|
Bivalvia | Unionoida | Unionoidea | Margaritiferidae | Margaritiferinae genus Margaritifera Schumacher, 1816: cladomics
23 valid species
Cladomics Summary
number of species = 23
analyzed in 181 cladograms in 106 publications
maximum number of terminals in an analysis = 8, mean = 2.33
Basic Monophyly
number of cladograms that test basic monophyly (i.e., more than 1 included species) = 87
maximum number of included terminals = 8, mean = 3.77
proportion of cladograms that support basic monophyly = 0.94
Pairwise Cladogram Clustering
The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Huff et al., 2004 [Publ 2697], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 8 |
10 |
2 |
8 |
4 |
2 |
2 |
0.0 |
*** Top Monophyly Cladogram ***
+-------- Margaritifera_margaritifera **
|
| +----- Cumberlandia_monodonta
--+ |
| | +-- Pseudunio_auricularius
| | |
+--+ +-- Margaritifera_dahurica **
| |
| +-- Margaritifera_middendorffi[1] **
| |
| +-- Margaritifera_middendorffi[2] **
+--+
+-- Margaritifera_falcata[1] **
|
+-- Margaritifera_falcata[2] **
|
+-- Margaritifera_falcata[3] **
|
+-- Margaritifera_falcata[4] **
Lyubas et al., 2019 [Publ 3963], fig. 10
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
7 |
9 |
6 |
2 |
11.0 |
*** Top Sister-Group Cladogram ***
+-------- Cumberlandia_monodonta
|
+--------+ +----- Pseudunio_marocanus
| | |
| +--+ +-- Pseudunio_auricularius
| | |
| | +-- Pseudunio_flabellata
| +--+
--+ +-- Pseudunio_flabellatiformis
| |
| +-- Pseudunio_homsensis
|
| +-- Margaritifera_margaritifera **
| +-----------+
| | +-- Margaritifera_dahurica **
+--+
| +----------- Margaritifera_falcata **
+--+
| +-------- Margaritifera_laevis **
+--+
| +----- Margaritifera_middendorffi **
+--+
| +-- Margaritifera_marrianae **
+--+
+-- Margaritifera_hembeli **
Huff et al., 2004 [Publ 2697], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
9 |
3 |
7 |
7 |
2 |
2 |
9.0 |
+-------- Margaritifera_dahurica **
|
+-------- Margaritifera_margaritifera[1] **
|
+-------- Margaritifera_margaritifera[2] **
|
--+-------- Margaritifera_margaritifera[3] **
|
| +-- Pseudunio_auricularius
+-----+
| +-- Cumberlandia_monodonta
|
| +----- Margaritifera_falcata **
+--+
| +-- Margaritifera_middendorffi **
+--+
+-- Margaritifera_marrianae **
Bolotov et al., 2015 [Publ 3646], fig. 2A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
9 |
5 |
7 |
3 |
2 |
2 |
11.0 |
+----------- Margaritifera_margaritifera **
|
| +----- Margaritifera_dahurica[1] **
| |
+-----+----- Margaritifera_dahurica[2] **
--+ |
| | +-- Pseudunio_auricularius
| +--+
| +-- Cumberlandia_monodonta
|
| +-------- Margaritifera_falcata **
+--+
| +----- Margaritifera_laevis **
+--+
| +-- Margaritifera_middendorffi **
+--+
+-- Margaritifera_marrianae **
Araujo et al., 2016 [Publ 3746], fig. 1,2b
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
7 |
6 |
3 |
2 |
11.0 |
+----- Cumberlandia_monodonta
+--------+
| | +-- Pseudunio_marocanus
| +--+
| +-- Pseudunio_auricularius
--+
| +-- Margaritifera_dahurica **
| +--------+
| | +-- Margaritifera_margaritifera **
+--+
| +-------- Margaritifera_falcata **
| |
+--+ +-- Margaritifera_middendorffi **
| +--+
| | +-- Margaritifera_laevis **
+--+
| +-- Margaritifera_marrianae **
+--+
+-- Margaritifera_hembeli **
Araujo et al., 2016 [Publ 3746], fig. 2a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
7 |
4 |
3 |
2 |
11.0 |
+-- Margaritifera_dahurica **
+--------+
| +-- Margaritifera_margaritifera **
--+
| +-------- Margaritifera_falcata **
| |
+--+ +-- Margaritifera_laevis **
| +--+
| | +-- Margaritifera_middendorffi **
+--+
| +-- Margaritifera_marrianae **
+--+
+-- Margaritifera_hembeli **
Huang et al., 2018 [Publ 3828], fig. 3, 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
7 |
22 |
3 |
2 |
11.0 |
+----- Cumberlandia_monodonta
+-----------+
| | +-- Pseudunio_auricularius
| +--+
| +-- Pseudunio_marocanus
--+
| +-- Margaritifera_margaritifera **
| +-----------+
| | +-- Margaritifera_dahurica **
+--+
| +----------- Margaritifera_falcata **
+--+
| +-------- Margaritifera_laevis **
+--+
| +----- Margaritifera_hembeli **
+--+
| +-- Margaritifera_middendorffi **
+--+
+-- Margaritifera_marrianae **
Lopes-Lima et al., 2018 [Publ 3829], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
7 |
21 |
4 |
2 |
11.0 |
+-------- Cumberlandia_monodonta
+-----+
| | +----- Pseudunio_marocanus
| +--+
| | +-- Pseudunio_auricularius
| +--+
--+ +-- Pseudunio_homsensis
|
| +-- Margaritifera_dahurica **
| +--------+
| | +-- Margaritifera_margaritifera **
+--+
| +-------- Margaritifera_falcata **
| |
+--+ +-- Margaritifera_middendorffi **
| +--+
| | +-- Margaritifera_laevis **
+--+
| +-- Margaritifera_hembeli **
+--+
+-- Margaritifera_marrianae **
Lopes-Lima et al., 2018 [Publ 3829], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
7 |
21 |
4 |
2 |
11.0 |
+-------- Cumberlandia_monodonta
+--------+
| | +----- Pseudunio_marocanus
| +--+
| | +-- Pseudunio_auricularius
| +--+
--+ +-- Pseudunio_homsensis
|
| +-- Margaritifera_margaritifera **
| +-----------+
| | +-- Margaritifera_dahurica **
+--+
| +----------- Margaritifera_falcata **
