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Page last updated
Wed 05 Jun 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Unionoida | Unionoidea | Unionidae | Unioninae | Anodontini | Alasmidontina

genus Pyganodon Crosse & Fischer, 1894: cladomics
     5 valid species

Cladomics Summary

number of species = 5

analyzed in 175 cladograms in 102 publications
maximum number of terminals in an analysis = 8, mean = 1.39

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 32
maximum number of included terminals = 8, mean = 3.12
proportion of cladograms that support basic monophyly = 0.81


Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ tree


Cyr et al., 2007 [Publ 2745], fig. 2C

iT tT bi riS oT roT roS iD
8 8 3 8 1 0 0 0.0

*** Top Monophyly Cladogram ***

        +-- Pyganodon_grandis **     
  +-----+                         
  |     +-- Pyganodon[1] **          
  |                               
--+  +----- Pyganodon[3] **          
  |  |                            
  |  |  +-- Pyganodon_cataracta[1] **
  +--+  |                         
     |  +-- Pyganodon_fragilis[1] ** 
     |  |                         
     +--+-- Pyganodon[2] **          
        |                         
        +-- Pyganodon_cataracta[2] **
        |                         
        +-- Pyganodon_fragilis[2] ** 

Whelan et al., 2023 [Publ 4287], fig. 2

iT tT bi riS oT roT roS iD
3 3 1 3 41 33 12 9.0

*** Top Sister-Group Cladogram ***

                       +----- Utterbackia_peninsularis[2]     
                    +--+                                      
                    |  |  +-- Utterbackia_peninsularis[1]     
                    |  +--+                                   
  +-----------------+     +-- Utterbackia_imbecillis          
  |                 |                                         
  |                 |  +----- Pyganodon_grandis[2] **            
  |                 +--+                                      
  |                    |  +-- Pyganodon_grandis[1] **            
  |                    +--+                                   
  |                       +-- Pyganodon_lacustris **             
  |                                                           
  |                       +-- Utterbackiana_couperiana        
  |                    +--+                                   
  |                    |  +-- Utterbackiana_heardi            
  |              +-----+                                      
  |              |     |  +-- Utterbackiana_suborbiculata     
--+              |     +--+                                   
  |              |        +-- Utterbackiana_hartfieldorum     
  |  +-----------+                                            
  |  |           |     +----- Lasmigona_costata               
  |  |           |  +--+                                      
  |  |           |  |  |  +-- Pseudodontoideus_subvexus       
  |  |           |  |  +--+                                   
  |  |           +--+     +-- Pseudodontoideus_connasaugaensis
  |  |              |                                         
  |  |              |  +----- Lasmigona_complanata            
  |  |              +--+                                      
  |  |                 |  +-- Arcidens_confragosus            
  |  |                 +--+                                   
  |  |                    +-- Arcidens_wheeleri               
  +--+                                                        
     |     +----------------- Pressodonta_viridis             
     |     |                                                  
     |  +--+              +-- Anodontoides_denigratus         
     |  |  |  +-----------+                                   
     |  |  |  |           +-- Anodontoides_ferussacianus      
     |  |  +--+                                               
     |  |     |  +----------- Strophitus_undulatus            
     |  |     +--+                                            
     |  |        |  +-------- Strophitus_pascagoulaensis[2]   
     |  |        +--+                                         
     |  |           |  +----- Strophitus_radiatus             
     +--+           +--+                                      
        |              |  +-- Strophitus_pascagoulaensis[1]   
        |              +--+                                   
        |                 +-- Strophitus_williamsi            
        |                                                     
        |              +----- Prolasmidonta_heterodon         
        |  +-----------+                                      
        |  |           |  +-- Platynaias_subviridis           
        |  |           +--+                                   
        |  |              +-- Platynaias_compressa            
        +--+                                                  
           |              +-- Alasmidonta_holstonia           
           |  +-----------+                                   
           |  |           +-- Alasminota_etowaensis           
           |  |                                               
           +--+           +-- Alasmidonta_raveneliana         
              |  +--------+                                   
              |  |        +-- Alasmidonta_marginata           
              |  |                                            
              +--+        +-- Alasmidonta_varicosa            
                 |  +-----+                                   
                 |  |     +-- Alasmidonta_uwharriensis        
                 +--+                                         
                    |  +----- Alasmidonta_undulata            
                    +--+                                      
                       |  +-- Alasmidonta_triangulata         
                       +--+                                   
                          +-- Alasmidonta_arcula              

Zanatta et al., 2007 [Publ 2698], fig. 1A

iT tT bi riS oT roT roS iD
5 5 0 5 11 9 4 8.0
     +-- Utterbackia_peggyae   
  +--+                         
  |  +-- Utterbackia_imbecillis
  |                            
--+  +-- Pyganodon_gibbosa **     
  |  |                         
  |  +-- Pyganodon_cataracta **   
  |  |                         
  +--+-- Pyganodon_fragilis **    
     |                         
     +-- Pyganodon_grandis **     
     |                         
     +-- Pyganodon_lacustris **   

Zanatta et al., 2007 [Publ 2698], fig. 1B

iT tT bi riS oT roT roS iD
5 5 3 5 11 9 4 7.0
              +-- Utterbackia_peggyae   
  +-----------+                         
  |           +-- Utterbackia_imbecillis
--+                                     
  |  +----------- Pyganodon_lacustris **   
  +--+                                  
     |  +-------- Pyganodon_grandis **     
     +--+                               
        |  +----- Pyganodon_fragilis **    
        +--+                            
           |  +-- Pyganodon_gibbosa **     
           +--+                         
              +-- Pyganodon_cataracta **   

Hoeh, 1990 [Publ 2717], fig. 2

iT tT bi riS oT roT roS iD
5 5 3 5 11 9 4 7.0
              +-- Utterbackia_imbecillis
  +-----------+                         
  |           +-- Utterbackia_peggyae   
--+                                     
  |  +----------- Pyganodon_lacustris **   
  +--+                                  
     |  +-------- Pyganodon_grandis **     
     +--+                               
        |  +----- Pyganodon_fragilis **    
        +--+                            
           |  +-- Pyganodon_cataracta **   
           +--+                         
              +-- Pyganodon_gibbosa **     

Hoeh, 1990 [Publ 2717], fig. 3

iT tT bi riS oT roT roS iD
5 16 2 5 11 9 4 18.0
  +----- Pyganodon_fragilis **         
  |                                 
  +----- Utterbackia_imbecillis     
  |                                 
  +----- Utterbackia_peggyae        
  |                                 
  +----- Anodonta_cygnea            
  |                                 
  +----- Anodonta_kennerlyi         
  |                                 
  +----- Utterbackiana_couperiana[1]
  |                                 
--+----- Utterbackiana_couperiana[2]
  |                                 
  +----- Utterbackiana_suborbiculata
  |                                 
  |  +-- Lasmigona_costata          
  +--+                              
  |  +-- Utterbackiana_implicata    
  |                                 
  |  +-- Lasmigona_complanata       
  |  |                              
  |  +-- Platynaias_compressa       
  |  |                              
  |  +-- Pyganodon_cataracta **        
  +--+                              
     +-- Pyganodon_gibbosa **          
     |                              
     +-- Pyganodon_grandis **          
     |                              
     +-- Pyganodon_lacustris **        

