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Page last updated
Mon 06 May 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Margaritiferidae | Margaritiferinae

genus Margaritifera Schumacher, 1816: cladomics
     23 valid species

Cladomics Summary

number of species = 23

analyzed in 193 cladograms in 111 publications
maximum number of terminals in an analysis = 8, mean = 2.31

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 96
maximum number of included terminals = 8, mean = 3.62
proportion of cladograms that support basic monophyly = 0.95


Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ tree


Huff et al., 2004 [Publ 2697], fig. 3

iT tT bi riS oT roT roS iD
8 10 2 8 4 2 2 0.0

*** Top Monophyly Cladogram ***

  +-------- Margaritifera_margaritifera **  
  |                                      
  |  +----- Cumberlandia_monodonta       
--+  |                                   
  |  |  +-- Pseudunio_auricularius       
  |  |  |                                
  +--+  +-- Margaritifera_dahurica **       
     |  |                                
     |  +-- Margaritifera_middendorffi[1] **
     |  |                                
     |  +-- Margaritifera_middendorffi[2] **
     +--+                                
        +-- Margaritifera_falcata[1] **     
        |                                
        +-- Margaritifera_falcata[2] **     
        |                                
        +-- Margaritifera_falcata[3] **     
        |                                
        +-- Margaritifera_falcata[4] **     

Lyubas et al., 2019 [Publ 3963], fig. 10

iT tT bi riS oT roT roS iD
7 7 5 7 9 6 2 11.0

*** Top Sister-Group Cladogram ***

           +-------- Cumberlandia_monodonta     
           |                                    
  +--------+  +----- Pseudunio_marocanus        
  |        |  |                                 
  |        +--+  +-- Pseudunio_auricularius     
  |           |  |                              
  |           |  +-- Pseudunio_flabellata       
  |           +--+                              
--+              +-- Pseudunio_flabellatiformis 
  |              |                              
  |              +-- Pseudunio_homsensis        
  |                                             
  |              +-- Margaritifera_margaritifera **
  |  +-----------+                              
  |  |           +-- Margaritifera_dahurica **     
  +--+                                          
     |  +----------- Margaritifera_falcata **      
     +--+                                       
        |  +-------- Margaritifera_laevis **       
        +--+                                    
           |  +----- Margaritifera_middendorffi ** 
           +--+                                 
              |  +-- Margaritifera_marrianae **    
              +--+                              
                 +-- Margaritifera_hembeli **      

Huff et al., 2004 [Publ 2697], fig. 2

iT tT bi riS oT roT roS iD
7 9 3 7 7 2 2 9.0
  +-------- Margaritifera_dahurica **        
  |                                       
  +-------- Margaritifera_margaritifera[1] **
  |                                       
  +-------- Margaritifera_margaritifera[2] **
  |                                       
--+-------- Margaritifera_margaritifera[3] **
  |                                       
  |     +-- Pseudunio_auricularius        
  +-----+                                 
  |     +-- Cumberlandia_monodonta        
  |                                       
  |  +----- Margaritifera_falcata **         
  +--+                                    
     |  +-- Margaritifera_middendorffi **    
     +--+                                 
        +-- Margaritifera_marrianae **       

Bolotov et al., 2015 [Publ 3646], fig. 2A

iT tT bi riS oT roT roS iD
7 9 5 7 3 2 2 11.0
  +----------- Margaritifera_margaritifera **
  |                                       
  |     +----- Margaritifera_dahurica[1] **  
  |     |                                 
  +-----+----- Margaritifera_dahurica[2] **  
--+     |                                 
  |     |  +-- Pseudunio_auricularius     
  |     +--+                              
  |        +-- Cumberlandia_monodonta     
  |                                       
  |  +-------- Margaritifera_falcata **      
  +--+                                    
     |  +----- Margaritifera_laevis **       
     +--+                                 
        |  +-- Margaritifera_middendorffi ** 
        +--+                              
           +-- Margaritifera_marrianae **    

Araujo et al., 2016 [Publ 3746], fig. 1,2b

iT tT bi riS oT roT roS iD
7 7 5 7 6 3 2 11.0
           +----- Cumberlandia_monodonta     
  +--------+                                 
  |        |  +-- Pseudunio_marocanus        
  |        +--+                              
  |           +-- Pseudunio_auricularius     
--+                                          
  |           +-- Margaritifera_dahurica **     
  |  +--------+                              
  |  |        +-- Margaritifera_margaritifera **
  +--+                                       
     |  +-------- Margaritifera_falcata **      
     |  |                                    
     +--+     +-- Margaritifera_middendorffi ** 
        |  +--+                              
        |  |  +-- Margaritifera_laevis **       
        +--+                                 
           |  +-- Margaritifera_marrianae **    
           +--+                              
              +-- Margaritifera_hembeli **      

Araujo et al., 2016 [Publ 3746], fig. 2a

iT tT bi riS oT roT roS iD
7 7 5 7 4 3 2 11.0
           +-- Margaritifera_dahurica **     
  +--------+                              
  |        +-- Margaritifera_margaritifera **
--+                                       
  |  +-------- Margaritifera_falcata **      
  |  |                                    
  +--+     +-- Margaritifera_laevis **       
     |  +--+                              
     |  |  +-- Margaritifera_middendorffi ** 
     +--+                                 
        |  +-- Margaritifera_marrianae **    
        +--+                              
           +-- Margaritifera_hembeli **      

Huang et al., 2018 [Publ 3828], fig. 3, 5

iT tT bi riS oT roT roS iD
7 7 5 7 22 3 2 11.0
              +----- Cumberlandia_monodonta     
  +-----------+                                 
  |           |  +-- Pseudunio_auricularius     
  |           +--+                              
  |              +-- Pseudunio_marocanus        
--+                                             
  |              +-- Margaritifera_margaritifera **
  |  +-----------+                              
  |  |           +-- Margaritifera_dahurica **     
  +--+                                          
     |  +----------- Margaritifera_falcata **      
     +--+                                       
        |  +-------- Margaritifera_laevis **       
        +--+                                    
           |  +----- Margaritifera_hembeli **      
           +--+                                 
              |  +-- Margaritifera_middendorffi ** 
              +--+                              
                 +-- Margaritifera_marrianae **    

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

iT tT bi riS oT roT roS iD
7 7 5 7 21 4 2 11.0
        +-------- Cumberlandia_monodonta     
  +-----+                                    
  |     |  +----- Pseudunio_marocanus        
  |     +--+                                 
  |        |  +-- Pseudunio_auricularius     
  |        +--+                              
--+           +-- Pseudunio_homsensis        
  |                                          
  |           +-- Margaritifera_dahurica **     
  |  +--------+                              
  |  |        +-- Margaritifera_margaritifera **
  +--+                                       
     |  +-------- Margaritifera_falcata **      
     |  |                                    
     +--+     +-- Margaritifera_middendorffi ** 
        |  +--+                              
        |  |  +-- Margaritifera_laevis **       
        +--+                                 
           |  +-- Margaritifera_hembeli **      
           +--+                              
              +-- Margaritifera_marrianae **    