+--+
| +-------- Margaritifera_laevis **
+--+
| +----- Margaritifera_middendorffi **
+--+
| +-- Margaritifera_hembeli **
+--+
+-- Margaritifera_marrianae **
Huang et al., 2019 [Publ 3843], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
4 |
7 |
197 |
4 |
2 |
11.0 |
+----------- Cumberlandia_monodonta
|
| +----- Pseudunio_marocanus
+-----+
| | +-- Pseudunio_auricularius
| +--+
--+ +-- Pseudunio_homsensis
|
| +-- Margaritifera_dahurica **
| +-----+
| | +-- Margaritifera_margaritifera **
| |
+--+ +----- Margaritifera_laevis **
| |
| | +-- Margaritifera_falcata **
+--+--+
| +-- Margaritifera_middendorffi **
|
| +-- Margaritifera_hembeli **
+--+
+-- Margaritifera_marrianae **
Huang et al., 2019 [Publ 3843], fig. S3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
7 |
194 |
4 |
2 |
11.0 |
+-------- Cumberlandia_monodonta
+-----+
| | +----- Pseudunio_marocanus
| +--+
| | +-- Pseudunio_auricularius
| +--+
--+ +-- Pseudunio_homsensis
|
| +-- Margaritifera_dahurica **
| +--------+
| | +-- Margaritifera_margaritifera **
+--+
| +-------- Margaritifera_falcata **
| |
+--+ +-- Margaritifera_hembeli **
| +--+
| | +-- Margaritifera_marrianae **
+--+
| +-- Margaritifera_laevis **
+--+
+-- Margaritifera_middendorffi **
Huang et al., 2019 [Publ 3843], fig. S4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
3 |
7 |
195 |
4 |
2 |
10.0 |
+-------------- Cumberlandia_monodonta
|
| +----- Pseudunio_marocanus
--+ +-----+
| | | +-- Pseudunio_auricularius
| | +--+
| | +-- Pseudunio_homsensis
+--+
| +-------- Margaritifera_falcata **
| |
| | +-- Margaritifera_dahurica **
| +-----+
+--+ +-- Margaritifera_margaritifera **
|
| +----- Margaritifera_laevis **
| |
+--+----- Margaritifera_middendorffi **
|
| +-- Margaritifera_hembeli **
+--+
+-- Margaritifera_marrianae **
Wu et al., 2020 [Publ 4121], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
7 |
27 |
3 |
2 |
11.0 |
+-- Margaritifera_dahurica **
+--------+
| +-- Margaritifera_margaritifera **
--+
| +-------- Margaritifera_falcata **
| |
+--+ +-- Margaritifera_hembeli **
| +--+
| | +-- Margaritifera_marrianae **
+--+
| +-- Margaritifera_laevis **
+--+
+-- Margaritifera_middendorffi **
Wu et al., 2020 [Publ 4121], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
7 |
27 |
3 |
2 |
11.0 |
+----- Cumberlandia_monodonta
+--------+
| | +-- Pseudunio_auricularius
| +--+
| +-- Pseudunio_marocanus
--+
| +-- Margaritifera_dahurica **
| +--------+
| | +-- Margaritifera_margaritifera **
+--+
| +-------- Margaritifera_falcata **
| |
+--+ +-- Margaritifera_hembeli **
| +--+
| | +-- Margaritifera_marrianae **
+--+
| +-- Margaritifera_laevis **
+--+
+-- Margaritifera_middendorffi **
Sano et al., 2020 [Publ 4138], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
7 |
66 |
3 |
2 |
11.0 |
+-- Margaritifera_dahurica **
+-----------+
| +-- Margaritifera_margaritifera **
--+
| +----------- Margaritifera_falcata **
+--+
| +-------- Margaritifera_middendorffi **
+--+
| +----- Margaritifera_laevis **
+--+
| +-- Margaritifera_hembeli **
+--+
+-- Margaritifera_marrianae **
Sano et al., 2020 [Publ 4138], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
7 |
65 |
3 |
2 |
11.0 |
+-- Margaritifera_margaritifera **
+-----------+
| +-- Margaritifera_dahurica **
--+
| +----------- Margaritifera_falcata **
+--+
| +-------- Margaritifera_middendorffi **
+--+
| +----- Margaritifera_laevis **
+--+
| +-- Margaritifera_hembeli **
+--+
+-- Margaritifera_marrianae **
Sano et al., 2020 [Publ 4138], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 7 |
7 |
5 |
7 |
65 |
3 |
2 |
11.0 |
+-- Margaritifera_margaritifera **
+--------+
| +-- Margaritifera_dahurica **
--+
| +-------- Margaritifera_falcata **
| |
+--+ +-- Margaritifera_laevis **
| +--+
| | +-- Margaritifera_middendorffi **
+--+
| +-- Margaritifera_hembeli **
+--+
+-- Margaritifera_marrianae **
Takeuchi et al., 2015 [Publ 3661], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
6 |
5 |
3 |
2 |
11.0 |
+----------------- Pseudunio_auricularius[1]
|
+----------------- Pseudunio_auricularius[2]
--+
| +-------------- Cumberlandia_monodonta
| |
+--+ +-- Margaritifera_margaritifera **
| +--------+
| | +-- Margaritifera_dahurica **
+--+
| +-------- Margaritifera_falcata **
+--+
| +----- Margaritifera_laevis **
+--+
| +-- Margaritifera_middendorffi **
+--+
+-- Margaritifera_marrianae **
Bolotov et al., 2016 [Publ 3743], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
6 |
24 |
3 |
2 |
11.0 |
+-- Margaritifera_dahurica **
+--------+
| +-- Margaritifera_margaritifera **
--+
| +-------- Margaritifera_falcata **
+--+
| +----- Margaritifera_laevis **
+--+
| +-- Margaritifera_middendorffi **
+--+
+-- Margaritifera_marrianae **
Bolotov et al., 2016 [Publ 3743], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
6 |
24 |
3 |
2 |
11.0 |
+-- Margaritifera_margaritifera **
+--------+
| +-- Margaritifera_dahurica **
--+
| +-------- Margaritifera_falcata **
+--+
| +----- Margaritifera_laevis **
+--+
| +-- Margaritifera_marrianae **
+--+
+-- Margaritifera_middendorffi **
Bolotov et al., 2017 [Publ 3761], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
