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Page last updated
Wed 06 Nov 2024

The Freshwater Mussels (Unionoida) of the World (and other less consequential bivalves)

FM(U)otW(aolcb) is the web version of the MUSSEL Project Database. Follow the links to browse the data or use the custom Google search field. Either way, you win!

Cladomics pages are experimental — even more so than the rest of this site!

The purpose of the cladomics pages for each genus and family-group level taxon is to provide statistics and references indicating the extent to which published phylogenetic work supports the classification used here.

The Cladomics Summary provides an overview of the number of studies that have included terminal species representing this taxon as well as an indication of how frequently it has been recovered as monophyletic.

Following the summary, the available published clades representing the taxon are listed.

For more information, visit the Cladomics FARQ (frequently asked rhetorical questions) page. Abbreviations for the various statistics are explained on the FARQ!

Search the database pages:

Bivalvia | Unionoida | Unionoidea | Unionidae | Ambleminae

tribe LAMPSILINI Ihering, 1901: cladomics
     31 genera

Cladomics Summary

number of species = 186
number of subtaxa (rank: genus) = 31

analyzed in 256 cladograms in 151 publications
maximum number of terminals in an analysis = 48, mean = 5.81
maximum number of subtaxa (genus) in an analysis = 23, mean = 3.9

Basic Monophyly

number of cladograms that test basic monophyly (i.e., more than 1 included species) = 187
maximum number of included terminals = 48, mean = 7.58
maximum number of subtaxa (genus) = 23, mean = 4.97
proportion of cladograms that support basic monophyly = 0.76

Rigorous Monophyly

number of cladograms that test rigorous monophyly = 160
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (genus) taxon and at least one rigorous outgroup (tribe).
maximum number of included terminals = 48, mean = 7.84
maximum number of subtaxa (genus) = 23, mean = 5.34
proportion of cladograms that support rigorous monophyly = 0.72


Pairwise Cladogram Clustering

The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.


PCoA Ordination


NJ tree


Campbell et al., 2005 [Publ 2660], fig. 1

iT tT bi riS oT roT roS iD
43 43 46 23 59 47 4 0.0

*** Top Monophyly Cladogram ***

  +----------------------------------------- Quadrulini                    
  |                                                                        
  |  +-------------------------------------- Pleurobemini                  
--+  |                                                                     
  |  |                                   +-- Plectomerus                   
  |  |  +--------------------------------+                                 
  +--+  |                                +-- Reginaia                      
     |  |                                                                  
     |  |                                +-- Popenaiadini                  
     |  |  +-----------------------------+                                 
     +--+  |                             +-- Amblema                       
        |  |                                                               
        |  |                       +-------- Toxolasma **                     
        |  |  +--------------------+                                       
        |  |  |                    |  +----- Obliquaria **                    
        |  |  |                    +--+                                    
        +--+  |                       |  +-- Cyrtonaias **                    
           |  |                       +--+                                 
           |  |                          +-- Glebula **                       
           |  |                                                            
           |  |              +-------------- Potamilus **                     
           |  |              |                                             
           |  |              |     +-------- Medionidus_acutissimus **        
           +--+              |  +--+                                       
              |  +-----------+  |  |  +----- Lemiox **                        
              |  |           |  |  +--+                                    
              |  |           |  |     |  +-- Dromus **                        
              |  |           |  |     +--+                                 
              |  |           +--+        +-- Cyprogenia **                    
              |  |              |                                          
              |  |              |        +-- Sagittunio **                    
              |  |              |     +--+                                 
              |  |              |     |  +-- Medionidus_conradicus **         
              +--+              +-----+                                    
                 |                    |  +-- Ptychobranchus **                
                 |                    +--+                                 
                 |                       +-- Venustaconcha_ellipsiformis__2 **
                 |                                                         
                 |                       +-- Ellipsaria **                    
                 |  +--------------------+                                 
                 |  |                    +-- Truncilla **                     
                 |  |                                                      
                 |  |  +-------------------- Villosa **                       
                 +--+  |                                                   
                    |  |                 +-- Obovaria_olivaria **             
                    |  |        +--------+                                 
                    |  |        |        +-- Venustaconcha_ellipsiformis__1 **
                    |  |  +-----+                                          
                    +--+  |     |  +-------- Epioblasma **                    
                       |  |     +--+                                       
                       |  |        |  +----- Obovaria_unicolor **             
                       |  |        +--+                                    
                       |  |           |  +-- Obovaria_arkansasensis **        
                       |  |           +--+                                 
                       |  |              +-- Obovaria_subrotunda **           
                       +--+                                                
                          |        +-------- Hamiota **                       
                          |        |                                       
                          |  +-----+     +-- Lampsilis_cardium **             
                          |  |     |  +--+                                 
                          |  |     |  |  +-- Lampsilis_ovata **               
                          |  |     +--+                                    
                          |  |        |  +-- Ortmanniana_pectorosa **         
                          +--+        +--+                                 
                             |           +-- Lampsilis_ornata **              
                             |                                             
                             |           +-- Ligumia **                       
                             |  +--------+                                 
                             |  |        +-- Cambarunio **                    
                             +--+                                          
                                |  +-------- Lampsilis_teres **               
                                +--+                                       
                                   |  +----- Ortmanniana_ligamentina **       
                                   +--+                                    
                                      |  +-- Lampsilis_siliquoidea **         
                                      +--+                                 
                                         +-- Leaunio **                       

Inoue et al., 2018 [Publ 3858], fig. 1_3a-c

iT tT bi riS oT roT roS iD
11 12 0 8 93 71 4 51.0

*** Top Sister-Group Cladogram ***

  +-------- Reginaia           
  |                            
  +-------- Amblema            
  |                            
  |     +-- Quadrulini         
  +-----+                      
  |     +-- Pleurobemini       
--+                            
  |  +----- Popenaiadini       
  |  |                         
  |  |  +-- Dromus **             
  |  |  |                      
  |  |  +-- Epioblasma **         
  |  |  |                      
  +--+  +-- Glebula **            
     |  |                      
     |  +-- Lampsilis_ovata **    
     |  |                      
     |  +-- Lampsilis_virescens **
     |  |                      
     |  +-- Lemiox **             
     +--+                      
        +-- Obliquaria **         
        |                      
        +-- Obovaria_olivaria **  
        |                      
        +-- Obovaria_unicolor **  
        |                      
        +-- Plectomerus        
        |                      
        +-- Potamilus_alatus **   
        |                      
        +-- Potamilus_capax **    

Zanatta & Murphy, 2006 [Publ 2672], fig. 1

iT tT bi riS oT roT roS iD
48 48 45 23 8 6 3 47.0
                                   +----- Pleurobemini           
  +--------------------------------+                             
  |                                |  +-- Amblemini              
  |                                +--+                          
  |                                   +-- Popenaiadini           
--+                                                              
  |                                   +-- Cyrtonaias **             
  |  +--------------------------------+                          
  |  |                                +-- Glebula **                
  |  |                                                           
  +--+  +-------------------------------- Paetulunio **             
     |  |                                                        
     |  |                             +-- Obliquaria **             
     +--+  +--------------------------+                          
        |  |                          +-- Toxolasma **              
        |  |                                                     
        |  |  +-------------------------- Sagittunio **             
        +--+  |                                                  
           |  |              +----------- Potamilus **              
           |  |              |                                   
           |  |  +-----------+        +-- Medionidus_conradicus **  
           |  |  |           |  +-----+                          
           +--+  |           |  |     +-- Ptychobranchus **         
              |  |           +--+                                
              |  |              |  +----- Medionidus_acutissimus ** 
              |  |              |  |                             
              |  |              +--+----- Lemiox **                 
              |  |                 |                             
              |  |                 |  +-- Dromus **                 
              +--+                 +--+                          
                 |                    +-- Cyprogenia **             
                 |                                               
                 |                 +----- Lampsilis_teres **        
                 |  +--------------+                             
                 |  |              |  +-- Truncilla **              
                 |  |              +--+                          
                 |  |                 +-- Ellipsaria **             
                 |  |                                            
                 +--+  +----------------- Leaunio **                
                    |  |                                         
                    |  |     +----------- Hamiota **                
                    |  |     |                                   
                    |  |  +--+        +-- Lampsilis_fasciola **     
                    +--+  |  |  +-----+                          
                       |  |  |  |     +-- Ortmanniana_pectorosa **  
                       |  |  +--+                                
                       |  |     |  +----- Lampsilis_ornata **       
                       |  |     +--+                             
                       |  |        |  +-- Lampsilis_ovata **        
                       +--+        +--+                          
                          |           +-- Lampsilis_cardium **      
                          |                                      
                          |        +----- Venustaconcha **          
                          |  +-----+                             
                          |  |     |  +-- Obovaria **               
                          |  |     +--+                          
                          +--+        +-- Epioblasma **             
                             |                                   
                             |  +-------- Lampsilis_siliquoidea **  
                             +--+                                
                                |  +----- Ortmanniana_ligamentina **
                                +--+                             
                                   |  +-- Cambarunio **             
                                   +--+                          
                                      +-- Ligumia **                

Zanatta & Murphy, 2006 [Publ 2672], fig. 2

iT tT bi riS oT roT roS iD
48 48 42 23 9 7 3 45.0
  +-------------------------------------------- Pleurobemini           
  |                                                                    
  |  +----------------------------------------- Amblemini              
--+  |                                                                 
  |  +----------------------------------------- Popenaiadini           
  |  |                                                                 
  |  |                                      +-- Paetulunio **             
  +--+  +-----------------------------------+                          
     |  |                                   +-- Toxolasma **              
     |  |                                                              
     |  |                                +----- Obliquaria **             
     |  |  +-----------------------------+                             
     +--+  |                             |  +-- Glebula **                
        |  |                             +--+                          
        |  |                                +-- Cyrtonaias **             
        |  |                                                           
        |  |                             +----- Ptychobranchus **         
        |  |                          +--+                             
        +--+                          |  |  +-- Dromus **                 
           |                          |  +--+                          
           |  +-----------------------+     +-- Cyprogenia **             
           |  |                       |                                
           |  |                       |  +----- Medionidus_conradicus **  
           |  |                       +--+                             
           |  |                          |  +-- Medionidus_acutissimus ** 
           |  |                          +--+                          
           +--+                             +-- Lemiox **                 
              |                                                        
              |  +----------------------------- Sagittunio **             
              |  |                                                     
              |  +----------------------------- Potamilus **              
              |  |                                                     
              |  |                          +-- Truncilla **              
              +--+  +-----------------------+                          
                 |  |                       +-- Ellipsaria **             
                 |  |                                                  
                 |  |                       +-- Ortmanniana_pectorosa **  
                 |  |                 +-----+                          
                 |  |                 |     +-- Lampsilis_fasciola **     
                 +--+  +--------------+                                
                    |  |              |  +----- Lampsilis_ornata **       
                    |  |              +--+                             
                    |  |                 |  +-- Lampsilis_ovata **        
                    |  |                 +--+                          
                    |  |                    +-- Lampsilis_cardium **      
                    +--+                                               
                       |                    +-- Ligumia **                
                       |                 +--+                          
                       |                 |  +-- Cambarunio **             
                       |  +--------------+                             
                       |  |              |  +-- Lampsilis_teres **        
                       |  |              +--+                          
                       +--+                 +-- Hamiota **                
                          |                                            
                          |  +----------------- Leaunio **                
                          |  |                                         
                          +--+  +-------------- Ortmanniana_ligamentina **
                             |  |                                      
                             +--+  +----------- Lampsilis_siliquoidea **  
                                |  |                                   
                                +--+  +-------- Obovaria_olivaria **      
                                   |  |                                
                                   |  +-------- Venustaconcha **          
                                   +--+                                
                                      +-------- Epioblasma **             
                                      |                                
                                      |  +----- Obovaria_unicolor **      
                                      +--+                             
                                         |  +-- Obovaria_subrotunda **    
                                         +--+                          
                                            +-- Obovaria_arkansasensis ** 

Campbell et al., 2005 [Publ 2660], fig. 2

iT tT bi riS oT roT roS iD
43 44 37 23 59 47 4 38.0
  +-------------------------- Quadrulini                    
  |                                                         
  |  +----------------------- Pleurobemini                  
--+  |                                                      
  |  |                 +----- Popenaiadini                  
  |  |  +--------------+                                    
  |  |  |              |  +-- Amblema                       
  +--+  |              +--+                                 
     |  |                 +-- Reginaia                      
     |  |                                                   
     |  |                 +-- Plectomerus                   
     |  |  +--------------+                                 
     +--+  |              +-- Toxolasma **                     
        |  |                                                
        |  |           +----- Obliquaria **                    
        |  |  +--------+                                    
        |  |  |        |  +-- Cyrtonaias **                    
        |  |  |        +--+                                 
        |  |  |           +-- Glebula **                       
        |  |  |                                             
        +--+  |  +----------- Sagittunio **                    
           |  |  |                                          
           |  |  +----------- Leaunio **                       
           |  |  |                                          
           |  |  +----------- Potamilus **                     
           |  |  |                                          
           |  |  +----------- Hamiota **                       
           |  |  |                                          
           |  |  |        +-- Ellipsaria **                    
           +--+  +--------+                                 
              |  |        +-- Truncilla **                     
              |  |                                          
              |  |     +----- Lampsilis_teres **               
              |  +-----+                                    
              |  |     |  +-- Ortmanniana_ligamentina **       
              |  |     +--+                                 
              |  |        +-- Lampsilis_siliquoidea **         
              |  |                                          
              |  |     +----- Villosa **                       
              |  +-----+                                    
              |  |     |  +-- Ligumia **                       
              |  |     +--+                                 
              |  |        +-- Cambarunio **                    
              |  |                                          
              +--+  +-------- Ortmanniana_pectorosa **         
                 +--+                                       
                 |  |  +----- Lampsilis_ornata **              
                 |  +--+                                    
                 |     |  +-- Lampsilis_cardium **             
                 |     +--+                                 
                 |        +-- Lampsilis_ovata **               
                 |                                          
                 |  +-------- Obovaria_olivaria **             
                 |  |                                       
                 |  +-------- Venustaconcha_ellipsiformis__2 **
                 +--+                                       
                 |  +-------- Epioblasma **                    
                 |  |                                       
                 |  |  +----- Obovaria_unicolor **             
                 |  +--+                                    
                 |     |  +-- Obovaria_arkansasensis **        
                 |     +--+                                 
                 |        +-- Obovaria_subrotunda **           
                 |                                          
                 |        +-- Ptychobranchus **                
                 |  +-----+                                 
                 |  |     +-- Venustaconcha_ellipsiformis__1 **
                 |  |                                       
                 +--+  +----- Medionidus_conradicus **         
                    |  |                                    
                    |  |  +-- Lemiox **                        
                    +--+--+                                 
                       |  +-- Medionidus_acutissimus **        
                       |                                    
                       |  +-- Dromus **                        
                       +--+                                 
                          +-- Cyprogenia **                    

Smith et al., 2019 [Publ 3951], fig. 2

iT tT bi riS oT roT roS iD
36 37 36 20 5 5 3 58.0
  +-------------------------------------- Quadrulini             
  |                                                              
  |                                   +-- Amblema                
--+  +--------------------------------+                          
  |  |                                +-- Reginaia               
  |  |                                                           
  +--+  +-------------------------------- Plectomerus            
     |  |                                                        
     |  |  +----------------------------- Toxolasma **              
     +--+  |                                                     
        |  |                       +----- Popenaiadini           
        |  |  +--------------------+                             
        +--+  |                    |  +-- Glebula **                
           |  |                    +--+                          
           |  |                       +-- Cyrtonaias **             
           |  |                                                  
           +--+  +----------------------- Obliquaria **             
              |  |                                               
              |  |                    +-- Medionidus **             
              |  |                 +--+                          
              |  |                 |  +-- Lemiox **                 
              +--+  +--------------+                             
                 |  |              |  +-- Cyprogenia **             
                 |  |              +--+                          
                 |  |                 +-- Ptychobranchus **         
                 |  |                                            
                 +--+           +-------- Potamilus **              
                    |  +--------+                                
                    |  |        |  +----- Atlanticoncha **          
                    |  |        +--+                             
                    |  |           |  +-- Ellipsaria **             
                    |  |           +--+                          
                    +--+              +-- Truncilla **              
                       |                                         
                       |              +-- Hamiota **                
                       |  +-----------+                          
                       |  |           +-- Villosa **                
                       |  |                                      
                       +--+  +----------- Ligumia **                
                          |  |                                   
                          |  |     +----- Venustaconcha **          
                          +--+  +--+                             
                             |  |  |  +-- Obovaria **               
                             |  |  +--+                          
                             +--+     +-- Epioblasma **             
                                |                                
                                |  +----- Ortmanniana_pectorosa **  
                                +--+                             
                                   |  +-- Ortmanniana_ligamentina **
                                   +--+                          
                                      +-- Lampsilis **              

Inoue et al., 2020 [Publ 3941], fig. 3

iT tT bi riS oT roT roS iD
27 27 27 7 15 14 3 57.0
  +-------------------------------------------- Quadrulini           
  |                                                                  
--+  +----------------------------------------- Pleurobemini         
  |  |                                                               
  +--+  +-------------------------------------- Amblemini            
     |  |                                                            
     +--+  +----------------------------------- Toxolasma **            
        |  |                                                         
        +--+  +-------------------------------- Potamilus **            
           |  |                                                      
           |  |                             +-- Ellipsaria **           
           +--+  +--------------------------+                        
              |  |                          +-- Truncilla **            
              |  |                                                   
              |  |  +-------------------------- Leaunio **              
              +--+  |                                                
                 |  |     +-------------------- Lampsilis_bracteata **  
                 |  |     |                                          
                 |  |  +--+  +----------------- Lampsilis_fasciola **   
                 |  |  |  |  |                                       
                 +--+  |  +--+  +-------------- Lampsilis_ornata **     
                    |  |     |  |                                    
                    |  |     +--+           +-- Lampsilis_ovata **      
                    |  |        |  +--------+                        
                    |  |        |  |        +-- Lampsilis_cariosa **    
                    |  |        +--+                                 
                    |  |           |  +-------- Lampsilis_cardium__3 ** 
                    +--+           +--+                              
                       |              |  +----- Lampsilis_cardium__2 ** 
                       |              +--+                           
                       |                 |  +-- Lampsilis_satura **     
                       |                 +--+                        
                       |                    +-- Lampsilis_cardium__1 ** 
                       |                                             
                       |                    +-- Lampsilis_higginsii **  
                       |     +--------------+                        
                       |     |              +-- Ortmanniana **          
                       +-----+                                       
                             |  +-------------- Lampsilis_teres **      
                             |  |                                    
                             +--+           +-- Lampsilis_radiata **    
                                |  +--------+                        
                                |  |        +-- Lampsilis_siliquoidea **
                                +--+                                 
                                   |  +-------- Lampsilis_bergmanni **  
                                   +--+                              
                                      |  +----- Lampsilis_hydiana **    
                                      +--+                           
                                         |  +-- Lampsilis_virescens **  
                                         +--+                        
                                            +-- Lampsilis_straminea **  

