabagina
| |
+--+ +-- Scabies_crispata
| +-----------+
| | +-- Scabies_mandarinus
+--+
| +----------- Scabies_nucleus
| |
+--+ +-- Scabies_humilis
| +-----+
| | +-- Scabies_pilata
+--+
| +----- Scabies_phaselus
+--+
| +-- Scabies_anceps
+--+
+-- Scabies_scobinatus
Bolotov et al., 2022 [Publ 4239], fig. S2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
0 |
1 |
54 |
54 |
1 |
29.5 |
+-- Pseudodon_lenyanensis **
|
+-- Nyeinchanconcha_nyeinchani **
+-----------------------------------------------+
| +-- Indopseudodon_kayinensis **
| |
| +-- Sundadontina_tanintharyiensis **
|
| +----------- Parreysia_keralaensis
| +-----------------------------------+
| | | +-------- Parreysia_corrugata
| | +--+
| | | +----- Parreysia_nagpoorensis
| | +--+
--+ | | +-- Parreysia_plagiosoma
| | +--+
| | +-- Parreysia_rakhinensis
| |
| | +----------- Trapezidens_scutum
| | +-----+
| | | | +-------- Trapezidens_exolescens
| | | +--+
| | | | +----- Trapezidens_yeti
| | | +--+
| | | | +-- Trapezidens_dolichorhynchus
+--+ | +--+
| +--------------------------+ +-- Trapezidens_angustior
| | |
| | | +----- Lamellidens_savadiensis
| | | +--------+
| | | | | +-- Lamellidens_corrianus
| | | | +--+
| | | | +-- Lamellidens_nongyangensis
| | +--+
| | | +-- Lamellidens_marginalis
| | | +--------+
| | | | +-- Lamellidens_ferrugineus
| | +--+
| | | +-------- Lamellidens_lamellatus
| | +--+
| | | +----- Lamellidens_jenkinsianus
+--+ +--+
| | +-- Lamellidens_mainwaringi
| +--+
| +-- Lamellidens_generosus
|
| +----------- Leoparreysia_burmana
| +-----------------------------+
| | | +-------- Leoparreysia_whitteni
| | +--+
| | | +----- Leoparreysia_olivacea
| | +--+
| | | +-- Leoparreysia_canefrii
| | +--+
| | +-- Leoparreysia_tavoyensis
| |
+--+ +-------- Nitia_teretiuscula
| +-----------------------------+
| | | +----- Coelatura_aegyptiaca
| | +--+
| | | +-- Coelatura_kunenensis
| | +--+
| | +-- Nyassunio_nyassaensis
| |
| | +-- Indonaia_cylindrica
| | +--------+
+--+ | +-- Indonaia_gratiosa
| |
| +-----------------------+ +----- Indonaia_shurtleffiana
| | | +--+
| | | | | +-- Indonaia_bonneaudii
| | | | +--+
| | +--+ +-- Indonaia_rugosa
| | |
| | | +----- Indonaia_andersoniana
| | +--+
+--+ | +-- Indonaia_caerulea
| +--+
| +-- Indonaia_subclathrata
|
| +-------------------------------- Indochinella_pugio
| |
| | +----------- Radiatula_crispisulcata
| | |
| | +-----------------+ +-- Radiatula_chaudhurii
+--+ | | +-----+
| | | | +-- Radiatula_myitkyinae
| | +--+
| | | +----- Radiatula_mouhoti
| | +--+
+--+ | +-- Radiatula_myitthanensis
| +--+
| +-- Radiatula_chindwinensis
|
| +-------------------------- Scabiellus_songkramensis
| |
+--+ +----------------------- Harmandia_somboriensis
| |
+--+ +-------------------- Unionetta_fabagina
| |
+--+ +----------------- Scabies_crispata
| |
+--+ +-- Scabies_humilis
| +-----------+
| | +-- Scabies_mandarinus
+--+
| +----------- Scabies_nucleus
+--+
| +-------- Scabies_pilata
+--+
| +----- Scabies_anceps
+--+
| +-- Scabies_phaselus
+--+
+-- Scabies_scobinatus
Bolotov et al., 2022 [Publ 4252], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
0 |
1 |
57 |
57 |
1 |
29.5 |
+-- Pseudodon_lenyanensis **
|
+-- Nyeinchanconcha_nyeinchani **
+--------------------------------------------+
| +-- Indopseudodon_kayinensis **
| |
| +-- Sundadontina_tanintharyiensis **
|
| +----------- Trapezidens_mogaungensis
| |
| +-----+ +-------- Trapezidens_exolescens
| | | |
| | +--+ +----- Trapezidens_scutum
| | | |
| | +--+ +-- Trapezidens_angustior
| | | |
| +--------------------------+ +--+-- Trapezidens_dolichorhynchus
| | | |
--+ | | +-- Trapezidens_yeti
| | |
| | | +-------------- Lamellidens_marginalis
| | | |
| | | | +----------- Lamellidens_ferrugineus
| | +--+ |
| | | | +-------- Lamellidens_generosus
| | | +--+
| | | | | +----- Lamellidens_mainwaringi
| | +--+ +--+
| | | | +-- Lamellidens_lamellatus
| | | +--+
| | | +-- Lamellidens_jenkinsianus
| | |
| | | +-------- Lamellidens_nongyangensis
+--+ +--+
| | +----- Lamellidens_savadiensis
| +--+
| | +-- Lamellidens_corrianus
| +--+
| +-- Lamellidens_chindwinensis
|
| +----------- Parreysia_keralaensis
| +-----------------------------+
| | | +-------- Parreysia_corrugata
| | +--+
| | | +----- Parreysia_rakhinensis
| | +--+
| | | +-- Parreysia_nagpoorensis
| | +--+
| | +-- Parreysia_plagiosoma
| |
| | +-- Leoparreysia_burmana
+--+ +-----+
| | +-- Leoparreysia_whitteni
| +-----------------------------+
| | | +----- Leoparreysia_olivacea
| | +--+
| | | +-- Leoparreysia_canefrii
| | +--+
| | +-- Leoparreysia_tavoyensis
| |
| | +-------- Nitia_teretiuscula
+--+ +--------------------------+
| | | +----- Coelatura_aegyptiaca
| | +--+
| | | +-- Nyassunio_nyassaensis
| | +--+
| | +-- Coelatura_kunenensis
| |
| | +----------------- Indonaia_cylindrica
| | |
+--+ +--------------+ +-------------- Indonaia_gratiosa
| | | |
| | +--+ +----------- Indonaia_andersoniana
| | | |
| | +--+ +-- Indonaia_caerulea
| | | +-----+
| | | | +-- Indonaia_subclathrata
| | +--+
| | | +----- Indonaia_rugosa
+--+ +--+
| | +-- Indonaia_bonneaudii
| +--+
| +-- Indonaia_shurtleffiana
|
| +----------------------------- Indochinella_pugio
| |
| | +----------- Radiatula_crispisulcata
| | |
| | | +----- Radiatula_chaudhurii
+--+ +--------------+ +--+
| | | | | +-- Radiatula_myitkyinae
| | | | +--+
| | +--+ +-- Radiatula_kachinensis
| | |
| | | +----- Radiatula_chindwinensis
+--+ +--+
| | +-- Radiatula_myitthanensis
| +--+
| +-- Radiatula_mouhoti
|
| +----------------------- Scabiellus_songkramensis
| |
+--+ +-------------------- Harmandia_somboriensis
| |
+--+ +----------------- Unionetta_fabagina
| |
+--+ +-- Scabies_crispata
| +-----------+
| | +-- Scabies_mandarinus
+--+
| +----------- Scabies_nucleus
| |
+--+ +-- Scabies_humilis
| +-----+
| | +-- Scabies_pilata
+--+
| +----- Scabies_phaselus
+--+
| +-- Scabies_anceps
+--+
+-- Scabies_scobinatus
Bolotov et al., 2022 [Publ 4252], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
0 |
1 |
57 |
57 |
1 |
29.5 |
+-- Pseudodon_lenyanensis **
|
+-- Nyeinchanconcha_nyeinchani **
+-----------------------------------------------+
| +-- Indopseudodon_kayinensis **
| |
| +-- Sundadontina_tanintharyiensis **
|
| +-- Parreysia_keralaensis
| |
| +-- Parreysia_corrugata
| |
| +--------------------------------------------+-- Parreysia_plagiosoma
| | |
| | +-- Parreysia_rakhinensis
--+ | |
| | +-- Parreysia_nagpoorensis
| |
| | +-------------- Trapezidens_mogaungensis
| | +--+
| | | | +----------- Trapezidens_scutum
| | | +--+
| | | | +-------- Trapezidens_exolescens
| | | +--+
| | | | +----- Trapezidens_yeti
| | | +--+
| | | | +-- Trapezidens_dolichorhynchus
| | +--------------------------+ +--+
+--+ | | +-- Trapezidens_angustior
| | |
| | | +-------- Lamellidens_savadiensis
| | | +-----+
| | | | | +----- Lamellidens_chindwinensis
| | | | +--+
| | | | | +-- Lamellidens_corrianus
| | | | +--+
| | +--+ +-- Lamellidens_nongyangensis
| | |
| | | +-- Lamellidens_marginalis
| | | +--------+
| | | | +-- Lamellidens_ferrugineus
| | +--+
| | | +-------- Lamellidens_generosus
| | +--+
| | | +----- Lamellidens_mainwaringi
+--+ +--+
| | +-- Lamellidens_jenkinsianus
| +--+
| +-- Lamellidens_lamellatus
|
| +-- Leoparreysia_burmana
| |
| +-- Leoparreysia_canefrii
| |
| +--------------------------------------+-- Leoparreysia_olivacea
| | |
| | +-- Leoparreysia_tavoyensis
| | |
| | +-- Leoparreysia_whitteni
| |
| | +-- Coelatura_aegyptiaca
| | |
| | +-- Coelatura_kunenensis
+--+ +-----------------------------------+
| | +-- Nitia_teretiuscula
| | |
| | +-- Nyassunio_nyassaensis
| |
| | +-- Indonaia_cylindrica
| | +--------+
| | | +-- Indonaia_gratiosa
| | |
+--+ +-----------------------+ +----- Indonaia_shurtleffiana
| | | +--+
| | | | | +-- Indonaia_bonneaudii
| | | | +--+
| | +--+ +-- Indonaia_rugosa
| | |
| | | +----- Indonaia_andersoniana
| | +--+
| | | +-- Indonaia_caerulea
+--+ +--+
| +-- Indonaia_subclathrata
|
| +-------------------------------- Indochinella_pugio
| |
| | +----- Radiatula_chaudhurii
| | +-----+
| | | | +-- Radiatula_kachinensis
| | | +--+
+--+ +-----------------+ +-- Radiatula_myitkyinae
| | |
| | | +-------- Radiatula_crispisulcata
| | +--+
| | | +----- Radiatula_mouhoti
| | +--+
+--+ | +-- Radiatula_myitthanensis
| +--+
| +-- Radiatula_chindwinensis
|
| +-------------------------- Scabiellus_songkramensis
| |
+--+ +----------------------- Harmandia_somboriensis