6 |
135 |
3 |
2 |
11.0 |
+-- Margaritifera_dahurica **
+--------+
| +-- Margaritifera_margaritifera **
--+
| +-------- Margaritifera_falcata **
+--+
| +----- Margaritifera_middendorffi **
+--+
| +-- Margaritifera_laevis **
+--+
+-- Margaritifera_marrianae **
Bolotov et al., 2017 [Publ 3761], fig. S2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
6 |
135 |
3 |
2 |
11.0 |
+-- Margaritifera_dahurica **
+--------+
| +-- Margaritifera_margaritifera **
--+
| +-------- Margaritifera_falcata **
+--+
| +----- Margaritifera_middendorffi **
+--+
| +-- Margaritifera_laevis **
+--+
+-- Margaritifera_marrianae **
Vikhrev et al., 2018 [Publ 3812], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
6 |
7 |
4 |
2 |
11.0 |
+-------- Cumberlandia_monodonta
+-----+
| | +----- Pseudunio_marocanus
| +--+
| | +-- Pseudunio_auricularius
| +--+
--+ +-- Pseudunio_homsensis
|
| +-- Margaritifera_margaritifera **
| +--------+
| | +-- Margaritifera_dahurica **
+--+
| +-------- Margaritifera_falcata **
+--+
| +----- Margaritifera_laevis **
+--+
| +-- Margaritifera_middendorffi **
+--+
+-- Margaritifera_marrianae **
Jeratthitikul et al., 2022 [Publ 4260], fig. S1, 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
6 |
70 |
3 |
2 |
11.0 |
+-- Pseudunio_auricularius
+-----------+
| +-- Pseudunio_marocanus
|
--+ +-- Margaritifera_margaritifera **
| +--------+
| | +-- Margaritifera_dahurica **
+--+
| +-------- Margaritifera_falcata **
+--+
| +----- Margaritifera_middendorffi **
+--+
| +-- Margaritifera_laevis **
+--+
+-- Margaritifera_marrianae **
Jeratthitikul et al., 2022 [Publ 4260], fig. S2, 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 6 |
6 |
4 |
6 |
70 |
3 |
2 |
11.0 |
+-- Margaritifera_dahurica **
+--------+
| +-- Margaritifera_margaritifera **
--+
| +-------- Margaritifera_falcata **
+--+
| +----- Margaritifera_marrianae **
+--+
| +-- Margaritifera_middendorffi **
+--+
+-- Margaritifera_laevis **
Huff et al., 2004 [Publ 2697], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
5 |
6 |
2 |
2 |
10.0 |
+----------- Pseudunio_auricularius
|
+----------- Cumberlandia_monodonta
--+
| +-- Margaritifera_dahurica **
| +-----+
| | +-- Margaritifera_margaritifera **
+--+
| +----- Margaritifera_falcata **
+--+
| +-- Margaritifera_middendorffi **
+--+
+-- Margaritifera_marrianae **
Huff et al., 2004 [Publ 2697], fig. 5-6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
5 |
7 |
2 |
2 |
10.0 |
+-- Pseudunio_auricularius
+--------+
| +-- Cumberlandia_monodonta
|
--+ +-- Margaritifera_dahurica **
| +-----+
| | +-- Margaritifera_margaritifera **
+--+
| +----- Margaritifera_falcata **
+--+
| +-- Margaritifera_middendorffi **
+--+
+-- Margaritifera_marrianae **
Inoue et al., 2013 [Publ 3579], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
5 |
4 |
3 |
2 |
10.0 |
+-------------- Cumberlandia_monodonta
|
| +-- Pseudunio_auricularius
--+ +--------+
| | +-- Pseudunio_marocanus
| |
+--+ +-- Margaritifera_dahurica **
| +-----+
| | +-- Margaritifera_margaritifera **
+--+
| +----- Margaritifera_falcata **
+--+
| +-- Margaritifera_laevis **
+--+
+-- Margaritifera_marrianae **
Bolotov et al., 2015 [Publ 3646], fig. 2B
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
5 |
4 |
3 |
2 |
10.0 |
+-------------- Cumberlandia_monodonta
|
| +-- Pseudunio_auricularius
--+ +--------+
| | +-- Pseudunio_marocanus
| |
+--+ +-- Margaritifera_dahurica **
| +-----+
| | +-- Margaritifera_margaritifera **
+--+
| +----- Margaritifera_falcata **
+--+
| +-- Margaritifera_middendorffi **
+--+
+-- Margaritifera_laevis **
Takeuchi et al., 2015 [Publ 3661], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 5 |
5 |
3 |
5 |
5 |
3 |
2 |
10.0 |
+-------------- Pseudunio_auricularius
|
--+ +----------- Cumberlandia_monodonta
| |
+--+ +-- Margaritifera_dahurica **
| +-----+
| | +-- Margaritifera_margaritifera **
+--+
| +----- Margaritifera_falcata **
+--+
| +-- Margaritifera_middendorffi **
+--+
+-- Margaritifera_laevis **
Araujo et al., 2009 [Publ 2395], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
4 |
4 |
3 |
2 |
9.0 |
+----------- Cumberlandia_monodonta
|
| +-- Pseudunio_marocanus
--+ +-----+
| | +-- Pseudunio_auricularius
| |
+--+ +-- Margaritifera_margaritifera **
| +--+
| | +-- Margaritifera_dahurica **
+--+
| +-- Margaritifera_falcata **
+--+
+-- Margaritifera_middendorffi **
Walker et al., 2006 [Publ 2666], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
4 |
34 |
2 |
2 |
9.0 |
+----------- Cumberlandia_monodonta
|
--+ +-------- Pseudunio_auricularius
| |
+--+ +-- Margaritifera_margaritifera **
| +--+
| | +-- Margaritifera_dahurica **
+--+
| +-- Margaritifera_falcata **
+--+
+-- Margaritifera_middendorffi **
Sano et al., 2017 [Publ 3799], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
2 |
4 |
23 |
0 |
0 |
10.0 |
+-------- Margaritifera_laevis[2] **
--+
| +----- Margaritifera_laevis[1] **
+--+
| +-- Margaritifera_middendorffi **
+--+
+-- Margaritifera_laevis[3] **
Bolotov et al., 2020 [Publ 4084], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
4 |
0 |
4 |
32 |
0 |
0 |
9.0 |
+-- Margaritifera_dahurica **
|