Burlakova et al., 2019 [Publ 3899], fig. 1

iT tT bi riS oT roT roS iD
26 30 18 18 7 7 4 56.0
  +-------------------------- Quadrulini           
  |                                                
  |  +----------------------- Plectomerus          
--+  |                                             
  |  |  +-------------------- Popenaiadini         
  |  |  |                                          
  |  |  +-------------------- Pleurobemini         
  +--+  |                                          
     |  |                 +-- Cyrtonaias **           
     |  |              +--+                        
     |  |              |  +-- Glebula **              
     |  +--------------+                           
     |  |              |  +-- Reginaia             
     +--+              +--+                        
        |                 +-- Obliquaria **           
        |                                          
        |  +----------------- Toxolasma **            
        |  |                                       
        |  |  +-------------- Potamilus **            
        |  |  |                                    
        |  |  |           +-- Ellipsaria **           
        |  |  +-----------+                        
        +--+  |           +-- Truncilla **            
           |  |                                    
           |  |        +----- Medionidus **           
           |  +--------+                           
           |  |        |  +-- Lemiox **               
           |  |        +--+                        
           +--+           +-- Dromus **               
              |                                    
              |  +----------- Cambarunio **           
              |  |                                 
              |  +----------- Leaunio **              
              |  |                                 
              |  +----------- Villosa **              
              |  |                                 
              |  |        +-- Sagittunio **           
              |  +--------+                        
              +--+        +-- Lampsilis_ovata **      
                 |                                 
                 |     +----- Venustaconcha **        
                 +-----+                           
                 |     |  +-- Epioblasma **           
                 |     +--+                        
                 |        +-- Obovaria **             
                 |                                 
                 |  +-------- Lampsilis_teres **      
                 +--+                              
                    |  +----- Lampsilis_siliquoidea **
                    +--+                           
                       |  +-- Lampsilis_hydiana **    
                       +--+                        
                          +-- Lampsilis_straminea **  

Inoue et al., 2020 [Publ 3941], fig. 4

iT tT bi riS oT roT roS iD
26 26 26 7 15 14 3 61.0
  +----------------------------------------------- Quadrulini           
  |                                                                     
--+  +-------------------------------------------- Reginaia             
  |  |                                                                  
  +--+  +----------------------------------------- Pleurobemini         
     |  |                                                               
     +--+  +-------------------------------------- Amblema              
        |  |                                                            
        +--+  +----------------------------------- Toxolasma **            
           |  |                                                         
           +--+  +-------------------------------- Ellipsaria **           
              |  |                                                      
              +--+  +----------------------------- Truncilla **            
                 |  |                                                   
                 |  |  +-------------------------- Potamilus **            
                 +--+  |                                                
                    |  |  +----------------------- Leaunio **              
                    |  |  |                                             
                    +--+  |     +----------------- Lampsilis_bracteata **  
                       |  |     |                                       
                       |  |  +--+              +-- Lampsilis_ornata **     
                       |  |  |  |  +-----------+                        
                       +--+  |  |  |           +-- Lampsilis_fasciola **   
                          |  |  +--+                                    
                          |  |     |  +----------- Lampsilis_ovata **      
                          |  |     +--+                                 
                          |  |        |  +-------- Lampsilis_cardium__3 ** 
                          |  |        +--+                              
                          +--+           |  +----- Lampsilis_cardium__2 ** 
                             |           +--+                           
                             |              |  +-- Lampsilis_satura **     
                             |              +--+                        
                             |                 +-- Lampsilis_cardium__1 ** 
                             |                                          
                             |     +-------------- Lampsilis_teres **      
                             |     |                                    
                             |     |           +-- Lampsilis_higginsii **  
                             +-----+  +--------+                        
                                   |  |        +-- Ortmanniana **          
                                   |  |                                 
                                   +--+        +-- Lampsilis_radiata **    
                                      |  +-----+                        
                                      |  |     +-- Lampsilis_siliquoidea **
                                      |  |                              
                                      +--+     +-- Lampsilis_hydiana **    
                                         |  +--+                        
                                         |  |  +-- Lampsilis_bergmanni **  
                                         +--+                           
                                            |  +-- Lampsilis_virescens **  
                                            +--+                        
                                               +-- Lampsilis_straminea **  

Smith et al., 2020 [Publ 4129], fig. 2-3

iT tT bi riS oT roT roS iD
20 20 18 8 8 8 4 52.0
  +-------------------------- Quadrulini   
  |                                        
--+  +----------------------- Pleurobemini 
  |  |                                     
  +--+  +-------------------- Amblemini    
     |  |                                  
     +--+  +----------------- Popenaiadini 
        |  |                               
        +--+  +-------------- Obliquaria **   
           |  |                            
           +--+  +----------- Cyrtonaias **   
              |  |                         
              |  |        +-- Pachynaias **   
              +--+  +-----+                
                 |  |     +-- Lampsilis **    
                 |  |                      
                 +--+     +-- Ellipsaria **   
                    |  +--+                
                    |  |  +-- Truncilla **    
                    +--+                   
                       |  +-- Atlanticoncha **
                       +--+                
                          +-- Potamilus **    

Smith et al., 2020 [Publ 4129], fig. 1

iT tT bi riS oT roT roS iD
20 20 18 8 8 8 4 52.0
  +-------------------------- Quadrulini   
  |                                        
--+  +----------------------- Pleurobemini 
  |  |                                     
  +--+  +-------------------- Amblemini    
     |  |                                  
     +--+  +----------------- Popenaiadini 
        |  |                               
        +--+  +-------------- Obliquaria **   
           |  |                            
           +--+  +----------- Cyrtonaias **   
              |  |                         
              |  |        +-- Pachynaias **   
              +--+  +-----+                
                 |  |     +-- Lampsilis **    
                 |  |                      
                 +--+     +-- Ellipsaria **   
                    |  +--+                
                    |  |  +-- Truncilla **    
                    +--+                   
                       |  +-- Atlanticoncha **
                       +--+                
                          +-- Potamilus **    

Porto-Hannes et al., 2019 [Publ 3969], fig. 2

iT tT bi riS oT roT roS iD
17 17 15 6 1 1 1 54.0
  +-------------------------- Amblemini            
  |                                                
  |  +----------------------- Obliquaria **           
--+  |                                             
  |  |                    +-- Cambarunio **           
  |  |  +-----------------+                        
  |  |  |                 +-- Leaunio **              
  +--+  |                                          
     |  |        +----------- Lampsilis_teres **      
     |  |        |                                 
     |  +--------+  +-------- Ortmanniana **          
     |  |        |  |                              
     |  |        +--+     +-- Lampsilis_hydiana **    
     +--+           |  +--+                        
        |           |  |  +-- Lampsilis_virescens **  
        |           +--+                           
        |              |  +-- Lampsilis_siliquoidea **
        |              +--+                        
        |                 +-- Lampsilis_radiata **    
        |                                          
        |  +----------------- Obovaria **             
        |  |                                       
        +--+  +-------------- Lampsilis_bracteata **  
           |  |                                    
           +--+  +----------- Lampsilis_ornata **     
              |  |                                 
              +--+  +-------- Lampsilis_ovata **      
                 |  |                              
                 +--+  +----- Lampsilis_cardium__3 ** 
                    |  |                           
                    +--+  +-- Lampsilis_satura **     
                       |  |                        
                       +--+-- Lampsilis_cardium__1 ** 
                          |                        
                          +-- Lampsilis_cardium__2 ** 

Porto-Hannes et al., 2019 [Publ 3969], fig. 3

iT tT bi riS oT roT roS iD
17 18 13 6 1 1 1 52.0
  +-------------------- Obliquaria **           
  |                                          
  +-------------------- Amblemini            
  |                                          
  |  +----------------- Lampsilis_teres **      
  |  |                                       
  |  +----------------- Obovaria **             
--+  |                                       
  |  |              +-- Leaunio **              
  |  +--------------+                        
  |  |              +-- Cambarunio **           
  |  |                                       
  |  |        +-------- Ortmanniana **          
  |  |        |                              
  +--+--------+     +-- Lampsilis_hydiana **    
     |        |  +--+                        
     |        |  |  +-- Lampsilis_virescens **  
     |        +--+                           
     |           |  +-- Lampsilis_siliquoidea **
     |           +--+                        
     |              +-- Lampsilis_radiata **    
     |                                       
     |  +-------------- Lampsilis_bracteata **  
     |  |                                    
     +--+  +----------- Lampsilis_ornata **     
        |  |                                 
        +--+  +-------- Lampsilis_ovata **      
           |  |                              
           +--+  +----- Lampsilis_cardium__3 ** 
              |  |                           
              +--+  +-- Lampsilis_satura **     
                 |  |                        
                 +--+-- Lampsilis_cardium__1 ** 
                    |                        
                    +-- Lampsilis_cardium__2 ** 

Boyer et al., 2011 [Publ 2718], fig. 4

iT tT bi riS oT roT roS iD
16 22 18 10 22 13 3 49.0
  +-------------------------------- Quadrulina           
  |                                                      
--+  +----------------------------- Megalonaias          
  |  |                                                   
  +--+  +-------------------------- Toxolasma **            
     |  |                                                
     +--+  +----------------------- Amblema              
        |  |                                             
        |  |  +-------------------- Pleurobemini         
        +--+  |                                          
           |  |                 +-- Reginaia             
           |  +-----------------+                        
           +--+                 +-- Obliquaria **           
              |                                          
              |  +----------------- Potamilus **            
              |  |                                       
              |  |              +-- Ellipsaria **           
              +--+  +-----------+                        
                 |  |           +-- Truncilla **            
                 |  |                                    
                 +--+  +----------- Lampsilis_cardium **    
                    |  |                                 
                    |  |     +----- Ligumia **              
                    |  +-----+                           
                    +--+     |  +-- Obovaria **             
                       |     +--+                        
                       |        +-- Epioblasma **           
                       |                                 
                       |  +-------- Lampsilis_teres **      
                       +--+                              
                          |  +----- Lampsilis_siliquoidea **
                          +--+                           
                             |  +-- Ortmanniana **          
                             +--+                        
                                +-- Lampsilis_higginsii **  

Grobler et al., 2011 [Publ 3359], fig. 2

iT tT bi riS oT roT roS iD
16 16 12 10 1 1 1 45.0
  +----------------- Amblemini            
  |                                       
  |        +-------- Lemiox **               
  |  +-----+                              
--+  |     |  +----- Medionidus **           
  |  |     +--+                           
  |  |        |  +-- Dromus **               
  |  |        +--+                        
  +--+           +-- Cyprogenia **           
     |                                    
     |  +----------- Potamilus **            
     |  |                                 
     |  |     +----- Lampsilis_siliquoidea **
     |  |  +--+                           
     +--+  |  |  +-- Lampsilis_higginsii **  
        |  |  +--+                        
        |  |     +-- Ortmanniana **          
        |  |                              
        |  |  +----- Lampsilis_fasciola **   
        +--+--+                           
           |  |  +-- Lampsilis_cardium **    
           |  +--+                        
           |     +-- Lampsilis_ovata **      
           |                              
           |     +-- Cambarunio **           
           |     |                        
           +-----+-- Ligumia **              
                 |                        
                 +-- Epioblasma **           

Inoue et al., 2020 [Publ 3941], fig. 5

iT tT bi riS oT roT roS iD
16 16 14 2 0 0 0 51.0
  +-- Truncilla **
--+            
  +-- Lampsilis **

Boyer et al., 2011 [Publ 2718], fig. 3

iT tT bi riS oT roT roS iD
16 16 9 10 19 13 3 50.0
  +-------------------- Reginaia             
  |                                          
--+                 +-- Pleurobemini         
  |  +--------------+                        
  |  |              +-- Quadrulini           
  +--+                                       
     |  +-------------- Amblema              
     |  |                                    
     +--+  +----------- Toxolasma **            
        |  |                                 
        |  |  +-------- Obliquaria **           
        +--+  |                              
           |  |  +----- Obovaria **             
           |  |  |                           
           |  |  +----- Lampsilis_cardium **    
           +--+  |                           
              |  +----- Potamilus **            
              |  |                           
              |  |  +-- Epioblasma **           
              |  +--+                        
              |  |  +-- Lampsilis_siliquoidea **
              +--+                           
                 |  +-- Ortmanniana **          
                 +--+                        
                 |  +-- Lampsilis_higginsii **  
                 |                           
                 |  +-- Ligumia **              
                 +--+                        
                 |  +-- Lampsilis_teres **      
                 |                           
                 |  +-- Ellipsaria **           
                 +--+                        
                    +-- Truncilla **            

Szumowski et al., 2012 [Publ 3569], fig. 4

iT tT bi riS oT roT roS iD
15 15 11 10 1 1 1 48.0
  +-------------------------- Amblemini            
  |                                                
--+  +----------------------- Ellipsaria **           
  |  |                                             
  +--+  +-------------------- Truncilla **            
     |  |                                          
     +--+  +----------------- Obliquaria **           
        |  |                                       
        |  |  +-------------- Potamilus **            
        +--+  |                                    
           |  |        +----- Lampsilis_ornata **     
           |  |  +-----+                           
           |  |  |     |  +-- Lampsilis_cardium **    
           +--+  |     +--+                        
              |  |        +-- Lampsilis_ovata **      
              |  |                                 
              |  |  +-------- Lampsilis_teres **      
              |  +--+                              
              +--+  |  +----- Lampsilis_siliquoidea **
                 |  +--+                           
                 |     |  +-- Ortmanniana **          
                 |     +--+                        
                 |        +-- Lampsilis_higginsii **  
                 |                                 
                 |     +----- Hamiota **              
                 |     |                           
                 +-----+  +-- Epioblasma **           
                       |  |                        
                       +--+-- Cambarunio **           
                          |                        
                          +-- Ligumia **              

Campbell & Lydeard, 2012 [Publ 3309], fig. 3

iT tT bi riS oT roT roS iD
15 16 14 11 58 58 4 48.0
  +----------------------------------- Quadrulini            
  |                                                          
--+  +-------------------------------- Pleurobemini          
  |  |                                                       
  |  |                             +-- Amblema               
  +--+  +--------------------------+                         
     |  |                          +-- Reginaia              
     |  |                                                    
     +--+  +-------------------------- Popenaiadini          
        |  |                                                 
        |  |                       +-- Glebula **               
        +--+  +--------------------+                         
           |  |                    +-- Paetulunio **            
           |  |                                              
           +--+                    +-- Toxolasma **             
              |  +-----------------+                         
              |  |                 +-- Plectomerus           
              |  |                                           
              +--+  +----------------- Obliquaria **            
                 |  |                                        
                 |  |           +----- Potamilus **             
                 +--+  +--------+                            
                    |  |        |  +-- Dromus **                
                    |  |        +--+                         
                    +--+           +-- Lemiox **                
                       |                                     
                       |  +----------- Lampsilis_ovata **       
                       |  |                                  
                       +--+  +-------- Lampsilis_virescens **   
                          |  |                               
                          +--+     +-- Epioblasma **            
                             |  +--+                         
                             |  |  +-- Obovaria_arkansasensis **
                             +--+                            
                                |  +-- Obovaria_olivaria **     
                                +--+                         
                                   +-- Venustaconcha **         

Campbell & Lydeard, 2012 [Publ 3309], fig. 1-2

iT tT bi riS oT roT roS iD
15 15 9 11 58 58 4 51.0
  +-------------------------- Quadrulini            
  |                                                 
--+  +----------------------- Pleurobemini          
  |  |                                              
  |  |                    +-- Plectomerus           
  +--+  +-----------------+                         
     |  |                 +-- Reginaia              
     |  |                                           
     |  |                 +-- Popenaiadini          
     +--+  +--------------+                         
        |  |              +-- Amblema               
        |  |                                        
        |  |  +-------------- Potamilus_alatus **      
        |  |  |                                     
        +--+  +-------------- Potamilus_capax **       
           |  |                                     
           |  +-------------- Glebula **               
           |  |                                     
           |  |           +-- Toxolasma **             
           |  +-----------+                         
           +--+           +-- Paetulunio **            
              |                                     
              |           +-- Dromus **                
              +-----------+                         
              |           +-- Lemiox **                
              |                                     
              |           +-- Lampsilis_ovata **       
              |  +--------+                         
              |  |        +-- Obliquaria **            
              +--+                                  
                 |  +-------- Lampsilis_virescens **   
                 |  |                               
                 +--+     +-- Epioblasma **            
                    |  +--+                         
                    |  |  +-- Obovaria_arkansasensis **
                    +--+                            
                       |  +-- Obovaria_olivaria **     
                       +--+                         
                          +-- Venustaconcha **         

Huang et al., 2019 [Publ 3843], fig. 2

iT tT bi riS oT roT roS iD
14 15 10 12 184 11 3 49.0
  +-------------------------- Quadrulini        
  |                                             
--+  +----------------------- Pleurobemini      
  |  |                                          
  +--+  +-------------------- Amblema           
     |  |                                       
     |  |              +----- Toxolasma **         
     +--+  +-----------+                        
        |  |           |  +-- Obliquaria **        
        |  |           +--+                     
        |  |              +-- Plectomerus       
        +--+                                    
           |  +-------------- Potamilus_leptodon **
           |  |                                 
           |  |  +----------- Potamilus_alatus **  
           +--+  |                              
              |  |  +-------- Venustaconcha **     
              |  |  |                           
              +--+  +-------- Leaunio **           
                 |  |                           
                 |  |     +-- Ellipsaria **        
                 |  +-----+                     
                 +--+     +-- Truncilla **         
                    |                           
                    |     +-- Medionidus **        
                    +-----+                     
                    |     +-- Ptychobranchus **    
                    |                           
                    |  +----- Cambarunio **        
                    +--+                        
                       |  +-- Ortmanniana **       
                       +--+                     
                          +-- Lampsilis **         