Hoeh, 1990 [Publ 2717], fig. 4

iT tT bi riS oT roT roS iD
5 5 3 5 11 9 4 9.0
           +-- Utterbackia_imbecillis
  +--------+                         
  |        +-- Utterbackia_peggyae   
--+                                  
  |  +-------- Pyganodon_fragilis **    
  |  |                               
  +--+     +-- Pyganodon_cataracta **   
     |  +--+                         
     |  |  +-- Pyganodon_gibbosa **     
     +--+                            
        |  +-- Pyganodon_lacustris **   
        +--+                         
           +-- Pyganodon_grandis **     

Cyr et al., 2007 [Publ 2745], fig. 2A

iT tT bi riS oT roT roS iD
5 5 2 5 1 0 0 5.0
  +-------- Pyganodon_grandis **  
  |                            
--+-------- Pyganodon[2] **       
  |                            
  |  +----- Pyganodon[1] **       
  +--+                         
     |  +-- Pyganodon_cataracta **
     +--+                      
        +-- Pyganodon_fragilis ** 

Cyr et al., 2007 [Publ 2745], fig. 2B

iT tT bi riS oT roT roS iD
5 5 3 5 1 0 0 5.0
        +-- Pyganodon_grandis **  
  +-----+                      
  |     +-- Pyganodon[1] **       
--+                            
  |  +----- Pyganodon[2] **       
  +--+                         
     |  +-- Pyganodon_cataracta **
     +--+                      
        +-- Pyganodon_fragilis ** 

Cyr et al., 2007 [Publ 2745], fig. 2D

iT tT bi riS oT roT roS iD
5 5 2 5 0 0 0 5.0
  +-------- Pyganodon_grandis **  
  |                            
--+-------- Pyganodon[2] **       
  |                            
  |  +----- Pyganodon[1] **       
  +--+                         
     |  +-- Pyganodon_cataracta **
     +--+                      
        +-- Pyganodon_fragilis ** 

Hoeh et al., 1995 [Publ 3391], fig. 2

iT tT bi riS oT roT roS iD
4 4 1 4 3 3 1 6.0
     +----- Utterbackia_peggyae     
  +--+                              
  |  |  +-- Utterbackia_imbecillis  
  |  +--+                           
--+     +-- Utterbackia_peninsularis
  |                                 
  |  +----- Pyganodon_lacustris **     
  |  |                              
  +--+----- Pyganodon_grandis **       
     |                              
     |  +-- Pyganodon_cataracta **     
     +--+                           
        +-- Pyganodon_fragilis **      

Inoue et al., 2014 [Publ 3635], fig. 3B

iT tT bi riS oT roT roS iD
4 7 2 4 4 3 2 8.0
  +-------- Pyganodon_grandis **    
  |                              
  +-------- Pyganodon_cataracta **  
  |                              
--+-------- Pyganodon_fragilis **   
  |                              
  +-------- Pyganodon **            
  |                              
  |  +----- Platynaias_subviridis
  +--+                           
     |  +-- Arcidens_confragosus 
     +--+                        
        +-- Arcidens_wheeleri    

Krebs et al., 2019 [Publ 4092], fig. 4

iT tT bi riS oT roT roS iD
3 3 1 3 0 0 0 5.0
  +----- Pyganodon_grandis **  
--+                         
  |  +-- Pyganodon_fragilis ** 
  +--+                      
     +-- Pyganodon_cataracta **

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

iT tT bi riS oT roT roS iD
2 2 0 2 28 3 3 --
        +-- Alasmidonta_marginata
  +-----+                        
  |     +-- Lasmigona_costata    
--+                              
  |  +----- Strophitus_undulatus 
  +--+                           
     |  +-- Pyganodon_grandis **    
     +--+                        
        +-- Pyganodon_fragilis **   

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

iT tT bi riS oT roT roS iD
2 2 0 2 28 3 3 --
        +-- Alasmidonta_marginata
  +-----+                        
  |     +-- Lasmigona_costata    
--+                              
  |  +----- Strophitus_undulatus 
  +--+                           
     |  +-- Pyganodon_grandis **    
     +--+                        
        +-- Pyganodon_fragilis **   

Walker et al., 2006 [Publ 2666], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 13 0 0 --
  +-- Pyganodon_fragilis **
--+                     
  +-- Pyganodon_grandis ** 

Walker et al., 2006 [Publ 2666], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 13 0 0 --
  +-- Pyganodon_fragilis **
--+                     
  +-- Pyganodon_grandis ** 

Walker et al., 2006 [Publ 2666], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 2 13 0 0 --
  +-- Pyganodon_fragilis **
--+                     
  +-- Pyganodon_grandis ** 

Walker et al., 2006 [Publ 2666], fig. 6

iT tT bi riS oT roT roS iD
2 2 0 2 36 0 0 --
  +-- Pyganodon_fragilis **
--+                     
  +-- Pyganodon_grandis ** 

Walker et al., 2006 [Publ 2666], fig. 6-M

iT tT bi riS oT roT roS iD
2 2 0 2 13 0 0 --
  +-- Pyganodon_fragilis **
--+                     
  +-- Pyganodon_grandis ** 

Hoeh et al., 2002 [Publ 2673], fig. 1-F

iT tT bi riS oT roT roS iD
2 2 0 2 7 0 0 --
  +-- Pyganodon_fragilis **
--+                     
  +-- Pyganodon_grandis ** 

Hoeh et al., 2002 [Publ 2673], fig. 1-M

iT tT bi riS oT roT roS iD
2 2 0 2 7 0 0 --
  +-- Pyganodon_fragilis **
--+                     
  +-- Pyganodon_grandis ** 

Hoeh et al., 1996 [Publ 2694], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 3 0 0 --
  +-- Pyganodon_fragilis **
--+                     
  +-- Pyganodon_grandis ** 

King et al., 1999 [Publ 2699], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 7 7 4 --
              +-- Pyganodon_grandis **      
  +-----------+                          
  |           +-- Pyganodon_fragilis **     
  |                                      
--+        +----- Lasmigona_costata      
  |  +-----+                             
  |  |     |  +-- Lasmigona_complanata   
  |  |     +--+                          
  +--+        +-- Lasmigona_alabamensis  
     |                                   
     |  +-------- Strophitus_undulatus   
     +--+                                
        |  +----- Prolasmidonta_heterodon
        +--+                             
           |  +-- Platynaias_subviridis  
           +--+                          
              +-- Platynaias_compressa   

Rosenberg et al., 1994 [Publ 2719], fig. 3

iT tT bi riS oT roT roS iD
2 10 0 2 17 1 1 11.0
  +-- Pyganodon_cataracta **   
  |                         
  +-- Pyganodon_grandis **     
  |                         
  +-- Utterbackia_imbecillis
  |                         
  +-- Lampsilis_teres       
  |                         
  +-- Lampsilis_straminea   
--+                         
  +-- Megalonaias_nervosa   
  |                         
  +-- Obliquaria_reflexa    
  |                         
  +-- Theliderma_cylindrica 
  |                         
  +-- Quadrula_quadrula     
  |                         
  +-- Uniomerus_tetralasmus 

Rosenberg et al., 1997 [Publ 2720], fig. 2A

iT tT bi riS oT roT roS iD
2 17 0 2 21 1 1 18.0
  +-- Gonidea_angulata      
  |                         
  +-- Pyganodon_grandis **     
  |                         
  +-- Utterbackia_imbecillis
  |                         
  +-- Pyganodon_cataracta **   
  |                         
  +-- Lampsilis_straminea   
  |                         
  +-- Lampsilis_teres       
  |                         
  +-- Megalonaias_nervosa   
  |                         
  +-- Obliquaria_reflexa    
  |                         
--+-- Theliderma_cylindrica 
  |                         
  +-- Quadrula_quadrula     
  |                         
  +-- Uniomerus_tetralasmus 
  |                         
  +-- Amblema_plicata       
  |                         
  +-- Elliptio_complanata   
  |                         
  +-- Fusconaia_cerina      
  |                         
  +-- Plectomerus_dombeyanus
  |                         
  +-- Pleurobema_cordatum   
  |                         
  +-- Unio_pictorum         

