Unio pustulosus Lea, 1831
Lea, 1831, Trans. Amer. Phil. Soc. 4 [O. 1]: 76 [86], pl. 7, fig. 7.
Type(s): SYNTYPES ANSP_43058, Ohio.

Unio pustulosus — Lea (1831), Ravenel (1834), Jay (1835), Ferussac (1835), ..., Singley (1893), Call (1895), Pilsbry & Rhoads (1896), de Gregorio (1914)
Margarita (Unio) pustulosus — Lea (1836), Lea (1838)
Unio bullatus — Conrad (1838), Conrad (1853)
Unio (Theliderma) pustulosa — Swainson (1840)
Margaron (Unio) pustulosus — Lea (1852), Lea (1870)
Unio (Quadrula) pustulosus — Mörch (1853)
Unio (Rotundaria) pustulosus — Adams & Adams (1857)
Unio postulosus — Lapham (1860)
Quadrula (sec. Theliderma) pustulosa — Simpson (1900), Simpson (1914)
Unio (s.s.) pustulosus — Call (1900)
Quadrula pustulosa — Ortmann (1909), Ortmann (1909), Ortmann (1909), Ortmann (1910), ..., Inoue et al. (2018), Marshall et al. (2018), Gasparini et al. (2020), Klymus et al. (2020)
Quadrula (Bullata) pustulosa — Frierson (1927)
Quadrula (s.s.) pustulosus — F.C. Baker (1928)
Quadrula (Pustulosa) pustulosa pustulosa — Haas (1969)
Quadrula bullata — Morrison (1969)
Quadrula pustulosa pustulosa — Turgeon et al. (1988), Williams et al. (1993), Turgeon et al. (1998), Lydeard et al. (1999), Watters et al. (2009), ITIS (2012)
Amphinaias pustulosa — Graf & Cummings (2007), Cummings & Graf (2010), IUCN (2011), Roe & Boyer (2015)
Rotundaria pustulosa — Campbell & Lydeard (2012)
Cyclonaias pustulosa — Williams et al. (2017), Johnson et al. (2018), Lopes-Lima et al. (2019), Graf & Cummings (2021), FMCS (2021), Bieler (2021), Hopper et al. (2023), Freeman et al. (2025)
Pustulosa pustulosa — Neemuchwala et al. (2023)

Unio refulgens Lea, 1868
Lea, 1868, Proc. Acad. Nat. Sci. 20: 145.
Lea, 1868, Jour. Acad. Nat. Sci. 6 [O. 12]: 317 [77], pl. 51, fig. 130.
Type(s): FIGURED SYNTYPE USNM_84290, Enterprise, Lauderdale Co., Miss. [Alabama].

Unio refulgens — Lea (1868), B.H. Wright (1888)
Margaron (Unio) refulgens — Lea (1870)
Unio pustulosus — Call (1895)
Quadrula (sec. Theliderma) refulgens — Simpson (1900), Simpson (1914)
Quadrula refulgens — Ortmann (1912), Turgeon et al. (1988), Williams et al. (1993), Vidrine (1993), ..., Haag (2012), ITIS (2012), Szumowski et al. (2012), Inoue et al. (2018)
Quadrula (Bullata) pustulosa-refulgens — Frierson (1927)
Quadrula (Pustulosa) pustulosa refulgens — Haas (1969)
Quadrula pustulosa — Burch (1975), R.I. Johnson (1980)
Amphinaias refulgens — IUCN (1996), Graf & Cummings (2007), Cummings & Graf (2010)
Rotundaria refulgens — Campbell & Lydeard (2012), Campbell & Lydeard (2012)
Cyclonaias refulgens — Williams et al. (2017)
Cyclonaias pustulosa — Johnson et al. (2018), Lopes-Lima et al. (2019), Graf & Cummings (2021), FMCS (2021)
Cyclonaias (Quadrula) refulgens — Porto-Hannes et al. (2021)

Unio mortoni Conrad, 1835
Conrad, 1835, Monogr. Fam. Unionidae 1(1): 11, pl. 6, fig. 1.
Conrad, 1853, Proc. Acad. Nat. Sci. 6: 252.
Type(s): HOLOTYPE ANSP_10287A, Bayou Teche, Louisiana.