Huang et al., 2019 [Publ 3843], fig. S3

iT tT bi riS oT roT roS iD
14 15 13 12 182 11 3 50.0
  +----------------------------------- Quadrulini        
  |                                                      
--+  +-------------------------------- Pleurobemini      
  |  |                                                   
  +--+  +----------------------------- Amblema           
     |  |                                                
     +--+  +-------------------------- Toxolasma **         
        |  |                                             
        +--+                       +-- Obliquaria **        
           |  +--------------------+                     
           |  |                    +-- Plectomerus       
           +--+                                          
              |  +-------------------- Potamilus_leptodon **
              |  |                                       
              +--+  +----------------- Potamilus_alatus **  
                 |  |                                    
                 |  |              +-- Medionidus **        
                 +--+  +-----------+                     
                    |  |           +-- Ptychobranchus **    
                    |  |                                 
                    +--+           +-- Ellipsaria **        
                       |  +--------+                     
                       |  |        +-- Truncilla **         
                       +--+                              
                          |  +-------- Leaunio **           
                          |  |                           
                          +--+     +-- Venustaconcha **     
                             |  +--+                     
                             |  |  +-- Cambarunio **        
                             +--+                        
                                |  +-- Ortmanniana **       
                                +--+                     
                                   +-- Lampsilis **         

Huang et al., 2019 [Publ 3843], fig. S4

iT tT bi riS oT roT roS iD
14 21 8 12 184 11 3 54.0
  +-------- Amblema           
  |                           
  +-------- Toxolasma **         
  |                           
  +-------- Pleurobemini      
  |                           
  |     +-- Obliquaria **        
  +-----+                     
  |     +-- Plectomerus       
--+                           
  |  +----- Ellipsaria **        
  |  |                        
  |  +----- Potamilus_leptodon **
  |  |                        
  |  +----- Medionidus **        
  |  |                        
  |  +----- Potamilus_alatus **  
  |  |                        
  |  +----- Ptychobranchus **    
  +--+                        
     +----- Truncilla **         
     |                        
     +----- Venustaconcha **     
     |                        
     +----- Cambarunio **        
     |                        
     +----- Leaunio **           
     |                        
     |  +-- Ortmanniana **       
     +--+                     
        +-- Lampsilis **         

Chapman et al., 2008 [Publ 2667], fig. 1

iT tT bi riS oT roT roS iD
13 13 11 11 8 7 4 47.0
  +----------------------------------------- Quadrulini         
  |                                                             
--+  +-------------------------------------- Pleurobemini       
  |  |                                                          
  +--+  +----------------------------------- Amblema            
     |  |                                                       
     +--+  +-------------------------------- Plectomerus        
        |  |                                                    
        +--+  +----------------------------- Popenaiadini       
           |  |                                                 
           +--+  +-------------------------- Toxolasma **          
              |  |                                              
              +--+  +----------------------- Obliquaria **         
                 |  |                                           
                 |  |                    +-- Cyrtonaias **         
                 +--+  +-----------------+                      
                    |  |                 +-- Glebula **            
                    |  |                                        
                    +--+                 +-- Ptychobranchus **     
                       |  +--------------+                      
                       |  |              +-- Lemiox **             
                       |  |                                     
                       +--+              +-- Venustaconcha **      
                          |  +-----------+                      
                          |  |           +-- Obovaria **           
                          +--+                                  
                             |  +----------- Hamiota **            
                             +--+                               
                                |  +-------- Lampsilis_ovata **    
                                +--+                            
                                   |  +----- Ortmanniana **        
                                   +--+                         
                                      |  +-- Lampsilis_straminea **
                                      +--+                      
                                         +-- Lampsilis_hydiana **  

Chapman et al., 2008 [Publ 2744], fig. 2

iT tT bi riS oT roT roS iD
13 13 11 11 8 7 4 47.0
  +----------------------------------------- Quadrulini         
  |                                                             
--+  +-------------------------------------- Pleurobemini       
  |  |                                                          
  +--+  +----------------------------------- Amblema            
     |  |                                                       
     +--+  +-------------------------------- Plectomerus        
        |  |                                                    
        +--+  +----------------------------- Popenaiadini       
           |  |                                                 
           +--+  +-------------------------- Toxolasma **          
              |  |                                              
              +--+  +----------------------- Obliquaria **         
                 |  |                                           
                 |  |                    +-- Cyrtonaias **         
                 +--+  +-----------------+                      
                    |  |                 +-- Glebula **            
                    |  |                                        
                    +--+                 +-- Ptychobranchus **     
                       |  +--------------+                      
                       |  |              +-- Lemiox **             
                       |  |                                     
                       +--+              +-- Venustaconcha **      
                          |  +-----------+                      
                          |  |           +-- Obovaria **           
                          +--+                                  
                             |  +----------- Hamiota **            
                             +--+                               
                                |  +-------- Lampsilis_ovata **    
                                +--+                            
                                   |  +----- Ortmanniana **        
                                   +--+                         
                                      |  +-- Lampsilis_straminea **
                                      +--+                      
                                         +-- Lampsilis_hydiana **  

Breton et al., 2011 [Publ 3376], fig. 3

iT tT bi riS oT roT roS iD
12 12 10 8 25 6 3 53.0
  +-------------------------- Quadrulini   
  |                                        
--+  +----------------------- Pleurobemini 
  |  |                                     
  +--+  +-------------------- Amblemini    
     |  |                                  
     +--+  +----------------- Toxolasma **    
        |  |                               
        +--+  +-------------- Ellipsaria **   
           |  |                            
           +--+  +----------- Truncilla **    
              |  |                         
              +--+        +-- Lemiox **       
                 |  +-----+                
                 |  |     +-- Potamilus **    
                 +--+                      
                    |  +----- Lampsilis **    
                    +--+                   
                       |  +-- Cambarunio **   
                       +--+                
                          +-- Venustaconcha **

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A

iT tT bi riS oT roT roS iD
12 12 6 9 18 5 3 49.0
                 +-- Quadrulini           
  +--------------+                        
  |              +-- Pleurobemini         
  |                                       
--+  +-------------- Amblemini            
  |  |                                    
  |  |  +----------- Ligumia **              
  |  |  |                                 
  +--+  +----------- Truncilla **            
     |  |                                 
     |  |        +-- Ptychobranchus **       
     |  +--------+                        
     +--+        +-- Sagittunio **           
        |                                 
        |        +-- Lampsilis_siliquoidea **
        |  +-----+                        
        |  |     +-- Epioblasma **           
        |  |                              
        +--+  +----- Ortmanniana **          
           |  |                           
           |  +----- Cambarunio **           
           +--+                           
              +----- Leaunio **              
              |                           
              |  +-- Lampsilis_cardium **    
              +--+                        
                 +-- Lampsilis_fasciola **   

Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B

iT tT bi riS oT roT roS iD
12 12 10 9 18 5 3 53.0
                          +-- Quadrulini           
  +-----------------------+                        
  |                       +-- Pleurobemini         
--+                                                
  |  +----------------------- Amblemini            
  |  |                                             
  +--+  +-------------------- Truncilla **            
     |  |                                          
     +--+  +----------------- Ligumia **              
        |  |                                       
        |  |              +-- Ptychobranchus **       
        +--+  +-----------+                        
           |  |           +-- Sagittunio **           
           |  |                                    
           +--+  +----------- Lampsilis_siliquoidea **
              |  |                                 
              |  |     +----- Ortmanniana **          
              +--+  +--+                           
                 |  |  |  +-- Lampsilis_cardium **    
                 |  |  +--+                        
                 +--+     +-- Lampsilis_fasciola **   
                    |                              
                    |  +----- Epioblasma **           
                    +--+                           
                       |  +-- Cambarunio **           
                       +--+                        
                          +-- Leaunio **              

Pfeiffer et al., 2019 [Publ 3866], fig. 1

iT tT bi riS oT roT roS iD
12 13 11 11 23 22 4 51.0
  +----------------------------------- Quadrulini             
  |                                                           
--+  +-------------------------------- Pleurobemini           
  |  |                                                        
  +--+  +----------------------------- Amblemini              
     |  |                                                     
     |  |                          +-- Popenaias              
     +--+  +-----------------------+                          
        |  |                       +-- Psoronaias             
        |  |                                                  
        +--+                       +-- Cyrtonaias_tampicoensis **
           |  +--------------------+                          
           |  |                    +-- Glebula **                
           |  |                                               
           +--+  +-------------------- Potamilus **              
              |  |                                            
              |  |                 +-- Truncilla **              
              +--+  +--------------+                          
                 |  |              +-- Ptychobranchus **         
                 |  |                                         
                 +--+  +-------------- Pachynaias **             
                    |  |                                      
                    |  |           +-- Cyrtonaias_explicata **   
                    +--+  +--------+                          
                       |  |        +-- Sphenonaias            
                       |  |                                   
                       +--+        +-- Lampsilis **              
                          |  +-----+                          
                          |  |     +-- Ortmanniana **            
                          +--+                                
                             |  +----- Cambarunio **             
                             +--+                             
                                |  +-- Epioblasma **             
                                +--+                          
                                   +-- Ligumia **                

Inoue et al., 2020 [Publ 4128], fig. 2

iT tT bi riS oT roT roS iD
12 13 6 11 14 14 3 48.0
  +-------------------------- Quadrulini             
  |                                                  
--+  +----------------------- Reginaia               
  |  |                                               
  +--+  +-------------------- Amblema                
     |  |                                            
     |  |                 +-- Popenaias              
     +--+  +--------------+                          
        |  |              +-- Psoronaias             
        |  |                                         
        +--+              +-- Glebula **                
           |  +-----------+                          
           |  |           +-- Cyrtonaias_tampicoensis **
           |  |                                      
           +--+  +----------- Potamilus **              
              |  |                                   
              |  |        +-- Ptychobranchus **         
              +--+  +-----+                          
                 |  |     +-- Truncilla **              
                 |  |                                
                 |  |  +----- Ortmanniana **            
                 +--+  |                             
                    |  +----- Epioblasma **             
                    |  |                             
                    |  +----- Lampsilis **              
                    |  |                             
                    +--+----- Ligumia **                
                       |                             
                       +----- Cambarunio **             
                       |                             
                       +----- Pachynaias **             
                       |                             
                       |  +-- Cyrtonaias_explicata **   
                       +--+                          
                          +-- Sphenonaias            

Chase et al., 2018 [Publ 3847], fig. 4

iT tT bi riS oT roT roS iD
12 12 10 8 26 5 3 53.0
  +----------------- Toxolasma **    
  |                               
--+  +-------------- Venustaconcha **
  |  |                            
  +--+           +-- Lemiox **       
     |  +--------+                
     |  |        +-- Lampsilis **    
     +--+                         
        |  +-------- Potamilus **    
        +--+                      
           |  +----- Cambarunio **   
           +--+                   
              |  +-- Ellipsaria **   
              +--+                
                 +-- Truncilla **    

Wu et al., 2023 [Publ 4339], fig. 4_S5-aa

iT tT bi riS oT roT roS iD
11 11 9 5 85 6 4 49.0
  +----------------------------- Quadrulini           
  |                                                   
--+  +-------------------------- Pleurobemini         
  |  |                                                
  +--+  +----------------------- Amblemini            
     |  |                                             
     +--+  +-------------------- Popenaiadini         
        |  |                                          
        +--+  +----------------- Toxolasma **            
           |  |                                       
           +--+  +-------------- Potamilus **            
              |  |                                    
              +--+  +----------- Leaunio **              
                 |  |                                 
                 |  |     +----- Lampsilis_ornata **     
                 +--+  +--+                           
                    |  |  |  +-- Lampsilis_cardium **    
                    |  |  +--+                        
                    +--+     +-- Lampsilis_cariosa **    
                       |                              
                       |  +----- Venustaconcha **        
                       +--+                           
                          |  +-- Lampsilis_powellii **   
                          +--+                        
                             +-- Lampsilis_siliquoidea **

Wu et al., 2023 [Publ 4339], fig. 4_S5-nt

iT tT bi riS oT roT roS iD
11 11 9 5 85 6 4 49.0
  +----------------------------- Quadrulini           
  |                                                   
--+  +-------------------------- Pleurobemini         
  |  |                                                
  +--+  +----------------------- Amblemini            
     |  |                                             
     +--+  +-------------------- Popenaiadini         
        |  |                                          
        +--+  +----------------- Toxolasma **            
           |  |                                       
           +--+  +-------------- Potamilus **            
              |  |                                    
              +--+  +----------- Leaunio **              
                 |  |                                 
                 |  |     +----- Lampsilis_ornata **     
                 +--+  +--+                           
                    |  |  |  +-- Lampsilis_cardium **    
                    |  |  +--+                        
                    +--+     +-- Lampsilis_cariosa **    
                       |                              
                       |  +----- Venustaconcha **        
                       +--+                           
                          |  +-- Lampsilis_powellii **   
                          +--+                        
                             +-- Lampsilis_siliquoidea **

Krebs et al., 2003 [Publ 2691], fig. 3

iT tT bi riS oT roT roS iD
11 12 10 8 22 10 3 53.0
                          +-- Megalonaias          
  +-----------------------+                        
  |                       +-- Quadrulina           
  |                                                
  |              +----------- Elliptoideus         
  |  +-----------+                                 
--+  |           |  +-------- Pleurobema           
  |  |           +--+                              
  |  |              |  +----- Uniomerus            
  |  |              +--+                           
  |  |                 |  +-- Fusconaia            
  +--+                 +--+                        
     |                    +-- Eurynia              
     |                                             
     |  +-------------------- Amblema              
     |  |                                          
     |  |              +----- Toxolasma **            
     |  |  +-----------+                           
     +--+  |           |  +-- Plectomerus          
        |  |           +--+                        
        |  |              +-- Obliquaria **           
        |  |                                       
        +--+           +----- Ptychobranchus **       
           |  +--------+                           
           |  |        |  +-- Potamilus_fragilis **   
           |  |        +--+                        
           |  |           +-- Medionidus **           
           +--+                                    
              |           +-- Villosa **              
              |  +--------+                        
              |  |        +-- Ellipsaria **           
              +--+                                 
                 |  +-------- Lampsilis_teres **      
                 +--+                              
                    |  +----- Lampsilis_siliquoidea **
                    +--+                           
                       |  +-- Potamilus_purpuratus ** 
                       +--+                        
                          +-- Potamilus_alatus **     

Wu et al., 2024 [Publ 4350], fig. 8b_S2

iT tT bi riS oT roT roS iD
11 11 8 5 81 6 4 48.0
  +----------------------------- Quadrulini           
  |                                                   
--+  +-------------------------- Pleurobemini         
  |  |                                                
  +--+  +----------------------- Amblemini            
     |  |                                             
     +--+  +-------------------- Popenaiadini         
        |  |                                          
        +--+  +----------------- Toxolasma **            
           |  |                                       
           +--+  +-------------- Potamilus **            
              |  |                                    
              +--+  +----------- Leaunio **              
                 |  |                                 
                 +--+  +-------- Venustaconcha **        
                    |  |                              
                    |  |     +-- Lampsilis_powellii **   
                    +--+-----+                        
                       |     +-- Lampsilis_siliquoidea **
                       |                              
                       |  +----- Lampsilis_ornata **     
                       +--+                           
                          |  +-- Lampsilis_cardium **    
                          +--+                        
                             +-- Lampsilis_cariosa **    

Wu et al., 2024 [Publ 4350], fig. 8a_S1

iT tT bi riS oT roT roS iD
11 11 9 5 81 6 4 49.0
  +----------------------------- Quadrulini           
  |                                                   
--+  +-------------------------- Pleurobemini         
  |  |                                                
  +--+  +----------------------- Amblemini            
     |  |                                             
     +--+  +-------------------- Popenaiadini         
        |  |                                          
        +--+  +----------------- Toxolasma **            
           |  |                                       
           +--+  +-------------- Potamilus **            
              |  |                                    
              +--+  +----------- Leaunio **              
                 |  |                                 
                 |  |     +----- Lampsilis_ornata **     
                 +--+  +--+                           
                    |  |  |  +-- Lampsilis_cardium **    
                    |  |  +--+                        
                    +--+     +-- Lampsilis_cariosa **    
                       |                              
                       |  +----- Venustaconcha **        
                       +--+                           
                          |  +-- Lampsilis_powellii **   
                          +--+                        
                             +-- Lampsilis_siliquoidea **

Roe, 2013 [Publ 3571], fig. 4

iT tT bi riS oT roT roS iD
11 11 9 6 0 0 0 48.0
  +----------- Obliquaria **    
  |                          
--+----------- Lampsilis **     
  |                          
  |  +-------- Medionidus **    
  +--+                       
     |  +----- Ptychobranchus **
     +--+                    
        |  +-- Cyprogenia **    
        +--+                 
           +-- Dromus **        

Inoue et al., 2020 [Publ 4128], fig. 3

iT tT bi riS oT roT roS iD
11 12 11 10 14 14 3 54.0
  +----------------------------------------- Quadrulini             
  |                                                                 
--+  +-------------------------------------- Reginaia               
  |  |                                                              
  +--+  +----------------------------------- Amblema                
     |  |                                                           
     +--+                                +-- Popenaias              
        |  +-----------------------------+                          
        |  |                             +-- Psoronaias             
        +--+                                                        
           |  +----------------------------- Cyrtonaias_tampicoensis **
           |  |                                                     
           +--+  +-------------------------- Ptychobranchus **         
              |  |                                                  
              +--+  +----------------------- Ligumia **                
                 |  |                                               
                 +--+  +-------------------- Potamilus **              
                    |  |                                            
                    +--+  +----------------- Pachynaias **             
                       |  |                                         
                       |  |              +-- Epioblasma **             
                       +--+  +-----------+                          
                          |  |           +-- Truncilla **              
                          |  |                                      
                          +--+           +-- Ortmanniana **            
                             |  +--------+                          
                             |  |        +-- Lampsilis **              
                             +--+                                   
                                |  +-------- Cambarunio **             
                                +--+                                
                                   |  +----- Sphenonaias_microdon__2
                                   +--+                             
                                      |  +-- Cyrtonaias_explicata **   
                                      +--+                          
                                         +-- Sphenonaias_microdon__1

