Bivalvia | Unionoida | Unionoidea | Unionidae subfamily UNIONINAE Rafinesque, 1820: cladomics 5 tribes
Cladomics Summary
number of species = 241
number of subtaxa (rank: tribe) = 5
analyzed in 379 cladograms in 210 publications
maximum number of terminals in an analysis = 69, mean = 10.27
maximum number of subtaxa (tribe) in an analysis = 5, mean = 2.0
Basic Monophyly
number of cladograms that test basic monophyly (i.e., more than 1 included species) = 332
maximum number of included terminals = 69, mean = 11.58
maximum number of subtaxa (tribe) = 5, mean = 2.14
proportion of cladograms that support basic monophyly = 0.92
Rigorous Monophyly
number of cladograms that test rigorous monophyly = 159
(Rigorous monophyly refers to cladograms with terminal species representing more than one subtaxon (tribe) taxon and at least one rigorous outgroup (subfamily).
maximum number of included terminals = 69, mean = 15.55
maximum number of subtaxa (tribe) = 5, mean = 3.09
proportion of cladograms that support rigorous monophyly = 0.87
Pairwise Cladogram Clustering
The graphs below depict in two dimensions all pair-wise ingroup distances within the cladome. See the Cladmoics FARQ for more information.
Wu et al., 2023 [Publ 4339], fig. 4_S5-aa
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
47 |
47 |
45 |
5 |
49 |
38 |
3 |
0.0 |
*** Top Monophyly Cladogram ***
+-------------------- Ambleminae
--+
| +----------------- Gonideinae
+--+
| +-------------- Parreysiinae
+--+
| +----------- Lepidodesmini **
+--+
| +-------- Unionini **
+--+
| +----- Anodontini **
+--+
| +-- Acuticostini **
+--+
+-- Lanceolariini **
Huang et al., 2019 [Publ 3843], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
44 |
44 |
35 |
5 |
154 |
115 |
3 |
50.0 |
*** Top Sister-Group Cladogram ***
+-------------------- Parreysiinae
|
| +-------- Acuticostini **
| |
--+ +--------+ +----- Lepidodesmini **
| | | |
| | +--+----- Unionini **
| | |
| | | +-- Lanceolariini **
+--+ +--+
| +-- Anodontini **
|
| +-- Rectidentini
| +-----------+
| | +-- Contradentini
+--+
| +----------- Ambleminae
+--+
| +-------- Chamberlainiini
+--+
| +----- Lamprotulini
+--+
| +-- Gonideini
+--+
+-- Pseudodontini
Lopes-Lima et al., 2020 [Publ 4080], fig. 2-4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
69 |
69 |
57 |
5 |
51 |
40 |
3 |
65.0 |
+----------------- Parreysiinae
|
| +----- Acuticostini **
| |
| +----- Lepidodesmini **
| +--------+
--+ | +----- Unionini **
| | |
| | | +-- Lanceolariini **
| | +--+
| | +-- Anodontini **
| |
+--+ +-- Lens
| |
| +-- Ensidens
| |
| +--------+-- Physunio
| | |
| | +-- Rectidens
| | |
+--+ +-- Yaukthwa
|
| +-------- Ambleminae
| |
+--+ +-- Chamberlainiini
| +--+
| | +-- Lamprotulini
+--+
| +-- Pseudodontini
+--+
+-- Gonideini
Wu et al., 2024 [Publ 4379], fig. 5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
52 |
52 |
51 |
4 |
2 |
0 |
0 |
27.0 |
+-------- Lepidodesmini **
--+
| +----- Unionini **
+--+
| +-- Lanceolariini **
+--+
+-- Anodontini **
Wu et al., 2023 [Publ 4339], fig. 4_S5-nt
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
47 |
47 |
43 |
5 |
49 |
38 |
3 |
16.0 |
+-------------------- Ambleminae
--+
| +----------------- Gonideinae
+--+
| +-------------- Parreysiinae
+--+
| +----------- Lepidodesmini **
+--+
| +-------- Unionini **
+--+
| +----- Anodontini **
+--+
| +-- Acuticostini **
+--+
+-- Lanceolariini **
Wu et al., 2024 [Publ 4350], fig. 8b_S2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
47 |
47 |
44 |
5 |
45 |
39 |
3 |
13.0 |
+-------------------- Ambleminae
--+
| +----------------- Gonideinae
+--+
| +-------------- Parreysiinae
+--+
| +----------- Lepidodesmini **
+--+
| +-------- Unionini **
+--+
| +----- Anodontini **
+--+
| +-- Acuticostini **
+--+
+-- Lanceolariini **
Wu et al., 2024 [Publ 4350], fig. 8a_S1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
47 |
47 |
44 |
5 |
45 |
39 |
3 |
13.0 |
+-------------------- Ambleminae
--+
| +----------------- Gonideinae
+--+
| +-------------- Parreysiinae
+--+
| +----------- Lepidodesmini **
+--+
| +-------- Unionini **
+--+
| +----- Anodontini **
+--+
| +-- Acuticostini **
+--+
+-- Lanceolariini **
Huang et al., 2019 [Publ 3843], fig. S4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
44 |
44 |
21 |
5 |
154 |
115 |
3 |
53.0 |
+-------------- Parreysiinae
|
| +----------- Chamberlainia
| |
| +----------- Megalonaias
| |
| +----------- Uniomerus
| |
| +----------- Quadrulina
| |
| +----------- Sinohyriopsis
| |
| +----------- Lamprotulini
| |
| +----------- Gonideini
| |
| +----------- Pseudodontini
| |
| | +-- Rectidentini
--+ +--------+
| | +-- Contradentini
| |
| | +-------- Amblema
| | |
| | +-------- Toxolasma
| | |
| | +-------- Pleurobemini
| | |
| | | +-- Obliquaria
| | +-----+
| | | +-- Plectomerus
| +--+
| | | +----- Ellipsaria
| | | |
| | | +----- Potamilus_leptodon
| | | |
| | | +----- Medionidus
| | | |
| | | +----- Potamilus_alatus
+--+ | |
| | +----- Ptychobranchus
| +--+
| +----- Truncilla
| |
| +----- Venustaconcha
| |
| +----- Cambarunio
| |
| +----- Leaunio
| |
| | +-- Ortmanniana
| +--+
| +-- Lampsilis
|
| +----- Acuticostini **
| |
| +----- Alasmidonta **
| |
| +----- Anodonta_anatina **
| |
| +----- Anemina **
| |
| +----- Anodonta_cygnea **
| |
| +----- Cristaria **
| |
| +----- Anodonta_nuttalliana **
| |
| +----- Lasmigona **
| |
| +----- Lepidodesmini **
| |
| +----- Pseudanodonta **
| |
| +----- Pyganodon **
| |
+-----+----- Simpsonaias **
|
+----- Strophitus **
|
+----- Schistodesmus **
|
+----- Utterbackia **
|
+----- Lanceolariini **
|
+----- Unio **
|
+----- Aculamprotula **
|
| +-- Pletholophus **
+--+
| +-- Sinanodonta **
|
| +-- Cuneopsis_heudei **
| |
| +-- Cuneopsis_celtiformis **
| |
+--+-- Tchangsinaia **
|
+-- Cuneopsis_rufescens **
|
+-- Nodularia **
Huang et al., 2019 [Publ 3843], fig. S3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
43 |
43 |
40 |
5 |
153 |
114 |
3 |
50.0 |
+----------------------- Parreysiinae
|
--+ +-------------------- Pseudodontini
| |
| | +-- Rectidentini
+--+ +--------------+
| | +-- Contradentini
| |
| | +-------- Chamberlainiini
+--+ +-----+
| | | +----- Ambleminae
| | +--+
| | | +-- Lamprotulini
+--+ +--+
| +-- Gonideini
|
| +----------- Acuticostini **
+--+
| +-------- Unionini **
+--+
| +----- Lepidodesmini **
+--+
| +-- Lanceolariini **
+--+
+-- Anodontini **
Whelan et al., 2023 [Publ 4287], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
41 |
41 |
42 |
1 |
0 |
0 |
0 |
68.0 |
-- (Anodontini
Wu et al., 2022 [Publ 4238], fig. 2d
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
39 |
39 |
37 |
5 |
0 |
0 |
0 |
24.0 |
+----------- Lepidodesmini **
--+
| +-------- Unionini **
+--+
| +----- Lanceolariini **
+--+
| +-- Acuticostini **
+--+
+-- Anodontini **
Wu et al., 2022 [Publ 4238], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
36 |
36 |
34 |
5 |
38 |
29 |
2 |
19.0 |
+----------------- Ambleminae
--+
| +-------------- Gonideinae
+--+
| +----------- Lepidodesmini **
+--+
| +-------- Unionini **
+--+
| +----- Anodontini **
+--+
| +-- Acuticostini **
+--+
+-- Lanceolariini **
Sano et al., 2020 [Publ 4138], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
36 |
36 |
37 |
4 |
33 |
10 |
3 |
43.0 |
+----------------------- Parreysiinae
|
| +-- Ambleminae
--+ +-----------------+
| | +-- Gonideinae
| |
+--+ +-- Buldowskia **
| +--------------+
| | +-- Anemina **
| |
+--+ +----- Pletholophus **
| +--------+
| | | +-- Beringiana **
| | +--+
+--+ +-- Sinanodonta **
|
| +----------- Cristaria **
| |
+--+ +-- Anodontina **
| +-----+
| | +-- Alasmidontina **
+--+
| +----- Lanceolariini **
+--+
| +-- Lepidodesmini **
+--+
+-- Unionini **
Sano et al., 2020 [Publ 4138], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
36 |
36 |
38 |
4 |
33 |
10 |
3 |
43.0 |
+----------------------- Parreysiinae
|
| +-------- Chamberlainiini
| +-----------+
--+ | | +----- Ambleminae
| | +--+
| | | +-- Lamprotulini
| | +--+
+--+ +-- Gonideini
|
| +-- Buldowskia **
| +--------------+
| | +-- Anemina **
| |
+--+ +----- Sinanodonta **
| +--------+
| | | +-- Pletholophus **
| | +--+
+--+ +-- Beringiana **
|
| +----------- Cristaria **
| |
+--+ +-- Anodontina **
| +-----+
| | +-- Alasmidontina **
+--+
| +----- Lanceolariini **
+--+
| +-- Lepidodesmini **
+--+
+-- Unionini **
Sano et al., 2020 [Publ 4138], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
36 |
36 |
38 |
4 |
33 |
10 |
3 |
43.0 |
+----------------------- Parreysiinae
|
| +-- Ambleminae
--+ +-----------------+
| | +-- Gonideinae
| |
+--+ +-- Buldowskia **
| +--------------+
| | +-- Anemina **
| |
+--+ +----- Sinanodonta **
| +--------+
| | | +-- Pletholophus **
| | +--+
+--+ +-- Beringiana **
|
| +----------- Cristaria **
| |
+--+ +-- Anodontina **
| +-----+
| | +-- Alasmidontina **
+--+
| +----- Lepidodesmini **
+--+
| +-- Lanceolariini **
+--+
+-- Unionini **
Wu et al., 2021 [Publ 4366], fig. 3-ML
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
34 |
34 |
32 |
5 |
40 |
29 |
2 |
23.0 |
+----------------- Ambleminae
--+
| +-------------- Gonideinae
+--+
| +----------- Lepidodesmini **
+--+
| +-------- Unionini **
+--+
| +----- Anodontini **
+--+
| +-- Acuticostini **
+--+
+-- Lanceolariini **
Wu et al., 2021 [Publ 4366], fig. 3-BI
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
34 |
34 |
32 |
5 |
40 |
29 |
2 |
25.0 |
+----------------- Ambleminae
--+
| +-------------- Gonideinae
+--+
| +----------- Lepidodesmini **
+--+
| +-------- Unionini **
+--+
| +----- Anodontini **
+--+
| +-- Acuticostini **
+--+
+-- Lanceolariini **
Wu et al., 2018 [Publ 3836], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
33 |
33 |
28 |
5 |
20 |
18 |
2 |
39.0 |
+----------------- Quadrulini
|
+----------------- Pleurobemini
|
| +-- Amblemini
--+--------------+
| +-- Lampsilini
|
| +-- Lamprotulini
| +-----------+
| | +-- Gonideini
+--+
| +----------- Chamberlainiini
| |
+--+ +-------- Lepidodesmini **
| |
+--+-------- Unionini **
|
| +----- Anodontini **
+--+
| +-- Acuticostini **
+--+
+-- Lanceolariini **
Wu et al., 2021 [Publ 4366], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
33 |
33 |
31 |
5 |
36 |
25 |
2 |
24.0 |
+----------------- Ambleminae
--+
| +-------------- Gonideinae
+--+
| +----------- Lepidodesmini **
+--+
| +-------- Unionini **
+--+
| +----- Anodontini **
+--+
| +-- Acuticostini **
+--+
+-- Lanceolariini **
Konopleva et al., 2023 [Publ 4269], fig. S1_4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
33 |
33 |
31 |
5 |
23 |
22 |
3 |
50.0 |
+-------------- Parreysiinae
|
--+ +-- Ambleminae
| +--------+
| | +-- Gonideinae
+--+
| +-------- Unionini **
| |
+--+ +-- Lepidodesmini **
| +--+
| | +-- Lanceolariini **
+--+
| +-- Acuticostini **
+--+
+-- Anodontini **
Chen et al., 2023 [Publ 4302], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
31 |
31 |
27 |
5 |
9 |
7 |
3 |
53.0 |
+-- Gonideinae
+-----------+
| +-- Ambleminae
--+
| +----------- Parreysiinae
| |
+--+ +-------- Acuticostini **
| |
+--+ +----- Lepidodesmini **
| |
+--+----- Unionini **
|
| +-- Lanceolariini **
+--+
+-- Anodontini **
Huang et al., 2019 [Publ 3843], fig. S1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
31 |
31 |
40 |
5 |
14 |
12 |
1 |
61.0 |
+----------------- Gonideinae
|
--+ +-------------- Anemina **
| |
+--+ +----------- Sinanodonta **
| |
+--+ +-------- Acuticostini **
| |
+--+ +-- Lepidodesmini **
| +--+
| | +-- Cristaria **
+--+
| +-- Lanceolariini **
+--+
+-- Unionini **
Chen et al., 2023 [Publ 4302], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
31 |
31 |
23 |
5 |