Unio mortoni — Conrad (1835), Küster (1852), Conrad (1853), B.H. Wright (1888)
Margarita (Unio) turgidus — Lea (1836), Lea (1838)
Unio turgidus — Hanley (1843), Reeve (1864)
Margaron (Unio) turgidus — Lea (1852), Lea (1870)
Unio mortonii — Singley (1893)
Unio pustulosus — Call (1895)
Quadrula (sec. Theliderma) mortoni — Simpson (1900), Simpson (1914)
Quadrula mortoni — Ortmann (1912), Vidrine (1993), Howells et al. (1996), Serb et al. (2003), Bogan (2010), Pfeiffer et al. (2016), Marshall et al. (2018)
Quadrula (Bullata) pustulosa-mortoni — Frierson (1927)
Quadrula (Pustulosa) pustulosa mortoni — Haas (1969)
Quadrula pustulosa mortoni — Turgeon et al. (1988), Williams et al. (1993), Turgeon et al. (1998), ITIS (2012), Szumowski et al. (2012)
Quadrula pustulosa — Parmalee & Bogan (1998), Williams et al. (2008), Haag (2012)
Cyclonaias mortoni — Williams et al. (2017)
Cyclonaias pustulosa — Johnson et al. (2018), Lopes-Lima et al. (2019), FMCS (2021)

Unio houstonensis Lea, 1859
Lea, 1859, Proc. Acad. Nat. Sci. 11: 155.
Lea, 1860, Jour. Acad. Nat. Sci. 4 [O. 8]: 358 [40], pl. 60, fig. 183.
Type(s): HOLOTYPE USNM_85768, Rutersville, Texas.

Unio houstonensis — Lea (1859), Sowerby (1868), B.H. Wright (1888), Singley (1893)
Margaron (Unio) houstonensis — Lea (1870)
Quadrula (sec. Theliderma) houstonensis — Simpson (1900), Simpson (1914)
Quadrula (Bullata) houstonensis — Frierson (1927)
Quadrula houstonensis — Turgeon et al. (1988), Williams et al. (1993), Howells et al. (1996), Turgeon et al. (1998), Bogan (2010), Breton et al. (2011), Haag (2012), ITIS (2012), Pfeiffer et al. (2016), Chase et al. (2018)
Pleurobema riddelli — Vidrine (1993)
Amphinaias houstonensis — IUCN (1996), Graf & Cummings (2007), Cummings & Graf (2010)
Cyclonaias houstonensis — Williams et al. (2017)
Cyclonaias pustulosa — Johnson et al. (2018), Lopes-Lima et al. (2019), Graf & Cummings (2021), FMCS (2021)

Unio aureus Lea, 1859
Lea, 1859, Proc. Acad. Nat. Sci. 11: 112.
Lea, 1862, Jour. Acad. Nat. Sci. 5 [O. 9]: 195 [17], pl. 26, fig. 264.
Type(s): HOLOTYPE USNM_84572, Texas.

Unio aureus — Lea (1859), Sowerby (1868), B.H. Wright (1888), Paetel (1890), Singley (1893)
Margaron (Unio) aureus — Lea (1870)
Quadrula (sec. Theliderma) aurea — Simpson (1900), Simpson (1914)
Quadrula (Bullata) aureus — Frierson (1927)
Quadrula (Pustulosa) aurea — Haas (1969)
Quadrula aurea — Burch (1975), Turgeon et al. (1988), Williams et al. (1993), Howells et al. (1996), Turgeon et al. (1998), Serb et al. (2003), Bogan (2010), Haag (2012), ITIS (2012), Szumowski et al. (2012)
Amphinaias aurea — Graf & Cummings (2007), Cummings & Graf (2010)
Cyclonaias aurea — Williams et al. (2017)
Cyclonaias pustulosa — Johnson et al. (2018), Lopes-Lima et al. (2019), Graf & Cummings (2021), FMCS (2021)

Obliquaria (Quadrula) retusa Rafinesque, 1820
Rafinesque, 1818, Amer. Mon. Mag. 3: 355 [as ‘Potamilus retusus’]. [nomen nudum]
Rafinesque, 1819, Jour. Phys. Chim. Hist. Nat. 88: 427 [as ‘U. (Obovaria) retusa’]. [nomen nudum]
Rafinesque, 1820, Ann. Gén. Sci. Phys. 5 (13): 306, pl. 81, figs. 19-20 [also as ‘Unio retusa’].
Rafinesque, 1864, [in] Binney & Tryon, reprint: 22, 29, 50.
(renamed Unio premorsus Rafinesque, 1831)
Type(s): NEOTYPE ANSP_20220, Green River.