Stodola et al., 2021 [Publ 4266], fig. 4

iT tT bi riS oT roT roS iD
10 10 7 2 0 0 0 54.0
              +-- Ortmanniana **           
              |                         
  +-----------+-- Lampsilis_abrupta **     
  |           |                         
  |           +-- Lampsilis_higginsii **   
--+                                     
  |  +----------- Lampsilis_sietmani **    
  |  |                                  
  |  |        +-- Lampsilis_siliquoidea ** 
  +--+  +-----+                         
     |  |     +-- Lampsilis_radiata **     
     |  |                               
     +--+  +----- Lampsilis_hydiana__2 **  
        |  |                            
        |  +----- Lampsilis_straminea__2 **
        |  |                            
        +--+  +-- Lampsilis_hydiana__1 **  
           +--+                         
           |  +-- Lampsilis_bergmanni **   
           |                            
           |  +-- Lampsilis_straminea__1 **
           +--+                         
              +-- Lampsilis_virescens **   

Roe et al., 2001 [Publ 2689], fig. 3a

iT tT bi riS oT roT roS iD
10 10 8 5 0 0 0 44.0
  +----- Ligumia **   
  |                
  +----- Obliquaria **
--+                
  +----- Lampsilis ** 
  |                
  |  +-- Villosa **   
  +--+             
     +-- Hamiota **   

Inoue et al., 2013 [Publ 3528], fig. 3

iT tT bi riS oT roT roS iD
10 13 10 6 7 5 3 52.0
              +-- Pleurobemini
  +-----------+               
  |           +-- Quadrulini  
  |                           
--+           +-- Paetulunio **  
  |  +--------+               
  |  |        +-- Amblemini   
  +--+                        
     |  +-------- Lampsilis **   
     |  |                     
     +--+  +----- Leaunio **     
        |  |                  
        +--+----- Obovaria **    
           |                  
           |  +-- Villosa **     
           +--+               
              +-- Cambarunio **  

Turner et al., 2000 [Publ 2722], fig. 2

iT tT bi riS oT roT roS iD
9 9 7 3 5 3 3 52.0
                       +-- Amblemini               
  +--------------------+                           
  |                    +-- Pleurobemini            
--+                                                
  |  +-------------------- Medionidus **              
  |  |                                             
  +--+  +----------------- Lampsilis_teres **         
     |  |                                          
     |  +----------------- Villosa **                 
     +--+                                          
        |              +-- Lampsilis_satura **        
        |  +-----------+                           
        |  |           +-- Lampsilis_cardium **       
        +--+                                       
           |  +----------- Lampsilis_reeveiana **     
           |  |                                    
           +--+  +-------- Lampsilis_hydiana__1 **    
              |  |                                 
              +--+-------- Lampsilis_hydiana__2 **    
                 |                                 
                 |  +----- Lampsilis_siliquoidea__2 **
                 +--+                              
                    |  +-- Lampsilis_siliquoidea__1 **
                    +--+                           
                       +-- Lampsilis_powellii **      

Porto-Hannes et al., 2021 [Publ 4227], fig. 3

iT tT bi riS oT roT roS iD
9 9 7 4 6 6 3 53.0
  +-------------------- Quadrula_quadrula    
  |                                          
  +-------------------- Quadrula_verrucosa   
--+                                          
  |  +----------------- Pleurobemini         
  |  |                                       
  |  |  +-------------- Amblemini            
  +--+  |                                    
     |  |           +-- Obovaria **             
     |  |  +--------+                        
     +--+  |        +-- Ortmanniana **          
        |  |                                 
        |  |        +-- Obliquaria **           
        |  +--------+                        
        +--+        +-- Lampsilis_ornata **     
           |                                 
           |     +----- Lampsilis_radiata__2 ** 
           |  +--+                           
           |  |  |  +-- Lampsilis_siliquoidea **
           |  |  +--+                        
           +--+     +-- Lampsilis_radiata__1 ** 
              |                              
              |  +----- Lampsilis_virescens **  
              +--+                           
                 |  +-- Lampsilis_hydiana **    
                 +--+                        
                    +-- Lampsilis_straminea **  

Roe & Lydeard, 1998 [Publ 2693], fig. 3

iT tT bi riS oT roT roS iD
9 10 5 3 1 1 1 49.0
  +-------------- Pleurobemini           
  |                                      
  +-------------- Obliquaria **             
--+                                      
  |  +----------- Potamilus_capax **        
  |  |                                   
  |  +----------- Lampsilis **              
  +--+                                   
     |  +-------- Potamilus_fragilis **     
     |  |                                
     |  |  +----- Potamilus_inflatus **     
     +--+  |                             
        |  |  +-- Potamilus_ohiensis **     
        |  +--+                          
        +--+  +-- Potamilus_amphichaenus ** 
           |                             
           |  +-- Potamilus_purpuratus__1 **
           |  |                          
           +--+-- Potamilus_purpuratus__2 **
              |                          
              +-- Potamilus_alatus **       

Lydeard et al., 1996 [Publ 2656], fig. 4

iT tT bi riS oT roT roS iD
9 9 7 7 19 9 3 48.0
                                +-- Megalonaias         
  +-----------------------------+                       
  |                             +-- Quadrulina          
  |                                                     
  |                       +-------- Elliptoideus        
--+  +--------------------+                             
  |  |                    |  +----- Uniomerus           
  |  |                    +--+                          
  |  |                       |  +-- Eurynia             
  +--+                       +--+                       
     |                          +-- Pleurobema          
     |                                                  
     |  +-------------------------- Amblema             
     +--+                                               
        |  +----------------------- Plectomerus         
        +--+                                            
           |  +-------------------- Obliquaria **          
           +--+                                         
              |  +----------------- Medionidus **          
              +--+                                      
                 |  +-------------- Ptychobranchus **      
                 +--+                                   
                    |  +----------- Potamilus_fragilis **  
                    +--+                                
                       |  +-------- Potamilus_purpuratus **
                       +--+                             
                          |  +----- Ellipsaria **          
                          +--+                          
                             |  +-- Villosa **             
                             +--+                       
                                +-- Lampsilis **           

Wu et al., 2021 [Publ 4366], fig. 3-BI

iT tT bi riS oT roT roS iD
9 9 7 5 65 6 4 47.0
  +----------------------------- Quadrulini           
  |                                                   
--+  +-------------------------- Pleurobemini         
  |  |                                                
  +--+  +----------------------- Amblemini            
     |  |                                             
     +--+  +-------------------- Popenaiadini         
        |  |                                          
        +--+  +----------------- Toxolasma **            
           |  |                                       
           +--+  +-------------- Potamilus **            
              |  |                                    
              +--+  +----------- Leaunio **              
                 |  |                                 
                 +--+        +-- Lampsilis_ornata **     
                    |  +-----+                        
                    |  |     +-- Lampsilis_cardium **    
                    +--+                              
                       |  +----- Venustaconcha **        
                       +--+                           
                          |  +-- Lampsilis_siliquoidea **
                          +--+                        
                             +-- Lampsilis_powellii **   

Wu et al., 2022 [Publ 4238], fig. 3

iT tT bi riS oT roT roS iD
9 9 7 5 65 6 4 47.0
  +----------------------------- Quadrulini           
  |                                                   
--+  +-------------------------- Pleurobemini         
  |  |                                                
  +--+  +----------------------- Amblemini            
     |  |                                             
     +--+  +-------------------- Popenaiadini         
        |  |                                          
        +--+  +----------------- Toxolasma **            
           |  |                                       
           +--+  +-------------- Potamilus **            
              |  |                                    
              +--+  +----------- Leaunio **              
                 |  |                                 
                 +--+        +-- Lampsilis_cardium **    
                    |  +-----+                        
                    |  |     +-- Lampsilis_ornata **     
                    +--+                              
                       |  +----- Venustaconcha **        
                       +--+                           
                          |  +-- Lampsilis_siliquoidea **
                          +--+                        
                             +-- Lampsilis_powellii **   

Wu et al., 2021 [Publ 4366], fig. 3-ML

iT tT bi riS oT roT roS iD
9 9 7 5 65 6 4 47.0
  +----------------------------- Quadrulini           
  |                                                   
--+  +-------------------------- Pleurobemini         
  |  |                                                
  +--+  +----------------------- Amblemini            
     |  |                                             
     +--+  +-------------------- Popenaiadini         
        |  |                                          
        +--+  +----------------- Toxolasma **            
           |  |                                       
           +--+  +-------------- Potamilus **            
              |  |                                    
              +--+  +----------- Leaunio **              
                 |  |                                 
                 +--+        +-- Lampsilis_ornata **     
                    |  +-----+                        
                    |  |     +-- Lampsilis_cardium **    
                    +--+                              
                       |  +----- Venustaconcha **        
                       +--+                           
                          |  +-- Lampsilis_siliquoidea **
                          +--+                        
                             +-- Lampsilis_powellii **   

Mejia-Trujillo & Smith, 2022 [Publ 4274], fig. 2-F

iT tT bi riS oT roT roS iD
9 15 13 4 6 6 4 51.0
     +----------- Toxolasma **            
  +--+                                 
  |  |  +-------- Popenaiadini         
  |  +--+                              
  |     |  +----- Amblemini            
  |     +--+                           
--+        |  +-- Quadrulini           
  |        +--+                        
  |           +-- Pleurobemini         
  |                                    
  |  +----------- Potamilus **            
  |  |                                 
  +--+        +-- Lampsilis_cardium **    
     |  +-----+                        
     |  |     +-- Lampsilis_ornata **     
     +--+                              
        |  +----- Venustaconcha **        
        +--+                           
           |  +-- Lampsilis_siliquoidea **
           +--+                        
              +-- Lampsilis_powellii **   

Dai et al., 2023 [Publ 4285], fig. 5

iT tT bi riS oT roT roS iD
9 9 7 5 44 6 4 47.0
  +----------------------------- Quadrulini           
  |                                                   
--+  +-------------------------- Pleurobemini         
  |  |                                                
  +--+  +----------------------- Amblemini            
     |  |                                             
     +--+  +-------------------- Popenaiadini         
        |  |                                          
        +--+  +----------------- Toxolasma **            
           |  |                                       
           +--+  +-------------- Potamilus **            
              |  |                                    
              +--+  +----------- Leaunio **              
                 |  |                                 
                 +--+        +-- Lampsilis_cardium **    
                    |  +-----+                        
                    |  |     +-- Lampsilis_ornata **     
                    +--+                              
                       |  +----- Venustaconcha **        
                       +--+                           
                          |  +-- Lampsilis_powellii **   
                          +--+                        
                             +-- Lampsilis_siliquoidea **

Lydeard et al., 1996 [Publ 2656], fig. 3

iT tT bi riS oT roT roS iD
9 10 7 7 18 9 3 52.0
                          +-- Megalonaias         
  +-----------------------+                       
  |                       +-- Quadrulina          
  |                                               
  |                    +----- Elliptoideus        
--+                    |                          
  |  +-----------------+  +-- Eurynia             
  |  |                 |  |                       
  |  |                 +--+-- Pleurobema          
  |  |                    |                       
  +--+                    +-- Uniomerus           
     |                                            
     |  +-------------------- Amblema             
     |  |                                         
     +--+                 +-- Plectomerus         
        |  +--------------+                       
        |  |              +-- Obliquaria **          
        +--+                                      
           |  +-------------- Potamilus_fragilis **  
           |  |                                   
           +--+  +----------- Ptychobranchus **      
              |  |                                
              +--+  +-------- Medionidus **          
                 |  |                             
                 +--+  +----- Potamilus_purpuratus **
                    |  |                          
                    +--+----- Lampsilis **           
                       |                          
                       |  +-- Ellipsaria **          
                       +--+                       
                          +-- Villosa **             

Wu et al., 2021 [Publ 4366], fig. 4

iT tT bi riS oT roT roS iD
8 8 6 4 61 4 4 45.0
  +-------------------------- Quadrulini           
  |                                                
--+  +----------------------- Pleurobemini         
  |  |                                             
  +--+  +-------------------- Amblemini            
     |  |                                          
     +--+  +----------------- Popenaiadini         
        |  |                                       
        +--+  +-------------- Toxolasma **            
           |  |                                    
           +--+  +----------- Potamilus **            
              |  |                                 
              +--+        +-- Lampsilis_ornata **     
                 |  +-----+                        
                 |  |     +-- Lampsilis_cardium **    
                 +--+                              
                    |  +----- Venustaconcha **        
                    +--+                           
                       |  +-- Lampsilis_siliquoidea **
                       +--+                        
                          +-- Lampsilis_powellii **   

Hoeh et al., 2009 [Publ 2716], fig. 4

iT tT bi riS oT roT roS iD
8 13 10 8 38 5 2 51.0
                 +----- Pleurobemini  
  +--------------+                    
  |              |  +-- Obliquaria **    
  |              +--+                 
--+                 +-- Quadrulini    
  |                                   
  |  +----------------- Ptychobranchus **
  +--+                                
     |  +-------------- Truncilla **     
     +--+                             
        |  +----------- Ortmanniana **   
        +--+                          
           |  +-------- Lampsilis **     
           +--+                       
              |  +----- Cambarunio **    
              +--+                    
                 |  +-- Ligumia **       
                 +--+                 
                    +-- Epioblasma **    

Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11

iT tT bi riS oT roT roS iD
8 11 9 8 17 5 3 55.0
              +----- Eurynia       
  +-----------+                    
  |           |  +-- Cambarunio **    
  |           +--+                 
  |              +-- Ptychobranchus **
--+                                
  |              +-- Amblemini     
  |  +-----------+                 
  |  |           +-- Pleurobema    
  +--+                             
     |  +----------- Truncilla **     
     |  |                          
     +--+        +-- Lampsilis **     
        |  +-----+                 
        |  |     +-- Epioblasma **    
        +--+                       
           |  +----- Ligumia **       
           +--+                    
              |  +-- Obliquaria **    
              +--+                 
                 +-- Ortmanniana **   

Huang et al., 2002 [Publ 2657], fig. 1

iT tT bi riS oT roT roS iD
8 9 7 7 26 6 3 52.0
  +----------------------------- Megalonaias         
  |                                                  
  |                       +----- Elliptoideus        
--+  +--------------------+                          
  |  |                    |  +-- Uniomerus           
  |  |                    +--+                       
  +--+                       +-- Pleurobema          
     |                                               
     |  +----------------------- Amblema             
     |  |                                            
     +--+                    +-- Obliquaria **          
        |  +-----------------+                       
        |  |                 +-- Plectomerus         
        +--+                                         
           |  +----------------- Medionidus **          
           +--+                                      
              |  +-------------- Potamilus_fragilis **  
              +--+                                   
                 |  +----------- Ptychobranchus **      
                 +--+                                
                    |  +-------- Potamilus_purpuratus **
                    +--+                             
                       |  +----- Lampsilis **           
                       +--+                          
                          |  +-- Villosa **             
                          +--+                       
                             +-- Ellipsaria **          

Huang et al., 2002 [Publ 2657], fig. 2

iT tT bi riS oT roT roS iD
8 10 2 7 26 6 3 52.0
  +----------- Megalonaias         
  |                                
  |     +----- Elliptoideus        
  |  +--+                          
--+  |  |  +-- Uniomerus           
  |  |  +--+                       
  |  |     +-- Pleurobema          
  |  |                             
  |  |  +----- Potamilus_fragilis **  
  +--+  |                          
     |  +----- Medionidus **          
     |  |                          
     |  +----- Amblema             
     |  |                          
     |  |  +-- Obliquaria **          
     +--+--+                       
        |  +-- Plectomerus         
        |                          
        |  +-- Villosa **             
        |  |                       
        |  +-- Ellipsaria **          
        |  |                       
        +--+-- Lampsilis **           
           |                       
           +-- Potamilus_purpuratus **
           |                       
           +-- Ptychobranchus **      

Graf & Cummings, 2006 [Publ 1968], fig. 2

iT tT bi riS oT roT roS iD
8 8 4 8 39 6 3 49.0
  +----------------------- Quadrulini    
  |                                      
--+  +-------------------- Amblemini     
  |  |                                   
  +--+  +----------------- Pleurobemini  
     |  |                                
     +--+  +-------------- Obliquaria **    
        |  |                             
        |  +-------------- Truncilla **     
        +--+                             
           +-------------- Ptychobranchus **
           |                             
           |  +----------- Ligumia **       
           +--+                          
              |  +-------- Cambarunio **    
              +--+                       
                 |  +----- Epioblasma **    
                 +--+                    
                    |  +-- Lampsilis **     
                    +--+                 
                       +-- Ortmanniana **   

Keogh & Simons, 2019 [Publ 3935], fig. S1

iT tT bi riS oT roT roS iD
8 8 6 3 2 2 2 52.0
        +-- Amblemini   
  +-----+               
  |     +-- Pleurobemini
--+                     
  |  +----- Potamilus **   
  +--+                  
     |  +-- Leaunio **     
     +--+               
        +-- Lampsilis **   

Fukata & Iigo, 2020 [Publ 4106], fig. 1

iT tT bi riS oT roT roS iD
8 8 6 4 46 3 2 48.0
  +-------------------- Quadrulini           
  |                                          
--+  +----------------- Pleurobemini         
  |  |                                       
  +--+  +-------------- Potamilus **            
     |  |                                    
     +--+  +----------- Leaunio **              
        |  |                                 
        +--+        +-- Lampsilis_cardium **    
           |  +-----+                        
           |  |     +-- Lampsilis_ornata **     
           +--+                              
              |  +----- Venustaconcha **        
              +--+                           
                 |  +-- Lampsilis_powellii **   
                 +--+                        
                    +-- Lampsilis_siliquoidea **

Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1

iT tT bi riS oT roT roS iD
8 8 6 4 11 6 4 45.0
  +-------------------------- Quadrulini           
  |                                                
--+  +----------------------- Pleurobemini         
  |  |                                             
  +--+  +-------------------- Amblemini            
     |  |                                          
     +--+  +----------------- Popenaiadini         
        |  |                                       
        +--+  +-------------- Toxolasma **            
           |  |                                    
           +--+  +----------- Potamilus **            
              |  |                                 
              +--+        +-- Lampsilis_cardium **    
                 |  +-----+                        
                 |  |     +-- Lampsilis_ornata **     
                 +--+                              
                    |  +----- Venustaconcha **        
                    +--+                           
                       |  +-- Lampsilis_powellii **   
                       +--+                        
                          +-- Lampsilis_siliquoidea **

Hoeh et al., 2009 [Publ 2716], fig. 3

iT tT bi riS oT roT roS iD
8 14 12 8 39 6 3 52.0
              +-------- Amblemini     
  +-----------+                       
  |           |  +----- Pleurobemini  
  |           +--+                    
  |              |  +-- Obliquaria **    
--+              +--+                 
  |                 +-- Quadrulini    
  |                                   
  |  +----------------- Ptychobranchus **
  +--+                                
     |  +-------------- Truncilla **     
     +--+                             
        |  +----------- Ortmanniana **   
        +--+                          
           |  +-------- Lampsilis **     
           +--+                       
              |  +----- Cambarunio **    
              +--+                    
                 |  +-- Ligumia **       
                 +--+                 
                    +-- Epioblasma **    