+-- Margaritifera_margaritifera **
--+
+-- Margaritifera_laevis **
|
+-- Margaritifera_middendorffi **
Bolotov et al., 2020 [Publ 4100], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 4 |
5 |
0 |
4 |
116 |
0 |
0 |
10.0 |
+-- Gibbosula_laosensis
|
+-- Margaritifera_dahurica **
|
--+-- Margaritifera_margaritifera **
|
+-- Margaritifera_laevis **
|
+-- Margaritifera_middendorffi **
Walker et al., 2006 [Publ 2666], fig. 6-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
0 |
3 |
12 |
1 |
1 |
9.0 |
+----- Cumberlandia_monodonta
|
--+ +-- Margaritifera_margaritifera **
| |
+--+-- Margaritifera_dahurica[1] **
|
+-- Margaritifera_dahurica[2] **
Breton et al., 2011 [Publ 3376], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
3 |
36 |
1 |
1 |
9.0 |
+-------- Cumberlandia_monodonta
--+
| +----- Margaritifera_margaritifera **
+--+
| +-- Margaritifera_falcata **
+--+
+-- Margaritifera_marrianae **
Mock et al., 2013 [Publ 3598], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
3 |
0 |
0 |
0 |
9.0 |
+----- Margaritifera_margaritifera **
--+
| +-- Margaritifera_falcata **
+--+
+-- Margaritifera_laevis **
Amaro et al., 2017 [Publ 3791], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
17 |
15 |
3 |
15 |
0 |
0 |
20.0 |
+-------------------- Margaritifera_margaritifera[2] **
|
| +-- Margaritifera_margaritifera[1] **
| +--------------+
--+ | +-- Margaritifera_falcata **
| |
| | +-- Lasmigona_costata
| | +-----+
| | | +-- Platynaias_compressa
+--+ +-----+
| | | +----- Pyganodon_grandis
| | +--+
| | | +-- Utterbackia_peninsularis
| | +--+
| | +-- Utterbackia_imbecillis
+--+
| +----- Fusconaia_flava
| +-----+
| | | +-- Pronodularia_japanensis
| | +--+
| | +-- Sinosolenaia_carinata
+--+
| +----- Toxolasma_parvum
| +--+
| | | +-- Venustaconcha_ellipsiformis
| | +--+
+--+ +-- Lampsilis_siliquoidea
|
| +----- Quadrula_quadrula[2]
+--+
| +-- Quadrula_quadrula[1]
+--+
+-- Quadrula_nobilis
Huang et al., 2018 [Publ 3828], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
3 |
40 |
2 |
2 |
9.0 |
+----------- Cumberlandia_monodonta
--+
| +-------- Pseudunio_marocanus
+--+
| +----- Margaritifera_falcata **
+--+
| +-- Margaritifera_margaritifera **
+--+
+-- Margaritifera_dahurica **
Huang et al., 2019 [Publ 3843], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
3 |
54 |
2 |
2 |
9.0 |
+----------- Cumberlandia_monodonta
--+
| +-------- Pseudunio_marocanus
+--+
| +----- Margaritifera_falcata **
+--+
| +-- Margaritifera_margaritifera **
+--+
+-- Margaritifera_dahurica **
Chase et al., 2018 [Publ 3847], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
3 |
36 |
1 |
1 |
9.0 |
+-------- Cumberlandia_monodonta
--+
| +----- Margaritifera_falcata **
+--+
| +-- Margaritifera_marrianae **
+--+
+-- Margaritifera_margaritifera **
Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
3 |
7 |
2 |
2 |
9.0 |
+----------- Cumberlandia_monodonta
--+
| +-------- Pseudunio_marocanus
+--+
| +----- Margaritifera_falcata **
+--+
| +-- Margaritifera_margaritifera **
+--+
+-- Margaritifera_dahurica **
Guerra et al., 2019 [Publ 4168], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
3 |
38 |
3 |
2 |
9.0 |
+----------- Cumberlandia_monodonta
|
--+ +-- Pseudunio_marocanus
| +-----+
| | +-- Pseudunio_auricularius
+--+
| +----- Margaritifera_falcata **
+--+
| +-- Margaritifera_dahurica **
+--+
+-- Margaritifera_margaritifera **
Guerra et al., 2019 [Publ 4168], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
3 |
38 |
3 |
2 |
9.0 |
+----------- Cumberlandia_monodonta
|
--+ +-- Pseudunio_marocanus
| +-----+
| | +-- Pseudunio_auricularius
+--+
| +----- Margaritifera_falcata **
+--+
| +-- Margaritifera_dahurica **
+--+
+-- Margaritifera_margaritifera **
Wu et al., 2022 [Publ 4238], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 3 |
3 |
1 |
3 |
71 |
2 |
2 |
9.0 |
+----------- Cumberlandia_monodonta
--+
| +-------- Pseudunio_marocanus
+--+
| +----- Margaritifera_falcata **
+--+
| +-- Margaritifera_dahurica **
+--+
+-- Margaritifera_margaritifera **
Lydeard et al., 1996 [Publ 2656], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
25 |
1 |
1 |
-- |
+----- Cumberlandia_monodonta
--+
| +-- Margaritifera_margaritifera **
+--+
+-- Margaritifera_falcata **
Lydeard et al., 1996 [Publ 2656], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
26 |
1 |
1 |
-- |
+----- Cumberlandia_monodonta
--+
| +-- Margaritifera_margaritifera **
+--+
+-- Margaritifera_falcata **
Krebs et al., 2003 [Publ 2691], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
32 |
0 |
0 |
-- |
+-- Margaritifera_margaritifera **
--+
+-- Margaritifera_falcata **
Rosenberg et al., 1994 [Publ 2719], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
17 |
1 |
1 |
-- |
+-- Margaritifera_falcata **
--+
+-- Margaritifera_margaritifera **
Rosenberg et al., 1997 [Publ 2720], fig. 2A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
21 |
1 |
1 |
-- |
+-- Margaritifera_falcata **
--+
+-- Margaritifera_margaritifera **
Rosenberg et al., 1997 [Publ 2720], fig. 2B
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
21 |
1 |
1 |
-- |
+-- Margaritifera_falcata **
--+