Obliquaria (Quadrula) retusa — Rafinesque (1820)
Potamilus retusus — Rafinesque (1818)
Unio (Obovaria) retusa — Rafinesque (1819)
Obliquaria (Quadrula) premorsus — Rafinesque (1831)
Unio retusus — Ferussac (1835)
Obovaria retusa — Simpson (1900), Simpson (1914)
Quadrula pustulosa — Vanatta (1915), Ortmann & Walker (1922), Williams et al. (2008)
Obliquaria retusa — Burch (1975)
Quadrula pustulosa pustulosa — Watters et al. (2009)
Cyclonaias pustulosa — Johnson et al. (2018), Lopes-Lima et al. (2019)

Unio vallatus Lea, 1868
Lea, 1868, Proc. Acad. Nat. Sci. 20: 145.
Lea, 1868, Jour. Acad. Nat. Sci. 6 [O. 12]: 315 [75], pl. 50, fig. 128.
Type(s): HOLOTYPE (MONOTYPY) USNM_84275, Alabama R., Ala.

Unio vallatus — Lea (1868), B.H. Wright (1888)
Margaron (Unio) vallatus — Lea (1870)
Unio pustulosus — Call (1895)
Quadrula (sec. Theliderma) vallata — Simpson (1900), Simpson (1914)
Quadrula (Bullata) pustulosa-vallata — Frierson (1927)
Quadrula (Pustulosa) pustulosa vallata — Haas (1969)
Quadrula pustulosa — Burch (1975)
Quadrula asperata — Williams et al. (2008)
Cyclonaias pustulosa — Lopes-Lima et al. (2019)

Obliquaria (Quadrula) bullata Rafinesque, 1820
Rafinesque, 1820, Ann. Gén. Sci. Phys. 5 (13): 307 [also as ‘U. bullata’].
Rafinesque, 1864, [in] Binney & Tryon, reprint: 51.
(homonym of Obliquaria (Quadrula) flexuosa var. bullata Rafinesque, 1820)
Type(s): NEOTYPE ANSP_20250, Kentucky River.

Obliquaria (Quadrula) bullata — Rafinesque (1820)
Unio bullatus — Ferussac (1835), Conrad (1838), Küster (1852), Conrad (1853)
Quadrula bullatus — Agassiz (1852)
Unio bullata — Reeve (1864)
Unio (sec. Rotundaria) bullatus — Paetel (1890)
Quadrula pustulosa pernodosa — Vanatta (1915)
Quadrula pustulosa — Ortmann (1918), Ortmann & Walker (1922), Burch (1975), R.I. Johnson (1980), Parmalee & Bogan (1998), Williams et al. (2008)
Quadrula (Bullata) pustulosa — Frierson (1927)
Quadrula (Pustulosa) pustulosa pustulosa — Haas (1969)
Quadrula bullata — Morrison (1969)
Quadrula pustulosa pustulosa — Watters et al. (2009)
Cyclonaias pustulosa — Johnson et al. (2018), Lopes-Lima et al. (2019)

Unio premorsus Rafinesque, 1831
Rafinesque, 1831, Continuation: 4.
Rafinesque, 1864, [in] Binney & Tryon, reprint: 77.
(new name for Obliquaria (Quadrula) retusa Rafinesque, 1820)

Unio premorsus — Rafinesque (1831)
Obliquaria (Quadrula) premorsus — Rafinesque (1831)
Pleurobema premorsa — Morrison (1969)
Quadrula pustulosa — Williams et al. (2008)
Quadrula pustulosa pustulosa — Watters et al. (2009)
Cyclonaias pustulosa — Johnson et al. (2018), Lopes-Lima et al. (2019)

Unio prasinus Conrad, 1834
Conrad, 1834, New Fresh-Water Shells: 44, pl. 3, fig.1.
Conrad, 1837, Monogr. Fam. Unionidae 1(9): 79, pl. 44, fig. 2.
Conrad, 1853, Proc. Acad. Nat. Sci. 6: 255.

Unio prasinus — Conrad (1834), Ferussac (1835), Conrad (1837), Conrad (1853), Reeve (1864), B.H. Wright (1888)
Margarita (Unio) schoolcraftensis — Lea (1836), Lea (1838)
Unio schoolcraftensis — Hanley (1843)
Margaron (Unio) schoolcraftensis — Lea (1852)
Margaron (Unio) schoolcraftii — Lea (1870)
Unio (sec. Rotundaria) bullatus vr. prasinus — Paetel (1890)
Unio pustulosus — Call (1895)
Quadrula (sec. Theliderma) pustulosa — Simpson (1900), Simpson (1914)
Quadrula pustulosa prasina — Ortmann & Walker (1922)
Quadrula (Bullata) pustulosa-prasina — Frierson (1927)
Quadrula (s.s.) pustulosus prasina — F.C. Baker (1928)
Quadrula (Pustulosa) pustulosa prasina — Haas (1969)
Quadrula pustulosa — Burch (1975), Parmalee & Bogan (1998), Williams et al. (2008)
Quadrula quadrula — R.I. Johnson (1980)
Quadrula pustulosa pustulosa — Watters et al. (2009)
Cyclonaias pustulosa — Johnson et al. (2018), Lopes-Lima et al. (2019)