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

iT tT bi riS oT roT roS iD
7 7 5 7 21 4 2 11.0
           +-------- Cumberlandia_monodonta     
  +--------+                                    
  |        |  +----- Pseudunio_marocanus        
  |        +--+                                 
  |           |  +-- Pseudunio_auricularius     
  |           +--+                              
--+              +-- Pseudunio_homsensis        
  |                                             
  |              +-- Margaritifera_margaritifera **
  |  +-----------+                              
  |  |           +-- Margaritifera_dahurica **     
  +--+                                          
     |  +----------- Margaritifera_falcata **      
     +--+                                       
        |  +-------- Margaritifera_laevis **       
        +--+                                    
           |  +----- Margaritifera_middendorffi ** 
           +--+                                 
              |  +-- Margaritifera_hembeli **      
              +--+                              
                 +-- Margaritifera_marrianae **    

Huang et al., 2019 [Publ 3843], fig. 2

iT tT bi riS oT roT roS iD
7 7 4 7 197 4 2 11.0
  +----------- Cumberlandia_monodonta     
  |                                       
  |     +----- Pseudunio_marocanus        
  +-----+                                 
  |     |  +-- Pseudunio_auricularius     
  |     +--+                              
--+        +-- Pseudunio_homsensis        
  |                                       
  |        +-- Margaritifera_dahurica **     
  |  +-----+                              
  |  |     +-- Margaritifera_margaritifera **
  |  |                                    
  +--+  +----- Margaritifera_laevis **       
     |  |                                 
     |  |  +-- Margaritifera_falcata **      
     +--+--+                              
        |  +-- Margaritifera_middendorffi ** 
        |                                 
        |  +-- Margaritifera_hembeli **      
        +--+                              
           +-- Margaritifera_marrianae **    

Huang et al., 2019 [Publ 3843], fig. S3

iT tT bi riS oT roT roS iD
7 7 5 7 194 4 2 11.0
        +-------- Cumberlandia_monodonta     
  +-----+                                    
  |     |  +----- Pseudunio_marocanus        
  |     +--+                                 
  |        |  +-- Pseudunio_auricularius     
  |        +--+                              
--+           +-- Pseudunio_homsensis        
  |                                          
  |           +-- Margaritifera_dahurica **     
  |  +--------+                              
  |  |        +-- Margaritifera_margaritifera **
  +--+                                       
     |  +-------- Margaritifera_falcata **      
     |  |                                    
     +--+     +-- Margaritifera_hembeli **      
        |  +--+                              
        |  |  +-- Margaritifera_marrianae **    
        +--+                                 
           |  +-- Margaritifera_laevis **       
           +--+                              
              +-- Margaritifera_middendorffi ** 

Huang et al., 2019 [Publ 3843], fig. S4

iT tT bi riS oT roT roS iD
7 7 3 7 195 4 2 10.0
  +-------------- Cumberlandia_monodonta     
  |                                          
  |        +----- Pseudunio_marocanus        
--+  +-----+                                 
  |  |     |  +-- Pseudunio_auricularius     
  |  |     +--+                              
  |  |        +-- Pseudunio_homsensis        
  +--+                                       
     |  +-------- Margaritifera_falcata **      
     |  |                                    
     |  |     +-- Margaritifera_dahurica **     
     |  +-----+                              
     +--+     +-- Margaritifera_margaritifera **
        |                                    
        |  +----- Margaritifera_laevis **       
        |  |                                 
        +--+----- Margaritifera_middendorffi ** 
           |                                 
           |  +-- Margaritifera_hembeli **      
           +--+                              
              +-- Margaritifera_marrianae **    

Wu et al., 2020 [Publ 4121], fig. 2

iT tT bi riS oT roT roS iD
7 7 5 7 27 3 2 11.0
           +-- Margaritifera_dahurica **     
  +--------+                              
  |        +-- Margaritifera_margaritifera **
--+                                       
  |  +-------- Margaritifera_falcata **      
  |  |                                    
  +--+     +-- Margaritifera_hembeli **      
     |  +--+                              
     |  |  +-- Margaritifera_marrianae **    
     +--+                                 
        |  +-- Margaritifera_laevis **       
        +--+                              
           +-- Margaritifera_middendorffi ** 

Wu et al., 2020 [Publ 4121], fig. S1

iT tT bi riS oT roT roS iD
7 7 5 7 27 3 2 11.0
           +----- Cumberlandia_monodonta     
  +--------+                                 
  |        |  +-- Pseudunio_auricularius     
  |        +--+                              
  |           +-- Pseudunio_marocanus        
--+                                          
  |           +-- Margaritifera_dahurica **     
  |  +--------+                              
  |  |        +-- Margaritifera_margaritifera **
  +--+                                       
     |  +-------- Margaritifera_falcata **      
     |  |                                    
     +--+     +-- Margaritifera_hembeli **      
        |  +--+                              
        |  |  +-- Margaritifera_marrianae **    
        +--+                                 
           |  +-- Margaritifera_laevis **       
           +--+                              
              +-- Margaritifera_middendorffi ** 

Sano et al., 2020 [Publ 4138], fig. 2

iT tT bi riS oT roT roS iD
7 7 5 7 66 3 2 11.0
              +-- Margaritifera_dahurica **     
  +-----------+                              
  |           +-- Margaritifera_margaritifera **
--+                                          
  |  +----------- Margaritifera_falcata **      
  +--+                                       
     |  +-------- Margaritifera_middendorffi ** 
     +--+                                    
        |  +----- Margaritifera_laevis **       
        +--+                                 
           |  +-- Margaritifera_hembeli **      
           +--+                              
              +-- Margaritifera_marrianae **    

Sano et al., 2020 [Publ 4138], fig. 3

iT tT bi riS oT roT roS iD
7 7 5 7 65 3 2 11.0
              +-- Margaritifera_margaritifera **
  +-----------+                              
  |           +-- Margaritifera_dahurica **     
--+                                          
  |  +----------- Margaritifera_falcata **      
  +--+                                       
     |  +-------- Margaritifera_middendorffi ** 
     +--+                                    
        |  +----- Margaritifera_laevis **       
        +--+                                 
           |  +-- Margaritifera_hembeli **      
           +--+                              
              +-- Margaritifera_marrianae **    

Sano et al., 2020 [Publ 4138], fig. 4

iT tT bi riS oT roT roS iD
7 7 5 7 65 3 2 11.0
           +-- Margaritifera_margaritifera **
  +--------+                              
  |        +-- Margaritifera_dahurica **     
--+                                       
  |  +-------- Margaritifera_falcata **      
  |  |                                    
  +--+     +-- Margaritifera_laevis **       
     |  +--+                              
     |  |  +-- Margaritifera_middendorffi ** 
     +--+                                 
        |  +-- Margaritifera_hembeli **      
        +--+                              
           +-- Margaritifera_marrianae **    

Takeuchi et al., 2015 [Publ 3661], fig. 3

iT tT bi riS oT roT roS iD
6 6 4 6 5 3 2 11.0
  +----------------- Pseudunio_auricularius[1]  
  |                                             
  +----------------- Pseudunio_auricularius[2]  
--+                                             
  |  +-------------- Cumberlandia_monodonta     
  |  |                                          
  +--+           +-- Margaritifera_margaritifera **
     |  +--------+                              
     |  |        +-- Margaritifera_dahurica **     
     +--+                                       
        |  +-------- Margaritifera_falcata **      
        +--+                                    
           |  +----- Margaritifera_laevis **       
           +--+                                 
              |  +-- Margaritifera_middendorffi ** 
              +--+                              
                 +-- Margaritifera_marrianae **    