+-- Margaritifera_margaritifera **
Fonseca et al., 2016 [Publ 3775], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
30 |
0 |
0 |
-- |
+-- Margaritifera_falcata **
--+
+-- Margaritifera_dahurica **
Amaro et al., 2017 [Publ 3791], fig. 3A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
10 |
0 |
0 |
-- |
+-- Margaritifera_falcata **
--+
+-- Margaritifera_margaritifera **
Amaro et al., 2017 [Publ 3791], fig. 3B
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
10 |
0 |
0 |
-- |
+-- Margaritifera_falcata **
--+
+-- Margaritifera_margaritifera **
Lopes-Lima et al., 2017 [Publ 3792], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
31 |
1 |
1 |
-- |
+----- Pseudunio_marocanus
--+
| +-- Margaritifera_falcata **
+--+
+-- Margaritifera_dahurica **
Wen et al., 2017 [Publ 3795], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
14 |
0 |
0 |
-- |
+-- Margaritifera_dahurica **
--+
+-- Margaritifera_falcata **
Sano et al., 2017 [Publ 3799], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
34 |
0 |
0 |
-- |
+-- Margaritifera_laevis **
--+
+-- Margaritifera_middendorffi **
Feng et al., 2016 [Publ 3800], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
13 |
0 |
0 |
-- |
+-- Margaritifera_dahurica **
--+
+-- Margaritifera_falcata **
Konopleva et al., 2017 [Publ 3807], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
29 |
0 |
0 |
-- |
+-- Margaritifera_dahurica **
--+
+-- Margaritifera_laevis **
Wu et al., 2018 [Publ 3836], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
51 |
0 |
0 |
-- |
+-- Margaritifera_dahurica **
--+
+-- Margaritifera_falcata **
Aunins et al., 2018 [Publ 3863], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
39 |
1 |
1 |
-- |
+----- Cumberlandia_monodonta
--+
| +-- Margaritifera_falcata **
+--+
+-- Margaritifera_dahurica **
Wu et al., 2019 [Publ 3864], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
45 |
1 |
1 |
-- |
+----- Cumberlandia_monodonta
--+
| +-- Margaritifera_dahurica **
+--+
+-- Margaritifera_falcata **
Cha et al., 2018 [Publ 3916], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
37 |
1 |
1 |
-- |
+----- Pseudunio_marocanus
--+
| +-- Margaritifera_falcata **
+--+
+-- Margaritifera_dahurica **
Froufe et al., 2019 [Publ 3936], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
43 |
1 |
1 |
-- |
+----- Pseudunio_marocanus
--+
| +-- Margaritifera_dahurica **
+--+
+-- Margaritifera_falcata **
Froufe et al., 2019 [Publ 3936], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
43 |
1 |
1 |
-- |
+----- Pseudunio_marocanus
--+
| +-- Margaritifera_dahurica **
+--+
+-- Margaritifera_falcata **
Pfeiffer et al., 2019 [Publ 3937], fig. 4-5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
55 |
0 |
0 |
-- |
+-- Margaritifera_margaritifera **
--+
+-- Margaritifera_hembeli **
Pfeiffer et al., 2019 [Publ 3937], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
55 |
0 |
0 |
-- |
+-- Margaritifera_margaritifera **
--+
+-- Margaritifera_hembeli **
Lopes-Lima et al., 2020 [Publ 4080], fig. 10
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
12 |
0 |
0 |
-- |
+-- Margaritifera_middendorffi **
--+
+-- Margaritifera_laevis **
Lopes-Lima et al., 2020 [Publ 4080], fig. 11
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
10 |
0 |
0 |
-- |
+-- Margaritifera_laevis **
--+
+-- Margaritifera_middendorffi **
Lopes-Lima et al., 2020 [Publ 4080], fig. 12
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
7 |
0 |
0 |
-- |
+-- Margaritifera_laevis **
--+
+-- Margaritifera_middendorffi **
Lopes-Lima et al., 2020 [Publ 4080], fig. 13
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
8 |
0 |
0 |
-- |
+-- Margaritifera_laevis **
--+
+-- Margaritifera_middendorffi **
Lopes-Lima et al., 2020 [Publ 4080], fig. 14
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
4 |
0 |
0 |
-- |
+-- Margaritifera_laevis **
--+
+-- Margaritifera_middendorffi **
Lopes-Lima et al., 2020 [Publ 4080], fig. 16
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
7 |
0 |
0 |
-- |
+-- Margaritifera_laevis **
--+
+-- Margaritifera_middendorffi **
Lopes-Lima et al., 2020 [Publ 4080], fig. 17
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
13 |
0 |
0 |
-- |
+-- Margaritifera_laevis **
--+
+-- Margaritifera_middendorffi **
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
120 |
0 |
0 |
-- |
+-- Margaritifera_laevis **
--+
+-- Margaritifera_middendorffi **
Lopes-Lima et al., 2020 [Publ 4080], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
7 |
0 |
0 |
-- |
+-- Margaritifera_laevis **
--+
+-- Margaritifera_middendorffi **
Lopes-Lima et al., 2020 [Publ 4080], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
10 |
0 |
0 |
-- |
+-- Margaritifera_laevis **
--+
+-- Margaritifera_middendorffi **
Lopes-Lima et al., 2020 [Publ 4080], fig. 7
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
6 |
0 |
0 |
-- |
+-- Margaritifera_laevis **
--+
+-- Margaritifera_dahurica **
Lopes-Lima et al., 2020 [Publ 4080], fig. 8
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
5 |
0 |
0 |
-- |
+-- Margaritifera_laevis **
--+
+-- Margaritifera_middendorffi **