9 |
7 |
3 |
54.0 |
+-- Gonideinae
+--------------+
| +-- Ambleminae
|
--+ +-------------- Parreysiinae
| |
| | +-- Anemina **
+--+ +--------+
| | +-- Amuranodonta **
| |
| | +----- Pletholophus **
+--+ +--+
| | | +-- Sinanodonta **
| | +--+
| | +-- Beringiana **
| |
+--+ +----- Lanceolaria_gladiola **
| |
| +----- Acuticostini **
| |
| +----- Lepidodesmini **
| |
+--+----- Lanceolaria_oxyrhyncha **
|
+----- Cristaria **
|
+----- Unionini **
|
| +-- Alasmidontina **
+--+
+-- Anodontina **
Bogan et al., 2023 [Publ 4295], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
29 |
29 |
26 |
2 |
2 |
0 |
0 |
62.0 |
+-- Unionini **
--+
+-- Anodontini **
Wu et al., 2020 [Publ 4121], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
28 |
28 |
26 |
5 |
22 |
14 |
2 |
26.0 |
+----------------- Ambleminae
--+
| +-------------- Gonideinae
+--+
| +----------- Lepidodesmini **
+--+
| +-------- Unionini **
+--+
| +----- Anodontini **
+--+
| +-- Acuticostini **
+--+
+-- Lanceolariini **
Lyubas et al., 2022 [Publ 4255], fig. 7
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
28 |
28 |
25 |
2 |
3 |
3 |
2 |
61.0 |
+-------- Parreysiinae
--+
| +----- Gonideinae
+--+
| +-- Anodontini **
+--+
+-- Unionini **
Whelan et al., 2023 [Publ 4287], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
28 |
28 |
27 |
1 |
0 |
0 |
0 |
66.0 |
-- (Anodontini
Lyubas et al., 2022 [Publ 4255], fig. S1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
28 |
28 |
19 |
2 |
3 |
3 |
2 |
61.0 |
+-------- Parreysiinae
--+
| +----- Gonideinae
+--+
| +-- Anodontini **
+--+
+-- Unionini **
Lopes-Lima et al., 2017 [Publ 3745], fig. 1-2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
28 |
28 |
25 |
3 |
52 |
40 |
3 |
46.0 |
+----- Unionini **
--+
| +-- Lanceolariini **
+--+
+-- Anodontini **
Huang et al., 2019 [Publ 3843], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
28 |
28 |
26 |
5 |
14 |
12 |
1 |
54.0 |
+-- Lamprotulini
+-----------------------------------+
| +-- Gonideini
--+
| +----------------------------------- Chamberlainiini
| |
+--+ +-------------------------------- Anemina **
| |
+--+ +----------------------------- Acuticostini **
| |
+--+ +-------------------------- Sinanodonta **
| |
+--+ +----------------------- Aculamprotula **
| |
+--+ +-- Cristaria **
| +-----------------+
| | +-- Lanceolariini **
+--+
| +----------------- Lepidodesmini **
+--+
| +-------------- Schistodesmus **
+--+
| +----------- Nodularia **
+--+
| +-------- Cuneopsis_rufescens **
+--+
| +----- Tchangsinaia **
+--+
| +-- Cuneopsis_heudei **
+--+
+-- Cuneopsis_celtiformis **
Fukata & Iigo, 2020 [Publ 4106], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
28 |
28 |
26 |
5 |
26 |
23 |
2 |
30.0 |
+-- Ambleminae
+-----------+
| +-- Gonideinae
--+
| +----------- Lepidodesmini **
+--+
| +-------- Lanceolariini **
+--+
| +----- Unionini **
+--+
| +-- Acuticostini **
+--+
+-- Anodontini **
Aunins et al., 2018 [Publ 3863], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
27 |
27 |
25 |
5 |
14 |
11 |
2 |
23.0 |
+----------------- Ambleminae
--+
| +-------------- Gonideinae
+--+
| +----------- Lepidodesmini **
+--+
| +-------- Unionini **
+--+
| +----- Anodontini **
+--+
| +-- Acuticostini **
+--+
+-- Lanceolariini **
Huang et al., 2019 [Publ 3843], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
27 |
27 |
25 |
5 |
30 |
20 |
2 |
28.0 |
+-- Gonideinae
+-----------+
| +-- Ambleminae
--+
| +----------- Lepidodesmini **
+--+
| +-------- Unionini **
+--+
| +----- Lanceolariini **
+--+
| +-- Acuticostini **
+--+
+-- Anodontini **
Dai et al., 2024 [Publ 4385], fig. 3-ML
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
27 |
27 |
25 |
1 |
4 |
2 |
1 |
64.0 |
+-- Parreysiinae
--+
+-- Anodontini **
Bolotov et al., 2017 [Publ 3761], fig. S1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
27 |
27 |
25 |
3 |
112 |
87 |
3 |
49.0 |
+-------------------- Parreysiinae
|
| +-- Rectidentini
--+ +--------------+
| | +-- Contradentini
| |
+--+ +-------------- Pseudodontini
| |
| | +----- Unionini **
+--+ +-----+
| | | +-- Lanceolariini **
| | +--+
+--+ +-- Anodontini **
|
| +-------- Ambleminae
+--+
| +----- Gonideini
+--+
| +-- Chamberlainiini
+--+
+-- Lamprotulini
Bolotov et al., 2017 [Publ 3761], fig. S2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
27 |
27 |
18 |
3 |
112 |
87 |
3 |
52.0 |
+-------------------- Parreysiinae
|
| +-- Contradentini
| +-----------+
| | +-- Rectidentini
--+ +--+
| | | +----------- Ambleminae
| | +--+
| | | +-------- Pseudodontini
| | +--+
| | | +----- Gonideini
+--+ +--+
| | +-- Chamberlainiini
| +--+
| +-- Lamprotulini
|
| +----------- Unionini **
| |
+-----+ +-------- Lanceolariini **
| |
| +-------- Cristariina **
+--+
| +----- Anodonta_anatina **
| |
| +----- Pseudanodonta **
+--+
+----- Alasmidontina **
|
| +-- Anodonta_cygnea **
+--+
+-- Anodonta_nuttalliana **
Dai et al., 2024 [Publ 4358], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
27 |
27 |
25 |
5 |
25 |
23 |
2 |
34.0 |
+----------------- Ambleminae
--+
| +-------------- Gonideinae
+--+
| +----------- Lepidodesmini **
+--+
| +-------- Unionini **
+--+
| +----- Lanceolariini **
+--+
| +-- Acuticostini **
+--+
+-- Anodontini **
Bolotov et al., 2017 [Publ 3764], fig. S1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
27 |
27 |
2 |
3 |
98 |
98 |
3 |
55.0 |
+----------------- Parreysiinae
|
| +-------------- Pseudodontini
| |
--+ | +-- Rectidentini
| | +--------+
| | | +-- Contradentini
| | |
| | | +----- Ambleminae
+--+ | |
| | | +-- Chamberlainia
| | | |
| | | +-- Sinohyriopsis
| | | |
| | | +-- Lamprotula_caveata
| | +--+ |
| | | | +-- Lamprotula_leaii
+--+ | | |
| | | +-- Potomida
| | | |
| | | +-- Pronodularia
| | | |
| | +--+-- Gonidea
| | |
| | +-- Leguminaia_wheatleyi
| | |
| | +-- Leguminaia
| | |
| | +-- Microcondylaea
| | |
+--+ +-- Sinosolenaia_carinata
| |
| +-- Sinosolenaia_oleivora
| |
| +-- Sinosolenaia_recognita
|
| +----- Unionini **
| |
| | +-- Alasmidonta **
| | |
| | +-- Anodonta_anatina **
| | |
| | +-- Anodonta_cygnea **
| | |
| | +-- Anodonta_nuttalliana **
+--+ |
| +-- Lasmigona **
| |
| +-- Pseudanodonta **
| |
| +-- Pyganodon **
| |
| +-- Simpsonaias **
| |
| +-- Strophitus **
+--+
+-- Sinanodonta_lauta **
|
+-- Cristaria_plicata **
|
+-- Cristaria_radiata **
|
+-- Pletholophus **
|
+-- Sinanodonta_lucida **
|
+-- Sinanodonta_jourdyi **
|
+-- Lanceolaria_gladiola **
|
+-- Lanceolaria_grayii__1 **
|
+-- Lanceolaria_grayii__2 **
Whelan et al., 2023 [Publ 4287], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
27 |
27 |
25 |
1 |
0 |
0 |
0 |
65.0 |
-- (Anodontini
Dai et al., 2024 [Publ 4385], fig. 3-BI
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
27 |
27 |
24 |
1 |
4 |
2 |
1 |
64.0 |
+-- Parreysiinae
--+
+-- Anodontini **
Wu et al., 2019 [Publ 3864], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
27 |
27 |
25 |
5 |
20 |
14 |
2 |
25.0 |
+----------------- Ambleminae
--+
| +-------------- Gonideinae
+--+
| +----------- Lepidodesmini **
+--+
| +-------- Unionini **
+--+
| +----- Anodontini **
+--+
| +-- Acuticostini **
+--+
+-- Lanceolariini **
Jeratthitikul et al., 2022 [Publ 4260], fig. S1_1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
26 |
26 |
24 |
3 |
50 |
40 |
2 |
46.0 |
+-------------------- Pseudodontini
|
--+ +-- Contradentini
| +--------------+
| | +-- Rectidentini
+--+
| +-------------- Gonideini
| |
+--+ +-- Chamberlainiini
| +--------+
| | +-- Lamprotulini
+--+
| +-------- Ambleminae
+--+
| +----- Unionini **
+--+
| +-- Lanceolariini **
+--+
+-- Anodontini **
Wu et al., 2021 [Publ 4366], fig. 6
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
26 |
26 |
24 |
5 |
65 |
48 |
3 |
41.0 |
+-------------------- Parreysiinae
|
| +----------- Acuticostini **
| +-----+
--+ | | +-------- Unionini **
| | +--+
| | | +----- Lanceolariini **
| | +--+
| | | +-- Lepidodesmini **
+--+ +--+
| +-- Anodontini **
|
| +-- Contradentini
| +-----------+
| | +-- Rectidentini
+--+
| +----------- Ambleminae
+--+
| +-------- Chamberlainiini
+--+
| +----- Lamprotulini
+--+
| +-- Pseudodontini
+--+
+-- Gonideini
Dai et al., 2024 [Publ 4386], fig. 6
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
26 |
26 |
25 |
5 |
14 |
14 |
2 |
37.0 |
+----------------- Ambleminae
--+
| +-------------- Gonideinae
+--+
| +----------- Lepidodesmini **
+--+
| +-------- Unionini **
+--+
| +----- Lanceolariini **
+--+
| +-- Acuticostini **
+--+
+-- Anodontini **
Wu et al., 2021 [Publ 4366], fig. 2_I
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
26 |
26 |
24 |
5 |
65 |
48 |
3 |
39.0 |
+-------------------- Parreysiinae
|
| +-------- Acuticostini **
| |
--+ +--------+ +-- Lepidodesmini **
| | | +--+
| | | | +-- Unionini **
| | +--+
| | | +-- Lanceolariini **
+--+ +--+
| +-- Anodontini **
|
| +-- Rectidentini
| +-----------+
| | +-- Contradentini
+--+
| +----------- Ambleminae
+--+
| +-------- Chamberlainiini
+--+
| +----- Lamprotulini
+--+
| +-- Pseudodontini
+--+
+-- Gonideini
Jeratthitikul et al., 2022 [Publ 4260], fig. S2_2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
26 |
26 |
24 |
3 |
50 |
40 |
2 |
48.0 |
+-------------------- Pseudodontini
|
--+ +-- Rectidentini
| +--------------+
| | +-- Contradentini
+--+
| +-------------- Gonideini
| |
+--+ +-- Chamberlainiini
| +--------+
| | +-- Lamprotulini
+--+
| +-------- Ambleminae
+--+
| +----- Unionini **
+--+
| +-- Lanceolariini **
+--+
+-- Anodontini **
Soroka & Burzynski, 2017 [Publ 3837], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
24 |
24 |
22 |
4 |
0 |
0 |
0 |
32.0 |
+-------- Anodontini **
--+
| +----- Lanceolariini **
+--+
| +-- Lepidodesmini **
+--+
+-- Unionini **
Froufe et al., 2019 [Publ 3936], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
23 |
23 |
21 |
4 |
22 |
19 |
2 |
27.0 |
+-------------- Ambleminae
--+
| +----------- Gonideinae
+--+
| +-------- Lepidodesmini **
+--+
| +----- Unionini **
+--+
| +-- Lanceolariini **
+--+
+-- Anodontini **
Froufe et al., 2019 [Publ 3936], fig. 6
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
23 |
23 |
21 |
4 |
22 |
19 |
2 |
27.0 |
+-------------- Ambleminae
--+
| +----------- Gonideinae
+--+
| +-------- Lepidodesmini **
+--+
| +----- Unionini **
+--+
| +-- Lanceolariini **
+--+
+-- Anodontini **
Zieritz et al., 2021 [Publ 4231], fig. 3-F_5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
23 |
23 |
21 |
4 |
26 |
23 |
2 |
27.0 |
+-------------- Ambleminae
--+
| +----------- Gonideinae
+--+
| +-------- Lepidodesmini **
+--+
| +----- Unionini **
+--+
| +-- Lanceolariini **
+--+
+-- Anodontini **
Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_AHE
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
22 |
22 |
20 |
3 |
0 |
0 |
0 |
52.0 |
+----- Unionini **
--+
| +-- Lanceolariini **
+--+
+-- Anodontini **
Liu et al., 2023 [Publ 4310], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
22 |
22 |
20 |
4 |
2 |
2 |
1 |
49.0 |
+-------------- Gonideinae
|
--+ +----------- Acuticostini **
| |
+--+ +-- Lanceolaria_lanceolata **
| +-----+
| | +-- Lanceolaria_grayii **
+--+
| +----- Lanceolaria_gladiola **
+--+
| +-- Anodontini **
+--+
+-- Unionini **
Liu et al., 2023 [Publ 4310], fig. 5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
22 |
22 |
15 |
4 |
2 |
2 |
1 |
49.0 |
+----------- Gonideinae
|
--+ +-------- Acuticostini **
| |
+--+ +----- Lanceolaria_grayii **
| |
| +----- Lanceolaria_gladiola **
+--+
+----- Lanceolaria_lanceolata **
|
| +-- Anodontini **
+--+
+-- Unionini **
Cha et al., 2018 [Publ 3916], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
22 |