Bolotov et al., 2016 [Publ 3743], fig. 2

iT tT bi riS oT roT roS iD
6 6 4 6 24 3 2 11.0
           +-- Margaritifera_dahurica **     
  +--------+                              
  |        +-- Margaritifera_margaritifera **
--+                                       
  |  +-------- Margaritifera_falcata **      
  +--+                                    
     |  +----- Margaritifera_laevis **       
     +--+                                 
        |  +-- Margaritifera_middendorffi ** 
        +--+                              
           +-- Margaritifera_marrianae **    

Bolotov et al., 2016 [Publ 3743], fig. 4

iT tT bi riS oT roT roS iD
6 6 4 6 24 3 2 11.0
           +-- Margaritifera_margaritifera **
  +--------+                              
  |        +-- Margaritifera_dahurica **     
--+                                       
  |  +-------- Margaritifera_falcata **      
  +--+                                    
     |  +----- Margaritifera_laevis **       
     +--+                                 
        |  +-- Margaritifera_marrianae **    
        +--+                              
           +-- Margaritifera_middendorffi ** 

Bolotov et al., 2017 [Publ 3761], fig. S1

iT tT bi riS oT roT roS iD
6 6 4 6 135 3 2 11.0
           +-- Margaritifera_dahurica **     
  +--------+                              
  |        +-- Margaritifera_margaritifera **
--+                                       
  |  +-------- Margaritifera_falcata **      
  +--+                                    
     |  +----- Margaritifera_middendorffi ** 
     +--+                                 
        |  +-- Margaritifera_laevis **       
        +--+                              
           +-- Margaritifera_marrianae **    

Bolotov et al., 2017 [Publ 3761], fig. S2

iT tT bi riS oT roT roS iD
6 6 4 6 135 3 2 11.0
           +-- Margaritifera_dahurica **     
  +--------+                              
  |        +-- Margaritifera_margaritifera **
--+                                       
  |  +-------- Margaritifera_falcata **      
  +--+                                    
     |  +----- Margaritifera_middendorffi ** 
     +--+                                 
        |  +-- Margaritifera_laevis **       
        +--+                              
           +-- Margaritifera_marrianae **    

Vikhrev et al., 2018 [Publ 3812], fig. 3

iT tT bi riS oT roT roS iD
6 6 4 6 7 4 2 11.0
        +-------- Cumberlandia_monodonta     
  +-----+                                    
  |     |  +----- Pseudunio_marocanus        
  |     +--+                                 
  |        |  +-- Pseudunio_auricularius     
  |        +--+                              
--+           +-- Pseudunio_homsensis        
  |                                          
  |           +-- Margaritifera_margaritifera **
  |  +--------+                              
  |  |        +-- Margaritifera_dahurica **     
  +--+                                       
     |  +-------- Margaritifera_falcata **      
     +--+                                    
        |  +----- Margaritifera_laevis **       
        +--+                                 
           |  +-- Margaritifera_middendorffi ** 
           +--+                              
              +-- Margaritifera_marrianae **    

Jeratthitikul et al., 2022 [Publ 4260], fig. S1, 1

iT tT bi riS oT roT roS iD
6 6 4 6 70 3 2 11.0
              +-- Pseudunio_auricularius     
  +-----------+                              
  |           +-- Pseudunio_marocanus        
  |                                          
--+           +-- Margaritifera_margaritifera **
  |  +--------+                              
  |  |        +-- Margaritifera_dahurica **     
  +--+                                       
     |  +-------- Margaritifera_falcata **      
     +--+                                    
        |  +----- Margaritifera_middendorffi ** 
        +--+                                 
           |  +-- Margaritifera_laevis **       
           +--+                              
              +-- Margaritifera_marrianae **    

Jeratthitikul et al., 2022 [Publ 4260], fig. S2, 2

iT tT bi riS oT roT roS iD
6 6 4 6 70 3 2 11.0
           +-- Margaritifera_dahurica **     
  +--------+                              
  |        +-- Margaritifera_margaritifera **
--+                                       
  |  +-------- Margaritifera_falcata **      
  +--+                                    
     |  +----- Margaritifera_marrianae **    
     +--+                                 
        |  +-- Margaritifera_middendorffi ** 
        +--+                              
           +-- Margaritifera_laevis **       

Huff et al., 2004 [Publ 2697], fig. 4

iT tT bi riS oT roT roS iD
5 5 3 5 6 2 2 10.0
  +----------- Pseudunio_auricularius     
  |                                       
  +----------- Cumberlandia_monodonta     
--+                                       
  |        +-- Margaritifera_dahurica **     
  |  +-----+                              
  |  |     +-- Margaritifera_margaritifera **
  +--+                                    
     |  +----- Margaritifera_falcata **      
     +--+                                 
        |  +-- Margaritifera_middendorffi ** 
        +--+                              
           +-- Margaritifera_marrianae **    

Huff et al., 2004 [Publ 2697], fig. 5-6

iT tT bi riS oT roT roS iD
5 5 3 5 7 2 2 10.0
           +-- Pseudunio_auricularius     
  +--------+                              
  |        +-- Cumberlandia_monodonta     
  |                                       
--+        +-- Margaritifera_dahurica **     
  |  +-----+                              
  |  |     +-- Margaritifera_margaritifera **
  +--+                                    
     |  +----- Margaritifera_falcata **      
     +--+                                 
        |  +-- Margaritifera_middendorffi ** 
        +--+                              
           +-- Margaritifera_marrianae **    

Inoue et al., 2013 [Publ 3579], fig. 3

iT tT bi riS oT roT roS iD
5 5 3 5 4 3 2 10.0
  +-------------- Cumberlandia_monodonta     
  |                                          
  |           +-- Pseudunio_auricularius     
--+  +--------+                              
  |  |        +-- Pseudunio_marocanus        
  |  |                                       
  +--+        +-- Margaritifera_dahurica **     
     |  +-----+                              
     |  |     +-- Margaritifera_margaritifera **
     +--+                                    
        |  +----- Margaritifera_falcata **      
        +--+                                 
           |  +-- Margaritifera_laevis **       
           +--+                              
              +-- Margaritifera_marrianae **    

Bolotov et al., 2015 [Publ 3646], fig. 2B

iT tT bi riS oT roT roS iD
5 5 3 5 4 3 2 10.0
  +-------------- Cumberlandia_monodonta     
  |                                          
  |           +-- Pseudunio_auricularius     
--+  +--------+                              
  |  |        +-- Pseudunio_marocanus        
  |  |                                       
  +--+        +-- Margaritifera_dahurica **     
     |  +-----+                              
     |  |     +-- Margaritifera_margaritifera **
     +--+                                    
        |  +----- Margaritifera_falcata **      
        +--+                                 
           |  +-- Margaritifera_middendorffi ** 
           +--+                              
              +-- Margaritifera_laevis **       