Unio pernodosus Lea, 1845
Lea, 1845, Proc. Amer. Phil. Soc. 4: 163.
Lea, 1848, Trans. Amer. Phil. Soc. 10 [O. 4]: 71 [45], pl. 3, fig. 8.
Type(s): SYNTYPE USNM_84298, N.C.

Unio pernodosus — Lea (1845), Conrad (1853), Küster (1861), Reeve (1864), Lewis (1871), B.H. Wright (1888)
Margaron (Unio) pernodosus — Lea (1852), Lea (1870)
Unio pustulosus — Call (1895)
Quadrula (sec. Theliderma) pustulosa var. pernodosa — Simpson (1900), Simpson (1914)
Quadrula pernodosa — Ihering (1901)
Quadrula pustulosa pernodosa — Vanatta (1915)
Quadrula pustulosa — Ortmann (1918), Ortmann & Walker (1922), Burch (1975), Parmalee & Bogan (1998), Williams et al. (2008)
Quadrula (Bullata) pustulosa-asperata — Frierson (1927)
Quadrula (Pustulosa) pustulosa pernodosa — Haas (1969)
Quadrula pustulosa pustulosa — Watters et al. (2009)
Cyclonaias pustulosa — Johnson et al. (2018), Lopes-Lima et al. (2019)

Unio pauperculus Lea, 1861
Lea, 1861, Proc. Acad. Nat. Sci. 13: 39.
Lea, 1862, Jour. Acad. Nat. Sci. 5 [O. 8]: 99 [103], pl. 15, fig. 247.
Type(s): HOLOTYPE USNM_85763, Columbus, Miss.

Unio pauperculus — Lea (1861), B.H. Wright (1888)
Margaron (Unio) pauperculus — Lea (1870)
Quadrula (sec. Theliderma) paupercula — Simpson (1900), Simpson (1914)
Quadrula (Bullata) pustulosa-paupercula — Frierson (1927)
Quadrula (Pustulosa) pustulosa paupercula — Haas (1969)
Quadrula asperata — Williams et al. (2008)
Cyclonaias pustulosa — Lopes-Lima et al. (2019)

Unio verrucosa Valenciennes, 1827
Valenciennes, 1827, Receuil Observ. Zool. 2(11-12): 231, pl. 53, fig. 2.
(homonym of Unio verrucosus Barnes, 1823)

Unio verrucosa — Valenciennes (1827)
Unio verrucosus — Lea (1829), Ferussac (1835)
Margarita (Unio) pustulosus — Lea (1836)
Margaron (Unio) pustulosus — Lea (1852), Lea (1870)
Quadrula (sec. Theliderma) pustulosa — Simpson (1900), Simpson (1914)
Quadrula (Bullata) pustulosa — Frierson (1927)
Quadrula (Pustulosa) pustulosa pustulosa — Haas (1969)
Quadrula pustulosa — Parmalee & Bogan (1998)
Quadrula pustulosa pustulosa — Watters et al. (2009)
Cyclonaias pustulosa — Johnson et al. (2018), Lopes-Lima et al. (2019)

Unio nodulosus Say, 1834
Say, 1834, Amer. Conch. 6: ??.
(misidentified reference to Mya nodulosa Wood, 1815)

Unio nodulosus — Say (1834)
Quadrula (sec. Theliderma) pustulosa — Simpson (1900), Simpson (1914)
Quadrula pustulosa — Utterback (1915), Oesch (1984), Howells et al. (1996), Parmalee & Bogan (1998), Williams et al. (2008)
Quadrula (Bullata) pustulosa — Frierson (1927)
Quadrula (Pustulosa) pustulosa pustulosa — Haas (1969)
Quadrula pustulosa pustulosa — Watters et al. (2009)
Cyclonaias pustulosa — Johnson et al. (2018), Lopes-Lima et al. (2019)

Unio schoolcraftensis Lea, 1834
Lea, 1834, Trans. Amer. Phil. Soc. 5 [O. 1]: 37 [149], pl. 3, fig. 9.