Hewitt et al., 2019 [Publ 3962], fig. 2

iT tT bi riS oT roT roS iD
7 7 8 2 3 1 1 50.0
  +----------------------- Pleurobemini         
  |                                             
--+  +-------------------- Lampsilis_siliquoidea **
  |  |                                          
  |  |  +----------------- Lampsilis_fasciola **   
  +--+  |                                       
     |  |              +-- Epioblasma **           
     |  +--------------+                        
     +--+              +-- Lampsilis_radiata **    
        |                                       
        |  +-------------- Lampsilis_ornata **     
        |  |                                    
        +--+           +-- Lampsilis_cardium__1 ** 
           |  +--------+                        
           |  |        +-- Lampsilis_ovata__1 **   
           +--+                                 
              |  +-------- Lampsilis_cardium__6 ** 
              |  |                              
              +--+  +----- Lampsilis_cardium__5 ** 
                 |  |                           
                 +--+  +-- Lampsilis_cardium__2 ** 
                    |  |                        
                    |  +-- Lampsilis_ovata__2 **   
                    +--+                        
                       +-- Lampsilis_cardium__3 ** 
                       |                        
                       +-- Lampsilis_cardium__4 ** 

Serb et al., 2003 [Publ 2659], fig. 3

iT tT bi riS oT roT roS iD
7 29 27 6 22 22 3 63.0
  +-------------------------- Lampsilis **   
  |                                       
--+  +----------------------- Potamilus **   
  |  |                                    
  |  |                 +----- Medionidus **  
  +--+  +--------------+                  
     |  |              |  +-- Dromus **      
     |  |              +--+               
     +--+                 +-- Cyprogenia **  
        |                                 
        |  +----------------- Obliquaria **  
        |  |                              
        +--+  +-------------- Amblema     
           |  |                           
           |  |        +----- Reginaia    
           +--+  +-----+                  
              |  |     |  +-- Elliptoideus
              |  |     +--+               
              +--+        +-- Plectomerus 
                 |                        
                 |  +-------- Quadrulini  
                 +--+                     
                    |  +----- Hemistena   
                    +--+                  
                       |  +-- Elliptio    
                       +--+               
                          +-- Fusconaia   

Serb & Barnhart, 2008 [Publ 2674], fig. 3

iT tT bi riS oT roT roS iD
7 7 10 5 0 0 0 45.0
  +-------------------------- Lampsilis **             
  |                                                 
--+  +----------------------- Obliquaria **            
  |  |                                              
  +--+  +-------------------- Ptychobranchus **        
     |  |                                           
     +--+                 +-- Cyprogenia_aberti__6 **  
        |  +--------------+                         
        |  |              +-- Dromus **                
        +--+                                        
           |  +-------------- Cyprogenia_aberti__4 **  
           |  |                                     
           +--+  +----------- Cyprogenia_aberti__3 **  
              |  |                                  
              +--+  +-------- Cyprogenia_aberti__5 **  
                 |  |                               
                 +--+  +----- Cyprogenia_stegaria__2 **
                    |  |                            
                    +--+----- Cyprogenia_aberti__2 **  
                       |                            
                       |  +-- Cyprogenia_stegaria__1 **
                       +--+                         
                          +-- Cyprogenia_aberti__1 **  

Serb, 2006 [Publ 2678], fig. 2

iT tT bi riS oT roT roS iD
7 7 7 5 0 0 0 46.0
  +-------------- Obliquaria **            
  |                                     
  |           +-- Ptychobranchus **        
  +-----------+                         
--+           +-- Lampsilis **             
  |                                     
  |           +-- Cyprogenia_aberti__6 **  
  |  +--------+                         
  |  |        +-- Dromus **                
  +--+                                  
     |  +-------- Cyprogenia_aberti__5 **  
     |  |                               
     +--+  +----- Cyprogenia_aberti__4 **  
        |  |                            
        |  |  +-- Cyprogenia_stegaria__1 **
        +--+  |                         
           |  +-- Cyprogenia_stegaria__2 **
           |  |                         
           +--+-- Cyprogenia_aberti__1 **  
              |                         
              +-- Cyprogenia_aberti__2 **  
              |                         
              +-- Cyprogenia_aberti__3 **  

Buhay et al., 2002 [Publ 2661], fig. 2A

iT tT bi riS oT roT roS iD
7 7 3 4 0 0 0 48.0
  +----- Lampsilis ** 
  |                
--+----- Epioblasma **
  |                
  |  +-- Cambarunio **
  +--+             
     +-- Villosa **   

Serb, 2006 [Publ 2678], fig. 4

iT tT bi riS oT roT roS iD
7 7 8 5 0 0 0 44.0
  +-------------------- Obliquaria **            
  |                                           
  +-------------------- Lampsilis **             
--+                                           
  |  +----------------- Ptychobranchus **        
  |  |                                        
  +--+              +-- Cyprogenia_aberti__5 **  
     |  +-----------+                         
     |  |           +-- Dromus **                
     +--+                                     
        |  +----------- Cyprogenia_aberti__4 **  
        |  |                                  
        +--+  +-------- Cyprogenia_aberti__3 **  
           |  |                               
           +--+  +----- Cyprogenia_stegaria__2 **
              |  |                            
              +--+----- Cyprogenia_aberti__2 **  
                 |                            
                 |  +-- Cyprogenia_stegaria__1 **
                 +--+                         
                    +-- Cyprogenia_aberti__1 **  

Jones & Neves, 2010 [Publ 2646], fig. 16

iT tT bi riS oT roT roS iD
7 7 5 1 0 0 0 50.0
-- (Epioblasma

Serb et al., 2003 [Publ 2659], fig. 2

iT tT bi riS oT roT roS iD
7 29 20 6 22 22 3 62.0
  +----------- Lampsilis **   
  |                        
  |  +-------- Obliquaria **  
  |  |                     
  |  +-------- Amblema     
  |  |                     
--+  +-------- Reginaia    
  |  |                     
  |  +-------- Potamilus **   
  |  |                     
  |  +-------- Quadrulini  
  |  |                     
  +--+     +-- Elliptoideus
     +-----+               
     |     +-- Plectomerus 
     |                     
     |  +----- Medionidus **  
     +--+                  
     |  |  +-- Dromus **      
     |  +--+               
     |     +-- Cyprogenia **  
     |                     
     |  +----- Hemistena   
     +--+                  
        |  +-- Elliptio    
        +--+               
           +-- Fusconaia   

Serb & Barnhart, 2008 [Publ 2674], fig. 4

iT tT bi riS oT roT roS iD
7 7 9 5 0 0 0 44.0
  +----------------------- Obliquaria **            
  |                                              
  +----------------------- Lampsilis **             
--+                                              
  |  +-------------------- Ptychobranchus **        
  |  |                                           
  +--+                 +-- Cyprogenia_aberti__6 **  
     |  +--------------+                         
     |  |              +-- Dromus **                
     +--+                                        
        |  +-------------- Cyprogenia_aberti__5 **  
        |  |                                     
        +--+  +----------- Cyprogenia_aberti__4 **  
           |  |                                  
           +--+  +-------- Cyprogenia_aberti__3 **  
              |  |                               
              +--+  +----- Cyprogenia_stegaria__2 **
                 |  |                            
                 +--+----- Cyprogenia_aberti__2 **  
                    |                            
                    |  +-- Cyprogenia_stegaria__1 **
                    +--+                         
                       +-- Cyprogenia_aberti__1 **  

Porto-Hannes et al., 2021 [Publ 4227], fig. 4

iT tT bi riS oT roT roS iD
7 7 3 3 4 4 3 50.0
  +-------------- Quadrula    
  |                           
--+-------------- Pustulosa   
  |                           
  |  +----------- Pleurobemini
  +--+                        
     |  +-------- Amblemini   
     +--+                     
        |  +----- Obovaria **    
        +--+                  
           |  +-- Ortmanniana ** 
           +--+               
              +-- Lampsilis **   

Jones & Neves, 2010 [Publ 2646], fig. 17

iT tT bi riS oT roT roS iD
7 7 1 1 0 0 0 49.0
-- (Epioblasma

Huang et al., 2019 [Publ 3843], fig. 3

iT tT bi riS oT roT roS iD
7 7 5 5 50 3 2 45.0
  +----------------- Quadrulini   
--+                               
  |  +-------------- Pleurobemini 
  +--+                            
     |  +----------- Toxolasma **    
     +--+                         
        |  +-------- Potamilus **    
        +--+                      
           |  +----- Leaunio **      
           +--+                   
              |  +-- Venustaconcha **
              +--+                
                 +-- Lampsilis **    

Buhay et al., 2002 [Publ 2661], fig. 2B

iT tT bi riS oT roT roS iD
7 7 5 4 0 0 0 47.0
  +----- Lampsilis ** 
  |                
--+----- Epioblasma **
  |                
  |  +-- Cambarunio **
  +--+             
     +-- Villosa **   

Serb, 2006 [Publ 2678], fig. 3

iT tT bi riS oT roT roS iD
7 7 8 5 0 0 0 44.0
  +-------------------- Obliquaria **            
  |                                           
  +-------------------- Lampsilis **             
--+                                           
  |  +----------------- Ptychobranchus **        
  |  |                                        
  +--+              +-- Cyprogenia_aberti__5 **  
     |  +-----------+                         
     |  |           +-- Dromus **                
     +--+                                     
        |  +----------- Cyprogenia_aberti__4 **  
        |  |                                  
        +--+  +-------- Cyprogenia_aberti__3 **  
           |  |                               
           +--+  +----- Cyprogenia_stegaria__2 **
              |  |                            
              +--+----- Cyprogenia_aberti__2 **  
                 |                            
                 |  +-- Cyprogenia_stegaria__1 **
                 +--+                         
                    +-- Cyprogenia_aberti__1 **  

Hoeh et al., 2009 [Publ 2716], fig. 5

iT tT bi riS oT roT roS iD
7 13 7 7 39 6 3 50.0
           +-- Obliquaria **    
  +--------+                 
  |        +-- Quadrulini    
  |                          
--+  +-------- Amblemini     
  |  |                       
  |  +-------- Pleurobemini  
  +--+                       
     |  +----- Truncilla **     
     |  |                    
     |  +----- Ptychobranchus **
     +--+                    
        |  +-- Ortmanniana **   
        |  |                 
        |  +-- Cambarunio **    
        +--+                 
           +-- Ligumia **       
           |                 
           +-- Epioblasma **    

Bogan & Hoeh, 2000 [Publ 2650], fig. 1

iT tT bi riS oT roT roS iD
6 11 5 6 28 5 3 49.0
  +-------- Toxolasma **   
  |                     
  +-------- Amblemini   
  |                     
  |     +-- Glebula **     
  +-----+               
--+     +-- Cyrtonaias **  
  |                     
  |     +-- Quadrulini  
  +-----+               
  |     +-- Pleurobemini
  |                     
  |  +----- Potamilus **   
  +--+                  
     |  +-- Ortmanniana ** 
     +--+               
        +-- Ligumia **     

Dai et al., 2024 [Publ 4358], fig. 4

iT tT bi riS oT roT roS iD
6 6 4 5 46 6 4 46.0
  +----------------------- Quadrulini   
--+                                     
  |  +-------------------- Pleurobemini 
  +--+                                  
     |  +----------------- Amblemini    
     +--+                               
        |  +-------------- Popenaiadini 
        +--+                            
           |  +----------- Toxolasma **    
           +--+                         
              |  +-------- Potamilus **    
              +--+                      
                 |  +----- Leaunio **      
                 +--+                   
                    |  +-- Lampsilis **    
                    +--+                
                       +-- Venustaconcha **

Keogh & Simons, 2019 [Publ 3935], fig. 4

iT tT bi riS oT roT roS iD
6 6 6 1 0 0 0 50.0
-- (Lampsilis

Hoeh et al., 2002 [Publ 2653], fig. 2

iT tT bi riS oT roT roS iD
6 11 8 6 25 5 3 50.0
           +-- Glebula **     
  +--------+               
  |        +-- Cyrtonaias **  
  |                        
--+  +-------- Toxolasma **   
  |  |                     
  |  |  +----- Potamilus **   
  |  +--+                  
  +--+  |  +-- Ligumia **     
     |  +--+               
     |     +-- Ortmanniana ** 
     |                     
     |  +----- Amblemini   
     +--+                  
        |  +-- Quadrulini  
        +--+               
           +-- Pleurobemini

Hoeh et al., 2001 [Publ 2652], fig. 14f2

iT tT bi riS oT roT roS iD
6 28 16 6 25 5 3 69.0
  +-------------------- Amblemini              
  |                                            
  +-------------------- Coelaturini            
  |                                            
  +-------------------- Eurynia                
  |                                            
  +-------------------- Fusconaia              
  |                                            
  +-------------------- Gonideini              
  |                                            
  +-------------------- Pleurobema             
  |                                            
  +-------------------- Quadrulini             
--+                                            
  |        +----------- Ortmanniana **            
  +--------+                                   
  |        |  +-------- Glebula **                
  |        +--+                                
  |           |  +----- Potamilus **              
  |           +--+                             
  |              |  +-- Ligumia **                
  |              +--+                          
  |                 +-- Toxolasma **              
  |                                            
  |  +----------------- Cyrtonaias **             
  |  |                                         
  +--+  +-------------- Unionini               
     |  |                                      
     |  |  +----------- Anodontini             
     +--+  |                                   
        |  |     +----- Mutela                 
        |  |     |                             
        +--+  +--+  +-- Anodontites_guanarensis
           |  |  |  |                          
           |  |  +--+-- Anodontites_trigona    
           |  |     |                          
           +--+     +-- Monocondylaea          
              |                                
              |  +----- Rhipidodontini         
              |  |                             
              |  +----- Echyridella            
              +--+                             
                 +----- Lortiella              
                 |                             
                 |  +-- Castaliini             
                 +--+                          
                    +-- Velesunio              

Pavan-Kumar et al., 2022 [Publ 4263], fig. S5

iT tT bi riS oT roT roS iD
6 6 4 4 33 1 1 45.0
  +-------------- Quadrulini           
  |                                    
--+  +----------- Toxolasma **            
  |  |                                 
  +--+  +-------- Potamilus **            
     |  |                              
     +--+     +-- Lampsilis_cardium **    
        |  +--+                        
        |  |  +-- Lampsilis_ornata **     
        +--+                           
           |  +-- Lampsilis_siliquoidea **
           +--+                        
              +-- Venustaconcha **        

Chen et al., 2020 [Publ 4170], fig. 1

iT tT bi riS oT roT roS iD
6 16 14 3 28 1 1 58.0
                    +-- Anodontini           
              +-----+                        
              |     +-- Potamilus_alatus **     
  +-----------+                              
  |           |  +----- Lamprotulini         
  |           +--+                           
  |              |  +-- Chamberlainiini      
--+              +--+                        
  |                 +-- Gonideini            
  |                                          
  |  +----------------- Gibbosula            
  +--+                                       
     |  +-------------- Quadrulini           
     +--+                                    
        |  +----------- Toxolasma **            
        +--+                                 
           |  +-------- Lampsilis_ornata **     
           +--+                              
              |  +----- Potamilus_leptodon **   
              +--+                           
                 |  +-- Lampsilis_siliquoidea **
                 +--+                        
                    +-- Lampsilis_powellii **   

Pavan-Kumar et al., 2022 [Publ 4263], fig. 4

iT tT bi riS oT roT roS iD
6 6 4 4 30 1 1 45.0
  +-------------- Quadrulini           
  |                                    
--+  +----------- Toxolasma **            
  |  |                                 
  +--+  +-------- Potamilus **            
     |  |                              
     +--+     +-- Lampsilis_cardium **    
        |  +--+                        
        |  |  +-- Lampsilis_ornata **     
        +--+                           
           |  +-- Lampsilis_siliquoidea **
           +--+                        
              +-- Venustaconcha **        

Roe & Hoeh, 2003 [Publ 2655], fig. 4f2

iT tT bi riS oT roT roS iD
6 12 9 6 25 5 3 57.0
  +----------------------- Fusconaia  
  |                                   
  +----------------------- Amblemini  
--+                                   
  |  +-------------------- Quadrulini 
  |  |                                
  +--+  +----------------- Pleurobema 
     |  |                             
     |  |  +-------------- Eurynia    
     +--+  |                          
        |  |        +----- Cyrtonaias ** 
        |  +--------+                 
        +--+        |  +-- Unionini   
           |        +--+              
           |           +-- Anodontini 
           |                          
           |  +----------- Ortmanniana **
           +--+                       
              |  +-------- Glebula **    
              +--+                    
                 |  +----- Potamilus **  
                 +--+                 
                    |  +-- Ligumia **    
                    +--+              
                       +-- Toxolasma **  

Pfeiffer et al., 2016 [Publ 3660], fig. 2

iT tT bi riS oT roT roS iD
6 6 4 6 32 31 3 47.0
  +----------------------------- Amblemini            
  |                                                   
  |                          +-- Cyrtonaias **           
  |                    +-----+                        
  |                    |     +-- Glebula **              
--+                    |                              
  |  +-----------------+     +-- Lampsilis **            
  |  |                 |  +--+                        
  |  |                 |  |  +-- Leaunio **              
  |  |                 +--+                           
  |  |                    |  +-- Truncilla **            
  +--+                    +--+                        
     |                       +-- Potamilus **            
     |                                                
     |  +----------------------- Elliptoideus         
     |  |                                             
     +--+  +-------------------- Quadrulini           
        |  |                                          
        +--+  +----------------- Elliptio             
           |  |                                       
           |  |  +-------------- Parvaspina           
           +--+  |                                    
              |  +-------------- Plethobasus          
              |  |                                    
              |  |           +-- Hemistena            
              +--+-----------+                        
                 |           +-- Eurynia              
                 |                                    
                 |  +----------- Fusconaia            
                 |  |                                 
                 |  |     +----- Pleurobema_riddellii 
                 +--+  +--+                           
                    |  |  |  +-- Pleurobema_sintoxia  
                    |  |  +--+                        
                    +--+     +-- Pleurobema_cordatum  
                       |                              
                       |  +----- Pleuronaia           
                       |  |                           
                       +--+  +-- Pleurobema_pyriforme 
                          |  |                        
                          +--+-- Pleurobema_strodeanum
                             |                        
                             +-- Pleurobema_clava     

