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Page last updated
Wed 01 Jul 2020

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

The nominal species (available and otherwise) in this synonymy are listed by usage rather than strictly by publication date. The primary sorting order is by the most recent year (since 1969) that the species name was treated as valid, and then nominal species are sorted secondarily by publication year.

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Ambleminae | Quadrulini | Cyclonaias

species Cyclonaias pustulosa (Lea, 1831): synonymy
     16 specimens available.

Unio pustulosus Lea, 1831
Lea, 1831, Trans. Amer. Phil. Soc. 4 [O. 1]: 76 [86], pl. 7, fig. 7.

Unio pustulosusLea (1831), Ravenel (1834), Jay (1835), Ferussac (1835), Hanley (1843), Singley (1893), Call (1895), de Gregorio (1914)
Margarita (Unio) pustulosusLea (1836)
Unio bullatusConrad (1838), Conrad (1853)
Unio (Theliderma) pustulosaSwainson (1840)
Margaron (Unio) pustulosusLea (1852), Lea (1870)
Unio (Quadrula) pustulosusMörch (1853)
Unio (Rotundaria) pustulosusAdams & Adams (1857)
Quadrula (sec. Theliderma) pustulosaSimpson (1900), Simpson (1914)
Unio (s.s.) pustulosusCall (1900)
Quadrula pustulosaOrtmann (1909), Ortmann (1910), Ortmann (1912), Utterback (1915), Ortmann & Walker (1922), Ortmann (1924), Ortmann (1925), Murray & Leonard (1962), Burch (1975), Mathiak (1979), Gordon et al. (1979), Mackie et al. (1980), R.I. Johnson (1980), Davis & Fuller (1981), Oesch (1984), Cummings & Mayer (1992), Vidrine (1993), Howells et al. (1996), Strayer & Jirka (1997), Parmalee & Bogan (1998), Johnson et al. (1998), Graf (2002), Bogan (2002), Cicerello & Schuster (2003), Sietman (2003), Serb et al. (2003), Jones et al. (2005), Henley et al. (2006), Williams et al. (2008), Bogan (2010), Boyer et al. (2011), Haag (2012), McMurray et al. (2012), Szumowski et al. (2012), Jones (2015), Inoue et al. (2018), Marshall et al. (2018)
Quadrula (Bullata) pustulosaFrierson (1927)
Quadrula (s.s.) pustulosusF.C. Baker (1928)
Quadrula (Pustulosa) pustulosa pustulosaHaas (1969)
Quadrula bullataMorrison (1969)
Quadrula pustulosa pustulosaTurgeon et al. (1988), Williams et al. (1993), Turgeon et al. (1998), Lydeard et al. (1999), Watters et al. (2009), ITIS (2012)
Amphinaias pustulosaGraf & Cummings (2007), Cummings & Graf (2010), IUCN (2011)
Rotundaria pustulosaCampbell & Lydeard (2012)
Cyclonaias pustulosaWilliams et al. (2017), Johnson et al. (2018), Lopes-Lima et al. (2019)

Unio mortoni Conrad, 1836
Conrad, 1836, Monogr. Fam. Unionidae 1(2): 11, pl. 6, fig. 1.
Conrad, 1853, Proc. Acad. Nat. Sci. 6: 252.
Type(s): HOLOTYPE ANSP_10287A, Bayou Teche, Louisiana.

Unio mortoniConrad (1836), Küster (1852), Conrad (1853)
Margarita (Unio) turgidusLea (1836)
Unio turgidusHanley (1843), Reeve (1864)
Margaron (Unio) turgidusLea (1852), Lea (1870)
Unio mortoniiSingley (1893)
Unio pustulosusCall (1895)
Quadrula (sec. Theliderma) mortoniSimpson (1900), Simpson (1914)
Quadrula mortoniOrtmann (1912), Vidrine (1993), Howells et al. (1996), Serb et al. (2003), Bogan (2010), Pfeiffer et al. (2016), Marshall et al. (2018)
Quadrula (Bullata) pustulosa-mortoniFrierson (1927)
Quadrula (Pustulosa) pustulosa mortoniHaas (1969)
Quadrula pustulosa mortoniTurgeon et al. (1988), Williams et al. (1993), Turgeon et al. (1998), ITIS (2012), Szumowski et al. (2012)
Quadrula pustulosaParmalee & Bogan (1998), Williams et al. (2008), Haag (2012)
Cyclonaias mortoniWilliams et al. (2017)
Cyclonaias pustulosaJohnson et al. (2018), Lopes-Lima et al. (2019)