Unio schoolcraftensis — Lea (1834), Ferussac (1835), Hanley (1843), Lapham (1860), Reeve (1864)
Margarita (Unio) schoolcraftensis — Lea (1836), Lea (1838)
Unio prasinus — Conrad (1837), Conrad (1853)
Margaron (Unio) schoolcraftensis — Lea (1852)
Unio (Quadrula) schoolcraftensis — Adams & Adams (1857)
Margaron (Unio) schoolcraftii — Lea (1870)
Unio schoolcraftii — B.H. Wright (1888)
Unio (sec. Rotundaria) bullatus vr. schoolcraftensis — Paetel (1890)
Unio pustulosus — Call (1895)
Quadrula (sec. Theliderma) pustulosa — Simpson (1900)
Quadrula (sec. Theliderma) pustulosa var. schoolcraftensis — Simpson (1914)
Quadrula pustulosa schoolcraftensis — Utterback (1915)
Quadrula pustulosa prasina — Ortmann & Walker (1922)
Quadrula (Bullata) pustulosa-prasina — Frierson (1927)
Quadrula (s.s.) pustulosus prasina — F.C. Baker (1928)
Quadrula (Pustulosa) pustulosa prasina — Haas (1969)
Quadrula pustulosa — R.I. Johnson (1980), Parmalee & Bogan (1998), Williams et al. (2008)
Quadrula pustulosa pustulosa — Watters et al. (2009)
Cyclonaias pustulosa — Johnson et al. (2018), Lopes-Lima et al. (2019), Bieler (2021)

Unio dorfuillianus Lea, 1836
Lea, 1836, Synopsis Fam. Naiades: addenda. [nomen nudum]
(incorrect original spelling of Unio dorfeuillianus Lea, 1838)

Unio dorfuillianus — Lea (1836)

Unio bullatus Conrad, 1838
‘Rafinesque’ of Authors.
Conrad, 1838, Monogr. Fam. Unionidae 1(10): 82, pl. 45, fig. 2.
Lapham, 1852, Trans. WI Agri. Soc. 2: 369.
Reeve, 1864, Conch. Icon. 16: pl. 12, sp. 43.
(reference to Obliquaria (Quadrula) bullata Rafinesque, 1820)

Unio bullatus — Conrad (1838), Küster (1852)
Unio pustulosus — Ferussac (1835), Call (1895)
Margaron (Unio) pustulosus — Lea (1852), Lea (1870)
Quadrula (sec. Theliderma) pustulosa — Simpson (1900), Simpson (1914)
Quadrula (s.s.) pustulosus — F.C. Baker (1928)
Quadrula pustulosa pustulosa — Watters et al. (2009)

Unio dorfeuillianus Lea, 1838
Lea, 1836, Synopsis Fam. Naiades: addenda [as ‘dorfuillianus’]. [nomen nudum]
Lea, 1838, Trans. Amer. Phil. Soc. 6 [O. 2]: 73, pl. 17, fig. 54.
(correct name for Unio dorfuillianus Lea, 1836)
Type(s): HOLOTYPE USNM_84215, Wabash River, Ind.

Unio dorfeuillianus — Lea (1838), Hanley (1843), Conrad (1853), Reeve (1864), B.H. Wright (1888), Singley (1893)
Margarita (Unio) dorfeuillianus — Lea (1836), Lea (1838)
Margaron (Unio) dorfeuillianus — Lea (1852), Lea (1870)
Unio (Quadrula) dorfeuillianus — Adams & Adams (1857)
Unio (sec. Rotundaria) bullatus vr. dorfeuillianus — Paetel (1890)
Unio pustulosus — Call (1895)
Quadrula (sec. Theliderma) pustulosa — Simpson (1900), Simpson (1914)
Unio (s.s.) pustulosus — Call (1900)
Quadrula (Bullata) nodulata — Frierson (1927)
Quadrula (s.s.) pustulosus — F.C. Baker (1928)
Quadrula pustulosa — Murray & Leonard (1962), Oesch (1984), Howells et al. (1996), Parmalee & Bogan (1998), Williams et al. (2008)
Quadrula pustulosa pustulosa — Watters et al. (2009)
Cyclonaias pustulosa — Johnson et al. (2018), Lopes-Lima et al. (2019)

Unio turgidus Lea, 1838
Lea, 1836, Synopsis Fam. Naiades: 16. [nomen nudum]
Lea, 1838, Trans. Amer. Phil. Soc. 6 [O. 2]: 11, pl. 5, fig. 11.
Type(s): HOLOTYPE USNM_84277, near New Orleans, La.