| |
+--+ +-------------------- Unionetta_fabagina
| |
+--+ +----------------- Scabies_crispata
| |
+--+ +-- Scabies_humilis
| +-----------+
| | +-- Scabies_mandarinus
+--+
| +----------- Scabies_nucleus
+--+
| +-------- Scabies_pilata
+--+
| +----- Scabies_anceps
+--+
| +-- Scabies_phaselus
+--+
+-- Scabies_scobinatus
Zhou et al., 2007 [Publ 2658], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
4 |
2 |
1 |
18 |
17 |
2 |
31.0 |
+----- Acuticosta_chinensis
+--------+
| | +-- Nodularia_douglasiae
| +--+
+-----+ +-- Cuneopsis_celtiformis
| |
| | +----------- Aculamprotula_tortuosa
| +--+
| | +-------- Aculamprotula_scripta
| +--+
| | +----- Aculamprotula_fibrosa
| +--+
| | +-- Aculamprotula_zonata
--+ +--+
| +-- Aculamprotula_tientsinensis
|
| +-------- Lamprotula_leaii **
| +--------+
| | | +----- Lamprotula_cornuumlunae **
| | +--+
| | | +-- Gibbosula_rochechouartii
| | +--+
+--+ +-- Lamprotula_caveata **
|
| +-- Theliderma_intermedia
| +-----------+
| | +-- Theliderma_sparsa
+--+
| +----------- Quadrula_quadrula
+--+
| +-------- Pustulosa_infucata
+--+
| +----- Quadrula_verrucosa
+--+
| +-- Pustulosa_kieneriana
+--+
+-- Pustulosa_succissa
Walker et al., 2006 [Publ 2666], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
10 |
8 |
3 |
12 |
9 |
2 |
33.0 |
+----- Anodonta
+-----------------+
| | +-- Pyganodon_fragilis
| +--+
| +-- Pyganodon_grandis
--+
| +-------------------- Pronodularia_japanensis **
| |
| | +-- Gonidea_angulata **
+--+ +--------------+
| | +-- Pseudodon_vondembuschianus **
| |
+--+ +-- Fusconaia_flava
| +-----------+
| | +-- Cumberlandia_monodonta
+--+
| +----------- Cyrtonaias_tampicoensis
+--+
| +-------- Potamilus_purpuratus
+--+
| +----- Ligumia_recta
+--+
| +-- Ortmanniana_ligamentina
+--+
+-- Lampsilis_teres
Walker et al., 2006 [Publ 2666], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
12 |
10 |
2 |
12 |
9 |
2 |
36.0 |
+-------------------------- Pseudodon_vondembuschianus **
|
--+ +----------------------- Gonidea_angulata **
| |
+--+ +-------------------- Inversidens **
| |
| | +----- Anodonta
+--+ +-----------+
| | | +-- Pyganodon_fragilis
| | +--+
+--+ +-- Pyganodon_grandis
|
| +-------------- Fusconaia_flava
+--+
| +----------- Potamilus_purpuratus
+--+
| +-------- Cyrtonaias_tampicoensis
+--+
| +----- Ligumia_recta
+--+
| +-- Ortmanniana_ligamentina
+--+
+-- Lampsilis_teres
Walker et al., 2006 [Publ 2666], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
12 |
10 |
2 |
12 |
9 |
2 |
36.0 |
+-------------------------- Pseudodon_vondembuschianus **
|
--+ +----------------------- Gonidea_angulata **
| |
+--+ +-------------------- Inversidens **
| |
| | +----- Anodonta
+--+ +-----------+
| | | +-- Pyganodon_fragilis
| | +--+
+--+ +-- Pyganodon_grandis
|
| +-------------- Fusconaia_flava
+--+
| +----------- Cyrtonaias_tampicoensis
+--+
| +-------- Potamilus_purpuratus
+--+
| +----- Ligumia_recta
+--+
| +-- Ortmanniana_ligamentina
+--+
+-- Lampsilis_teres
Shirai et al., 2010 [Publ 3353], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
0 |
0 |
0 |
29.0 |
+----- Pronodularia_japanensis **
--+
| +-- Sinohyriopsis_cumingii **
+--+
+-- Sinohyriopsis_schlegelii **
Froufe et al., 2015 [Publ 3664], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
5 |
5 |
2 |
28.0 |
+----- Anodonta_anatina
+-----+
| | +-- Utterbackia_peninsularis
| +--+
| +-- Pyganodon_grandis
--+
| +-- Venustaconcha_ellipsiformis
| +-----+
| | +-- Quadrula_quadrula
+--+
| +----- Sinosolenaia_carinata **
+--+
| +-- Potomida_littoralis **
+--+
+-- Pronodularia_japanensis **
Plazzi et al., 2011 [Publ 3676], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
4 |
1 |
2 |
15 |
14 |
2 |
30.0 |
+-------------- Lanceolaria_grayii
|
| +-- Pyganodon_grandis
+-----------+
| +-- Pseudanodonta_complanata
|
| +-- Anodonta_anatina
+-----------+
| +-- Anodonta_cygnea
|
| +----- Unio_tumidus
--+--------+
| | +-- Unio_crassus
| +--+
| +-- Unio_pictorum
|
| +----- Alathyria_jacksoni
| |
| +-----+----- Pronodularia_japanensis **
| | |
| | | +-- Sinohyriopsis_cumingii **
| | +--+
+--+ +-- Sinohyriopsis_schlegelii **
|
| +-------- Quadrula_quadrula
| |
+--+-------- Parvaspina_collina
|
| +----- Lampsilis_ornata
+--+
| +-- Epioblasma_rangiana
+--+
+-- Venustaconcha_ellipsiformis
Sheng et al., 2014 [Publ 3704], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
5 |
5 |
2 |
29.0 |
+----------- Lampsilis_ornata
|
--+ +-- Venustaconcha_ellipsiformis
| +-----+
| | +-- Quadrula_quadrula
+--+
| +----- Lamprotula_leaii **
+--+
| +-- Sinohyriopsis_cumingii **
+--+
+-- Sinohyriopsis_schlegelii **
Fonseca et al., 2016 [Publ 3775], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
6 |
6 |
2 |
28.0 |
+-- Venustaconcha_ellipsiformis
+-----------+
| +-- Quadrula_quadrula
|
--+ +----- Sinosolenaia_carinata **
| +-----+
| | | +-- Potomida_littoralis **
| | +--+
+--+ +-- Pronodularia_japanensis **
|
| +-------- Unio_delphinus
+--+
| +----- Anodonta_anatina
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Pyganodon_grandis
Lopes-Lima et al., 2017 [Publ 3792], fig. 2-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
7 |
6 |
2 |
28.0 |
+-- Venustaconcha_ellipsiformis
+-----------+
| +-- Quadrula_quadrula
|
--+ +----- Sinosolenaia_carinata **
| +-----+
| | | +-- Potomida_littoralis **
| | +--+
+--+ +-- Pronodularia_japanensis **
|
| +-------- Unio_delphinus
+--+
| +----- Anodonta_anatina
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Pyganodon_grandis
Kano et al., 2019 [Publ 4194], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
2 |
7 |
6 |
1 |
29.0 |
+----- Sinanodonta_lauta
+-----+
| | +-- Pletholophus_tenuis
| +--+
| +-- Beringiana_japonica
--+
| +----- Inversidens_brandtii **
| +--+
| | | +-- Obovalis_omiensis **
| | +--+
+--+ +-- Pronodularia_japanensis **
|
| +----- Lanceolaria_grayii
+--+
| +-- Inversiunio_yanagawensis
+--+
+-- Nodularia_nipponensis
Smith, 2021 [Publ 4331], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
3 |
17 |
13 |
3 |
27.0 |
+----- Sinohyriopsis_cumingii **
+-----+
| | +-- Lens_contradens **
| +--+
+--------+ +-- Lamprotula_leaii **
| |
| | +-------- Scabies_phaselus
| +--+
| | +----- Cristaria_plicata
| +--+
| | +-- Pyganodon_grandis
--+ +--+
| +-- Utterbackia_peninsularis
|
| +-- Uniomerus_tetralasmus
| +--------------+
| | +-- Megalonaias_nervosa
| |
+--+ +-- Elliptio_crassidens
| +-----------+
| | +-- Elliptio_complanata
+--+
| +----------- Amblema_plicata
+--+
| +-------- Potamilus_streckersoni
+--+
| +----- Venustaconcha_ellipsiformis
+--+
| +-- Lampsilis_siliquoidea
+--+
+-- Lampsilis_cardium
Nagel et al., 1998 [Publ 1946], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
9 |
0 |
2 |
7 |
7 |
1 |
33.0 |
+----------------- Microcondylaea_bonellii **
|
--+ +-------------- Potomida_littoralis **
| |
| | +----- Pseudanodonta_complanata
+--+ +-----+
| | | +-- Anodonta_anatina
| | +--+
+--+ +-- Anodonta_cygnea
|
| +-------- Unio_crassus
+--+
| +----- Unio_tumidus
+--+
| +-- Unio_pictorum
+--+
+-- Unio_mancus
Nagel et al., 1998 [Publ 1946], fig. 9
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
5 |
0 |
2 |
4 |
3 |
1 |
30.0 |
+----------- Potomida **
--+
| +-------- Microcondylaea **
+--+
| +----- Unio
+--+
| +-- Anodonta
+--+
+-- Pseudanodonta
Walker et al., 2006 [Publ 2666], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
36 |
8 |
2 |
-- |
+-- Gonidea_angulata **
+-----------+
| +-- Sundadontina_tumida **
--+
| +----------- Cyrtonaias_tampicoensis
| |
+--+----------- Fusconaia_flava
|
| +-------- Potamilus_purpuratus
+--+
| +----- Lampsilis_teres
+--+
| +-- Ligumia_recta
+--+
+-- Ortmanniana_ligamentina
Walker et al., 2006 [Publ 2666], fig. 6-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
13 |
8 |
2 |
-- |
+-- Gonidea_angulata **
+-----------------+
| +-- Sundadontina_tumida **
|
--+ +-- Pyganodon_fragilis
| +--------------+
| | +-- Pyganodon_grandis
+--+
| +-------------- Fusconaia_flava
+--+
| +----------- Potamilus_purpuratus
+--+
| +-------- Cyrtonaias_tampicoensis
+--+
| +----- Ligumia_recta
+--+
| +-- Lampsilis_teres
+--+
+-- Ortmanniana_ligamentina
Zhang et al., 2004 [Publ 2735], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
12 |
12 |
1 |
-- |
+-- Lamprotula_caveata **
+-----------------------+
| +-- Sinohyriopsis_cumingii **
|
--+ +-- Sinanodonta_woodiana
| +--------------------+
| | +-- Cristaria_plicata
| |
+--+ +----- Acuticosta_chinensis