Takeuchi et al., 2015 [Publ 3661], fig. 2

iT tT bi riS oT roT roS iD
5 5 3 5 5 3 2 10.0
  +-------------- Pseudunio_auricularius     
  |                                          
--+  +----------- Cumberlandia_monodonta     
  |  |                                       
  +--+        +-- Margaritifera_dahurica **     
     |  +-----+                              
     |  |     +-- Margaritifera_margaritifera **
     +--+                                    
        |  +----- Margaritifera_falcata **      
        +--+                                 
           |  +-- Margaritifera_middendorffi ** 
           +--+                              
              +-- Margaritifera_laevis **       

Araujo et al., 2009 [Publ 2395], fig. 3

iT tT bi riS oT roT roS iD
4 4 2 4 4 3 2 9.0
  +----------- Cumberlandia_monodonta     
  |                                       
  |        +-- Pseudunio_marocanus        
--+  +-----+                              
  |  |     +-- Pseudunio_auricularius     
  |  |                                    
  +--+     +-- Margaritifera_margaritifera **
     |  +--+                              
     |  |  +-- Margaritifera_dahurica **     
     +--+                                 
        |  +-- Margaritifera_falcata **      
        +--+                              
           +-- Margaritifera_middendorffi ** 

Walker et al., 2006 [Publ 2666], fig. 6

iT tT bi riS oT roT roS iD
4 4 2 4 34 2 2 9.0
  +----------- Cumberlandia_monodonta     
  |                                       
--+  +-------- Pseudunio_auricularius     
  |  |                                    
  +--+     +-- Margaritifera_margaritifera **
     |  +--+                              
     |  |  +-- Margaritifera_dahurica **     
     +--+                                 
        |  +-- Margaritifera_falcata **      
        +--+                              
           +-- Margaritifera_middendorffi ** 

Sano et al., 2017 [Publ 3799], fig. 1

iT tT bi riS oT roT roS iD
4 4 2 4 23 0 0 10.0
  +-------- Margaritifera_laevis[2] **   
--+                                   
  |  +----- Margaritifera_laevis[1] **   
  +--+                                
     |  +-- Margaritifera_middendorffi **
     +--+                             
        +-- Margaritifera_laevis[3] **   

Bolotov et al., 2020 [Publ 4084], fig. 1

iT tT bi riS oT roT roS iD
4 4 0 4 32 0 0 9.0
  +-- Margaritifera_dahurica **     
  |                              
  +-- Margaritifera_margaritifera **
--+                              
  +-- Margaritifera_laevis **       
  |                              
  +-- Margaritifera_middendorffi ** 

Bolotov et al., 2020 [Publ 4100], fig. 1

iT tT bi riS oT roT roS iD
4 5 0 4 116 0 0 10.0
  +-- Gibbosula_laosensis        
  |                              
  +-- Margaritifera_dahurica **     
  |                              
--+-- Margaritifera_margaritifera **
  |                              
  +-- Margaritifera_laevis **       
  |                              
  +-- Margaritifera_middendorffi ** 

Walker et al., 2006 [Publ 2666], fig. 6-M

iT tT bi riS oT roT roS iD
3 3 0 3 12 1 1 9.0
  +----- Cumberlandia_monodonta     
  |                                 
--+  +-- Margaritifera_margaritifera **
  |  |                              
  +--+-- Margaritifera_dahurica[1] **  
     |                              
     +-- Margaritifera_dahurica[2] **  

Breton et al., 2011 [Publ 3376], fig. 3

iT tT bi riS oT roT roS iD
3 3 1 3 36 1 1 9.0
  +-------- Cumberlandia_monodonta     
--+                                    
  |  +----- Margaritifera_margaritifera **
  +--+                                 
     |  +-- Margaritifera_falcata **      
     +--+                              
        +-- Margaritifera_marrianae **    

Mock et al., 2013 [Publ 3598], fig. 3

iT tT bi riS oT roT roS iD
3 3 1 3 0 0 0 9.0
  +----- Margaritifera_margaritifera **
--+                                 
  |  +-- Margaritifera_falcata **      
  +--+                              
     +-- Margaritifera_laevis **       

Amaro et al., 2017 [Publ 3791], fig. 2

iT tT bi riS oT roT roS iD
3 17 15 3 15 0 0 20.0
  +-------------------- Margaritifera_margaritifera[2] **
  |                                                   
  |                 +-- Margaritifera_margaritifera[1] **
  |  +--------------+                                 
--+  |              +-- Margaritifera_falcata **         
  |  |                                                
  |  |              +-- Lasmigona_costata             
  |  |        +-----+                                 
  |  |        |     +-- Platynaias_compressa          
  +--+  +-----+                                       
     |  |     |  +----- Pyganodon_grandis             
     |  |     +--+                                    
     |  |        |  +-- Utterbackia_peninsularis      
     |  |        +--+                                 
     |  |           +-- Utterbackia_imbecillis        
     +--+                                             
        |        +----- Fusconaia_flava               
        |  +-----+                                    
        |  |     |  +-- Pronodularia_japanensis       
        |  |     +--+                                 
        |  |        +-- Sinosolenaia_carinata         
        +--+                                          
           |     +----- Toxolasma_parvum              
           |  +--+                                    
           |  |  |  +-- Venustaconcha_ellipsiformis   
           |  |  +--+                                 
           +--+     +-- Lampsilis_siliquoidea         
              |                                       
              |  +----- Quadrula_quadrula[2]          
              +--+                                    
                 |  +-- Quadrula_quadrula[1]          
                 +--+                                 
                    +-- Quadrula_nobilis              

Huang et al., 2018 [Publ 3828], fig. 6

iT tT bi riS oT roT roS iD
3 3 1 3 40 2 2 9.0
  +----------- Cumberlandia_monodonta     
--+                                       
  |  +-------- Pseudunio_marocanus        
  +--+                                    
     |  +----- Margaritifera_falcata **      
     +--+                                 
        |  +-- Margaritifera_margaritifera **
        +--+                              
           +-- Margaritifera_dahurica **     

Huang et al., 2019 [Publ 3843], fig. 3

iT tT bi riS oT roT roS iD
3 3 1 3 54 2 2 9.0
  +----------- Cumberlandia_monodonta     
--+                                       
  |  +-------- Pseudunio_marocanus        
  +--+                                    
     |  +----- Margaritifera_falcata **      
     +--+                                 
        |  +-- Margaritifera_margaritifera **
        +--+                              
           +-- Margaritifera_dahurica **     

Chase et al., 2018 [Publ 3847], fig. 4

iT tT bi riS oT roT roS iD
3 3 1 3 36 1 1 9.0
  +-------- Cumberlandia_monodonta     
--+                                    
  |  +----- Margaritifera_falcata **      
  +--+                                 
     |  +-- Margaritifera_marrianae **    
     +--+                              
        +-- Margaritifera_margaritifera **

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1

iT tT bi riS oT roT roS iD
3 3 1 3 7 2 2 9.0
  +----------- Cumberlandia_monodonta     
--+                                       
  |  +-------- Pseudunio_marocanus        
  +--+                                    
     |  +----- Margaritifera_falcata **      
     +--+                                 
        |  +-- Margaritifera_margaritifera **
        +--+                              
           +-- Margaritifera_dahurica **     