Rosenberg et al., 1997 [Publ 2720], fig. 2B

iT tT bi riS oT roT roS iD
2 17 0 2 21 1 1 18.0
  +-- Gonidea_angulata      
  |                         
  +-- Pyganodon_grandis **     
  |                         
  +-- Utterbackia_imbecillis
  |                         
  +-- Pyganodon_cataracta **   
  |                         
  +-- Lampsilis_straminea   
  |                         
  +-- Lampsilis_teres       
  |                         
  +-- Megalonaias_nervosa   
  |                         
  +-- Obliquaria_reflexa    
  |                         
--+-- Theliderma_cylindrica 
  |                         
  +-- Quadrula_quadrula     
  |                         
  +-- Uniomerus_tetralasmus 
  |                         
  +-- Amblema_plicata       
  |                         
  +-- Elliptio_complanata   
  |                         
  +-- Fusconaia_cerina      
  |                         
  +-- Plectomerus_dombeyanus
  |                         
  +-- Aculamprotula_tortuosa
  |                         
  +-- Unio_pictorum         

Breton et al., 2011 [Publ 3376], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 37 10 5 --
           +-- Platynaias_compressa       
        +--+                              
        |  +-- Platynaias_subviridis      
  +-----+                                 
  |     |  +-- Alasmidonta_marginata      
  |     +--+                              
  |        +-- Alasmidonta_undulata       
--+                                       
  |        +-- Pyganodon_lacustris **        
  |  +-----+                              
  |  |     +-- Pyganodon_grandis **          
  +--+                                    
     |  +----- Utterbackia_peninsularis[2]
     +--+                                 
        |  +-- Utterbackia_peninsularis[1]
        +--+                              
           +-- Utterbackia_imbecillis     

Wu et al., 2018 [Publ 3836], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 51 3 3 --
  +-------- Strophitus_undulatus 
  |                              
--+  +----- Platynaias_compressa 
  |  |                           
  +--+----- Alasmidonta_marginata
     |                           
     |  +-- Pyganodon_cataracta **  
     +--+                        
        +-- Pyganodon_grandis **    

Chase et al., 2018 [Publ 3847], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 2 37 10 5 --
              +-- Pyganodon_lacustris **        
        +-----+                              
        |     +-- Pyganodon_grandis **          
  +-----+                                    
  |     |  +----- Utterbackia_peninsularis[2]
  |     +--+                                 
  |        |  +-- Utterbackia_peninsularis[1]
  |        +--+                              
--+           +-- Utterbackia_imbecillis     
  |                                          
  |           +-- Lasmigona_costata          
  |  +--------+                              
  |  |        +-- Lasmigona_complanata       
  |  |                                       
  +--+        +-- Alasmidonta_undulata       
     |  +-----+                              
     |  |     +-- Alasmidonta_marginata      
     +--+                                    
        |  +----- Strophitus_undulatus       
        +--+                                 
           |  +-- Platynaias_compressa       
           +--+                              
              +-- Platynaias_subviridis      

Inoue et al., 2018 [Publ 3858], fig. 1, 3a-c

iT tT bi riS oT roT roS iD
2 21 0 2 195 15 8 23.0
  +-- Alasmidonta_marginata      
  |                              
  +-- Prolasmidonta_heterodon    
  |                              
  +-- Alasmidonta_undulata       
  |                              
  +-- Beringiana_beringiana      
  |                              
  +-- Anodonta_kennerlyi         
  |                              
  +-- Anodonta_nuttalliana       
  |                              
  +-- Anodonta_oregonensis       
  |                              
  +-- Arcidens_confragosus       
  |                              
  +-- Arcidens_wheeleri          
  |                              
  +-- Lasmigona_complanata       
  |                              
--+-- Platynaias_compressa       
  |                              
  +-- Lasmigona_costata          
  |                              
  +-- Platynaias_decorata        
  |                              
  +-- Platynaias_subviridis      
  |                              
  +-- Pyganodon_grandis **          
  |                              
  +-- Pyganodon_lacustris **        
  |                              
  +-- Pseudodontoideus_subvexus  
  |                              
  +-- Strophitus_undulatus       
  |                              
  +-- Utterbackia_imbecillis     
  |                              
  +-- Utterbackia_peninsularis[1]
  |                              
  +-- Utterbackia_peninsularis[2]

Whelan et al., 2023 [Publ 4287], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 2 27 25 10 --
     +-------- Strophitus_pascagoulaensis
  +--+                                   
  |  |  +----- Strophitus_undulatus      
  |  +--+                                
  |     |  +-- Strophitus_radiatus       
  |     +--+                             
--+        +-- Strophitus_williamsi      
  |                                      
  |        +-- Pyganodon_grandis **         
  |  +-----+                             
  |  |     +-- Pyganodon_cataracta **       
  +--+                                   
     |  +----- Utterbackia_peninsularis  
     +--+                                
        |  +-- Anodontoides_ferussacianus
        +--+                             
           +-- Anodontoides_denigratus   

Graf & Cummings, 2006 [Publ 1968], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 46 3 3 --
  +----- Alasmidonta_marginata
--+                           
  |  +-- Pyganodon_grandis **    
  +--+                        
     +-- Strophitus_undulatus 

Graf, 2000 [Publ 2647], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 17 0 0 --
-- Pyganodon

Graf & Ó Foighil, 2000 [Publ 2649], fig. 1, 2

iT tT bi riS oT roT roS iD
1 1 0 1 11 0 0 --
-- Pyganodon_grandis

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 33 1 1 --
  +-- Strophitus_undulatus
--+                       
  +-- Pyganodon_grandis **   

Graf, 2002 [Publ 2651], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 17 1 1 --
  +-- Alasmidonta_marginata
--+                        
  +-- Pyganodon_grandis **    

Hoeh et al., 2001 [Publ 2652], fig. 14.2

iT tT bi riS oT roT roS iD
1 1 0 1 30 1 1 --
-- Pyganodon_grandis

Hoeh et al., 2001 [Publ 2652], fig. 14.4

iT tT bi riS oT roT roS iD
1 1 0 1 30 1 1 --
  +-- Strophitus_undulatus
--+                       
  +-- Pyganodon_grandis **   

Hoeh et al., 2002 [Publ 2653], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 30 1 1 --
  +-- Pyganodon_grandis **   
--+                       
  +-- Strophitus_undulatus

Roe & Hoeh, 2003 [Publ 2655], fig. 4.2

iT tT bi riS oT roT roS iD
1 1 0 1 30 1 1 --
-- Pyganodon_grandis

Roe & Hoeh, 2003 [Publ 2655], fig. 4.3

iT tT bi riS oT roT roS iD
1 1 0 1 30 1 1 --
  +-- Pyganodon_grandis **   
--+                       
  +-- Strophitus_undulatus

Campbell et al., 2005 [Publ 2660], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 115 3 3 --
  +----- Pyganodon_grandis **        
--+                               
  |  +-- Lasmigona_costata        
  +--+                            
     +-- Pseudodontoideus_subvexus