Unio turgidus — Lea (1838), Hanley (1843), Reeve (1864), B.H. Wright (1888), Pilsbry & Rhoads (1896)
Margarita (Unio) turgidus — Lea (1836), Lea (1838)
Margaron (Unio) turgidus — Lea (1852), Lea (1870)
Unio mortoni — Conrad (1853)
Unio (Quadrula) turgidus — Mörch (1853)
Unio (Rotundaria) turgidus — Adams & Adams (1857)
Unio turgidis — Singley (1893)
Unio pustulosus — Call (1895)
Quadrula (sec. Theliderma) mortoni — Simpson (1900), Simpson (1914)
Quadrula (Bullata) pustulosa-mortoni — Frierson (1927)
Quadrula (Pustulosa) pustulosa mortoni — Haas (1969)
Quadrula mortoni — Vidrine (1993), Howells et al. (1996)
Quadrula pustulosa — Parmalee & Bogan (1998), Williams et al. (2008)
Cyclonaias pustulosa — Johnson et al. (2018), Lopes-Lima et al. (2019)

Unio nodiferus Conrad, 1841
Conrad, 1841, Proc. Acad. Nat. Sci. 1: 19.
Conrad, 1850, Jour. Acad. Nat. Sci. 1: 277, pl. 38, figs. 4, 8.
Conrad, 1853, Proc. Acad. Nat. Sci. 6: 253.
Type(s): LECTOTYPE ANSP_10285, Jackson, La.

Unio nodiferus — Conrad (1841), Conrad (1853)
Margaron (Unio) schoolcraftensis — Lea (1852)
Unio schoolcraftensis — Reeve (1864)
Margaron (Unio) schoolcraftii — Lea (1870)
Unio nodiferous — Singley (1893)
Quadrula (sec. Theliderma) nodifera — Simpson (1900), Simpson (1914)
Quadrula (Bullata) pustulosa-mortoni — Frierson (1927)
Quadrula (Pustulosa) pustulosa mortoni — Haas (1969)
Quadrula pustulosa — Burch (1975)
Quadrula mortoni — Vidrine (1993), Howells et al. (1996)
Cyclonaias pustulosa — Johnson et al. (2018), Lopes-Lima et al. (2019)

Unio postulosus Lapham, 1860
‘Lea’ Lapham, 1860, Proc. Acad. Nat. Sci. 12: 155.
(error for Unio pustulosus Lea, 1831)

Unio postulosus — Lapham (1860)

Unio sphaericus Lea, 1868
Lea, 1868, Proc. Acad. Nat. Sci. 20: 145 [as ‘sphæricus’].
Lea, 1868, Jour. Acad. Nat. Sci. 6 [O. 12]: 319 [79], pl. 51, fig. 132 [as ‘sphæricus’].
Type(s): FIGURED SYNTYPE USNM_84271, Pearl River, Jackson, Miss.

Unio sphaericus — Lea (1868), Singley (1893)
Margaron (Unio) spæricus — Lea (1870)
Unio sphæricus — B.H. Wright (1888), Pilsbry & Rhoads (1896)
Unio pustulosus — Call (1895)
Quadrula (sec. Theliderma) sphaerica — Simpson (1900), Simpson (1914)
Quadrula sphaerica — Ortmann (1912)
Quadrula pustulosa — Ortmann (1918), Burch (1975)
Quadrula (Bullata) pustulosa-refulgens — Frierson (1927)
Quadrula mortoni — Vidrine (1993), Howells et al. (1996)
Cyclonaias pustulosa — Johnson et al. (2018), Lopes-Lima et al. (2019)

Margaron (Unio) spaericus Lea, 1870
‘Lea’ Lea, 1870, Synopsis Fam. Naiades: 33 [as ‘spæricus’].
(error for Unio sphaericus Lea, 1868)

Margaron (Unio) spaericus — Lea (1870)

Unio schoolcraftii Lea, 1870
‘Lea’ of Authors.
Lea, 1870, Synopsis Fam. Naiades: 33.
B.H. Wright, 1888, Check List N. Amer. Unionidae etc.: [5].
(error for Unio schoolcraftensis Lea, 1834)

Unio schoolcraftii — Lea (1870)
Unio pustulosus — Call (1895)
Quadrula (sec. Theliderma) pustulosa — Simpson (1900)
Quadrula (sec. Theliderma) pustulosa var. schoolcraftensis — Simpson (1914)
Quadrula (Bullata) pustulosa-prasina — Frierson (1927)
Quadrula (Pustulosa) pustulosa prasina — Haas (1969)
Quadrula pustulosa — Vidrine (1993), Parmalee & Bogan (1998), Williams et al. (2008)
Quadrula pustulosa pustulosa — Watters et al. (2009)
Cyclonaias pustulosa — Bieler (2021)

Unio cahabensis Lea, 1871
Lea, 1871, Proc. Acad. Nat. Sci. 23: 190.
Lea, 1874, Jour. Acad. Nat. Sci. 8 [O. 13]: 17 [21], pl. 5, fig. 14.
Type(s): FIGURED SYNTYPE USNM_84204, Cahaba R., Ala.