Unio houstonensis Lea, 1859
Lea, 1859, Proc. Acad. Nat. Sci. 11: 155.
Lea, 1860, Jour. Acad. Nat. Sci. 4 [O. 8]: 358 [40], pl. 60, fig. 183.
Type(s): HOLOTYPE USNM_85768, Rutersville, Texas.

Unio houstonensisLea (1859), Sowerby (1868), Singley (1893)
Margaron (Unio) houstonensisLea (1870)
Quadrula (sec. Theliderma) houstonensisSimpson (1900), Simpson (1914)
Quadrula (Bullata) houstonensisFrierson (1927)
Quadrula houstonensisTurgeon et al. (1988), Williams et al. (1993), Howells et al. (1996), Turgeon et al. (1998), Bogan (2010), Breton et al. (2011), Haag (2012), ITIS (2012), Pfeiffer et al. (2016), Chase et al. (2018)
Pleurobema riddelliVidrine (1993)
Amphinaias houstonensisIUCN (1996), Graf & Cummings (2007), Cummings & Graf (2010)
Cyclonaias houstonensisWilliams et al. (2017)
Cyclonaias pustulosaJohnson et al. (2018), Lopes-Lima et al. (2019)

Unio refulgens Lea, 1868
Lea, 1868, Proc. Acad. Nat. Sci. 20: 145.
Lea, 1868, Jour. Acad. Nat. Sci. 6 [O. 12]: 317 [77], pl. 51, fig. 130.
Type(s): HOLOTYPE USNM_84290, Enterprise, Lauderdale Co., Miss. [Alabama].

Unio refulgensLea (1868)
Margaron (Unio) refulgensLea (1870)
Unio pustulosusCall (1895)
Quadrula (sec. Theliderma) refulgensSimpson (1900), Simpson (1914)
Quadrula refulgensOrtmann (1912), Turgeon et al. (1988), Williams et al. (1993), Vidrine (1993), Turgeon et al. (1998), Curole & Kocher (2002), Serb et al. (2003), Jones et al. (2005), Curole & Kocher (2005), Chapman et al. (2008), Chapman et al. (2008), Williams et al. (2008), Bogan (2010), Haag (2012), ITIS (2012), Szumowski et al. (2012), Inoue et al. (2018)
Quadrula (Bullata) pustulosa-refulgensFrierson (1927)
Quadrula (Pustulosa) pustulosa refulgensHaas (1969)
Quadrula pustulosaBurch (1975), R.I. Johnson (1980)
Amphinaias refulgensIUCN (1996), Graf & Cummings (2007), Cummings & Graf (2010)
Rotundaria refulgensCampbell & Lydeard (2012), Campbell & Lydeard (2012)
Cyclonaias refulgensWilliams et al. (2017)
Cyclonaias pustulosaJohnson et al. (2018), Lopes-Lima et al. (2019)

Unio aureus Lea, 1859
Lea, 1859, Proc. Acad. Nat. Sci. 11: 112.
Lea, 1862, Jour. Acad. Nat. Sci. 5 [O. 9]: 195 [17], pl. 26, fig. 264.
Type(s): HOLOTYPE USNM_84572, Texas.

Unio aureusLea (1859), Sowerby (1868), Paetel (1890), Singley (1893)
Margaron (Unio) aureusLea (1870)
Quadrula (sec. Theliderma) aureaSimpson (1900), Simpson (1914)
Quadrula (Bullata) aureusFrierson (1927)
Quadrula (Pustulosa) aureaHaas (1969)
Quadrula aureaBurch (1975), Turgeon et al. (1988), Williams et al. (1993), Howells et al. (1996), Turgeon et al. (1998), Serb et al. (2003), Bogan (2010), Haag (2012), ITIS (2012), Szumowski et al. (2012)
Amphinaias aureaGraf & Cummings (2007), Cummings & Graf (2010)
Cyclonaias aureaWilliams et al. (2017)
Cyclonaias pustulosaJohnson et al. (2018), Lopes-Lima et al. (2019)

Quadrula archeri Frierson, 1905
Frierson, 1905, Nautilus 19(2): 13, pl. 1, figs. 1-2.
Type(s): HOLOTYPE UMMZ_113538, Tallapoosa River, Tallassee, Elmore, Alabama.