| +--------------+
| | | +-- Lanceolaria_grayii
| | +--+
+--+ +-- Lanceolaria_lanceolata
|
| +----------------- Tchangsinaia_piscicula
+--+
| +-------------- Cuneopsis_heudei
+--+
| +----------- Nodularia_douglasiae
+--+
| +-------- Pseudocuneopsis_capitata
+--+
| +----- Cuneopsis_rufescens
+--+
| +-- Schistodesmus_lampreyanus
+--+
+-- Schistodesmus_spinosus
Plazzi & Passamonti, 2010 [Publ 3354], fig. 2-3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
2 |
2 |
2 |
-- |
+----- Sinanodonta_woodiana
|
--+----- Lampsilis_ornata
|
| +-- Sinohyriopsis_cumingii **
+--+
+-- Pronodularia_japanensis **
Breton et al., 2011 [Publ 3376], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
15 |
0 |
2 |
37 |
32 |
2 |
40.0 |
+----- Quadrula_quadrula
+--------------------------+
| | +-- Cyclonaias_tuberculata
| +--+
| +-- Pustulosa_pustulosa
|
| +-------------------------- Pronodularia_japanensis **
| |
| +--+ +----------------------- Sinohyriopsis_cumingii **
| | | |
| | +--+ +-------------------- Cristaria_plicata
| | | |
--+ | | | +-- Lasmigona_complanata
| | +--+ +--------------+
| | | | +-- Lasmigona_costata
| | | |
| | +--+ +-------------- Strophitus_undulatus
| | | |
| | | | +-- Platynaias_compressa
| | | | +--+
| | +--+ | +-- Platynaias_subviridis
| | | +-----+
| | | | | +-- Alasmidonta_marginata
+--+ | | +--+
| | | +-- Alasmidonta_undulata
| +--+
| | +-- Pyganodon_lacustris
| | +-----+
| | | +-- Pyganodon_grandis
| +--+
| | +----- Utterbackia_peninsularis[2]
| +--+
| | +-- Utterbackia_peninsularis[1]
| +--+
| +-- Utterbackia_imbecillis
|
| +-------------------------- Megalonaias_nervosa
| |
+--+ +----------------------- Fusconaia_flava
| |
+--+ +-------------------- Reginaia_ebenus
| |
| | +-------- Toxolasma_paulum[2]
+--+ +--------+
| | | +----- Toxolasma_lividum
| | +--+
| | | +-- Toxolasma_parvum
+--+ +--+
| +-- Toxolasma_paulum[1]
|
| +-------------- Ellipsaria_lineolata
| |
+--+ +----------- Truncilla_macrodon
| |
| | +-- Lemiox_rimosus
+--+ +-----+
| | +-- Potamilus_metnecktayi
| |
+--+ +-- Cambarunio_iris
| +--+
| | +-- Venustaconcha_ellipsiformis
+--+
| +-- Lampsilis_powellii
| |
+--+-- Lampsilis_ornata
|
+-- Lampsilis_siliquoidea
Jiang et al., 2010 [Publ 3438], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
2 |
2 |
2 |
-- |
+----- Lampsilis_ornata
|
--+----- Cristaria_plicata
|
| +-- Pronodularia_japanensis **
+--+
+-- Sinohyriopsis_cumingii **
Chen et al., 2012 [Publ 3624], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
5 |
5 |
2 |
-- |
+----- Quadrula_quadrula
+--+
| | +-- Lampsilis_ornata
| +--+
| +-- Venustaconcha_ellipsiformis
--+
| +-- Pyganodon_grandis
| +--+
| | +-- Cristaria_plicata
+--+
| +-- Sinohyriopsis_cumingii **
+--+
+-- Lamprotula_leaii **
Huang et al., 2013 [Publ 3629], fig. 3-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
4 |
4 |
2 |
-- |
+-- Pyganodon_grandis
+-----+
| +-- Utterbackia_peninsularis
|
--+ +-- Sinosolenaia_carinata **
| +--+
| | +-- Pronodularia_japanensis **
+--+
| +-- Quadrula_quadrula
+--+
+-- Venustaconcha_ellipsiformis
Xue et al., 2016 [Publ 3659], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
14 |
14 |
2 |
-- |
+-- Sinohyriopsis_cumingii **
+--+
| +-- Lamprotula_leaii **
+-----------+
| | +-- Lampsilis_ornata
| +--+
| +-- Venustaconcha_ellipsiformis
|
| +-- Cristaria_plicata
| +-----+
--+ | +-- Platynaias_compressa
| |
| +-----+ +-- Anemina_arcaeformis
| | | +--+
| | | | +-- Anemina_euscaphys
| | +--+
| | | +-- Sinanodonta_lucida
+--+ +--+
| +-- Sinanodonta_woodiana
|
| +-- Lanceolaria_lanceolata
| +--------+
| | +-- Lanceolaria_grayii
+--+
| +-------- Aculamprotula_tortuosa
+--+
| +----- Schistodesmus_lampreyanus
+--+
| +-- Nodularia_douglasiae
+--+
+-- Unio_pictorum
Wei et al., 2016 [Publ 3665], fig. 0
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
4 |
4 |
2 |
-- |
+-- Sinohyriopsis_cumingii **
+--+
| +-- Sinohyriopsis_schlegelii **
|
--+ +-- Cristaria_plicata
| |
| +-- Lampsilis_ornata
+--+
+-- Pyganodon_grandis
|
+-- Quadrula_quadrula
Peng et al., 2012 [Publ 3689], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
1 |
1 |
1 |
-- |
+----- Cristaria_plicata
--+
| +-- Sinohyriopsis_schlegelii **
+--+
+-- Sinohyriopsis_cumingii **
Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
15 |
0 |
2 |
24 |
24 |
3 |
40.0 |
+-------------------------------- Lamellidens_corrianus
|
| +-------------------- Nodularia_douglasiae
| |
| +--------+ +----------------- Anodonta_cygnea
| | | |
| | +--+ +-------------- Unio_pictorum[2]
--+ | | |
| | +--+ +-- Unio_pictorum[1]
| | | +--------+
| | | | +-- Unio_caffer
| | +--+
| | | +-------- Anodonta
| | | |
+--+ +--+ +----- Lasmigona_costata
| | |
| +--+ +-- Strophitus_undulatus
| | |
| +--+-- Alasmidonta_marginata
| |
| +-- Pyganodon_grandis
|
| +-------------------------- Pronodularia_japanensis **
| |
| | +-- Quadrula_quadrula
+--+ +--------------------+
| | +-- Quadrula_verrucosa
| |
+--+ +-------------------- Gonidea_angulata **
| |
| | +----- Eurynia_dilatata
+--+ +-----------+
| | | +-- Cambarunio_iris
| | +--+
| | +-- Ptychobranchus_fasciolaris
+--+
| +-- Amblema_plicata
| +-----------+
| | +-- Pleurobema_sintoxia
+--+
| +----------- Truncilla_truncata
| |
+--+ +-- Lampsilis_cardium
| +-----+
| | +-- Epioblasma_triquetra
+--+
| +----- Ligumia_recta
+--+
| +-- Obliquaria_reflexa
+--+
+-- Ortmanniana_ligamentina
Wang et al., 2016 [Publ 3714], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
1 |
1 |
1 |
-- |
+----- Unio_tumidus
--+
| +-- Sinohyriopsis_schlegelii **
+--+
+-- Sinohyriopsis_cumingii **
Lopes-Lima et al., 2017 [Publ 3734], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
15 |
12 |
1 |
-- |
+-- Potomida_littoralis **
+--------------+
| +-- Microcondylaea_bonellii **
|
| +----- Anodonta_anatina
--+ +--------+
| | | +-- Pseudanodonta_complanata
| | +--+
| | +-- Anodonta_cygnea
+--+
| +-- Unio_tumidus
| +--------+
| | +-- Unio_gibbus
| |
+--+ +-- Unio_crassus
| +-----+
| | +-- Unio_tumidiformis
+--+
| +----- Unio_ravoisieri
| |
+--+ +-- Unio_mancus
| |
| +-- Unio_pictorum
+--+
+-- Unio_delphinus
|
+-- Unio_elongatulus
Guerra et al., 2017 [Publ 3772], fig. 2a-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
6 |
4 |
2 |
-- |
+-- Sinosolenaia_carinata **
+--+
| +-- Pronodularia_japanensis **
|
| +-- Utterbackia_peninsularis
--+--+
| +-- Pyganodon_grandis
|
| +-- Venustaconcha_ellipsiformis
+--+
+-- Quadrula_quadrula
Guerra et al., 2017 [Publ 3772], fig. 2b-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
6 |
4 |
2 |
-- |
+-- Sinosolenaia_carinata **
+--+
| +-- Pronodularia_japanensis **
|
| +-- Utterbackia_peninsularis
--+--+
| +-- Pyganodon_grandis
|
| +-- Venustaconcha_ellipsiformis
+--+
+-- Quadrula_quadrula
Amaro et al., 2017 [Publ 3791], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
16 |
12 |
2 |
-- |
+----- Fusconaia_flava
+-----+
| | +-- Pronodularia_japanensis **
| +--+
| +-- Sinosolenaia_carinata **
--+
| +----- Toxolasma_parvum
| +--+
| | | +-- Venustaconcha_ellipsiformis
| | +--+
+--+ +-- Lampsilis_siliquoidea
|
| +----- Quadrula_quadrula[2]
+--+
| +-- Quadrula_quadrula[1]
+--+
+-- Quadrula_nobilis
Amaro et al., 2017 [Publ 3791], fig. 2-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
8 |
8 |
2 |
-- |
+-- Fusconaia_flava
+--+
| +-- Quadrula_quadrula
+-----+
| | +-- Lampsilis_siliquoidea
| +--+
| +-- Venustaconcha_ellipsiformis
--+
| +-- Pronodularia_japanensis **
| +-----+
| | +-- Sinosolenaia_carinata **
+--+
| +----- Pyganodon_grandis
| |
+--+ +-- Lasmigona_costata
| |
+--+-- Sinanodonta_woodiana
|
+-- Utterbackia_peninsularis
Amaro et al., 2017 [Publ 3791], fig. 3A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
10 |
7 |
2 |
-- |
+-------- Platynaias_compressa
+--+
| | +----- Pyganodon_grandis
| +--+
| | +-- Utterbackia_imbecillis
| +--+
--+ +-- Utterbackia_peninsularis
|
| +-- Pronodularia_japanensis **
| +-----+
| | +-- Sinosolenaia_carinata **
+--+
| +----- Quadrula_quadrula
+--+
| +-- Toxolasma_parvum
+--+
+-- Venustaconcha_ellipsiformis
Amaro et al., 2017 [Publ 3791], fig. 3A-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
5 |
4 |
2 |
-- |
+----------- Quadrula_quadrula
|
--+ +-- Pronodularia_japanensis **
| +-----+
| | +-- Sinosolenaia_carinata **
+--+
| +----- Venustaconcha_ellipsiformis
+--+
| +-- Pyganodon_grandis
+--+
+-- Utterbackia_peninsularis
Amaro et al., 2017 [Publ 3791], fig. 3B