22 |
20 |
4 |
17 |
14 |
2 |
29.5 |
+-------------- Ambleminae
--+
| +----------- Gonideinae
+--+
| +-------- Lepidodesmini **
+--+
| +----- Unionini **
+--+
| +-- Lanceolariini **
+--+
+-- Anodontini **
Lopes-Lima et al., 2024 [Publ 4376], fig. 4-5_MtG
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
22 |
22 |
20 |
3 |
0 |
0 |
0 |
52.0 |
+----- Unionini **
--+
| +-- Lanceolariini **
+--+
+-- Anodontini **
Sano et al., 2020 [Publ 4138], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
22 |
22 |
5 |
5 |
12 |
8 |
1 |
61.0 |
+-------- Gonideinae
|
| +----- Lanceolaria_grayii__1 **
| |
| +----- Lanceolaria_grayii__2 **
| |
| +----- Cristaria_plicata__1 **
| |
| +----- Cristaria_plicata__2 **
| |
| +----- Sinanodonta_lauta **
| |
| +----- Beringiana_japonica **
--+ |
| +----- Beringiana_fukuharai__1 **
| |
| +----- Beringiana_beringiana **
| |
| +----- Beringiana_fukuharai__2 **
| |
| +----- Beringiana_fukuharai__3 **
| |
| +----- Buldowskia_iwakawai__1 **
| |
| +----- Buldowskia_iwakawai__2 **
| |
| +----- Sinanodonta_tumens__1 **
+--+
+----- Sinanodonta_tumens__2 **
|
+----- Pletholophus **
|
+----- Anemina **
|
+----- Lanceolaria_lanceolata **
|
+----- Cristaria_plicata__3 **
|
+----- Lanceolaria_gladiola **
|
+----- Sinanodonta_angula **
|
+----- Lepidodesmini **
|
| +-- Aculamprotula **
+--+
| +-- Acuticostini **
|
| +-- Cuneopsis **
+--+
| +-- Nodularia **
|
| +-- Schistodesmus **
+--+
+-- Inversiunio **
Sano et al., 2017 [Publ 3799], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
21 |
21 |
22 |
5 |
12 |
8 |
1 |
57.0 |
+----------------- Gonideinae
|
--+ +-------------- Anodontini **
| |
| | +-- Tchangsinaia **
+--+ +--------+
| | +-- Nodularia **
| |
+--+ +-- Schistodesmus **
| +-----+
| | +-- Acuticostini **
+--+
| +----- Inversiunio **
+--+
| +-- Lepidodesmini **
+--+
+-- Lanceolariini **
Wu et al., 2023 [Publ 4296], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
21 |
21 |
19 |
4 |
2 |
2 |
1 |
46.0 |
+----------- Gonideinae
--+
| +-------- Acuticostini **
+--+
| +----- Lanceolariini **
+--+
| +-- Anodontini **
+--+
+-- Unionini **
Inoue et al., 2018 [Publ 3858], fig. 1_3a-c
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
20 |
20 |
0 |
1 |
84 |
83 |
2 |
59.0 |
+----- Anodontini **
--+
| +-- Gonideinae
+--+
+-- Ambleminae
Lopes-Lima et al., 2017 [Publ 3792], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
19 |
19 |
17 |
3 |
14 |
11 |
2 |
35.0 |
+----------- Ambleminae
--+
| +-------- Gonideinae
+--+
| +----- Anodontini **
+--+
| +-- Lanceolariini **
+--+
+-- Unionini **
Huang et al., 2018 [Publ 3828], fig. 6
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
19 |
19 |
17 |
2 |
24 |
14 |
2 |
33.0 |
+-- Ambleminae
+--+
| +-- Gonideinae
--+
| +-- Unionini **
+--+
+-- Anodontini **
Lopes-Lima et al., 2021 [Publ 4217], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
19 |
19 |
17 |
2 |
6 |
6 |
1 |
54.0 |
+----- Gonideinae
--+
| +-- Anodontini **
+--+
+-- Unionini **
Fonseca et al., 2016 [Publ 3775], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
19 |
19 |
17 |
3 |
13 |
11 |
2 |
35.0 |
+----------- Ambleminae
--+
| +-------- Gonideinae
+--+
| +----- Anodontini **
+--+
| +-- Lanceolariini **
+--+
+-- Unionini **
Bolotov et al., 2020 [Publ 4084], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
19 |
19 |
17 |
3 |
17 |
13 |
2 |
50.0 |
+-------- Gonideinae
--+
| +----- Unionini **
+--+
| +-- Lanceolariini **
+--+
+-- Anodontini **
Wang et al., 2018 [Publ 3865], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
18 |
18 |
16 |
4 |
14 |
14 |
2 |
32.0 |
+-- Ambleminae
+--------+
| +-- Gonideinae
--+
| +-------- Unionini **
+--+
| +----- Anodontini **
+--+
| +-- Acuticostini **
+--+
+-- Lanceolariini **
Wang et al., 2018 [Publ 3846], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
18 |
18 |
16 |
3 |
14 |
14 |
2 |
34.0 |
+-- Ambleminae
+-----+
| +-- Gonideinae
--+
| +----- Anodontini **
+--+
| +-- Lanceolariini **
+--+
+-- Unionini **
Riccardi et al., 2020 [Publ 4089], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
18 |
18 |
16 |
3 |
2 |
0 |
0 |
39.0 |
+----- Anodontini **
--+
| +-- Lanceolariini **
+--+
+-- Unionini **
Smith et al., 2018 [Publ 3790], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
18 |
18 |
17 |
1 |
0 |
0 |
0 |
58.0 |
-- (Anodontini
Chong et al., 2008 [Publ 2668], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
18 |
18 |
10 |
2 |
0 |
0 |
0 |
56.0 |
+----------------- Anodonta_cygnea **
|
+----------------- Pyganodon **
|
+----------------- Alasminota **
|
+----------------- Anodonta_anatina **
|
+----------------- Unionini **
|
+----------------- Cristariina **
|
| +-- Anodonta_nuttalliana **
--+--------------+
| +-- Anodonta_californiensis **
|
| +-- Pseudodontoideus **
| |
+--------------+-- Alasmidonta **
| |
| +-- Lasmigona **
|
| +-------------- Pseudanodonta **
| |
| | +----- Strophitus **
| +--------+
+--+ | +-- Prolasmidonta **
| +--+
| +-- Platynaias **
|
| +----------- Anodonta_oregonensis__4 **
+--+
| +-------- Anodonta_oregonensis__3 **
+--+
| +----- Anodonta_oregonensis__2 **
+--+
| +-- Anodonta_oregonensis__1 **
+--+
+-- Anodonta_kennerlyi **
Wu et al., 2024 [Publ 4388], fig. 6
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
18 |
18 |
16 |
4 |
36 |
34 |
3 |
35.0 |
+----------------- Ambleminae
--+
| +-------------- Gonideinae
+--+
| +----------- Parreysiinae
+--+
| +-------- Lepidodesmini **
+--+
| +----- Unionini **
+--+
| +-- Lanceolariini **
+--+
+-- Anodontini **
Sano et al., 2022 [Publ 4311], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
18 |
18 |
16 |
1 |
0 |
0 |
0 |
55.0 |
-- (Cristariina
Araujo et al., 2018 [Publ 3779], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
17 |
17 |
13 |
2 |
1 |
0 |
0 |
51.0 |
+-- Anodontini **
--+
+-- Unionini **
Dai et al., 2023 [Publ 4285], fig. 5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
17 |
17 |
15 |
5 |
36 |
34 |
2 |
35.0 |
+----------------- Ambleminae
--+
| +-------------- Gonideinae
+--+
| +----------- Lepidodesmini **
+--+
| +-------- Unionini **
+--+
| +----- Anodontini **
+--+
| +-- Acuticostini **
+--+
+-- Lanceolariini **
Wu et al., 2022 [Publ 4238], fig. 2e
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
17 |
17 |
15 |
3 |
0 |
0 |
0 |
35.0 |
+----- Anodontini **
--+
| +-- Lanceolariini **
+--+
+-- Unionini **
Wu et al., 2024 [Publ 4379], fig. 4_6
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
17 |
17 |
16 |
2 |
5 |
3 |
1 |
48.0 |
+----------- Gonideinae
--+
| +-------- Aculamprotula **
+--+
| +----- Anodontini **
+--+
| +-- Schistodesmus **
+--+
+-- Nodularia **
Pavan-Kumar et al., 2022 [Publ 4263], fig. S5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
17 |
17 |
15 |
5 |
22 |
20 |
3 |
40.0 |
+-- Ambleminae
+--------------------+
| +-- Gonideinae
--+
| +-------------------- Parreysiinae
| |
+--+ +----------------- Anodontini **
| |
| | +-- Acuticostini **
+--+ +-----------+
| | +-- Lanceolariini **
| |
+--+ +-- Lepidodesmini **
| +--------+
| | +-- Aculamprotula **
+--+
| +-------- Unio **
+--+
| +----- Schistodesmus **
+--+
| +-- Nodularia **
+--+
+-- Tchangsinaia **
Wu et al., 2020 [Publ 4147], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
17 |
17 |
15 |
2 |
12 |
11 |
2 |
37.0 |
+-------- Ambleminae
--+
| +----- Gonideinae
+--+
| +-- Anodontini **
+--+
+-- Unionini **
Bolotov et al., 2018 [Publ 3826], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
17 |
17 |
16 |
3 |
66 |
66 |
3 |
46.0 |
+----------------- Parreysiinae
|
| +-------------- Pseudodontini
--+ |
| | +-- Rectidentini
| | +--------+
+--+ | +-- Contradentini
| |
| | +----- Unionini **
| | +--+
| | | | +-- Lanceolariini **
+--+ | +--+
| | +-- Anodontini **
| |
| | +-- Lamprotula
| | |
| | +-- Potomida
+--+ |
| +-- Pronodularia
| |
| +-- Gonidea
| |
| +-- Leguminaia
| |
| +-- Microcondylaea
| |
+-----+-- Amblemini
|
+-- Ortmanniana
|
+-- Lampsilis
|
+-- Cambarunio
|
+-- Elliptio
|
+-- Pleurobema
|
+-- Quadrulini
Chase et al., 2018 [Publ 3847], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
16 |
16 |
15 |
2 |
22 |
17 |
1 |
43.0 |
+-- Unionini **
--+
+-- Anodontini **
Dai et al., 2024 [Publ 4386], fig. 5-ML
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
16 |
16 |
14 |
5 |
5 |
3 |
1 |
52.0 |
+-------------- Parreysiinae
|
--+ +----------- Anodontina **
| |
+--+ +-- Acuticostini **
| +-----+
| | +-- Cristariina **
+--+
| +----- Lepidodesmini **
+--+
| +-- Lanceolariini **
+--+
+-- Unionini **
Dai et al., 2024 [Publ 4386], fig. 5-BI
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
16 |
16 |
11 |
5 |
5 |
3 |
1 |
52.0 |
+-------------------- Parreysiinae
|
| +----------------- Acuticostini **
--+ |
| +----------------- Cristariina **
| |
| +----------------- Anodontina **
+--+
| +-------------- Lanceolariini **
| |
| | +-- Aculamprotula **
| +-----------+
+--+ +-- Lepidodesmini **
|
| +----------- Unio **
| |
+--+ +-------- Middendorffinaia **
| |
+--+ +----- Cuneopsis **
| |
+--+----- Nodularia **
|
| +-- Inversiunio **
+--+
+-- Schistodesmus **
Zanatta et al., 2007 [Publ 2698], fig. 1A
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
16 |
16 |
10 |
1 |
0 |
0 |
0 |
55.0 |
-- (Anodontini
Kim et al., 2021 [Publ 4197], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
16 |
17 |
15 |
4 |
1 |
1 |
1 |
38.0 |
+-- Gonideinae
+--------------+
| +-- Lepidodesmini **
--+
| +-------------- Aculamprotula **
| |
+--+ +-- Acuticostini **
| +--------+
| | +-- Lanceolariini **
+--+
| +-------- Unio **
+--+
| +----- Nodularia **
+--+
| +-- Schistodesmus **
+--+
+-- Cuneopsis **
Fukata & Iigo, 2019 [Publ 4108], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
16 |
16 |
14 |
3 |
0 |
0 |
0 |
41.0 |
+----- Aculamprotula **
+--------+
| | +-- Acuticostini **
| +--+
--+ +-- Lanceolariini **
|
| +----------- Unio **
| |
+--+ +-------- Pseudocuneopsis **
| |
+--+ +-- Schistodesmus **
| +--+
| | +-- Cuneopsis **
+--+
| +-- Tchangsinaia **
+--+
+-- Nodularia **
Zanatta et al., 2007 [Publ 2698], fig. 1B
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
16 |
16 |
13 |
1 |
0 |
0 |
0 |
55.0 |
-- (Anodontini
Hoeh, 1990 [Publ 2717], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
15 |
15 |
13 |
1 |
0 |
0 |
0 |
54.0 |
-- (Anodontini
Zhou et al., 2007 [Publ 2658], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
15 |
21 |
19 |
4 |
8 |
5 |
1 |
55.0 |
+----------------- Anodontini **
|
| +----------- Chamberlainiini
| +--+
--+ | | +-------- Lamprotula_leaii
| | +--+
| | | +----- Lamprotula_caveata
| | +--+
| | | +-- Gibbosulinae
+--+ +--+
| +-- Lamprotula_cornuumlunae
|
| +----- Schistodesmus **
| +--+
| | | +-- Lanceolariini **
| | +--+
+-----+ +-- Acuticostini **
|
| +----- Aculamprotula **
+--+
| +-- Tchangsinaia **
+--+
+-- Nodularia **
Hoeh, 1990 [Publ 2717], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
15 |
15 |
2 |
1 |
0 |
0 |
0 |
56.0 |
-- (Anodontini
Chen et al., 2020 [Publ 4170], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
15 |
34 |
32 |
3 |
19 |
18 |
2 |
73.0 |
+----- Prolasmidonta **
+--+
| | +-- Alasmidonta **
| +--+
+-----------------+ +-- Anemina **
| |
| | +----- Pseudodontini
| +--+
| | +-- Lamprotula_caveata
| +--+
| +-- Chamberlainia
|
| +----- Schistodesmus **
| +-----+
| | | +-- Lepidodesmini **
--+ | +--+
| | +-- Aculamprotula **
| +-----------+
| | | +----- Platynaias **
| | | +--+
| | | | | +-- Utterbackia_imbecillis **
| | | | +--+
| | +--+ +-- Pyganodon **
| | |
| | | +----- Sinanodonta **
| | +--+
| | | +-- Cristaria **
+--+ +--+
| +-- Anodontina **
|
| +-- Utterbackia_peninsularis **