Hoeh et al., 2001 [Publ 2652], fig. 14f4

iT tT bi riS oT roT roS iD
6 6 4 6 25 5 3 46.0
  +----------------- Amblemini   
  |                              
  |              +-- Quadrulini  
--+  +-----------+               
  |  |           +-- Pleurobemini
  |  |                           
  +--+           +-- Cyrtonaias **  
     |  +--------+               
     |  |        +-- Glebula **     
     +--+                        
        |  +-------- Toxolasma **   
        +--+                     
           |  +----- Potamilus **   
           +--+                  
              |  +-- Ligumia **     
              +--+               
                 +-- Ortmanniana ** 

Roe & Hoeh, 2003 [Publ 2655], fig. 4f3

iT tT bi riS oT roT roS iD
6 6 4 6 25 5 3 48.0
  +----------------- Amblemini   
  |                              
  |              +-- Quadrulini  
--+  +-----------+               
  |  |           +-- Pleurobemini
  |  |                           
  +--+           +-- Cyrtonaias **  
     |  +--------+               
     |  |        +-- Glebula **     
     +--+                        
        |  +-------- Toxolasma **   
        +--+                     
           |  +----- Ortmanniana ** 
           +--+                  
              |  +-- Ligumia **     
              +--+               
                 +-- Potamilus **   

Krebs et al., 2013 [Publ 3515], fig. 2A

iT tT bi riS oT roT roS iD
5 5 3 2 0 0 0 50.0
  +-------- Lampsilis_radiata **  
  |                            
--+     +-- Lampsilis_ovata **    
  |  +--+                      
  |  |  +-- Ortmanniana **        
  +--+                         
     |  +-- Lampsilis_hydiana **  
     +--+                      
        +-- Lampsilis_straminea **

Mejia-Trujillo & Smith, 2022 [Publ 4274], fig. 2-M

iT tT bi riS oT roT roS iD
5 5 3 3 1 1 1 48.0
  +-------- Quadrulini   
--+                      
  |  +----- Potamilus **    
  +--+                   
     |  +-- Venustaconcha **
     +--+                
        +-- Lampsilis **    

Huang et al., 2018 [Publ 3828], fig. 6

iT tT bi riS oT roT roS iD
5 5 3 4 38 1 1 44.0
  +----------- Quadrulini   
--+                         
  |  +-------- Toxolasma **    
  +--+                      
     |  +----- Potamilus **    
     +--+                   
        |  +-- Lampsilis **    
        +--+                
           +-- Venustaconcha **

Pfeiffer et al., 2019 [Publ 3937], fig. 4-5

iT tT bi riS oT roT roS iD
5 5 3 5 52 10 4 49.0
  +----------------------- Quadrulini  
--+                                    
  |  +-------------------- Pleurobemini
  +--+                                 
     |  +----------------- Amblemini   
     +--+                              
        |  +-------------- Popenaiadini
        +--+                           
           |  +----------- Obliquaria **  
           +--+                        
              |  +-------- Cyrtonaias **  
              +--+                     
                 |  +----- Leaunio **     
                 +--+                  
                    |  +-- Lampsilis **   
                    +--+               
                       +-- Pachynaias **  

Curole & Kocher, 2005 [Publ 2682], fig. 1

iT tT bi riS oT roT roS iD
5 9 7 4 6 4 2 52.0
  +----------------- Glebula **            
  |                                     
--+              +-- Amblema            
  |  +-----------+                      
  |  |           +-- Quadrulini         
  +--+                                  
     |  +----------- Plectomerus        
     +--+                               
        |  +-------- Leaunio **            
        +--+                            
           |  +----- Lampsilis_straminea **
           +--+                         
              |  +-- Lampsilis_teres **    
              +--+                      
                 +-- Potamilus **          

Zieritz et al., 2021 [Publ 4231], fig. 3-F_5

iT tT bi riS oT roT roS iD
5 5 3 4 44 1 1 44.0
  +----------- Quadrulini   
--+                         
  |  +-------- Toxolasma **    
  +--+                      
     |  +----- Potamilus **    
     +--+                   
        |  +-- Venustaconcha **
        +--+                
           +-- Lampsilis **    

Walker et al., 2006 [Publ 2666], fig. 6-M

iT tT bi riS oT roT roS iD
5 5 3 5 9 1 1 46.0
  +-------------- Pleurobemini
--+                           
  |  +----------- Potamilus **   
  +--+                        
     |  +-------- Cyrtonaias **  
     +--+                     
        |  +----- Ligumia **     
        +--+                  
           |  +-- Lampsilis **   
           +--+               
              +-- Ortmanniana ** 

Wen et al., 2017 [Publ 3795], fig. 3

iT tT bi riS oT roT roS iD
5 5 3 4 11 1 1 44.0
  +----------- Quadrulini   
--+                         
  |  +-------- Toxolasma **    
  +--+                      
     |  +----- Potamilus **    
     +--+                   
        |  +-- Lampsilis **    
        +--+                
           +-- Venustaconcha **

Buhay et al., 2002 [Publ 2661], fig. 1

iT tT bi riS oT roT roS iD
5 5 3 2 0 0 0 45.0
  +-- Lampsilis_siliquoidea__1 **
  |                           
--+-- Lampsilis_siliquoidea__2 **
  |                           
  +-- Epioblasma **              

Walker et al., 2006 [Publ 2666], fig. 3

iT tT bi riS oT roT roS iD
5 5 3 5 10 1 1 46.0
  +-------------- Pleurobemini
--+                           
  |  +----------- Potamilus **   
  +--+                        
     |  +-------- Cyrtonaias **  
     +--+                     
        |  +----- Ligumia **     
        +--+                  
           |  +-- Ortmanniana ** 
           +--+               
              +-- Lampsilis **   

Wang et al., 2018 [Publ 3865], fig. 2

iT tT bi riS oT roT roS iD
5 5 3 4 27 1 1 44.0
  +----------- Quadrulini   
--+                         
  |  +-------- Toxolasma **    
  +--+                      
     |  +----- Potamilus **    
     +--+                   
        |  +-- Venustaconcha **
        +--+                
           +-- Lampsilis **    

Saxon et al., 2023 [Publ 4338], fig. 4

iT tT bi riS oT roT roS iD
5 5 3 2 0 0 0 49.0
  +-- Ellipsaria **
--+             
  +-- Truncilla ** 

Campbell & Lydeard, 2012 [Publ 3308], fig. 3

iT tT bi riS oT roT roS iD
5 36 35 4 38 38 4 73.0
  +-------------------------------- Quadrulini          
  |                                                     
  |                 +-------------- Elliptoideus        
  |  +--------------+                                   
  |  |              |  +----------- Plectomerus         
--+  |              +--+                                
  |  |                 |  +-------- Obovaria_unicolor **   
  |  |                 +--+                             
  |  |                    |  +----- Obovaria_olivaria **   
  |  |                    +--+                          
  |  |                       |  +-- Lampsilis **           
  +--+                       +--+                       
     |                          +-- Lemiox **              
     |                                                  
     |                    +-------- Cyrtonaias **          
     |  +-----------------+                             
     |  |                 |  +----- Reginaia            
     |  |                 +--+                          
     |  |                    |  +-- Popenaiadini        
     +--+                    +--+                       
        |                       +-- Amblema             
        |                                               
        |  +----------------------- Elliptio            
        |  |                                            
        +--+  +-------------------- Eurynia             
           |  |                                         
           |  |  +----------------- Parvaspina          
           +--+  |                                      
              |  |              +-- Hemistena           
              |  +--------------+                       
              +--+              +-- Plethobasus         
                 |                                      
                 |  +-------------- Pleuronaia          
                 |  |                                   
                 |  |           +-- Pleurobema_cordatum 
                 +--+  +--------+                       
                    |  |        +-- Fusconaia           
                    |  |                                
                    +--+        +-- Pleurobema_stabile  
                       |  +-----+                       
                       |  |     +-- Pleurobema_sintoxia 
                       +--+                             
                          |  +----- Pleurobema_decisum  
                          +--+                          
                             |  +-- Pleurobema_clava    
                             +--+                       
                                +-- Pleurobema_pyriforme

Pfeiffer et al., 2019 [Publ 3937], fig. 6

iT tT bi riS oT roT roS iD
5 5 3 5 52 10 4 49.0
  +-------------------- Quadrulini  
  |                                 
--+                 +-- Pleurobemini
  |  +--------------+               
  |  |              +-- Amblemini   
  +--+                              
     |  +-------------- Popenaiadini
     +--+                           
        |  +----------- Obliquaria **  
        +--+                        
           |  +-------- Cyrtonaias **  
           +--+                     
              |  +----- Leaunio **     
              +--+                  
                 |  +-- Lampsilis **   
                 +--+               
                    +-- Pachynaias **  

Wang et al., 2018 [Publ 3846], fig. 2

iT tT bi riS oT roT roS iD
5 5 3 4 27 1 1 44.0
  +----------- Quadrulini   
--+                         
  |  +-------- Toxolasma **    
  +--+                      
     |  +----- Potamilus **    
     +--+                   
        |  +-- Venustaconcha **
        +--+                
           +-- Lampsilis **    

Wu et al., 2020 [Publ 4121], fig. 3

iT tT bi riS oT roT roS iD
5 5 3 4 45 1 1 44.0
  +----------- Quadrulini   
--+                         
  |  +-------- Toxolasma **    
  +--+                      
     |  +----- Potamilus **    
     +--+                   
        |  +-- Venustaconcha **
        +--+                
           +-- Lampsilis **    

Walker et al., 2006 [Publ 2666], fig. 2

iT tT bi riS oT roT roS iD
5 5 3 5 10 1 1 44.0
  +----------- Cyrtonaias ** 
--+                       
  |  +-------- Potamilus **  
  +--+                    
     |  +----- Ligumia **    
     +--+                 
        |  +-- Ortmanniana **
        +--+              
           +-- Lampsilis **  

Froufe et al., 2019 [Publ 3936], fig. 3

iT tT bi riS oT roT roS iD
5 5 3 4 40 1 1 44.0
  +----------- Quadrulini   
--+                         
  |  +-------- Toxolasma **    
  +--+                      
     |  +----- Potamilus **    
     +--+                   
        |  +-- Lampsilis **    
        +--+                
           +-- Venustaconcha **

Walker et al., 2006 [Publ 2666], fig. 6

iT tT bi riS oT roT roS iD
5 6 3 5 31 1 1 47.0
  +----------- Cyrtonaias **  
  |                        
--+----------- Pleurobemini
  |                        
  |  +-------- Potamilus **   
  +--+                     
     |  +----- Lampsilis **   
     +--+                  
        |  +-- Ligumia **     
        +--+               
           +-- Ortmanniana ** 

Campbell & Lydeard, 2012 [Publ 3308], fig. 1

iT tT bi riS oT roT roS iD
5 8 6 4 37 37 4 46.0
  +-------------------- Quadrulini  
  |                                 
--+  +----------------- Pleurobemini
  |  |                              
  |  |              +-- Popenaiadini
  +--+  +-----------+               
     |  |           +-- Amblema     
     |  |                           
     +--+           +-- Cyrtonaias **  
        |  +--------+               
        |  |        +-- Reginaia    
        +--+                        
           |  +-------- Plectomerus 
           +--+                     
              |  +----- Lemiox **      
              +--+                  
                 |  +-- Lampsilis **   
                 +--+               
                    +-- Obovaria **    

Hoeh et al., 2002 [Publ 2673], fig. 1-M

iT tT bi riS oT roT roS iD
5 5 3 5 4 1 1 46.0
  +-------------- Pleurobemini
--+                           
  |  +----------- Potamilus **   
  +--+                        
     |  +-------- Cyrtonaias **  
     +--+                     
        |  +----- Ligumia **     
        +--+                  
           |  +-- Lampsilis **   
           +--+               
              +-- Ortmanniana ** 

Campbell & Lydeard, 2012 [Publ 3308], fig. 2

iT tT bi riS oT roT roS iD
5 6 3 4 37 37 4 45.0
  +-------------------- Quadrulini  
  |                                 
--+  +----------------- Pleurobemini
  |  |                              
  +--+              +-- Reginaia    
     |  +-----------+               
     |  |           +-- Amblema     
     +--+                           
        |  +----------- Popenaiadini
        |  |                        
        +--+  +-------- Cyrtonaias **  
           |  |                     
           +--+-------- Plectomerus 
              |                     
              |  +----- Lemiox **      
              +--+                  
                 |  +-- Lampsilis **   
                 +--+               
                    +-- Obovaria **    

Walker et al., 2006 [Publ 2666], fig. 4

iT tT bi riS oT roT roS iD
5 5 3 5 10 1 1 44.0
  +-------------- Pleurobemini
--+                           
  |  +----------- Cyrtonaias **  
  +--+                        
     |  +-------- Potamilus **   
     +--+                     
        |  +----- Ligumia **     
        +--+                  
           |  +-- Ortmanniana ** 
           +--+               
              +-- Lampsilis **   

Wu et al., 2019 [Publ 3864], fig. 3

iT tT bi riS oT roT roS iD
5 5 3 4 42 1 1 44.0
  +----------- Quadrulini   
--+                         
  |  +-------- Toxolasma **    
  +--+                      
     |  +----- Potamilus **    
     +--+                   
        |  +-- Lampsilis **    
        +--+                
           +-- Venustaconcha **

Campbell & Lydeard, 2012 [Publ 3308], fig. 4

iT tT bi riS oT roT roS iD
5 8 5 4 37 37 4 46.0
  +----------------- Quadrulini  
  |                              
--+  +-------------- Pleurobemini
  |  |                           
  |  |           +-- Popenaiadini
  +--+  +--------+               
     |  |        +-- Amblema     
     |  |                        
     +--+  +-------- Plectomerus 
        |  |                     
        |  |     +-- Cyrtonaias **  
        +--+-----+               
           |     +-- Reginaia    
           |                     
           |  +----- Lemiox **      
           +--+                  
              |  +-- Lampsilis **   
              +--+               
                 +-- Obovaria **    

Curole & Kocher, 2005 [Publ 2682], fig. 1-M

iT tT bi riS oT roT roS iD
5 5 3 4 7 4 2 49.0
  +-------------- Quadrulini 
  |                          
--+  +----------- Plectomerus
  |  |                       
  +--+  +-------- Amblema    
     |  |                    
     +--+     +-- Potamilus **  
        |  +--+              
        |  |  +-- Glebula **    
        +--+                 
           |  +-- Leaunio **    
           +--+              
              +-- Lampsilis **  

Jones et al., 2006 [Publ 2681], fig. 2

iT tT bi riS oT roT roS iD
5 5 4 1 0 0 0 49.0
-- (Epioblasma

Lane et al., 2016 [Publ 3735], fig. 3

iT tT bi riS oT roT roS iD
5 5 1 3 0 0 0 49.0
  +-- Paetulunio **   
  |                
--+-- Epioblasma **   
  |                
  +-- Venustaconcha **

Froufe et al., 2019 [Publ 3936], fig. 6

iT tT bi riS oT roT roS iD
5 5 3 4 40 1 1 44.0
  +----------- Quadrulini   
--+                         
  |  +-------- Toxolasma **    
  +--+                      
     |  +----- Potamilus **    
     +--+                   
        |  +-- Lampsilis **    
        +--+                
           +-- Venustaconcha **

Hoeh et al., 2002 [Publ 2673], fig. 1-F

iT tT bi riS oT roT roS iD
5 5 3 5 4 1 1 48.0
  +-------------- Pleurobemini
--+                           
  |  +----------- Cyrtonaias **  
  +--+                        
     |  +-------- Ortmanniana ** 
     +--+                     
        |  +----- Lampsilis **   
        +--+                  
           |  +-- Potamilus **   
           +--+               
              +-- Ligumia **     

Wu et al., 2021 [Publ 4366], fig. 6

iT tT bi riS oT roT roS iD
4 4 2 4 87 2 2 45.0
           +-- Quadrulini  
  +--------+               
  |        +-- Pleurobemini
--+                        
  |  +-------- Obliquaria **  
  +--+                     
     |  +----- Potamilus **   
     +--+                  
        |  +-- Lampsilis **   
        +--+               
           +-- Truncilla **   

Wu et al., 2023 [Publ 4339], fig. 5

iT tT bi riS oT roT roS iD
4 4 2 4 15 2 2 47.0
           +-- Pleurobemini
  +--------+               
  |        +-- Quadrulini  
--+                        
  |  +-------- Obliquaria **  
  +--+                     
     |  +----- Truncilla **   
     +--+                  
        |  +-- Lampsilis **   
        +--+               
           +-- Potamilus **   

Cha et al., 2018 [Publ 3916], fig. 1

iT tT bi riS oT roT roS iD
4 4 2 3 35 1 1 45.0
  +-------- Quadrulini
--+                   
  |  +----- Toxolasma ** 
  +--+                
     |  +-- Lampsilis ** 
     +--+             
        +-- Potamilus ** 

Smith, 2021 [Publ 4331], fig. 1

iT tT bi riS oT roT roS iD
4 4 2 3 16 5 3 46.0
  +-------------- Quadrulini   
--+                            
  |  +----------- Pleurobemini 
  +--+                         
     |  +-------- Amblemini    
     +--+                      
        |  +----- Potamilus **    
        +--+                   
           |  +-- Venustaconcha **
           +--+                
              +-- Lampsilis **    

Wu et al., 2021 [Publ 4366], fig. 2_I

iT tT bi riS oT roT roS iD
4 4 2 4 87 2 2 45.0
  +-------------- Quadrulini  
--+                           
  |  +----------- Pleurobemini
  +--+                        
     |  +-------- Obliquaria **  
     +--+                     
        |  +----- Potamilus **   
        +--+                  
           |  +-- Lampsilis **   
           +--+               
              +-- Truncilla **   

Gangloff et al., 2013 [Publ 3551], fig. 2

iT tT bi riS oT roT roS iD
4 4 2 3 0 0 0 49.0
  +----- Venustaconcha **
--+                   
  |  +-- Ligumia **      
  +--+                
     +-- Sagittunio **   

Zanatta & Harris, 2013 [Publ 3530], fig. 2

iT tT bi riS oT roT roS iD
4 4 2 3 1 0 0 48.0
  +----- Lampsilis **    
--+                   
  |  +-- Epioblasma **   
  +--+                
     +-- Venustaconcha **