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
10 |
7 |
2 |
-- |
+-- Toxolasma_parvum
+-----+
| +-- Venustaconcha_ellipsiformis
--+
| +----- Quadrula_quadrula
+--+
| +-- Pronodularia_japanensis **
+--+
+-- Sinosolenaia_carinata **
Amaro et al., 2017 [Publ 3791], fig. 3B-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
5 |
4 |
2 |
-- |
+-------- Venustaconcha_ellipsiformis
|
--+ +-- Utterbackia_peninsularis
| +--+
| | +-- Pyganodon_grandis
+--+
| +-- Pronodularia_japanensis **
+--+
+-- Sinosolenaia_carinata **
Feng et al., 2016 [Publ 3800], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
13 |
11 |
2 |
-- |
+-- Sinosolenaia_carinata **
+-----+
| +-- Sinosolenaia_oleivora **
+-----+
| | +----- Quadrula_quadrula
| +--+
| | +-- Potamilus_leptodon
--+ +--+
| +-- Lampsilis_ornata
|
| +----------- Tchangsinaia_piscicula
| |
+--+ +-------- Pyganodon_grandis
| |
+--+ +-- Anemina_arcaeformis
| +--+
| | +-- Anemina_euscaphys
+--+
| +-- Sinanodonta_lucida
+--+
+-- Sinanodonta_woodiana
Froufe et al., 2017 [Publ 3802], fig. 7
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
8 |
7 |
1 |
-- |
+-- Microcondylaea_bonellii **
+-----------+
| +-- Potomida_littoralis **
|
--+ +----- Unio_mancus
| +-----+
| | | +-- Unio_elongatulus
| | +--+
+--+ +-- Unio_pictorum
|
| +-------- Sinanodonta_woodiana
+--+
| +----- Anodonta_anatina
+--+
| +-- Anodonta_exulcerata
+--+
+-- Anodonta_cygnea
Huang et al., 2018 [Publ 3828], fig. 3, 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
27 |
4 |
3 |
-- |
+-- Potomida_littoralis **
+-----+
| +-- Lamprotula_leaii **
|
--+ +-- Lampsilis_cardium
| +--+
| | +-- Scabies_crispata
+--+
| +-- Unio_pictorum
+--+
+-- Aculamprotula_tientsinensis
Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
17 |
15 |
2 |
-- |
+----- Lanceolaria_grayii
+-----------------------+
| | +-- Chamberlainia_somsakpanhai **
| +--+
| +-- Lamprotula_leaii **
--+
| +-- Uniomerus_tetralasmus
| +-----------------------+
| | +-- Quadrula_quadrula
| |
+--+ +-- Elliptio_complanata
| +--------------------+
| | +-- Pleurobema_oviforme
+--+
| +-------------------- Amblema_plicata
| |
+--+ +----------------- Popenaias_popeii
| |
+--+ +-------------- Toxolasma_parvum
| |
| | +-- Potamilus_alatus
+--+ +--------+
| | +-- Potamilus_leptodon
| |
+--+ +-- Lampsilis_cardium
| +-----+
| | +-- Lampsilis_ornata
+--+
| +----- Venustaconcha_ellipsiformis
+--+
| +-- Lampsilis_powellii
+--+
+-- Lampsilis_siliquoidea
Sonowal et al., 2021 [Publ 4221], fig. 3a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
13 |
7 |
3 |
-- |
+-- Lamprotula_leaii **
--+
+-- Potomida_littoralis **
Sonowal et al., 2021 [Publ 4221], fig. 3b
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
9 |
0 |
1 |
13 |
7 |
3 |
35.0 |
+-- Aculamprotula_tientsinensis
|
+-- Lamprotula_leaii **
|
+-- Lampsilis_cardium
|
+-- Potomida_littoralis **
|
--+-- Unio_pictorum
|
+-- Anodonta_cygnea
|
+-- Unio_ravoisieri
|
+-- Lamellidens_mainwaringi
|
+-- Elliptio_complanata
Sonowal et al., 2021 [Publ 4221], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
9 |
0 |
1 |
14 |
8 |
3 |
35.0 |
+-------------------------- Lamellidens_mainwaringi
--+
| +----------------------- Lamprotula_leaii **
+--+
| +-------------------- Potomida_littoralis **
+--+
| +----------------- Lampsilis_cardium
+--+
| +-------------- Elliptio_complanata
+--+
| +----------- Aculamprotula_tientsinensis
+--+
| +-------- Unio_pictorum
+--+
| +----- Anodonta_cygnea
+--+
| +-- Unio_ravoisieri
+--+
+-- Lamellidens_corrianus
Wu et al., 2023 [Publ 4296], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
21 |
21 |
1 |
-- |
+-- Sinosolenaia_oleivora **
+--------------------------------+
| +-- Lamprotula_leaii **
|
--+ +-------------------------------- Acuticosta_chinensis
| |
| | +----- Lanceolaria_gladiola
| | +-----------------------+
+--+ | | +-- Lanceolaria_lanceolata
| | +--+
| | +-- Lanceolaria_grayii
| |
+--+ +-------- Pseudanodonta_complanata
| +-----------------+
| | | +----- Lasmigona_costata
| | +--+
| | | +-- Strophitus_undulatus
| | +--+
+--+ +-- Pyganodon_grandis
|
| +----------------------- Unio_tumidus
| |
| | +-- Aculamprotula_scripta
+--+ +-----------------+
| | +-- Aculamprotula_tientsinensis
| |
+--+ +-- Cuneopsis_heudei
| +--------------+
| | +-- Cuneopsis_rufescens
| |
+--+ +----- Nodularia_douglasiae
| +--------+
| | | +-- Schistodesmus_lampreyanus
| | +--+
+--+ +-- Schistodesmus_spinosus
|
| +----------- Tchangsinaia_piscicula
+--+
| +-------- Inversiunio_yanagawensis
+--+
| +----- Pseudocuneopsis_capitata
+--+
| +-- Pseudocuneopsis_yangshuoensis
+--+
+-- Pseudocuneopsis_sichuanensis
Chen et al., 2023 [Publ 4302], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
38 |
36 |
3 |
-- |
+-- Pronodularia_japanensis **
+--+
| +-- Lamprotula_leaii **
+-----------------------------+
| | +-- Lampsilis_siliquoidea
| +--+
| +-- Quadrula_quadrula
|
| +-- Coelatura_aegyptiaca
--+ |
| +-----------------------------+-- Indonaia_andersoniana
| | |
| | +-- Parreysia_nagpoorensis
| |
| | +-- Anemina_arcaeformis
+--+ +--------------------------+
| | +-- Amuranodonta_kijaensis
| |
| | +----- Pletholophus_tenuis
| | +--------------------+
| | | | +-- Sinanodonta_jourdyi
+--+ | +--+
| | +-- Beringiana_beringiana
| |
| | +----------------------- Lanceolaria_gladiola
| | |
| | +----------------------- Acuticosta_chinensis
+--+ |
| +----------------------- Lepidodesma_languilati
| |
| +----------------------- Lanceolaria_oxyrhyncha
| |
| +----------------------- Cristaria_plicata
| |
| | +-- Alasmidonta_marginata
+--+ +--+
| | +-- Lasmigona_costata
+-----------------+
| | +-- Anodonta_anatina
| +--+
| +-- Pseudanodonta_complanata
|
| +-------------------- Aculamprotula_fibrosa
| |
| | +----- Middendorffinaia_mongolica
| | +-----------+
+--+ | | +-- Nodularia_douglasiae
| | +--+
| | +-- Nodularia_breviconcha
| |
| | +-- Cuneopsis_celtiformis
+--+ +-----------+
| | +-- Cuneopsis_heudei
| |
| | +-- Diaurora_aurorea
| | +--+
| | | +-- Diaurora_laeve
| +--------+
+--+ | +-- Schistodesmus_lampreyanus
| +--+
| +-- Schistodesmus_spinosus
|
| +-- Pseudocuneopsis_sichuanensis
| +--------+
| | +-- Pseudocuneopsis_capitata
| |
+--+ +-- Inversiunio_yanagawensis
| +-----+
| | +-- Inversiunio_reinianus
+--+
| +----- Tchangsinaia_piscicula
+--+
| +-- Unio_pictorum
+--+
+-- Unio_crassus
Chen et al., 2023 [Publ 4302], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
1 |
38 |
36 |
3 |
-- |
+-- Pronodularia_japanensis **
+--+
| +-- Lamprotula_leaii **
+--------------------------+
| | +-- Lampsilis_siliquoidea
| +--+
| +-- Quadrula_quadrula
|
| +----- Parreysia_nagpoorensis
--+ +-----------------------+
| | | +-- Coelatura_aegyptiaca
| | +--+
| | +-- Indonaia_andersoniana
| |
| | +-------------------------- Acuticosta_chinensis
+--+ |
| | +----------------------- Lepidodesma_languilati
| | |
| | | +-- Lanceolaria_gladiola
| | | +--------------+
| | | | +-- Lanceolaria_oxyrhyncha
| | | |
+--+ | | +-- Alasmidonta_marginata
| +-----+ +--+
| | | | +-- Lasmigona_costata
| | | +--------+
| | | | | +-- Anodonta_anatina
| | | | +--+
| | +--+ +-- Pseudanodonta_complanata
| | |
| | | +----------- Cristaria_plicata
+--+ | |
| +--+ +-- Anemina_arcaeformis
| | +-----+
| | | +-- Amuranodonta_kijaensis
| +--+
| | +----- Pletholophus_tenuis
| +--+
| | +-- Sinanodonta_jourdyi
| +--+
| +-- Beringiana_beringiana
|
| +-------------------- Aculamprotula_fibrosa
| |
| | +----- Middendorffinaia_mongolica
| | +-----------+
+--+ | | +-- Nodularia_douglasiae
| | +--+
| | +-- Nodularia_breviconcha
| |
| | +-- Cuneopsis_celtiformis
+--+ +-----------+
| | +-- Cuneopsis_heudei
| |
| | +-- Diaurora_aurorea
| | +--+
| | | +-- Diaurora_laeve
| +--------+
+--+ | +-- Schistodesmus_lampreyanus
| +--+
| +-- Schistodesmus_spinosus
|
| +-- Pseudocuneopsis_sichuanensis
| +--------+
| | +-- Pseudocuneopsis_capitata
| |
+--+ +-- Inversiunio_yanagawensis
| +-----+
| | +-- Inversiunio_reinianus
+--+
| +----- Tchangsinaia_piscicula
+--+
| +-- Unio_pictorum
+--+
+-- Unio_crassus
Pfarrer et al., 2022 [Publ 4305], fig. 2-5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
10 |
10 |
1 |
-- |
+-- Potomida_littoralis **
+--------------+
| +-- Microcondylaea_bonellii **
|
| +----------- Unio_tumidus
| +--+
--+ | | +-------- Unio_crassus
| | +--+
| | | +----- Unio_elongatulus
| | +--+
| | | +-- Unio_mancus
+--+ +--+
| +-- Unio_pictorum
|
| +-- Cristaria_plicata