| +-----+
| | +-- Potamilus_alatus
| +-----------+
| | | +----- Lamprotula_leaii
| | +--+
| | | +-- Sinohyriopsis
+--+ +--+
| +-- Gonideini
|
| +----------------- Gibbosulinae
+--+
| +-------------- Quadrulini
+--+
| +----------- Toxolasma
+--+
| +-------- Lampsilis_ornata
+--+
| +----- Potamilus_leptodon
+--+
| +-- Lampsilis_siliquoidea
+--+
+-- Lampsilis_powellii
Pavan-Kumar et al., 2022 [Publ 4263], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
15 |
15 |
13 |
5 |
21 |
19 |
3 |
38.0 |
+-------------------- Ambleminae
--+
| +----------------- Gonideinae
+--+
| +-------------- Parreysiinae
+--+
| +----------- Lepidodesmini **
+--+
| +-------- Unionini **
+--+
| +----- Lanceolariini **
+--+
| +-- Acuticostini **
+--+
+-- Anodontini **
Hoeh, 1990 [Publ 2717], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
15 |
15 |
13 |
1 |
0 |
0 |
0 |
54.0 |
-- (Anodontini
Liu et al., 2017 [Publ 3939], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
14 |
15 |
12 |
4 |
1 |
1 |
1 |
47.0 |
+-- Lanceolariini **
+-----+
| +-- Unionini **
--+
| +----- Acuticostini **
+--+
| +-- Anodontini **
+--+
+-- Gonideinae
Lopes-Lima et al., 2024 [Publ 4376], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
14 |
14 |
12 |
1 |
0 |
0 |
0 |
55.0 |
-- (Unio
Huang et al., 2002 [Publ 2657], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
14 |
14 |
12 |
5 |
20 |
18 |
2 |
51.0 |
+----------- Sinosolenaia
+--+
| | +-------- Lamprotulini
| +--+
| | +----- Chamberlainiini
| +--+
| | +-- Ptychorhynchus
--+ +--+
| +-- Ambleminae
|
| +----------- Anodontini **
| |
| | +----- Acuticostini **
+--+ +--+
| | | +-- Nodularia **
| | +--+
+--+ +-- Tchangsinaia **
|
| +----- Lanceolariini **
+--+
| +-- Schistodesmus **
+--+
+-- Lepidodesmini **
Prié & Puillandre, 2014 [Publ 3529], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
14 |
14 |
12 |
2 |
3 |
2 |
2 |
51.0 |
+-------- Ambleminae
--+
| +----- Gonideinae
+--+
| +-- Anodontini **
+--+
+-- Unionini **
Inoue et al., 2014 [Publ 3635], fig. 3A
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
14 |
14 |
13 |
1 |
6 |
5 |
2 |
52.0 |
+-------- Amblemini
|
+-------- Pleurobemini
--+
+-------- Lampsilini
|
| +----- Quadrulini
+--+
| +-- Gonideinae
+--+
+-- Anodontini **
Froufe et al., 2016 [Publ 3738], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
14 |
14 |
12 |
1 |
0 |
0 |
0 |
53.0 |
-- (Unionini
Huang et al., 2002 [Publ 2657], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
14 |
14 |
5 |
5 |
20 |
18 |
2 |
48.0 |
+-- Ptychorhynchus
|
+-- Chamberlainiini
|
+--------+-- Lamprotulini
| |
| +-- Sinosolenaia
| |
| +-- Ambleminae
|
| +-------- Strophitus **
--+ |
| +-------- Schistodesmus **
| |
| +-------- Lepidodesmini **
| |
| +-------- Acuticostini **
| |
| +-------- Cristariina **
+--+
+-------- Lanceolariini **
|
| +-- Nodularia **
+-----+
| +-- Tchangsinaia **
|
| +----- Utterbackia **
| |
+--+----- Utterbackiana **
|
| +-- Alasmidonta **
+--+
+-- Lasmigona **
Guerra et al., 2019 [Publ 4168], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
13 |
13 |
11 |
3 |
28 |
8 |
2 |
38.0 |
+-------------- Ambleminae
|
--+ +----------- Gonideinae
| |
+--+ +-------- Aculamprotula **
| |
+--+ +-- Schistodesmus **
| +--+
| | +-- Unio **
+--+
| +-- Lanceolariini **
+--+
+-- Anodontini **
Sayenko et al., 2021 [Publ 4189], fig. 6
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
13 |
13 |
11 |
1 |
0 |
0 |
0 |
52.0 |
-- (Unionini
Sano et al., 2017 [Publ 3799], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
13 |
13 |
14 |
3 |
8 |
4 |
1 |
52.0 |
+-------- Gonideinae
--+
| +----- Anodontini **
+--+
| +-- Lanceolariini **
+--+
+-- Unionini **
Pfeiffer & Graf, 2013 [Publ 3531], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
13 |
13 |
10 |
3 |
21 |
18 |
3 |
43.0 |
+----- Anodontini **
--+
| +-- Acuticostini **
+--+
+-- Unionini **
Douda et al., 2024 [Publ 4364], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
13 |
13 |
12 |
1 |
1 |
0 |
0 |
53.0 |
-- (Cristariina
Guerra et al., 2019 [Publ 4168], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
13 |
13 |
11 |
3 |
28 |
8 |
2 |
38.0 |
+----------- Ambleminae
--+
| +-------- Gonideinae
+--+
| +----- Anodontini **
+--+
| +-- Lanceolariini **
+--+
+-- Unionini **
Wu et al., 2018 [Publ 4143], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
13 |
13 |
11 |
2 |
9 |
9 |
2 |
39.0 |
+-- Ambleminae
+--+
| +-- Gonideinae
--+
| +-- Unionini **
+--+
+-- Anodontini **
Breton et al., 2011 [Publ 3376], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
12 |
12 |
11 |
1 |
25 |
20 |
2 |
49.0 |
+-------------- Quadrulina
|
| +----- Lamprotulini
--+ +-----+
| | | +-- Chamberlainiini
| | +--+
+--+ +-- Anodontini **
|
| +-------- Megalonaias
+--+
| +----- Pleurobemini
+--+
| +-- Amblemini
+--+
+-- Lampsilini
Dai et al., 2024 [Publ 4386], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
12 |
12 |
10 |
3 |
1 |
1 |
1 |
54.0 |
+----------------- Parreysiinae
|
--+ +-------------- Unio **
| |
| | +----- Aculamprotula **
+--+ +-----+
| | | +-- Anodontini **
| | +--+
+--+ +-- Lepidodesmini **
|
| +-------- Middendorffinaia **
+--+
| +----- Inversiunio **
+--+
| +-- Cuneopsis **
+--+
+-- Nodularia **
Choi et al., 2020 [Publ 4169], fig. 2a
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
12 |
12 |
11 |
1 |
0 |
0 |
0 |
51.0 |
-- (Unionini
Froufe et al., 2017 [Publ 3802], fig. 6
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
12 |
12 |
9 |
1 |
0 |
0 |
0 |
48.0 |
-- (Anodontini
Bogan et al., 2023 [Publ 4295], fig. 6
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
12 |
12 |
9 |
1 |
1 |
0 |
0 |
52.0 |
-- (Cristariina
Lopes-Lima et al., 2020 [Publ 4080], fig. 10
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
12 |
12 |
8 |
1 |
2 |
0 |
0 |
52.0 |
-- (Cristariina
Xue et al., 2016 [Publ 3659], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
12 |
12 |
10 |
3 |
4 |
4 |
2 |
44.0 |
+-- Gonideinae
+-----+
| +-- Ambleminae
--+
| +----- Anodontini **
+--+
| +-- Lanceolariini **
+--+
+-- Unionini **
Sano et al., 2022 [Publ 4311], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
12 |
12 |
12 |
1 |
0 |
0 |
0 |
53.0 |
-- (Cristariina
Wu et al., 2016 [Publ 3711], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
12 |
12 |
10 |
2 |
9 |
9 |
2 |
40.0 |
+-- Ambleminae
+--+
| +-- Gonideinae
--+
| +-- Unionini **
+--+
+-- Anodontini **
Wu et al., 2024 [Publ 4379], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
12 |
12 |
10 |
1 |
0 |
0 |
0 |
54.0 |
-- (Unionini
Zhang et al., 2004 [Publ 2735], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
12 |
12 |
10 |
4 |
2 |
2 |
1 |
46.0 |
+----------- Gonideinae
--+
| +-------- Anodontini **
+--+
| +----- Unionini **
+--+
| +-- Acuticostini **
+--+
+-- Lanceolariini **
Lopes-Lima et al., 2017 [Publ 3734], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
12 |
12 |
8 |
2 |
5 |
2 |
1 |
47.0 |
+----- Gonideinae
--+
| +-- Anodontini **
+--+
+-- Unionini **
Wu et al., 2023 [Publ 4339], fig. 5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
11 |
11 |
9 |
2 |
8 |
6 |
1 |
42.0 |
+----- Ambleminae
--+
| +-- Lepidodesmini **
+--+
+-- Anodontini **
Burzynski & Soroka, 2018 [Publ 3862], fig. 5a
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
10 |
10 |
7 |
3 |
10 |
8 |
2 |
40.0 |
+----------- Ambleminae
--+
| +-------- Gonideinae
+--+
| +----- Anodontini **
+--+
| +-- Lanceolariini **
+--+
+-- Unionini **
Lopes-Lima et al., 2021 [Publ 4217], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
10 |
10 |
8 |
1 |
0 |
0 |
0 |
51.0 |
-- (Unio
Klishko et al., 2019 [Publ 3886], fig. 9
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
10 |
10 |
7 |
2 |
1 |
0 |
0 |
45.0 |
+-- Anodontini **
--+
+-- Unionini **
Zhou et al., 2007 [Publ 2658], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
10 |
21 |
19 |
3 |
11 |
10 |
2 |
59.0 |
+-------------------- Sinanodonta **
|
--+ +----------------- Cristaria **
| |
| | +-------- Aculamprotula **
+--+ +-----+
| | | +----- Acuticostini **
| | +--+
| | | +-- Nodularia **
+--+ +--+
| +-- Cuneopsis **
|
| +----------- Ambleminae
+--+
| +-------- Lamprotula_leaii
+--+
| +----- Lamprotula_cornuumlunae
+--+
| +-- Gibbosulinae
+--+
+-- Lamprotula_caveata
Lopes-Lima et al., 2020 [Publ 4080], fig. 6
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
10 |
10 |
8 |
1 |
2 |
0 |
0 |
51.0 |
-- (Cristariina
Guerra et al., 2017 [Publ 3772], fig. 2a
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
10 |
10 |
8 |
2 |
15 |
9 |
2 |
41.0 |
+-- Ambleminae
+--+
| +-- Gonideinae
--+
| +-- Unionini **
+--+
+-- Anodontini **
Huang et al., 2019 [Publ 3843], fig. 3-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
10 |
10 |
8 |
3 |
13 |
10 |
2 |
41.0 |
+----------- Ambleminae
--+
| +-------- Gonideinae
+--+
| +----- Anodontini **
+--+
| +-- Lanceolariini **
+--+
+-- Unionini **
Guerra et al., 2019 [Publ 4168], fig. 1-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
10 |
10 |
8 |
3 |
12 |
7 |
2 |
41.0 |
+----------- Ambleminae
--+
| +-------- Gonideinae
+--+
| +----- Anodontini **
+--+
| +-- Lanceolariini **
+--+
+-- Unionini **
Krebs et al., 2003 [Publ 2691], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
10 |
10 |
5 |
1 |
23 |
21 |
1 |
52.5 |
+-- Anodontini **
--+
+-- Ambleminae
Pfarrer et al., 2022 [Publ 4305], fig. 2-5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
10 |
10 |
8 |
2 |
2 |
2 |
1 |
37.0 |
+----- Gonideinae
--+
| +-- Unionini **
+--+
+-- Anodontini **
Guerra et al., 2017 [Publ 3772], fig. 2b
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
10 |
10 |
8 |
2 |
15 |
9 |
2 |
41.0 |
+----------- Ambleminae
--+
| +-------- Gonideinae
+--+
| +----- Unio **
+--+
| +-- Aculamprotula **
+--+
+-- Anodontini **
Lopes-Lima et al., 2020 [Publ 4080], fig. 11
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
10 |
10 |
8 |
2 |
2 |
0 |
0 |
49.0 |
+-- Unionini **
--+
+-- Lanceolariini **
Reis et al., 2013 [Publ 3580], fig. 5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
10 |
10 |
8 |
2 |
5 |
3 |
2 |
44.0 |
+-------- Ambleminae
--+
| +----- Gonideinae
+--+
| +-- Anodontini **
+--+
+-- Unionini **
Burzynski & Soroka, 2018 [Publ 3862], fig. 5b
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
10 |
10 |
8 |
3 |
0 |
0 |
0 |
39.0 |
+----- Lanceolariini **
--+
| +-- Anodontini **
+--+
+-- Unionini **
Guerra et al., 2019 [Publ 4168], fig. 2-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
10 |
10 |
8 |
3 |
12 |
7 |
2 |
41.0 |
+----------- Ambleminae
--+
| +-------- Gonideinae
+--+
| +----- Anodontini **
+--+
| +-- Lanceolariini **
+--+
+-- Unionini **
Zhou et al., 2016 [Publ 3710], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
9 |
9 |
7 |
3 |
11 |
10 |
2 |
42.0 |
+----- Anodontini **
--+
| +-- Lepidodesmini **
+--+
+-- Unionini **
Boyer et al., 2011 [Publ 2718], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
9 |
9 |
6 |
1 |
29 |
29 |
1 |
52.0 |
+-- Anodontini **
--+
+-- Ambleminae
Reis et al., 2013 [Publ 3580], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
9 |
9 |
7 |
2 |
5 |
3 |
2 |
46.0 |
+-------- Ambleminae
--+
| +----- Gonideinae
+--+
| +-- Anodontini **
+--+
+-- Unionini **
Jadhav & Jamkhedkar, 2009 [Publ 3697], fig. 11
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
9 |
9 |
7 |
2 |
16 |
16 |
3 |
54.0 |
+-------------------------------- Parreysiinae
|
| +-------------- Nodularia **
| |
| +--------------+ +----------- Anodonta_cygnea **
--+ | | |
| | +--+ +-------- Unio_pictorum__2 **
| | | |
| | +--+ +-- Unio_pictorum__1 **
| | | +--+
| | | | +-- Unio_caffer **