Lopes-Lima et al., 2020 [Publ 4080], fig. 9
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
4 |
0 |
0 |
-- |
+-- Margaritifera_laevis **
--+
+-- Margaritifera_middendorffi **
Wu et al., 2020 [Publ 4121], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
48 |
1 |
1 |
-- |
+----- Cumberlandia_monodonta
--+
| +-- Margaritifera_falcata **
+--+
+-- Margaritifera_dahurica **
Sano et al., 2020 [Publ 4138], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
46 |
0 |
0 |
-- |
+-- Margaritifera_middendorffi **
--+
+-- Margaritifera_laevis **
Wu et al., 2020 [Publ 4147], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
12 |
0 |
0 |
-- |
+-- Margaritifera_margaritifera **
--+
+-- Margaritifera_falcata **
Zieritz et al., 2021 [Publ 4231], fig. 3-F, 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
47 |
1 |
1 |
-- |
+----- Pseudunio_marocanus
--+
| +-- Margaritifera_falcata **
+--+
+-- Margaritifera_dahurica **
Pavan-Kumar et al., 2022 [Publ 4263], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
34 |
0 |
0 |
-- |
+-- Margaritifera_dahurica **
--+
+-- Margaritifera_falcata **
Pavan-Kumar et al., 2022 [Publ 4263], fig. S5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 2 |
2 |
0 |
2 |
37 |
0 |
0 |
-- |
+-- Margaritifera_dahurica **
--+
+-- Margaritifera_falcata **
Nagel et al., 1998 [Publ 1946], fig. 9
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
5 |
0 |
0 |
-- |
-- Margaritifera
Graf & Cummings, 2006 [Publ 1968], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
46 |
1 |
1 |
-- |
+-- Margaritifera_margaritifera **
--+
+-- Cumberlandia_monodonta
Graf, 2000 [Publ 2647], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
17 |
0 |
0 |
-- |
-- Margaritifera
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
29 |
1 |
1 |
-- |
+-- Margaritifera_margaritifera **
--+
+-- Cumberlandia_monodonta
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
29 |
1 |
1 |
-- |
+-- Margaritifera_margaritifera **
--+
+-- Cumberlandia_monodonta
Bogan & Hoeh, 2000 [Publ 2650], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
33 |
1 |
1 |
-- |
+-- Cumberlandia_monodonta
--+
+-- Margaritifera_margaritifera **
Hoeh et al., 2001 [Publ 2652], fig. 14.2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
30 |
1 |
1 |
-- |
+-- Cumberlandia_monodonta
--+
+-- Margaritifera_margaritifera **
Hoeh et al., 2001 [Publ 2652], fig. 14.4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
30 |
1 |
1 |
-- |
+-- Cumberlandia_monodonta
--+
+-- Margaritifera_margaritifera **
Hoeh et al., 2002 [Publ 2653], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
30 |
1 |
1 |
-- |
+-- Margaritifera_margaritifera **
--+
+-- Cumberlandia_monodonta
Roe & Hoeh, 2003 [Publ 2655], fig. 4.2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
30 |
1 |
1 |
-- |
+-- Cumberlandia_monodonta
--+
+-- Margaritifera_margaritifera **
Roe & Hoeh, 2003 [Publ 2655], fig. 4.3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
30 |
1 |
1 |
-- |
+-- Cumberlandia_monodonta
--+
+-- Margaritifera_margaritifera **
Huang et al., 2002 [Publ 2657], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
33 |
1 |
1 |
-- |
-- Margaritifera_margaritifera
Huang et al., 2002 [Publ 2657], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
33 |
1 |
1 |
-- |
-- Margaritifera_margaritifera
Zhou et al., 2007 [Publ 2658], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
22 |
1 |
1 |
-- |
+-- Margaritifera_margaritifera **
--+
+-- Cumberlandia_monodonta
Campbell et al., 2005 [Publ 2660], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
116 |
1 |
1 |
-- |
+-- Margaritifera_margaritifera **
--+
+-- Cumberlandia_monodonta
Campbell et al., 2005 [Publ 2660], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
114 |
1 |
1 |
-- |
+-- Cumberlandia_monodonta
--+
+-- Margaritifera_margaritifera **
Curole & Kocher, 2005 [Publ 2682], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
10 |
0 |
0 |
-- |
-- Margaritifera_hembeli
Curole & Kocher, 2005 [Publ 2682], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
11 |
0 |
0 |
-- |
-- Margaritifera_hembeli
Källersjö et al., 2005 [Publ 2684], fig. 1A
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
6 |
0 |
0 |
-- |
-- Margaritifera_margaritifera
Källersjö et al., 2005 [Publ 2684], fig. 1B-F
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
6 |
0 |
0 |
-- |
-- Margaritifera_margaritifera
Machordom et al., 2003 [Publ 2688], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
2 |
1 |
1 |
-- |
+-- Margaritifera_margaritifera **
--+
+-- Pseudunio_auricularius
Machordom et al., 2003 [Publ 2688], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
3 |
2 |
2 |
-- |
+----- Cumberlandia_monodonta
--+
| +-- Margaritifera_margaritifera **
+--+
+-- Pseudunio_auricularius
Hoeh et al., 1998 [Publ 2695], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
4 |
0 |
0 |
-- |
-- Margaritifera_margaritifera
Hoeh et al., 1998 [Publ 2695], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
4 |
0 |
0 |
-- |
-- Margaritifera_margaritifera
Taylor et al., 2007 [Publ 2696], fig. 1-3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
3 |
0 |