| +-----+
| | +-- Sinanodonta_woodiana
+-----+
| +----- Anodonta_anatina
+--+
| +-- Anodonta_exulcerata
+--+
+-- Anodonta_cygnea
Liu et al., 2023 [Publ 4310], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
22 |
22 |
1 |
-- |
+-- Sinosolenaia_oleivora **
+--------------------------------------+
| +-- Lamprotula_leaii **
|
--+ +-------------------------------------- Acuticosta_chinensis
| |
| | +-- Lanceolaria_lanceolata
+--+ +--------------------------------+
| | +-- Lanceolaria_grayii
| |
+--+ +-------------------------------- Lanceolaria_gladiola
| |
| | +-------- Pseudanodonta_complanata
| | +--------------------+
+--+ | | +----- Lasmigona_costata
| | +--+
| | | +-- Strophitus_undulatus
| | +--+
+--+ +-- Pyganodon_grandis
|
| +-------------------------- Unio_tumidus
| |
| | +-- Aculamprotula_scripta
+--+ +--------------------+
| | +-- Aculamprotula_tientsinensis
| |
+--+ +-- Cuneopsis_heudei
| +-----------------+
| | +-- Cuneopsis_rufescens
| |
+--+ +----- Nodularia_douglasiae
| +-----------+
| | | +-- Schistodesmus_lampreyanus
| | +--+
+--+ +-- Schistodesmus_spinosus
|
| +-------------- Tchangsinaia_piscicula
+--+
| +----------- Inversiunio_yanagawensis
+--+
| +-------- Pseudocuneopsis_capitata
+--+
| +----- Pseudocuneopsis_sichuanensis
+--+
| +-- Pseudocuneopsis_wuana
+--+
+-- Pseudocuneopsis_yangshuoensis
Liu et al., 2023 [Publ 4310], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
22 |
22 |
1 |
-- |
+-- Sinosolenaia_oleivora **
+--------------------------+
| +-- Lamprotula_leaii **
|
--+ +-------------------------- Acuticosta_chinensis
| |
| | +----------------------- Lanceolaria_grayii
| | |
+--+ +----------------------- Lanceolaria_gladiola
| |
| +----------------------- Lanceolaria_lanceolata
| |
+--+ +-------- Lasmigona_costata
| +-----------+
| | | +----- Pseudanodonta_complanata
| | +--+
| | | +-- Pyganodon_grandis
| | +--+
| | +-- Strophitus_undulatus
+--+
| +----------------- Unio_tumidus
| |
| | +-- Aculamprotula_tientsinensis
| +--------------+
| | +-- Aculamprotula_scripta
| |
+--+ +-- Cuneopsis_rufescens
| +-----------+
| | +-- Cuneopsis_heudei
| |
| | +----- Nodularia_douglasiae
| +--------+
+--+ | +-- Schistodesmus_lampreyanus
| +--+
| +-- Schistodesmus_spinosus
|
| +----------- Tchangsinaia_piscicula
| |
+--+----------- Inversiunio_yanagawensis
|
| +-------- Pseudocuneopsis_capitata
+--+
| +----- Pseudocuneopsis_sichuanensis
+--+
| +-- Pseudocuneopsis_wuana
+--+
+-- Pseudocuneopsis_yangshuoensis
Araujo et al., 2009 [Publ 2589], fig. 7A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
11 |
10 |
2 |
-- |
+----------------- Amblema_plicata
|
--+ +-------------- Potomida_littoralis **
| |
+--+ +----------- Nodularia_douglasiae
| |
| | +-- Unio_gibbus
+--+ |
| +-----+-- Unio_tumidus
| | |
| | +-- Anodonta_cygnea
+--+
| +----- Unio_pictorum[1]
| |
| +----- Unio_pictorum[2]
+--+
+----- Unio_mancus
|
| +-- Unio_tumidiformis
+--+
+-- Unio_crassus
Araujo et al., 2009 [Publ 2589], fig. 7B
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
11 |
10 |
2 |
-- |
+----------------------- Potomida_littoralis **
|
--+ +-------------------- Amblema_plicata
| |
+--+ +----------------- Anodonta_cygnea
| |
+--+ +-------------- Unio_gibbus
| |
+--+ +----------- Unio_tumidus
| |
| | +-- Unio_tumidiformis
+--+ +-----+
| | +-- Unio_crassus
| |
+--+ +-- Unio_pictorum[1]
| +--+
| | +-- Unio_mancus
+--+
| +-- Unio_pictorum[2]
+--+
+-- Unio_caffer
Araujo et al., 2009 [Publ 2589], fig. 7C
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
10 |
9 |
2 |
-- |
+----------------- Potomida_littoralis **
|
--+ +-------------- Amblema_plicata
| |
+--+ +----------- Anodonta_cygnea
| |
| | +-- Unio_gibbus
+--+ +-----+
| | +-- Unio_tumidus
| |
+--+ +-- Unio_tumidiformis
| +--+
| | +-- Unio_crassus
+--+
| +-- Unio_pictorum[1]
| |
+--+-- Unio_pictorum[2]
|
+-- Unio_mancus
Bogan & Hoeh, 2000 [Publ 2650], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
33 |
17 |
3 |
-- |
-- Gonidea_angulata
Hoeh et al., 2001 [Publ 2652], fig. 14.2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
30 |
17 |
3 |
-- |
-- Gonidea_angulata
Hoeh et al., 2001 [Publ 2652], fig. 14.4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
30 |
17 |
3 |
-- |
+----------------------- Gonidea_angulata **
|
| +-------- Unio_pictorum
| |
| +-----------+-------- Unio_tumidus
--+ | |
| | | +----- Anodonta_cygnea
| | +--+
| | | +-- Strophitus_undulatus
| | +--+
+--+ +-- Pyganodon_grandis
|
| +----------------- Amblema_plicata
| |
| | +-------- Quadrula_quadrula
| | +-----+
+--+ | | +----- Eurynia_dilatata
| | +--+
| | | +-- Fusconaia_flava
| | +--+
+--+ +-- Pleurobema_clava
|
| +-- Cyrtonaias_tampicoensis
| +--------+
| | +-- Glebula_rotundata
+--+
| +-------- Toxolasma_lividum
+--+
| +----- Potamilus_alatus
+--+
| +-- Ligumia_recta
+--+
+-- Ortmanniana_ligamentina
Hoeh et al., 2002 [Publ 2653], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
30 |
17 |
3 |
-- |
+-------------- Gonidea_angulata **
+-----+
| | +----------- Unio_pictorum
| +--+
| | +-------- Unio_tumidus
| +--+
| | +----- Anodonta_cygnea
| +--+
--+ | +-- Pyganodon_grandis
| +--+
| +-- Strophitus_undulatus
|
| +-- Glebula_rotundata
| +--------------+
| | +-- Cyrtonaias_tampicoensis
| |
+--+ +-------------- Toxolasma_lividum
| |
| | +----- Potamilus_alatus
| +--------+
+--+ | +-- Ligumia_recta
| +--+
| +-- Ortmanniana_ligamentina
|
| +----------- Amblema_plicata
+--+
| +-------- Quadrula_quadrula
+--+
| +----- Eurynia_dilatata
+--+
| +-- Pleurobema_clava
+--+
+-- Fusconaia_flava
Roe & Hoeh, 2003 [Publ 2655], fig. 4.2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
30 |
17 |
3 |
-- |
+----------------------------- Gonidea_angulata **
|
--+ +-------------------------- Coelatura_aegyptiaca
| |
+--+ +----------------------- Fusconaia_flava
| |
| +----------------------- Amblema_plicata
+--+
| +-------------------- Quadrula_quadrula
| |
| | +----------------- Pleurobema_clava
+--+ |
| | +-------------- Eurynia_dilatata
| | |
+--+ | +----------- Ortmanniana_ligamentina
| +--+
| | | +-------- Glebula_rotundata
| | +--+
| | | +----- Potamilus_alatus
+--+ +--+
| | +-- Ligumia_recta
| +--+
| +-- Toxolasma_lividum
|
| +----------- Cyrtonaias_tampicoensis
| |
+--+ +-- Unio_pictorum
| +-----+
| | +-- Unio_tumidus
+--+
| +----- Pyganodon_grandis
+--+
| +-- Anodonta_cygnea
+--+
+-- Strophitus_undulatus
Roe & Hoeh, 2003 [Publ 2655], fig. 4.3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
30 |
17 |
3 |
-- |
+-- Unio_pictorum
+-----+
| +-- Unio_tumidus
+--------------+
| | +----- Anodonta_cygnea
| +--+
| | +-- Pyganodon_grandis
| +--+
--+ +-- Strophitus_undulatus
|
| +-------------------- Gonidea_angulata **
| |
| | +----------------- Amblema_plicata
+--+ |
| | +-------- Quadrula_quadrula
| | +-----+
+--+ | | +----- Eurynia_dilatata
| | +--+
| | | +-- Fusconaia_flava
| | +--+
+--+ +-- Pleurobema_clava
|
| +-- Cyrtonaias_tampicoensis
| +--------+
| | +-- Glebula_rotundata
+--+
| +-------- Toxolasma_lividum
+--+
| +----- Ortmanniana_ligamentina
+--+
| +-- Ligumia_recta
+--+
+-- Potamilus_alatus
Chapman et al., 2008 [Publ 2667], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
20 |
20 |
1 |
-- |
+-------------------------------------------- Pronodularia_japanensis **
|
| +-- Quadrula_quadrula
--+ +--------------------------------------+
| | +-- Pustulosa_pustulosa
| |
+--+ +-- Pleurobema_sintoxia
| +-----------------------------------+
| | +-- Fusconaia_flava
+--+
| +----------------------------------- Amblema_plicata
| |
+--+ +-------------------------------- Plectomerus_dombeyanus
| |
+--+ +----------------------------- Popenaias_popeii
| |
+--+ +-------------------------- Toxolasma_lividum
| |
+--+ +----------------------- Obliquaria_reflexa
| |
| | +-- Cyrtonaias_tampicoensis
+--+ +-----------------+
| | +-- Glebula_rotundata
| |
+--+ +-- Ptychobranchus_fasciolaris
| +--------------+
| | +-- Lemiox_rimosus
| |
+--+ +-- Venustaconcha_ellipsiformis
| +-----------+
| | +-- Obovaria_olivaria
+--+
| +----------- Hamiota_subangulata
+--+
| +-------- Lampsilis_ovata
+--+
| +----- Ortmanniana_ligamentina
+--+
| +-- Lampsilis_straminea
+--+
+-- Lampsilis_hydiana
Machordom et al., 2003 [Publ 2688], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
2 |
0 |
0 |
-- |
-- Potomida_littoralis
Machordom et al., 2003 [Publ 2688], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