+--+ +--+
| | +-- Anodonta **
| +--+
| +-- Alasmidontina **
|
| +-------------------------- Lamprotulini
| |
+--+ +----------------------- Quadrulini
| |
+--+ +-------------------- Gonideini
| |
| | +----- Eurynia
+--+ +-----------+
| | | +-- Cambarunio
| | +--+
| | +-- Ptychobranchus
+--+
| +-- Amblemini
| +-----------+
| | +-- Pleurobema
+--+
| +----------- Truncilla
| |
+--+ +-- Lampsilis
| +-----+
| | +-- Epioblasma
+--+
| +----- Ligumia
+--+
| +-- Obliquaria
+--+
+-- Ortmanniana
Soroka, 2010 [Publ 3404], fig. 3a
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
9 |
9 |
7 |
2 |
0 |
0 |
0 |
46.0 |
+----- Cristariina **
--+
| +-- Anodontina **
+--+
+-- Unionini **
Ouyang et al., 2011 [Publ 3408], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
9 |
9 |
6 |
4 |
4 |
4 |
1 |
49.0 |
+-------------- Gonideinae
|
--+ +----------- Anodontini **
| |
+--+ +-- Nodularia **
| +-----+
| | +-- Cuneopsis **
+--+
| +----- Lanceolaria_grayii **
| |
+--+----- Acuticostini **
|
| +-- Lanceolaria_gladiola **
+--+
+-- Aculamprotula **
King et al., 1999 [Publ 2699], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
9 |
9 |
7 |
1 |
0 |
0 |
0 |
50.0 |
-- (Alasmidontina
Khalloufi et al., 2011 [Publ 2748], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
9 |
9 |
6 |
1 |
2 |
2 |
2 |
48.0 |
+----- Ambleminae
--+
| +-- Gonideinae
+--+
+-- Unionini **
Plazzi et al., 2011 [Publ 3676], fig. 5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
9 |
18 |
11 |
3 |
9 |
8 |
2 |
55.0 |
+----------- Cristariina **
|
| +-------- Lanceolariini **
| |
--+ +-------- Anodonta **
| |
| +-------- Unionini **
| |
+--+ +-- Alasmidontina **
+-----+
| +-- Pseudanodonta **
|
| +----- Ambleminae
| |
+--+ +-- Velesunioninae
| |
+--+-- Lamprotulini
|
+-- Chamberlainiini
Soroka, 2010 [Publ 3404], fig. 3b
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
9 |
9 |
5 |
2 |
0 |
0 |
0 |
46.0 |
+----- Anodonta_anatina **
|
+----- Anodonta_cygnea **
--+
+----- Pseudanodonta **
|
| +-- Cristariina **
+--+
+-- Unionini **
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2A
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
8 |
25 |
19 |
2 |
22 |
17 |
1 |
64.0 |
+----- Unionini **
--+
| +-- Anodontini **
+--+
+-- Ambleminae
Kondakov et al., 2018 [Publ 3914], fig. 5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
8 |
8 |
6 |
1 |
0 |
0 |
0 |
49.0 |
-- (Cristariina
Araujo et al., 2009 [Publ 2589], fig. 7A
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
8 |
8 |
4 |
2 |
3 |
2 |
2 |
51.0 |
+----------------- Ambleminae
|
--+ +-------------- Gonideinae
| |
+--+ +----------- Nodularia **
| |
| | +-- Unio_gibbus **
+--+ |
| +-----+-- Unio_tumidus **
| | |
| | +-- Anodontini **
+--+
| +----- Unio_pictorum__1 **
| |
| +----- Unio_pictorum__2 **
+--+
+----- Unio_mancus **
|
| +-- Unio_tumidiformis **
+--+
+-- Unio_crassus **
Marrone et al., 2019 [Publ 3907], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
8 |
8 |
4 |
1 |
0 |
0 |
0 |
48.0 |
-- (Unio
Feng et al., 2016 [Publ 3800], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
8 |
13 |
11 |
2 |
7 |
5 |
2 |
50.0 |
+-- Anodontina **
+--------+
| +-- Utterbackia **
|
--+ +-- Gonideinae
| +-----+
| | +-- Ambleminae
+--+
| +----- Unionini **
+--+
| +-- Pyganodon **
+--+
+-- Cristariina **
Mabuchi & Nishida, 2020 [Publ 4120], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
8 |
8 |
8 |
3 |
4 |
4 |
1 |
50.0 |
+----------- Gonideinae
--+
| +-------- Anodontini **
+--+
| +----- Nodularia **
+--+
| +-- Lanceolariini **
+--+
+-- Inversiunio **
Bauer et al., 2021 [Publ 4365], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
8 |
8 |
5 |
1 |
0 |
0 |
0 |
49.0 |
-- (Cristariina
Araujo et al., 2009 [Publ 2589], fig. 7B
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
8 |
8 |
7 |
2 |
3 |
2 |
2 |
48.0 |
+-------- Gonideinae
--+
| +----- Ambleminae
+--+
| +-- Anodontini **
+--+
+-- Unionini **
Bespalaya et al., 2018 [Publ 3811], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
8 |
8 |
7 |
1 |
2 |
0 |
0 |
50.0 |
-- (Cristariina
Soroka, 2010 [Publ 3404], fig. 5a
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
8 |
8 |
6 |
2 |
0 |
0 |
0 |
47.0 |
+-- Unionini **
--+
+-- Anodontini **
Manendo et al., 2008 [Publ 2669], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
8 |
8 |
6 |
2 |
5 |
5 |
1 |
44.0 |
+----- Ambleminae
--+
| +-- Unionini **
+--+
+-- Anodontini **
Vikhrev et al., 2017 [Publ 3810], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
8 |
8 |
6 |
1 |
2 |
0 |
0 |
49.0 |
-- (Cristariina
Tomilova et al., 2020 [Publ 4097], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
8 |
8 |
5 |
2 |
0 |
0 |
0 |
44.0 |
+-- Unionini **
|
--+-- Cristariina **
|
+-- Anodontina **
Pfeiffer et al., 2019 [Publ 3937], fig. 4-5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
8 |
8 |
6 |
3 |
49 |
42 |
3 |
43.0 |
+-------------- Ambleminae
--+
| +----------- Gonideinae
+--+
| +-------- Parreysiinae
+--+
| +----- Lanceolariini **
+--+
| +-- Unionini **
+--+
+-- Anodontini **
Klishko et al., 2018 [Publ 3945], fig. 8
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
8 |
8 |
5 |
1 |
0 |
0 |
0 |
45.0 |
-- (Unionini
Lopes-Lima et al., 2020 [Publ 4080], fig. 13
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
8 |
8 |
6 |
1 |
2 |
0 |
0 |
51.0 |
-- (Unionini
Graf & Ó Foighil, 2000 [Publ 2648], fig. 2B
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
8 |
25 |
23 |
2 |
22 |
17 |
1 |
64.0 |
+----- Unionini **
--+
| +-- Anodontini **
+--+
+-- Ambleminae
Pfeiffer et al., 2019 [Publ 3937], fig. 6
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
8 |
8 |
6 |
3 |
49 |
42 |
3 |
43.0 |
+----------- Ambleminae
|
--+----------- Gonideinae
|
| +-------- Parreysiinae
+--+
| +----- Anodontini **
+--+
| +-- Lanceolariini **
+--+
+-- Unionini **
Soroka, 2010 [Publ 3404], fig. 5b
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
8 |
8 |
5 |
2 |
0 |
0 |
0 |
46.0 |
+-------- Anodonta_anatina **
|
--+ +----- Cristariina **
| |
+--+----- Unionini **
|
| +-- Anodonta_cygnea **
+--+
+-- Pseudanodonta **
Huang et al., 2018 [Publ 3828], fig. 6-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
8 |
8 |
6 |
2 |
9 |
7 |
2 |
41.0 |
+-------- Ambleminae
--+
| +----- Gonideinae
+--+
| +-- Anodontini **
+--+
+-- Unionini **
Smith et al., 2023 [Publ 4291], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
7 |
7 |
5 |
1 |
0 |
0 |
0 |
53.0 |
-- (Alasmidontina
Araujo et al., 2009 [Publ 2589], fig. 7C
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
7 |
7 |
5 |
2 |
3 |
2 |
2 |
47.0 |
+-------- Gonideinae
--+
| +----- Ambleminae
+--+
| +-- Anodontini **
+--+
+-- Unionini **
Zieritz et al., 2021 [Publ 4231], fig. 3-M_4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
7 |
7 |
5 |
2 |
16 |
15 |
2 |
40.0 |
+-------- Ambleminae
--+
| +----- Gonideinae
+--+
| +-- Unionini **
+--+
+-- Anodontini **
Soroka, 2010 [Publ 3404], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
7 |
7 |
4 |
2 |
0 |
0 |
0 |
45.0 |
+----- Cristariina **
--+
| +-- Unionini **
+--+
+-- Anodontina **
Lopes-Lima et al., 2020 [Publ 4080], fig. 5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
7 |
7 |
5 |
1 |
2 |
0 |
0 |
46.0 |
-- (Unionini
Nagel et al., 1998 [Publ 1946], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
7 |
7 |
5 |
2 |
2 |
2 |
1 |
42.0 |
+-------- Gonideini
--+
| +----- Lamprotulini
+--+
| +-- Anodontini **
+--+
+-- Unionini **
Hoeh et al., 1995 [Publ 3391], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
7 |
7 |
4 |
1 |
0 |
0 |
0 |
48.0 |
-- (Alasmidontina
Zieritz et al., 2012 [Publ 3673], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
7 |
8 |
4 |
2 |
1 |
0 |
0 |
46.0 |
+----- Anodontina **
|
--+ +-- Cristariina **
| |
+--+-- Margaritiferinae
|
+-- Unionini **
Wen et al., 2017 [Publ 3795], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
7 |
7 |
5 |
3 |
9 |
7 |
2 |
44.0 |
+-------------- Ambleminae
--+
| +----------- Gonideinae
+--+
| +-------- Anodontini **
+--+
| +----- Aculamprotula **
+--+
| +-- Lepidodesmini **
+--+
+-- Unio **
Lopes-Lima et al., 2021 [Publ 4217], fig. 6
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
7 |
7 |
5 |
1 |
0 |
0 |
0 |
48.0 |
-- (Unionini
Inoue et al., 2014 [Publ 3635], fig. 3B
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
7 |
7 |
2 |
1 |
1 |
1 |
1 |
51.0 |
+-- Ambleminae
--+
+-- Anodontini **
Froufe et al., 2019 [Publ 3936], fig. 3-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
7 |
7 |
5 |
2 |
12 |
11 |
2 |
40.0 |
+-------- Ambleminae
--+
| +----- Gonideinae
+--+
| +-- Unionini **
+--+
+-- Anodontini **
Zhou et al., 2017 [Publ 3788], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
7 |
17 |
15 |
3 |
10 |
10 |
2 |
53.0 |
+-------- Anodontini **
|
--+ +-- Lanceolariini **
| +--+
| | +-- Unionini **
+--+
| +-- Ambleminae
+--+
+-- Gonideinae
Froufe et al., 2017 [Publ 3802], fig. 7
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
7 |
7 |
5 |
2 |
3 |
2 |
1 |
42.0 |
+----- Gonideinae
--+
| +-- Unionini **
+--+
+-- Anodontini **
Huang et al., 2013 [Publ 3629], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
7 |
7 |
5 |
2 |
9 |
8 |
2 |
40.0 |
+-- Unionini **
+--+
| +-- Anodontini **
--+
| +-- Ambleminae
+--+
+-- Gonideinae
Froufe et al., 2019 [Publ 3936], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
7 |
7 |
5 |
2 |
12 |
11 |
2 |
40.0 |
+-------- Ambleminae
--+
| +----- Gonideinae
+--+
| +-- Unionini **
+--+
+-- Anodontini **
Choi et al., 2020 [Publ 4169], fig. 5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
7 |
7 |
5 |
1 |
3 |
3 |
1 |
50.0 |
+-- Parreysiinae
--+
+-- Unionini **
Soroka, 2010 [Publ 3404], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
7 |
7 |
5 |
2 |
0 |
0 |
0 |
46.0 |
+-- Anodonta_anatina **
+-----+
| +-- Unionini **
--+
| +----- Cristariina **
+--+
| +-- Anodonta_cygnea **
+--+
+-- Pseudanodonta **
Hewitt et al., 2018 [Publ 3758], fig. 5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
6 |
6 |
3 |
1 |
3 |
2 |
1 |
49.0 |
+-- Ambleminae
--+
+-- Anodontini **
Hoeh et al., 2009 [Publ 2716], fig. 5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
6 |
6 |
4 |
2 |
40 |
18 |
3 |
49.0 |
+----------- Pseudodontini
|
--+ +-------- Contradentini
| |
+--+ +-- Unionini **
| +--+
| | +-- Anodontini **
+--+
| +-- Gonideini
+--+
+-- Ambleminae
Bolotov et al., 2016 [Publ 3732], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
6 |
6 |
5 |
1 |
2 |
0 |
0 |
50.0 |
-- (Sinanodonta
Campbell et al., 2005 [Publ 2660], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
6 |
6 |
3 |
1 |
96 |
94 |
2 |
49.0 |
+-- Chamberlainiini
+-----+
| +-- Anodontini **
--+
| +----- Ambleminae
+--+
| +-- Gonideini
+--+
+-- Lamprotulini
Klishko et al., 2017 [Publ 3946], fig. 8
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
6 |
6 |
4 |
1 |
0 |
0 |
0 |
45.0 |
-- (Unionini
King et al., 1999 [Publ 2699], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
6 |
6 |
3 |
1 |
0 |
0 |
0 |
49.0 |
-- (Alasmidontina
Bogan et al., 2023 [Publ 4295], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
6 |
6 |
4 |
1 |
1 |
0 |
0 |
50.0 |
-- (Cristariina
Källersjö et al., 2005 [Publ 2684], fig. 1A
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
6 |
7 |
2 |
2 |
1 |
0 |
0 |
45.0 |
+-- Margaritiferinae
|
--+-- Anodontini **
|
+-- Unionini **
Soroka, 2010 [Publ 3404], fig. 3a-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
6 |
6 |
4 |
2 |
0 |
0 |
0 |
47.0 |
+----- Anodontina **
--+
| +-- Cristariina **
+--+
+-- Unionini **
Hoeh et al., 2009 [Publ 2716], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