Bolotov et al., 2018 [Publ 3826], fig. 2

iT tT bi riS oT roT roS iD
3 13 0 3 80 4 3 55.0
  +-- Lamprotula    
  |                 
  +-- Potomida      
  |                 
  +-- Pronodularia  
  |                 
  +-- Gonidea       
  |                 
  +-- Leguminaia    
  |                 
  +-- Microcondylaea
  |                 
--+-- Amblemini     
  |                 
  +-- Ortmanniana **   
  |                 
  +-- Lampsilis **     
  |                 
  +-- Cambarunio **    
  |                 
  +-- Elliptio      
  |                 
  +-- Pleurobema    
  |                 
  +-- Quadrulini    

Lydeard et al., 2000 [Publ 2654], fig. 2b

iT tT bi riS oT roT roS iD
3 3 1 2 11 9 3 46.0
        +-- Lampsilis **   
  +-----+               
  |     +-- Obovaria **    
--+                     
  |  +----- Amblemini   
  +--+                  
     |  +-- Quadrulini  
     +--+               
        +-- Pleurobemini

Rosenberg et al., 1997 [Publ 2720], fig. 2A

iT tT bi riS oT roT roS iD
3 17 0 2 19 9 3 59.0
  +-- Gonideini          
  |                      
  +-- Pyganodon_grandis  
  |                      
  +-- Utterbackia        
  |                      
  +-- Pyganodon_cataracta
  |                      
  +-- Lampsilis_straminea **
  |                      
  +-- Lampsilis_teres **    
  |                      
  +-- Megalonaias        
  |                      
  +-- Obliquaria **         
  |                      
--+-- Theliderma         
  |                      
  +-- Quadrula           
  |                      
  +-- Uniomerus          
  |                      
  +-- Amblema            
  |                      
  +-- Elliptio           
  |                      
  +-- Fusconaia          
  |                      
  +-- Plectomerus        
  |                      
  +-- Pleurobema         
  |                      
  +-- Unionini           

Fonseca et al., 2016 [Publ 3775], fig. 1

iT tT bi riS oT roT roS iD
3 3 1 3 29 1 1 46.0
  +-------- Quadrulini   
--+                      
  |  +----- Toxolasma **    
  +--+                   
     |  +-- Lampsilis **    
     +--+                
        +-- Venustaconcha **

Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2

iT tT bi riS oT roT roS iD
3 3 1 3 77 6 3 48.0
           +-- Quadrulini  
  +--------+               
  |        +-- Pleurobemini
--+                        
  |  +-------- Amblemini   
  +--+                     
     |  +----- Cambarunio **  
     +--+                  
        |  +-- Lampsilis **   
        +--+               
           +-- Ortmanniana ** 

Plazzi et al., 2011 [Publ 3676], fig. 5

iT tT bi riS oT roT roS iD
3 3 1 3 15 2 2 47.0
  +-------- Quadrulini   
  |                      
--+-------- Pleurobemini 
  |                      
  |  +----- Lampsilis **    
  +--+                   
     |  +-- Epioblasma **   
     +--+                
        +-- Venustaconcha **

Lopes-Lima et al., 2017 [Publ 3792], fig. 2

iT tT bi riS oT roT roS iD
3 3 1 3 30 1 1 46.0
  +-------- Quadrulini   
--+                      
  |  +----- Toxolasma **    
  +--+                   
     |  +-- Lampsilis **    
     +--+                
        +-- Venustaconcha **

Lydeard et al., 2000 [Publ 2654], fig. 2a

iT tT bi riS oT roT roS iD
3 3 1 2 11 9 3 46.0
  +-- Lampsilis **
--+            
  +-- Obovaria ** 

Dai et al., 2024 [Publ 4386], fig. 6

iT tT bi riS oT roT roS iD
3 3 1 2 37 1 1 48.0
  +----- Quadrulini   
--+                   
  |  +-- Venustaconcha **
  +--+                
     +-- Lampsilis **    

Lydeard et al., 2000 [Publ 2654], fig. 2c

iT tT bi riS oT roT roS iD
3 3 1 2 11 9 3 48.0
  +----- Obovaria_unicolor **
--+                       
  |  +-- Obovaria_olivaria **
  +--+                    
     +-- Lampsilis **        

Guerra et al., 2017 [Publ 3772], fig. 2b

iT tT bi riS oT roT roS iD
3 3 1 3 22 1 1 46.0
  +-------- Quadrulini   
--+                      
  |  +----- Toxolasma **    
  +--+                   
     |  +-- Venustaconcha **
     +--+                
        +-- Lampsilis **    

Zhou et al., 2016 [Publ 3710], fig. 1

iT tT bi riS oT roT roS iD
3 3 1 3 17 1 1 46.0
  +-------- Quadrulini   
--+                      
  |  +----- Toxolasma **    
  +--+                   
     |  +-- Venustaconcha **
     +--+                
        +-- Lampsilis **    

Pfeiffer & Graf, 2015 [Publ 3620], fig. 2

iT tT bi riS oT roT roS iD
3 3 1 3 37 2 2 46.0
  +----------- Quadrulini
--+                      
  |  +-------- Amblemini 
  +--+                   
     |  +----- Potamilus ** 
     +--+                
        |  +-- Lampsilis ** 
        +--+             
           +-- Truncilla ** 

Huang et al., 2013 [Publ 3629], fig. 3

iT tT bi riS oT roT roS iD
3 3 1 3 13 1 1 46.0
  +-------- Quadrulini   
--+                      
  |  +----- Toxolasma **    
  +--+                   
     |  +-- Lampsilis **    
     +--+                
        +-- Venustaconcha **

Bolotov et al., 2020 [Publ 4084], fig. 1

iT tT bi riS oT roT roS iD
3 7 0 3 33 4 3 50.0
  +-- Amblemini  
  |              
  +-- Ortmanniana **
  |              
  +-- Lampsilis **  
  |              
--+-- Cambarunio ** 
  |              
  +-- Elliptio   
  |              
  +-- Pleurobema 
  |              
  +-- Quadrulini 

Rosenberg et al., 1997 [Publ 2720], fig. 2B

iT tT bi riS oT roT roS iD
3 17 0 2 19 8 3 59.0
  +-- Gonideini          
  |                      
  +-- Pyganodon_grandis  
  |                      
  +-- Utterbackia        
  |                      
  +-- Pyganodon_cataracta
  |                      
  +-- Lampsilis_straminea **
  |                      
  +-- Lampsilis_teres **    
  |                      
  +-- Megalonaias        
  |                      
  +-- Obliquaria **         
  |                      
--+-- Theliderma         
  |                      
  +-- Quadrula           
  |                      
  +-- Uniomerus          
  |                      
  +-- Amblema            
  |                      
  +-- Elliptio           
  |                      
  +-- Fusconaia          
  |                      
  +-- Plectomerus        
  |                      
  +-- Aculamprotula      
  |                      
  +-- Unio               

Moyer & Díaz-Ferguson, 2012 [Publ 3375], fig. 3

iT tT bi riS oT roT roS iD
3 3 2 1 3 0 0 51.0
-- (Lampsilis

Guerra et al., 2017 [Publ 3772], fig. 2a

iT tT bi riS oT roT roS iD
3 3 1 3 22 1 1 46.0
  +-------- Quadrulini   
--+                      
  |  +----- Toxolasma **    
  +--+                   
     |  +-- Venustaconcha **
     +--+                
        +-- Lampsilis **    

Amaro et al., 2017 [Publ 3791], fig. 2

iT tT bi riS oT roT roS iD
3 3 1 3 13 3 2 46.0
  +-------- Quadrulini   
--+                      
  |  +----- Toxolasma **    
  +--+                   
     |  +-- Venustaconcha **
     +--+                
        +-- Lampsilis **    

Rosenberg et al., 1994 [Publ 2719], fig. 3

iT tT bi riS oT roT roS iD
3 10 0 2 16 8 3 54.0
  +-- Pyganodon_cataracta
  |                      
  +-- Pyganodon_grandis  
  |                      
  +-- Utterbackia        
  |                      
  +-- Lampsilis_teres **    
  |                      
  +-- Lampsilis_straminea **
--+                      
  +-- Megalonaias        
  |                      
  +-- Obliquaria **         
  |                      
  +-- Theliderma         
  |                      
  +-- Quadrula           
  |                      
  +-- Uniomerus          

Aunins et al., 2018 [Publ 3863], fig. 1

iT tT bi riS oT roT roS iD
3 3 1 3 38 1 1 46.0
  +-------- Quadrulini   
--+                      
  |  +----- Toxolasma **    
  +--+                   
     |  +-- Lampsilis **    
     +--+                
        +-- Venustaconcha **

Bolotov et al., 2017 [Publ 3761], fig. S1

iT tT bi riS oT roT roS iD
3 3 1 3 136 6 3 48.0
           +-- Quadrulini  
  +--------+               
  |        +-- Pleurobemini
--+                        
  |  +-------- Amblemini   
  +--+                     
     |  +----- Ortmanniana ** 
     +--+                  
        |  +-- Lampsilis **   
        +--+               
           +-- Cambarunio **  

Chase et al., 2018 [Publ 3847], fig. 5

iT tT bi riS oT roT roS iD
3 3 1 2 7 0 0 48.0
  +-- Potamilus **
--+            
  +-- Lampsilis **

Smith & Johnson, 2020 [Publ 4229], fig. 2

iT tT bi riS oT roT roS iD
3 3 1 1 0 0 0 48.0
-- (Potamilus

Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1

iT tT bi riS oT roT roS iD
3 3 1 3 73 6 3 48.0
           +-- Quadrulini  
  +--------+               
  |        +-- Pleurobemini
--+                        
  |  +-------- Amblemini   
  +--+                     
     |  +----- Ortmanniana ** 
     +--+                  
        |  +-- Cambarunio **  
        +--+               
           +-- Lampsilis **   

Bolotov et al., 2017 [Publ 3761], fig. S2

iT tT bi riS oT roT roS iD
3 3 1 3 136 6 3 48.0
           +-- Quadrulini  
  +--------+               
  |        +-- Pleurobemini
--+                        
  |  +-------- Amblemini   
  +--+                     
     |  +----- Cambarunio **  
     +--+                  
        |  +-- Ortmanniana ** 
        +--+               
           +-- Lampsilis **   

Bolotov et al., 2017 [Publ 3764], fig. S1

iT tT bi riS oT roT roS iD
3 9 0 3 122 6 3 52.0
  +-- Amblemini         
  |                     
  +-- Ortmanniana **       
  |                     
  +-- Lampsilis **         
  |                     
  +-- Cambarunio **        
  |                     
--+-- Elliptio          
  |                     
  +-- Eurynia           
  |                     
  +-- Pleurobema        
  |                     
  +-- Quadrula_quadrula 
  |                     
  +-- Quadrula_verrucosa

Lydeard et al., 2000 [Publ 2654], fig. 2d

iT tT bi riS oT roT roS iD
3 3 1 2 11 9 3 46.0
  +-------- Amblemini   
  |                     
--+     +-- Lampsilis **   
  |  +--+               
  |  |  +-- Obovaria **    
  +--+                  
     |  +-- Quadrulini  
     +--+               
        +-- Pleurobemini

Guerra et al., 2019 [Publ 4168], fig. 1

iT tT bi riS oT roT roS iD
3 3 1 3 38 1 1 46.0
  +-------- Quadrulini   
--+                      
  |  +----- Toxolasma **    
  +--+                   
     |  +-- Venustaconcha **
     +--+                
        +-- Potamilus **    

Guerra et al., 2019 [Publ 4168], fig. 2

iT tT bi riS oT roT roS iD
3 3 1 3 38 1 1 46.0
  +-------- Quadrulini   
--+                      
  |  +----- Toxolasma **    
  +--+                   
     |  +-- Venustaconcha **
     +--+                
        +-- Potamilus **    

Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2

iT tT bi riS oT roT roS iD
3 3 1 3 73 6 3 48.0
           +-- Quadrulini  
  +--------+               
  |        +-- Pleurobemini
--+                        
  |  +-------- Amblemini   
  +--+                     
     |  +----- Ortmanniana ** 
     +--+                  
        |  +-- Lampsilis **   
        +--+               
           +-- Cambarunio **  

Huang et al., 2019 [Publ 3843], fig. 3-M

iT tT bi riS oT roT roS iD
2 2 0 2 21 1 1 --
  +----- Quadrulini   
--+                   
  |  +-- Venustaconcha **
  +--+                
     +-- Potamilus **    

Amaro et al., 2017 [Publ 3791], fig. 3A

iT tT bi riS oT roT roS iD
2 2 0 2 10 1 1 --
  +----- Quadrulini   
--+                   
  |  +-- Toxolasma **    
  +--+                
     +-- Venustaconcha **

Amaro et al., 2017 [Publ 3791], fig. 2-M

iT tT bi riS oT roT roS iD
2 2 0 2 8 2 2 --
     +-- Pleurobemini 
  +--+                
  |  +-- Quadrulini   
--+                   
  |  +-- Lampsilis **    
  +--+                
     +-- Venustaconcha **

Burlakova et al., 2012 [Publ 3486], fig. 1

iT tT bi riS oT roT roS iD
2 39 0 2 39 39 3 78.0
  +----------- Amblema     
  |                        
--+        +-- Lampsilis **   
  |  +-----+               
  |  |     +-- Plectomerus 
  +--+                     
     |  +----- Popenaiadini
     +--+                  
        |  +-- Cyrtonaias **  
        +--+               
           +-- Pleurobemini

Feng et al., 2016 [Publ 3800], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 13 1 1 --
  +----- Quadrulini
--+                
  |  +-- Potamilus ** 
  +--+             
     +-- Lampsilis ** 

Chase et al., 2018 [Publ 3861], fig. 1-M

iT tT bi riS oT roT roS iD
2 2 0 1 9 1 1 --
  +-- Quadrulini
--+             
  +-- Lampsilis ** 

Manendo et al., 2008 [Publ 2669], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 11 3 2 --
     +-- Lampsilis **   
  +--+               
  |  +-- Sagittunio **  
--+                  
  |  +-- Amblemini   
  +--+               
     +-- Pleurobemini

Xue et al., 2016 [Publ 3659], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 14 0 0 --
  +-- Lampsilis **    
--+                
  +-- Venustaconcha **

Amaro et al., 2017 [Publ 3791], fig. 3B

iT tT bi riS oT roT roS iD
2 2 0 2 10 1 1 --
  +-- Toxolasma **    
--+                
  +-- Venustaconcha **

Wu et al., 2018 [Publ 4143], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 20 1 1 --
  +----- Quadrulini   
--+                   
  |  +-- Venustaconcha **
  +--+                
     +-- Lampsilis **    

Pfeiffer & Graf, 2013 [Publ 3531], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 32 3 3 --
  +----------- Quadrulini  
--+                        
  |  +-------- Pleurobemini
  +--+                     
     |  +----- Amblemini   
     +--+                  
        |  +-- Lampsilis **   
        +--+               
           +-- Truncilla **   

Lopes-Lima et al., 2019 [Publ 3901], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 27 23 3 --
  +-------------- Megalonaias 
  |                           
--+           +-- Uniomerus   
  |  +--------+               
  |  |        +-- Quadrulina  
  +--+                        
     |  +-------- Pleurobemini
     +--+                     
        |  +----- Amblemini   
        +--+                  
           |  +-- Lampsilis **   
           +--+               
              +-- Potamilus **   

Chen et al., 2012 [Publ 3624], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 2 5 1 1 --
  +----- Quadrulini   
--+                   
  |  +-- Lampsilis **    
  +--+                
     +-- Venustaconcha **

Zhou et al., 2017 [Publ 3788], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 15 1 1 --
  +----- Quadrulini
--+                
  |  +-- Potamilus ** 
  +--+             
     +-- Lampsilis ** 

Guerra et al., 2019 [Publ 4168], fig. 2-M

iT tT bi riS oT roT roS iD
2 2 0 2 20 1 1 --
  +----- Quadrulini   
--+                   
  |  +-- Venustaconcha **
  +--+                
     +-- Potamilus **    

Wen et al., 2017 [Publ 3795], fig. 3-M

iT tT bi riS oT roT roS iD
2 2 0 2 4 1 1 --
  +----- Quadrulini   
--+                   
  |  +-- Potamilus **    
  +--+                
     +-- Venustaconcha **

Lopes-Lima et al., 2019 [Publ 3901], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 26 22 3 --
  +-------- Amblemini   
  |                     
--+     +-- Lampsilis **   
  |  +--+               
  |  |  +-- Potamilus **   
  +--+                  
     |  +-- Pleurobemini
     +--+               
        +-- Quadrulini  

Chase et al., 2018 [Publ 3861], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 1 9 1 1 --
  +-- Quadrulini
--+             
  +-- Lampsilis ** 

Guerra et al., 2019 [Publ 4168], fig. 1-M

iT tT bi riS oT roT roS iD
2 2 0 2 20 1 1 --
  +----- Quadrulini   
--+                   
  |  +-- Venustaconcha **
  +--+                
     +-- Potamilus **    

Lopes-Lima et al., 2019 [Publ 3901], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 2 24 20 3 --
  +----------- Quadrulini  
--+                        
  |  +-------- Amblemini   
  +--+                     
     |  +----- Pleurobemini
     +--+                  
        |  +-- Lampsilis **   
        +--+               
           +-- Potamilus **   

Burzynski & Soroka, 2018 [Publ 3862], fig. 5a

iT tT bi riS oT roT roS iD
2 2 0 2 18 1 1 --
  +----- Quadrulini   
--+                   
  |  +-- Potamilus **    
  +--+                
     +-- Venustaconcha **

Froufe et al., 2019 [Publ 3936], fig. 3-M

iT tT bi riS oT roT roS iD
2 2 0 2 17 1 1 --
  +----- Quadrulini   
--+                   
  |  +-- Potamilus **    
  +--+                
     +-- Venustaconcha **

Huang et al., 2018 [Publ 3828], fig. 6-M

iT tT bi riS oT roT roS iD
2 2 0 2 15 1 1 --
  +----- Quadrulini   
--+                   
  |  +-- Venustaconcha **
  +--+                
     +-- Potamilus **    

Graf, 2002 [Publ 2651], fig. 2

iT tT bi riS oT roT roS iD
2 2 0 2 16 2 2 --
     +-- Lampsilis **   
  +--+               
  |  +-- Obliquaria **  
--+                  
  |  +-- Pleurobemini
  +--+               
     +-- Amblemini   

Whelan et al., 2011 [Publ 3380], fig. 1-ML

iT tT bi riS oT roT roS iD
2 2 0 2 34 3 3 --
  +----------- Pleurobemini
--+                        
  |  +-------- Quadrulini  
  +--+                     
     |  +----- Amblemini   
     +--+                  
        |  +-- Truncilla **   
        +--+               
           +-- Lampsilis **   