Guerra et al., 2019 [Publ 4168], fig. 1

iT tT bi riS oT roT roS iD
3 3 1 3 38 3 2 9.0
  +----------- Cumberlandia_monodonta     
  |                                       
--+        +-- Pseudunio_marocanus        
  |  +-----+                              
  |  |     +-- Pseudunio_auricularius     
  +--+                                    
     |  +----- Margaritifera_falcata **      
     +--+                                 
        |  +-- Margaritifera_dahurica **     
        +--+                              
           +-- Margaritifera_margaritifera **

Guerra et al., 2019 [Publ 4168], fig. 2

iT tT bi riS oT roT roS iD
3 3 1 3 38 3 2 9.0
  +----------- Cumberlandia_monodonta     
  |                                       
--+        +-- Pseudunio_marocanus        
  |  +-----+                              
  |  |     +-- Pseudunio_auricularius     
  +--+                                    
     |  +----- Margaritifera_falcata **      
     +--+                                 
        |  +-- Margaritifera_dahurica **     
        +--+                              
           +-- Margaritifera_margaritifera **

Wu et al., 2022 [Publ 4238], fig. 3

iT tT bi riS oT roT roS iD
3 3 1 3 71 2 2 9.0
  +----------- Cumberlandia_monodonta     
--+                                       
  |  +-------- Pseudunio_marocanus        
  +--+                                    
     |  +----- Margaritifera_falcata **      
     +--+                                 
        |  +-- Margaritifera_dahurica **     
        +--+                              
           +-- Margaritifera_margaritifera **

Wu et al., 2023 [Publ 4339], fig. 4, S5-aa

iT tT bi riS oT roT roS iD
3 3 1 3 93 2 2 9.0
  +----------- Cumberlandia_monodonta     
--+                                       
  |  +-------- Pseudunio_marocanus        
  +--+                                    
     |  +----- Margaritifera_falcata **      
     +--+                                 
        |  +-- Margaritifera_dahurica **     
        +--+                              
           +-- Margaritifera_margaritifera **

Wu et al., 2023 [Publ 4339], fig. 4, S5-nt

iT tT bi riS oT roT roS iD
3 3 1 3 94 2 2 9.0
  +----------- Cumberlandia_monodonta     
--+                                       
  |  +-------- Pseudunio_marocanus        
  +--+                                    
     |  +----- Margaritifera_falcata **      
     +--+                                 
        |  +-- Margaritifera_dahurica **     
        +--+                              
           +-- Margaritifera_margaritifera **

Lydeard et al., 1996 [Publ 2656], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 25 1 1 --
  +----- Cumberlandia_monodonta     
--+                                 
  |  +-- Margaritifera_margaritifera **
  +--+                              
     +-- Margaritifera_falcata **      

Lydeard et al., 1996 [Publ 2656], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 2 26 1 1 --
  +----- Cumberlandia_monodonta     
--+                                 
  |  +-- Margaritifera_margaritifera **
  +--+                              
     +-- Margaritifera_falcata **      

Krebs et al., 2003 [Publ 2691], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 32 0 0 --
  +-- Margaritifera_margaritifera **
--+                              
  +-- Margaritifera_falcata **      

Rosenberg et al., 1994 [Publ 2719], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 17 1 1 --
  +-- Margaritifera_falcata **      
--+                              
  +-- Margaritifera_margaritifera **

Rosenberg et al., 1997 [Publ 2720], fig. 2A

iT tT bi riS oT roT roS iD
2 2 0 2 21 1 1 --
  +-- Margaritifera_falcata **      
--+                              
  +-- Margaritifera_margaritifera **

Rosenberg et al., 1997 [Publ 2720], fig. 2B

iT tT bi riS oT roT roS iD
2 2 0 2 21 1 1 --
  +-- Margaritifera_falcata **      
--+                              
  +-- Margaritifera_margaritifera **

Fonseca et al., 2016 [Publ 3775], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 30 0 0 --
  +-- Margaritifera_falcata ** 
--+                         
  +-- Margaritifera_dahurica **

Amaro et al., 2017 [Publ 3791], fig. 3A

iT tT bi riS oT roT roS iD
2 2 0 2 10 0 0 --
  +-- Margaritifera_falcata **      
--+                              
  +-- Margaritifera_margaritifera **

Amaro et al., 2017 [Publ 3791], fig. 3B

iT tT bi riS oT roT roS iD
2 2 0 2 10 0 0 --
  +-- Margaritifera_falcata **      
--+                              
  +-- Margaritifera_margaritifera **

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 31 1 1 --
  +----- Pseudunio_marocanus   
--+                            
  |  +-- Margaritifera_falcata ** 
  +--+                         
     +-- Margaritifera_dahurica **

Wen et al., 2017 [Publ 3795], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 14 0 0 --
  +-- Margaritifera_dahurica **
--+                         
  +-- Margaritifera_falcata ** 

Sano et al., 2017 [Publ 3799], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 34 0 0 --
  +-- Margaritifera_laevis **      
--+                             
  +-- Margaritifera_middendorffi **

Feng et al., 2016 [Publ 3800], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 13 0 0 --
  +-- Margaritifera_dahurica **
--+                         
  +-- Margaritifera_falcata ** 

Konopleva et al., 2017 [Publ 3807], fig. 6

iT tT bi riS oT roT roS iD
2 2 0 2 29 0 0 --
  +-- Margaritifera_dahurica **
--+                         
  +-- Margaritifera_laevis **  

Wu et al., 2018 [Publ 3836], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 51 0 0 --
  +-- Margaritifera_dahurica **
--+                         
  +-- Margaritifera_falcata ** 

Aunins et al., 2018 [Publ 3863], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 39 1 1 --
  +----- Cumberlandia_monodonta
--+                            
  |  +-- Margaritifera_falcata ** 
  +--+                         
     +-- Margaritifera_dahurica **

Wu et al., 2019 [Publ 3864], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 45 1 1 --
  +----- Cumberlandia_monodonta
--+                            
  |  +-- Margaritifera_dahurica **
  +--+                         
     +-- Margaritifera_falcata ** 

Cha et al., 2018 [Publ 3916], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 37 1 1 --
  +----- Pseudunio_marocanus   
--+                            
  |  +-- Margaritifera_falcata ** 
  +--+                         
     +-- Margaritifera_dahurica **

Froufe et al., 2019 [Publ 3936], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 43 1 1 --
  +----- Pseudunio_marocanus   
--+                            
  |  +-- Margaritifera_dahurica **
  +--+                         
     +-- Margaritifera_falcata ** 

Froufe et al., 2019 [Publ 3936], fig. 6

iT tT bi riS oT roT roS iD
2 2 0 2 43 1 1 --
  +----- Pseudunio_marocanus   
--+                            
  |  +-- Margaritifera_dahurica **
  +--+                         
     +-- Margaritifera_falcata ** 

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iT tT bi riS oT roT roS iD
2 2 0 2 55 0 0 --
  +-- Margaritifera_margaritifera **
--+                              
  +-- Margaritifera_hembeli **      