6 |
6 |
4 |
2 |
40 |
18 |
3 |
49.0 |
+----------- Pseudodontini
|
--+ +-------- Contradentini
| |
+--+ +-- Unionini **
| +--+
| | +-- Anodontini **
+--+
| +-- Gonideini
+--+
+-- Ambleminae
Lydeard et al., 1996 [Publ 2656], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
6 |
6 |
4 |
1 |
22 |
18 |
1 |
49.5 |
+-- Anodontini **
--+
+-- Ambleminae
Gomes-dos-Santos et al., 2019 [Publ 4042], fig. 8
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
6 |
6 |
4 |
1 |
0 |
0 |
0 |
44.0 |
-- (Anodontini
Burzynski et al., 2017 [Publ 3774], fig. 1-top
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
6 |
6 |
4 |
2 |
7 |
7 |
2 |
41.0 |
+-- Unionini **
+--+
| +-- Anodontini **
--+
| +-- Ambleminae
+--+
+-- Gonideinae
Graf & Cummings, 2006 [Publ 1968], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
6 |
6 |
4 |
2 |
41 |
19 |
3 |
49.0 |
+-------- Pseudodontini
|
| +----- Parreysiinae
--+ |
| +----- Contradentini
| |
+--+ +-- Unionini **
+--+
| +-- Anodontini **
|
| +-- Gonideini
+--+
+-- Ambleminae
Lopes-Lima et al., 2020 [Publ 4080], fig. 12
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
6 |
6 |
4 |
1 |
2 |
0 |
0 |
51.0 |
-- (Unionini
Soroka, 2010 [Publ 3404], fig. 8b
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
6 |
6 |
4 |
2 |
0 |
0 |
0 |
43.0 |
+----- Cristariina **
--+
| +-- Anodontina **
+--+
+-- Unionini **
Källersjö et al., 2005 [Publ 2684], fig. 1B-F
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
6 |
7 |
4 |
2 |
1 |
0 |
0 |
46.0 |
+-- Margaritiferinae
|
--+-- Anodontini **
|
+-- Unionini **
Soroka, 2010 [Publ 3404], fig. 3b-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
6 |
6 |
4 |
2 |
0 |
0 |
0 |
45.0 |
+----- Anodontina **
--+
| +-- Cristariina **
+--+
+-- Unionini **
Boyer et al., 2011 [Publ 2718], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
6 |
6 |
2 |
1 |
29 |
29 |
1 |
48.0 |
+-- Anodontini **
--+
+-- Ambleminae
Burzynski et al., 2017 [Publ 3774], fig. 1-bottom
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
6 |
6 |
4 |
2 |
7 |
7 |
2 |
41.0 |
+-- Unionini **
+--+
| +-- Anodontini **
--+
| +-- Ambleminae
+--+
+-- Gonideinae
Riccardi et al., 2020 [Publ 4089], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
6 |
6 |
4 |
1 |
0 |
0 |
0 |
43.0 |
-- (Anodontini
Campbell et al., 2005 [Publ 2660], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
6 |
6 |
4 |
1 |
96 |
94 |
2 |
48.0 |
+-------- Chamberlainiini
|
--+-------- Anodontini **
|
| +----- Ambleminae
+--+
| +-- Gonideini
+--+
+-- Lamprotulini
Dai et al., 2024 [Publ 4358], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
6 |
6 |
4 |
1 |
0 |
0 |
0 |
48.0 |
-- (Unionini
Hoeh et al., 2009 [Publ 2716], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
6 |
6 |
4 |
2 |
41 |
19 |
3 |
49.0 |
+-- Unionini **
+--------+
| +-- Anodontini **
--+
| +-------- Contradentini
+--+
| +----- Ambleminae
+--+
| +-- Gonideini
+--+
+-- Pseudodontini
Lydeard et al., 1996 [Publ 2656], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
6 |
6 |
4 |
1 |
21 |
18 |
1 |
49.0 |
+-- Anodontini **
--+
+-- Ambleminae
Choi et al., 2020 [Publ 4169], fig. 3a
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
6 |
6 |
5 |
1 |
0 |
0 |
0 |
48.0 |
-- (Unionini
Rosenberg et al., 1997 [Publ 2720], fig. 2B
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
6 |
18 |
1 |
2 |
16 |
12 |
2 |
59.5 |
+----- Anodontina **
|
| +-- Gonideinae
| |
| +-- Pyganodon_grandis **
| |
| +-- Utterbackia **
| |
| +-- Pyganodon_cataracta **
--+ |
| +-- Lampsilis_straminea
| |
| +-- Lampsilis_teres
| |
| +-- Megalonaias
| |
| +-- Obliquaria
| |
+--+-- Theliderma
|
+-- Quadrula
|
+-- Uniomerus
|
+-- Amblema
|
+-- Elliptio
|
+-- Fusconaia
|
+-- Plectomerus
|
+-- Aculamprotula **
|
+-- Unio **
Soroka, 2010 [Publ 3404], fig. 8a
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
6 |
6 |
4 |
2 |
0 |
0 |
0 |
41.0 |
+-- Unionini **
--+
+-- Anodontini **
Lopes-Lima et al., 2020 [Publ 4080], fig. 7
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
6 |
6 |
4 |
1 |
2 |
0 |
0 |
47.0 |
-- (Cristariina
Kano et al., 2019 [Publ 4194], fig. 6
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
6 |
9 |
7 |
3 |
4 |
3 |
1 |
52.0 |
+-------- Anodontini **
--+
| +----- Gonideinae
+--+
| +-- Lanceolariini **
+--+
+-- Unionini **
Roe & Hoeh, 2003 [Publ 2655], fig. 4f3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
3 |
2 |
26 |
13 |
3 |
44.0 |
+-------- Parreysiinae
|
--+ +-- Unionini **
| +--+
| | +-- Anodontini **
+--+
| +-- Gonideinae
+--+
+-- Ambleminae
Chase et al., 2018 [Publ 3861], fig. 1-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
3 |
2 |
6 |
4 |
2 |
42.0 |
+-- Gonideinae
+--+
| +-- Ambleminae
--+
| +-- Unionini **
+--+
+-- Anodontini **
Bogan & Hoeh, 2000 [Publ 2650], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
19 |
12 |
2 |
29 |
13 |
3 |
58.0 |
+-------- Anodontini **
|
--+ +----- Gonideinae
| |
+--+----- Unionini **
|
| +-- Margaritiferinae
+--+
+-- Ambleminae
Cyr et al., 2007 [Publ 2745], fig. 2B
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
4 |
1 |
0 |
0 |
0 |
48.0 |
-- (Anodontini
Dyduch-Falniowska & Koziol, 1989 [Publ 3814], fig. 5_7
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
3 |
2 |
0 |
0 |
0 |
44.0 |
+-- Anodontini **
--+
+-- Unionini **
Wu et al., 2020 [Publ 4147], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
3 |
1 |
9 |
6 |
2 |
44.0 |
+----- Ambleminae
--+
| +-- Gonideinae
+--+
+-- Unionini **
Chase et al., 2018 [Publ 3861], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
3 |
2 |
6 |
4 |
2 |
42.0 |
+-------- Ambleminae
--+
| +----- Gonideinae
+--+
| +-- Unionini **
+--+
+-- Anodontini **
Prie et al., 2012 [Publ 3487], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
3 |
1 |
1 |
0 |
0 |
46.0 |
-- (Unio
Roe & Hoeh, 2003 [Publ 2655], fig. 4f2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
3 |
2 |
26 |
13 |
3 |
44.0 |
+----------------------------- Gonideinae
|
--+ +-------------------------- Parreysiinae
| |
+--+ +----------------------- Fusconaia
| |
| +----------------------- Amblemini
+--+
| +-------------------- Quadrulini
| |
+--+ +----------------- Pleurobema
| |
| | +-------------- Eurynia
+--+ |
| | +----- Cyrtonaias
| +--------+
+--+ | +-- Unionini **
| +--+
| +-- Anodontini **
|
| +----------- Ortmanniana
+--+
| +-------- Glebula
+--+
| +----- Potamilus
+--+
| +-- Ligumia
+--+
+-- Toxolasma
Soroka & Burzynski, 2018 [Publ 3762], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
3 |
2 |
2 |
2 |
1 |
42.0 |
+----- Ambleminae
--+
| +-- Unionini **
+--+
+-- Anodontini **
Nagel et al., 1998 [Publ 1946], fig. 7
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
4 |
1 |
0 |
0 |
0 |
48.0 |
-- (Anodontini
Lopes-Lima et al., 2020 [Publ 4080], fig. 8
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
3 |
1 |
2 |
0 |
0 |
46.0 |
-- (Cristariina
Wu et al., 2020 [Publ 4121], fig. S1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
3 |
1 |
29 |
6 |
3 |
44.0 |
+-- Ambleminae
+--+
| +-- Parreysiinae
--+
| +-- Gonideinae
+--+
+-- Unionini **
Bashê, 2022 [Publ 4265], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
4 |
1 |
1 |
0 |
0 |
49.0 |
-- (Unio
Rosenberg et al., 1997 [Publ 2720], fig. 2A
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
18 |
1 |
2 |
17 |
13 |
2 |
60.0 |
+----- Anodontina **
|
| +-- Gonideinae
| |
| +-- Pyganodon_grandis **
| |
| +-- Utterbackia **
| |
| +-- Pyganodon_cataracta **
--+ |
| +-- Lampsilis_straminea
| |
| +-- Lampsilis_teres
| |
| +-- Megalonaias
| |
| +-- Obliquaria
| |
+--+-- Theliderma
|
+-- Quadrula
|
+-- Uniomerus
|
+-- Amblema
|
+-- Elliptio
|
+-- Fusconaia
|
+-- Plectomerus
|
+-- Pleurobema
|
+-- Unionini **
Hoeh et al., 2002 [Publ 2653], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
3 |
2 |
26 |
13 |
3 |
46.0 |
+----------- Ambleminae
--+
| +-------- Gonideinae
+--+
| +----- Unio_pictorum **
+--+
| +-- Unio_tumidus **
+--+
+-- Anodontini **
Bogatov et al., 2018 [Publ 3859], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
3 |
2 |
1 |
0 |
0 |
46.0 |
+-------- Unio_tumidus **
--+
| +----- Anodontini **
+--+
| +-- Unio_crassus **
+--+
+-- Unio_pictorum **
Nagel et al., 1996 [Publ 3400], fig. 351
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
4 |
1 |
0 |
0 |
0 |
48.0 |
-- (Anodontini
Soroka, 2010 [Publ 3404], fig. 2-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
3 |
2 |
0 |
0 |
0 |
46.0 |
+----- Cristariina **
--+
| +-- Anodontina **
+--+
+-- Unionini **
Soroka & Burzynski, 2018 [Publ 3762], fig. 3-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
3 |
2 |
2 |
2 |
1 |
42.0 |
+----- Ambleminae
--+
| +-- Unionini **
+--+
+-- Anodontini **
Soroka, 2010 [Publ 3404], fig. 9a
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
3 |
1 |
0 |
0 |
0 |
49.0 |
-- (Cristariina
Mabuchi et al., 2021 [Publ 4233], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
3 |
1 |
0 |
0 |
0 |
46.0 |
-- (Unionini
Amaro et al., 2017 [Publ 3791], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
3 |
1 |
11 |
8 |
2 |
43.0 |
+-------- Anodontini **
|
--+ +-- Pleurobemini
| +--+
| | +-- Gonideinae
+--+
| +-- Lampsilini
+--+
+-- Quadrulini
Cyr et al., 2007 [Publ 2745], fig. 2C
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
4 |
1 |
0 |
0 |
0 |
48.0 |
-- (Anodontini
Hoeh et al., 2001 [Publ 2652], fig. 14f2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
15 |
10 |
2 |
26 |
13 |
3 |
54.0 |
+-------------------- Amblemini
|
+-------------------- Parreysiinae
|
+-------------------- Eurynia
|
+-------------------- Fusconaia
|
+-------------------- Gonideinae
|
+-------------------- Pleurobema
|
+-------------------- Quadrulini
--+
| +----------- Ortmanniana
+--------+
| | +-------- Glebula
| +--+
| | +----- Potamilus
| +--+
| | +-- Ligumia
| +--+
| +-- Toxolasma
|
| +----------------- Cyrtonaias
| |
+--+ +-------------- Unionini **
| |
| | +----------- Anodontini **
+--+ |
| | +----- Iridininae
| | |
+--+ +--+ +-- Anodontites_guanarensis
| | | |
| | +--+-- Anodontites_trigona
| | |
+--+ +-- Monocondylaeinae
|
| +----- Rhipidodontini
| |
| +----- Echyridella
+--+
+----- Lortiella
|
| +-- Castaliini
+--+
+-- Velesunio
Wu et al., 2020 [Publ 4121], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
3 |
1 |
29 |
6 |
3 |
44.0 |
+-- Ambleminae
+--+
| +-- Parreysiinae
--+
| +-- Gonideinae
+--+
+-- Unionini **
Bogatov et al., 2018 [Publ 3859], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
3 |
2 |
1 |
0 |
0 |
46.0 |
+-------- Unio_crassus **
--+
| +----- Unio_pictorum **
+--+
| +-- Unio_tumidus **
+--+
+-- Anodontini **
Bogan et al., 2023 [Publ 4295], fig. 5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
3 |
1 |
1 |
0 |
0 |
50.0 |
-- (Cristariina
Hoeh et al., 2001 [Publ 2652], fig. 14f4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
5 |
5 |
2 |
2 |
26 |
13 |
3 |
45.0 |
+-------- Gonideinae
|
--+ +----- Ambleminae
| |
+--+ +-- Unio_pictorum **
| |
+--+-- Unio_tumidus **
|
+-- Anodontini **
Soroka, 2010 [Publ 3404], fig. 9b
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
1 |
0 |
0 |
0 |
49.0 |
-- (Cristariina
Rosenberg et al., 1994 [Publ 2719], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
15 |
2 |
2 |
15 |
12 |
2 |
56.0 |
+-------- Gonideinae
|
| +-- Elliptio
| |
| +-- Fusconaia
| |
--+ +--+-- Amblemini
| | |
| | +-- Pleurobema
| | |
| | +-- Unionini **
| |
| | +-- Pyganodon_cataracta **
+--+ |
| +-- Pyganodon_grandis **
| |
| +-- Utterbackia **
| |
| +-- Lampsilis_teres
| |
| +-- Lampsilis_straminea
+--+
+-- Megalonaias
|
+-- Obliquaria
|
+-- Theliderma
|