Campbell et al., 2005 [Publ 2660], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 113 3 3 --
-- Pyganodon_grandis

Chong et al., 2008 [Publ 2668], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 20 7 6 --
-- Pyganodon_fragilis

Manendo et al., 2008 [Publ 2669], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 12 1 1 --
  +-- Platynaias_subviridis
--+                        
  +-- Pyganodon_grandis **    

Zanatta & Murphy, 2006 [Publ 2672], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 55 0 0 --
-- Pyganodon_grandis

Zanatta & Murphy, 2006 [Publ 2672], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 56 0 0 --
-- Pyganodon_grandis

Curole & Kocher, 2005 [Publ 2682], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 10 0 0 --
-- Pyganodon_grandis

Curole & Kocher, 2005 [Publ 2682], fig. 1-M

iT tT bi riS oT roT roS iD
1 1 0 1 11 1 1 --
  +-- Utterbackiana_implicata
--+                          
  +-- Pyganodon_grandis **      

Krebs et al., 2003 [Publ 2691], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 33 8 6 --
-- Pyganodon_grandis

Krebs, 2004 [Publ 2692], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 4 1 1 --
  +-- Lasmigona_costata
--+                    
  +-- Pyganodon_grandis **

Krebs, 2004 [Publ 2692], fig. 2-M

iT tT bi riS oT roT roS iD
1 1 0 1 3 1 1 --
  +-- Lasmigona_costata
--+                    
  +-- Pyganodon_grandis **

Hoeh et al., 2009 [Publ 2716], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 46 3 3 --
  +----- Alasmidonta_marginata
--+                           
  |  +-- Strophitus_undulatus 
  +--+                        
     +-- Pyganodon_grandis **    

Hoeh et al., 2009 [Publ 2716], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 45 3 3 --
  +----- Alasmidonta_marginata
--+                           
  |  +-- Strophitus_undulatus 
  +--+                        
     +-- Pyganodon_grandis **    

Hoeh et al., 2009 [Publ 2716], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 45 3 3 --
  +----- Alasmidonta_marginata
--+                           
  |  +-- Strophitus_undulatus 
  +--+                        
     +-- Pyganodon_grandis **    

Boyer et al., 2011 [Publ 2718], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 34 5 4 --
  +----- Alasmidonta_marginata
  |                           
  +----- Pyganodon_grandis **    
  |                           
--+  +-- Lasmigona_complanata 
  +--+                        
  |  +-- Lasmigona_costata    
  |                           
  |  +-- Platynaias_compressa 
  +--+                        
     +-- Strophitus_undulatus 

Boyer et al., 2011 [Publ 2718], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 37 8 7 --
  +-------- Utterbackia_imbecillis    
  |                                   
--+-------- Alasmidonta_marginata     
  |                                   
  |  +----- Pyganodon_grandis **         
  +--+                                
     |  +-- Anodontoides_ferussacianus
     +--+                             
        +-- Platynaias_compressa      

Whelan et al., 2011 [Publ 3380], fig. 1 ML

iT tT bi riS oT roT roS iD
1 1 0 1 36 1 1 --
  +-- Pyganodon_grandis **
--+                    
  +-- Lasmigona_costata

Whelan et al., 2011 [Publ 3380], fig. 1 MP

iT tT bi riS oT roT roS iD
1 1 0 1 36 1 1 --
  +-- Pyganodon_grandis **
--+                    
  +-- Lasmigona_costata

Inoue et al., 2013 [Publ 3528], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 16 0 0 --
-- Pyganodon_grandis

Zanatta & Harris, 2013 [Publ 3530], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 4 0 0 --
-- Pyganodon_grandis

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 34 2 2 --
  +----- Strophitus_undulatus
--+                          
  |  +-- Platynaias_compressa
  +--+                       
     +-- Pyganodon_grandis **   

Inoue et al., 2013 [Publ 3579], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 8 0 0 --
-- Pyganodon_grandis

Marshall et al., 2014 [Publ 3593], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 13 0 0 --
-- Pyganodon_grandis

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 39 1 1 --
  +-- Pyganodon_grandis **
--+                    
  +-- Lasmigona_costata

Chen et al., 2012 [Publ 3624], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 6 0 0 --
-- Pyganodon_grandis

Huang et al., 2013 [Publ 3629], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 15 3 2 --
  +-------- Platynaias_compressa    
--+                                 
  |  +----- Pyganodon_grandis **       
  +--+                              
     |  +-- Utterbackia_imbecillis  
     +--+                           
        +-- Utterbackia_peninsularis

Huang et al., 2013 [Publ 3629], fig. 3-M

iT tT bi riS oT roT roS iD
1 1 0 1 5 1 1 --
  +-- Pyganodon_grandis **       
--+                           
  +-- Utterbackia_peninsularis

Inoue et al., 2014 [Publ 3635], fig. 3A

iT tT bi riS oT roT roS iD
1 1 0 1 20 14 7 --
     +-------- Pyganodon_grandis **          
  +--+                                    
  |  |  +----- Utterbackia_peninsularis[2]
  |  +--+                                 
  |     |  +-- Utterbackia_imbecillis     
  |     +--+                              
--+        +-- Utterbackia_peninsularis[1]
  |                                       
  |        +-- Platynaias_compressa       
  |  +-----+                              
  |  |     +-- Platynaias_subviridis      
  +--+                                    
     |  +----- Alasmidonta_marginata      
     +--+                                 
        |  +-- Alasmidonta_undulata       
        +--+                              
           +-- Alasmidonta_varicosa       

Froufe et al., 2015 [Publ 3664], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 11 1 1 --
  +-- Utterbackia_peninsularis
--+                           
  +-- Pyganodon_grandis **       

Froufe et al., 2015 [Publ 3664], fig. 1-M

iT tT bi riS oT roT roS iD
1 1 0 1 7 1 1 --
  +-- Utterbackia_peninsularis
--+                           
  +-- Pyganodon_grandis **       

Wei et al., 2016 [Publ 3665], fig. 0

iT tT bi riS oT roT roS iD
1 1 0 1 5 0 0 --
-- Pyganodon_grandis

Plazzi et al., 2011 [Publ 3676], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 17 0 0 --
-- Pyganodon_grandis

Soroka, 2008 [Publ 3695], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 3 0 0 --
-- Pyganodon_grandis

Soroka, 2008 [Publ 3695], fig. 1-M

iT tT bi riS oT roT roS iD
1 1 0 1 3 0 0 --
-- Pyganodon_grandis

Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11

iT tT bi riS oT roT roS iD
1 1 0 1 25 3 3 --
  +----- Lasmigona_costata    
  |                           
--+  +-- Strophitus_undulatus 
  |  |                        
  +--+-- Alasmidonta_marginata
     |                        
     +-- Pyganodon_grandis **    

Sheng et al., 2014 [Publ 3704], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 7 0 0 --
-- Pyganodon_grandis

Zhou et al., 2016 [Publ 3710], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 19 3 2 --
  +-------- Platynaias_compressa    
--+                                 
  |  +----- Pyganodon_grandis **       
  +--+                              
     |  +-- Utterbackia_imbecillis  
     +--+                           
        +-- Utterbackia_peninsularis

Wu et al., 2016 [Publ 3711], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 20 2 1 --
  +----- Pyganodon_grandis **       
--+                              
  |  +-- Utterbackia_peninsularis
  +--+                           
     +-- Utterbackia_imbecillis  