Quadrula archeriFrierson (1905), Burch (1975)
Quadrula (sec. Theliderma) archeriSimpson (1914)
Quadrula (Bullata) archeriFrierson (1927)
Quadrula (Pustulosa) archeriHaas (1969)
Amphinaias archeriGraf & Cummings (2007), Cummings & Graf (2010)
Quadrula asperataWilliams et al. (2008)
Cyclonaias archeriWilliams et al. (2017)
Cyclonaias pustulosaLopes-Lima et al. (2019)

Obliquaria (Quadrula) retusa Rafinesque, 1820
Rafinesque, 1818, Amer. Mon. Mag. 3: 355 [as ‘Potamilus retusus’]. [nomen nudum]
Rafinesque, 1819, Jour. Phys. Chim. Hist. Nat. 88: 427 [as ‘U. (Obovaria) retusa’]. [nomen nudum]
Rafinesque, 1820, Ann. Gén. Sci. Phys. 5 (13): 306, pl. 81, figs. 19-20.
Rafinesque, 1864, [in] Binney & Tryon, reprint: 22, 29, 50.
(renamed Unio premorsus Rafinesque, 1831)
Type(s): NEOTYPE ANSP_20220, Green River.

Obliquaria (Quadrula) retusaRafinesque (1820)
Potamilus retususRafinesque (1818)
Unio (Obovaria) retusaRafinesque (1819)
Obliquaria (Quadrula) premorsusRafinesque (1831)
Unio retususFerussac (1835)
Obovaria retusaSimpson (1900), Simpson (1914)
Quadrula pustulosaOrtmann & Walker (1922), Williams et al. (2008)
Obliquaria retusaBurch (1975)
Quadrula pustulosa pustulosaWatters et al. (2009)
Cyclonaias pustulosaJohnson et al. (2018), Lopes-Lima et al. (2019)

Unio vallatus Lea, 1868
Lea, 1868, Jour. Acad. Nat. Sci. 6 [O. 12]: 315.
Lea, 1868, Jour. Acad. Nat. Sci. 6 [O. 12]: 315 [75], pl. 50, fig. 128.
Type(s): HOLOTYPE USNM_84275, Alabama R., Ala.

Unio vallatusLea (1868)
Margaron (Unio) vallatusLea (1870)
Unio pustulosusCall (1895)
Quadrula (sec. Theliderma) vallataSimpson (1900), Simpson (1914)
Quadrula (Bullata) pustulosa-vallataFrierson (1927)
Quadrula (Pustulosa) pustulosa vallataHaas (1969)
Quadrula pustulosaBurch (1975)
Quadrula asperataWilliams et al. (2008)
Cyclonaias pustulosaLopes-Lima et al. (2019)

Obliquaria (Quadrula) bullata Rafinesque, 1820
Rafinesque, 1820, Ann. Gén. Sci. Phys. 5 (13): 307.
Rafinesque, 1864, [in] Binney & Tryon, reprint: 51.
(homonym of Obliquaria (Quadrula) flexuosa var. bullata Rafinesque, 1820)
Type(s): NEOTYPE ANSP_20250, Kentucky River.

Obliquaria (Quadrula) bullataRafinesque (1820)
Unio bullatusFerussac (1835), Conrad (1838), Küster (1852), Conrad (1853)
Quadrula bullatusAgassiz (1852)
Unio bullataReeve (1864)
Unio (sec. Rotundaria) bullatusPaetel (1890)
Quadrula pustulosaOrtmann & Walker (1922), Burch (1975), R.I. Johnson (1980), Parmalee & Bogan (1998), Williams et al. (2008)
Quadrula (Bullata) pustulosaFrierson (1927)
Quadrula (Pustulosa) pustulosa pustulosaHaas (1969)
Quadrula bullataMorrison (1969)
Quadrula pustulosa pustulosaWatters et al. (2009)
Cyclonaias pustulosaJohnson et al. (2018), Lopes-Lima et al. (2019)