Unio cahabensis — Lea (1871), B.H. Wright (1888)
Unio pustulosus — Call (1895)
Quadrula (sec. Theliderma) cahabensis — Simpson (1900), Simpson (1914)
Quadrula (Bullata) pustulosa-vallata — Frierson (1927)
Quadrula (Pustulosa) pustulosa vallata — Haas (1969)
Quadrula pustulosa — Burch (1975)
Quadrula asperata — Williams et al. (2008)
Cyclonaias pustulosa — Lopes-Lima et al. (2019)

Unio nodiferous Singley, 1893
‘Conrad’ Singley, 1893, Geol. Surv. Texas (1892): 319.
(error for Unio nodiferus Conrad, 1841)

Unio nodiferous — Singley (1893)

Unio turgidis Singley, 1893
‘Lea’ Singley, 1893, Geol. Surv. Texas (1892): 319.
(error for Unio turgidus Lea, 1838)

Unio turgidis — Singley (1893)

Unio pustulosus Call, 1895
‘Lea’ Call, 1895, Trans. Acad. Sci. St. Louis 7: 42, pl. 15, figs. 3-4.
(reference to Unio pustulosus Lea, 1831)

Unio pustulosus — Call (1895)
Quadrula (sec. Theliderma) refulgens — Simpson (1900), Simpson (1914)

Quadrula pustulosa asperatus Utterback, 1915
‘Lea’ Utterback, 1915, Amer. Midl. Nat. 4(4): 134, pl. 17, figs. 42a-b.
(misidentified reference to Unio asperatus Lea, 1861)

Quadrula pustulosa asperatus — Utterback (1915)
Quadrula pustulosa — Parmalee & Bogan (1998)
Quadrula pustulosa pustulosa — Watters et al. (2009)

Unio petrinus Frierson, 1927
‘Call (Mss)’ Frierson, 1927, Check List N. Amer. Naiades: 49.
(intro'd as syn of Unio aureus Lea, 1859)

Unio petrinus — Frierson (1927)
Quadrula (Bullata) aureus — Frierson (1927)
Cyclonaias pustulosa — Johnson et al. (2018)

Taxonomic History (since 1900)