+-- Quadrula
|
+-- Uniomerus
Cyr et al., 2007 [Publ 2745], fig. 2D
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
1 |
0 |
0 |
0 |
49.0 |
-- (Pyganodon
Burzynski & Soroka, 2018 [Publ 3862], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
1 |
0 |
0 |
0 |
43.0 |
-- (Anodontini
Sonowal et al., 2021 [Publ 4221], fig. 3a
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
0 |
2 |
11 |
5 |
3 |
46.0 |
+----- Pleurobemini
|
+----- Lampsilini
--+
| +-- Anodontini **
| |
| +-- Unio_ravoisieri **
+--+
+-- Unio_pictorum **
|
+-- Aculamprotula **
Mezhzherin et al., 2014 [Publ 3705], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
1 |
0 |
0 |
0 |
43.0 |
-- (Anodontini
Amaro et al., 2017 [Publ 3791], fig. 3A
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
1 |
8 |
5 |
2 |
43.0 |
+----- Anodontini **
--+
| +-- Gonideinae
+--+
+-- Ambleminae
Sonowal et al., 2021 [Publ 4221], fig. 3b
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
9 |
0 |
2 |
11 |
5 |
3 |
51.0 |
+-- Aculamprotula **
|
+-- Lamprotula
|
+-- Lampsilini
|
+-- Potomida
|
--+-- Unio_pictorum **
|
+-- Anodontini **
|
+-- Unio_ravoisieri **
|
+-- Parreysiinae
|
+-- Pleurobemini
Sonowal et al., 2021 [Publ 4221], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
5 |
3 |
2 |
12 |
6 |
3 |
48.0 |
+-------------------------- Lamellidens_mainwaringi
--+
| +----------------------- Lamprotula
+--+
| +-------------------- Potomida
+--+
| +----------------- Lampsilini
+--+
| +-------------- Pleurobemini
+--+
| +----------- Aculamprotula **
+--+
| +-------- Unio_pictorum **
+--+
| +----- Anodontini **
+--+
| +-- Unio_ravoisieri **
+--+
+-- Lamellidens_corrianus
Lopes-Lima et al., 2020 [Publ 4080], fig. 9
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
1 |
2 |
0 |
0 |
48.0 |
-- (Cristariina
Araujo et al., 2017 [Publ 3801], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
2 |
10 |
9 |
3 |
47.0 |
+-------------- Parreysiinae
|
--+ +----------- Ambleminae
| |
+--+ +-------- Gonideinae
| |
+--+ +-- Schistodesmus **
| +--+
| | +-- Aculamprotula **
+--+
| +-- Anodontini **
+--+
+-- Unio **
Fonseca et al., 2016 [Publ 3775], fig. 1-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
2 |
5 |
5 |
2 |
43.0 |
+-------- Ambleminae
--+
| +----- Gonideinae
+--+
| +-- Unionini **
+--+
+-- Anodontini **
Wu et al., 2024 [Publ 4388], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
3 |
1 |
5 |
5 |
1 |
49.0 |
+-- Unionini **
--+
+-- Gonideinae
Klishko et al., 2016 [Publ 3733], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
1 |
0 |
0 |
0 |
46.0 |
-- (Cristariina
Soroka, 2010 [Publ 3404], fig. 9c
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
3 |
1 |
0 |
0 |
0 |
50.0 |
-- (Cristariina
Klishko et al., 2014 [Publ 3666], fig. 8
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
1 |
0 |
0 |
0 |
46.0 |
-- (Cristariina
Wu et al., 2021 [Publ 4215], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
2 |
20 |
20 |
2 |
48.0 |
+-------- Parreysiinae
--+
| +----- Gonideinae
+--+
| +-- Acuticostini **
+--+
+-- Unionini **
Amaro et al., 2017 [Publ 3791], fig. 2-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
1 |
1 |
6 |
6 |
2 |
47.0 |
+----- Ambleminae
--+
| +-- Gonideinae
+--+
+-- Anodontini **
Soroka, 2005 [Publ 2726], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
1 |
1 |
1 |
1 |
45.0 |
+-- Gonideinae
--+
+-- Unionini **
Sayenko et al., 2017 [Publ 3760], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
1 |
0 |
0 |
0 |
48.0 |
-- (Cristariina
Amaro et al., 2017 [Publ 3791], fig. 3B
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
1 |
8 |
5 |
2 |
43.0 |
+-------- Anodontini **
--+
| +----- Lampsilini
+--+
| +-- Quadrulini
+--+
+-- Gonideinae
Cyr et al., 2007 [Publ 2745], fig. 2A
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
1 |
1 |
1 |
1 |
49.0 |
+-- Ambleminae
--+
+-- Anodontini **
Chase et al., 2018 [Publ 3847], fig. 5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
2 |
6 |
4 |
2 |
43.0 |
+-------- Gonideinae
--+
| +----- Ambleminae
+--+
| +-- Anodontini **
+--+
+-- Unionini **
Froufe et al., 2017 [Publ 3802], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
1 |
0 |
0 |
0 |
43.0 |
-- (Anodontini
Graf, 2002 [Publ 2651], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
2 |
14 |
8 |
2 |
46.0 |
+----------- Pseudodontini
|
--+ +-------- Contradentini
| |
+--+ +-- Unionini **
| +--+
| | +-- Anodontini **
+--+
| +-- Gonideini
+--+
+-- Ambleminae
Nagel et al., 1998 [Publ 1946], fig. 8
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
1 |
1 |
0 |
0 |
0 |
45.0 |
-- (Unio
Whelan et al., 2011 [Publ 3380], fig. 1-ML
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
2 |
32 |
20 |
3 |
46.0 |
+-------------- Parreysiinae
|
| +-- Pseudodontini
--+ +--------+
| | +-- Contradentini
| |
+--+ +-- Anodontini **
| +-----+
| | +-- Unionini **
+--+
| +----- Gonideini
+--+
| +-- Lamprotulini
+--+
+-- Ambleminae
Mezhzherin et al., 2014 [Publ 3705], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
1 |
0 |
0 |
0 |
45.0 |
-- (Anodontini
Lopes-Lima et al., 2021 [Publ 4217], fig. 11
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
1 |
0 |
0 |
0 |
45.0 |
-- (Anodonta
Bulakhova et al., 2023 [Publ 4325], fig. 5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
1 |
0 |
0 |
0 |
49.0 |
-- (Beringiana
Whelan et al., 2011 [Publ 3380], fig. 1-MP
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
2 |
32 |
20 |
3 |
46.0 |
+----------------- Parreysiinae
|
--+ +-------------- Contradentini
| |
+--+ +----------- Pseudodontini
| |
+--+ +-- Anodontini **
| +-----+
| | +-- Unionini **
+--+
| +----- Gonideini
+--+
| +-- Lamprotulini
+--+
+-- Ambleminae
Lopes-Lima et al., 2017 [Publ 3792], fig. 2-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
4 |
4 |
2 |
2 |
6 |
5 |
2 |
43.0 |
+-------- Ambleminae
--+
| +----- Gonideinae
+--+
| +-- Unionini **
+--+
+-- Anodontini **
Lopes-Lima et al., 2016 [Publ 3757], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
1 |
0 |
0 |
0 |
44.0 |
-- (Anodontina
Walker et al., 2006 [Publ 2666], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
1 |
12 |
9 |
2 |
46.5 |
+----------- Pseudodontini
--+
| +-------- Gonidea
+--+
| +----- Inversidens
+--+
| +-- Anodontini **
+--+
+-- Ambleminae
Froufe et al., 2015 [Publ 3664], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
1 |
9 |
9 |
2 |
44.0 |
+-- Gonideini
+-----+
| +-- Lamprotulini
--+
| +----- Chamberlainiini
+--+
| +-- Ambleminae
+--+
+-- Anodontini **
Nagel et al., 1998 [Publ 1946], fig. 9
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
2 |
3 |
2 |
1 |
45.5 |
+-------- Lamprotulini
--+
| +----- Gonideini
+--+
| +-- Unionini **
+--+
+-- Anodontini **
Soroka, 2010 [Publ 3404], fig. 8b-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
2 |
0 |
0 |
0 |
44.0 |
+-- Anodontini **
--+
+-- Unionini **
Bolotov et al., 2016 [Publ 3743], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
2 |
27 |
1 |
1 |
46.0 |
+----- Ambleminae
--+
| +-- Anodontini **
+--+
+-- Unionini **
Pfeiffer & Graf, 2015 [Publ 3620], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
2 |
37 |
27 |
3 |
45.0 |
+-------- Parreysiinae
|
--+ +-- Unionini **
| +--+
| | +-- Anodontini **
+--+
| +-- Ambleminae
+--+
+-- Gonideinae
Walker et al., 2006 [Publ 2666], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
1 |
12 |
9 |
2 |
46.5 |
-- (Anodontini
Dai et al., 2024 [Publ 4389], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
1 |
44 |
42 |
1 |
44.0 |
+-- Anodontini **
--+
+-- Gonideinae
Soroka, 2010 [Publ 3404], fig. 8a-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
2 |
0 |
0 |
0 |
44.0 |
+-- Anodontini **
--+
+-- Unionini **
Soroka, 2008 [Publ 3695], fig. 1-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
1 |
1 |
1 |
1 |
44.0 |
+-- Ambleminae
--+
+-- Anodontini **
Kilikowska et al., 2020 [Publ 4335], fig. 4a
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
1 |
0 |
0 |
0 |
46.0 |
-- (Unio
Walker et al., 2006 [Publ 2666], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
1 |
12 |
9 |
2 |
46.5 |
+----------- Pseudodontini
--+
| +-------- Gonidea
+--+
| +----- Inversidens
+--+
| +-- Anodontini **
+--+
+-- Ambleminae
Soroka, 2008 [Publ 3695], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
1 |
1 |
1 |
1 |
44.0 |
+-- Ambleminae
--+
+-- Anodontini **
Hinzmann et al., 2013 [Publ 3541], fig. 5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
1 |
0 |
0 |
0 |
44.0 |
-- (Anodontina
Moyer & Díaz-Ferguson, 2012 [Publ 3375], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
1 |
3 |
3 |
1 |
47.0 |
+-- Anodontini **
--+
+-- Ambleminae
Smith, 2021 [Publ 4331], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
1 |
17 |
13 |
3 |
44.0 |
+-------- Ambleminae
--+
| +----- Gonideinae
+--+
| +-- Parreysiinae
+--+
+-- Anodontini **
Wen et al., 2017 [Publ 3795], fig. 3-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
1 |
3 |
3 |
1 |
44.0 |
+-- Anodontini **
--+
+-- Ambleminae
Bolotov et al., 2016 [Publ 3743], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
2 |
27 |
1 |
1 |
46.0 |
+----- Ambleminae
--+
| +-- Anodontini **
+--+
+-- Unionini **
Froufe et al., 2015 [Publ 3664], fig. 1-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
1 |
5 |
5 |
2 |
44.0 |
+----- Anodontini **
--+
| +-- Ambleminae
+--+
+-- Gonideinae
Gustafson & Iwamoto, 2005 [Publ 2741], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
1 |
2 |
1 |
1 |
47.0 |
+-- Gonideinae
--+
+-- Anodontini **
Krebs et al., 2019 [Publ 4092], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
1 |
0 |
0 |
0 |
48.0 |
-- (Pyganodon
Froufe et al., 2014 [Publ 3592], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
3 |
3 |
1 |
1 |
0 |
0 |
0 |
46.0 |
-- (Anodontina
Lopes-Lima et al., 2019 [Publ 3901], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
1 |
24 |
22 |
1 |
-- |
+-- Anodontini **
--+
+-- Ambleminae
Xia et al., 2016 [Publ 3712], fig. 7B
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
3 |
0 |
1 |
1 |
1 |
1 |
46.0 |
+----- Sinanodonta **
--+
| +-- Cristaria **
+--+
+-- Gonideinae
Huang et al., 2013 [Publ 3629], fig. 3-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
1 |
4 |
4 |
2 |
-- |
+----- Anodontini **
--+
| +-- Gonideinae
+--+
+-- Ambleminae
Sheng et al., 2014 [Publ 3704], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
8 |
0 |
1 |
6 |
6 |
2 |
51.0 |
+-------------- Cristariina **
--+
| +----------- Alasmidontina **
+--+
| +-------- Lampsilis
+--+
| +----- Gonideinae
+--+
| +-- Venustaconcha
+--+
+-- Quadrulini
Amaro et al., 2017 [Publ 3791], fig. 3A-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
1 |
5 |
4 |
2 |
-- |
+-------- Quadrulini
--+
| +----- Gonideinae
+--+
| +-- Lampsilini
+--+
+-- Anodontini **
Walker et al., 2006 [Publ 2666], fig. 6-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
1 |
12 |
8 |
2 |
-- |
+----- Gonideinae
--+
| +-- Anodontini **
+--+
+-- Ambleminae
Zieritz et al., 2024 [Publ 4381], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
44 |
42 |
3 |
-- |
+----------------- Ambleminae
|
| +-------- Parreysiinae
| |
| +-----+ +-- Anodontini **
--+ | | +--+
| | | | +-- Unionini **
| | +--+
| | | +-- Chamberlainia
| | +--+
+--+ +-- Discomya
|
| +----- Ctenodesmini
| +-----+
| | | +-- Rectidentini
| | +--+
| | +-- Contradentini
+--+
| +-- Schepmaniini
| +-----+
| | +-- Pseudodontini
+--+
| +----- Caudiculatus
+--+
| +-- Potomida
+--+
+-- Gonideini
Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
4 |
1 |
1 |
-- |
+----- Gonideinae
--+
| +-- Unionini **
+--+
+-- Anodontini **