Unio premorsus Rafinesque, 1831
Rafinesque, 1831, Continuation: 4.
Rafinesque, 1864, [in] Binney & Tryon, reprint: 77.
(new name for Obliquaria (Quadrula) retusa Rafinesque, 1820)

Unio premorsusRafinesque (1831)
Obliquaria (Quadrula) premorsusRafinesque (1831)
Pleurobema premorsaMorrison (1969)
Quadrula pustulosaWilliams et al. (2008)
Quadrula pustulosa pustulosaWatters et al. (2009)
Cyclonaias pustulosaJohnson et al. (2018), Lopes-Lima et al. (2019)

Unio prasinus Conrad, 1834
Conrad, 1834, New Fresh-Water Shells: 44, pl. 3, fig.1.
Conrad, 1837, Monogr. Fam. Unionidae 1(9): 79, pl. 44, fig. 2.
Conrad, 1853, Proc. Acad. Nat. Sci. 6: 255.

Unio prasinusConrad (1834), Ferussac (1835), Conrad (1837), Conrad (1853), Reeve (1864)
Margarita (Unio) schoolcraftensisLea (1836)
Unio schoolcraftensisHanley (1843)
Margaron (Unio) schoolcraftensisLea (1852)
Margaron (Unio) schoolcraftiiLea (1870)
Unio (sec. Rotundaria) bullatus vr. prasinusPaetel (1890)
Unio pustulosusCall (1895)
Quadrula (sec. Theliderma) pustulosaSimpson (1900), Simpson (1914)
Quadrula pustulosa prasinaOrtmann & Walker (1922)
Quadrula (Bullata) pustulosa-prasinaFrierson (1927)
Quadrula (s.s.) pustulosus prasinaF.C. Baker (1928)
Quadrula (Pustulosa) pustulosa prasinaHaas (1969)
Quadrula pustulosaBurch (1975), Parmalee & Bogan (1998), Williams et al. (2008)
Quadrula quadrulaR.I. Johnson (1980)
Quadrula pustulosa pustulosaWatters et al. (2009)
Cyclonaias pustulosaJohnson et al. (2018), Lopes-Lima et al. (2019)

Unio pernodosus Lea, 1845
Lea, 1845, Proc. Amer. Phil. Soc. 4: 163.
Lea, 1848, Trans. Amer. Phil. Soc. 10 [O. 4]: 71 [45], pl. 3, fig. 8.
Type(s): SYNTYPE USNM_84298, N.C.

Unio pernodosusLea (1845), Conrad (1853), Küster (1861), Reeve (1864)
Margaron (Unio) pernodosusLea (1852), Lea (1870)
Unio pustulosusCall (1895)
Quadrula (sec. Theliderma) pustulosa var. pernodosaSimpson (1900), Simpson (1914)
Quadrula pernodosaIhering (1901)
Quadrula pustulosaOrtmann & Walker (1922), Burch (1975), Parmalee & Bogan (1998), Williams et al. (2008)
Quadrula (Bullata) pustulosa-asperataFrierson (1927)
Quadrula (Pustulosa) pustulosa pernodosaHaas (1969)
Quadrula pustulosa pustulosaWatters et al. (2009)
Cyclonaias pustulosaJohnson et al. (2018), Lopes-Lima et al. (2019)

Unio pauperculus Lea, 1861
Lea, 1861, Proc. Acad. Nat. Sci. 13: 39.
Lea, 1862, Jour. Acad. Nat. Sci. 5 [O. 8]: 99 [103], pl. 15, fig. 247.
Type(s): HOLOTYPE USNM_85763, Columbus, Miss.