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iT tT bi riS oT roT roS iD
2 2 0 2 55 0 0 --
  +-- Margaritifera_margaritifera **
--+                              
  +-- Margaritifera_hembeli **      

Lopes-Lima et al., 2020 [Publ 4080], fig. 10

iT tT bi riS oT roT roS iD
2 2 0 2 12 0 0 --
  +-- Margaritifera_middendorffi **
--+                             
  +-- Margaritifera_laevis **      

Lopes-Lima et al., 2020 [Publ 4080], fig. 11

iT tT bi riS oT roT roS iD
2 2 0 2 10 0 0 --
  +-- Margaritifera_laevis **      
--+                             
  +-- Margaritifera_middendorffi **

Lopes-Lima et al., 2020 [Publ 4080], fig. 12

iT tT bi riS oT roT roS iD
2 2 0 2 7 0 0 --
  +-- Margaritifera_laevis **      
--+                             
  +-- Margaritifera_middendorffi **

Lopes-Lima et al., 2020 [Publ 4080], fig. 13

iT tT bi riS oT roT roS iD
2 2 0 2 8 0 0 --
  +-- Margaritifera_laevis **      
--+                             
  +-- Margaritifera_middendorffi **

Lopes-Lima et al., 2020 [Publ 4080], fig. 14

iT tT bi riS oT roT roS iD
2 2 0 2 4 0 0 --
  +-- Margaritifera_laevis **      
--+                             
  +-- Margaritifera_middendorffi **

Lopes-Lima et al., 2020 [Publ 4080], fig. 16

iT tT bi riS oT roT roS iD
2 2 0 2 7 0 0 --
  +-- Margaritifera_laevis **      
--+                             
  +-- Margaritifera_middendorffi **

Lopes-Lima et al., 2020 [Publ 4080], fig. 17

iT tT bi riS oT roT roS iD
2 2 0 2 13 0 0 --
  +-- Margaritifera_laevis **      
--+                             
  +-- Margaritifera_middendorffi **

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iT tT bi riS oT roT roS iD
2 2 0 2 120 0 0 --
  +-- Margaritifera_laevis **      
--+                             
  +-- Margaritifera_middendorffi **

Lopes-Lima et al., 2020 [Publ 4080], fig. 5

iT tT bi riS oT roT roS iD
2 2 0 2 7 0 0 --
  +-- Margaritifera_laevis **      
--+                             
  +-- Margaritifera_middendorffi **

Lopes-Lima et al., 2020 [Publ 4080], fig. 6

iT tT bi riS oT roT roS iD
2 2 0 2 10 0 0 --
  +-- Margaritifera_laevis **      
--+                             
  +-- Margaritifera_middendorffi **

Lopes-Lima et al., 2020 [Publ 4080], fig. 7

iT tT bi riS oT roT roS iD
2 2 0 2 6 0 0 --
  +-- Margaritifera_laevis **  
--+                         
  +-- Margaritifera_dahurica **

Lopes-Lima et al., 2020 [Publ 4080], fig. 8

iT tT bi riS oT roT roS iD
2 2 0 2 5 0 0 --
  +-- Margaritifera_laevis **      
--+                             
  +-- Margaritifera_middendorffi **

Lopes-Lima et al., 2020 [Publ 4080], fig. 9

iT tT bi riS oT roT roS iD
2 2 0 2 4 0 0 --
  +-- Margaritifera_laevis **      
--+                             
  +-- Margaritifera_middendorffi **

Wu et al., 2020 [Publ 4121], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 48 1 1 --
  +----- Cumberlandia_monodonta
--+                            
  |  +-- Margaritifera_falcata ** 
  +--+                         
     +-- Margaritifera_dahurica **

Sano et al., 2020 [Publ 4138], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 46 0 0 --
  +-- Margaritifera_middendorffi **
--+                             
  +-- Margaritifera_laevis **      

Wu et al., 2020 [Publ 4147], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 2 12 0 0 --
  +-- Margaritifera_margaritifera **
--+                              
  +-- Margaritifera_falcata **      

Zieritz et al., 2021 [Publ 4231], fig. 3-F, 5

iT tT bi riS oT roT roS iD
2 2 0 2 47 1 1 --
  +----- Pseudunio_marocanus   
--+                            
  |  +-- Margaritifera_falcata ** 
  +--+                         
     +-- Margaritifera_dahurica **

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 2 34 0 0 --
  +-- Margaritifera_dahurica **
--+                         
  +-- Margaritifera_falcata ** 

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iT tT bi riS oT roT roS iD
2 2 0 2 37 0 0 --
  +-- Margaritifera_dahurica **
--+                         
  +-- Margaritifera_falcata ** 

Wu et al., 2023 [Publ 4339], fig. 5

iT tT bi riS oT roT roS iD
2 2 0 2 17 0 0 --
  +-- Margaritifera_margaritifera **
--+                              
  +-- Margaritifera_dahurica **     

Wu et al., 2024 [Publ 4350], fig. 5a

iT tT bi riS oT roT roS iD
2 2 0 2 36 0 0 --
  +-- Margaritifera_dahurica **     
--+                              
  +-- Margaritifera_margaritifera **

Wu et al., 2024 [Publ 4350], fig. 5b

iT tT bi riS oT roT roS iD
2 2 0 2 36 0 0 --
  +-- Margaritifera_dahurica **     
--+                              
  +-- Margaritifera_margaritifera **

Wu et al., 2024 [Publ 4350], fig. 5c

iT tT bi riS oT roT roS iD
2 2 0 2 36 0 0 --
  +-- Margaritifera_dahurica **     
--+                              
  +-- Margaritifera_margaritifera **

Wu et al., 2024 [Publ 4350], fig. 6-7

iT tT bi riS oT roT roS iD
2 2 0 2 36 0 0 --
  +-- Margaritifera_dahurica **     
--+                              
  +-- Margaritifera_margaritifera **

Wu et al., 2024 [Publ 4350], fig. 8a, S1

iT tT bi riS oT roT roS iD
2 2 0 2 90 0 0 --
  +-- Margaritifera_dahurica **     
--+                              
  +-- Margaritifera_margaritifera **

Wu et al., 2024 [Publ 4350], fig. 8b, S2

iT tT bi riS oT roT roS iD
2 2 0 2 90 0 0 --
  +-- Margaritifera_dahurica **     
--+                              
  +-- Margaritifera_margaritifera **

Nagel et al., 1998 [Publ 1946], fig. 9

iT tT bi riS oT roT roS iD
1 1 0 1 5 0 0 --
-- Margaritifera

Graf & Cummings, 2006 [Publ 1968], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 46 1 1 --
  +-- Margaritifera_margaritifera **
--+                              
  +-- Cumberlandia_monodonta     

Graf, 2000 [Publ 2647], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 17 0 0 --
-- Margaritifera

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

iT tT bi riS oT roT roS iD
1 1 0 1 29 1 1 --
  +-- Margaritifera_margaritifera **
--+                              
  +-- Cumberlandia_monodonta     

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

iT tT bi riS oT roT roS iD
1 1 0 1 29 1 1 --
  +-- Margaritifera_margaritifera **
--+                              
  +-- Cumberlandia_monodonta     