3 |
0 |
0 |
-- |
-- Potomida_littoralis
Rosenberg et al., 1994 [Publ 2719], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
18 |
15 |
2 |
-- |
+-------- Gonidea_angulata **
|
| +-- Elliptio_complanata
| |
| +-- Fusconaia_cerina
| |
--+ +--+-- Plectomerus_dombeyanus
| | |
| | +-- Pleurobema_cordatum
| | |
| | +-- Unio_pictorum
| |
| | +-- Pyganodon_cataracta
+--+ |
| +-- Pyganodon_grandis
| |
| +-- Utterbackia_imbecillis
| |
| +-- Lampsilis_teres
| |
| +-- Lampsilis_straminea
+--+
+-- Megalonaias_nervosa
|
+-- Obliquaria_reflexa
|
+-- Theliderma_cylindrica
|
+-- Quadrula_quadrula
|
+-- Uniomerus_tetralasmus
Rosenberg et al., 1997 [Publ 2720], fig. 2A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
22 |
17 |
2 |
-- |
+----- Anodonta_anatina
|
| +-- Gonidea_angulata **
| |
| +-- Pyganodon_grandis
| |
| +-- Utterbackia_imbecillis
| |
| +-- Pyganodon_cataracta
--+ |
| +-- Lampsilis_straminea
| |
| +-- Lampsilis_teres
| |
| +-- Megalonaias_nervosa
| |
| +-- Obliquaria_reflexa
| |
+--+-- Theliderma_cylindrica
|
+-- Quadrula_quadrula
|
+-- Uniomerus_tetralasmus
|
+-- Amblema_plicata
|
+-- Elliptio_complanata
|
+-- Fusconaia_cerina
|
+-- Plectomerus_dombeyanus
|
+-- Pleurobema_cordatum
|
+-- Unio_pictorum
Rosenberg et al., 1997 [Publ 2720], fig. 2B
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
22 |
17 |
2 |
-- |
+----- Anodonta
|
| +-- Gonidea_angulata **
| |
| +-- Pyganodon_grandis
| |
| +-- Utterbackia_imbecillis
| |
| +-- Pyganodon_cataracta
--+ |
| +-- Lampsilis_straminea
| |
| +-- Lampsilis_teres
| |
| +-- Megalonaias_nervosa
| |
| +-- Obliquaria_reflexa
| |
+--+-- Theliderma_cylindrica
|
+-- Quadrula_quadrula
|
+-- Uniomerus_tetralasmus
|
+-- Amblema_plicata
|
+-- Elliptio_complanata
|
+-- Fusconaia_cerina
|
+-- Plectomerus_dombeyanus
|
+-- Aculamprotula_tortuosa
|
+-- Unio_pictorum
Soroka, 2005 [Publ 2726], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
4 |
4 |
1 |
-- |
+----------- Pronodularia_japanensis **
--+
| +-------- Unio_tumidus
+--+
| +----- Unio_crassus
+--+
| +-- Unio_caffer
+--+
+-- Unio_pictorum
Gustafson & Iwamoto, 2005 [Publ 2741], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
4 |
3 |
1 |
-- |
+-------- Gonidea_angulata **
--+
| +----- Beringiana_beringiana
+--+
| +-- Anodonta_oregonensis
+--+
+-- Anodonta_nuttalliana
Chapman et al., 2008 [Publ 2744], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
20 |
20 |
1 |
-- |
+-------------------------------------------- Pronodularia_japanensis **
|
| +-- Quadrula_quadrula
--+ +--------------------------------------+
| | +-- Pustulosa_pustulosa
| |
+--+ +-- Pleurobema_sintoxia
| +-----------------------------------+
| | +-- Fusconaia_flava
+--+
| +----------------------------------- Amblema_plicata
| |
+--+ +-------------------------------- Plectomerus_dombeyanus
| |
+--+ +----------------------------- Popenaias_popeii
| |
+--+ +-------------------------- Toxolasma_lividum
| |
+--+ +----------------------- Obliquaria_reflexa
| |
| | +-- Cyrtonaias_tampicoensis
+--+ +-----------------+
| | +-- Glebula_rotundata
| |
+--+ +-- Ptychobranchus_fasciolaris
| +--------------+
| | +-- Lemiox_rimosus
| |
+--+ +-- Venustaconcha_ellipsiformis
| +-----------+
| | +-- Obovaria_olivaria
+--+
| +----------- Hamiota_subangulata
+--+
| +-------- Lampsilis_ovata
+--+
| +----- Ortmanniana_ligamentina
+--+
| +-- Lampsilis_straminea
+--+
+-- Lampsilis_hydiana
Khalloufi et al., 2011 [Publ 2748], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
10 |
10 |
2 |
-- |
+----------------- Amblema_plicata
|
--+ +-------------- Potomida_littoralis **
| |
| | +----- Unio_tumidus
+--+ +-----+
| | | +-- Unio_gibbus
| | +--+
| | +-- Unio_durieui
+--+
| +-- Unio_tumidiformis
| +-----+
| | +-- Unio_crassus
+--+
| +----- Unio
| |
+--+----- Unio_pictorum
|
| +-- Unio_delphinus
+--+
+-- Unio_mancus
Plazzi & Passamonti, 2010 [Publ 3354], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
1 |
1 |
1 |
-- |
+-- Pronodularia_japanensis **
--+
+-- Lampsilis_ornata
Inoue et al., 2013 [Publ 3528], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
16 |
16 |
2 |
-- |
+-------------------------- Gonidea_angulata **
|
+-------------------------- Pyganodon_grandis
|
--+ +-- Eurynia_dilatata
| +--------------------+
| | +-- Quadrula_quadrula
| |
| | +-------- Paetulunio_fabalis
+--+ +-----------+
| | | +----- Amblema_plicata
| | +--+
| | | +-- Reginaia_ebenus
+--+ +--+
| +-- Reginaia_rotulata
|
| +----------------- Lampsilis_ornata
| |
+--+ +-------------- Leaunio_vanuxemensis
| |
| | +-- Villosa_villosa
+--+-----------+
| +-- Cambarunio_iris
|
| +----------- Obovaria_olivaria
+--+
| +-------- Obovaria_arkansasensis
+--+
| +----- Obovaria_retusa
+--+
| +-- Obovaria_unicolor
+--+
+-- Obovaria_subrotunda
Prié & Puillandre, 2014 [Publ 3529], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
16 |
15 |
2 |
-- |
+-------------------------- Amblema_plicata
|
| +----------------------- Potomida_littoralis **
--+ |
| | +----- Anodonta_anatina
| | +--------------+
+--+ | | +-- Pseudanodonta_complanata
| | +--+
| | +-- Anodonta_cygnea
| |
+--+ +----- Unio_tumidus
| +-----------+
| | | +-- Unio_gibbus
| | +--+
| | +-- Unio_durieui
+--+
| +----- Unio_tumidiformis
| +--------+
| | | +-- Unio_crassus
| | +--+
+--+ +-- Unio_courtillieri
|
| +----------- Unio_delphinus
+--+
| +-------- Unio_elongatulus
+--+
| +----- Unio_mancus
+--+
| +-- Unio_pictorum
+--+
+-- Unio_ravoisieri
Reis et al., 2013 [Publ 3580], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
13 |
11 |
2 |
-- |
+-- Amblema_plicata
+-----------------+
| +-- Quadrula_nobilis
|
--+ +----------------- Potomida_littoralis **
| |
| | +----- Anodonta_cygnea
+--+ +--------+
| | | +-- Anodonta_anatina
| | +--+
+--+ +-- Pseudanodonta_complanata
|
| +----------- Unio_tumidus
| |
+--+ +-- Unio_tumidiformis
| +-----+
| | +-- Unio_crassus
+--+
| +----- Unio_delphinus
+--+
| +-- Unio_mancus
+--+
+-- Unio_pictorum
Reis et al., 2013 [Publ 3580], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
15 |
13 |
2 |
-- |
+-- Amblema_plicata
+--------------------+
| +-- Theliderma_metanevra
|
--+ +-------------------- Potomida_littoralis **
| |
| | +-- Anodonta_anatina
+--+ |
| +--------------+-- Anodonta_cygnea
| | |
| | +-- Pseudanodonta_complanata
+--+
| +-------------- Unio_tumidus
| |
+--+ +-- Unio_tumidiformis
| +--------+
| | +-- Unio_crassus
+--+
| +-------- Unio
| |
+--+ +-- Unio_delphinus
| +--+
| | +-- Unio_mancus
+--+
| +-- Unio_pictorum
+--+
+-- Unio_caffer
Inoue et al., 2014 [Publ 3635], fig. 3A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
20 |
19 |
2 |
-- |
+----------------------- Quadrula_quadrula
|
| +-------------------- Gonidea_angulata **
--+ |
| | +----------- Pseudodontoideus_subvexus
| | +-----+
+--+ | | +-------- Lasmigona_costata
| | +--+
| | | +----- Lasmigona_complanata
| | +--+
| | | +-- Arcidens_confragosus
+--+ +--+
| +-- Arcidens_wheeleri
|
| +-------------- Strophitus_undulatus
| |
| | +-------- Pyganodon_grandis
| | +--+
+--+ | | +----- Utterbackia_peninsularis[2]
| | +--+
| | | +-- Utterbackia_imbecillis
| | +--+
+--+ +-- Utterbackia_peninsularis[1]
|
| +-- Platynaias_compressa
| +-----+
| | +-- Platynaias_subviridis
+--+
| +----- Alasmidonta_marginata
+--+
| +-- Alasmidonta_undulata
+--+
+-- Alasmidonta_varicosa
Xia et al., 2016 [Publ 3712], fig. 7B
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
2 |
2 |
1 |
-- |
+----- Sinanodonta_woodiana
--+
| +-- Cristaria_plicata
+--+
+-- Sinohyriopsis_cumingii **
Araujo et al., 2016 [Publ 3746], fig. 1,2b
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
12 |
0 |
0 |
-- |
-- Potomida_littoralis
Välilä et al., 2017 [Publ 3793], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
2 |
0 |
0 |
-- |
-- Potomida_littoralis
Wen et al., 2017 [Publ 3795], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
15 |
13 |
2 |
-- |
+----------- Quadrula_quadrula
|
+--+ +-------- Toxolasma_parvum
| | |
| +--+ +-- Lampsilis_ornata
| | +--+
| | | +-- Venustaconcha_ellipsiformis
| +--+
--+ | +-- Potamilus_leptodon
| +--+
| +-- Potamilus_alatus
|
| +----------- Lamprotula_leaii **
| |
| | +----- Anodonta_anatina
+--+ +--+
| | | +-- Sinanodonta_woodiana
| | +--+
| | +-- Cristaria_plicata
+--+
| +-- Aculamprotula_coreana
| +--+
| | +-- Aculamprotula_tortuosa
+--+
| +-- Lepidodesma_languilati
+--+
+-- Unio_pictorum