Unio pauperculusLea (1861)
Margaron (Unio) pauperculusLea (1870)
Quadrula (sec. Theliderma) pauperculaSimpson (1900), Simpson (1914)
Quadrula (Bullata) pustulosa-pauperculaFrierson (1927)
Quadrula (Pustulosa) pustulosa pauperculaHaas (1969)
Quadrula asperataWilliams et al. (2008)
Cyclonaias pustulosaLopes-Lima et al. (2019)

Unio verrucosa Valenciennes, 1827
Valenciennes, 1827, Recueil Observ. Zool. Anat. Comp. 2(11-12): 231, pl. 53, fig. 2.
(homonym of Unio verrucosus Barnes, 1823)

Unio verrucosaValenciennes (1827)
Unio verrucosusLea (1829), Ferussac (1835)
Margarita (Unio) pustulosusLea (1836)
Margaron (Unio) pustulosusLea (1852), Lea (1870)
Quadrula (sec. Theliderma) pustulosaSimpson (1900), Simpson (1914)
Quadrula (Bullata) pustulosaFrierson (1927)
Quadrula (Pustulosa) pustulosa pustulosaHaas (1969)
Quadrula pustulosaParmalee & Bogan (1998)
Quadrula pustulosa pustulosaWatters et al. (2009)
Cyclonaias pustulosaJohnson et al. (2018), Lopes-Lima et al. (2019)

Unio nodulosus Say, 1834
Say, 1834, Amer. Conch. 6: ??.
(misidentified reference to Mya nodulosa Wood, 1815)

Unio nodulosusSay (1834)
Quadrula (sec. Theliderma) pustulosaSimpson (1900), Simpson (1914)
Quadrula pustulosaUtterback (1915), Oesch (1984), Howells et al. (1996), Parmalee & Bogan (1998), Williams et al. (2008)
Quadrula (Bullata) pustulosaFrierson (1927)
Quadrula (Pustulosa) pustulosa pustulosaHaas (1969)
Quadrula pustulosa pustulosaWatters et al. (2009)
Cyclonaias pustulosaJohnson et al. (2018), Lopes-Lima et al. (2019)

Unio schoolcraftensis Lea, 1834
Lea, 1834, Trans. Amer. Phil. Soc. 5 [O. 1]: 37 [149], pl. 3, fig. 9.

Unio schoolcraftensisLea (1834), Ferussac (1835), Hanley (1843), Reeve (1864)
Margarita (Unio) schoolcraftensisLea (1836)
Unio prasinusConrad (1837), Conrad (1853)
Margaron (Unio) schoolcraftensisLea (1852)
Unio (Quadrula) schoolcraftensisAdams & Adams (1857)
Margaron (Unio) schoolcraftiiLea (1870)
Unio (sec. Rotundaria) bullatus vr. schoolcraftensisPaetel (1890)
Unio pustulosusCall (1895)
Quadrula (sec. Theliderma) pustulosaSimpson (1900)
Quadrula (sec. Theliderma) pustulosa var. schoolcraftensisSimpson (1914)
Quadrula pustulosa schoolcraftensisUtterback (1915)
Quadrula pustulosa prasinaOrtmann & Walker (1922)
Quadrula (Bullata) pustulosa-prasinaFrierson (1927)
Quadrula (s.s.) pustulosus prasinaF.C. Baker (1928)
Quadrula (Pustulosa) pustulosa prasinaHaas (1969)
Quadrula pustulosaR.I. Johnson (1980), Parmalee & Bogan (1998), Williams et al. (2008)
Quadrula pustulosa pustulosaWatters et al. (2009)
Cyclonaias pustulosaJohnson et al. (2018), Lopes-Lima et al. (2019)

Unio dorfuillianus Lea, 1836
Lea, 1836, Synopsis Fam. Naiades: addenda. [nomen nudum]
(incorrect original spelling of Unio dorfeuillianus Lea, 1838)

Unio dorfuillianusLea (1836)

Unio bullatus Conrad, 1838
‘Rafinesque’ of Authors.
Conrad, 1838, Monogr. Fam. Unionidae 1(10): 82, pl. 45, fig. 2.
Lapham, 1852, Trans. WI Agri. Soc. 2: 369.
Reeve, 1864, Conch. Icon. 16: pl. 12, sp. 43.
(reference to Obliquaria (Quadrula) bullata Rafinesque, 1820)

Unio bullatusConrad (1838), Küster (1852)
Unio pustulosusFerussac (1835), Call (1895)
Margaron (Unio) pustulosusLea (1852), Lea (1870)
Quadrula (sec. Theliderma) pustulosaSimpson (1900), Simpson (1914)
Quadrula (s.s.) pustulosusF.C. Baker (1928)
Quadrula pustulosa pustulosaWatters et al. (2009)

Unio dorfeuillianus Lea, 1838
Lea, 1836, Synopsis Fam. Naiades: addenda [as ‘dorfuillianus’]. [nomen nudum]
Lea, 1838, Trans. Amer. Phil. Soc. 6 [O. 2]: 73, pl. 17, fig. 54.
(correct name for Unio dorfuillianus Lea, 1836)
Type(s): HOLOTYPE USNM_84215, Wabash River, Ind.