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 33 1 1 --
  +-- Cumberlandia_monodonta     
--+                              
  +-- Margaritifera_margaritifera **

Hoeh et al., 2001 [Publ 2652], fig. 14.2

iT tT bi riS oT roT roS iD
1 1 0 1 30 1 1 --
  +-- Cumberlandia_monodonta     
--+                              
  +-- Margaritifera_margaritifera **

Hoeh et al., 2001 [Publ 2652], fig. 14.4

iT tT bi riS oT roT roS iD
1 1 0 1 30 1 1 --
  +-- Cumberlandia_monodonta     
--+                              
  +-- Margaritifera_margaritifera **

Hoeh et al., 2002 [Publ 2653], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 30 1 1 --
  +-- Margaritifera_margaritifera **
--+                              
  +-- Cumberlandia_monodonta     

Roe & Hoeh, 2003 [Publ 2655], fig. 4.2

iT tT bi riS oT roT roS iD
1 1 0 1 30 1 1 --
  +-- Cumberlandia_monodonta     
--+                              
  +-- Margaritifera_margaritifera **

Roe & Hoeh, 2003 [Publ 2655], fig. 4.3

iT tT bi riS oT roT roS iD
1 1 0 1 30 1 1 --
  +-- Cumberlandia_monodonta     
--+                              
  +-- Margaritifera_margaritifera **

Huang et al., 2002 [Publ 2657], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 33 1 1 --
-- Margaritifera_margaritifera

Huang et al., 2002 [Publ 2657], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 33 1 1 --
-- Margaritifera_margaritifera

Zhou et al., 2007 [Publ 2658], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 22 1 1 --
  +-- Margaritifera_margaritifera **
--+                              
  +-- Cumberlandia_monodonta     

Campbell et al., 2005 [Publ 2660], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 115 1 1 --
  +-- Margaritifera_margaritifera **
--+                              
  +-- Cumberlandia_monodonta     

Campbell et al., 2005 [Publ 2660], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 113 1 1 --
  +-- Cumberlandia_monodonta     
--+                              
  +-- Margaritifera_margaritifera **

Curole & Kocher, 2005 [Publ 2682], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 10 0 0 --
-- Margaritifera_hembeli

Curole & Kocher, 2005 [Publ 2682], fig. 1-M

iT tT bi riS oT roT roS iD
1 1 0 1 11 0 0 --
-- Margaritifera_hembeli

Källersjö et al., 2005 [Publ 2684], fig. 1A

iT tT bi riS oT roT roS iD
1 1 0 1 6 0 0 --
-- Margaritifera_margaritifera

Källersjö et al., 2005 [Publ 2684], fig. 1B-F

iT tT bi riS oT roT roS iD
1 1 0 1 6 0 0 --
-- Margaritifera_margaritifera

Machordom et al., 2003 [Publ 2688], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 2 1 1 --
  +-- Margaritifera_margaritifera **
--+                              
  +-- Pseudunio_auricularius     

Machordom et al., 2003 [Publ 2688], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 3 2 2 --
  +----- Cumberlandia_monodonta     
--+                                 
  |  +-- Margaritifera_margaritifera **
  +--+                              
     +-- Pseudunio_auricularius     

Hoeh et al., 1998 [Publ 2695], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 4 0 0 --
-- Margaritifera_margaritifera

Hoeh et al., 1998 [Publ 2695], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 4 0 0 --
-- Margaritifera_margaritifera

Taylor et al., 2007 [Publ 2696], fig. 1-3

iT tT bi riS oT roT roS iD
1 1 0 1 3 0 0 --
-- Margaritifera_margaritifera

Curole & Kocher, 2002 [Publ 2712], fig. 1-F

iT tT bi riS oT roT roS iD
1 1 0 1 3 0 0 --
-- Margaritifera_hembeli

Hoeh et al., 2009 [Publ 2716], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 46 1 1 --
  +-- Margaritifera_margaritifera **
--+                              
  +-- Cumberlandia_monodonta     

Hoeh et al., 2009 [Publ 2716], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 45 1 1 --
  +-- Margaritifera_margaritifera **
--+                              
  +-- Cumberlandia_monodonta     

Hoeh et al., 2009 [Publ 2716], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 45 1 1 --
  +-- Margaritifera_margaritifera **
--+                              
  +-- Cumberlandia_monodonta     

Turner et al., 2000 [Publ 2722], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 15 0 0 --
-- Margaritifera_margaritifera

Campbell, 2000 [Publ 2724], fig. 1-5

iT tT bi riS oT roT roS iD
1 1 0 1 2 0 0 --
-- Margaritifera_margaritifera

Gustafson & Iwamoto, 2005 [Publ 2741], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 4 0 0 --
-- Margaritifera_falcata

Whelan et al., 2011 [Publ 3380], fig. 1 ML

iT tT bi riS oT roT roS iD
1 1 0 1 36 1 1 --
  +-- Margaritifera_margaritifera **
--+                              
  +-- Cumberlandia_monodonta     

Whelan et al., 2011 [Publ 3380], fig. 1 MP

iT tT bi riS oT roT roS iD
1 1 0 1 36 1 1 --
  +-- Margaritifera_margaritifera **
--+                              
  +-- Cumberlandia_monodonta     

Prie et al., 2012 [Publ 3487], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 5 0 0 --
-- Margaritifera_margaritifera

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 34 0 0 --
-- Margaritifera_margaritifera

Reis et al., 2013 [Publ 3580], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 13 0 0 --
-- Margaritifera_margaritifera

Reis et al., 2013 [Publ 3580], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 15 0 0 --
-- Margaritifera_margaritifera

Graf et al., 2014 [Publ 3591], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 26 0 0 --
-- Margaritifera_margaritifera

Graf et al., 2015 [Publ 3619], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 19 0 0 --
-- Margaritifera_margaritifera

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 39 1 1 --
  +-- Margaritifera_margaritifera **
--+                              
  +-- Cumberlandia_monodonta     

Bieler et al., 2014 [Publ 3622], fig. 26, 27, 29, 30

iT tT bi riS oT roT roS iD
1 1 0 1 4 0 0 --
-- Margaritifera_margaritifera

Bieler et al., 2014 [Publ 3622], fig. 28

iT tT bi riS oT roT roS iD
1 1 0 1 4 0 0 --
-- Margaritifera_margaritifera

Huang et al., 2013 [Publ 3629], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 15 0 0 --
-- Margaritifera_falcata

González & Giribet, 2015 [Publ 3632], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 5 0 0 --
-- Margaritifera_margaritifera

González & Giribet, 2015 [Publ 3632], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 5 0 0 --
-- Margaritifera_margaritifera

González & Giribet, 2015 [Publ 3632], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 5 0 0 --
-- Margaritifera_margaritifera

González & Giribet, 2015 [Publ 3632], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 5 0 0 --
-- Margaritifera_margaritifera

Giribet & Distel, 2003 [Publ 3670], fig. 3.5

iT tT bi riS oT roT roS iD
1 1 0 1 5 0 0 --
-- Margaritifera_margaritifera

Zieritz et al., 2012 [Publ 3673], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 7 0 0 --
-- Margaritifera_margaritifera

Zhou et al., 2016 [Publ 3710], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 19 0 0 --
-- Margaritifera_falcata