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iT tT bi riS oT roT roS iD
1 1 0 1 80 4 4 --
        +-- Pyganodon_grandis **    
  +-----+                        
  |     +-- Simpsonaias_ambigua  
--+                              
  |  +----- Strophitus_undulatus 
  +--+                           
     |  +-- Alasmidonta_marginata
     +--+                        
        +-- Lasmigona_costata    

Hewitt et al., 2018 [Publ 3758], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 8 5 4 --
  +-------- Pyganodon_grandis **    
  |                              
  |     +-- Platynaias_subviridis
--+-----+                        
  |     +-- Strophitus_undulatus 
  |                              
  |  +----- Lasmigona_costata    
  +--+                           
     |  +-- Arcidens_wheeleri    
     +--+                        
        +-- Lasmigona_complanata 

Bolotov et al., 2017 [Publ 3761], fig. S1

iT tT bi riS oT roT roS iD
1 1 0 1 140 4 4 --
  +----- Simpsonaias_ambigua 
--+                          
  |  +-- Pyganodon_grandis **   
  +--+                       
     +-- Strophitus_undulatus

Bolotov et al., 2017 [Publ 3761], fig. S2

iT tT bi riS oT roT roS iD
1 1 0 1 140 4 4 --
  +----- Pyganodon_grandis **    
  |                           
  +----- Simpsonaias_ambigua  
--+                           
  +----- Strophitus_undulatus 
  |                           
  |  +-- Alasmidonta_marginata
  +--+                        
     +-- Lasmigona_costata    

Soroka & Burzynski, 2018 [Publ 3762], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 6 1 1 --
  +-- Pyganodon_grandis **       
--+                           
  +-- Utterbackia_peninsularis

Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M

iT tT bi riS oT roT roS iD
1 1 0 1 6 1 1 --
  +-- Pyganodon_grandis **       
--+                           
  +-- Utterbackia_peninsularis

Bolotov et al., 2017 [Publ 3764], fig. S1

iT tT bi riS oT roT roS iD
1 1 0 1 131 4 4 --
-- Pyganodon_grandis

Guerra et al., 2017 [Publ 3772], fig. 2a

iT tT bi riS oT roT roS iD
1 1 0 1 24 4 2 --
        +-- Platynaias_subviridis   
  +-----+                           
  |     +-- Platynaias_compressa    
--+                                 
  |  +----- Pyganodon_grandis **       
  +--+                              
     |  +-- Utterbackia_peninsularis
     +--+                           
        +-- Utterbackia_imbecillis  

Guerra et al., 2017 [Publ 3772], fig. 2a-M

iT tT bi riS oT roT roS iD
1 1 0 1 7 1 1 --
  +-- Utterbackia_peninsularis
--+                           
  +-- Pyganodon_grandis **       

Guerra et al., 2017 [Publ 3772], fig. 2b

iT tT bi riS oT roT roS iD
1 1 0 1 24 4 2 --
        +-- Platynaias_subviridis   
  +-----+                           
  |     +-- Platynaias_compressa    
--+                                 
  |  +----- Pyganodon_grandis **       
  +--+                              
     |  +-- Utterbackia_peninsularis
     +--+                           
        +-- Utterbackia_imbecillis  

Guerra et al., 2017 [Publ 3772], fig. 2b-M

iT tT bi riS oT roT roS iD
1 1 0 1 7 1 1 --
  +-- Utterbackia_peninsularis
--+                           
  +-- Pyganodon_grandis **       

Burzynski et al., 2017 [Publ 3774], fig. 1 bottom

iT tT bi riS oT roT roS iD
1 1 0 1 12 1 1 --
  +-- Utterbackia_peninsularis
--+                           
  +-- Pyganodon_grandis **       

Burzynski et al., 2017 [Publ 3774], fig. 1 top

iT tT bi riS oT roT roS iD
1 1 0 1 12 1 1 --
  +-- Utterbackia_peninsularis
--+                           
  +-- Pyganodon_grandis **       

Fonseca et al., 2016 [Publ 3775], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 31 3 2 --
  +-------- Platynaias_compressa    
--+                                 
  |  +----- Pyganodon_grandis **       
  +--+                              
     |  +-- Utterbackia_peninsularis
     +--+                           
        +-- Utterbackia_imbecillis  

Fonseca et al., 2016 [Publ 3775], fig. 1-M

iT tT bi riS oT roT roS iD
1 1 0 1 8 1 1 --
  +-- Utterbackia_peninsularis
--+                           
  +-- Pyganodon_grandis **       

Araujo et al., 2018 [Publ 3779], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 18 1 1 --
  +-- Pyganodon_grandis **    
--+                        
  +-- Alasmidonta_marginata

Smith et al., 2018 [Publ 3790], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 18 17 7 --
                 +-- Utterbackia_peninsularis        
  +--------------+                                   
  |              +-- Pyganodon_grandis **               
  |                                                  
  |              +-- Anodontoides_ferussacianus      
  |     +--------+                                   
  |     |        +-- Anodontoides_denigratus         
  |     |                                            
--+  +--+     +----- Strophitus_pascagoulaensis      
  |  |  |  +--+                                      
  |  |  |  |  |  +-- Strophitus_radiatus             
  |  |  |  |  +--+                                   
  |  |  +--+     +-- Strophitus_williamsi            
  |  |     |                                         
  |  |     |  +----- Strophitus_howellsi[2]          
  |  |     +--+                                      
  +--+        |  +-- Strophitus_undulatus            
     |        +--+                                   
     |           +-- Strophitus_howellsi[1]          
     |                                               
     |           +-- Alasmidonta_triangulata         
     |  +--------+                                   
     |  |        +-- Alasminota_etowaensis           
     |  |                                            
     +--+        +-- Pseudodontoideus_connasaugaensis
        |  +-----+                                   
        |  |     +-- Pseudodontoideus_subvexus       
        |  |                                         
        +--+     +-- Utterbackiana_couperiana        
           |  +--+                                   
           |  |  +-- Utterbackiana_heardi            
           +--+                                      
              |  +-- Utterbackiana_hartfieldorum     
              +--+                                   
                 +-- Utterbackiana_suborbiculata     

Amaro et al., 2017 [Publ 3791], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 17 4 3 --
        +-- Lasmigona_costata       
  +-----+                           
  |     +-- Platynaias_compressa    
--+                                 
  |  +----- Pyganodon_grandis **       
  +--+                              
     |  +-- Utterbackia_peninsularis
     +--+                           
        +-- Utterbackia_imbecillis  

Amaro et al., 2017 [Publ 3791], fig. 2-M

iT tT bi riS oT roT roS iD
1 1 0 1 9 2 2 --
-- Pyganodon_grandis

Amaro et al., 2017 [Publ 3791], fig. 3A

iT tT bi riS oT roT roS iD
1 1 0 1 11 3 2 --
  +-------- Platynaias_compressa    
--+                                 
  |  +----- Pyganodon_grandis **       
  +--+                              
     |  +-- Utterbackia_imbecillis  
     +--+                           
        +-- Utterbackia_peninsularis

Amaro et al., 2017 [Publ 3791], fig. 3A-M

iT tT bi riS oT roT roS iD
1 1 0 1 6 1 1 --
  +-- Pyganodon_grandis **       
--+                           
  +-- Utterbackia_peninsularis

Amaro et al., 2017 [Publ 3791], fig. 3B

iT tT bi riS oT roT roS iD
1 1 0 1 11 3 2 --
  +-------- Platynaias_compressa    
--+                                 
  |  +----- Pyganodon_grandis **       
  +--+                              
     |  +-- Utterbackia_imbecillis  
     +--+                           
        +-- Utterbackia_peninsularis