Lopes-Lima et al., 2018 [Publ 3829], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
27 |
2 |
2 |
-- |
+----- Potomida_littoralis **
--+
| +-- Lampsilis_cardium
+--+
+-- Unio_pictorum
Lopes-Lima et al., 2018 [Publ 3829], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
27 |
2 |
2 |
-- |
+----- Potomida_littoralis **
--+
| +-- Lampsilis_cardium
+--+
+-- Unio_pictorum
Chase et al., 2018 [Publ 3847], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
9 |
7 |
2 |
-- |
+----------- Sinosolenaia_carinata **
|
| +----- Potamilus_alatus
--+ +--+
| | | +-- Lampsilis_siliquoidea
| | +--+
| | +-- Lampsilis_powellii
+--+
| +-- Anodonta_anatina
| +--+
| | +-- Utterbackia_peninsularis
+--+
| +-- Unio_delphinus
+--+
+-- Unio_tumidus
Inoue et al., 2018 [Publ 3858], fig. 1, 3a-c
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
204 |
203 |
2 |
-- |
+-- Alasmidonta_marginata
|
+-- Prolasmidonta_heterodon
|
+-- Alasmidonta_undulata
|
+-- Beringiana_beringiana
|
+-- Anodonta_kennerlyi
|
+-- Anodonta_nuttalliana
|
+-- Anodonta_oregonensis
|
+-- Arcidens_confragosus
|
+-- Arcidens_wheeleri
|
+-- Lasmigona_complanata
|
+--------------------------------------------------+-- Platynaias_compressa
| |
| +-- Lasmigona_costata
| |
| +-- Platynaias_decorata
| |
| +-- Platynaias_subviridis
| |
| +-- Pyganodon_grandis
| |
| +-- Pyganodon_lacustris
| |
| +-- Pseudodontoideus_subvexus
| |
| +-- Strophitus_undulatus
| |
| +-- Utterbackia_imbecillis
| |
--+ +-- Utterbackia_peninsularis[1]
| |
| +-- Utterbackia_peninsularis[2]
|
| +-------------------------------------------------- Gonidea_angulata **
| |
| | +-- Reginaia_ebenus
| | +--------------------------------------------+
| | | +-- Reginaia_rotulata
| | |
| | | +-- Amblema_plicata
| | +--------------------------------------------+
| | | +-- Amblema_elliottii
| | |
| | | +----- Popenaias_popeii
| | | |
| | | | +-- Dromus_dromas
| | | | |
+--+ | | +-- Epioblasma_rangiana
| | | |
| +-----------------------------------------+ +-- Glebula_rotundata
| | | |
| | | +-- Lampsilis_ovata
| | | |
| | | +-- Lampsilis_virescens
| | | |
| | | +-- Lemiox_rimosus
| | +--+
| | +-- Obliquaria_reflexa
| | |
| | +-- Obovaria_olivaria
| | |
+--+ +-- Obovaria_unicolor
| |
| +-- Plectomerus_dombeyanus
| |
| +-- Potamilus_alatus
| |
| +-- Potamilus_capax
|
| +-- Uniomerus_declivis
| +--+
| | +-- Uniomerus_tetralasmus
| |
| | +-- Cyclonaias_tuberculata
| +--------------------------------------+ |
| | | +-- Pustulosa_pustulosa[1]
| | | |
| | | +-- Theliderma_metanevra
| | +--+
| | +-- Pustulosa_pustulosa[2]
| | |
| | +-- Quadrula_quadrula
| | |
| | +-- Quadrula_verrucosa
| |
| | +----------------------------------------- Elliptoideus_sloatianus
+--+ |
| | +-- Elliptio_lanceolata
| | +-----------------------------------+
| | | +-- Parvaspina_collina
| | |
| | | +----------------------------------- Plethobasus_cyphyus
| | | |
| | | +----------------------------------- Eurynia_dilatata
| | | |
| | | +----------------------------------- Hemistena_lata
| | | |
| | | | +-- Elliptio_complanata
+--+ | | |
| | | +-- Elliptio_hopetonensis
| | | |
| | | +-- Elliptio_producta[1]
| | | |
| | | +-- Elliptio_shepardiana
| | | |
| | | +-- Elliptio_angustata
| | | |
| | | +-- Elliptio_arca
| | | |
| | | +-- Elliptio_arctata
| | | |
+--+ | +-- Elliptio_jayensis
| | |
| | +-- Elliptio_crassidens
| | |
| +--------------------------------+-- Elliptio_dariensis
| | |
| | +-- Elliptio_fisheriana
| | |
| | +-- Elliptio_folliculata
| | |
| | +-- Elliptio_fumata
| | |
| | +-- Elliptio_icterina
| | |
| | +-- Elliptio_mcmichaeli
| | |
| | +-- Elliptio_producta[2]
| | |
| | +-- Elliptio_pullata
| | |
| | +-- Elliptio_purpurella
| | |
| | +-- Elliptio_congaraea
| |
+--+ +-- Pleuronaia_barnesiana
| |
| +-----------------------------+-- Pleuronaia_dolabelloides
| | |
| | +-- Pleuronaia_gibber
| |
| | +-------------------------- Pleurobema_plenum
| | |
| | | +----------------------- Pleurobema_stabile
| | | |
| | | | +----- Pleurobema_pyriforme
| | +--+ | +--+
| | | | | | | +-- Pleurobema_georgianum[2]
| | | | | | +--+
| | | | | +-----------+ +-- Pleurobema_athearni
| | | | | | |
| | | +--+ | | +----- Pleurobema_decisum
| | | | | | |
| | | | | +--+ +-- Pleurobema_georgianum[1]
| | | | | | |
| | | | | +--+-- Pleurobema_hanleyianum[1]
| | | | | |
| | | | | +-- Pleurobema_hanleyianum[2]
| | | | |
| | | +--+ +-------------- Pleurobema_beadleianum[2]
| | | | |
| | | | +--+ +----------- Pleurobema_strodeanum[2]
| | | | | | |
| | | | | +--+----------- Pleurobema_rubellum
| | | | | |
| | | | | | +-------- Pleurobema_beadleianum[1]
| | | | | +--+
| | | | | | +----- Pleurobema_taitianum
| | | | | +--+
| | | | | | +-- Pleurobema_perovatum
| | | +--+ +--+
| | | | +-- Pleurobema_strodeanum[1]
| | | |
| | | | +----- Pleurobema_oviforme[6]
| | | | |
| | | | +--+----- Pleurobema_clava[3]
| | | | | |
| | | | | | +-- Pleurobema_oviforme[5]
| | | | | +--+
| | | +--------+ +-- Pleurobema_clava[2]
+--+ | |
| | | +----- Pleurobema_oviforme[2]
| | | |
| | | +----- Pleurobema_oviforme[3]
| | +--+
| | +----- Pleurobema_oviforme[4]
| | |
| | | +-- Pleurobema_clava[1]
| | +--+
| | +-- Pleurobema_oviforme[1]
| |
| | +-------------------------- Pleurobema_cordatum
| | |
| | | +----- Fusconaia_subrotunda
| | | +-----------------+
| +--+ | | +-- Fusconaia_cor
| | | | +--+
| | | | +-- Fusconaia_cuneolus
| | | |
| | | | +-------------------- Fusconaia_ozarkensis
| | +--+ |
| | | | +-- Fusconaia_escambia
| | | | +--------------+
| | | | | +-- Fusconaia_burkei
| | | | |
| | | | | +----- Fusconaia_flava[27]
| | | | | |
| | +--+ | | +-- Fusconaia_iheringi
| | | | +--+
| | | | | +-- Fusconaia_mitchelli
| | | +-----------+
| | | | | +-- Fusconaia_askewi[1]
| | | | | |
| | | | | +-- Fusconaia_askewi[2]
| | | | | |
| | +--+ | +-- Fusconaia_askewi[3]
| | | +--+
| | | +-- Fusconaia_askewi[4]
| | | |
| | | +-- Fusconaia_askewi[5]
| | | |
| | | +-- Fusconaia_chunii
| | |
+--+ | +-------------- Fusconaia_masoni
| | |
| | | +----------- Fusconaia_flava[26]
| | | |
| | | | +-- Fusconaia_cerina[8]
| +--+ | +-----+
| | | | +-- Fusconaia_flava[25]
| | | |
| | | | +----- Fusconaia_flava[5]
| | | | |
| | | | +----- Fusconaia_flava[6]
| +--+ | |
| | | +----- Fusconaia_flava[7]
| | | |
| | | +----- Fusconaia_flava[8]
| | | |
| | | +----- Fusconaia_cerina[4]
| | | |
| | | +----- Fusconaia_flava[9]
| | | |
| +--+ +----- Fusconaia_flava[10]
| | |
| | +----- Fusconaia_flava[11]
| | |
| | +----- Fusconaia_flava[12]
| | |
| | +----- Fusconaia_flava[13]
| | |
| | +----- Fusconaia_flava[14]
| | |
| | +----- Fusconaia_flava[15]
| | |
| | +----- Fusconaia_flava[16]
| | |
| | +----- Fusconaia_flava[17]
| +--+
| +----- Fusconaia_flava[18]
| |
| +----- Fusconaia_flava[19]
| |
| +----- Fusconaia_flava[20]
| |
| +----- Fusconaia_flava[21]
| |
| +----- Fusconaia_flava[22]
| |
| +----- Fusconaia_flava[23]
| |
| +----- Fusconaia_flava[24]
| |
| +----- Fusconaia_cerina[5]
| |
| +----- Fusconaia_cerina[6]
| |
| +----- Fusconaia_cerina[7]
| |
| | +-- Fusconaia_flava[1]
| | |
| | +-- Fusconaia_flava[2]
| | |
| | +-- Fusconaia_flava[3]
| | |
| +--+-- Fusconaia_flava[4]
| |
| +-- Fusconaia_cerina[1]
| |
| +-- Fusconaia_cerina[2]
| |
| +-- Fusconaia_cerina[3]
|
| +----------------- Pleurobema_riddellii
| |
| | +-------------- Pleurobema_sintoxia[21]
| | |
| | | +----------- Pleurobema_sintoxia[19]
| | | |
| | | +----------- Pleurobema_sintoxia[20]
| | | |
+-----------+ | | +----- Pleurobema_sintoxia[18]
| | | |
| | | +----- Pleurobema_rubrum[19]
| | | |
| | | +----- Pleurobema_rubrum[20]
| | +-----+
| | | +----- Pleurobema_rubrum[21]
| | | |
| | | | +-- Pleurobema_rubrum[17]
+--+ | | |
| | | +-- Pleurobema_sintoxia[16]
| | +--+
| | +-- Pleurobema_sintoxia[17]
| | |
| | +-- Pleurobema_rubrum[18]
| |
| | +-------- Pleurobema[4]
| | |
| | +-------- Pleurobema[5]
| | |
| | +-------- Pleurobema_sintoxia[1]
| | |
| | +-------- Pleurobema[8]
| | |
| | +-------- Pleurobema[9]
+--+ |
| +-------- Pleurobema[10]
| |
| +-------- Pleurobema_sintoxia[3]
| |
| +-------- Pleurobema_sintoxia[4]
| |
| +-------- Pleurobema_sintoxia[5]
| |
| +-------- Pleurobema_sintoxia[6]
| |
| +-------- Pleurobema_sintoxia[7]
| |
| +-------- Pleurobema_sintoxia[8]
| |
| +-------- Pleurobema_sintoxia[9]
| |
| +-------- Pleurobema_sintoxia[10]
| |
| +-------- Pleurobema_sintoxia[11]
| |
| +-------- Pleurobema_sintoxia[12]
| |
| +-------- Pleurobema_sintoxia[13]