0 |
-- |
-- Margaritifera_margaritifera
Curole & Kocher, 2002 [Publ 2712], fig. 1-F
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
3 |
0 |
0 |
-- |
-- Margaritifera_hembeli
Hoeh et al., 2009 [Publ 2716], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
46 |
1 |
1 |
-- |
+-- Margaritifera_margaritifera **
--+
+-- Cumberlandia_monodonta
Hoeh et al., 2009 [Publ 2716], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
45 |
1 |
1 |
-- |
+-- Margaritifera_margaritifera **
--+
+-- Cumberlandia_monodonta
Hoeh et al., 2009 [Publ 2716], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
45 |
1 |
1 |
-- |
+-- Margaritifera_margaritifera **
--+
+-- Cumberlandia_monodonta
Turner et al., 2000 [Publ 2722], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
15 |
0 |
0 |
-- |
-- Margaritifera_margaritifera
Campbell, 2000 [Publ 2724], fig. 1-5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
2 |
0 |
0 |
-- |
-- Margaritifera_margaritifera
Gustafson & Iwamoto, 2005 [Publ 2741], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
4 |
0 |
0 |
-- |
-- Margaritifera_falcata
Whelan et al., 2011 [Publ 3380], fig. 1 ML
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
36 |
1 |
1 |
-- |
+-- Margaritifera_margaritifera **
--+
+-- Cumberlandia_monodonta
Whelan et al., 2011 [Publ 3380], fig. 1 MP
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
36 |
1 |
1 |
-- |
+-- Margaritifera_margaritifera **
--+
+-- Cumberlandia_monodonta
Prie et al., 2012 [Publ 3487], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
5 |
0 |
0 |
-- |
-- Margaritifera_margaritifera
Pfeiffer & Graf, 2013 [Publ 3531], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
34 |
0 |
0 |
-- |
-- Margaritifera_margaritifera
Reis et al., 2013 [Publ 3580], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
13 |
0 |
0 |
-- |
-- Margaritifera_margaritifera
Reis et al., 2013 [Publ 3580], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
15 |
0 |
0 |
-- |
-- Margaritifera_margaritifera
Graf et al., 2014 [Publ 3591], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
26 |
0 |
0 |
-- |
-- Margaritifera_margaritifera
Graf et al., 2015 [Publ 3619], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
19 |
0 |
0 |
-- |
-- Margaritifera_margaritifera
Pfeiffer & Graf, 2015 [Publ 3620], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
39 |
1 |
1 |
-- |
+-- Margaritifera_margaritifera **
--+
+-- Cumberlandia_monodonta
Bieler et al., 2014 [Publ 3622], fig. 26, 27, 29, 30
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
4 |
0 |
0 |
-- |
-- Margaritifera_margaritifera
Bieler et al., 2014 [Publ 3622], fig. 28
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
4 |
0 |
0 |
-- |
-- Margaritifera_margaritifera
Huang et al., 2013 [Publ 3629], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
15 |
0 |
0 |
-- |
-- Margaritifera_falcata
González & Giribet, 2015 [Publ 3632], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
5 |
0 |
0 |
-- |
-- Margaritifera_margaritifera
González & Giribet, 2015 [Publ 3632], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
5 |
0 |
0 |
-- |
-- Margaritifera_margaritifera
González & Giribet, 2015 [Publ 3632], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
5 |
0 |
0 |
-- |
-- Margaritifera_margaritifera
González & Giribet, 2015 [Publ 3632], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
5 |
0 |
0 |
-- |
-- Margaritifera_margaritifera
Giribet & Distel, 2003 [Publ 3670], fig. 3.5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
5 |
0 |
0 |
-- |
-- Margaritifera_margaritifera
Zieritz et al., 2012 [Publ 3673], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
7 |
0 |
0 |
-- |
-- Margaritifera_margaritifera
Zhou et al., 2016 [Publ 3710], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
19 |
0 |
0 |
-- |
-- Margaritifera_falcata
Bolotov et al., 2016 [Publ 3732], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
7 |
0 |
0 |
-- |
-- Margaritifera_dahurica
Lopes-Lima et al., 2017 [Publ 3734], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
16 |
1 |
1 |
-- |
+-- Pseudunio_auricularius
--+
+-- Margaritifera_margaritifera **
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
80 |
1 |
1 |
-- |
+-- Cumberlandia_monodonta
--+
+-- Margaritifera_margaritifera **
Hewitt et al., 2018 [Publ 3758], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
8 |
0 |
0 |
-- |
-- Margaritifera_margaritifera
Guerra et al., 2017 [Publ 3772], fig. 2a
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
24 |
1 |
1 |
-- |
+-- Margaritifera_falcata **
--+
+-- Cumberlandia_monodonta
Guerra et al., 2017 [Publ 3772], fig. 2b
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
24 |
1 |
1 |
-- |
+-- Margaritifera_falcata **
--+
+-- Cumberlandia_monodonta
Amaro et al., 2017 [Publ 3791], fig. 3A-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
6 |
0 |
0 |
-- |
-- Margaritifera_margaritifera
Amaro et al., 2017 [Publ 3791], fig. 3B-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
6 |
0 |
0 |
-- |
-- Margaritifera_margaritifera
Välilä et al., 2017 [Publ 3793], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