Unio dorfeuillianusLea (1838), Hanley (1843), Conrad (1853), Reeve (1864), Singley (1893)
Margarita (Unio) dorfeuillianusLea (1836)
Margaron (Unio) dorfeuillianusLea (1852), Lea (1870)
Unio (Quadrula) dorfeuillianusAdams & Adams (1857)
Unio (sec. Rotundaria) bullatus vr. dorfeuillianusPaetel (1890)
Unio pustulosusCall (1895)
Quadrula (sec. Theliderma) pustulosaSimpson (1900), Simpson (1914)
Unio (s.s.) pustulosusCall (1900)
Quadrula (Bullata) nodulataFrierson (1927)
Quadrula (s.s.) pustulosusF.C. Baker (1928)
Quadrula pustulosaMurray & Leonard (1962), Oesch (1984), Howells et al. (1996), Parmalee & Bogan (1998), Williams et al. (2008)
Quadrula pustulosa pustulosaWatters et al. (2009)
Cyclonaias pustulosaJohnson et al. (2018), Lopes-Lima et al. (2019)

Unio turgidus Lea, 1838
Lea, 1836, Synopsis Fam. Naiades: 16. [nomen nudum]
Lea, 1838, Trans. Amer. Phil. Soc. 6 [O. 2]: 11, pl. 5, fig. 11.
Type(s): HOLOTYPE USNM_84277, near New Orleans, La.

Unio turgidusLea (1838), Hanley (1843), Reeve (1864)
Margarita (Unio) turgidusLea (1836)
Margaron (Unio) turgidusLea (1852), Lea (1870)
Unio mortoniConrad (1853)
Unio (Quadrula) turgidusMörch (1853)
Unio (Rotundaria) turgidusAdams & Adams (1857)
Unio turgidisSingley (1893)
Unio pustulosusCall (1895)
Quadrula (sec. Theliderma) mortoniSimpson (1900), Simpson (1914)
Quadrula (Bullata) pustulosa-mortoniFrierson (1927)
Quadrula (Pustulosa) pustulosa mortoniHaas (1969)
Quadrula mortoniVidrine (1993), Howells et al. (1996)
Quadrula pustulosaParmalee & Bogan (1998), Williams et al. (2008)
Cyclonaias pustulosaJohnson et al. (2018), Lopes-Lima et al. (2019)

Unio nodiferus Conrad, 1841
Conrad, 1841, Proc. Acad. Nat. Sci. 1: 19.
Conrad, 1850, Jour. Acad. Nat. Sci. 1: 277, pl. 38, figs. 4, 8.
Conrad, 1853, Proc. Acad. Nat. Sci. 6: 253.
Type(s): LECTOTYPE ANSP_10285, Jackson, La.

Unio nodiferusConrad (1841), Conrad (1853)
Margaron (Unio) schoolcraftensisLea (1852)
Unio schoolcraftensisReeve (1864)
Margaron (Unio) schoolcraftiiLea (1870)
Unio nodiferousSingley (1893)
Quadrula (sec. Theliderma) nodiferaSimpson (1900), Simpson (1914)
Quadrula (Bullata) pustulosa-mortoniFrierson (1927)
Quadrula (Pustulosa) pustulosa mortoniHaas (1969)
Quadrula pustulosaBurch (1975)
Quadrula mortoniVidrine (1993), Howells et al. (1996)
Cyclonaias pustulosaJohnson et al. (2018), Lopes-Lima et al. (2019)

Unio sphaericus Lea, 1868
Lea, 1868, Proc. Acad. Nat. Sci. 20: 145 [as ‘sphæricus’].
Lea, 1868, Jour. Acad. Nat. Sci. 6 [O. 12]: 319 [79], pl. 51, fig. 132 [as ‘sphæricus’].
Type(s): HOLOTYPE USNM_84271, Pearl River, Jackson, Miss.