| |
| +-------- Pleurobema_sintoxia[14]
| |
| +-------- Pleurobema_sintoxia[15]
| |
+--+-------- Pleurobema_rubrum[4]
|
+-------- Pleurobema_rubrum[5]
|
+-------- Pleurobema_rubrum[6]
|
+-------- Pleurobema_rubrum[7]
|
+-------- Pleurobema_rubrum[8]
|
+-------- Pleurobema_rubrum[9]
|
+-------- Pleurobema_rubrum[10]
|
+-------- Pleurobema_rubrum[11]
|
+-------- Pleurobema_rubrum[12]
|
+-------- Pleurobema_rubrum[13]
|
+-------- Pleurobema_rubrum[14]
|
+-------- Pleurobema_rubrum[15]
|
+-------- Pleurobema_rubrum[16]
|
| +-- Pleurobema[6]
+-----+
| +-- Pleurobema_rubrum[1]
|
| +-- Pleurobema_sintoxia[2]
+-----+
| +-- Pleurobema_rubrum[2]
|
| +-- Pleurobema[7]
+-----+
| +-- Pleurobema_rubrum[3]
|
| +----- Pleurobema[3]
+--+
| +-- Pleurobema[1]
+--+
+-- Pleurobema[2]
Chase et al., 2018 [Publ 3861], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
10 |
8 |
2 |
-- |
+----- Quadrula_quadrula
+--------+
| | +-- Lampsilis_powellii
| +--+
--+ +-- Lampsilis_siliquoidea
|
| +----------- Lamprotula_leaii **
| |
+--+ +-- Unio_delphinus
| +-----+
| | +-- Unio_tumidus
+--+
| +----- Anodonta_anatina
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Pyganodon_grandis
Chase et al., 2018 [Publ 3861], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
10 |
8 |
2 |
-- |
+-------- Lamprotula_leaii **
+--+
| | +----- Quadrula_quadrula
| +--+
| | +-- Lampsilis_powellii
| +--+
--+ +-- Lampsilis_siliquoidea
|
| +-- Unio_delphinus
| +-----+
| | +-- Unio_tumidus
+--+
| +----- Anodonta_anatina
+--+
| +-- Utterbackia_peninsularis
+--+
+-- Pyganodon_grandis
Pfeiffer et al., 2019 [Publ 3866], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
38 |
38 |
1 |
-- |
+-------------------------------------- Gonidea_angulata **
|
| +----------- Megalonaias_nervosa
| +-----------------------+
| | | +-------- Uniomerus_declivis
| | +--+
--+ | | +----- Quadrula_verrucosa
| | +--+
| | | +-- Cyclonaias_tuberculata
| | +--+
| | +-- Quadrula_quadrula
| |
| | +-------------- Elliptoideus_sloatianus
+--+ +-----------------+
| | | +----------- Elliptio_fumata
| | +--+
| | | +-------- Pleuronaia_barnesiana
| | +--+
| | | +----- Pleurobema_riddellii
| | +--+
| | | +-- Fusconaia_flava
| | +--+
| | +-- Pleurobema_strodeanum
| |
+--+ +-- Amblema_plicata
| +--+
| | +-- Amblema_neislerii
| +-----------------------+
| | | +-- Reginaia_ebenus
| | +--+
| | +-- Reginaia_rotulata
| |
| | +----------------- Popenaias_popeii
| | |
| | +--------+ +-------------- Psoronaias_guatemalensis[2]
| | | | |
+--+ | +--+ +----------- Psoronaias_guatemalensis[1]
| | | |
| | +--+ +-------- Psoronaias_semigranosa[3]
| | | |
| | | | +----- Psoronaias_semigranosa[2]
| | +--+ |
| | | +----- Psoronaias_usumasintae[2]
| | | |
| | +--+----- Psoronaias_percompressa
+--+ |
| | +-- Psoronaias_semigranosa[1]
| | |
| +--+-- Psoronaias_profunda
| |
| +-- Psoronaias_usumasintae[1]
|
| +-- Cyrtonaias_tampicoensis
| +--------------------+
| | +-- Glebula_rotundata
| |
+--+ +-------------------- Potamilus_alatus
| |
| | +-- Truncilla_truncata
+--+ +--------------+
| | +-- Ptychobranchus_fasciolaris
| |
+--+ +-------------- Pachynaias_spheniopsis
| |
| | +-- Cyrtonaias_explicata
+--+ +--------+
| | +-- Sphenonaias_microdon
| |
+--+ +-- Lampsilis_cardium
| +-----+
| | +-- Ortmanniana_ligamentina
+--+
| +----- Cambarunio_iris
+--+
| +-- Epioblasma_triquetra
+--+
+-- Ligumia_recta
Liu et al., 2017 [Publ 3939], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
16 |
16 |
1 |
-- |
+----- Acuticosta_chinensis
--+
| +-- Anemina_arcaeformis
+--+
+-- Sinosolenaia_oleivora **
Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
9 |
2 |
1 |
-- |
+----- Potomida_littoralis **
--+
| +-- Unio_delphinus
+--+
+-- Anodonta_anatina
Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
5 |
2 |
1 |
-- |
+----- Potomida_littoralis **
--+
| +-- Unio_delphinus
+--+
+-- Anodonta_anatina
Kim et al., 2021 [Publ 4197], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
14 |
14 |
1 |
-- |
+-- Sinohyriopsis_schlegelii **
+-----------------+
| +-- Lepidodesma_languilati
|
--+ +----------------- Aculamprotula_tientsinensis
| |
| | +-------- Acuticosta_chinensis
| | +-----+
+--+ | | +----- Lanceolaria_gladiola
| | +--+
| | | +-- Lanceolaria_grayii
| | +--+
| | +-- Lanceolaria_lanceolata
+--+
| +-------- Unio_tumidus
| +--+
| | | +----- Unio_crassus
| | +--+
| | | +-- Unio_delphinus
| | +--+
+--+ +-- Unio_pictorum
|
| +-- Nodularia_douglasiae
| +--+
| | +-- Nodularia_breviconcha
+-----+
| +-- Schistodesmus_lampreyanus
+--+
+-- Cuneopsis_heudei
Zhang et al., 2013 [Publ 4214], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
3 |
2 |
1 |
-- |
-- Lamprotula_leaii
Lyubas et al., 2022 [Publ 4255], fig. 7
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
30 |
30 |
2 |
-- |
+-- Parreysia_corrugata
+-----------------------------------+
| +-- Parreysia_keralaensis
|
| +----------------------------------- Microcondylaea_bonellii **
--+ |
| | +----------- Pseudanodonta_complanata
| | +--------------------+
| | | | +-------- Anodonta_anatina
+--+ | +--+
| | | +----- Anodonta_cygnea
| | +--+
| | | +-- Anodonta_exulcerata
| | +--+
+--+ +-- Anodonta_seddoni
|
| +----------------------------- Unio_tumidus
| |
| | +-- Unio_durieui
| | +-----------------------+
| | | +-- Unio_gibbus
+--+ |
| | +-------------- Unio_damascensis
| | |
| | +-------------- Unio_sesirmensis
| | +--------+
+--+ | | +----------- Unio_ionicus
| | | |
| | +--+ +-- Unio_bruguierianus
| | | +-----+
| | | | +-- Unio_tumidiformis
| | +--+
| | | +----- Unio_crassus[2]
+--+ +--+
| | +-- Unio_carneus
| +--+
| +-- Unio_crassus[1]
|
| +----- Unio_caffer
| +--------------+
| | | +-- Unio_terminalis
| | +--+
+--+ +-- Unio_delicatus
|
| +----------------- Unio_pictorum
| |
+--+ +-------------- Unio_eucirrus
| |
+--+ +-- Unio_elongatulus
| +--------+
| | +-- Unio_mancus
+--+
| +-------- Unio_ravoisieri
| |
+--+ +-- Unio_delphinus
| +--+
| | +-- Unio_foucauldianus
+--+
| +-- Unio_hueti
+--+
+-- Unio_tigridis
Lyubas et al., 2022 [Publ 4255], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
30 |
30 |
2 |
-- |
+-- Parreysia_corrugata
+-----------------------+
| +-- Parreysia_keralaensis
|
| +----------------------- Microcondylaea_bonellii **
--+ |
| | +----------- Anodonta_anatina
| | +--------+
| | | | +-------- Pseudanodonta_complanata
| | | +--+
+--+ | | +----- Anodonta_exulcerata
| | +--+
| | | +-- Anodonta_seddoni
| | +--+
| | +-- Anodonta_cygnea
| |
+--+ +----------------- Unio_tumidus
| |
| | +-- Unio_durieui
| +--------------+
| | +-- Unio_gibbus
| |
| | +----------- Unio_sesirmensis
| | |
| | +--+ +-------- Unio_damascensis
+--+ | | |
| | | | +----- Unio_ionicus
| | +--+ |
| | | | +-- Unio_bruguierianus
| | | +--+
| | +--+ +-- Unio_tumidiformis
| | |
| | | +-- Unio_carneus
| | | |
| | +--+-- Unio_crassus[1]
+--+ |
| +-- Unio_crassus[2]
|
| +-------- Unio_elongatulus
| |
| +-------- Unio_mancus
| |
| | +-- Unio_pictorum
| +-----+
| | +-- Unio_eucirrus
| |
+-----+ +----- Unio_caffer
+--+
| | +-- Unio_terminalis
| +--+
| +-- Unio_delicatus
|
| +----- Unio_ravoisieri
| |
| | +-- Unio_delphinus
+--+--+
| +-- Unio_foucauldianus
|
| +-- Unio_hueti
+--+
+-- Unio_tigridis
Neemuchwala et al., 2023 [Publ 4292], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
26 |
26 |
1 |
-- |
+-------------------------------- Gonidea_angulata **
|
| +-------- Pleurobema_strodeanum
| +--------------------+
--+ | | +----- Amblema_plicata
| | +--+
| | | +-- Lampsilis_cardium
| | +--+
+--+ +-- Popenaias_popeii
|
| +-------------------------- Megalonaias_nervosa
| |
| | +-------- Uniomerus_carolinianus
| | +--------------+
+--+ | | +----- Uniomerus_columbensis
| | +--+
| | | +-- Uniomerus_tetralasmus
| | +--+
| | +-- Uniomerus_declivis
+--+
| +-------- Theliderma_cylindrica
| +-----------+
| | | +----- Theliderma_metanevra
| | +--+
| | | +-- Theliderma_sparsa
| | +--+
+--+ +-- Theliderma_johnsoni
|
| +----------------- Cyclonaias_tuberculata
| |
| | +-------- Quadrula_nobilis
| | +-----+
+--+ | | +----- Quadrula_verrucosa
| | +--+
| | | +-- Quadrula_fragosa
| | +--+
| | +-- Quadrula_quadrula
+--+
| +----- Pustulosa_nodulata
| +-----+
| | | +-- Pustulosa_necki
| | +--+
| | +-- Pustulosa_petrina
+--+
| +-- Pustulosa_kieneriana
| +-----+
| | +-- Pustulosa_pustulosa
+--+
| +----- Pustulosa_succissa
+--+
| +-- Pustulosa_infucata
+--+
+-- Pustulosa_kleiniana
| 