Bolotov et al., 2016 [Publ 3732], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 7 0 0 --
-- Margaritifera_dahurica

Lopes-Lima et al., 2017 [Publ 3734], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 16 1 1 --
  +-- Pseudunio_auricularius     
--+                              
  +-- Margaritifera_margaritifera **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iT tT bi riS oT roT roS iD
1 1 0 1 80 1 1 --
  +-- Cumberlandia_monodonta     
--+                              
  +-- Margaritifera_margaritifera **

Hewitt et al., 2018 [Publ 3758], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 8 0 0 --
-- Margaritifera_margaritifera

Guerra et al., 2017 [Publ 3772], fig. 2a

iT tT bi riS oT roT roS iD
1 1 0 1 24 1 1 --
  +-- Margaritifera_falcata ** 
--+                         
  +-- Cumberlandia_monodonta

Guerra et al., 2017 [Publ 3772], fig. 2b

iT tT bi riS oT roT roS iD
1 1 0 1 24 1 1 --
  +-- Margaritifera_falcata ** 
--+                         
  +-- Cumberlandia_monodonta

Amaro et al., 2017 [Publ 3791], fig. 3A-M

iT tT bi riS oT roT roS iD
1 1 0 1 6 0 0 --
-- Margaritifera_margaritifera

Amaro et al., 2017 [Publ 3791], fig. 3B-M

iT tT bi riS oT roT roS iD
1 1 0 1 6 0 0 --
-- Margaritifera_margaritifera

Välilä et al., 2017 [Publ 3793], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 2 1 1 --
  +-- Margaritifera_margaritifera **
--+                              
  +-- Pseudunio_auricularius     

Froufe et al., 2017 [Publ 3802], fig. 7

iT tT bi riS oT roT roS iD
1 1 0 1 9 0 0 --
-- Margaritifera_margaritifera

Combosch et al., 2017 [Publ 3806], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 14 0 0 --
-- Margaritifera_margaritifera

Combosch et al., 2017 [Publ 3806], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 14 0 0 --
-- Margaritifera_margaritifera

Vikhrev et al., 2017 [Publ 3810], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 9 0 0 --
-- Margaritifera_dahurica

Bespalaya et al., 2018 [Publ 3811], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 9 0 0 --
-- Margaritifera_dahurica

Pfeiffer et al., 2018 [Publ 3827], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 28 0 0 --
-- Margaritifera_margaritifera

Huang et al., 2019 [Publ 3843], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 43 0 0 --
-- Margaritifera_dahurica

Huang et al., 2019 [Publ 3843], fig. 3-M

iT tT bi riS oT roT roS iD
1 1 0 1 22 1 1 --
  +-- Pseudunio_marocanus        
--+                              
  +-- Margaritifera_margaritifera **

Huang et al., 2019 [Publ 3843], fig. S1

iT tT bi riS oT roT roS iD
1 1 0 1 81 0 0 --
-- Margaritifera_dahurica

Bogatov et al., 2018 [Publ 3859], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 5 0 0 --
-- Margaritifera_margaritifera

Bogatov et al., 2018 [Publ 3859], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 5 0 0 --
-- Margaritifera_margaritifera

Klishko et al., 2019 [Publ 3886], fig. 9

iT tT bi riS oT roT roS iD
1 1 0 1 10 0 0 --
-- Margaritifera_dahurica

Konopleva et al., 2019 [Publ 3900], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 51 0 0 --
-- Margaritifera_dahurica

Lopes-Lima et al., 2019 [Publ 3901], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 27 1 1 --
  +-- Margaritifera_falcata ** 
--+                         
  +-- Cumberlandia_monodonta

Lopes-Lima et al., 2019 [Publ 3901], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 28 1 1 --
  +-- Cumberlandia_monodonta
--+                         
  +-- Margaritifera_falcata ** 

Lopes-Lima et al., 2019 [Publ 3901], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 25 1 1 --
  +-- Margaritifera_falcata ** 
--+                         
  +-- Cumberlandia_monodonta

Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1-M

iT tT bi riS oT roT roS iD
1 1 0 1 5 2 2 --
  +----- Margaritifera_margaritifera **
--+                                 
  |  +-- Pseudunio_marocanus        
  +--+                              
     +-- Cumberlandia_monodonta     

Hewitt et al., 2019 [Publ 3962], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 15 0 0 --
-- Margaritifera_margaritifera

Klunzinger et al., 2021 [Publ 4086], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 5 0 0 --
-- Margaritifera_margaritifera

Lemer et al., 2019 [Publ 4093], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 5 0 0 --
-- Margaritifera_margaritifera

Wu et al., 2020 [Publ 4147], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 29 0 0 --
-- Margaritifera_falcata

Guerra et al., 2019 [Publ 4168], fig. 1-M

iT tT bi riS oT roT roS iD
1 1 0 1 21 2 2 --
  +----- Cumberlandia_monodonta     
--+                                 
  |  +-- Margaritifera_margaritifera **
  +--+                              
     +-- Pseudunio_marocanus        

Guerra et al., 2019 [Publ 4168], fig. 2-M

iT tT bi riS oT roT roS iD
1 1 0 1 21 2 2 --
  +----- Pseudunio_marocanus        
--+                                 
  |  +-- Margaritifera_margaritifera **
  +--+                              
     +-- Cumberlandia_monodonta     

Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 18 1 1 --
  +-- Margaritifera_falcata ** 
--+                         
  +-- Cumberlandia_monodonta

Kano et al., 2019 [Publ 4194], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 9 0 0 --
-- Margaritifera_falcata

Zhang et al., 2013 [Publ 4214], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 3 0 0 --
-- Margaritifera_margaritifera

Zieritz et al., 2021 [Publ 4218], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 31 1 1 --
  +-- Margaritifera_margaritifera **
--+                              
  +-- Pseudunio_auricularius     

Bolotov et al., 2022 [Publ 4250], fig. 8

iT tT bi riS oT roT roS iD
1 1 0 1 48 0 0 --
-- Margaritifera_dahurica

Dai et al., 2023 [Publ 4285], fig. 3-BI

iT tT bi riS oT roT roS iD
1 1 0 1 44 0 0 --
-- Margaritifera_margaritifera

Dai et al., 2023 [Publ 4285], fig. 3-ML

iT tT bi riS oT roT roS iD
1 1 0 1 44 0 0 --
-- Margaritifera_margaritifera

Dai et al., 2023 [Publ 4285], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 52 0 0 --
-- Margaritifera_dahurica

Bogan et al., 2023 [Publ 4295], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 30 0 0 --
-- Margaritifera_laevis

Chen et al., 2023 [Publ 4302], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 39 0 0 --
-- Margaritifera_dahurica

Chen et al., 2023 [Publ 4302], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 39 0 0 --
-- Margaritifera_dahurica

Smith, 2021 [Publ 4331], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 19 0 0 --
-- Margaritifera_margaritifera

Dai et al., 2024 [Publ 4358], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 51 0 0 --
-- Margaritifera_dahurica

Douda et al., 2024 [Publ 4364], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 14 0 0 --
-- Margaritifera_middendorffi

Page updated: 08:51:57 Mon 06 May 2024

 
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