Amaro et al., 2017 [Publ 3791], fig. 3B-M

iT tT bi riS oT roT roS iD
1 1 0 1 6 1 1 --
  +-- Utterbackia_peninsularis
--+                           
  +-- Pyganodon_grandis **       

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 32 3 2 --
  +-------- Platynaias_compressa    
--+                                 
  |  +----- Pyganodon_grandis **       
  +--+                              
     |  +-- Utterbackia_peninsularis
     +--+                           
        +-- Utterbackia_imbecillis  

Lopes-Lima et al., 2017 [Publ 3792], fig. 2-M

iT tT bi riS oT roT roS iD
1 1 0 1 9 1 1 --
  +-- Utterbackia_peninsularis
--+                           
  +-- Pyganodon_grandis **       

Wen et al., 2017 [Publ 3795], fig. 3-M

iT tT bi riS oT roT roS iD
1 1 0 1 5 1 1 --
  +-- Utterbackia_peninsularis
--+                           
  +-- Pyganodon_grandis **       

Feng et al., 2016 [Publ 3800], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 14 1 1 --
-- Pyganodon_grandis

Bolotov et al., 2018 [Publ 3826], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 84 4 4 --
  +----------- Pyganodon_grandis **    
--+                                 
  |  +-------- Simpsonaias_ambigua  
  +--+                              
     |  +----- Strophitus_undulatus 
     +--+                           
        |  +-- Alasmidonta_marginata
        +--+                        
           +-- Lasmigona_costata    

Huang et al., 2018 [Publ 3828], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 42 3 2 --
  +-------- Platynaias_compressa    
--+                                 
  |  +----- Pyganodon_grandis **       
  +--+                              
     |  +-- Utterbackia_imbecillis  
     +--+                           
        +-- Utterbackia_peninsularis

Huang et al., 2018 [Publ 3828], fig. 6-M

iT tT bi riS oT roT roS iD
1 1 0 1 16 1 1 --
  +-- Pyganodon_grandis **       
--+                           
  +-- Utterbackia_peninsularis

Soroka & Burzynski, 2017 [Publ 3837], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 23 3 2 --
  +-------- Platynaias_compressa    
--+                                 
  |  +----- Pyganodon_grandis **       
  +--+                              
     |  +-- Utterbackia_imbecillis  
     +--+                           
        +-- Utterbackia_peninsularis

Huang et al., 2019 [Publ 3843], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 204 6 5 --
  +----- Pyganodon_grandis **       
--+                              
  |  +-- Utterbackia_imbecillis  
  +--+                           
     +-- Utterbackia_peninsularis

Huang et al., 2019 [Publ 3843], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 56 3 2 --
  +-------- Lasmigona_costata       
--+                                 
  |  +----- Pyganodon_grandis **       
  +--+                              
     |  +-- Utterbackia_imbecillis  
     +--+                           
        +-- Utterbackia_peninsularis

Huang et al., 2019 [Publ 3843], fig. 3-M

iT tT bi riS oT roT roS iD
1 1 0 1 22 1 1 --
  +-- Pyganodon_grandis **       
--+                           
  +-- Utterbackia_peninsularis

Huang et al., 2019 [Publ 3843], fig. S3

iT tT bi riS oT roT roS iD
1 1 0 1 201 6 5 --
  +----- Pyganodon_grandis **       
--+                              
  |  +-- Utterbackia_imbecillis  
  +--+                           
     +-- Utterbackia_peninsularis

Huang et al., 2019 [Publ 3843], fig. S4

iT tT bi riS oT roT roS iD
1 1 0 1 202 6 5 --
-- Pyganodon_grandis

Wang et al., 2018 [Publ 3846], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 31 1 1 --
  +-- Pyganodon_grandis **   
--+                       
  +-- Platynaias_compressa

Chase et al., 2018 [Publ 3861], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 10 1 1 --
  +-- Utterbackia_peninsularis
--+                           
  +-- Pyganodon_grandis **       

Chase et al., 2018 [Publ 3861], fig. 1-M

iT tT bi riS oT roT roS iD
1 1 0 1 10 1 1 --
  +-- Utterbackia_peninsularis
--+                           
  +-- Pyganodon_grandis **       

Burzynski & Soroka, 2018 [Publ 3862], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 3 1 1 --
  +-- Utterbackia_peninsularis
--+                           
  +-- Pyganodon_grandis **       

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

iT tT bi riS oT roT roS iD
1 1 0 1 19 1 1 --
  +-- Utterbackia_peninsularis
--+                           
  +-- Pyganodon_grandis **       

Burzynski & Soroka, 2018 [Publ 3862], fig. 5b

iT tT bi riS oT roT roS iD
1 1 0 1 9 1 1 --
  +-- Utterbackia_peninsularis
--+                           
  +-- Pyganodon_grandis **       

Aunins et al., 2018 [Publ 3863], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 40 4 3 --
        +-- Prolasmidonta_heterodon 
  +-----+                           
  |     +-- Alasmidonta_varicosa    
--+                                 
  |  +----- Pyganodon_grandis **       
  +--+                              
     |  +-- Utterbackia_imbecillis  
     +--+                           
        +-- Utterbackia_peninsularis

Wu et al., 2019 [Publ 3864], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 46 3 2 --
  +-------- Platynaias_compressa    
--+                                 
  |  +----- Pyganodon_grandis **       
  +--+                              
     |  +-- Utterbackia_peninsularis
     +--+                           
        +-- Utterbackia_imbecillis  

Wang et al., 2018 [Publ 3865], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 31 1 1 --
  +-- Pyganodon_grandis **   
--+                       
  +-- Platynaias_compressa

Lopes-Lima et al., 2019 [Publ 3901], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 27 0 0 --
-- Pyganodon_grandis

Lopes-Lima et al., 2019 [Publ 3901], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 28 0 0 --
-- Pyganodon_grandis

Lopes-Lima et al., 2019 [Publ 3901], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 25 0 0 --
-- Pyganodon_grandis

Cha et al., 2018 [Publ 3916], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 38 3 2 --
  +-------- Platynaias_compressa    
--+                                 
  |  +----- Pyganodon_grandis **       
  +--+                              
     |  +-- Utterbackia_peninsularis
     +--+                           
        +-- Utterbackia_imbecillis  

Froufe et al., 2019 [Publ 3936], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 44 3 2 --
  +-------- Lasmigona_complanata    
--+                                 
  |  +----- Pyganodon_grandis **       
  +--+                              
     |  +-- Utterbackia_peninsularis
     +--+                           
        +-- Utterbackia_imbecillis  

Froufe et al., 2019 [Publ 3936], fig. 3-M

iT tT bi riS oT roT roS iD
1 1 0 1 18 1 1 --
  +-- Utterbackia_peninsularis
--+                           
  +-- Pyganodon_grandis **       

Froufe et al., 2019 [Publ 3936], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 18 1 1 --
  +-- Utterbackia_peninsularis
--+                           
  +-- Pyganodon_grandis **       

Froufe et al., 2019 [Publ 3936], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 44 3 2 --
  +-------- Lasmigona_complanata    
--+                                 
  |  +----- Pyganodon_grandis **       
  +--+                              
     |  +-- Utterbackia_peninsularis
     +--+                           
        +-- Utterbackia_imbecillis  

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iT tT bi riS oT roT roS iD
1 1 0 1 56 0 0 --
-- Pyganodon_grandis

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 56 0 0 --
-- Pyganodon_grandis