Whelan et al., 2011 [Publ 3380], fig. 1-MP

iT tT bi riS oT roT roS iD
2 2 0 2 34 3 3 --
  +-------- Quadrulini  
  |                     
--+-------- Pleurobemini
  |                     
  |  +----- Amblemini   
  +--+                  
     |  +-- Lampsilis **   
     +--+               
        +-- Truncilla **   

Wu et al., 2016 [Publ 3711], fig. 1

iT tT bi riS oT roT roS iD
2 2 0 2 19 1 1 --
  +----- Quadrulini   
--+                   
  |  +-- Venustaconcha **
  +--+                
     +-- Lampsilis **    

Wu et al., 2020 [Publ 4147], fig. 3

iT tT bi riS oT roT roS iD
2 2 0 2 27 1 1 --
  +----- Quadrulini   
--+                   
  |  +-- Toxolasma **    
  +--+                
     +-- Venustaconcha **

Froufe et al., 2019 [Publ 3936], fig. 4

iT tT bi riS oT roT roS iD
2 2 0 2 17 1 1 --
  +----- Quadrulini   
--+                   
  |  +-- Potamilus **    
  +--+                
     +-- Venustaconcha **

Sheng et al., 2014 [Publ 3704], fig. 2

iT tT bi riS oT roT roS iD
2 6 0 2 6 1 1 50.0
  +-------- Lampsilis **      
  |                        
--+     +-- Venustaconcha **  
  |  +--+                  
  |  |  +-- Quadrulini     
  +--+                     
     |  +-- Lamprotulini   
     +--+                  
        +-- Chamberlainiini

Zieritz et al., 2021 [Publ 4231], fig. 3-M_4

iT tT bi riS oT roT roS iD
2 2 0 2 21 1 1 --
  +----- Quadrulini   
--+                   
  |  +-- Venustaconcha **
  +--+                
     +-- Potamilus **    

Campbell et al., 2008 [Publ 2675], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 25 25 2 --
  +-- Cyrtonaias **  
  |               
--+-- Amblemini   
  |               
  +-- Pleurobemini

Zieritz et al., 2024 [Publ 4381], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 45 1 1 --
  +-- Amblemini
--+            
  +-- Lampsilis **

Sonowal et al., 2021 [Publ 4221], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 15 1 1 --
-- Lampsilis_cardium

Plazzi & Passamonti, 2010 [Publ 3354], fig. 2-3

iT tT bi riS oT roT roS iD
1 1 0 1 3 0 0 --
-- Lampsilis_ornata

Bolotov et al., 2016 [Publ 3743], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 29 0 0 --
-- Lampsilis_cardium

Metzger et al., 2018 [Publ 3952], fig. 1C

iT tT bi riS oT roT roS iD
1 1 0 1 2 1 1 --
  +-- Pleurobemini
--+               
  +-- Lampsilis **   

Giribet & Distel, 2003 [Publ 3670], fig. 3f5

iT tT bi riS oT roT roS iD
1 1 0 1 5 1 1 --
-- Lampsilis_cardium

Campbell et al., 2008 [Publ 2675], fig. 6

iT tT bi riS oT roT roS iD
1 1 0 1 25 25 2 --
  +-- Cyrtonaias **  
  |               
--+-- Amblemini   
  |               
  +-- Pleurobemini

Neemuchwala et al., 2023 [Publ 4292], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 26 25 4 --
  +----------- Quadrulini  
--+                        
  |  +-------- Pleurobemini
  +--+                     
     |  +----- Amblemini   
     +--+                  
        |  +-- Lampsilis **   
        +--+               
           +-- Popenaiadini

Patterson et al., 2021 [Publ 4222], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 3 3 1 --
  +-- Villosa **  
--+            
  +-- Amblemini

Sonowal et al., 2021 [Publ 4221], fig. 3b

iT tT bi riS oT roT roS iD
1 1 0 1 14 1 1 --
-- Lampsilis_cardium

Xue et al., 2016 [Publ 3659], fig. 1-M

iT tT bi riS oT roT roS iD
1 1 0 1 1 1 1 --
  +-- Quadrulini   
--+                
  +-- Venustaconcha **

Araujo et al., 2017 [Publ 3801], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 13 1 1 --
  +-- Amblemini
--+            
  +-- Lampsilis **

Campbell et al., 2008 [Publ 2675], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 24 24 3 --
  +----------------------- Quadrulini               
  |                                                 
  +----------------------- Amblemini                
  |                                                 
  |  +-------------------- Cyrtonaias **               
  |  |                                              
  |  +-------------------- Eurynia                  
  |  |                                              
  |  +-------------------- Fusconaia_cerina         
--+  |                                              
  |  +-------------------- Fusconaia_flava          
  |  |                                              
  |  +-------------------- Hemistena                
  |  |                                              
  |  +-------------------- Plethobasus              
  |  |                                              
  |  +-------------------- Parvaspina               
  |  |                                              
  +--+-------------------- Pleurobema_cordatum      
     |                                              
     +-------------------- Pleuronaia               
     |                                              
     +-------------------- Pleurobema_oviforme__2   
     |                                              
     +-------------------- Pleurobema_stabile       
     |                                              
     +-------------------- Elliptio                 
     |                                              
     |  +----------------- Pleurobema_decisum       
     |  |                                           
     |  +----------------- Pleurobema_sintoxia      
     |  |                                           
     |  +----------------- Pleurobema_clava         
     +--+                                           
        +----------------- Pleurobema_oviforme__1   
        |                                           
        |  +-------------- Pleurobema_beadleianum   
        |  |                                        
        +--+  +----------- Pleurobema_pyriforme     
           |  |                                     
           +--+  +-------- Pleurobema_rubellum      
              |  |                                  
              |  +-------- Pleurobema_hanleyianum__2
              +--+                                  
                 +-------- Pleurobema_georgianum__2 
                 |                                  
                 |  +----- Pleurobema_hanleyianum__1
                 +--+                               
                    |  +-- Pleurobema_georgianum__1 
                    +--+                            
                       +-- Pleurobema_athearni      

Amaro et al., 2017 [Publ 3791], fig. 3B-M

iT tT bi riS oT roT roS iD
1 1 0 1 6 1 1 --
-- Venustaconcha_ellipsiformis

Curole & Kocher, 2002 [Publ 2712], fig. 1b-M

iT tT bi riS oT roT roS iD
1 1 0 1 3 2 1 --
-- Lampsilis_teres

Wu et al., 2020 [Publ 4147], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 13 0 0 --
-- Lampsilis_cardium

Konopleva et al., 2023 [Publ 4269], fig. S1_4

iT tT bi riS oT roT roS iD
1 1 0 1 55 1 1 --
  +-- Lampsilis **
--+            
  +-- Amblemini

Lopes-Lima et al., 2018 [Publ 3829], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 27 0 0 --
-- Lampsilis_cardium

Wei et al., 2016 [Publ 3665], fig. 0

iT tT bi riS oT roT roS iD
1 1 0 1 5 1 1 --
-- Lampsilis_ornata

Huff et al., 2004 [Publ 2697], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 11 0 0 --
-- Lampsilis_cardium

Takeuchi et al., 2015 [Publ 3661], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 9 0 0 --
-- Lampsilis_cardium

Amaro et al., 2017 [Publ 3791], fig. 3A-M

iT tT bi riS oT roT roS iD
1 1 0 1 6 1 1 --
-- Venustaconcha_ellipsiformis

Huff et al., 2004 [Publ 2697], fig. 5-6

iT tT bi riS oT roT roS iD
1 1 0 1 11 0 0 --
-- Lampsilis_cardium

Guerra et al., 2017 [Publ 3772], fig. 2a-M

iT tT bi riS oT roT roS iD
1 1 0 1 7 1 1 --
  +-- Venustaconcha **
--+                
  +-- Quadrulini   

Krebs, 2004 [Publ 2692], fig. 2-M

iT tT bi riS oT roT roS iD
1 1 0 1 3 1 1 --
  +-- Lampsilis **   
--+               
  +-- Pleurobemini

Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4

iT tT bi riS oT roT roS iD
1 1 0 1 119 3 3 --
  +-- Amblemini   
  |               
  +-- Pleurobemini
--+               
  +-- Lampsilis **   
  |               
  +-- Quadrulini  

Zieritz et al., 2021 [Publ 4218], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 31 1 1 --
  +-- Amblemini
--+            
  +-- Lampsilis **

Vikhrev et al., 2018 [Publ 3812], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 12 0 0 --
-- Lampsilis_cardium

Huff et al., 2004 [Publ 2697], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 10 0 0 --
-- Lampsilis_cardium

Plazzi & Passamonti, 2010 [Publ 3354], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 1 0 0 --
-- Lampsilis_ornata

Jiang et al., 2010 [Publ 3438], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 3 0 0 --
-- Lampsilis_ornata

Lopes-Lima et al., 2018 [Publ 3829], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 27 0 0 --
-- Lampsilis_cardium

Burzynski et al., 2017 [Publ 3774], fig. 1-bottom

iT tT bi riS oT roT roS iD
1 1 0 1 12 1 1 --
  +-- Quadrulini   
--+                
  +-- Venustaconcha **

Inoue et al., 2014 [Publ 3635], fig. 3A

iT tT bi riS oT roT roS iD
1 1 0 1 19 3 3 --
-- Lampsilis_ornata

Combosch et al., 2017 [Publ 3806], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 14 0 0 --
-- Lampsilis_cardium

Baker et al., 2004 [Publ 2685], fig. 2-3

iT tT bi riS oT roT roS iD
1 1 0 1 8 0 0 --
-- Epioblasma_triquetra

Sonowal et al., 2021 [Publ 4221], fig. 3a

iT tT bi riS oT roT roS iD
1 1 0 1 14 1 1 --
-- Lampsilis_cardium

Froufe et al., 2015 [Publ 3664], fig. 1-M

iT tT bi riS oT roT roS iD
1 1 0 1 7 1 1 --
  +-- Venustaconcha **
--+                
  +-- Quadrulini   

Takeuchi et al., 2015 [Publ 3661], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 9 0 0 --
-- Lampsilis_cardium

Combosch et al., 2017 [Publ 3806], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 14 0 0 --
-- Lampsilis_cardium

Burzynski et al., 2017 [Publ 3774], fig. 1-top

iT tT bi riS oT roT roS iD
1 1 0 1 12 1 1 --
  +-- Quadrulini   
--+                
  +-- Venustaconcha **

Burdick & White, 2007 [Publ 2679], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 29 29 2 --
  +-------------------------------- Lampsilis **             
  |                                                       
--+  +----------------------------- Parvaspina            
  |  |                                                    
  |  |  +-------------------------- Amblemini             
  +--+  |                                                 
     |  |                    +----- Eurynia               
     |  |  +-----------------+                            
     +--+  |                 |  +-- Plethobasus           
        |  |                 +--+                         
        |  |                    +-- Elliptio              
        |  |                                              
        |  |  +-------------------- Pleurobema_plenum     
        +--+  |                                           
           |  +-------------------- Pleuronaia            
           |  |                                           
           |  |                 +-- Pleurobema_cordatum__1
           |  |           +-----+                         
           |  |           |     +-- Fusconaia             
           +--+  +--------+                               
              |  |        |  +----- Pleurobema_sintoxia__1
              |  |        |  |                            
              |  |        +--+  +-- Pleurobema_cordatum__2
              |  |           |  |                         
              |  |           +--+-- Pleurobema_sintoxia__2
              |  |              |                         
              +--+              +-- Pleurobema_sintoxia__3
                 |                                        
                 |  +-------------- Pleurobema_stabile    
                 |  |                                     
                 |  |           +-- Pleurobema_rubellum   
                 |  |     +-----+                         
                 |  |     |     +-- Pleurobema_strodeanum 
                 +--+  +--+                               
                    |  |  |  +----- Pleurobema_beadleianum
                    |  |  +--+                            
                    |  |     |  +-- Pleurobema_perovatum  
                    |  |     +--+                         
                    +--+        +-- Pleurobema_taitianum  
                       |                                  
                       |        +-- Pleurobema_clava      
                       |  +-----+                         
                       |  |     +-- Pleurobema_oviforme   
                       |  |                               
                       +--+     +-- Pleurobema_decisum    
                          |  +--+                         
                          |  |  +-- Pleurobema_hanleyianum
                          +--+                            
                             |  +-- Pleurobema_pyriforme  
                             +--+                         
                                +-- Pleurobema_georgianum 

Huang et al., 2013 [Publ 3629], fig. 3-M

iT tT bi riS oT roT roS iD
1 1 0 1 5 1 1 --
  +-- Quadrulini   
--+                
  +-- Venustaconcha **

Sano et al., 2020 [Publ 4138], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 68 0 0 --
-- Lampsilis_cardium

Guerra et al., 2017 [Publ 3772], fig. 2b-M

iT tT bi riS oT roT roS iD
1 1 0 1 7 1 1 --
  +-- Venustaconcha **
--+                
  +-- Quadrulini   

Wu et al., 2018 [Publ 3836], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 52 8 3 --
  +-- Amblemini
--+            
  +-- Obovaria ** 

Froufe et al., 2015 [Publ 3664], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 11 1 1 --
  +-- Venustaconcha **
--+                
  +-- Quadrulini   

Lopes-Lima et al., 2017 [Publ 3792], fig. 2-M

iT tT bi riS oT roT roS iD
1 1 0 1 9 1 1 --
  +-- Venustaconcha **
--+                
  +-- Quadrulini   

Chen et al., 2023 [Publ 4302], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 39 1 1 --
  +-- Lampsilis ** 
--+             
  +-- Quadrulini

Chen et al., 2023 [Publ 4302], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 39 1 1 --
  +-- Lampsilis ** 
--+             
  +-- Quadrulini

Fonseca et al., 2016 [Publ 3775], fig. 1-M

iT tT bi riS oT roT roS iD
1 1 0 1 8 1 1 --
  +-- Venustaconcha **
--+                
  +-- Quadrulini   

Graf, 2000 [Publ 2647], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 16 1 1 --
-- Lampsilis

Cyr et al., 2007 [Publ 2745], fig. 2A

iT tT bi riS oT roT roS iD
1 1 0 1 4 0 0 --
-- Lampsilis_ornata

Lemer et al., 2019 [Publ 4093], fig. 1

iT tT bi riS oT roT roS iD
1 1 0 1 5 0 0 --
-- Lampsilis_cardium

Curole & Kocher, 2002 [Publ 2712], fig. 1a-M

iT tT bi riS oT roT roS iD
1 1 0 1 3 2 1 --
-- Lampsilis_teres

Wu et al., 2020 [Publ 4121], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 33 0 0 --
-- Lampsilis_cardium

Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M

iT tT bi riS oT roT roS iD
1 1 0 1 6 1 1 --
  +-- Venustaconcha **
--+                
  +-- Quadrulini   

Soroka & Burzynski, 2018 [Publ 3762], fig. 3

iT tT bi riS oT roT roS iD
1 1 0 1 6 1 1 --
  +-- Venustaconcha **
--+                
  +-- Quadrulini   

Wu et al., 2020 [Publ 4121], fig. S1

iT tT bi riS oT roT roS iD
1 1 0 1 33 0 0 --
-- Lampsilis_cardium

Huang et al., 2018 [Publ 3828], fig. 3_5

iT tT bi riS oT roT roS iD
1 1 0 1 28 0 0 --
-- Lampsilis_cardium

Campbell et al., 2008 [Publ 2675], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 24 24 3 --
  +----------- Quadrulini               
  |                                     
  +----------- Amblemini                
  |                                     
  |        +-- Elliptio_arctata         
  |  +-----+                            
  |  |     +-- Elliptio_crassidens      
  |  |                                  
  |  |  +----- Pleurobema_decisum       
--+  |  |                               
  |  |  +----- Pleurobema_rubellum      
  |  |  |                               
  |  |  +----- Pleurobema_georgianum__2 
  |  |  |                               
  |  |  +----- Pleurobema_stabile       
  |  |  |                               
  |  |  +----- Pleurobema_pyriforme     
  |  |  |                               
  +--+  +----- Pleurobema_oviforme__2   
     |  |                               
     |  +----- Pleurobema_hanleyianum__2
     |  |                               
     |  +----- Pleuronaia               
     |  |                               
     |  +----- Pleurobema_georgianum__3 
     |  |                               
     |  +----- Pleurobema_cordatum      
     |  |                               
     |  +----- Pleurobema_beadleianum   
     |  |                               
     |  +----- Hemistena                
     +--+                               
        +----- Fusconaia_flava          
        |                               
        +----- Fusconaia_cerina         
        |                               
        +----- Eurynia                  
        |                               
        +----- Elliptio_arca            
        |                               
        +----- Cyrtonaias **               
        |                               
        |  +-- Plethobasus              
        +--+                            
        |  +-- Parvaspina               
        |                               
        |  +-- Pleurobema_sintoxia      
        |  |                            
        +--+-- Pleurobema_oviforme__1   
        |  |                            
        |  +-- Pleurobema_clava         
        |                               
        |  +-- Pleurobema_athearni      
        |  |                            
        +--+-- Pleurobema_georgianum__1 
           |                            
           +-- Pleurobema_hanleyianum__1

Sano et al., 2020 [Publ 4138], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 68 0 0 --
-- Lampsilis_cardium

Krebs, 2004 [Publ 2692], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 4 2 2 --
  +-- Lampsilis **   
--+               
  +-- Pleurobemini

Curole & Kocher, 2002 [Publ 2712], fig. 1-F

iT tT bi riS oT roT roS iD
1 1 0 1 3 2 1 --
  +-- Lampsilis ** 
--+             
  +-- Quadrulini

Bolotov et al., 2016 [Publ 3743], fig. 2

iT tT bi riS oT roT roS iD
1 1 0 1 29 0 0 --
-- Lampsilis_cardium

Baker et al., 2003 [Publ 2690], fig. 5

iT tT bi riS oT roT roS iD
1 1 0 1 7 0 0 --
-- Epioblasma_triquetra

Sano et al., 2020 [Publ 4138], fig. 4

iT tT bi riS oT roT roS iD
1 1 0 1 68 0 0 --
-- Lampsilis_cardium

Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2

iT tT bi riS oT roT roS iD
1 1 0 1 10 1 1 --
  +-- Amblemini
--+            
  +-- Lampsilis **

Page updated: 08:34:02 Wed 06 Nov 2024

 
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