Unio sphaericusLea (1868), Singley (1893)
Margaron (Unio) spæricusLea (1870)
Unio pustulosusCall (1895)
Quadrula (sec. Theliderma) sphaericaSimpson (1900), Simpson (1914)
Quadrula sphaericaOrtmann (1912)
Quadrula (Bullata) pustulosa-refulgensFrierson (1927)
Quadrula pustulosaBurch (1975)
Quadrula mortoniVidrine (1993), Howells et al. (1996)
Cyclonaias pustulosaJohnson et al. (2018), Lopes-Lima et al. (2019)

Margaron (Unio) spaericus Lea, 1870
‘Lea’ Lea, 1870, Synopsis Fam. Naiades: 33 [as ‘spæricus’].
(error for Unio sphaericus Lea, 1868)

Margaron (Unio) spaericusLea (1870)

Unio schoolcraftii Lea, 1870
‘Lea’ Lea, 1870, Synopsis Fam. Naiades: 33.
(error for Unio schoolcraftensis Lea, 1834)

Unio schoolcraftiiLea (1870)
Unio pustulosusCall (1895)
Quadrula (sec. Theliderma) pustulosaSimpson (1900)
Quadrula (sec. Theliderma) pustulosa var. schoolcraftensisSimpson (1914)
Quadrula (Bullata) pustulosa-prasinaFrierson (1927)
Quadrula (Pustulosa) pustulosa prasinaHaas (1969)
Quadrula pustulosaVidrine (1993), Parmalee & Bogan (1998), Williams et al. (2008)
Quadrula pustulosa pustulosaWatters et al. (2009)

Unio cahabensis Lea, 1871
Lea, 1871, Proc. Acad. Nat. Sci. 23: 190.
Lea, 1874, Jour. Acad. Nat. Sci. 8 [O. 13]: 17 [21], pl. 5, fig. 14.
Type(s): HOLOTYPE USNM_84204, Cahaba R., Ala.

Unio cahabensisLea (1871)
Unio pustulosusCall (1895)
Quadrula (sec. Theliderma) cahabensisSimpson (1900), Simpson (1914)
Quadrula (Bullata) pustulosa-vallataFrierson (1927)
Quadrula (Pustulosa) pustulosa vallataHaas (1969)
Quadrula pustulosaBurch (1975)
Quadrula asperataWilliams et al. (2008)
Cyclonaias pustulosaLopes-Lima et al. (2019)

Unio nodiferous Singley, 1893
‘Conrad’ Singley, 1893, Geol. Surv. Texas (1892): 319.
(error for Unio nodiferus Conrad, 1841)

Unio nodiferousSingley (1893)

Unio turgidis Singley, 1893
‘Lea’ Singley, 1893, Geol. Surv. Texas (1892): 319.
(error for Unio turgidus Lea, 1838)

Unio turgidisSingley (1893)

Unio pustulosus Call, 1895
‘Lea’ Call, 1895, Trans. Acad. Sci. St. Louis 7: 42, pl. 15, figs. 3-4.
(reference to Unio pustulosus Lea, 1831)

Unio pustulosusCall (1895)
Quadrula (sec. Theliderma) refulgensSimpson (1900), Simpson (1914)

Quadrula pustulosa asperatus Utterback, 1915
‘Lea’ Utterback, 1915, Amer. Midl. Nat. 4(4): 134, pl. 17, figs. 42a-b.
(misidentified reference to Unio asperatus Lea, 1861)

Quadrula pustulosa asperatusUtterback (1915)
Quadrula pustulosaParmalee & Bogan (1998)
Quadrula pustulosa pustulosaWatters et al. (2009)

Unio petrinus Frierson, 1927
‘Call (Mss)’ Frierson, 1927, Check List N. Amer. Naiades: 49.
(intro'd as syn of Unio aureus Lea, 1859)

Unio petrinusFrierson (1927)
Quadrula (Bullata) aureusFrierson (1927)
Cyclonaias pustulosaJohnson et al. (2018)

Page updated: 16:16:21 Wed 01 Jul 2020

 
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