Hewitt et al., 2019 [Publ 3962], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 15 0 0 --
-- Pyganodon_grandis

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iT tT bi riS oT roT roS iD
1 1 0 1 121 4 4 --
  +----- Pyganodon_grandis **    
  |                           
  +----- Simpsonaias_ambigua  
--+                           
  +----- Strophitus_undulatus 
  |                           
  |  +-- Alasmidonta_marginata
  +--+                        
     +-- Lasmigona_costata    

Riccardi et al., 2019 [Publ 4089], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 19 3 2 --
  +-------- Platynaias_compressa    
--+                                 
  |  +----- Pyganodon_grandis **       
  +--+                              
     |  +-- Utterbackia_peninsularis
     +--+                           
        +-- Utterbackia_imbecillis  

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 53 3 3 --
     +-- Pyganodon_grandis **       
  +--+                           
  |  +-- Utterbackia_peninsularis
--+                              
  |  +-- Alasmidonta_varicosa    
  +--+                           
     +-- Prolasmidonta_heterodon 

Wu et al., 2020 [Publ 4121], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 49 3 2 --
  +-------- Platynaias_compressa    
--+                                 
  |  +----- Pyganodon_grandis **       
  +--+                              
     |  +-- Utterbackia_peninsularis
     +--+                           
        +-- Utterbackia_imbecillis  

Sano et al., 2020 [Publ 4138], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 72 5 4 --
     +----- Alasmidonta_varicosa    
  +--+                              
  |  |  +-- Prolasmidonta_heterodon 
  |  +--+                           
--+     +-- Platynaias_compressa    
  |                                 
  |  +----- Pyganodon_grandis **       
  +--+                              
     |  +-- Utterbackia_imbecillis  
     +--+                           
        +-- Utterbackia_peninsularis

Sano et al., 2020 [Publ 4138], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 71 5 4 --
     +----- Pyganodon_grandis **       
  +--+                              
  |  |  +-- Utterbackia_imbecillis  
  |  +--+                           
--+     +-- Utterbackia_peninsularis
  |                                 
  |  +----- Alasmidonta_varicosa    
  +--+                              
     |  +-- Platynaias_compressa    
     +--+                           
        +-- Prolasmidonta_heterodon 

Sano et al., 2020 [Publ 4138], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 71 5 4 --
     +----- Alasmidonta_varicosa    
  +--+                              
  |  |  +-- Prolasmidonta_heterodon 
  |  +--+                           
--+     +-- Platynaias_compressa    
  |                                 
  |  +----- Pyganodon_grandis **       
  +--+                              
     |  +-- Utterbackia_imbecillis  
     +--+                           
        +-- Utterbackia_peninsularis

Wu et al., 2018 [Publ 4143], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 21 2 1 --
  +----- Pyganodon_grandis **       
--+                              
  |  +-- Utterbackia_peninsularis
  +--+                           
     +-- Utterbackia_imbecillis  

Wu et al., 2020 [Publ 4147], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 29 2 1 --
  +----- Pyganodon_grandis **       
--+                              
  |  +-- Utterbackia_peninsularis
  +--+                           
     +-- Utterbackia_imbecillis  

Guerra et al., 2019 [Publ 4168], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 40 4 2 --
        +-- Platynaias_compressa    
  +-----+                           
  |     +-- Platynaias_subviridis   
--+                                 
  |  +----- Pyganodon_grandis **       
  +--+                              
     |  +-- Utterbackia_peninsularis
     +--+                           
        +-- Utterbackia_imbecillis  

Guerra et al., 2019 [Publ 4168], fig. 1-M

iT tT bi riS oT roT roS iD
1 1 0 1 21 1 1 --
  +-- Utterbackia_peninsularis
--+                           
  +-- Pyganodon_grandis **       

Guerra et al., 2019 [Publ 4168], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 40 4 2 --
        +-- Platynaias_compressa    
  +-----+                           
  |     +-- Platynaias_subviridis   
--+                                 
  |  +----- Pyganodon_grandis **       
  +--+                              
     |  +-- Utterbackia_peninsularis
     +--+                           
        +-- Utterbackia_imbecillis  

Guerra et al., 2019 [Publ 4168], fig. 2-M

iT tT bi riS oT roT roS iD
1 1 0 1 21 1 1 --
  +-- Utterbackia_peninsularis
--+                           
  +-- Pyganodon_grandis **       

Chen et al., 2020 [Publ 4170], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 33 5 4 --
  +----- Platynaias_compressa  
--+                            
  |  +-- Utterbackia_imbecillis
  +--+                         
     +-- Pyganodon_grandis **     

Zieritz et al., 2021 [Publ 4231], fig. 3-F, 5

iT tT bi riS oT roT roS iD
1 1 0 1 48 3 2 --
  +-------- Platynaias_compressa    
--+                                 
  |  +----- Pyganodon_grandis **       
  +--+                              
     |  +-- Utterbackia_imbecillis  
     +--+                           
        +-- Utterbackia_peninsularis

Zieritz et al., 2021 [Publ 4231], fig. 3-M, 4

iT tT bi riS oT roT roS iD
1 1 0 1 22 1 1 --
  +-- Utterbackia_peninsularis
--+                           
  +-- Pyganodon_grandis **       

Wu et al., 2022 [Publ 4238], fig. 2d

iT tT bi riS oT roT roS iD
1 1 0 1 38 3 2 --
  +-------- Platynaias_compressa    
--+                                 
  |  +----- Pyganodon_grandis **       
  +--+                              
     |  +-- Utterbackia_imbecillis  
     +--+                           
        +-- Utterbackia_peninsularis

Wu et al., 2022 [Publ 4238], fig. 2e

iT tT bi riS oT roT roS iD
1 1 0 1 16 1 1 --
  +-- Utterbackia_peninsularis
--+                           
  +-- Pyganodon_grandis **       

Wu et al., 2022 [Publ 4238], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 73 3 2 --
  +-------- Platynaias_compressa    
--+                                 
  |  +----- Pyganodon_grandis **       
  +--+                              
     |  +-- Utterbackia_imbecillis  
     +--+                           
        +-- Utterbackia_peninsularis

Jeratthitikul et al., 2022 [Publ 4260], fig. S1, 1

iT tT bi riS oT roT roS iD
1 1 0 1 75 4 4 --
        +-- Pyganodon_grandis **    
  +-----+                        
  |     +-- Simpsonaias_ambigua  
--+                              
  |  +----- Strophitus_undulatus 
  +--+                           
     |  +-- Alasmidonta_marginata
     +--+                        
        +-- Lasmigona_costata    

Jeratthitikul et al., 2022 [Publ 4260], fig. S2, 2

iT tT bi riS oT roT roS iD
1 1 0 1 75 4 4 --
        +-- Alasmidonta_marginata
  +-----+                        
  |     +-- Lasmigona_costata    
--+                              
  |  +----- Pyganodon_grandis **    
  +--+                           
     |  +-- Strophitus_undulatus 
     +--+                        
        +-- Simpsonaias_ambigua  

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 35 0 0 --
-- Pyganodon_grandis

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iT tT bi riS oT roT roS iD
1 1 0 1 38 0 0 --
-- Pyganodon_grandis

Dai et al., 2023 [Publ 4285], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 52 2 2 --
  +----- Platynaias_compressa  
--+                            
  |  +-- Pyganodon_grandis **     
  +--+                         
     +-- Utterbackia_imbecillis