Obovaria retusa — Simpson (1900), Simpson (1914)
Quadrula (sec. Theliderma) pustulosa — Simpson (1900), Simpson (1914)
Unio (s.s.) pustulosus — Call (1900)
Quadrula (sec. Theliderma) mortoni — Simpson (1900), Simpson (1914)
Quadrula (sec. Theliderma) nodifera — Simpson (1900), Simpson (1914)
Quadrula (sec. Theliderma) pustulosa var. pernodosa — Simpson (1900), Simpson (1914)
Quadrula (sec. Theliderma) aurea — Simpson (1900), Simpson (1914)
Quadrula (sec. Theliderma) houstonensis — Simpson (1900), Simpson (1914)
Quadrula (sec. Theliderma) paupercula — Simpson (1900), Simpson (1914)
Quadrula (sec. Theliderma) refulgens — Simpson (1900), Simpson (1914)
Quadrula (sec. Theliderma) sphaerica — Simpson (1900), Simpson (1914)
Quadrula (sec. Theliderma) vallata — Simpson (1900), Simpson (1914)
Quadrula (sec. Theliderma) cahabensis — Simpson (1900), Simpson (1914)
Quadrula pernodosa — Ihering (1901)
Quadrula pustulosa — Ortmann (1909), Ortmann (1909), Ortmann (1909), Ortmann (1910), ..., Inoue et al. (2018), Marshall et al. (2018), Gasparini et al. (2020), Klymus et al. (2020)
Quadrula mortoni — Ortmann (1912), Vidrine (1993), Howells et al. (1996), Serb et al. (2003), Bogan (2010), Pfeiffer et al. (2016), Marshall et al. (2018)
Quadrula refulgens — Ortmann (1912), Turgeon et al. (1988), Williams et al. (1993), Vidrine (1993), ..., Haag (2012), ITIS (2012), Szumowski et al. (2012), Inoue et al. (2018)
Quadrula sphaerica — Ortmann (1912)
Unio pustulosus — de Gregorio (1914)
Quadrula (sec. Theliderma) pustulosa var. schoolcraftensis — Simpson (1914)
Quadrula pustulosa pernodosa — Vanatta (1915)
Quadrula pustulosa schoolcraftensis — Utterback (1915)
Quadrula pustulosa asperatus — Utterback (1915)
Quadrula pustulosa prasina — Ortmann & Walker (1922)
Quadrula (Bullata) pustulosa — Frierson (1927)
Quadrula (Bullata) pustulosa-prasina — Frierson (1927)
Quadrula (Bullata) pustulosa-mortoni — Frierson (1927)
Quadrula (Bullata) nodulata — Frierson (1927)
Quadrula (Bullata) pustulosa-asperata — Frierson (1927)
Quadrula (Bullata) aureus — Frierson (1927)
Quadrula (Bullata) houstonensis — Frierson (1927)
Quadrula (Bullata) pustulosa-paupercula — Frierson (1927)
Quadrula (Bullata) pustulosa-refulgens — Frierson (1927)
Quadrula (Bullata) pustulosa-vallata — Frierson (1927)
Unio petrinus — Frierson (1927)
Quadrula (s.s.) pustulosus — F.C. Baker (1928)
Quadrula (s.s.) pustulosus prasina — F.C. Baker (1928)
Quadrula (Pustulosa) pustulosa pustulosa — Haas (1969)
Quadrula bullata — Morrison (1969)
Pleurobema premorsa — Morrison (1969)
Quadrula (Pustulosa) pustulosa prasina — Haas (1969)
Quadrula (Pustulosa) pustulosa mortoni — Haas (1969)
Quadrula (Pustulosa) pustulosa pernodosa — Haas (1969)
Quadrula (Pustulosa) aurea — Haas (1969)
Quadrula (Pustulosa) pustulosa paupercula — Haas (1969)
Quadrula (Pustulosa) pustulosa refulgens — Haas (1969)
Quadrula (Pustulosa) pustulosa vallata — Haas (1969)
Obliquaria retusa — Burch (1975)
Quadrula aurea — Burch (1975), Turgeon et al. (1988), Williams et al. (1993), Howells et al. (1996), Turgeon et al. (1998), Serb et al. (2003), Bogan (2010), Haag (2012), ITIS (2012), Szumowski et al. (2012)
Quadrula quadrula — R.I. Johnson (1980)
Quadrula pustulosa pustulosa — Turgeon et al. (1988), Williams et al. (1993), Turgeon et al. (1998), Lydeard et al. (1999), Watters et al. (2009), ITIS (2012)
Quadrula pustulosa mortoni — Turgeon et al. (1988), Williams et al. (1993), Turgeon et al. (1998), ITIS (2012), Szumowski et al. (2012)
Quadrula houstonensis — Turgeon et al. (1988), Williams et al. (1993), Howells et al. (1996), Turgeon et al. (1998), Bogan (2010), Breton et al. (2011), Haag (2012), ITIS (2012), Pfeiffer et al. (2016), Chase et al. (2018)
Pleurobema riddelli — Vidrine (1993)
Amphinaias houstonensis — IUCN (1996), Graf & Cummings (2007), Cummings & Graf (2010)
Amphinaias refulgens — IUCN (1996), Graf & Cummings (2007), Cummings & Graf (2010)
Amphinaias pustulosa — Graf & Cummings (2007), Cummings & Graf (2010), IUCN (2011), Roe & Boyer (2015)
Amphinaias aurea — Graf & Cummings (2007), Cummings & Graf (2010)
Quadrula asperata — Williams et al. (2008)
Rotundaria pustulosa — Campbell & Lydeard (2012)
Rotundaria refulgens — Campbell & Lydeard (2012), Campbell & Lydeard (2012)
Cyclonaias pustulosa — Williams et al. (2017), Johnson et al. (2018), Lopes-Lima et al. (2019), Graf & Cummings (2021), FMCS (2021), Bieler (2021), Hopper et al. (2023), Freeman et al. (2025)
Cyclonaias mortoni — Williams et al. (2017)
Cyclonaias aurea — Williams et al. (2017)
Cyclonaias houstonensis — Williams et al. (2017)
Cyclonaias refulgens — Williams et al. (2017)
Cyclonaias (Quadrula) refulgens — Porto-Hannes et al. (2021)
Pustulosa pustulosa — Neemuchwala et al. (2023)