2 |
1 |
1 |
-- |
+-- Margaritifera_margaritifera **
--+
+-- Pseudunio_auricularius
Froufe et al., 2017 [Publ 3802], fig. 7
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
9 |
0 |
0 |
-- |
-- Margaritifera_margaritifera
Combosch et al., 2017 [Publ 3806], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
14 |
0 |
0 |
-- |
-- Margaritifera_margaritifera
Combosch et al., 2017 [Publ 3806], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
14 |
0 |
0 |
-- |
-- Margaritifera_margaritifera
Vikhrev et al., 2017 [Publ 3810], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
9 |
0 |
0 |
-- |
-- Margaritifera_dahurica
Bespalaya et al., 2018 [Publ 3811], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
9 |
0 |
0 |
-- |
-- Margaritifera_dahurica
Pfeiffer et al., 2018 [Publ 3827], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
28 |
0 |
0 |
-- |
-- Margaritifera_margaritifera
Huang et al., 2019 [Publ 3843], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
43 |
0 |
0 |
-- |
-- Margaritifera_dahurica
Huang et al., 2019 [Publ 3843], fig. 3-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
22 |
1 |
1 |
-- |
+-- Pseudunio_marocanus
--+
+-- Margaritifera_margaritifera **
Huang et al., 2019 [Publ 3843], fig. S1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
74 |
0 |
0 |
-- |
-- Margaritifera_dahurica
Bogatov et al., 2018 [Publ 3859], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
5 |
0 |
0 |
-- |
-- Margaritifera_margaritifera
Bogatov et al., 2018 [Publ 3859], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
5 |
0 |
0 |
-- |
-- Margaritifera_margaritifera
Klishko et al., 2019 [Publ 3886], fig. 9
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
10 |
0 |
0 |
-- |
-- Margaritifera_dahurica
Konopleva et al., 2019 [Publ 3900], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
51 |
0 |
0 |
-- |
-- Margaritifera_dahurica
Lopes-Lima et al., 2019 [Publ 3901], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
27 |
1 |
1 |
-- |
+-- Margaritifera_falcata **
--+
+-- Cumberlandia_monodonta
Lopes-Lima et al., 2019 [Publ 3901], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
28 |
1 |
1 |
-- |
+-- Cumberlandia_monodonta
--+
+-- Margaritifera_falcata **
Lopes-Lima et al., 2019 [Publ 3901], fig. 4
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
25 |
1 |
1 |
-- |
+-- Margaritifera_falcata **
--+
+-- Cumberlandia_monodonta
Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
5 |
2 |
2 |
-- |
+----- Margaritifera_margaritifera **
--+
| +-- Pseudunio_marocanus
+--+
+-- Cumberlandia_monodonta
Hewitt et al., 2019 [Publ 3962], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
15 |
0 |
0 |
-- |
-- Margaritifera_margaritifera
Klunzinger et al., 2021 [Publ 4086], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
5 |
0 |
0 |
-- |
-- Margaritifera_margaritifera
Lemer et al., 2019 [Publ 4093], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
5 |
0 |
0 |
-- |
-- Margaritifera_margaritifera
Wu et al., 2020 [Publ 4147], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
29 |
0 |
0 |
-- |
-- Margaritifera_falcata
Guerra et al., 2019 [Publ 4168], fig. 1-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
21 |
2 |
2 |
-- |
+----- Cumberlandia_monodonta
--+
| +-- Margaritifera_margaritifera **
+--+
+-- Pseudunio_marocanus
Guerra et al., 2019 [Publ 4168], fig. 2-M
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
21 |
2 |
2 |
-- |
+----- Pseudunio_marocanus
--+
| +-- Margaritifera_margaritifera **
+--+
+-- Cumberlandia_monodonta
Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
18 |
1 |
1 |
-- |
+-- Margaritifera_falcata **
--+
+-- Cumberlandia_monodonta
Kano et al., 2019 [Publ 4194], fig. 6
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
9 |
0 |
0 |
-- |
-- Margaritifera_falcata
Zhang et al., 2013 [Publ 4214], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
3 |
0 |
0 |
-- |
-- Margaritifera_margaritifera
Zieritz et al., 2021 [Publ 4218], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
31 |
1 |
1 |
-- |
+-- Margaritifera_margaritifera **
--+
+-- Pseudunio_auricularius
Bolotov et al., 2022 [Publ 4250], fig. 8
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
48 |
0 |
0 |
-- |
-- Margaritifera_dahurica
Dai et al., 2023 [Publ 4285], fig. 3-BI
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
44 |
0 |
0 |
-- |
-- Margaritifera_margaritifera
Dai et al., 2023 [Publ 4285], fig. 3-ML
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
44 |
0 |
0 |
-- |
-- Margaritifera_margaritifera
Dai et al., 2023 [Publ 4285], fig. 5
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
52 |
0 |
0 |
-- |
-- Margaritifera_dahurica
Bogan et al., 2023 [Publ 4295], fig. 3
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
30 |
0 |
0 |
-- |
-- Margaritifera_laevis
Chen et al., 2023 [Publ 4302], fig. 1
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
39 |
0 |
0 |
-- |
-- Margaritifera_dahurica
Chen et al., 2023 [Publ 4302], fig. 2
| iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
| 1 |
1 |
0 |
1 |
39 |
0 |
0 |
-- |
-- Margaritifera_dahurica
| 