Soroka, 2010 [Publ 3404], fig. 5a-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
1 |
0 |
0 |
0 |
-- |
-- (Anodontini
Lopes-Lima et al., 2019 [Publ 3901], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
1 |
26 |
24 |
1 |
-- |
+-- Anodontini **
--+
+-- Ambleminae
Graf, 2000 [Publ 2647], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
4 |
0 |
2 |
15 |
4 |
2 |
48.0 |
+-------- Unionini **
--+
| +----- Pleurobemini
+--+
| +-- Anodontini **
+--+
+-- Lampsilini
Gomes-dos-Santos et al., 2019 [Publ 3943], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
8 |
1 |
1 |
-- |
+----- Gonideinae
--+
| +-- Unionini **
+--+
+-- Anodontini **
Huang et al., 2018 [Publ 3828], fig. 3_5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
1 |
27 |
4 |
3 |
-- |
+-------- Gonideinae
--+
| +----- Unionini **
+--+
| +-- Ambleminae
+--+
+-- Parreysiinae
Mock et al., 2004 [Publ 2686], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
1 |
0 |
0 |
0 |
-- |
-- (Anodontini
Hoeh et al., 2002 [Publ 2673], fig. 1-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
1 |
7 |
6 |
1 |
-- |
+-- Anodontini **
--+
+-- Ambleminae
Hoeh et al., 2002 [Publ 2673], fig. 1-F
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
1 |
7 |
6 |
1 |
-- |
+-- Anodontini **
--+
+-- Ambleminae
Chen et al., 2012 [Publ 3624], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
1 |
5 |
5 |
2 |
-- |
+----- Ambleminae
--+
| +-- Anodontini **
+--+
+-- Gonideinae
Huff et al., 2004 [Publ 2697], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
9 |
1 |
1 |
-- |
+----- Ambleminae
--+
| +-- Unionini **
+--+
+-- Anodontini **
Zhang et al., 2013 [Publ 4214], fig. 5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
4 |
0 |
1 |
2 |
1 |
1 |
48.0 |
+-------- Cristaria **
--+
| +----- Sinanodonta **
+--+
| +-- Margaritiferinae
+--+
+-- Gonideinae
Lopes-Lima et al., 2019 [Publ 3901], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
1 |
27 |
25 |
1 |
-- |
+-- Anodontini **
--+
+-- Ambleminae
Wei et al., 2016 [Publ 3665], fig. 0
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
4 |
0 |
1 |
4 |
4 |
2 |
47.0 |
+----- Gonideinae
|
--+ +-- Cristariina **
| |
| +-- Lampsilini
+--+
+-- Alasmidontina **
|
+-- Quadrulini
Guerra et al., 2017 [Publ 3772], fig. 2b-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
1 |
6 |
4 |
2 |
-- |
+-- Gonideinae
|
--+-- Anodontini **
|
+-- Ambleminae
Amaro et al., 2017 [Publ 3791], fig. 3B-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
1 |
5 |
4 |
2 |
-- |
+-------- Quadrulini
--+
| +----- Lampsilini
+--+
| +-- Anodontini **
+--+
+-- Gonideinae
Guerra et al., 2017 [Publ 3772], fig. 2a-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
1 |
6 |
4 |
2 |
-- |
+-- Gonideinae
|
--+-- Anodontini **
|
+-- Ambleminae
Taylor et al., 2007 [Publ 2696], fig. 1-3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
2 |
0 |
0 |
-- |
+-- Anodontini **
--+
+-- Unionini **
Soroka, 2010 [Publ 3404], fig. 5b-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
1 |
0 |
0 |
0 |
-- |
-- (Anodontini
Krebs, 2004 [Publ 2692], fig. 2-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
1 |
2 |
2 |
1 |
-- |
+-- Anodontini **
--+
+-- Ambleminae
Hoeh et al., 1996 [Publ 2694], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
1 |
2 |
1 |
1 |
-- |
+-- None__1
+--+
| +-- None__2
--+
| +-- Ambleminae
+--+
+-- Anodontini **
Huff et al., 2004 [Publ 2697], fig. 5-6
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
3 |
0 |
2 |
10 |
1 |
1 |
47.5 |
+-- Unionini **
|
--+-- Anodontini **
|
+-- Ambleminae
Curole & Kocher, 2005 [Publ 2682], fig. 1-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
1 |
10 |
9 |
1 |
-- |
+-- Anodontini **
--+
+-- Ambleminae
Huff et al., 2004 [Publ 2697], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
10 |
1 |
1 |
-- |
+----- Ambleminae
--+
| +-- Unionini **
+--+
+-- Anodontini **
Soroka, 2010 [Publ 3404], fig. 4-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
1 |
0 |
0 |
0 |
-- |
-- (Anodontini
Graf & Ó Foighil, 2000 [Publ 2649], fig. 1_2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
9 |
2 |
1 |
-- |
+----- Ambleminae
--+
| +-- Unionini **
+--+
+-- Anodontini **
Mock et al., 2004 [Publ 2686], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
1 |
0 |
0 |
0 |
-- |
-- (Anodontini
Krebs, 2004 [Publ 2692], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
1 |
3 |
3 |
1 |
-- |
+-------- Quadrulini
--+
| +----- Anodontini **
+--+
| +-- Lampsilini
+--+
+-- Pleurobemini
Walker et al., 2006 [Publ 2666], fig. 6
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
1 |
34 |
8 |
2 |
-- |
-- (Pyganodon
Marshall et al., 2014 [Publ 3593], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
2 |
2 |
0 |
2 |
12 |
1 |
1 |
-- |
+-- Anodontini **
--+
+-- Unionini **
Combosch et al., 2017 [Publ 3806], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
14 |
2 |
2 |
-- |
+----- Unionini **
--+
| +-- Ambleminae
+--+
+-- Parreysiinae
Lydeard et al., 2000 [Publ 2654], fig. 2b
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
13 |
12 |
1 |
-- |
-- Anodonta_cygnea
González & Giribet, 2015 [Publ 3632], fig. 6
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
5 |
0 |
0 |
-- |
-- Unio_pictorum
Metzger et al., 2018 [Publ 3952], fig. 1C
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
2 |
2 |
1 |
-- |
+-- Anodontini **
--+
+-- Ambleminae
Zanatta & Harris, 2013 [Publ 3530], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
4 |
4 |
1 |
-- |
+-- Anodontini **
--+
+-- Ambleminae
Curole & Kocher, 2002 [Publ 2712], fig. 1a-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
3 |
3 |
1 |
-- |
+----- Quadrulini
--+
| +-- Anodontini **
+--+
+-- Lampsilini
Giribet & Distel, 2003 [Publ 3670], fig. 3f5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
5 |
2 |
1 |
-- |
+-- Pleurobemini
|
--+-- Lampsilini
|
+-- Unionini **
Inoue et al., 2020 [Publ 3941], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
41 |
41 |
1 |
-- |
+-- Anodontini **
--+
+-- Ambleminae
Hoeh et al., 1998 [Publ 2695], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
4 |
1 |
1 |
-- |
-- Anodonta_cygnea
Araujo et al., 2009 [Publ 2395], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
7 |
0 |
0 |
-- |
-- Unio
Bieler et al., 2014 [Publ 3622], fig. 26_27_29_30
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
4 |
0 |
0 |
-- |
-- Unio_pictorum
Zanatta & Murphy, 2006 [Publ 2672], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
55 |
54 |
1 |
-- |
+-- Anodontini **
--+
+-- Ambleminae
Combosch et al., 2017 [Publ 3806], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
14 |
2 |
2 |
-- |
+----- Unionini **
--+
| +-- Parreysiinae
+--+
+-- Ambleminae
Peng et al., 2012 [Publ 3689], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
2 |
2 |
1 |
-- |
+-- Anodontini **
--+
+-- Gonideinae
Mulvey et al., 1997 [Publ 3390], fig. 1_3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
4 |
4 |
1 |
-- |
+-- Anodontini **
--+
+-- Ambleminae
Lydeard et al., 2000 [Publ 2654], fig. 2d
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
13 |
12 |
1 |
-- |
-- Anodonta_cygnea
Curole & Kocher, 2002 [Publ 2712], fig. 1b-M
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
3 |
3 |
1 |
-- |
+----- Lampsilini
--+
| +-- Anodontini **
+--+
+-- Quadrulini
Inoue et al., 2013 [Publ 3579], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
8 |
0 |
0 |
-- |
-- Pyganodon_grandis
Lydeard et al., 2000 [Publ 2654], fig. 2c
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
13 |
12 |
1 |
-- |
-- Anodonta_cygnea
González & Giribet, 2015 [Publ 3632], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
5 |
0 |
0 |
-- |
-- Unio_pictorum
Wang et al., 2016 [Publ 3714], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
2 |
2 |
1 |
-- |
+-- Unionini **
--+
+-- Gonideinae
Lopes-Lima et al., 2018 [Publ 3829], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
27 |
2 |
2 |
-- |
+----- Gonideinae
--+
| +-- Ambleminae
+--+
+-- Unionini **
Zanatta & Murphy, 2006 [Publ 2672], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
56 |
55 |
1 |
-- |
-- Pyganodon_grandis
Graf et al., 2015 [Publ 3619], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
19 |
0 |
0 |
-- |
-- Unio_pictorum
Lydeard et al., 2000 [Publ 2654], fig. 2a
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
13 |
12 |
1 |
-- |
-- Anodonta_cygnea
Inoue et al., 2013 [Publ 3528], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
16 |
16 |
2 |
-- |
+-- Gonideinae
|
--+-- Anodontini **
|
+-- Ambleminae
Bolotov et al., 2015 [Publ 3646], fig. 2B
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
8 |
0 |
0 |
-- |
-- Unio_pictorum
Teiga-Teixeira et al., 2020 [Publ 4173], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
18 |
16 |
2 |
-- |
+----- Ambleminae
--+
| +-- Lanceolariini **
+--+
+-- Gonideinae
Campbell, 2000 [Publ 2724], fig. 1-5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
2 |
1 |
1 |
-- |
+-- Anodontini **
--+
+-- Ambleminae
González & Giribet, 2015 [Publ 3632], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
5 |
0 |
0 |
-- |
-- Unio_pictorum
Hewitt et al., 2019 [Publ 3962], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
9 |
8 |
1 |
-- |
+-- Anodontini **
--+
+-- Ambleminae
Bieler et al., 2014 [Publ 3622], fig. 28
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
4 |
0 |
0 |
-- |
-- Unio_pictorum
Jiang et al., 2010 [Publ 3438], fig. 5
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
3 |
3 |
2 |
-- |
+-- Ambleminae
|
--+-- Anodontini **
|
+-- Gonideinae
Inoue et al., 2020 [Publ 3941], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
40 |
40 |
1 |
-- |
+-- Anodontini **
--+
+-- Ambleminae
Hoeh et al., 1998 [Publ 2695], fig. 4
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
4 |
1 |
1 |
-- |
-- Anodonta_cygnea
Plazzi & Passamonti, 2010 [Publ 3354], fig. 2-3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
3 |
3 |
2 |
-- |
+-- Anodontini **
|
--+-- Ambleminae
|
+-- Gonideinae
Vikhrev et al., 2018 [Publ 3812], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
12 |
1 |
1 |
-- |
+-- Unionini **
--+
+-- Ambleminae
Graf et al., 2014 [Publ 3591], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
10 |
9 |
1 |
-- |
-- Unio_pictorum
Bolotov et al., 2015 [Publ 3646], fig. 2A
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
8 |
0 |
0 |
-- |
-- Unio_pictorum
González & Giribet, 2015 [Publ 3632], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
5 |
0 |
0 |
-- |
-- Unio_pictorum
Curole & Kocher, 2005 [Publ 2682], fig. 1
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
10 |
9 |
1 |
-- |
-- Pyganodon_grandis
Huff et al., 2004 [Publ 2697], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
7 |
0 |
0 |
-- |
-- Anodonta
Pfeiffer et al., 2016 [Publ 3660], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
37 |
37 |
1 |
-- |
+-- Anodontini **
--+
+-- Ambleminae
Pieri et al., 2018 [Publ 3867], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
19 |
19 |
1 |
-- |
+-- Anodontini **
--+
+-- Ambleminae
Turner et al., 2000 [Publ 2722], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
13 |
12 |
1 |
-- |
+-- Quadrulini
+-----+
| +-- Anodontini **
--+
| +----- Lampsilini
+--+
| +-- Amblemini
+--+
+-- Pleurobemini
Lopes-Lima et al., 2018 [Publ 3829], fig. 3
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
27 |
2 |
2 |
-- |
+----- Gonideinae
--+
| +-- Ambleminae
+--+
+-- Unionini **
Klunzinger et al., 2021 [Publ 4086], fig. 2
iT |
tT |
bi |
riS |
oT |
roT |
roS |
iD |
1 |
1 |
0 |
1 |
5 |
0 |
0 |
-- |
-- Unio_pictorum
|