Whelan et al., 2023 [Publ 4287], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 28 27 11 --
                          +-- Utterbackia_peninsularis        
  +-----------------------+                                   
  |                       +-- Pyganodon_grandis **               
  |                                                           
  |     +-------------------- Alasminota_etowaensis           
  |     |                                                     
  |     |              +----- Pressodonta_viridis             
  |  +--+  +-----------+                                      
  |  |  |  |           |  +-- Platynaias_subviridis           
  |  |  |  |           +--+                                   
--+  |  |  |              +-- Prolasmidonta_heterodon         
  |  |  +--+                                                  
  |  |     |              +-- Anodontoides_ferussacianus      
  |  |     |  +-----------+                                   
  |  |     |  |           +-- Anodontoides_denigratus         
  |  |     +--+                                               
  |  |        |  +----------- Strophitus_undulatus            
  |  |        +--+                                            
  |  |           |  +-------- Strophitus_pascagoulaensis[2]   
  |  |           +--+                                         
  +--+              |  +----- Strophitus_pascagoulaensis[1]   
     |              +--+                                      
     |                 |  +-- Strophitus_radiatus             
     |                 +--+                                   
     |                    +-- Strophitus_williamsi            
     |                                                        
     |                    +-- Arcidens_confragosus            
     |                 +--+                                   
     |                 |  +-- Arcidens_wheeleri               
     |        +--------+                                      
     |        |        |  +-- Pseudodontoideus_connasaugaensis
     |        |        +--+                                   
     |        |           +-- Pseudodontoideus_subvexus       
     +--------+                                               
              |           +-- Alasmidonta_marginata           
              |  +--------+                                   
              |  |        +-- Alasmidonta_raveneliana         
              |  |                                            
              +--+        +-- Alasmidonta_varicosa            
                 |  +-----+                                   
                 |  |     +-- Alasmidonta_uwharriensis        
                 +--+                                         
                    |  +----- Alasmidonta_undulata            
                    +--+                                      
                       |  +-- Alasmidonta_triangulata         
                       +--+                                   
                          +-- Alasmidonta_arcula              

Wu et al., 2023 [Publ 4296], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 22 2 2 --
  +----- Lasmigona_costata   
--+                          
  |  +-- Strophitus_undulatus
  +--+                       
     +-- Pyganodon_grandis **   

Liu et al., 2023 [Publ 4310], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 23 2 2 --
  +----- Lasmigona_costata   
--+                          
  |  +-- Strophitus_undulatus
  +--+                       
     +-- Pyganodon_grandis **   

Liu et al., 2023 [Publ 4310], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 23 2 2 --
  +-- Pyganodon_grandis **   
--+                       
  +-- Strophitus_undulatus

Smith, 2021 [Publ 4331], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 19 1 1 --
  +-- Pyganodon_grandis **       
--+                           
  +-- Utterbackia_peninsularis

Wu et al., 2023 [Publ 4339], fig. 4, S5-aa

iT tT bi riS oT roT roS iD
1 1 0 1 95 6 5 --
     +----- Pyganodon_grandis **       
  +--+                              
  |  |  +-- Utterbackia_peninsularis
  |  +--+                           
  |     +-- Utterbackia_imbecillis  
--+                                 
  |     +-- Alasmidonta_varicosa    
  |  +--+                           
  |  |  +-- Platynaias_compressa    
  +--+                              
     |  +-- Prolasmidonta_heterodon 
     +--+                           
        +-- Lasmigona_complanata    

Wu et al., 2023 [Publ 4339], fig. 4, S5-nt

iT tT bi riS oT roT roS iD
1 1 0 1 96 6 5 --
        +----- Pyganodon_grandis **       
  +-----+                              
  |     |  +-- Utterbackia_imbecillis  
  |     +--+                           
--+        +-- Utterbackia_peninsularis
  |                                    
  |  +-------- Lasmigona_complanata    
  +--+                                 
     |  +----- Prolasmidonta_heterodon 
     +--+                              
        |  +-- Alasmidonta_varicosa    
        +--+                           
           +-- Platynaias_compressa    

Wu et al., 2024 [Publ 4350], fig. 8a, S1

iT tT bi riS oT roT roS iD
1 1 0 1 91 6 5 --
        +----- Pyganodon_grandis **       
  +-----+                              
  |     |  +-- Utterbackia_imbecillis  
  |     +--+                           
--+        +-- Utterbackia_peninsularis
  |                                    
  |  +-------- Lasmigona_complanata    
  +--+                                 
     |  +----- Prolasmidonta_heterodon 
     +--+                              
        |  +-- Alasmidonta_varicosa    
        +--+                           
           +-- Platynaias_compressa    

Wu et al., 2024 [Publ 4350], fig. 8b, S2

iT tT bi riS oT roT roS iD
1 1 0 1 91 6 5 --
        +----- Pyganodon_grandis **       
  +-----+                              
  |     |  +-- Utterbackia_imbecillis  
  |     +--+                           
--+        +-- Utterbackia_peninsularis
  |                                    
  |  +-------- Lasmigona_complanata    
  +--+                                 
     |  +----- Prolasmidonta_heterodon 
     +--+                              
        |  +-- Alasmidonta_varicosa    
        +--+                           
           +-- Platynaias_compressa    

Dai et al., 2024 [Publ 4358], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 51 2 2 --
  +----- Platynaias_compressa  
--+                            
  |  +-- Pyganodon_grandis **     
  +--+                         
     +-- Utterbackia_imbecillis

Wu et al., 2021 [Publ 4366], fig. 2, I

iT tT bi riS oT roT roS iD
1 1 0 1 90 2 2 --
  +-- Pyganodon_grandis **   
--+                       
  +-- Strophitus_undulatus

Wu et al., 2021 [Publ 4366], fig. 3-BI

iT tT bi riS oT roT roS iD
1 1 0 1 73 6 4 --
        +----- Pyganodon_grandis **       
  +-----+                              
  |     |  +-- Utterbackia_peninsularis
  |     +--+                           
--+        +-- Utterbackia_imbecillis  
  |                                    
  |  +-------- Prolasmidonta_heterodon 
  +--+                                 
     |  +----- Alasmidonta_varicosa    
     +--+                              
        |  +-- Platynaias_subviridis   
        +--+                           
           +-- Platynaias_compressa    

Wu et al., 2021 [Publ 4366], fig. 3-ML

iT tT bi riS oT roT roS iD
1 1 0 1 73 6 4 --
        +----- Pyganodon_grandis **       
  +-----+                              
  |     |  +-- Utterbackia_peninsularis
  |     +--+                           
--+        +-- Utterbackia_imbecillis  
  |                                    
  |  +-------- Prolasmidonta_heterodon 
  +--+                                 
     |  +----- Alasmidonta_varicosa    
     +--+                              
        |  +-- Platynaias_subviridis   
        +--+                           
           +-- Platynaias_compressa    

Wu et al., 2021 [Publ 4366], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 68 5 4 --
     +----- Pyganodon_grandis **       
  +--+                              
  |  |  +-- Utterbackia_imbecillis  
  |  +--+                           
--+     +-- Utterbackia_peninsularis
  |                                 
  |  +----- Prolasmidonta_heterodon 
  +--+                              
     |  +-- Platynaias_compressa    
     +--+                           
        +-- Alasmidonta_varicosa    

Wu et al., 2021 [Publ 4366], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 90 2 2 --
  +-- Pyganodon_grandis **   
--+                       
  +-- Strophitus_undulatus

Page updated: 10:06:06 Wed 05 Jun 2024

 
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